306 lines
34 KiB
XML
306 lines
34 KiB
XML
<document id="A5AFA4E07C4C5393E5BBF0B62A3A6437" ID-DOI="10.11646/zootaxa.2390.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5305157" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629669061610" checkinUser="felipe" docAuthor="Kiel, Steffen, Campbell, Kathleen A. & Gaillard, Christian" docDate="2010" docId="C86DD84BFFEAFFBEFF79C8C8DE31FF78" docLanguage="en" docName="zt02390p048.pdf" docOrigin="Zootaxa 2390 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2390.1.2" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Bathymodiolus palmarensis Kiel & Campbell & Gaillard 2010, sp. nov." docType="treatment" docVersion="5" lastPageNumber="40" masterDocId="3454A033FFE7FFB0FFEEC85FDF23FF84" masterDocTitle="New and little known mollusks from ancient chemosynthetic environments" masterLastPageNumber="48" masterPageNumber="26" pageNumber="39" updateTime="1699093293136" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="C8CEDE8EF6C6BA359A6B6FE9E4A52E17" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9C78A9D755AACB6F7C6F46388FF33F11">
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<mods:title id="77631FCC61C1C6ED0A6313EAB84071C8">New and little known mollusks from ancient chemosynthetic environments</mods:title>
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</mods:titleInfo>
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<mods:name id="941519488931371D84C2449A7D05E887" type="personal">
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<mods:role id="7A1888061F6E670AF74773C31D7AD2D6">
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<mods:roleTerm id="CFF6CAF5F2B994EF0808B242E04A62A5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="02A8CBD5BA10745FBBB01900ED1F9F11">Kiel, Steffen</mods:namePart>
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<mods:affiliation id="3EAEB2649F9F5B0B2193B71A43B5CEE8">Institut für Geowissenschaften, Christian-Albrechts-Universität, Ludewig-Meyn-Str. 10, 24118 Kiel, Germany.</mods:affiliation>
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</mods:name>
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<mods:name id="EDEB7F87D5100387C5679427F96B8966" type="personal">
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<mods:role id="4CF9F14EB0542988D7F431904B9D17D7">
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<mods:roleTerm id="B19CA6BC48C663883AFCB001E231B718">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CB7A98A7EDB385D0002A8072D3137420">Campbell, Kathleen A.</mods:namePart>
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<mods:affiliation id="7F4CA32718AA8B3A75D3524F94A7D305">School of Geography, Geology and Environmental Science, University of Auckland, Private Bag 92019, Auckland 1142, New Zealand. E-mail: ka. campbell @ auckland. ac. nz & Department of Invertebrate Zoology and Geology, California Academy of Sciences, Golden Gate Park, San Francisco, California 94118, USA</mods:affiliation>
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</mods:name>
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<mods:name id="FC260956AF36784D8FC9C2B512532C39" type="personal">
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<mods:roleTerm id="1C702587EEF925550312CE167C414B89">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D8517344978AECE8B26191F9E5F8428C">Gaillard, Christian</mods:namePart>
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<mods:affiliation id="011BC3A4A894BA57BBFF5DAA4004260D">Université de Lyon, Lyon 1, UMR CNRS 5125 Paléoenvironnements et Paléobiosphère, 2 rue Raphaël Dubois, 69622 Villeurbanne Cedex, France. E-mail: Christian. Gaillard @ univ-lyon 1. fr</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="797E8D7240ED68FD71BB37568617FD49">text</mods:typeOfResource>
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<mods:title id="6A41E9F0EBFB648F745456ED6CF34135">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A936DC08F32DC50653D5C46A1C787A2A">
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<mods:date id="635119FBE597A89B4D2936D1AD0CDAFB">2010</mods:date>
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<mods:number id="8D8E5D1C52296A897B41FBA5C48ACD28">2010-03-05</mods:number>
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<mods:detail id="B5490E8A40A89366019140AC949AD29D" type="volume">
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<mods:number id="7DA16B126C6146769660FF7FE45BEFAD">2390</mods:number>
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</mods:detail>
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<mods:detail id="C696998048FC09A6E8BD4EAFDD2CC6B6" type="issue">
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<mods:number id="92593C30A4E652515A58E6C3D0D36BE1">1</mods:number>
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<mods:start id="5876A3CC38F414E9E0C671E9CBD08DC5">26</mods:start>
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<mods:end id="6C725E944A171320E40C8FFEEBC7DFC5">48</mods:end>
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<mods:url id="FFFE2D8ED95D571936E32458CF29E0D7">https://biotaxa.org/Zootaxa/article/view/zootaxa.2390.1.2</mods:url>
