<documentID-CLB-Dataset="5452"ID-DOI="http://dx.doi.org/10.3897/zookeys.1048.67564"ID-GBIF-Dataset="57d9ade8-67e7-4d65-9581-b85a80e88893"ID-PMC="PMC8292283"ID-Pensoft-Pub="1313-2970-1048-145"ID-Pensoft-UUID="93FE96A4127A5D8FAA3A39BCB3E4B729"ID-PubMed="34316267"ID-ZooBank="578795FD41BF4807B0016F7B5AC4D4B9"ModsDocID="1313-2970-1048-145"checkinTime="1626283267302"checkinUser="pensoft"docAuthor="Lantsov, Vladimir I. & Pilipenko, Valentin E."docDate="2021"docId="219A9352B86056889DBEB0947CC84DEA"docLanguage="en"docName="ZooKeys 1048: 145-175"docOrigin="ZooKeys 1048"docPubDate="2021-07-13"docSource="http://dx.doi.org/10.3897/zookeys.1048.67564"docTitle="Tipula (Lunatipula) caucasica Riedel 1920"docType="treatment"docVersion="6"id="93FE96A4127A5D8FAA3A39BCB3E4B729"lastPageNumber="145"masterDocId="93FE96A4127A5D8FAA3A39BCB3E4B729"masterDocTitle="A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group"masterLastPageNumber="175"masterPageNumber="145"pageNumber="145"updateTime="1732760741040"updateUser="ExternalLinkService">
<mods:titleid="8C4F0DEFE576778BF6BDDDCD420A0E4F">A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group</mods:title>
<mods:affiliationid="3FE40A84AEB044D17B806FC89436C83B">Tembotov Institute of Ecology of Mountain Territories of Russian Academy of Sciences, I. Armand str., 37 a, Nalchik 360051, Russia</mods:affiliation>
<figureCitationid="D1AEE56CBDA76565EB487F5CA859F6CB"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)."figureDoi="10.3897/zookeys.1048.67564.figure1"httpUri="https://binary.pensoft.net/fig/566410"pageId="0"pageNumber="145">Figs 1</figureCitation>
<figureCitationid="1E72E6E0E97DBC78D0654686D95E806A"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">, 2</figureCitation>
<figureCitationid="EFA4B421A52B3E40E9C9E416B14FFC41"captionStart="Figure 12"captionStartId="F12"captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure12"httpUri="https://binary.pensoft.net/fig/566421"pageId="0"pageNumber="145">, 12A-C</figureCitation>
<figureCitationid="689C96221C142D6038FD02928FD2A0DE"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure13"httpUri="https://binary.pensoft.net/fig/566422"pageId="0"pageNumber="145">, 13A</figureCitation>
<figureCitationid="0C06CB26A93A64C7C6BA7980CE4D8B3C"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure14"httpUri="https://binary.pensoft.net/fig/566423"pageId="0"pageNumber="145">, 14A,F</figureCitation>
<bibRefCitationid="2D9A557A94D89137B71C659365D1CCCF"DOI="https://doi.org/10.47640/978-5-00105-586-0_2020_292"author="Lantsov, VI"journalOrPublisher="Fauna of the Stavropol region"pageId="0"pageNumber="145"refId="B7"refString="Lantsov, VI, 2020. Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24-29, 2020. Proceedings of conference / Responsible editors: O.G.Ovchinnikova, I.V.Shamshev. St.Petersburg: Russian Entomological Society, LEMA, 292-296. https://doi.org/10.47640/978-5-00105-586-0_2020_292"title="Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24 - 29, 2020. Proceedings of conference / Responsible editors: O. G. Ovchinnikova, I. V. Shamshev. St. Petersburg: Russian Entomological Society, LEMA, 292 - 296."url="https://doi.org/10.47640/978-5-00105-586-0_2020_292"year="2020">Lantsov 2020</bibRefCitation>
<bibRefCitationid="C00EE5EA3D3DAAF0461D3F0F54A1E7A1"author="Oosterbroek, P"journalOrPublisher="Fauna of the Stavropol region"pageId="0"pageNumber="145"publicationUrl="http://ccw.naturalis.nl/detail.php"refId="B8"refString="Oosterbroek, P, 2021. Catalogue of the Craneflies of the World. http://ccw.naturalis.nl/detail.php"title="Catalogue of the Craneflies of the World."url="http://ccw.naturalis.nl/detail.php"year="2021">Oosterbroek 2021</bibRefCitation>
<materialsCitationid="01B46CDDDA165BCC3D4FB87E7AD98BF0"collectingDate="1932-05-06"collectingDateMax="2018-05-10"collectingDateMin="1932-05-06"collectorName="B. Rohdendorf, V. Lantsov, V. Pilipenko"country="Russia"county="Krasnodarskiy"determinerName="Ried"elevation="54"location="Pontica"municipality="Sotshi prov."specimenCount="37"specimenCount-female="16"specimenCount-male="21">
<materialsCitationid="3649214820E1FC95731B9423BC09711C"collectingDate="1909-06-29"collectingDateMax="2016-07-05"collectingDateMin="1909-06-29"collectorName="Mlokosevich, V. Lantsov"country="Russia"county="Tlyaratinsky district"determinerName="Ried"elevation="1792"location="river Dzhurmut"municipality="Salda"specimenCount="11"specimenCount-female="3"specimenCount-male="8"stateProvince="Dagestan">
