treatments-xml/data/21/9A/93/219A9352B86056889DBEB0947CC84DEA.xml

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<mods:title id="8C4F0DEFE576778BF6BDDDCD420A0E4F">A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group</mods:title>
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<mods:namePart id="DF607D222F5B7FBBB8D3D84465CC94CA">Lantsov, Vladimir I.</mods:namePart>
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<mods:affiliation id="3FE40A84AEB044D17B806FC89436C83B">Tembotov Institute of Ecology of Mountain Territories of Russian Academy of Sciences, I. Armand str., 37 a, Nalchik 360051, Russia</mods:affiliation>
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<mods:namePart id="3EE898C0A5B87D2987A955AA97F716D3">Pilipenko, Valentin E.</mods:namePart>
<mods:affiliation id="E002BDAE1B3115DB1C4E89622CEA437A">Lomonosov Moscow State University, GSP- 1, Leninskie Gory, Moscow 119991, Russia</mods:affiliation>
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<treatment id="219A9352B86056889DBEB0947CC84DEA" ID-GBIF-Taxon="183543016" LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA" httpUri="http://treatment.plazi.org/id/219A9352B86056889DBEB0947CC84DEA" lastPageNumber="145" pageId="0" pageNumber="145" scope_family="Tipulidae" scope_order="Diptera">
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<paragraph id="2D2361AC4AC602BF08CC6588E37FEEC4" pageId="0" pageNumber="145">
<taxonomicName id="F64CAF44DB56F1091DB992149B951A4B" ID-CoL="5757F" LSID="219A9352-B860-5688-9DBE-B0947CC84DEA" authority="Riedel, 1920" authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Lunatipula) caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica" subGenus="Lunatipula">Tipula (Lunatipula) caucasica Riedel, 1920</taxonomicName>
</paragraph>
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<paragraph id="119CA4D493E25E13A9FE2994F6C07F5F" pageId="0" pageNumber="145">
<figureCitation id="D1AEE56CBDA76565EB487F5CA859F6CB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">Figs 1</figureCitation>
<figureCitation id="1E72E6E0E97DBC78D0654686D95E806A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">, 2</figureCitation>
<figureCitation id="EFA4B421A52B3E40E9C9E416B14FFC41" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">, 12A-C</figureCitation>
<figureCitation id="689C96221C142D6038FD02928FD2A0DE" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">, 13A</figureCitation>
<figureCitation id="0C06CB26A93A64C7C6BA7980CE4D8B3C" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">, 14A,F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C8A472D2913B66C9543E1EEBC8253304" pageId="0" pageNumber="145" type="reference_group">
<paragraph id="16EE4517CBB6D389988BDC8DC309277B" pageId="0" pageNumber="145">
<taxonomicName id="25429DE3378DA263FF60F83917F55F63" authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Lunatipula) caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica" subGenus="Lunatipula">Tipula (Lunatipula) caucasica</taxonomicName>
<bibRefCitation id="6CBC262F868BC24BCC3AB65FDEA4288D" author="Riedel, MP" journalOrPublisher="Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere" pageId="0" pageNumber="145" pagination="13 - 22" refId="B9" refString="Riedel, MP, 1920. Nematocera Polyneura (Dipt.) aus dem Kaukasus. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 43: 13 - 22" title="Nematocera Polyneura (Dipt.) aus dem Kaukasus." volume="43" year="1920">Riedel 1920</bibRefCitation>
: 17 (type locality: Georgia, 
<normalizedToken id="7FAD04519F513450898C8FE297F739B7" originalValue="Mts’chet">Mts'chet</normalizedToken>
, prope Tiflis); 
<bibRefCitation id="4CE4D93F90437F4E23BFFF29CECAB291" author="Savchenko, EN" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="0" pageNumber="145" refId="B12" refString="Savchenko, EN, 1964. Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N.S. 89, Nasekomye Dvukrylye [Diptera] 2(4): 1-503. [in Russian]" title="Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N. S. 89, Nasekomye Dvukrylye [Diptera] 2 (4): 1 - 503. [in Russian]" year="1964">Savchenko 1964</bibRefCitation>
: 390; Oosterbroek, Theowald 1992:104; 
<bibRefCitation id="2D9A557A94D89137B71C659365D1CCCF" DOI="https://doi.org/10.47640/978-5-00105-586-0_2020_292" author="Lantsov, VI" journalOrPublisher="Fauna of the Stavropol region" pageId="0" pageNumber="145" refId="B7" refString="Lantsov, VI, 2020. Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24-29, 2020. Proceedings of conference / Responsible editors: O.G.Ovchinnikova, I.V.Shamshev. St.Petersburg: Russian Entomological Society, LEMA, 292-296. https://doi.org/10.47640/978-5-00105-586-0_2020_292" title="Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24 - 29, 2020. Proceedings of conference / Responsible editors: O. G. Ovchinnikova, I. V. Shamshev. St. Petersburg: Russian Entomological Society, LEMA, 292 - 296." url="https://doi.org/10.47640/978-5-00105-586-0_2020_292" year="2020">Lantsov 2020</bibRefCitation>
: 294; 
<bibRefCitation id="C00EE5EA3D3DAAF0461D3F0F54A1E7A1" author="Oosterbroek, P" journalOrPublisher="Fauna of the Stavropol region" pageId="0" pageNumber="145" publicationUrl="http://ccw.naturalis.nl/detail.php" refId="B8" refString="Oosterbroek, P, 2021. Catalogue of the Craneflies of the World. http://ccw.naturalis.nl/detail.php" title="Catalogue of the Craneflies of the World." url="http://ccw.naturalis.nl/detail.php" year="2021">Oosterbroek 2021</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="32EDE59A3AE9BC200BD7425EC70796EE" pageId="0" pageNumber="145" type="materials_examined">
<paragraph id="CA34F9A7F639F8658753919985C275E3" pageId="0" pageNumber="145">Material examined.</paragraph>
<paragraph id="419CACA693D6EB9BAF3DC83C54BFE5FE" pageId="0" pageNumber="145">
<materialsCitation id="1BAF8A84C0FB74ADD2F19ABCC91BBFCB" collectingDate="1958-05-23" collectingDateMax="1959-07-17" collectingDateMin="1958-05-23" collectorName="Kurnakov, Savenko" country="Georgia" location="Bani-Khevi" specimenCount="10" specimenCount-female="4" specimenCount-male="6">
<collectingCountry id="F04BD16626FE54781A145652155A9299" name="Georgia">Georgia</collectingCountry>
• 
<specimenCount id="798BBC960642D8A2C81A5BE5C6D0A57D" type="male">1 male</specimenCount>
, 
<specimenCount id="F56FB5673D73261699EA1AF7D3965700" type="female">3 females</specimenCount>
; 
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. 
<normalizedToken id="46DDE4ED08B5DB7959C348D8819EB81D" originalValue="Бани-Хеви">
<location id="01AD7850430DC2BAF6BC3CC4422828FD" LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:01AD7850430DC2BAF6BC3CC4422828FD" country="Georgia" name="Bani-Khevi">Bani-Khevi</location>
</normalizedToken>
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. 
<normalizedToken id="E26E24ABCD60385FB7CE1BC6136FFCEB" originalValue="Боржоми">Borzhomi</normalizedToken>
, 
<normalizedToken id="1360E41C1ED44BD324CA66C21D17E76A" originalValue="Аджаро-Имеретинск">Adzharo-Imeretinsk</normalizedToken>
. 
<normalizedToken id="C0372532CD7CA536A38CE3353E66C330" originalValue="хр">khr</normalizedToken>
. [Gorge Bani-Khevi near Borjomi, Adjaro-Imeretinsk. Ridge]; 
<collectingDate id="E81D38B0ECAEBF203ECF636E63967E89" value="1958-06-26">26 Jun. 1958</collectingDate>
; 
<collectorName id="0D08528761F7E3CE7C568536ECF25007">Kurnakov</collectorName>
leg.; ZISP • 
<specimenCount id="9A0C27C2FF34E9F974C98E38F45BE59B" type="male">1 male</specimenCount>
; 
<normalizedToken id="8E57D10062CE922AA5D8A172FDDE4222" originalValue="“Цхра-Цхаро">&quot;Tskhra-Tskharo</normalizedToken>
<normalizedToken id="58A72FF623A79DAC4A729271F301EF87" originalValue="близ">bliz</normalizedToken>
<normalizedToken id="936DB376F1FED08485A5F0F8DC010F01" originalValue="Бакуриани”">Bakuriani&quot;</normalizedToken>
[Tskhra-Tskharo near Bakuriani]; 
<collectingDate id="AE16448AA56C7053F3886A7F81EB5038" value="1958-06-29">29 Jun. 1958</collectingDate>
; 
<collectorName id="29643A7E87171E3E307EBFF346281C6F">Kurnakov</collectorName>
leg.; ZISP • 
<specimenCount id="FA70121FE531F6589E5BC404F92A56B5" type="male">1 male</specimenCount>
; 
<normalizedToken id="0F7028E5BB4893D8F37418DADF3D1DFA" originalValue="“Леберде">&quot;Leberde</normalizedToken>
, 
<normalizedToken id="4EB6D5E0F1B9DD4109F44DC1E7D2895E" originalValue="Менгрелия">Mengreliya</normalizedToken>
, 
<normalizedToken id="C78B7A9C62E82AE4F08F040EE4D0BCE5" originalValue="Грузия”">Gruziya&quot;</normalizedToken>
[Leberde, Mengrelia, 
<collectingCountry id="604B61943CBB6E70D89A85D448645196" name="Georgia">Georgia</collectingCountry>
]; 
<collectingDate id="48354CECEFA88482D758A4DEED963320" value="1959-07-17">17 Jul. 1959</collectingDate>
; 
<collectorName id="CA25EC5AA398F94C2C0D2871610C5973">Savenko</collectorName>
leg.; ZISP • 
<specimenCount id="898BE3BB558B8252B49ADE57E00A27B5" type="male">1 male</specimenCount>
; 
<normalizedToken id="A4A81C9ADA6F5D5A574137AA52EC6C3A" originalValue="“Аджаро-Имеретинск">&quot;Adzharo-Imeretinsk</normalizedToken>
. 