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</mods:location>
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<mods:classification id="50F99E41B050370FD6566D0166D9783E">journal article</mods:classification>
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<mods:identifier id="E063E3DA01977A92A2804D2784718860" type="DOI">10.11646/zootaxa.2390.1.2</mods:identifier>
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<mods:identifier id="6A3AFA0A8E7CE9C140708EBD3098A6D8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="921B7D5D2DAF30D59D2BF65AB156CEA4" type="Zenodo-Dep">5305157</mods:identifier>
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<treatment id="C86DD84BFFEAFFBEFF79C8C8DE31FF78" ID-DOI="http://doi.org/10.5281/zenodo.5312898" ID-Zenodo-Dep="5312898" LSID="urn:lsid:plazi:treatment:C86DD84BFFEAFFBEFF79C8C8DE31FF78" httpUri="http://treatment.plazi.org/id/C86DD84BFFEAFFBEFF79C8C8DE31FF78" lastPageId="14" lastPageNumber="40" pageId="13" pageNumber="39">
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<subSubSection id="08DE3AD6FFEAFFBDFF79C8C8DDC2FF35" box="[151,737,151,177]" pageId="13" pageNumber="39" type="nomenclature">
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<paragraph id="407B695DFFEAFFBDFF79C8C8DDC2FF35" blockId="13.[151,737,151,212]" box="[151,737,151,177]" pageId="13" pageNumber="39">
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<heading id="1B33DE31FFEAFFBDFF79C8C8DDC2FF35" bold="true" box="[151,737,151,177]" fontSize="11" level="1" pageId="13" pageNumber="39" reason="1">
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<emphasis id="72B0B54FFFEAFFBDFF79C8C8DDC2FF35" bold="true" box="[151,737,151,177]" pageId="13" pageNumber="39">
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<taxonomicName id="87C412DEFFEAFFBDFF79C8C8DD5FFF35" ID-CoL="KXBM" authority="Kiel & Campbell & Gaillard, 2010" authorityName="Kiel & Campbell & Gaillard" authorityYear="2010" box="[151,636,151,177]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="palmarensis" status="sp. nov.">
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<emphasis id="72B0B54FFFEAFFBDFF79C8C8DE6EFF35" bold="true" box="[151,333,151,177]" italics="true" pageId="13" pageNumber="39">Bathymodiolus</emphasis>
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(sensu lato)
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<emphasis id="72B0B54FFFEAFFBDFE04C8C8DD5FFF35" bold="true" box="[490,636,151,177]" italics="true" pageId="13" pageNumber="39">palmarensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="69830834FFEAFFBDFD6AC8C8DDC2FF35" box="[644,737,151,177]" pageId="13" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="08DE3AD6FFEAFFBDFF79C8E5DC79FEB8" pageId="13" pageNumber="39" type="description">
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<paragraph id="407B695DFFEAFFBDFF79C8E5DFCBFF50" blockId="13.[151,737,151,212]" box="[151,232,186,212]" pageId="13" pageNumber="39">
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(
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<figureCitation id="D8FF75D8FFEAFFBDFF71C8E5DFFDFF50" box="[159,222,186,212]" captionStart="FIGURE 8" captionStartId="13.[151,254,1893,1917]" captionTargetBox="[151,1436,1525,1872]" captionTargetId="figure-496@13.[151,1436,1525,1872]" captionTargetPageId="13" captionText="FIGURE 8. Bathymodiolus (s.l.) palmarensis n. sp. from a presumably Oligocene seep carbonate in northern Colombia. A, B. Holotype (USNM 533981), an elongate specimen showing shell remains on the anterior end. C. Smaller specimen (USNM 533982)." figureDoi="http://doi.org/10.5281/zenodo.5305179" httpUri="https://zenodo.org/record/5305179/files/figure.png" pageId="13" pageNumber="39">Fig 8</figureCitation>
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)
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</paragraph>
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<paragraph id="407B695DFFEAFFBDFF79C95ADD58FE98" blockId="13.[151,858,261,316]" box="[151,635,261,284]" pageId="13" pageNumber="39">
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2005 Bathymodiolin;
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<bibRefCitation id="245514ACFFEAFFBDFE6FC95ADD07FE98" author="Gill F. L. & Harding I. C. & Little C. T. S. & Todd J. A." box="[385,548,261,284]" pageId="13" pageNumber="39" pagination="191 - 209" refId="ref12112" refString="Gill F. L., Harding I. C., Little C. T. S. & Todd J. A. (2005) Palaeogene and Neogene cold seep communities in Barbados, Trinidad and Venezuela: An overview. Palaeogeography, Palaeoclimatology, Palaeoecology, 227, 191 - 209." type="journal article" year="2005">
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Gill
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<emphasis id="72B0B54FFFEAFFBDFE5FC95ADEFCFE98" box="[433,479,261,284]" italics="true" pageId="13" pageNumber="39">et al</emphasis>
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. 2005
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</bibRefCitation>
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, fig.