<paragraphid="95609981A3F096E4E722DA7E25C93141"pageId="0"pageNumber="145">Dark stripes medially on lighter background extending entire length of scutum. Inner gonostylus posteriorly with wedge-shaped projection directed anteriorly. Tergite 9 at apex with three rounded notches, middle one usually deepest and widest.</paragraph>
<figureCitationid="F7254C2631217FFDAA3E9CAD60A58083"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)."figureDoi="10.3897/zookeys.1048.67564.figure1"httpUri="https://binary.pensoft.net/fig/566410"pageId="0"pageNumber="145">1A, B</figureCitation>
<figureCitationid="131335BC300C72F80A8352AEC6FAD2B5"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)."figureDoi="10.3897/zookeys.1048.67564.figure1"httpUri="https://binary.pensoft.net/fig/566410"pageId="0"pageNumber="145">1B</figureCitation>
<figureCitationid="D0041866AED40C923DA756AE913EB6D6"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure13"httpUri="https://binary.pensoft.net/fig/566422"pageId="0"pageNumber="145">13A</figureCitation>
). Rostrum whitish dorsally, flanks rusty or brownish with whitish bristles; nasus distinct, with long, whitish yellow setae; palpus grey. Frons with rows of brown setae along inner edge of eye. Vertex with faint dark median line, posteriorly with dark grey striation, with group of small rusty setae arranged fan-like on either side of glabrous medial area. Genae rusty; wide dark median stripe ventrally on rostrum, continuing along gula midline. Pharyngeal foramen framed with long, rusty hair-like setae. Postgenae light grey.
<figureCitationid="93C4938AEB3B03997DB13FC8A796BEB1"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)."figureDoi="10.3897/zookeys.1048.67564.figure1"httpUri="https://binary.pensoft.net/fig/566410"pageId="0"pageNumber="145">1B</figureCitation>
) 13-segmented, if bent backwards just reaching wing base. Scape light brown or grey, depending on angle of rotation to light source, with slightly pronounced silvery pruinescence and brown horizontal striations. Pedicel yellow, sometimes brownish at base. Flagellomeres brownish, thickened at base, verticils subequal in length to corresponding flagellomeres.
<figureCitationid="98030BB52493A65E421A35D4361938D4"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)."figureDoi="10.3897/zookeys.1048.67564.figure1"httpUri="https://binary.pensoft.net/fig/566410"pageId="0"pageNumber="145">1B</figureCitation>
). Grey or blue-grey. Pronotum with dark grey median stripe. Katepisternum, anepisternum and meron grey, glabrous. Scutum with four longitudinal wide brownish stripes on lighter background; medial stripes extending length of scutum. Scutal lobes, scutellum and mediotergite grey with light bristles in dark brown sockets.
<figureCitationid="DAB0AE6769656783B2B62BA894CA52CB"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)."figureDoi="10.3897/zookeys.1048.67564.figure1"httpUri="https://binary.pensoft.net/fig/566410"pageId="0"pageNumber="145">1A</figureCitation>
<figureCitationid="07C01D93BF4479D10473F8F9580A2586"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)."figureDoi="10.3897/zookeys.1048.67564.figure1"httpUri="https://binary.pensoft.net/fig/566410"pageId="0"pageNumber="145">1A</figureCitation>
with coxae light grey with long light hairs. Coxae may appear darker when specimen is rotated. Trochanters yellow. Femora yellow at base, rest of femora, tibiae, and tarsi dark brown.
<figureCitationid="66384AB41CDAFD0D1702AB81EE339F07"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)."figureDoi="10.3897/zookeys.1048.67564.figure1"httpUri="https://binary.pensoft.net/fig/566410"pageId="0"pageNumber="145">1A</figureCitation>
<figureCitationid="D158E7B1ABDCF139A4BA625007649E8C"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2A</figureCitation>
). Grey or dark grey; tergites with dark lateral stripe narrowly interrupted with whitish at posterior margins, lateral margins with wider whitish edge.
adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8-paired appendages of sternite 8; as 9-apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s8-sternite 8; s9-sternite 9; t9-tergite 9. Scale bar: 1 mm.
<figureCitationid="F143AAFE19D6D48116527A757EDE5F34"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2</figureCitation>
<figureCitationid="0C698594C08317D33B664EA305BD8166"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure14"httpUri="https://binary.pensoft.net/fig/566423"pageId="0"pageNumber="145">14A, F</figureCitation>
<figureCitationid="13C5F868CFABD58E05E7C38AAF308642"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2B, C</figureCitation>
<figureCitationid="73DCA65EAC7E789D63125FC993CAD821"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2C, G</figureCitation>
<figureCitationid="CEDEBBFEB3267EDA8FDC45D04E4E8586"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2C, E, F</figureCitation>
<figureCitationid="68BD15B0A23C06B665A9B23AC3E513AE"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure14"httpUri="https://binary.pensoft.net/fig/566423"pageId="0"pageNumber="145">14A</figureCitation>
) transverse, lateral corner broadly rounded, at apex with three rounded notches, middle one usually deepest and widest, bounded laterally by two large dentate projections with blunt apices, clearly visible when viewed from above (Figs
<figureCitationid="F1F63223B7DF593BD71DC5FA9BFFC0BE"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2F</figureCitation>
<figureCitationid="6167167AAC9B2DBFB18323BEE5433D7E"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure14"httpUri="https://binary.pensoft.net/fig/566423"pageId="0"pageNumber="145">14A</figureCitation>
<figureCitationid="A914D678162A6001DD4FD0CEAD4CCED5"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2E</figureCitation>
<figureCitationid="003609FDC85B2CB33981D1F8680B96DD"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2G</figureCitation>
<figureCitationid="CFB0E5D3341219600D2D2C4917B65C7A"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure14"httpUri="https://binary.pensoft.net/fig/566423"pageId="0"pageNumber="145">14F</figureCitation>
<figureCitationid="D67862F4CBAF5B74D406914E33F38A01"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2G</figureCitation>
<figureCitationid="9D60B1CC9886DDF87A5C0B1FEE026680"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure14"httpUri="https://binary.pensoft.net/fig/566423"pageId="0"pageNumber="145">14F</figureCitation>
<figureCitationid="F546016BE96F66FC639EA1E44B577EA4"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2D</figureCitation>
as long as wide, with thin rigid yellow setae along edge, not obscuring gap in middle; long, glabrous red-brown spine arising from base, bent, and intersecting medially. Paired appendage of sternite 9 (Fig.
<figureCitationid="A503105C3C2FDC8DEF531EEF585B2E8D"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2K, L</figureCitation>
<figureCitationid="8600595E006826FA8776694E0B2B9D21"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2I</figureCitation>
<figureCitationid="8895FB6D37065D6EE18AD8E4D65F5C65"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure2"httpUri="https://binary.pensoft.net/fig/566411"pageId="0"pageNumber="145">2J</figureCitation>
Body length 17-21.5 mm, wing 15.5-18 mm. Antennae with scape and pedicel brownish yellow. Legs stocky, black to brown, yellow at base, femora grey. Wings with noticeable bluish tint; whitish area (lunule) of wings barely reaching base of discal cell. Female terminalia (Fig.
<figureCitationid="C8C0ADD2C829B5B5B7C562FC289E7E0E"captionStart="Figure 12"captionStartId="F12"captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure12"httpUri="https://binary.pensoft.net/fig/566421"pageId="0"pageNumber="145">12A-C</figureCitation>
) comparatively short. Tip of cercus upcurved, length of cercus ~ 0.8 mm. Hypogynial valve only slightly longer than width at base; length of hypogynial valve 0.2 mm. Sternite 9 and furca as in Figure
<figureCitationid="0147064F8F06BF3273099D49E842DEBC"captionStart="Figure 12"captionStartId="F12"captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm."figureDoi="10.3897/zookeys.1048.67564.figure12"httpUri="https://binary.pensoft.net/fig/566421"pageId="0"pageNumber="145">12A, C</figureCitation>
<paragraphid="49B380CCB67997FCADBF477D4C5DE53D"pageId="0"pageNumber="145">Adults were collected at altitudes ranging from sea level (Black Sea coast at 35 m) to 2011 m (in Dagestan).</paragraph>
<bibRefCitationid="0F41EB0EA2A18ED2D7BF271B7A2D4ECC"DOI="https://doi.org/10.47640/978-5-00105-586-0_2020_292"author="Lantsov, VI"journalOrPublisher="Fauna of the Stavropol region"pageId="0"pageNumber="145"refId="B7"refString="Lantsov, VI, 2020. Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24-29, 2020. Proceedings of conference / Responsible editors: O.G.Ovchinnikova, I.V.Shamshev. St.Petersburg: Russian Entomological Society, LEMA, 292-296. https://doi.org/10.47640/978-5-00105-586-0_2020_292"title="Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24 - 29, 2020. Proceedings of conference / Responsible editors: O. G. Ovchinnikova, I. V. Shamshev. St. Petersburg: Russian Entomological Society, LEMA, 292 - 296."url="https://doi.org/10.47640/978-5-00105-586-0_2020_292"year="2020">Lantsov 2020</bibRefCitation>
publication, neither the holotype nor the paratypes are indicated. Presumably, they are not indicated on the material itself. Because this material turned out to be inaccessible to the authors of this article - it is not yet known whether this material has survived at all - there was no opportunity to designate lectotype and paralectotypes.