<normalizedToken id="CBA6C93D638CAC6D46F63DE7E0B8D9C5" originalValue="хр">khr</normalizedToken>
. 
<normalizedToken id="351934EFA8EECAFD0AA748CA2DBAE9EA" originalValue="Горы">Gory</normalizedToken>
<normalizedToken id="EBF638CFE617CD8EFEE5834D9CF704FC" originalValue="у">u</normalizedToken>
<normalizedToken id="5C27B62C1A208E504BDE478CF3BA253A" originalValue="Саирме">Sairme</normalizedToken>
, 
<normalizedToken id="F26E5438AFAA18A74C1770D805ACF18F" originalValue="Грузия”">Gruziya&quot;</normalizedToken>
[Adjaro-Imeretinsk. Ridge, mountains near Sairme, 
<collectingCountry id="EEEC8040C30641D24EE8CD883964F5FE" name="Georgia">Georgia</collectingCountry>
]; 
<collectingDate id="04E728CCAC76E394491C975F855A032E" value="1958-05-23">23 May 1958</collectingDate>
; Kurnakov; ZISP • 
<specimenCount id="7E5F7E9E1C34F6F3388386D175852AF1" type="male">2 males</specimenCount>
, 
<specimenCount id="E630C49607729ED3396A158519A13CCF" type="female">1 female</specimenCount>
; 
<normalizedToken id="71FAE1BE9A541D78A8653520FF9770FC" originalValue="”ущ">&quot;ushch</normalizedToken>
. 
<normalizedToken id="CE53E1C7FE0D90D6FFDE4248FA5BB5CD" originalValue="Цваниат-Хеви">Tsvaniat-Khevi</normalizedToken>
<normalizedToken id="CA72EF6BD10458C348CE15395FF2DB12" originalValue="Аджаро-Имеретинск">Adzharo-Imeretinsk</normalizedToken>
. 
<normalizedToken id="183D32548D1C6421F5E6A1D75485D1F6" originalValue="хр">khr</normalizedToken>
</materialsCitation>
. 
<materialsCitation id="D1573BD3A67B32CEEE32EC107D9BFCF1" collectingDate="1958-05-20" collectingDateMax="1958-06-30" collectingDateMin="1958-05-20" collectorName="Kurnakov" country="Georgia" location="Oak-hornbeam forest" municipality="Gudautsk. District" specimenCount="4" specimenCount-male="4" stateProvince="Abkhazia">
<normalizedToken id="35B21298F224981251316C28C5FCC7EA" originalValue="Грузия”">Gruziya&quot;</normalizedToken>
[Tsvaniat-Khevi Gorge, Adjaro-Imeretinsk. Ridge, 
<collectingCountry id="3BE638E5A167D72305B4FC525D1EE2E0" name="Georgia">Georgia</collectingCountry>
]; 
<collectingDate id="E65F525167B86AA5AC6B55E7205D5B3E" value="1958-06-23">23 Jun. 1958</collectingDate>
; 
<collectorName id="B891E0C50EAB77FB249C1A8315E90245">Kurnakov</collectorName>
leg; / 
<normalizedToken id="CF77B7F60E35C1586C912454FC7F61CB" originalValue="“Высокотравье">&quot;Vysokotrave</normalizedToken>
<normalizedToken id="029BA5A93DF51E767DBBB04686D3A369" originalValue="у">u</normalizedToken>
<normalizedToken id="71C5F1489AAB26A97A024F1C25921E1D" originalValue="верхн">verkhn</normalizedToken>
. 
<normalizedToken id="C41A28745857E8F94C692BB1E89DEC54" originalValue="границы">granitsy</normalizedToken>
<normalizedToken id="CE8BF647EA41BDEA0EC344E00A551345" originalValue="леса">lesa</normalizedToken>
<normalizedToken id="5A68FEC560A90DA444E30B60346B1F24" originalValue="у">u</normalizedToken>
<normalizedToken id="365B33C156A09B8F37177644C25A3199" originalValue="ручья”">ruchya&quot;</normalizedToken>
[High grass at the edge of the border of the forest by the stream]; ZISP • 
<specimenCount id="ABA1693EEE1BDE4233FB4E83890EC99D" type="male">2 males</specimenCount>
; 
<normalizedToken id="01291BF8F31E1FB42F7D988B45E24550" originalValue="“с">&quot;s</normalizedToken>
. 
<normalizedToken id="E7E72258F4DCF5648BE9F34D0A0F1CD1" originalValue="Отхара">Otkhara</normalizedToken>
, 
<normalizedToken id="637008A87CA44E044C5DD96491A32E11" originalValue="Гудаутск">Gudautsk</normalizedToken>
. 
<normalizedToken id="DB56061A4DF5A5E904633AAA15F0DF5D" originalValue="р-н">r-n</normalizedToken>
, 
<normalizedToken id="F6A0EF6E8A4B30ADDBBD1627DB7B9590" originalValue="Абхазия">Abkhaziya</normalizedToken>
, 
<normalizedToken id="4CFA03AF7F5E916E992BA3C9C9F466B7" originalValue="бер">ber</normalizedToken>
. 
<normalizedToken id="1D3E9DBAFD297AD5885DFF7C1555BF92" originalValue="речки">rechki</normalizedToken>
, 
<normalizedToken id="A8721815CC8AC1DF6A3AC0C5EE42EDA0" originalValue="кусты.”">kusty.&quot;</normalizedToken>
[village Otkhara, 
<collectingMunicipality id="FCA5BA899F118CED8DD5E1D21DD4034D">Gudautsk. District</collectingMunicipality>
, 
<collectingRegion id="4C9008EB4A728104DE7BEA78AFDD5B9E" country="Georgia" name="Abkhazia">Abkhazia</collectingRegion>
, river bank, bushes]; 
<collectingDate id="BFEED71737436D10C7CB93813CF86B16" value="1958-06-30">30 Jun. 1958</collectingDate>
; Kurnakov; ZISP • 
<specimenCount id="93CC10ACEA97C90ED6307B33F29BFFB9" type="male">2 males</specimenCount>
; 
<normalizedToken id="D29FA5BD4BCC05BB8510732E1FF04A90" originalValue="“Абхазия">&quot;Abkhaziya</normalizedToken>
, 
<normalizedToken id="B527BFF9EA794E44586555283A2EB958" originalValue="с">s</normalizedToken>
. 
<normalizedToken id="E0770E623A2BE9E96E711C37665C4047" originalValue="Отхара">Otkhara</normalizedToken>
<normalizedToken id="25345B998E5162F74380BE7F4E6DCEC7" originalValue="предгорья">predgorya</normalizedToken>
<normalizedToken id="DF523E66B1199BAC41BD6B1B8D34A8BC" originalValue="Бзыбского">Bzybskogo</normalizedToken>
<normalizedToken id="E7AB9C4307EFFF875E2AEBC0F79E849A" originalValue="хр.”">khr.&quot;</normalizedToken>
[
<collectingRegion id="0C54A414E2372460484C09E40C54AB0C" country="Georgia" name="Abkhazia">Abkhazia</collectingRegion>
, village Otkhara foothills of the Bzybsk Ridge]; 
<collectingDate id="62AC0870951EE3E0B6EC9202FB1341F4" value="1958-05-20">20 May 1958</collectingDate>
; Kurnakov; / 
<normalizedToken id="82F40F7A9E423FADF9CAA3A0AEA524B4" originalValue="“Дубово-грабовый">&quot;Dubovo-grabovyi</normalizedToken>
<normalizedToken id="DF478865F1873F5C4836408E520B1755" originalValue="лес">les</normalizedToken>
, 
<normalizedToken id="F38E367D05A5ACCFA32BFCD2DC34549E" originalValue="кусты">kusty</normalizedToken>
Asalia&quot; [
<location id="61555C28CE56949C9D2B78972D9F174E" LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:61555C28CE56949C9D2B78972D9F174E" country="Georgia" municipality="Gudautsk. District" name="Oak-hornbeam forest" stateProvince="Abkhazia">Oak-hornbeam forest</location>
, bushes Asalia&quot;]; ZISP
</materialsCitation>
; 
<materialsCitation id="F103641D40EE8FE174AD4F2D8660F9A7" collectingDate="1964-06-15" collectorName="Pastukhov" country="Azerbaijan" elevation="200" location="Akhkemal Ridge. Azerb." municipality="Belokany" specimenCount="1" specimenCount-male="1">
<collectingCountry id="53C35C6F0D02A7D20B1BBC0BF85495D6" name="Azerbaijan">Azerbaijan</collectingCountry>
• 
<specimenCount id="74CF776C41C8C22D7FB56DCBD15294FE" type="male">1 male</specimenCount>
; 
<normalizedToken id="09E404D429E1082B00F324D0218C30CC" originalValue="“Белоканы">&quot;Belokany</normalizedToken>
, 
<normalizedToken id="DC720EA30E53687ED7A31642716FB907" originalValue="хр">khr</normalizedToken>
. 
<normalizedToken id="0229703FE767DA72903AA98CA52BA1B2" originalValue="Ахкемал">Akhkemal</normalizedToken>
. 
<normalizedToken id="971006906DE74D4121A5A9CD65881D54" originalValue="Азерб">Azerb</normalizedToken>
. 
<quantity id="28B4D253A5C7614D2F0DED95AB5CB482" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation id="003E9EF02AB03CC0E0B73AD4499E849C" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
200 
<normalizedToken id="7630B38CF4C10EBBA36AC9A77781C99F" originalValue="м”">m&quot;</normalizedToken>
</elevation>
</quantity>
[
<collectingMunicipality id="9870D9FECD2C11C0C3971856F57D9F3E">Belokany</collectingMunicipality>
, 
<location id="F13E52C84FF7BA3AFF9E97401FDB0E61" LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:F13E52C84FF7BA3AFF9E97401FDB0E61" country="Azerbaijan" municipality="Belokany" name="Akhkemal Ridge. Azerb.">Akhkemal Ridge. Azerb.</location>
[aidjan] alt. 