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<specimenCount id="56C2A2D4FFEAFFBDFDB4C95ADD58FE98" box="[602,635,261,284]" pageId="13" pageNumber="39" type="generic">9F.</specimenCount>
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</paragraph>
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<paragraph id="407B695DFFEAFFBDFF79C97ADC79FEB8" blockId="13.[151,858,261,316]" box="[151,858,292,316]" pageId="13" pageNumber="39">
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2007
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<taxonomicName id="87C412DEFFEAFFBDFF3DC97ADE50FEB8" authorityName="Kenk and Wilson" authorityYear="1985" box="[211,371,293,316]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="genus">
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<emphasis id="72B0B54FFFEAFFBDFF3DC97ADE50FEB8" box="[211,371,293,316]" italics="true" pageId="13" pageNumber="39">Bathymodiolus</emphasis>
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</taxonomicName>
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?
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<emphasis id="72B0B54FFFEAFFBDFE6BC97BDEE2FEB8" bold="true" box="[389,449,292,316]" pageId="13" pageNumber="39">n. sp.</emphasis>
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;
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<bibRefCitation id="245514ACFFEAFFBDFE21C97ADDFCFEB8" author="Kiel S. & Peckmann J." box="[463,735,293,316]" pageId="13" pageNumber="39" pagination="345 - 357" refId="ref13160" refString="Kiel S. & Peckmann J. (2007) Chemosymbiotic bivalves and stable carbon isotopes indicate hydrocarbon seepage at four unusual Cenozoic fossil localities. Lethaia, 40, 345 - 357." type="journal article" year="2007">Kiel and Peckmann 2007</bibRefCitation>
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, figs. 5E–I.
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</paragraph>
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</subSubSection>
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<subSubSection id="08DE3AD6FFEAFFBDFF79C935DE77FE2E" pageId="13" pageNumber="39" type="diagnosis">
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<paragraph id="407B695DFFEAFFBDFF79C935DE77FE2E" blockId="13.[151,1436,362,1471]" pageId="13" pageNumber="39">
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<emphasis id="72B0B54FFFEAFFBDFF79C935DE3EFE00" bold="true" box="[151,285,362,388]" pageId="13" pageNumber="39">Diagnosis:</emphasis>
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Bathymodioline shell with subterminal beak, straight dorsal and ventral margins, and pointed anterior margin.
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</paragraph>
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</subSubSection>
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<subSubSection id="08DE3AD6FFEAFFBDFF2AC9EADE58FD0E" pageId="13" pageNumber="39" type="materials_examined">
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<paragraph id="407B695DFFEAFFBDFF2AC9EADD21FE4B" blockId="13.[151,1436,362,1471]" box="[196,514,437,463]" pageId="13" pageNumber="39">
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<materialsCitation id="F0AC6300FFEAFFBDFF2AC9EADD21FE4B" box="[196,514,437,463]" collectionCode="USNM" pageId="13" pageNumber="39" specimenCode="USNM 533981" specimenCount="1" typeStatus="holotype">
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<emphasis id="72B0B54FFFEAFFBDFF2AC9EADE1CFE4B" bold="true" box="[196,319,437,463]" pageId="13" pageNumber="39">
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<typeStatus id="9F7FD7FFFFEAFFBDFF2AC9EADE19FE4B" box="[196,314,437,463]" pageId="13" pageNumber="39" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCode id="1062C126FFEAFFBDFEA9C9EADEDDFE4B" box="[327,510,437,463]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="39" type="Museum">USNM 533981</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="407B695DFFEAFFBDFF2AC985DD4CFE70" blockId="13.[151,1436,362,1471]" box="[196,623,474,500]" pageId="13" pageNumber="39">
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<materialsCitation id="F0AC6300FFEAFFBDFF2AC985DD4FFE70" box="[196,620,474,500]" collectionCode="USNM" pageId="13" pageNumber="39" specimenCode="USNM 533982-553984" specimenCount="1" typeStatus="paratype">
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<emphasis id="72B0B54FFFEAFFBDFF2AC985DE69FE70" bold="true" box="[196,330,474,500]" pageId="13" pageNumber="39">
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<typeStatus id="9F7FD7FFFFEAFFBDFF2AC985DE65FE70" box="[196,326,474,500]" pageId="13" pageNumber="39" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCode id="1062C126FFEAFFBDFEBCC985DD4FFE70" box="[338,620,474,500]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="39" type="Museum">USNM 533982-553984</specimenCode>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="407B695DFFEAFFBDFF2AC9A0DE58FD0E" blockId="13.[151,1436,362,1471]" pageId="13" pageNumber="39">
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<emphasis id="72B0B54FFFEAFFBDFF2AC9A0DE5AFD9D" bold="true" box="[196,377,511,537]" pageId="13" pageNumber="39">Type locality:</emphasis>
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<materialsCitation id="F0AC6300FFEAFFBDFE6CCA5FDE54FD0E" country="Colombia" county="Arroyo Piadras" latitude="10.616667" location="Oligocene" longLatPrecision="1297" longitude="-75.1" municipality="The town" pageId="13" pageNumber="39" specimenCount="1" stateProvince="Atlantico">
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USGS locality 11253,
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<collectingRegion id="8200A7BFFFEAFFBDFD70CA5FDC36FD9E" box="[670,789,512,538]" country="Colombia" name="Atlantico" pageId="13" pageNumber="39">Atlantico</collectingRegion>
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,
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<collectingCountry id="38D329CDFFEAFFBDFCCDCA5FDC83FD9E" box="[803,928,512,538]" name="Colombia" pageId="13" pageNumber="39">Colombia</collectingCountry>
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; considered to be of
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<location id="451B3F86FFEAFFBDFB46CA5FDA04FD9E" LSID="urn:lsid:plazi:treatment:C86DD84BFFEAFFBEFF79C8C8DE31FF78:451B3F86FFEAFFBDFB46CA5FDA04FD9E" box="[1192,1319,512,538]" country="Colombia" county="Arroyo Piadras" latitude="10.616667" longLatPrecision="1297" longitude="-75.1" municipality="The town" name="Oligocene" pageId="13" pageNumber="39" stateProvince="Atlantico">Oligocene</location>
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age.