<quantity id="8D90E6086CFEF3C790C0E6E064E2C83E" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation id="6ADEAF27F920AFF60DEB53046C409CD3" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
]; 
<collectingDate id="C67A6F14CBE8FCBFCE36F25F36310550" value="1964-06-15">15 Jun. 1964</collectingDate>
; 
<collectorName id="1CFE1508DF5798DD3A6E7AECF8678A95">Pastukhov</collectorName>
leg.; ZISP
</materialsCitation>
; 
<materialsCitation id="01B46CDDDA165BCC3D4FB87E7AD98BF0" collectingDate="1932-05-06" collectingDateMax="2018-05-10" collectingDateMin="1932-05-06" collectorName="B. Rohdendorf, V. Lantsov, V. Pilipenko" country="Russia" county="Krasnodarskiy" determinerName="Ried" elevation="54" location="Pontica" municipality="Sotshi prov." specimenCount="37" specimenCount-female="16" specimenCount-male="21">
<collectingCountry id="F9A46D03339107AD7A9EC95F23F31735" name="Russia">Russia</collectingCountry>
- 
<emphasis id="38160711BB6EBE7E13246A9828BE774A" bold="true" pageId="0" pageNumber="145">
<collectingCounty id="94195779E5BC35B016A425729B8C8104">Krasnodarskiy</collectingCounty>
kray
</emphasis>
• 
<specimenCount id="33617DC2614D50D10487AE93FBB63994" type="male">1 male</specimenCount>
; 
<collectingMunicipality id="6E4829BE2899108359F820FA9E747E59">Sotshi prov.</collectingMunicipality>
<location id="FD55ACAB2034F90A598D387613A1A4C0" LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:FD55ACAB2034F90A598D387613A1A4C0" country="Russia" county="Krasnodarskiy" municipality="Sotshi prov." name="Pontica">Pontica</location>
; 
<collectingDate id="DEE0BB13B87D10A57330B00C437B16BB" value="1932-05-06">6 May 1932</collectingDate>
; 
<collectorName id="B9E17A07C21F94D6FEC114D015BCEA2D">B.Rohdendorf</collectorName>
leg.; / &quot; 
<taxonomicName id="8CDABDBEDE172E357B04A5362808DB03" genus="T." lsidName="T." pageId="0" pageNumber="145" rank="genus">T.</taxonomicName>
<taxonomicName id="6AF660BCA6436DAF18582AC0516F4399" authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica">Tipula caucasica</taxonomicName>
<determinerName id="B068B0EEEB3041F4FA4223451B41BEB6">Ried</determinerName>
det. Lacksch.&quot; [terminalia in 
<collectingCountry id="637C76B9E836AB1203D4640A8647B5C1" name="Canada">Canada</collectingCountry>
balsam pinned with specimen]; ZISP • 
<specimenCount id="6752757B9CE17DB46DC2ABBF190F2D9B" type="female">1 female</specimenCount>
, Khosta, 
<location id="6D6A33030C8E654AED854694996317AC" LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:6D6A33030C8E654AED854694996317AC" country="Russia" county="Krasnodarskiy" municipality="Sotshi prov." name="Caucasian Reserve">Caucasian Reserve</location>
, Tiso-samshitovaya rosha [Yew-and-Boxwood Tree Grove]; 43°31'656&quot;N, 39°52'467&quot;E; alt. 
<quantity id="2F1E19DE08ED6979174377CDA90871CB" metricMagnitude="1" metricUnit="m" metricValue="5.4" unit="m" value="54.0">
<elevation id="9F04F6F0D0018CB2215EC1916120BC70" metricMagnitude="1" metricUnit="m" metricValue="5.4" unit="m" value="54.0">54 m</elevation>
</quantity>
; collected at light, 
<collectingDate id="954205E6186909D51B6FC2F95057DF0A" value="2018-05-08">8 May 2018</collectingDate>
; 
<collectorName id="98D7A7791675A68A9916B814F0050F88">V. Lantsov</collectorName>
leg.; IEMT • 
<specimenCount id="E11FFEBAD6ADDAA732EB45FA35BDF24D" type="male">2 males</specimenCount>
, 
<specimenCount id="ACF1157F0AD8CDAAAC377D0C7FFEBA8E" type="female">2 females</specimenCount>
; same collection data as for preceding, 43°31'688&quot;N, 39°52'561&quot;E; alt. 
<quantity id="C3EF91E8CF3FF46F50349E7F5B1E447B" metricMagnitude="1" metricUnit="m" metricValue="3.5" unit="m" value="35.0">
<elevation id="4AAC535616BB467C309813417938AC66" metricMagnitude="1" metricUnit="m" metricValue="3.5" unit="m" value="35.0">35 m</elevation>
</quantity>
; 
<collectingDate id="3474C1EB58B9E7F05A26FD089E5FD033" value="2018-05-08">8 May 2018</collectingDate>
; 
<collectorName id="1989613D25468D4BF74ACBA6DC7BCD3C">V. Lantsov</collectorName>
leg. / Lime-beech (
<taxonomicName id="77E4D81F35C0709348BD31DB4232A363" class="Dicotyledoneae" family="Tiliaceae" genus="Tilia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Tilia begoniifolia" order="Malvales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="begoniifolia">
<emphasis id="37B6455B2668792BD45C97672B787743" italics="true" pageId="0" pageNumber="145">Tilia begoniifolia</emphasis>
</taxonomicName>
and 
<taxonomicName id="739B19C1B4507F0CCB6BEB9E7F4A4304" authorityName="Lipsky" authorityYear="1898" class="Dicotyledoneae" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fagus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis id="760FCD16DFD6A8CDE17836716824FBE7" italics="true" pageId="0" pageNumber="145">Fagus orientalis</emphasis>
</taxonomicName>
) forest with yew (
<taxonomicName id="2307605B083C6EB578EE6CDCD18590DD" class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Taxus baccata" order="Taxales" pageId="0" pageNumber="145" phylum="Coniferophyta" rank="species" species="baccata">
<emphasis id="10408A390F728DFD9FFC98A862E45370" italics="true" pageId="0" pageNumber="145">Taxus baccata</emphasis>
</taxonomicName>
) and hornbeam (
<taxonomicName id="BC0A92116955A527D1A9AEA623D45D41" class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus betulus" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="betulus">
<emphasis id="DE9679F5516987D5FDB12EB1935A36A2" italics="true" pageId="0" pageNumber="145">Carpinus betulus</emphasis>
</taxonomicName>
) in tree layer and 
<normalizedToken id="89B484DCAF56415174F25F10D102B811" originalValue="butcher’s-broom">butcher's-broom</normalizedToken>
(
<taxonomicName id="75F4886D5E2DBCFFBF871FD192D4F3ED" authorityName="Yeo" authorityYear="1966" class="Monocotyledoneae" family="Liliaceae" genus="Ruscus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ruscus colchicus" order="Liliiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="colchicus">
<emphasis id="F87A7D96B8A825878B14B1C59F078098" italics="true" pageId="0" pageNumber="145">Ruscus colchicus</emphasis>
</taxonomicName>
- dominant and 
<taxonomicName id="06624896C8CCD9A9BF01E6D452A1A271" class="Monocotyledoneae" family="Liliaceae" genus="Ruscus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ruscus aculeatus" order="Liliiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="aculeatus">
<emphasis id="3F12ED80FF4932EACE11AC50DEDD39ED" italics="true" pageId="0" pageNumber="145">Ruscus aculeatus</emphasis>
</taxonomicName>
) in ground layer [one of the females was collected in the same community in a damp place near a yew log]; IEMT • 
<specimenCount id="99B9AEE5CABDD8174C8C511B4E9B1208" type="male">2 males</specimenCount>
, 
<specimenCount id="BD386A4A107159AD883F2679736C2D4A" type="female">2 females</specimenCount>
, same collection data as for preceding, 43°31'780&quot;N, 39°52'518&quot;E; alt. 
<quantity id="D368D8309251E5416D2A5D627D0CAC72" metricMagnitude="1" metricUnit="m" metricValue="9.8" unit="m" value="98.0">
<elevation id="D559A8BA41BAF6FD014CFBB7F3E3FF2B" metricMagnitude="1" metricUnit="m" metricValue="9.8" unit="m" value="98.0">98 m</elevation>
</quantity>
; 
<collectingDate id="6BB61FC4DD3E6027228DB921E1D0328F" value="2018-05-09">9 May 2018</collectingDate>
; 
<collectorName id="00785A4EBBFBB3BE4B28A5A2E37332F3">V. Lantsov</collectorName>
leg.; / Hornbeam - ash (
<taxonomicName id="45D480AB149D54758BB473550EAA91F1" authorityName="Mill" authorityYear="1768" class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis id="3720E0883461A82DB11D189BE5A4BB99" italics="true" pageId="0" pageNumber="145">Carpinus orientalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="09569D467EBC74653CAB6C7C28082E01" class="Dicotyledoneae" family="Oleaceae" genus="Fraxinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fraxinus excelsior" order="Oleales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="excelsior">
<emphasis id="150D6954E3F28CF4D6736466246797D9" italics="true" pageId="0" pageNumber="145">Fraxinus excelsior</emphasis>
</taxonomicName>
) forest with addition of yew (
<taxonomicName id="C50F8E22D55EAE95FF6AA2C89175776C" class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Taxus baccata" order="Taxales" pageId="0" pageNumber="145" phylum="Coniferophyta" rank="species" species="baccata">
<emphasis id="37FEAE5322127E94732CB4A2966812B4" italics="true" pageId="0" pageNumber="145">Taxus baccata</emphasis>
</taxonomicName>
) and lime (
<taxonomicName id="B8E1936EB24B9E7E8EDADE7EE51A1A47" class="Dicotyledoneae" family="Tiliaceae" genus="Tilia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Tilia begoniifolia" order="Malvales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="begoniifolia">
<emphasis id="A39BF965518E0D9C9C0644C39BEA0528" italics="true" pageId="0" pageNumber="145">Tilia begoniifolia</emphasis>
</taxonomicName>
) with blackberry (
<taxonomicName id="D3C5CE31F6CFF1969B649FC1E31763ED" class="Magnoliopsida" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rubus sanctus" order="Rosales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="sanctus">
<emphasis id="C14784283F26195EAC56AC1BE8C63D19" italics="true" pageId="0" pageNumber="145">Rubus sanctus</emphasis>
</taxonomicName>
) and fig (
<taxonomicName id="00EDA3BFC2AAF8598831FFAEA58E9AA9" class="Gastropoda" family="Ficidae" genus="Ficus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ficus carica" order="Littorinimorpha" pageId="0" pageNumber="145" phylum="Mollusca" rank="species" species="carica">
<emphasis id="F2FD0E373436B1B3E5075DBAD03E71B9" italics="true" pageId="0" pageNumber="145">Ficus carica</emphasis>
</taxonomicName>
) in shrub layer and with 
<normalizedToken id="649F26424E6656E657EC724E36F28218" originalValue="butcher’s-broom">butcher's-broom</normalizedToken>
(
<taxonomicName id="ADEB663ACC048453F16C267645BB7264" authorityName="Yeo" authorityYear="1966" class="Monocotyledoneae" family="Liliaceae" genus="Ruscus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ruscus colchicus" order="Liliiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="colchicus">
<emphasis id="3F35185B4A70A5CDB4A21D42E1D1B4F2" italics="true" pageId="0" pageNumber="145">Ruscus colchicus</emphasis>
</taxonomicName>
) in ground layer; ZISP • 
<specimenCount id="E82467E1779DBE892BD41B1083AE7884" type="male">4 males</specimenCount>
, 
<specimenCount id="7FC64B6ED4CF1E464C43C5BC308B45DA" type="female">4 females</specimenCount>
, same collection data as for preceding, 43°31'775&quot;N, 39°52'423&quot;E; alt. 
<quantity id="302BBB3559825A98FCBFC4607F1F3368" metricMagnitude="2" metricUnit="m" metricValue="1.01" unit="m" value="101.0">
<elevation id="D78808FBAA527B112A9B9F1BD6E3A664" metricMagnitude="2" metricUnit="m" metricValue="1.01" unit="m" value="101.0">101 m</elevation>
</quantity>
; 
<collectingDate id="F17E874E045C8EDF2B9CDB2AD7B97D58" value="2018-05-09">9 May 2018</collectingDate>
; 
<collectorName id="A101FFB8FC73C042D6ED77D43B194070">V. Lantsov</collectorName>
leg.; / Hornbeam (
<taxonomicName id="D538DED2DFF418ED12FA4225B67A99A2" authorityName="Mill" authorityYear="1768" class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis id="09CDC203985C932B7B87D67572021E49" italics="true" pageId="0" pageNumber="145">Carpinus orientalis</emphasis>
</taxonomicName>
) forest with 
<normalizedToken id="78F9BA3B826B8E5A0CC79BB73CBFA603" originalValue="butcher’s-broom">butcher's-broom</normalizedToken>
(
<taxonomicName id="698C22DE893074BD9EC512E95F716B2B" authorityName="Yeo" authorityYear="1966" class="Monocotyledoneae" family="Liliaceae" genus="Ruscus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ruscus colchicus" order="Liliiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="colchicus">
<emphasis id="7D43703D167F46A96B798C5175057F74" italics="true" pageId="0" pageNumber="145">Ruscus colchicus</emphasis>
</taxonomicName>
) and fern (
<taxonomicName id="50EE85E0E5DCC3004675AE5DB3F820F8" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium scolopendrium" order="Polypodiales" pageId="0" pageNumber="145" phylum="Polypodiopsida" rank="species" species="scolopendrium">
<emphasis id="DBBFAE401977A9C58804B40E2DAFF09B" italics="true" pageId="0" pageNumber="145">Asplenium scolopendrium</emphasis>
</taxonomicName>
) in ground layer; ZISP • 
<specimenCount id="BCB4CF0D58B911EAE881C941025FEF6D" type="male">1 male</specimenCount>
and 
<specimenCount id="E0850F62E90DED48CA477DA4C98F5C8B" type="female">1 female</specimenCount>
(in copula), 
<specimenCount id="8EA7E78D1CF698023271B413E0019CA9" type="female">1 female</specimenCount>
; same collection data as for preceding, 43°31'913&quot;N, 39°52'463&quot;E; alt. 
<quantity id="74A32F3D34223B403E1937A0477F3642" metricMagnitude="2" metricUnit="m" metricValue="1.53" unit="m" value="153.0">
<elevation id="8CE914ACD9BCD5F79E009E12CD1F595F" metricMagnitude="2" metricUnit="m" metricValue="1.53" unit="m" value="153.0">153 m</elevation>
</quantity>
; 
<collectingDate id="B8BFFF29BFDC4F3FAFEB7191C1864D9F" value="2018-05-09">9 May 2018</collectingDate>
, 
<collectorName id="7C768AB40F897299DD03625A84209443">V. Lantsov</collectorName>
leg. / Ash-lime (
<taxonomicName id="03A3568963E47912A06701DAE2E626A6" class="Dicotyledoneae" family="Oleaceae" genus="Fraxinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fraxinus excelsior" order="Oleales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="excelsior">
<emphasis id="F65EEA097EAA747BB730A8D8EE0E4744" italics="true" pageId="0" pageNumber="145">Fraxinus excelsior</emphasis>
</taxonomicName>
+ 
<taxonomicName id="8BBB207567A8D28BE07CD0AE59A0D065" class="Dicotyledoneae" family="Tiliaceae" genus="Tilia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Tilia begoniifolia" order="Malvales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="begoniifolia">
<emphasis id="919A15DB7ACF57025CA3BD59224F2728" italics="true" pageId="0" pageNumber="145">Tilia begoniifolia</emphasis>
</taxonomicName>
) forest with hornbeam (
<taxonomicName id="7DC60359D07FEB0E8A288269363F1373" class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus betulus" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="betulus">
<emphasis id="25720FE6CA3978BBA380EAD89B465487" italics="true" pageId="0" pageNumber="145">Carpinus betulus</emphasis>
</taxonomicName>
) and with black berry (
<taxonomicName id="F0F87F1F81CA84CD3815B37519BCE69F" class="Magnoliopsida" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rubus anatolicus" order="Rosales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="anatolicus">
<emphasis id="5738DF7F50BF786FA50546A945C178F8" italics="true" pageId="0" pageNumber="145">Rubus anatolicus</emphasis>
</taxonomicName>
) and miscellaneous herbs in ground layer; IEMT • 
<specimenCount id="16BB48AF574AF95C6359429CE68D3615" type="male">2 males</specimenCount>
, 
<specimenCount id="290B75EDCCF249782A0A967301EFD991" type="female">1 female</specimenCount>
, same collection data as for preceding, 43°31'825&quot;N, 39°52'653&quot;E; alt. 
<quantity id="301A7E2C811DCE15018801197C8E0959" metricMagnitude="1" metricUnit="m" metricValue="9.9" unit="m" value="99.0">
<elevation id="0DD5F0C226506958BE9812C31341BB24" metricMagnitude="1" metricUnit="m" metricValue="9.9" unit="m" value="99.0">99 m</elevation>
</quantity>
; 
<collectingDate id="9D7B0313778314879AAB0238D3AD8A25" value="2018-05-10">10 May 2018</collectingDate>
, 
<collectorName id="8C1E86CE120F8D5EC754CA59697D39B8">V. Lantsov</collectorName>
leg.; IEMT • 
<specimenCount id="200C7339E247A1C31D360479BB1B672E" type="male">1 male</specimenCount>
, same collection data as for preceding, 43°31'656&quot;N, 39°52'467&quot;E; alt. 
<quantity id="1CE80EE759DFA25C16EACBA9F5DE9CAA" metricMagnitude="1" metricUnit="m" metricValue="5.4" unit="m" value="54.0">
<elevation id="267B023065F41438DAEA7FE4B69E47E4" metricMagnitude="1" metricUnit="m" metricValue="5.4" unit="m" value="54.0">54 m</elevation>
</quantity>
; 
<collectorName id="C93495D3BA262AA2BF27D4F2554D39CC">V. Lantsov</collectorName>
leg.; ZISP • 
<specimenCount id="C409C24B2E38D5926748F1368EF39427" type="male">2 males</specimenCount>
, 
<specimenCount id="3052BE959E87F34E970232A2E9BC0C97" type="female">1 female</specimenCount>
, same collection data as for preceding, 43°32'257&quot;N, 39°52'682&quot;E; alt. 