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<location id="451B3F86FFEAFFBDFA83CA5FDABFFD9E" LSID="urn:lsid:plazi:treatment:C86DD84BFFEAFFBEFF79C8C8DE31FF78:451B3F86FFEAFFBDFA83CA5FDABFFD9E" box="[1389,1436,512,538]" country="Colombia" county="Arroyo Piadras" latitude="10.616667" longLatPrecision="1297" longitude="-75.1" municipality="The town" name="The" pageId="13" pageNumber="39" stateProvince="Atlantico">The</location>
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original locality description is “¼ mile N of junction Arroyo Piadras Palmar and Palmar-Molinera-Road,
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<collectingRegion id="8200A7BFFFEAFFBDFF79CA15DE2FFDE0" box="[151,268,586,612]" country="Colombia" name="Atlantico" pageId="13" pageNumber="39">Atlantico</collectingRegion>
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,
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<collectingCountry id="38D329CDFFEAFFBDFEF4CA15DEB6FDE0" box="[282,405,586,612]" name="Colombia" pageId="13" pageNumber="39">Colombia</collectingCountry>
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” and its exact location proved difficult to pin down.
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<collectingMunicipality id="A01FF327FFEAFFBDFBC9CA15DBB8FDE0" box="[1063,1179,586,612]" pageId="13" pageNumber="39">The town</collectingMunicipality>
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<collectingCounty id="A91A11D1FFEAFFBDFB4BCA15DA43FDE0" box="[1189,1376,586,612]" pageId="13" pageNumber="39">Arroyo Piadras</collectingCounty>
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is at
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<geoCoordinate id="25F00F9AFFEAFFBDFF79CA2FDFDDFD0E" box="[151,254,624,650]" degrees="10" direction="north" minutes="37" orientation="latitude" pageId="13" pageNumber="39" precision="925" value="10.616667">10°37’N</geoCoordinate>
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,
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<geoCoordinate id="25F00F9AFFEAFFBDFEE5CA2FDE54FD0E" box="[267,375,624,650]" degrees="75" direction="west" minutes="06" orientation="longitude" pageId="13" pageNumber="39" precision="925" value="-75.1">75°06’W</geoCoordinate>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="08DE3AD6FFEAFFBDFF2ACACADDE4FCC0" pageId="13" pageNumber="39" type="description">
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<paragraph id="407B695DFFEAFFBDFF2ACACADDE4FCC0" blockId="13.[151,1436,362,1471]" pageId="13" pageNumber="39">
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<emphasis id="72B0B54FFFEAFFBDFF2ACACADE45FD2B" bold="true" box="[196,358,661,687]" pageId="13" pageNumber="39">Description:</emphasis>
|
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Elongate modioliform shell, slightly inflated but with strong umbonal ridge, beaks in subterminal position; length/height-ratio 1:
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<quantity id="873CC4B8FFEAFFBDFD66CAE5DD9AFD50" box="[648,697,698,724]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="13" pageNumber="39" unit="in" value="2.0">2 in</quantity>
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smallest specimen, 1:2.5 in largest; surface with strong growth increments; dorsal margin straight, angular transition to posterodorsal margin at about two-thirds of total shell length; posterior margin broadly rounded; ventral margin straight; anterior margin pointed but rounded; hinge unknown, interior of shell with fine radial lines.