<quantity id="6AAB726E6B22823EE0DD1F795772EC4E" metricMagnitude="1" metricUnit="m" metricValue="9.4" unit="m" value="94.0">
<elevation id="52599582F8FF7A258CB27F8B10B71920" metricMagnitude="1" metricUnit="m" metricValue="9.4" unit="m" value="94.0">94 m</elevation>
</quantity>
; 
<collectingDate id="67D7568C0E40102B91022DD8CD9DE5A8" value="2018-05-10">10 May 2018</collectingDate>
, 
<collectorName id="2AA61F723875DD19D1832A938E9582F7">V. Lantsov</collectorName>
leg.; / beech (
<taxonomicName id="517767ED603B3CCA7FB42F590EB22666" authorityName="Lipsky" authorityYear="1898" class="Dicotyledoneae" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fagus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis id="62668EE1A5197B17B1BC98844A626974" italics="true" pageId="0" pageNumber="145">Fagus orientalis</emphasis>
</taxonomicName>
) forest with addition of yew (
<taxonomicName id="01687FD50732DABFAF98A1BCEEEFE74E" class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Taxus baccata" order="Taxales" pageId="0" pageNumber="145" phylum="Coniferophyta" rank="species" species="baccata">
<emphasis id="824D5B882B82F9EADF91AF2F72DB83FD" italics="true" pageId="0" pageNumber="145">Taxus baccata</emphasis>
</taxonomicName>
) and hornbeam (
<taxonomicName id="BBF86FE97EAEA4DD7D9DA52BD3E64FEE" class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus betulus" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="betulus">
<emphasis id="AE1A2DCB9318E93F613437BD15C27764" italics="true" pageId="0" pageNumber="145">Carpinus betulus</emphasis>
</taxonomicName>
) with bunch grass (
<taxonomicName id="9C9B5D421904A867C94CD0F6817AB117" class="Monocotyledoneae" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calamagrostis arundinacea" order="Graminales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="arundinacea">
<emphasis id="DA12626AD694CA6EB30F672EB8E662A3" italics="true" pageId="0" pageNumber="145">Calamagrostis arundinacea</emphasis>
</taxonomicName>
) in ground layer; IEMT • 
<specimenCount id="4B9B861CAC82CDA2206F3F43C2EA9F38" type="male">6 males</specimenCount>
, 
<specimenCount id="1765627C4DEB9388EE31D78F4E03DC9B" type="female">3 females</specimenCount>
; Krasnodar Territory, environs 
<location id="B80E7705F8EFD3C39DECBD708528B115" LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:B80E7705F8EFD3C39DECBD708528B115" country="Russia" county="Krasnodarskiy" municipality="Sotshi prov." name="Shakhe River">Shakhe River</location>
higher than Salokh-Aul; 
<collectingDate id="06FFD6774F4906FE2D49DE45E138BCB4" value="2011-06-05" valueMax="2011-06-08" valueMin="2011-06-05">5-8 Jun. 2011</collectingDate>
; 
<collectorName id="B4220E9BA5F11C125AE67BF7EE0302E4">V. Pilipenko</collectorName>
leg.; VPMC
</materialsCitation>
; 
<materialsCitation id="3649214820E1FC95731B9423BC09711C" collectingDate="1909-06-29" collectingDateMax="2016-07-05" collectingDateMin="1909-06-29" collectorName="Mlokosevich, V. Lantsov" country="Russia" county="Tlyaratinsky district" determinerName="Ried" elevation="1792" location="river Dzhurmut" municipality="Salda" specimenCount="11" specimenCount-female="3" specimenCount-male="8" stateProvince="Dagestan">
<collectingCountry id="6227592819229A44E6368A425A2EEAF6" name="Russia">Russia</collectingCountry>
- 
<emphasis id="0F8FD823F963C5614EEBF7EAEE1C8DF1" bold="true" pageId="0" pageNumber="145">
<collectingRegion id="784CAC58033E28D0E7FDC21FA093C382" country="Russia" name="Dagestan">Dagestan</collectingRegion>
</emphasis>
• 
<specimenCount id="F49C12875201CD2FA160D79E3DB89676" type="female">1 female</specimenCount>
, 
<normalizedToken id="B1C265140068DBC2457DB53CC220B04B" originalValue="“Хочал-Даг">&quot;Khochal-Dag</normalizedToken>
, 
<normalizedToken id="34E86427B67FD7A562993736EAA4A3F0" originalValue="альп">alp</normalizedToken>
. 
<normalizedToken id="AF55B56AA81F965667EAE03592B05642" originalValue="обл">obl</normalizedToken>
. 
<normalizedToken id="EB089C7787C3CA4C28403D1E14B99110" originalValue="Дагст">Dagst</normalizedToken>
. [
<normalizedToken id="B0D042F80DAE6106E20AF51AA3505957" originalValue="Дагестан]”">
<collectingRegion id="1EEC4703C519CA70CCB29BF3B3D27CB9" country="Russia" name="Dagestan">Dagestan</collectingRegion>
]&quot;
</normalizedToken>
[Hochal-Dag, alp. Region Dagst. [
<collectingRegion id="E7E6375689E6FD015643357DC1BC2D9A" country="Russia" name="Dagestan">Dagestan</collectingRegion>
]]; 
<collectingDate id="9B45761CB3B9968D5FFB49BA67B1EFCC" value="1909-06-29">29 Jun. 1909</collectingDate>
<collectorName id="7414F55CA780248268DC8526484843DD">Mlokosevich</collectorName>
leg.; ZISP • 
<specimenCount id="DDBF2A4626DE0B13E48EC892E688D703" type="female">1 female</specimenCount>
; 
<collectingCounty id="3512ACF129BC28E0B18EF229186374C9">Tlyaratinsky district</collectingCounty>
, near village 
<collectingMunicipality id="58EFD344479D3B8228CD799B486C04CD">Salda</collectingMunicipality>
, stream on the right bank of the 
<location id="10A03F8FDE20845170E21CD52183F14F" LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:10A03F8FDE20845170E21CD52183F14F" country="Russia" county="Tlyaratinsky district" municipality="Salda" name="river Dzhurmut" stateProvince="Dagestan">river Dzhurmut</location>
; 41°58'388&quot;N, 46°30'446&quot;E; alt. 
<quantity id="E1AA32326A92D888C20858D0C372ECEA" metricMagnitude="3" metricUnit="m" metricValue="1.7919999999999998" unit="m" value="1792.0">
<elevation id="CEB32F886BC5289B8FFAF26EB8378C72" metricMagnitude="3" metricUnit="m" metricValue="1.7919999999999998" unit="m" value="1792.0">1792 m</elevation>
</quantity>
; 
<collectingDate id="4263EBBF254A4505538FC6AAA4AECC06" value="2016-07-02">2 Jul. 2016</collectingDate>
, 
<collectorName id="86B97B9427A13F14230C5581F3044EF2">V. Lantsov</collectorName>
leg.; / &quot;Oak (
<taxonomicName id="F20FBF4639DDF13D129271304479DCF2" class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus iberica" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="iberica">
<emphasis id="24397A86DD8532126E4B3B4A246751AD" italics="true" pageId="0" pageNumber="145">Quercus iberica</emphasis>
</taxonomicName>
) sparse forest with 
<taxonomicName id="D34767CC80036259AF2EDA30B761BEC3" class="Dicotyledoneae" family="Salicaceae" genus="Populus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Populus tremula" order="Salicales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="tremula">
<emphasis id="857D991C6EDFC482408937756C83296F" italics="true" pageId="0" pageNumber="145">Populus tremula</emphasis>
</taxonomicName>
and 
<taxonomicName id="FCD43D4719BB3F8E4F8945168CF715D6" class="Dicotyledoneae" family="Aceraceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Acer platanoides" order="Sapindales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="platanoides">
<emphasis id="8BD1F5A23E5C4EBA8EC8C81CED81A00F" italics="true" pageId="0" pageNumber="145">Acer platanoides</emphasis>
</taxonomicName>
in tree layer, 
<emphasis id="5FFB1797A03505C2452C70A401F79F65" italics="true" pageId="0" pageNumber="145">Rosa canina</emphasis>
, 
<taxonomicName id="05581CF99B786529B413463D083B8496" class="Dicotyledoneae" family="Rosaceae" genus="Spiraea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Spiraea crenata" order="Rosales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="crenata">
<emphasis id="3C7E07968497B874A39E82AD272567B5" italics="true" pageId="0" pageNumber="145">Spiraea crenata</emphasis>
</taxonomicName>
and 
<taxonomicName id="3225F81816A15048AA602D55B6421140" class="Dicotyledoneae" family="Rosaceae" genus="Cotoneaster" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cotoneaster integerrimus" order="Rosales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="integerrimus">
<emphasis id="CD6CB0DE5C00552E7C2592A945D70D13" italics="true" pageId="0" pageNumber="145">Cotoneaster integerrimus</emphasis>
</taxonomicName>
in shrub, and clover (
<taxonomicName id="D8E2FF190F1F5ECA7268A9176F5A3479" class="Magnoliopsida" family="Fabaceae" genus="Trifolium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trifolium canescens" order="Fabales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="canescens">
<emphasis id="DF0BBFC67FE65C2E9D96477FA6C7EA3D" italics="true" pageId="0" pageNumber="145">Trifolium canescens</emphasis>
</taxonomicName>
) and miscellaneous herbs in ground layer; ZISP • 
<specimenCount id="8144FC4B979D33C884989A3E26723B88" type="male">2 males</specimenCount>
, 
<specimenCount id="190A2489EFF7A43CD866A6FC5797187E" type="female">1 female</specimenCount>
; near village Betelda, stream on the left bank of the 
<location id="FED2A7EFB21D4CAE46D06E4881F53C13" LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:FED2A7EFB21D4CAE46D06E4881F53C13" country="Russia" county="Tlyaratinsky district" municipality="Salda" name="river Dzhurmut" stateProvince="Dagestan">river Dzhurmut</location>
; 41°56'853&quot;N, 46°32'424&quot;E; alt. 
<quantity id="66BF105C848049E6BFF0DBAE3E982EFD" metricMagnitude="3" metricUnit="m" metricValue="1.85" unit="m" value="1850.0">
<elevation id="418A8F532A83C6740D7C0033CA053884" metricMagnitude="3" metricUnit="m" metricValue="1.85" unit="m" value="1850.0">1850 m</elevation>
</quantity>
; 
<collectingDate id="08AD0F1CA1453C265EAF1FB52803CFE2" value="2016-07-03">3 Jul. 2016</collectingDate>
, 
<collectorName id="4145A94E41CFAC9979B75198A118AEC0">V. Lantsov</collectorName>
leg.; / Willow (
<taxonomicName id="0C9982726325B8E95C7C0E0109891E61" class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Salix caprea" order="Aplousobranchia" pageId="0" pageNumber="145" phylum="Chordata" rank="species" species="caprea">
<emphasis id="0241CBF523B04FAFE2B7C485E75BD9BA" italics="true" pageId="0" pageNumber="145">Salix caprea</emphasis>
</taxonomicName>
) shrub with herbaceous layer (
<taxonomicName id="3712CCE3FCCEE3990E3D05317C7CDD1F" class="Dicotyledoneae" family="Onagraceae" genus="Epilobium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Epilobium montanum" order="Myrtiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="montanum">
<emphasis id="291A7A70C75094D8ED433CD4D2AA970E" italics="true" pageId="0" pageNumber="145">Epilobium montanum</emphasis>
</taxonomicName>
) (dominant), 
<taxonomicName id="C91EE9EA2D982DCB7AEB269EFDC7D08D" authorityName="Albov" authorityYear="1894" class="Dicotyledoneae" family="Brassicaceae" genus="Cardamine" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cardamine seidlitziana" order="Papaverales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="seidlitziana">
<emphasis id="268CBE946F324F12EBB1458CC1521FB3" italics="true" pageId="0" pageNumber="145">Cardamine seidlitziana</emphasis>
</taxonomicName>
, 
<taxonomicName id="ABCD40123A8757A1516B9CE359E739CE" class="Dicotyledoneae" family="Lamiaceae" genus="Mentha" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Mentha caucasica" order="Tubiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="caucasica">
<emphasis id="D724C029A1AC171F3497619AEB1353CB" italics="true" pageId="0" pageNumber="145">Mentha caucasica</emphasis>
</taxonomicName>
, 
<taxonomicName id="E92FE00553D9ABC391CB4DEDDA10DE42" class="Dicotyledoneae" family="Scrophulariaceae" genus="Veronica" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Veronica anagallis-aquatica" order="Tubiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="anagallis-aquatica">
<emphasis id="F8F83B34662C13C279C1D1716C756D02" italics="true" pageId="0" pageNumber="145">Veronica anagallis-aquatica</emphasis>
</taxonomicName>
and 
<taxonomicName id="45AFDF4A523C79B11AC7E753627A9C70" authorityName="Moench" authorityYear="1794" baseAuthorityName="L." class="Dicotyledoneae" family="Caryophyllaceae" genus="Myosoton" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Myosoton aquaticum" order="Centrospermae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="aquaticum">
<emphasis id="C2CF656C9629DCC86F6B161F74D9C839" italics="true" pageId="0" pageNumber="145">Myosoton aquaticum</emphasis>
</taxonomicName>
community near spring above the flood plain of the left bank of the river; ZISP • 
<specimenCount id="D606DA44164745935EC5C4B9BBD47105" type="male">2 males</specimenCount>
; near village Genekolob, on the high left bank of the river Dzhurmut; 41°57'392&quot;N, 46°31'276&quot;E; alt. 
<quantity id="316908049111412FE52577C9DBF3273A" metricMagnitude="3" metricUnit="m" metricValue="2.011" unit="m" value="2011.0">
<elevation id="2AF80D36C50A73F15758F69665A9C450" metricMagnitude="3" metricUnit="m" metricValue="2.011" unit="m" value="2011.0">2011 m</elevation>
</quantity>
; 
<collectingDate id="0324265756ADB0AA755A8B689A1C2950" value="2016-07-04">4 Jul. 2016</collectingDate>
, 
<collectorName id="C9473EA6C4C2B1F93FD6799E77C1B6DE">V. Lantsov</collectorName>
leg.; / Oak (
<taxonomicName id="DBFE7652306D7DC3F5399F394503A523" class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus iberica" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="iberica">
<emphasis id="327E97F8F4FD7D0820491FB521B4BDF0" italics="true" pageId="0" pageNumber="145">Quercus iberica</emphasis>
</taxonomicName>
) forest with mountain-ash (
<taxonomicName id="FB3AB81A548BC87D5B443C606F888F86" class="Dicotyledoneae" family="Rosaceae" genus="Sorbus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sorbus aucuparia" order="Rosales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="aucuparia">
<emphasis id="98EB4D7A213BB1CD0AEEAD1050F98658" italics="true" pageId="0" pageNumber="145">Sorbus aucuparia</emphasis>
</taxonomicName>
), wayfaring tree (
<taxonomicName id="9D4FF2D70F2C74F29FF2D8DD8D0EA1A4" class="Dicotyledoneae" family="Caprifoliaceae" genus="Viburnum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Viburnum lantana" order="Dipsacales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="lantana">
<emphasis id="FECEB63DA21915B94087701A85807F22" italics="true" pageId="0" pageNumber="145">Viburnum lantana</emphasis>
</taxonomicName>
) in shrub layer [the lower border of a forest and a forb-cereal meadow]; ZISP • 
<specimenCount id="B98BA3B8992E52C05FDBAF96BFA99679" type="male">2 males</specimenCount>
; same collection data as for preceding, 41°57'449&quot;N, 46°31'228&quot;E; alt. 
<quantity id="D7F260859CA2D585A2C109C104BDE553" metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">
<elevation id="80140657C8AFE356182D62E64555E666" metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">1977 m</elevation>
</quantity>
; 
<collectingDate id="AD0E142EBABD1802E3B663BB8AB21F94" value="2016-07-04">4 Jul. 2016</collectingDate>
, 
<collectorName id="2537B2DECB16726A4EF9144816110320">V. Lantsov</collectorName>
leg.; ZISP • 
<specimenCount id="BFCDE37F6244C78ECE128BC72AFD3870" type="male">2 males</specimenCount>
, between the villages of Gerel and Genekolob, the right bank of the river Dzhurmut, 41°55'619&quot;N, 46°34'468&quot;E; alt. 
<quantity id="2FC1947F5343C05AAF8A67AF397805E4" metricMagnitude="3" metricUnit="m" metricValue="1.925" unit="m" value="1925.0">
<elevation id="F705E748B508D84F55F10D2A0610F64F" metricMagnitude="3" metricUnit="m" metricValue="1.925" unit="m" value="1925.0">1925 m</elevation>
</quantity>
; 
<collectingDate id="9D86380AF381F8175DC2330B9B179BAE" value="2016-07-05">5 Jul. 2016</collectingDate>
, 
<collectorName id="0C5FD4F2FCF5ADEFD306CFD3DF1950C2">V. Lantsov</collectorName>
leg.; / Above the floodplain terrace with a rare herbaceous birch (
<taxonomicName id="2DC8A769655C5654183BF6FE8880CD5B" class="Dicotyledoneae" family="Betulaceae" genus="Betula" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Betula raddeana" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="raddeana">
<emphasis id="230329831FFFAAA9344CBDC0DE76163A" italics="true" pageId="0" pageNumber="145">Betula raddeana</emphasis>
</taxonomicName>
) forest with willow (
<taxonomicName id="4AE5D2183630A290B2132421A655D461" class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Salix caprea" order="Aplousobranchia" pageId="0" pageNumber="145" phylum="Chordata" rank="species" species="caprea">
<emphasis id="C07191EAEAF9513FDCADF2CA4929B794" italics="true" pageId="0" pageNumber="145">Salix caprea</emphasis>
</taxonomicName>
) (coppice forest); ZISP
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="922A217BA0B173B8F2086C4BBC21378D" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="1BB359AB7F2FE826F28FFD3B7686772B" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="95609981A3F096E4E722DA7E25C93141" pageId="0" pageNumber="145">Dark stripes medially on lighter background extending entire length of scutum. Inner gonostylus posteriorly with wedge-shaped projection directed anteriorly. Tergite 9 at apex with three rounded notches, middle one usually deepest and widest.</paragraph>
</subSubSection>
<subSubSection id="7B3F3E78BFE9CB360DCB03D4D1F463EC" pageId="0" pageNumber="145" type="redescription">
<paragraph id="25E9FFB7EF95F0F2C92EA7C893A93029" pageId="0" pageNumber="145">Redescription.</paragraph>
<paragraph id="9536DE9E84214C9B5EDF4FEC31C8202B" pageId="0" pageNumber="145">
Adult male (Fig. 
<figureCitation id="F7254C2631217FFDAA3E9CAD60A58083" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1A, B</figureCitation>
). General color grey. Body length 14-17 mm, wing 14-17.5 mm.
</paragraph>
<caption id="9CE7720B870B74F73FCE674D7F2E2D38" doi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145" start="Figure 1" startId="F1">
<paragraph id="D9330ECF39116EB0F662DDF46E4A09F4" pageId="0" pageNumber="145">
<emphasis id="7A51AD6D405BEAD8B4F6F75C9F4B8D05" bold="true" pageId="0" pageNumber="145">Figure 1.</emphasis>
<taxonomicName id="15C43F0E3CFF9C616AD5EBC0D3A5960C" lsidName="T. (Lunatipula) caucasica" pageId="0" pageNumber="145" rank="species" species="caucasica" subGenus="Lunatipula">T. (Lunatipula) caucasica</taxonomicName>
male habitus 
<emphasis id="ED70823C3641DDC2DD55F55910DA3638" bold="true" pageId="0" pageNumber="145">A</emphasis>
general lateral view 
<emphasis id="CC5FAEC19D978D1BBD1EB1C240FFB9FA" bold="true" pageId="0" pageNumber="145">B</emphasis>
head and thorax, dorsal view. Scale bars: 10 mm (
<emphasis id="3D5B2082DB3B330B29967D685F57CC70" bold="true" pageId="0" pageNumber="145">A</emphasis>
), 1 mm (
<emphasis id="B8118861645FB67702BA884F77AEEBB5" bold="true" pageId="0" pageNumber="145">B</emphasis>
).
</paragraph>
</caption>
<paragraph id="BDF01A8A972E95D2BB73E0A3F4E45E96" pageId="0" pageNumber="145">
<emphasis id="0676CC8626B2816A76A3A63FDE9D4B5F" bold="true" italics="true" pageId="0" pageNumber="145">Head.</emphasis>
Light grey, sometimes bluish (Figs 
<figureCitation id="131335BC300C72F80A8352AEC6FAD2B5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1B</figureCitation>
, 
<figureCitation id="D0041866AED40C923DA756AE913EB6D6" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">13A</figureCitation>
). Rostrum whitish dorsally, flanks rusty or brownish with whitish bristles; nasus distinct, with long, whitish yellow setae; palpus grey. Frons with rows of brown setae along inner edge of eye. Vertex with faint dark median line, posteriorly with dark grey striation, with group of small rusty setae arranged fan-like on either side of glabrous medial area. Genae rusty; wide dark median stripe ventrally on rostrum, continuing along gula midline. Pharyngeal foramen framed with long, rusty hair-like setae. Postgenae light grey.