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</paragraph>
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</subSubSection>
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<subSubSection id="08DE3AD6FFEAFFBDFF2ACB10DD14FA3B" pageId="13" pageNumber="39" type="discussion">
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<paragraph id="407B695DFFEAFFBDFF2ACB10DD14FA3B" blockId="13.[151,1436,362,1471]" pageId="13" pageNumber="39">
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<emphasis id="72B0B54FFFEAFFBDFF2ACB10DE60FCED" bold="true" box="[196,323,847,873]" pageId="13" pageNumber="39">Remarks:</emphasis>
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Many extant species of
|
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<taxonomicName id="87C412DEFFEAFFBDFD9ECB0FDC05FCED" authorityName="Kenk and Wilson" authorityYear="1985" box="[624,806,848,873]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="genus">
|
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<emphasis id="72B0B54FFFEAFFBDFD9ECB0FDC05FCED" box="[624,806,848,873]" italics="true" pageId="13" pageNumber="39">Bathymodiolus</emphasis>
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</taxonomicName>
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differ from
|
||
<taxonomicName id="87C412DEFFEAFFBDFC50CB0FDB95FCED" authority="Kiel & Campbell & Gaillard, 2010" authorityName="Kiel & Campbell & Gaillard" authorityYear="2010" box="[958,1206,848,874]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="palmarensis" status="sp. nov.">
|
||
<emphasis id="72B0B54FFFEAFFBDFC50CB0FDCF3FCED" box="[958,976,848,873]" italics="true" pageId="13" pageNumber="39">B</emphasis>
|
||
. (s.l.)
|
||
<emphasis id="72B0B54FFFEAFFBDFBCDCB0FDB95FCED" box="[1059,1206,848,873]" italics="true" pageId="13" pageNumber="39">palmarensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72B0B54FFFEAFFBDFB51CB10DA27FCED" bold="true" box="[1215,1284,847,873]" pageId="13" pageNumber="39">
|
||
<taxonomicNameLabel id="69830834FFEAFFBDFB51CB10DA27FCED" box="[1215,1284,847,873]" pageId="13" pageNumber="39" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by having a concave ventral margin and by having the beak in a less anterior position. Quite similar to
|
||
<taxonomicName id="87C412DEFFEAFFBDFB5FCB2ADABFFC0A" authorityName="Kiel & Campbell & Gaillard" authorityYear="2010" box="[1201,1436,885,911]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="palmarensis">
|
||
<emphasis id="72B0B54FFFEAFFBDFB5FCB2ADBE0FC0A" box="[1201,1219,885,910]" italics="true" pageId="13" pageNumber="39">B</emphasis>
|
||
. (s.l.)
|
||
<emphasis id="72B0B54FFFEAFFBDFAE0CB2ADABFFC0A" box="[1294,1436,885,910]" italics="true" pageId="13" pageNumber="39">palmarensis</emphasis>
|
||
</taxonomicName>
|
||
regarding these two characters is
|
||
<taxonomicName id="87C412DEFFEAFFBDFDC3CBC5DB5AFC30" authority="von Cosel, Comtet & Krylova, 1999" authorityName="von Cosel, Comtet & Krylova" authorityYear="1999" box="[557,1145,922,948]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="azoricus">
|
||
<emphasis id="72B0B54FFFEAFFBDFDC3CBC5DD97FC37" box="[557,692,922,947]" italics="true" pageId="13" pageNumber="39">B. azoricus</emphasis>
|
||
von
|
||
<bibRefCitation id="245514ACFFEAFFBDFD1DCBC5DB5AFC30" author="Cosel R. & Comtet T. & Krylova E. M." box="[755,1145,922,948]" pageId="13" pageNumber="39" pagination="218 - 248" refId="ref11657" refString="Cosel R. v., Comtet T. & Krylova E. M. (1999) Bathymodiolus (Bivalvia: Mytilidae) from hydrothermal vents on the Azores Triple Junction and the Logatchev hydrothermal field, Mid-Atlantic Ridge. The Veliger, 42, 218 - 248." type="journal article" year="1999">Cosel, Comtet & Krylova, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the Azores Triple Junction on the Mid-Atlantic Ridge, but it differs from that extant species by having a proportionally narrower anterior margin and by a less well-developed umbonal ridge. The same is true for
|
||
<taxonomicName id="87C412DEFFEAFFBDFB7CCBBADD02FBA0" authority="Gustafson, Turner, Lutz & Vrijenhoek, 1998" authorityName="Gustafson, Turner, Lutz & Vrijenhoek" authorityYear="1998" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="brooksi">
|
||
<emphasis id="72B0B54FFFEAFFBDFB7CCBBADA2CFC7A" box="[1170,1295,997,1022]" italics="true" pageId="13" pageNumber="39">B. brooksi</emphasis>
|
||
<bibRefCitation id="245514ACFFEAFFBDFAF6CBBADD02FBA0" author="Gustafson R. G. & Turner R. D. & Lutz R. A. & Vrijenhoek R. C." pageId="13" pageNumber="39" pagination="63 - 112" refId="ref12297" refString="Gustafson R. G., Turner R. D., Lutz R. A. & Vrijenhoek R. C. (1998) A new genus and five new species of mussels (Bivalvia: Mytilidae) from deep-sea sulfide / hydrocarbon seeps in the Gulf of Mexico. Malacologia, 40, 63 - 112." type="journal article" year="1998">Gustafson, Turner, Lutz & Vrijenhoek, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
. The East-Atlantic species
|
||
<taxonomicName id="87C412DEFFEAFFBDFC8ACC55DBD2FBA0" authority="von Cosel, 2002" authorityName="von Cosel" authorityYear="2002" box="[868,1265,1034,1060]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="mauretanicus">
|
||
<emphasis id="72B0B54FFFEAFFBDFC8ACC55DB06FBA7" box="[868,1061,1034,1059]" italics="true" pageId="13" pageNumber="39">B. mauretanicus</emphasis>
|
||
von Cosel, 2002
|
||
</taxonomicName>
|
||
, has the beaks in an even more anterior position than
|
||
<taxonomicName id="87C412DEFFEAFFBDFD84CC6FDC7FFBCD" authorityName="Kiel & Campbell & Gaillard" authorityYear="2010" box="[618,860,1072,1098]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="palmarensis">
|
||
<emphasis id="72B0B54FFFEAFFBDFD84CC6FDD5FFBCD" box="[618,636,1072,1097]" italics="true" pageId="13" pageNumber="39">B</emphasis>
|
||
. (s.l.)
|
||
<emphasis id="72B0B54FFFEAFFBDFD22CC6FDC7FFBCD" box="[716,860,1072,1097]" italics="true" pageId="13" pageNumber="39">palmarensis</emphasis>
|
||
</taxonomicName>
|
||
, but is shorter (has a lower height/length ratio), lacks internal radial ridges or striations, and usually has a concave ventral margin. Another species with the beak in an almost terminal position and a lower height/length ratio than
|
||
<taxonomicName id="87C412DEFFEAFFBDFC33CC24DBEBFB10" authorityName="Kiel & Campbell & Gaillard" authorityYear="2010" box="[989,1224,1146,1172]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="palmarensis">
|
||
<emphasis id="72B0B54FFFEAFFBDFC33CC24DCCCFB10" box="[989,1007,1147,1172]" italics="true" pageId="13" pageNumber="39">B</emphasis>
|
||
. (s.l.)
|
||
<emphasis id="72B0B54FFFEAFFBDFBD4CC24DBEBFB10" box="[1082,1224,1147,1172]" italics="true" pageId="13" pageNumber="39">palmarensis</emphasis>
|
||
</taxonomicName>
|
||
is the Japanese
|
||
<taxonomicName id="87C412DEFFEAFFBDFA6DCC24DD42FB3E" authority="Hashimoto & Okutani, 1994" authorityName="Hashimoto & Okutani" authorityYear="1994" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="platifrons">
|
||
<emphasis id="72B0B54FFFEAFFBDFA6DCC24DE2BFB3D" italics="true" pageId="13" pageNumber="39">B. platifrons</emphasis>
|
||
<bibRefCitation id="245514ACFFEAFFBDFEE1CCFFDD42FB3E" author="Hashimoto J. & Okutani T." box="[271,609,1184,1210]" pageId="13" pageNumber="39" pagination="61 - 83" refId="ref12357" refString="Hashimoto J. & Okutani T. (1994) Four new mytilid mussels associated with deepsea chemysynthetic communitites around Japan. Venus, 53, 61 - 83." type="journal article" year="1994">Hashimoto & Okutani, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
. The slightly older fossil species
|
||
<taxonomicName id="87C412DEFFEAFFBDFC07CCFFDEA1FB5B" authority="(Squires and Goedert, 1991)" baseAuthorityName="Squires and Goedert" baseAuthorityYear="1991" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="willapaensis">
|
||
<emphasis id="72B0B54FFFEAFFBDFC07CCFFDA10FB3D" box="[1001,1331,1184,1209]" italics="true" pageId="13" pageNumber="39">Bathymodiolus willapaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="245514ACFFEAFFBDFAACCCFFDE5AFB5B" author="Squires R. L. & Goedert J. L." pageId="13" pageNumber="39" pagination="412 - 416" refId="ref14493" refString="Squires R. L. & Goedert J. L. (1991) New Late Eocene Mollusks from localized limestone deposits formed by subductionrelated methane seeps, southwestern Washington. Journal of Paleontology, 65, 412 - 416." type="journal article" year="1991">Squires and Goedert,1991</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the Eocene and Oligocene of
|
||
<collectingRegion id="8200A7BFFFEAFFBDFCABCC9ADB00FB5B" box="[837,1059,1221,1247]" country="United States of America" name="Washington" pageId="13" pageNumber="39">Washington State</collectingRegion>
|
||
,
|
||
<collectingCountry id="38D329CDFFEAFFBDFBDCCC9ADB51FB5B" box="[1074,1138,1221,1247]" name="United States of America" pageId="13" pageNumber="39">USA</collectingCountry>
|
||
, also has the beaks in a subterminal position. It differs from
|
||
<taxonomicName id="87C412DEFFEAFFBDFDB2CCB4DC75FA80" authorityName="Kiel & Campbell & Gaillard" authorityYear="2010" box="[604,854,1258,1284]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="palmarensis">
|
||
<emphasis id="72B0B54FFFEAFFBDFDB2CCB4DD4DFA80" box="[604,622,1259,1284]" italics="true" pageId="13" pageNumber="39">B</emphasis>
|
||
. (s.l.)