</paragraph>
<paragraph id="11BD36574EF226305E441AAEF15D29FD" pageId="0" pageNumber="145">
<emphasis id="D9ECCFE6A31DF670818039085885CC72" bold="true" italics="true" pageId="0" pageNumber="145">Antenna</emphasis>
(Fig. 
<figureCitation id="93C4938AEB3B03997DB13FC8A796BEB1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1B</figureCitation>
) 13-segmented, if bent backwards just reaching wing base. Scape light brown or grey, depending on angle of rotation to light source, with slightly pronounced silvery pruinescence and brown horizontal striations. Pedicel yellow, sometimes brownish at base. Flagellomeres brownish, thickened at base, verticils subequal in length to corresponding flagellomeres.
</paragraph>
<paragraph id="85811C5CFC816FA199F5D268A5EBFFD2" pageId="0" pageNumber="145">
<emphasis id="97ECB26B3ABD4ACBD3A98FBA17D2C4F5" bold="true" italics="true" pageId="0" pageNumber="145">Thorax</emphasis>
(Fig. 
<figureCitation id="98030BB52493A65E421A35D4361938D4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1B</figureCitation>
). Grey or blue-grey. Pronotum with dark grey median stripe. Katepisternum, anepisternum and meron grey, glabrous. Scutum with four longitudinal wide brownish stripes on lighter background; medial stripes extending length of scutum. Scutal lobes, scutellum and mediotergite grey with light bristles in dark brown sockets.
</paragraph>
<paragraph id="5E8929B809A723EE1EBBFE3B4A317914" pageId="0" pageNumber="145">
<emphasis id="F17F9831087A94B91010EBA47513E3C8" bold="true" italics="true" pageId="0" pageNumber="145">Wings</emphasis>
(Fig. 
<figureCitation id="DAB0AE6769656783B2B62BA894CA52CB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1A</figureCitation>
) greyish with a bluish tinge in transmitted light, stigma brown; narrow whitish area (lunule) extending to base of discal cell.
</paragraph>
<paragraph id="39B90788A53C57CDB370B50702E8BF56" pageId="0" pageNumber="145">
<taxonomicName id="F23D019CCBCE4B1C7628C966A3D209E7" genus="Halteres" lsidName="Halteres" pageId="0" pageNumber="145" rank="genus">
<emphasis id="091ACE50F5F0D67D9EB13266588294EF" bold="true" italics="true" pageId="0" pageNumber="145">Halteres</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="07C01D93BF4479D10473F8F9580A2586" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1A</figureCitation>
) with light brown or dark-yellow stem, lighter yellow at base, covered with short small pale setae; knob dark brown to black.
</paragraph>
<paragraph id="404EC389CA2C6F68E0C85C4C7A555F30" pageId="0" pageNumber="145">
<emphasis id="8FFA7381AFA51E00B862FDD68AD24D2D" bold="true" italics="true" pageId="0" pageNumber="145">Legs</emphasis>
with coxae light grey with long light hairs. Coxae may appear darker when specimen is rotated. Trochanters yellow. Femora yellow at base, rest of femora, tibiae, and tarsi dark brown.
</paragraph>
<paragraph id="FBE6BCBB31D0EDB68E321F8AEF163434" pageId="0" pageNumber="145">
<emphasis id="1A665457247D903CB1A14190281F8752" bold="true" italics="true" pageId="0" pageNumber="145">Abdomen</emphasis>
(Figs 
<figureCitation id="66384AB41CDAFD0D1702AB81EE339F07" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1A</figureCitation>
, 
<figureCitation id="D158E7B1ABDCF139A4BA625007649E8C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2A</figureCitation>
). Grey or dark grey; tergites with dark lateral stripe narrowly interrupted with whitish at posterior margins, lateral margins with wider whitish edge.
</paragraph>
<caption id="26F5967C7FEF776C8989223E577BE33D" doi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145" start="Figure 2" startId="F2">
<paragraph id="287AB7372D4767FCC63F30731B1353E1" pageId="0" pageNumber="145">
<emphasis id="6975DCD9C97DFF8F6D0B812EBB39B35F" bold="true" pageId="0" pageNumber="145">Figure 2.</emphasis>
<taxonomicName id="E2159BC45D4B0B3E49CB5F6A52D86B80" lsidName="T. (Lunatipula) caucasica" pageId="0" pageNumber="145" rank="species" species="caucasica" subGenus="Lunatipula">T. (Lunatipula) caucasica</taxonomicName>
<emphasis id="A036F8E4C950AF4B3E1C91B9F64B32C5" bold="true" pageId="0" pageNumber="145">A</emphasis>
terminal segments of male abdomen, lateral view (dry) 
<emphasis id="374EAA43F9FC4339CAB952777E59AD5D" bold="true" pageId="0" pageNumber="145">B-L</emphasis>
male terminalia (after KOH 10% treatment) 
<emphasis id="854C5D81B67611463E373267E259FABA" bold="true" pageId="0" pageNumber="145">B</emphasis>
hypopygium, caudal view 
<emphasis id="C05C456A5673410D3939BE5AF1A71734" bold="true" pageId="0" pageNumber="145">C</emphasis>
hypopygium, lateral view 
<emphasis id="D8CDC7D8091C8F50505DC0B50563813E" bold="true" pageId="0" pageNumber="145">D</emphasis>
sternite 8, ventral view 
<emphasis id="223750BA34ECA8E3F07D8E4234FF8A66" bold="true" pageId="0" pageNumber="145">E</emphasis>
tergite 9, caudal view 
<emphasis id="47449F9D0815C7543CE2D339C7685359" bold="true" pageId="0" pageNumber="145">F</emphasis>
tergite 9, dorsal view 
<emphasis id="D4542170619C989F57F893E11A0B86C8" bold="true" pageId="0" pageNumber="145">G</emphasis>
inner and outer gonostylus, lateral view 
<emphasis id="0FC9A57C4F969A755538654F4F5BBA67" bold="true" pageId="0" pageNumber="145">H</emphasis>
inner and outer gonostylus, caudal view 
<emphasis id="48B6E1CDBB42D8E869571C30F6995864" bold="true" pageId="0" pageNumber="145">I</emphasis>
gonocoxal fragment, dorsal view 
<emphasis id="3F38C87E7FE66F77CBEF50AD86FA16FF" bold="true" pageId="0" pageNumber="145">J</emphasis>
semen pump and aedeagus, lateral view 
<emphasis id="CBB8A4A915B242D324301FC6F8764F3A" bold="true" pageId="0" pageNumber="145">K</emphasis>
adminiculum and paired apical appendages of sternite 9, caudal view 
<emphasis id="74D652F92E0928E0485383C2FCE1F6C0" bold="true" pageId="0" pageNumber="145">L</emphasis>
adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8-paired appendages of sternite 8; as 9-apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s8-sternite 8; s9-sternite 9; t9-tergite 9. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="35C7AF081AEC1BBB8DD476835C069C6C" pageId="0" pageNumber="145">
<emphasis id="51577C199F7823CFE6B5D1E9BDEF8A24" bold="true" italics="true" pageId="0" pageNumber="145">Terminalia</emphasis>
(Figs 
<figureCitation id="F143AAFE19D6D48116527A757EDE5F34" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2</figureCitation>
, 
<figureCitation id="0C698594C08317D33B664EA305BD8166" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14A, F</figureCitation>
). Hypopygium (Fig. 
<figureCitation id="13C5F868CFABD58E05E7C38AAF308642" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2B, C</figureCitation>
) not much thicker than rest of abdomen, brown with moderately dense grey pruinescence, with golden yellow hairs. Gonocoxite (Fig. 
<figureCitation id="73DCA65EAC7E789D63125FC993CAD821" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2C, G</figureCitation>
) with broad tooth ventrally. Tergite 9 (Figs 
<figureCitation id="CEDEBBFEB3267EDA8FDC45D04E4E8586" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2C, E, F</figureCitation>
; 
<figureCitation id="68BD15B0A23C06B665A9B23AC3E513AE" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14A</figureCitation>
) transverse, lateral corner broadly rounded, at apex with three rounded notches, middle one usually deepest and widest, bounded laterally by two large dentate projections with blunt apices, clearly visible when viewed from above (Figs 
<figureCitation id="F1F63223B7DF593BD71DC5FA9BFFC0BE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2F</figureCitation>
, 
<figureCitation id="6167167AAC9B2DBFB18323BEE5433D7E" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14A</figureCitation>
) (
<bibRefCitation id="77DA87644C9F03CC0596B918FD1039B9" author="Riedel, MP" journalOrPublisher="Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere" pageId="0" pageNumber="145" pagination="13 - 22" refId="B9" refString="Riedel, MP, 1920. Nematocera Polyneura (Dipt.) aus dem Kaukasus. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 43: 13 - 22" title="Nematocera Polyneura (Dipt.) aus dem Kaukasus." volume="43" year="1920">Riedel 1920</bibRefCitation>
; 
<bibRefCitation id="7A299F314BDE370170A2C54A2F6599C7" author="Savchenko, EN" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="0" pageNumber="145" refId="B12" refString="Savchenko, EN, 1964. Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N.S. 89, Nasekomye Dvukrylye [Diptera] 2(4): 1-503. [in Russian]" title="Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N. S. 89, Nasekomye Dvukrylye [Diptera] 2 (4): 1 - 503. [in Russian]" year="1964">Savchenko 1964</bibRefCitation>
). When viewed from behind, three pairs of very short projections are visible (Fig. 
<figureCitation id="A914D678162A6001DD4FD0CEAD4CCED5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2E</figureCitation>
). Outer gonostylus (Figs 
<figureCitation id="003609FDC85B2CB33981D1F8680B96DD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2G</figureCitation>
, 
<figureCitation id="CFB0E5D3341219600D2D2C4917B65C7A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14F</figureCitation>
) very small, wineglass shaped, elongated distally. Inner gonostylus posteriorly, with wedge-shaped projection (Figs 
<figureCitation id="D67862F4CBAF5B74D406914E33F38A01" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2G</figureCitation>
, 
<figureCitation id="9D60B1CC9886DDF87A5C0B1FEE026680" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14F</figureCitation>
). Paired appendage of sternite 8 (Fig. 
<figureCitation id="F546016BE96F66FC639EA1E44B577EA4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2D</figureCitation>
) with base ~ 2 
<normalizedToken id="CD8DFCFC1F7CA0CED574A15F45A444D4" originalValue="×">x</normalizedToken>
as long as wide, with thin rigid yellow setae along edge, not obscuring gap in middle; long, glabrous red-brown spine arising from base, bent, and intersecting medially. Paired appendage of sternite 9 (Fig. 
<figureCitation id="A503105C3C2FDC8DEF531EEF585B2E8D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2K, L</figureCitation>
) with thick pubescent apex. Gonocoxal fragment (Fig. 
<figureCitation id="8600595E006826FA8776694E0B2B9D21" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2I</figureCitation>
) flattened at the base. Semen pump and aedeagus as in Figure 
<figureCitation id="8895FB6D37065D6EE18AD8E4D65F5C65" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2J</figureCitation>
.
</paragraph>
<paragraph id="5B8D31E89D56C12F41DDA66C1C2BEDE4" pageId="0" pageNumber="145">
<emphasis id="BC7D4EBB07388B0A3EC35001AA729D32" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Body length 17-21.5 mm, wing 15.5-18 mm. Antennae with scape and pedicel brownish yellow. Legs stocky, black to brown, yellow at base, femora grey. Wings with noticeable bluish tint; whitish area (lunule) of wings barely reaching base of discal cell. Female terminalia (Fig. 
<figureCitation id="C8C0ADD2C829B5B5B7C562FC289E7E0E" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">12A-C</figureCitation>
) comparatively short. Tip of cercus upcurved, length of cercus ~ 0.8 mm. Hypogynial valve only slightly longer than width at base; length of hypogynial valve 0.2 mm. Sternite 9 and furca as in Figure 
<figureCitation id="0147064F8F06BF3273099D49E842DEBC" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">12A, C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="159DD335E4CBD27D1971C18FD9EA2A86" pageId="0" pageNumber="145" type="comparisons">
<paragraph id="F4B84A8C92B0EA111842CAB97F9B0B94" pageId="0" pageNumber="145">Comparison with closely related species.</paragraph>
<paragraph id="14857F2F90684305453A64CB96E3967D" pageId="0" pageNumber="145">
This species differs from other species of the 
<taxonomicName id="9845F2ADA43C313AF5836CFEBBE51271" authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="5B4BD6C9BF5EE7DB0073B20FDC7C6D3A" italics="true" pageId="0" pageNumber="145">Tipula caucasica</emphasis>
</taxonomicName>
group primarily by the presence of a large, anteriorly directed sharp wedge-shaped projection located posteriorly on the inner gonostylus.
</paragraph>
</subSubSection>
<subSubSection id="61111D4B4594DA187A5213E0E1280DFE" pageId="0" pageNumber="145" type="elevation">
<paragraph id="AE334B373537CC6BC2F1D7B127F243B7" pageId="0" pageNumber="145">Elevation.</paragraph>
<paragraph id="49B380CCB67997FCADBF477D4C5DE53D" pageId="0" pageNumber="145">Adults were collected at altitudes ranging from sea level (Black Sea coast at 35 m) to 2011 m (in Dagestan).</paragraph>
</subSubSection>
<subSubSection id="2F173B59AB9A4267EE74960BC8D6659B" pageId="0" pageNumber="145" type="flight period">
<paragraph id="660D16175C0DBDF3869C43AD68F53F78" pageId="0" pageNumber="145">Flight period.</paragraph>
<paragraph id="B499A24898E0D24B7511D88859FBA1C0" pageId="0" pageNumber="145">From end of April to the beginning of July.</paragraph>
</subSubSection>
<subSubSection id="4633EAF289602A0AD38DB7022F6B4F1D" pageId="0" pageNumber="145" type="habitat">
<paragraph id="03AEAF8C03B063D5BA564B1F087B71A2" pageId="0" pageNumber="145">Habitat.</paragraph>
<paragraph id="3214CE0159415522803BDD050B7E2751" pageId="0" pageNumber="145">
Specimens were captured in diverse forest habitats (see above) predominantly in mesophytic moderately moist deciduous or mixed communities with 
<taxonomicName id="19B39B6D704A7B8A760E17CF4D1CB3C0" authorityName="Lipsky" authorityYear="1898" class="Dicotyledoneae" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fagus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis id="D0CA29D9CB61A3DCB37235DE3C90B76F" italics="true" pageId="0" pageNumber="145">Fagus orientalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A9A594B04376A4263F58323FC130A813" class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Taxus baccata" order="Taxales" pageId="0" pageNumber="145" phylum="Coniferophyta" rank="species" species="baccata">
<emphasis id="9EE06763B85B556A90A0C227F2797652" italics="true" pageId="0" pageNumber="145">Taxus baccata</emphasis>
</taxonomicName>
, 
<taxonomicName id="D9B6B771D194DCCDC0998109CF843064" authorityName="Mill" authorityYear="1768" class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis id="D95D1FD6505B3C18C9E4CD8A9149D2C7" italics="true" pageId="0" pageNumber="145">Carpinus orientalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5EA998DC3271A1775117F1226FDBF4D4" class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus iberica" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="iberica">
<emphasis id="79CBC8953F48086B10096C788CF0BD2B" italics="true" pageId="0" pageNumber="145">Quercus iberica</emphasis>
</taxonomicName>
, 
<taxonomicName id="AA3FFDD25AB07FF1963196902AB4F2B9" class="Dicotyledoneae" family="Salicaceae" genus="Populus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Populus tremula" order="Salicales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="tremula">
<emphasis id="D262FD70C3D9EDD4D919134A544CF7FE" italics="true" pageId="0" pageNumber="145">Populus tremula</emphasis>
</taxonomicName>
, 
<taxonomicName id="70177B21707713A93464D64BC7192B0E" class="Dicotyledoneae" family="Aceraceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Acer platanoides" order="Sapindales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="platanoides">
<emphasis id="20A67BA0280657363B2306D99EBC4B12" italics="true" pageId="0" pageNumber="145">Acer platanoides</emphasis>
</taxonomicName>
, etc., sometimes in communities near springs with 
<taxonomicName id="4392BABF6125AE77294B85114976CDDA" class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Salix caprea" order="Aplousobranchia" pageId="0" pageNumber="145" phylum="Chordata" rank="species" species="caprea">
<emphasis id="190B21BC5A77A8754FFD2216EAB193B3" italics="true" pageId="0" pageNumber="145">Salix caprea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EB9674B728FA8CFEA20470A9AD4131D1" pageId="0" pageNumber="145" type="distribution">
<paragraph id="CEAFE60237FF2E634EA27B44C367629A" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="C136749CE5DA8F122F56A550167E06CB" pageId="0" pageNumber="145">
Endemic to the Caucasus. It was recently noted for Russia (Dagestan) for the first time (
<bibRefCitation id="0F41EB0EA2A18ED2D7BF271B7A2D4ECC" DOI="https://doi.org/10.47640/978-5-00105-586-0_2020_292" author="Lantsov, VI" journalOrPublisher="Fauna of the Stavropol region" pageId="0" pageNumber="145" refId="B7" refString="Lantsov, VI, 2020. Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24-29, 2020. Proceedings of conference / Responsible editors: O.G.Ovchinnikova, I.V.Shamshev. St.Petersburg: Russian Entomological Society, LEMA, 292-296. https://doi.org/10.47640/978-5-00105-586-0_2020_292" title="Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24 - 29, 2020. Proceedings of conference / Responsible editors: O. G. Ovchinnikova, I. V. Shamshev. St. Petersburg: Russian Entomological Society, LEMA, 292 - 296." url="https://doi.org/10.47640/978-5-00105-586-0_2020_292" year="2020">Lantsov 2020</bibRefCitation>
). Currently known from the West Caucasus (first record), East Caucasus and Transcaucasus.
</paragraph>
</subSubSection>
<subSubSection id="418129D99BF2535021DF1E85B2B8522A" pageId="0" pageNumber="145" type="remarks">
<paragraph id="21F5B4DCB5E7807686451DC274896DAD" pageId="0" pageNumber="145">Remarks.</paragraph>
<paragraph id="77A8543C738105495D1A9635989D4D14" pageId="0" pageNumber="145">
According to 
<bibRefCitation id="90B52E639F7EE0412F46229033EDEA11" author="Riedel, MP" journalOrPublisher="Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere" pageId="0" pageNumber="145" pagination="13 - 22" refId="B9" refString="Riedel, MP, 1920. Nematocera Polyneura (Dipt.) aus dem Kaukasus. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 43: 13 - 22" title="Nematocera Polyneura (Dipt.) aus dem Kaukasus." volume="43" year="1920">Riedel (1920)</bibRefCitation>
, a series of type specimens, four males, two females. 
<normalizedToken id="50DCAB4EAAFA1E674EDF7B34A6DCD0E7" originalValue="“Mts’chet">&quot;Mts'chet</normalizedToken>
prope Tiflis [Georgia, Tbilisi], 6. 5. 1913 (Mus. Caucas. Ph. Zaitzev)&quot; were deposited in the &quot;Museum Caucasicum in Tiflis&quot;. In the 
<normalizedToken id="828C01C462C88B5274B69C0AC0FDE32E" originalValue="author’s">author's</normalizedToken>
publication, neither the holotype nor the paratypes are indicated. Presumably, they are not indicated on the material itself. Because this material turned out to be inaccessible to the authors of this article - it is not yet known whether this material has survived at all - there was no opportunity to designate lectotype and paralectotypes.
</paragraph>
</subSubSection>
</treatment>
</document>