|
||
<emphasis id="72B0B54FFFEAFFBDFD2CCCB4DC75FA80" box="[706,854,1259,1284]" italics="true" pageId="13" pageNumber="39">palmarensis</emphasis>
|
||
</taxonomicName>
|
||
by having a slightly convex and shorter dorsal margin, which occupies only about half of the distance between beak and posterior end, whereas in
|
||
<taxonomicName id="87C412DEFFEAFFBDFAAACD4FDE05FACA" authorityName="Kiel & Campbell & Gaillard" authorityYear="2010" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="palmarensis">
|
||
<emphasis id="72B0B54FFFEAFFBDFAAACD4FDA75FAAD" box="[1348,1366,1296,1321]" italics="true" pageId="13" pageNumber="39">B</emphasis>
|
||
. (s.l.)
|
||
<emphasis id="72B0B54FFFEAFFBDFF79CD6ADE05FACA" box="[151,294,1333,1358]" italics="true" pageId="13" pageNumber="39">palmarensis</emphasis>
|
||
</taxonomicName>
|
||
the dorsal margin is straight and occupies two-thirds of the distance between beak and posterior end. There are several genetically distinct clades among the species traditionally placed in
|
||
<taxonomicName id="87C412DEFFEAFFBDFB06CD04DABFFAF0" authorityName="Kenk and Wilson" authorityYear="1985" box="[1256,1436,1371,1396]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="genus">
|
||
<emphasis id="72B0B54FFFEAFFBDFB06CD04DABFFAF0" box="[1256,1436,1371,1396]" italics="true" pageId="13" pageNumber="39">Bathymodiolus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="245514ACFFEAFFBDFF71CDDFDEB8FA1E" author="Miyazaki J. - I. & Shintaku M. & Kyuno A. & Fujiwara Y." box="[159,411,1408,1434]" pageId="13" pageNumber="39" pagination="527 - 535" refId="ref13745" refString="Miyazaki J. - I., Shintaku M., Kyuno A., Fujiwara Y. et al. (2004) Phylogenetic relationships of deep-sea mussels of the genus Bathymodiolus (Bivalvia: Mytilidae). Marine Biology, 144, 527 - 535." type="journal article" year="2004">
|
||
Miyazaki
|
||
<emphasis id="72B0B54FFFEAFFBDFEF6CDDFDE76FA1D" box="[280,341,1408,1433]" italics="true" pageId="13" pageNumber="39">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="245514ACFFEAFFBDFE46CDDFDDACFA1E" author="Iwasaki H. & Kyuno A. & Shintaku M. & Fujita Y." box="[424,655,1408,1434]" pageId="13" pageNumber="39" pagination="1111 - 1122" refId="ref12465" refString="Iwasaki H., Kyuno A., Shintaku M., Fujita Y. et al. (2006) Evolutionary relationships of deep-sea mussels inferred by mitochondrial DNA sequences. Marine Biology, 149, 1111 - 1122." type="journal article" year="2006">
|
||
Iwasaki
|
||
<emphasis id="72B0B54FFFEAFFBDFDE2CDDFDD6BFA1D" box="[524,584,1408,1433]" italics="true" pageId="13" pageNumber="39">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
; Jones
|
||
<emphasis id="72B0B54FFFEAFFBDFD0BCDDFDC01FA1D" box="[741,802,1408,1433]" italics="true" pageId="13" pageNumber="39">et al.</emphasis>
|
||
2006;
|
||
<bibRefCitation id="245514ACFFEAFFBDFC9BCDDFDB0EFA1E" author="Olu K. & Cosel R. & Hourdez S. & Carney S. L." box="[885,1069,1408,1434]" pageId="13" pageNumber="39" pagination="1890 - 1911" refId="ref14096" refString="Olu K., Cosel R. v., Hourdez S., Carney S. L. et al. (2007) Amphi-Atlantic cold-seep Bathymodiolus species complexes across the equatorial belt. Deep-Sea Research I, 54, 1890 - 1911." type="journal article" year="2007">
|
||
Olu
|
||
<emphasis id="72B0B54FFFEAFFBDFC44CDDFDCC5FA1D" box="[938,998,1408,1433]" italics="true" pageId="13" pageNumber="39">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
), hence we assign
|
||
<taxonomicName id="87C412DEFFEAFFBDFAE2CDDFDAB8FA1D" authorityName="Kiel & Campbell & Gaillard" authorityYear="2010" box="[1292,1435,1408,1433]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="palmarensis">
|
||
<emphasis id="72B0B54FFFEAFFBDFAE2CDDFDAB8FA1D" box="[1292,1435,1408,1433]" italics="true" pageId="13" pageNumber="39">palmarensis</emphasis>
|
||
</taxonomicName>
|
||
only to
|
||
<taxonomicName id="87C412DEFFEAFFBDFF1ECDFADE83FA3A" authorityName="Kenk and Wilson" authorityYear="1985" box="[240,416,1445,1470]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="genus">
|
||
<emphasis id="72B0B54FFFEAFFBDFF1ECDFADE83FA3A" box="[240,416,1445,1470]" italics="true" pageId="13" pageNumber="39">Bathymodiolus</emphasis>
|
||
</taxonomicName>
|
||
(sensu lato).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="14BB39D5FFEAFFBDFF79CF3ADE77F838" ID-DOI="http://doi.org/10.5281/zenodo.5305179" ID-Zenodo-Dep="5305179" httpUri="https://zenodo.org/record/5305179/files/figure.png" pageId="13" pageNumber="39" startId="13.[151,254,1893,1917]" targetBox="[151,1436,1525,1872]" targetPageId="13">
|
||
<paragraph id="407B695DFFEAFFBDFF79CF3ADE77F838" blockId="13.[151,1437,1893,1980]" pageId="13" pageNumber="39">
|
||
<emphasis id="72B0B54FFFEAFFBDFF79CF3ADE3AF8F9" bold="true" box="[151,281,1893,1917]" pageId="13" pageNumber="39">FIGURE 8.</emphasis>
|
||
<taxonomicName id="87C412DEFFEAFFBDFEF1CF3ADD5DF8F8" authority="Kiel & Campbell & Gaillard, 2010" authorityName="Kiel & Campbell & Gaillard" authorityYear="2010" box="[287,638,1893,1916]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="39" phylum="Mollusca" rank="species" species="palmarensis" status="sp. nov.">
|
||
<emphasis id="72B0B54FFFEAFFBDFEF1CF3ADE9CF8F8" box="[287,447,1893,1916]" italics="true" pageId="13" pageNumber="39">Bathymodiolus</emphasis>
|
||
(s.l.)
|
||
<emphasis id="72B0B54FFFEAFFBDFE13CF3ADD5DF8F8" box="[509,638,1893,1916]" italics="true" pageId="13" pageNumber="39">palmarensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72B0B54FFFEAFFBDFD6BCF3ADDE2F8F9" bold="true" box="[645,705,1893,1917]" pageId="13" pageNumber="39">
|
||
<taxonomicNameLabel id="69830834FFEAFFBDFD6BCF3ADDE2F8F9" box="[645,705,1893,1917]" pageId="13" pageNumber="39" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from a presumably Oligocene seep carbonate in northern Colombia. A, B. Holotype (USNM 533981), an elongate specimen showing shell remains on the anterior end. C. Smaller specimen (USNM 533982).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="08DE3AD6FFE9FFBEFF2AC8C8DB06FF36" box="[196,1061,151,178]" pageId="14" pageNumber="40" type="distribution">
|
||
<paragraph id="407B695DFFE9FFBEFF2AC8C8DB06FF36" blockId="14.[151,1436,151,252]" box="[196,1061,151,178]" pageId="14" pageNumber="40">
|
||
<emphasis id="72B0B54FFFE9FFBEFF2AC8C8DE45FF35" bold="true" box="[196,358,151,177]" pageId="14" pageNumber="40">Distribution:</emphasis>
|
||
<collectingCountry id="38D329CDFFE9FFBEFE83C8C7DEC2FF36" box="[365,481,152,178]" name="Colombia" pageId="14" pageNumber="40">Colombia</collectingCountry>
|
||
(Oligocene?) and possibly
|
||
<collectingCountry id="38D329CDFFE9FFBEFCCAC8C7DCBFFF36" box="[804,924,152,178]" name="Venezuela" pageId="14" pageNumber="40">Venezuela</collectingCountry>
|
||
(Miocene).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08DE3AD6FFE9FFBEFF2AC8E2DE31FF78" pageId="14" pageNumber="40" type="etymology">
|
||
<paragraph id="407B695DFFE9FFBEFF2AC8E2DE31FF78" blockId="14.[151,1436,151,252]" pageId="14" pageNumber="40">
|
||
<emphasis id="72B0B54FFFE9FFBEFF2AC8E2DE43FF53" bold="true" box="[196,352,189,215]" pageId="14" pageNumber="40">Etymology:</emphasis>
|
||
After its occurrence near the Arroyo Piadras Palmar and Palmar-Molinera-Road in
|
||
<collectingCountry id="38D329CDFFE9FFBEFF79C8BDDE2DFF78" box="[151,270,226,252]" name="Colombia" pageId="14" pageNumber="40">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |