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<document id="AA6EABE5EA134236BB56190BA4EA19A3" ID-DOI="10.3897/zookeys.1216.130017" ID-ZooBank="A6D95900-0A19-4778-9AA5-6D87622B419F" ID-publisher-id="130017" URI-arpha="F535348F-BBA1-5E6C-B72A-6E5D85153D3B" URI-zoobank="http://zoobank.org/A6D95900-0A19-4778-9AA5-6D87622B419F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1729263426519" checkinUser="pensoft" docAuthor="Lin, Shi-Shi, Chen, Hong-Hui, Li, Yuan-Hang, Peng, Zhao-Ning, Zeng, Zhao-Chi &amp; Wang, Jian" docDate="2024" docId="AF6296E2B1615E10819274EAB15B0D8F" docLanguage="en" docName="ZooKeys 1216: 1-15" docOrigin="ZooKeys 1216" docSource="https://zookeys.pensoft.net/article/130017/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Boulenophrys lichun Lin, Chen, Li, Peng, Zeng &amp; Wang, 2024, sp. nov." docType="treatment" docUuid="02303B1E-ED1A-482B-8BD3-B32BC0DB9D6B" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="15" masterDocId="F535348FBBA15E6CB72A6E5D85153D3B" masterDocTitle="A new Boulenophrys species (Anura, Megophryidae) from the coastal hills of eastern Fujian Province, China" masterLastPageNumber="15" masterPageNumber="1" pageNumber="1" updateTime="1729275296647" updateUser="admin">
<mods:mods id="DB059258D1432A65433F5D8F697B7703" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1776DFFB4084B876CE038DD1B504B179">A new Boulenophrys species (Anura, Megophryidae) from the coastal hills of eastern Fujian Province, China</mods:title>
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<mods:namePart id="0D444D80D12A233C29DAE001742E121C">Lin, Shi-Shi</mods:namePart>
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<mods:namePart id="5C5CDEE41ED381E14DB9D58CA70250D4">Chen, Hong-Hui</mods:namePart>
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<mods:affiliation id="A959A2515DC35F7265DAACEA8F102E17">Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
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<mods:namePart id="453780905578CABCCEF23CB41034E590">Li, Yuan-Hang</mods:namePart>
<mods:nameIdentifier id="B149A5D9D7D84E3A2B04BD7B40823559" type="ORCID">https://orcid.org/0009-0009-6543-9245</mods:nameIdentifier>
<mods:affiliation id="9C2FD728B39771066FD21DEFC9DE586D">Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
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<mods:namePart id="A096D8CB42C7B1AA2F1E4FE8E0613521">Peng, Zhao-Ning</mods:namePart>
<mods:nameIdentifier id="DAD9FEFB96B8ECBE6D71383DE5C668D8" type="ORCID">https://orcid.org/0009-0008-5103-2797</mods:nameIdentifier>
<mods:affiliation id="A07AB56C65E9297BD54CCDD9E38FC417">Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
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<mods:namePart id="1437B6BB3586E5CDF76251FCA1B307CE">Zeng, Zhao-Chi</mods:namePart>
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<mods:affiliation id="7EE8A4E22B521560D5FE74843FA82899">Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
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<mods:namePart id="625189FE1E1F13765CF750DAFB323DA4">Wang, Jian</mods:namePart>
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<mods:affiliation id="DB60CD9E43125B528AA6A59961B05EC1">Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
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<mods:part id="5C4E9DA83FCA6A69544659627AC30D4F">
<mods:date id="CC2A3594DCD12D4B9C3FA82C3AB60B72">2024</mods:date>
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<mods:number id="014598C9B6989C6A62617F27D997DA84">2024-10-18</mods:number>
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<mods:number id="ADAD9394930DFC601A0DF78EE41E1330">1216</mods:number>
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<mods:identifier id="4C78A532C394544B484E2DA595D87B7A" type="ZooBank">A6D95900-0A19-4778-9AA5-6D87622B419F</mods:identifier>
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<treatment id="AF6296E2B1615E10819274EAB15B0D8F" ID-arpha="AF6296E2-B161-5E10-8192-74EAB15B0D8F" ID-zoobank="https://zoobank.org/02303B1E-ED1A-482B-8BD3-B32BC0DB9D6B" LSID="urn:lsid:zoobank.org:act:02303B1E-ED1A-482B-8BD3-B32BC0DB9D6B" LSID-ZBK="urn:lsid:zoobank.org:act:02303B1E-ED1A-482B-8BD3-B32BC0DB9D6B" httpUri="http://treatment.plazi.org/id/AF6296E2B1615E10819274EAB15B0D8F">
<subSubSection id="96814DDC7AA93331346FF9933D1F0E71" type="nomenclature">
<paragraph id="69D4D87F05E4895B0794FFD3E5A0E8C3">
<taxonomicName id="031EEE677BC30E14167330E78F7415C6" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="A63568DF62BF8FF9D553CE15D72CB620" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AB90265A369E1B92AA371A987A520840" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="07B553921D823682B689CBC32F7EB794" type="description">
<paragraph id="735D68D77958EDBEB4ACA38809D44937">
<figureCitation id="F599BCED89E4BE8DE6D3B6529700F5B4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The male holotype (GEP a 214, A C) and the female paratype (GEP a 215, D F) of Boulenophrys lichun sp. nov. in life." figureDoi="10.3897/zookeys.1216.130017.figure3" httpUri="https://binary.pensoft.net/fig/1156127">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQWAG" type="material">
<paragraph id="21D4438E06133DD48BA415328542F275">
<heading id="568EEE22023837BC329BC2931F8A4E62" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="10CE74763BDA39EE1E025021F1F955B8">
<materialsCitation id="B6FBF24E33008A8740FF24CF314E1F38" collectingDate="2024-02-04" collectionCode="GEP" collectorName="Jian Wang &amp; Zhao-Chi Zeng &amp; Shi-Shi Lin &amp; Yuan-Hang Li" country="China" county="Ningde City" elevation="230" latitude="26.645775" location="Mt Nanji" longLatPrecision="1" longitude="119.519936" municipality="Jiaocheng District" specimenCount="1" specimenCount-male="1" stateProvince="Fujian Province" typeStatus="holotype">
<emphasis id="E2FB06A0ADA9E9B26D38B43C2E5E50E3" bold="true">
<emphasis id="76BD50B47D05FC388B9F6345C69110B7" bold="true" italics="true">
<typeStatus id="3305849CE846A782158678C629740D00" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="866218AF02769AFBB9A9DACA21F62C16" name="China">China</collectingCountry>
<specimenCount id="1C4F50874D03D620299222F7E511D6C8" count="1" type="male"></specimenCount>
;
<collectingRegion id="57C18B1DC8A182645A89589508C30801" country="China" name="Fujian">Fujian Province</collectingRegion>
,
<collectingCounty id="18B5719013D71DBA244D1CD520C1ABD8">Ningde City</collectingCounty>
,
<collectingMunicipality id="03FBAAB78139765F50EA5CA35A047F86">Jiaocheng District</collectingMunicipality>
,
<location id="DE6207C5BB6B7362B58F832C12364DDE" LSID="urn:lsid:plazi:treatment:AF6296E2B1615E10819274EAB15B0D8F:DE6207C5BB6B7362B58F832C12364DDE" country="China" county="Ningde City" latitude="26.645775" longLatPrecision="1" longitude="119.519936" municipality="Jiaocheng District" name="Mt Nanji" stateProvince="Fujian Province">Mt Nanji</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5WAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[119.519939,26.645774]}">
<geoCoordinate id="967A3B1B95D18BED9E947E10D946CD2C" degrees="26.645774" direction="north" orientation="latitude" precision="1" value="26.645775">26.645774 ° N</geoCoordinate>
,
<geoCoordinate id="B5852F7616BD278F00E3BD11444801B1" degrees="119.519939" direction="east" orientation="longitude" precision="1" value="119.519936">119.519939 ° E</geoCoordinate>
</named-content>
, ca.
<quantity id="FC66C952E3B80EB39AA032196B9736F9" metricMagnitude="2" metricUnit="m" metricValue="2.3" unit="m" value="230.0">
<elevation id="7F73EED5053DBABA8D574A6918FE80CE" metricMagnitude="2" metricUnit="m" metricValue="2.3" unit="m" value="230.0">230 m</elevation>
</quantity>
elev.;
<date id="37DFA94075086614C564684DE0F96D3C" value="2024-02-04">
<collectingDate id="D418BFA3AADC142ED963ED7E5274AD0F" value="2024-02-04">4 Feb. 2024</collectingDate>
</date>
;
<collectorName id="6592F6C511E42837DC501ED1AB3D863B">Jian Wang</collectorName>
,
<collectorName id="CB056EF3341E2EC4DEA8B8DE93A8ED9B">Zhao-Chi Zeng</collectorName>
,
<collectorName id="7862A6FC298FB98D24E057FE0E01FE3D">Shi-Shi Lin</collectorName>
and
<collectorName id="BC4564DB01F86975C5B25DD869221513">Yuan-Hang Li</collectorName>
leg.;
<abbrev id="ABBRID0EDXAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 214.
</materialsCitation>
</paragraph>
<caption id="D53DEA791A1D5574A1C7A2BD10DBDCB4" ID-DOI="10.3897/zookeys.1216.130017.figure3" ID-arpha="E7F83773-006E-5EB9-B15F-D9DEA63D520C" httpUri="https://binary.pensoft.net/fig/1156127" startId="F3">
<paragraph id="E48D97F35FD56BD62620E1DBECCDD6F8">
<label id="3BC83A9A52342898546438B577318BBE">Figure 3.</label>
</paragraph>
<paragraph id="12BAE93FD80B0CC445431A6B78B14A4A">
The male holotype (
<abbrev id="ABBRID0EPXAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 214,
<emphasis id="94AA5D543ABF3AF99FF99077798264DB" bold="true">A C</emphasis>
) and the female paratype (
<abbrev id="ABBRID0EVXAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 215,
<emphasis id="5A5EC990F6C6CAD258E0ECAE10B84E9F" bold="true">D F</emphasis>
) of
<taxonomicName id="FC53F6FE3F61AD931B21F4D494900294" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="03B7DAB662232C3C2B124D596F711A39" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AA4A5091AD29037F53D5DD0709140BF8" rank="species">sp. nov.</taxonomicNameLabel>
in life.
</paragraph>
</caption>
<paragraph id="C01E78590826CF741F64B2928A73A06F">
<materialsCitation id="F4229613C249F24AD2BA4425C5521C01" collectingDate="2024-02-04" collectionCode="CIB" collectorName="Jian Wang &amp; Zhao-Chi Zeng &amp; Shi-Shi Lin &amp; Yuan-Hang Li" country="China" county="Ningde City" elevation="230" latitude="26.645775" location="China" longLatPrecision="1" longitude="119.519936" municipality="Jiaocheng District" specimenCode="CIB 121428" specimenCount="4" specimenCount-male="4" stateProvince="Fujian" typeStatus="paratype">
<emphasis id="C807152715572C9EF308E5C33CFCC1C6" bold="true">
<emphasis id="46674839CCE3335187CFBCD3D48A3DDD" bold="true" italics="true">
<typeStatus id="5F162DE408EEC8397BE136C6DEA2C344" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="ECF24295A223CCBDA791656BB3C028F4" name="China">China</collectingCountry>
<specimenCount id="6389891E63B50A3285CC3E37C7F0BED2" count="4" type="male">4 ♂♂</specimenCount>
; same data as for holotype;
<specimenCode id="00EB43A2FF2439BCB54C08326EF8D9DA" collectionCode="CIB" country="China" httpUri="http://grbio.org/cool/vxkr-andy" name="Chengdu Institute of Biology" type="">
<named-content id="06FBE4BF760F565539D1073BF7E837C2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chengdu-institute-biology" xlink_title="Herpetological Museum, Chengdu Institute of Biology, the Chinese Academy of Sciences">CIB</named-content>
121428
</specimenCode>
[field number
<abbrev id="ABBRID0EWYAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 210],
<abbrev id="ABBRID0E1YAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 211213
</materialsCitation>
<materialsCitation id="A93161AD17461AE4A2530F73F0EDBBC8" collectingDate="2024-02-04" collectionCode="GEP" collectorName="Jian Wang &amp; Zhao-Chi Zeng &amp; Shi-Shi Lin &amp; Yuan-Hang Li" country="China" county="Ningde City" elevation="230" latitude="26.645775" location="Mt Nanji" longLatPrecision="1" longitude="119.519936" municipality="Jiaocheng District" specimenCount="1" specimenCount-female="1" stateProvince="Fujian" typeStatus="paratype">
<specimenCount id="8BB68CCCB1C31E4F9C04B8DDC299ADBA" count="1" type="female">1 ♀</specimenCount>
; same data as for holotype;
<abbrev id="ABBRID0E5YAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 215
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECZAG" type="etymology">
<paragraph id="92C180E593FEFBD7F072E258C6F2DB68">
<heading id="7ED79F399FBC8B64B52731ABE3D3991C" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="7CC4B3701B803939B310AD9505099C42">
The specific name
<taxonomicName id="03CB938BCBB375066139BB4B4CF96AF6" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun">
<emphasis id="0428019D5072279D5B7F1620B9528080" italics="true">lichun</emphasis>
</taxonomicName>
is derived from Chinese Pinyin Lì Chūn, i. e. 立春 in Chinese, which means the beginning of spring, the first of the 24 solar terms (24 节气) of
<collectingCountry id="2A711593687B42B6046BA10E20108634" name="China">China</collectingCountry>
. The specific name refers to the breeding season of the new species which begins around this period. The song of the new species heralds the spring of a year. The type specimens of the new species were also collected on “ Lichun ” of the Year 2024.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESZAG" type="diagnosis">
<paragraph id="C97CD7C6EB1230B85E3F8236CB3A244C">
<heading id="05A00138F2B06019A7FBCD3BC1E7A59B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D861EC12CD81CCA7509EAEC1C14206A1">
(1) small size (
<abbrev id="ABBRID0EYZAG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
33.537.0 mm in
<specimenCount id="B3FA198ABAE1F41F3359528EF1B1D817" count="5" type="adult">five adult</specimenCount>
males,
<abbrev id="ABBRID0E3ZAG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
<quantity id="8C2478430DFFD9D633B6AEFD8C0E8B2B" metricMagnitude="-2" metricUnit="m" metricValue="4.71" unit="mm" value="47.1">47.1 mm</quantity>
in a single adult female); (2) canthus rostralis well developed, tongue not notched posteriorly; (3) tympanum distinct; (4) vomerine ridges and vomerine teeth present; (5) dorsal skin rough and highly granular, discontinuous X-shaped ridge on center of dorsum, discontinuous dorsolateral ridges present, sparse large tubercles on flanks, dorsal limbs with discontinuous transverse ridges and tubercles, ventral skin with dense raised tubercles; (6) outer margin of upper eyelid with a small horn-like prominent tubercle, supratympanic fold distinct and narrow, curving posteroventrally to above arm; (7) two metacarpal tubercles distinct, inner one observably enlarged; relative finger lengths I &lt;
<collectionCode id="770B80BC54A6923ECE90FC29A8A6C3E3">II</collectionCode>
&lt;
<collectionCode id="472A53BC938581808472CAE4BAC9167C">IV</collectionCode>
&lt;
<collectionCode id="A0FFC6EAF11600DDC886766AE3037587">III</collectionCode>
; distinct subarticular tubercle at base of each finger; (8) heels not meeting when hindlimbs folded; tibio-tarsal articulation reaching shoulder to posterior corner of eye; (9) toes without webbing and lateral fringes, inner metatarsal tubercle long ovoid, outer one absent, relative toe length I &lt;
<collectionCode id="9DBF51B57765AFD268B53DD853F24268">II</collectionCode>
&lt;
<collectionCode id="48E44799D9EAF7E9336D7A9952500841" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
&lt;
<collectionCode id="289E46F80D7AC294F457DB3C85829EC5">III</collectionCode>
&lt;
<collectionCode id="7154D65FADFBB31C6602659F5226972E">IV</collectionCode>
; (10) dorsal surface yellowish-brown with irregular dark-brown patches, and dark-brown triangular marking between eyes, dorsal limbs and digits light brown with dark-brown transverse bands; and (11) dense nuptial spines on dorsal bases of fingers I and II in breeding adult males, subgular vocal sac present in males.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EA1AG" type="description">
<paragraph id="38647783CC98549DD21E6929F48569A4">
<heading id="7EE3B7352624C3E305EB223EB5A1AA44" reason="title">
Description of
<typeStatus id="68D46B223EBEDDD3688110C16F42D8F2" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="963E5247AF99523288BE7FD7EC43E54D">
Adult male. Body size small,
<abbrev id="ABBRID0EG1AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
37.0 mm. Head width larger than head length, HWD /
<abbrev id="ABBRID0EK1AG" xlink_title="head length, from tip of snout to the articulation of the jaw">HDL</abbrev>
1.04; snout rounded in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eyes moderate in size,
<abbrev id="ABBRID0EO1AG" xlink_title="eye diameter, from the anterior corner of the eye to posterior corner of the eye">ED</abbrev>
0.34 of
<abbrev id="ABBRID0ES1AG" xlink_title="head length, from tip of snout to the articulation of the jaw">HDL</abbrev>
, pupil vertical, near diamond-shaped; nostril obliquely ovoid; canthus rostralis well developed; loreal region slightly oblique; internasal distance slightly larger than interorbital distance; tympanic region oblique, tympanum distinct and visible in dorsal view; tympanum moderate in size, margin clear, upper margin in contact with supratympanic fold, lower margin in contact with upper lip,
<abbrev id="ABBRID0EW1AG" xlink_title="tympanum diameter, horizontal diameter of tympanum">TD</abbrev>
/
<abbrev id="ABBRID0E11AG" xlink_title="eye diameter, from the anterior corner of the eye to posterior corner of the eye">ED</abbrev>
0.55; large ovoid choanae at base of maxilla; vomerine ridge and vomerine teeth present, maxillary teeth present; margin of tongue rounded, not notched distally; presence of single subgular vocal sac.
</paragraph>
<paragraph id="73BBD0DAA8EECF113AA572B7D0BF29AA">
Forearm length 0.23 of
<abbrev id="ABBRID0EA2AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
, hand 0.24 of
<abbrev id="ABBRID0EE2AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
; webbing absent between fingers, lateral fringes absent, relative finger length I &lt;
<collectionCode id="B5A70161DD3E82EF72B3ADD5C48CF042">II</collectionCode>
&lt;
<collectionCode id="41ABB0E53CC8BCB5BECC4222F53F70A1">IV</collectionCode>
&lt;
<collectionCode id="7023D05580953F3F7FFC0C5B7DE5E5B2">III</collectionCode>
; tips of fingers slightly dilated, round; subarticular tubercles on base of fingers present, distinct; inner metacarpal tubercle observably enlarged, outer one slightly smaller; single nuptial pad bearing nuptial spines present on dorsal surface of first and second fingers, respectively. Hindlimbs short, tibio-tarsal articulation reaching forward to posterior conner of eye when hindlimb stretched along body; heels not meeting when flexed hindlimbs held at right angles to body axis; crus length 0.40 of
<abbrev id="ABBRID0EI2AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
and foot length 0.58 of
<abbrev id="ABBRID0EM2AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
; relative toe length I &lt;
<collectionCode id="7A7EE6D2EFBD63E7BD08A123412242C9">II</collectionCode>
&lt;
<collectionCode id="61DBC4FE9B23DBD317B27BBB5CDDDB57" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
&lt;
<collectionCode id="5DE27FC7CACE106DBCB5B0F21AF36BF8">III</collectionCode>
&lt;
<collectionCode id="901BCF7C4017A075DC4F1CA9E46A51AF">IV</collectionCode>
; tips of toes round and slightly dilated; toes without lateral fringes and webbing; subarticular tubercles on base of toes present and distinct; inner metatarsal tubercle long ovoid and lacking outer metatarsal tubercle.
</paragraph>
<paragraph id="4734D26911AA43CFC8B6127EC5A2F0AB">Dorsal skin rough and highly granular; dense large tubercles on flanks; single horn-like prominent tubercle on edge of upper eyelid; obvious supratympanic fold curving posteroventrally from posterior corner of eye to level above insertion of arm; upper lip, mandibular articulation, loreal, temporal region excluding tympanum, upper eyelid and surface around cloaca with conical tubercles; discontinuous X-shaped ridge on center of dorsum, discontinuous dorsolateral ridges present; ventral surface with dense raised tubercles; tubercles on ventral hindlimbs and around cloaca bearing tiny spines on their tips; small and distinct pectoral gland closer to axilla; single femoral gland positioned on posterior surface of thigh at midpoint between knee and cloaca.</paragraph>
</subSubSection>
<subSubSection id="SECID0ER2AG" type="Coloration of holotype">
<paragraph id="B6271C329E968C040FD7A209914A3CA2">
<heading id="D52D074E86447DB558C5C6BA45BD5C27" reason="title">
Coloration of
<typeStatus id="1471056AE28271617DBBAFED457CBB9A" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="801916717A651819C67249A8BD85D970">In life, dorsal surface of body yellowish-brown with irregular dark-brown patches, dark-brown X-shaped marking on center of dorsum, dark-brown triangular marking between eyes. A vertical dark-brown band present below eye. Dorsal surface of limbs with dark-brown transverse bands. Tubercles on edge of upper eyelids orange. Supratympanic fold light brown. Surface of throat and chest yellowish-brown with irregular dark brown and white patches and white and orange dots. Center of throat with black longitudinal band. Surface of abdomen white, mottled with orange dots and black patches. Surface of ventral limbs purple brown, with white mottling and dark-brown patches. Spines on tips of tubercles on ventral hindlimbs and area around cloaca black. Digits gray white; subarticular tubercles, inner and outer metacarpal tubercles and inner metatarsal tubercle grayish-brown. Pectoral glands and femoral glands white. Iris yellowish-brown with range mottling.</paragraph>
<paragraph id="F1239DB3829FDD7CE831755D55F2B2C9">In preservative, the dorsal surface of the body is dark brown, with markings and patches more distinct. Surface of chest, throat and limbs are dark brown, with dark-brown markings and patches more distinct, white patches and dots faded and orange dots absent. Color of pectoral glands and femoral glands faded.</paragraph>
</subSubSection>
<subSubSection id="SECID0EX2AG" type="variation">
<paragraph id="A05EE43468A52F5E89D8E035470F520A">
<heading id="24A90567E5E25DB12CD4257C7BBD9320" reason="title">Variation.</heading>
</paragraph>
<paragraph id="1CAB12BD573BE69772DAEAC1F0644AB8">
Morphometric variation is listed in Table
<tableCitation id="58EB5F81A86768F92996A51BE55598DE" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements (in mm) of voucher specimens of Boulenophrys lichun sp. nov.; * holotype." httpUri="http://table.plazi.org/id/D7A01796E20787A9657B8242C75A0570" tableUuid="D7A01796E20787A9657B8242C75A0570">2</tableCitation>
. Most of the
<typeStatus id="D6734D31C7CE3F4A0B4CD4DB892F682E" type="paratype">paratypes</typeStatus>
are similar to the
<typeStatus id="399D572516854E98B39EB3B11A099B0F" type="holotype">holotype</typeStatus>
in morphology and color pattern, except for the following: tibio-tarsal articulation reaching forward to posterior corner of eye when hindlimb stretched along body in the
<typeStatus id="94E92210034E424CD44299F6EDA9D9C7" type="holotype">holotype</typeStatus>
<abbrev id="ABBRID0EB3AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 214, while reaching to shoulder in female
<typeStatus id="79F10013B192518984DB5BD07B2D0039" type="paratype">paratype</typeStatus>
<abbrev id="ABBRID0EF3AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 215; absence of nuptial pads and spines in the female
<typeStatus id="F2E552D0D89127E1E74A1C3431F29F98" type="paratype">paratype</typeStatus>
; larger body size in the female
<typeStatus id="866F3D355B5033C4436C299E4493472A" type="paratype">paratype</typeStatus>
.
</paragraph>
<caption id="D7A01796E20787A9657B8242C75A0570" ID-Table-UUID="D7A01796E20787A9657B8242C75A0570" httpUri="http://table.plazi.org/id/D7A01796E20787A9657B8242C75A0570" startId="T2" targetIsTable="true">
<paragraph id="6CA7C23C57EC05ACCF370E0EFB2F8706">
<label id="67B894C971DF5AB609BA0172C86E490B">Table 2.</label>
</paragraph>
<paragraph id="805032C2F6223927388B992DB159BE75">
Measurements (in mm) of voucher specimens of
<taxonomicName id="E76D9F8CE35830C7FEE2E253D9182869" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="67AD94F17D3693F0D769D4E2FB0751CA" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="514146FED6A315B9242D5D6C95CFA703" rank="species">sp. nov.</taxonomicNameLabel>
; * holotype.
</paragraph>
</caption>
<paragraph id="A6C52B2D2377C1964F7CC05202432890">
<table id="TID0EMFBI" rules="all">
<tbody id="36E3B7183B13F1276238D087B8E649B8">
<tr id="F51F397C98409923553C8BA2D17A3C28">
<th id="049887F96C8FF5D6897CFA64A5C0B654" colspan="1" rowspan="1">Voucher</th>
<th id="DB8DC50175D957C19DCA3A7E692CC27F" colspan="1" rowspan="1">
<named-content id="8D527AA7A55CBB37457ADEE52051DCD2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chengdu-institute-biology" xlink_title="Herpetological Museum, Chengdu Institute of Biology, the Chinese Academy of Sciences">CIB</named-content>
121428
</th>
<th id="0CA4E6B5F1CF2906F196C049163B2365" colspan="1" rowspan="1">
<abbrev id="ABBRID0ER4AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 211
</th>
<th id="DDC60DC5F600A676CAF3F09830C5456D" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZ4AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 212
</th>
<th id="B44F3FD5CC718314C296736B2745EE22" colspan="1" rowspan="1">
<abbrev id="ABBRID0EB5AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 213
</th>
<th id="31B362FEEF17A1B10142106D4931C82D" colspan="1" rowspan="1">
<abbrev id="ABBRID0EJ5AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 214 *
</th>
<th id="5397D18DC1FAA4D6BDBB0CD662A57880" colspan="1" rowspan="1">
<abbrev id="ABBRID0ER5AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 215
</th>
</tr>
<tr id="C93855150C4F8713157657F22D11AED2">
<td id="E8EBC8994251B0808D08F12BAE518F40" colspan="1" rowspan="1">
<emphasis id="CE3EAEAFC816C38EC2841B687B901552" bold="true">Sex</emphasis>
</td>
<td id="72A185572C78C8A4DFA64AC659CD6D5F" colspan="1" rowspan="1">male</td>
<td id="26D17BEF9E42DB7C61A35A33203C12E0" colspan="1" rowspan="1">male</td>
<td id="9B5BC789F04E9028D8F638F1BF22B024" colspan="1" rowspan="1">male</td>
<td id="63F422351A8CB4296A15FC23F918F6CE" colspan="1" rowspan="1">male</td>
<td id="3D276470CE88E549A3BB40CA75EE3ADA" colspan="1" rowspan="1">male</td>
<td id="1FAD1EAFB39C9E20C34C70666F96724A" colspan="1" rowspan="1">female</td>
</tr>
<tr id="837206366F948802BB8652B46F2D8482">
<td id="0EA5A4DC01394BB6A47F650C3CABD6D5" colspan="1" rowspan="1">
<abbrev id="ABBRID0EU6AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">
<emphasis id="836BC9227213F70705FCAB622914323A" bold="true">SVL</emphasis>
</abbrev>
</td>
<td id="E6E77903A0E468DDB8446C91C6A4EE46" colspan="1" rowspan="1">36.7</td>
<td id="1E988A9285124D1F36A35DC8643EF954" colspan="1" rowspan="1">33.5</td>
<td id="5BC78BF7ACAEDAF5BA8846C52782AE72" colspan="1" rowspan="1">35.7</td>
<td id="01EDEA2094248502067A4C76EBFB3835" colspan="1" rowspan="1">34.6</td>
<td id="2C528598EFBD687E8B18FB9E221585E4" colspan="1" rowspan="1">37.0</td>
<td id="6DF0424C0454D1221495A78F8DB7DF51" colspan="1" rowspan="1">47.1</td>
</tr>
<tr id="75040B7A321018737F7F57E04C8B8C2A">
<td id="FDA8154537EB3E9739900077B83A8B82" colspan="1" rowspan="1">
<abbrev id="ABBRID0EQABG" xlink_title="head length, from tip of snout to the articulation of the jaw">
<emphasis id="542561CB99529E1956A708C77F33FFA5" bold="true">HDL</emphasis>
</abbrev>
</td>
<td id="C17C2254E2DA8E44C75600E2DA1BF5EA" colspan="1" rowspan="1">13.1</td>
<td id="098CD03A141193C5CCE97F89D2A3C4DB" colspan="1" rowspan="1">12.7</td>
<td id="86858C9A684984C08F3326BADFF7CE9C" colspan="1" rowspan="1">13.5</td>
<td id="9E8746D54D5DD09C535E869DD94321E1" colspan="1" rowspan="1">12.9</td>
<td id="327043E8B1B9DB37DA08D9F43E9DABA1" colspan="1" rowspan="1">13.3</td>
<td id="D398C48E384A5BF7921DCCA6C65613AF" colspan="1" rowspan="1">15.3</td>
</tr>
<tr id="4881AD5ED959143B2F629D0AD811F818">
<td id="FDC4C9E3DC04CE8FD83F93A0D782540B" colspan="1" rowspan="1">
<abbrev id="ABBRID0EMBBG" xlink_title="head width, head width at the commissure of the jaws">
<emphasis id="991DADDEF5EDC9129CBD39E218919338" bold="true">HDW</emphasis>
</abbrev>
</td>
<td id="F26CE8FC60C4807AA6DF6CB6EBF06916" colspan="1" rowspan="1">13.6</td>
<td id="2114F11BCD04FDBDDEAA1FDE10193939" colspan="1" rowspan="1">13.3</td>
<td id="DF4D73B960699E6397BE850D025F67F2" colspan="1" rowspan="1">13.8</td>
<td id="9EF3D3E4DDD716E8B4B9F3EF6858B17F" colspan="1" rowspan="1">13.6</td>
<td id="B7ECD76DE5F61A9DE8CC37BCE97CE6A2" colspan="1" rowspan="1">13.9</td>
<td id="0AEEABB8B490A8E567008C95306969C1" colspan="1" rowspan="1">16.5</td>
</tr>
<tr id="7FCB56D4F0089B5E93DF821E7E411AE5">
<td id="A2F06853CEE82191B6AB68D670C2537E" colspan="1" rowspan="1">
<abbrev id="ABBRID0EICBG" xlink_title="eye diameter, from the anterior corner of the eye to posterior corner of the eye">
<emphasis id="4C293E554603FEC204996F0FE9C36402" bold="true">ED</emphasis>
</abbrev>
</td>
<td id="895F6A2509CBFDF8688C910B66C6F3C3" colspan="1" rowspan="1">4.5</td>
<td id="4A98977A01EA7BD8248402DDCF7241EF" colspan="1" rowspan="1">4.0</td>
<td id="9371D4F35B0DFDEC61FBF8055AB5CE66" colspan="1" rowspan="1">4.5</td>
<td id="F168E37842191EF6BF89EA4CEC5C56C3" colspan="1" rowspan="1">4.5</td>
<td id="E37F63BBB44165519C3F2C0F37C041A7" colspan="1" rowspan="1">4.8</td>
<td id="5E848F9895BBA3F944C663DB6BBD9F64" colspan="1" rowspan="1">5.5</td>
</tr>
<tr id="DFA07F122FB37ADD221C553E7F04BF7E">
<td id="36DCF704884B0F227EDECA6007B94EEE" colspan="1" rowspan="1">
<abbrev id="ABBRID0EEDBG" xlink_title="tympanum diameter, horizontal diameter of tympanum">
<emphasis id="B9EF7459A0B992C8D0FD4C6243110E56" bold="true">TD</emphasis>
</abbrev>
</td>
<td id="9AEE1C580EA8877D13768E47BEBCF7CB" colspan="1" rowspan="1">2.3</td>
<td id="8E075FA5B85383DA471D5E01D003A313" colspan="1" rowspan="1">2.6</td>
<td id="6F829CFE7AB9EF9C4679628E60F44AAA" colspan="1" rowspan="1">2.5</td>
<td id="AFEB03FE6E9FF4BB16314642D5A283B6" colspan="1" rowspan="1">2.5</td>
<td id="5BCF0497C339322372C754DD8AF2FD36" colspan="1" rowspan="1">2.6</td>
<td id="EA68076DDF2CCEAD933F4E5712B47FFD" colspan="1" rowspan="1">2.8</td>
</tr>
<tr id="62E1EA674F983185FEC2B930B47F61F7">
<td id="B7C1B46FCADA13D8143CC42749DBD8A5" colspan="1" rowspan="1">
<abbrev id="ABBRID0EAEBG" xlink_title="tympanumeye distance, from anterior edge of tympanum to posterior corner of the eye">
<emphasis id="8F2278A59BA90DBFD5053590EE507221" bold="true">TED</emphasis>
</abbrev>
</td>
<td id="DC83CA011294231E55E87FCCC843C09B" colspan="1" rowspan="1">1.7</td>
<td id="1FD511F64C860832241AEB92B6864294" colspan="1" rowspan="1">1.7</td>
<td id="3DF903E38F468B722CAFE760DAC50849" colspan="1" rowspan="1">1.8</td>
<td id="C4A34397B0D15C6E4CD3746C77C0993D" colspan="1" rowspan="1">1.8</td>
<td id="D646DCA0A2286CFF1BFBBC0180A5D1E2" colspan="1" rowspan="1">1.7</td>
<td id="FDDDBC69FE2035E818EF1D19364C3E61" colspan="1" rowspan="1">2.4</td>
</tr>
<tr id="CEE70C15D1F6F47109B336F9006817C6">
<td id="A4C790331A905078FAB53957F2B2976E" colspan="1" rowspan="1">
<abbrev id="ABBRID0E3EBG" xlink_title="snout length, from tip of snout to the anterior corner of the eye">
<emphasis id="2370E57CA98AE47F32FF329618C8CE67" bold="true">SNT</emphasis>
</abbrev>
</td>
<td id="6858B6CBE3641A1C40D27D8A4FFC9EAE" colspan="1" rowspan="1">4.5</td>
<td id="F4B1F48E4D4E44055223C24A0DDAEFDF" colspan="1" rowspan="1">4.4</td>
<td id="76B55FEF75ECA495FD47452A8C5907B8" colspan="1" rowspan="1">4.5</td>
<td id="3DDEC2E3474C1FAAA1EDB4465233672A" colspan="1" rowspan="1">4.5</td>
<td id="A8B3AFBA28CB97B436968FC7F376B40B" colspan="1" rowspan="1">4.6</td>
<td id="97BC150C73E30FDE53F64D7041E550B1" colspan="1" rowspan="1">4.9</td>
</tr>
<tr id="FC4C843E00E54AD1782541ECB85B8B01">
<td id="CDDE20A5786956A1E1F609CC71AED899" colspan="1" rowspan="1">
<abbrev id="ABBRID0EYFBG" xlink_title="internasal distance, distance between nares">
<emphasis id="9F94FDE8684335AEF85B76AB29E82BEB" bold="true">IND</emphasis>
</abbrev>
</td>
<td id="34330699A17B440F3D918DFE135FEB16" colspan="1" rowspan="1">3.6</td>
<td id="E3AEBF782E4EBB2B9BC589AAD9031228" colspan="1" rowspan="1">3.7</td>
<td id="CE04950AC81AF8B94A52A8E03F8A108B" colspan="1" rowspan="1">3.8</td>
<td id="D5FA6308A105D4DD78AC199AB8C7EC99" colspan="1" rowspan="1">3.7</td>
<td id="68F11F770A1E7DB9383D16DF5606B650" colspan="1" rowspan="1">3.9</td>
<td id="8CD0BCCCE0802570410CDE3715D32810" colspan="1" rowspan="1">4.3</td>
</tr>
<tr id="ABA95591697C706BDFA6221857661B1C">
<td id="099301FB9BA322A17148FB7C8ABC38AF" colspan="1" rowspan="1">
<abbrev id="ABBRID0EUGBG" xlink_title="interorbital distance, minimum distance between upper eyelids">
<emphasis id="30F493A5A76AE8B3E65F400774021CF4" bold="true">IOD</emphasis>
</abbrev>
</td>
<td id="4466D45C2232AACA74FF5148DFB31C3A" colspan="1" rowspan="1">3.5</td>
<td id="5D4FCCB81CE9AA4D23B14293CB045461" colspan="1" rowspan="1">3.3</td>
<td id="D124B416B9D9BD8569BD86795AD69A3E" colspan="1" rowspan="1">3.4</td>
<td id="8130E6AA8A6CC9E5A985D3407CA8878A" colspan="1" rowspan="1">3.5</td>
<td id="D41D4374C0DD317EFAAE2FB3136677DD" colspan="1" rowspan="1">3.7</td>
<td id="E71BE03E633733E9906F22BE6892AD3C" colspan="1" rowspan="1">4.3</td>
</tr>
<tr id="336B68D02418E26419DE10C8CFD1483B">
<td id="E775385E74FD8C793800BDB0BE18E3D3" colspan="1" rowspan="1">
<emphasis id="12301992514B37BCD0C86BB595C2058E" bold="true">HDN</emphasis>
</td>
<td id="DEDDDE40B833D9603A3B3AB644E605AF" colspan="1" rowspan="1">8.7</td>
<td id="21F0A68966074FF0FEF070ECD68FB82C" colspan="1" rowspan="1">7.8</td>
<td id="5DA0C61BEBDB75FE9D28DB7456EF344B" colspan="1" rowspan="1">8.7</td>
<td id="7559EA8BA19814B5DC2E45390BBF28A9" colspan="1" rowspan="1">9.1</td>
<td id="4C1274E021587BA9D5D77CCAADFF9C8D" colspan="1" rowspan="1">9.2</td>
<td id="57EC47621EE5734CDBEFD8F6A8DA878E" colspan="1" rowspan="1">10.2</td>
</tr>
<tr id="918FFCA1FE96E3C58DEE294E981B81D1">
<td id="5EC2BC172CAFBD35CA53229B3983502E" colspan="1" rowspan="1">
<abbrev id="ABBRID0EJIBG" xlink_title="forearm or radiulna length, from the flexed elbow to the proximal border of the outer palmar tubercle">
<emphasis id="6066D2CAB027DA10A07B99B25BB43209" bold="true">RAD</emphasis>
</abbrev>
</td>
<td id="7AC0B388EC756766A2D6795467B1C77F" colspan="1" rowspan="1">8.6</td>
<td id="FA7C181065AD18C3A9613A34BE4A02CA" colspan="1" rowspan="1">7.0</td>
<td id="461EBCD62B7E08C4DA78827B2FBCCC7E" colspan="1" rowspan="1">8.5</td>
<td id="A4645527919747E813A8AD132A8A9A3A" colspan="1" rowspan="1">8.1</td>
<td id="40433DB7EF899C0A6764B9CE5FC0FA5D" colspan="1" rowspan="1">8.7</td>
<td id="D09CB509751A7E7713DBB2B7EDA3A337" colspan="1" rowspan="1">9.0</td>
</tr>
<tr id="12F9B569A327495B6762F32E1B89C8CA">
<td id="74A1A4509C09C0A0B5C6B1D6F191CB01" colspan="1" rowspan="1">
<abbrev id="ABBRID0EFJBG" xlink_title="foot length, from distal end of shank to the tip of digit IV">
<emphasis id="D1105FED716839ED0C96F79AC1DA206D" bold="true">FTL</emphasis>
</abbrev>
</td>
<td id="D90F4074B2AC6D12C3738D556D3A6749" colspan="1" rowspan="1">20.7</td>
<td id="31EFE29ADDA1FCDA5BDB07FE7DF37FCF" colspan="1" rowspan="1">18.9</td>
<td id="D91089EA322F744A56C24B1677DDD767" colspan="1" rowspan="1">20.7</td>
<td id="99ADC8395DAD6E4920B04E48E264C99E" colspan="1" rowspan="1">20.9</td>
<td id="3F65D824066F1FDE0D8DE065B40C984C" colspan="1" rowspan="1">21.4</td>
<td id="8EA6C31A474B8CE73B728BB77BFE6D8F" colspan="1" rowspan="1">24.1</td>
</tr>
<tr id="A674B4931E9612DB95BCA6A42EAC6810">
<td id="7F106DAA7E452DAC075FC199228C1D51" colspan="1" rowspan="1">
<abbrev id="ABBRID0EBKBG" xlink_title="crus or tibiofibula length, from the outer surface of the flexed knee to the heel">
<emphasis id="06732C834DDD5C076E6C87ED3E806710" bold="true">TIB</emphasis>
</abbrev>
</td>
<td id="0944694D1DD0B27046D3855EB6B05E84" colspan="1" rowspan="1">14.3</td>
<td id="3A71C93BE3D94444AA564639AE9B0D9F" colspan="1" rowspan="1">12.5</td>
<td id="B350CEE68CB969FEF88CC53FBFD58CDA" colspan="1" rowspan="1">14.3</td>
<td id="B4E17C505916B6D6944C5FD0B7C275F9" colspan="1" rowspan="1">14.5</td>
<td id="D449D805881DAB681CEBC614426DD1AA" colspan="1" rowspan="1">14.7</td>
<td id="A6FFF6F5BADB81EDD0BE2D14EEFF1B8D" colspan="1" rowspan="1">15.7</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXKBG" type="comparisons">
<paragraph id="E2D10ACCFAE6CB2A9EFF30D146914556">
<heading id="77B7C96CB03F25E56DA833B49ABC1E7F" reason="title">Comparisons.</heading>
</paragraph>
<paragraph id="9C7199A408517EE176B9C32B9FC26E86">
<taxonomicName id="2F96F083B9A1CA835D02B2C357A0E14D" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="D3CF4FF24269E5F976E881F7106F1499" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F21AFA049080D8E24DC92668B5C4ED35" rank="species">sp. nov.</taxonomicNameLabel>
can easily be distinguished from the following congeners by its heels not meeting when flexed hindlimbs held at right angles to body axis:
<taxonomicName id="24155DAE73375F7DB73E283FA5704FFD" baseAuthorityName="Li, Lu, Liu &amp; Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="anlongensis">
<emphasis id="D791F609DA31DB398A4DC915611DCD7E" italics="true">B. anlongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="A260512E2FCD102E46469FED8840F9A8" baseAuthorityName="Wu, Li, Liu, Wang &amp; Wu" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="baishanzuensis">
<emphasis id="F375BD0535FC10B8DD2D01B9A4237F9B" italics="true">B. baishanzuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="F90362AB635CA49ED4CD10A8921233C8" baseAuthorityName="Jiang, Fei &amp; Ye" baseAuthorityYear="2009" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="binlingensis">
<emphasis id="74F5F6E771ACAC72226A15F8298C857F" italics="true">B. binlingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6688A3C8E36AD7A6616BD9258705A2C0" baseAuthorityName="Shen" baseAuthorityYear="1994" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="caudoprocta">
<emphasis id="A00FA677354BBF047351254AE2341935" italics="true">B. caudoprocta</emphasis>
</taxonomicName>
,
<taxonomicName id="C37E60217285E6539A484F8A88382D96" baseAuthorityName="Wang &amp; Liu" baseAuthorityYear="2014" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cheni">
<emphasis id="24357EE2235C4A3229CB2A9F6211B644" italics="true">B. cheni</emphasis>
</taxonomicName>
,
<taxonomicName id="441143FBD9992F34D0AA3C7814CE9D8A" baseAuthorityName="Xu, Li, Liu, Wei &amp; Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="chishuiensis">
<emphasis id="69A7D4CA45A16C53783631761C8FB4E3" italics="true">B. chishuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="3C371B7DAAC3CAE8425AE343103CCBDB" baseAuthorityName="Luo, Wang, Wang, Lu, Wang, Deng &amp; Zhou" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="congjiangensis">
<emphasis id="932BF2820894C46DEC1C7D9D8B53FEE9" italics="true">B. congjiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="C2CDF35B4609D28429D1ED44775F7B7D" baseAuthorityName="Rao &amp; Yang" baseAuthorityYear="1997" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="daweimontis">
<emphasis id="7E7A9AD3D8287937CC06E1DA9C489F1A" italics="true">B. daweimontis</emphasis>
</taxonomicName>
,
<taxonomicName id="2CCD4CBBC50AB9405922F7C23F0D2109" baseAuthorityName="Zhang, Liang, Ran &amp; Shen" baseAuthorityYear="2012" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="fanjingmontis">
<emphasis id="BD44715D31CA96880B5D3B9019531AFB" italics="true">B. fanjingmontis</emphasis>
</taxonomicName>
,
<taxonomicName id="9DE4F65C33248229AF37875D9045EDD2" baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong &amp; Rowley" baseAuthorityYear="2018" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="fansipanensis">
<emphasis id="8E0C4281EEA4317734FA7CAE7CCE9025" italics="true">B. fansipanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="43137B08DB5B0C605B6216694A3C34E4" baseAuthorityName="Tapley, Cutaja, Nguyen, Portway, Mahony, Nguyen, Harding, Luong &amp; Rowley" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="frigida">
<emphasis id="8E02E1D6021ABE0C352BC42F0DDD0C66" italics="true">B. frigida</emphasis>
</taxonomicName>
,
<taxonomicName id="5C42BB3EF910F8EC74EC9249E0A19694" baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong &amp; Rowley" baseAuthorityYear="2018" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hoanglienensis">
<emphasis id="7B4A28A8C83CF3BAD350DD817F44002F" italics="true">B. hoanglienensis</emphasis>
</taxonomicName>
,
<taxonomicName id="A8F881E9AD2D7FD4FFD81E1654BE795C" baseAuthorityName="Liu, Li, Wei, Xu, Cheng, Wang &amp; Wu" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jiangi">
<emphasis id="A848CFD5C6A4E583DF722F5A0A1368B9" italics="true">B. jiangi</emphasis>
</taxonomicName>
,
<taxonomicName id="960CBF0D10D4E83F56F0D36D7CE07F1E" baseAuthorityName="Fei &amp; Ye" baseAuthorityYear="1983" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="0D227A7B98D1E3599A848B2CA95FCF6D" italics="true">B. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="3FB32EFE95D6B448943441155F021B1E" baseAuthorityName="Wang" baseAuthorityYear="2012" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jinggangensis">
<emphasis id="B0261B66457C902DE73CBFAFA820D8E0" italics="true">B. jinggangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="371645458C4617C6A9CFF29CC384607A" baseAuthorityName="Wang, Zeng, Lyu &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jiulianensis">
<emphasis id="527BE408C17010EDFAF105F230DBF444" italics="true">B. jiulianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="829653CC5551636E754EAAF9DA1EAFE7" baseAuthorityName="Li, Xu, Liu, Jiang, Wei &amp; Wang" baseAuthorityYear="2018" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="leishanensis">
<emphasis id="EC7666E852A285FFE1E7D6E35EBF502D" italics="true">B. leishanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="C78930BE650F7D6984BB566ABC92639F" baseAuthorityName="Zhang, Li, Xiao, Li, Pan, Wang, Zhang &amp; Zhou" baseAuthorityYear="2017" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="liboensis">
<emphasis id="09CD2B5EAC669EBF695605C4B72973A5" italics="true">B. liboensis</emphasis>
</taxonomicName>
,
<taxonomicName id="AF8D7268FBD6FA1C20392EA5D5FFE6E6" baseAuthorityName="Wang &amp; Yang" baseAuthorityYear="2014" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lini">
<emphasis id="D157C214704ADC6C87CD0A84F4C5251F" italics="true">B. lini</emphasis>
</taxonomicName>
,
<taxonomicName id="8733BC4ADDF7479D10DBEDC3D41556D0" baseAuthorityName="Shi, Li, Zhu, Jiang, Jiang &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lushuiensis">
<emphasis id="63960180AB95EC85A77FF4D8599D7258" italics="true">B. lushuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="15AA48C10E28FE7082945706B012B8D9" baseAuthorityName="Lyu, Wang &amp; Zhao" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="mirabilis">
<emphasis id="1DE5911DDDECC7AA2729A8B787661462" italics="true">B mirabilis</emphasis>
</taxonomicName>
,
<taxonomicName id="3455D116D283C8FBB700DD2D2C0C3F2F" baseAuthorityName="Wang, Lyu &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="mufumontana">
<emphasis id="AA613C0EA26D76017D11BFDF2E51C4B2" italics="true">B. mufumontana</emphasis>
</taxonomicName>
,
<taxonomicName id="1A7C94B076A33991E64AA5712EEC8CF9" baseAuthorityName="Lyu, Wang, Liu &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nanlingensis">
<emphasis id="5442AC70817C956A99B89115187AAFE3" italics="true">B. nanlingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="EB6306A121B5024F8B3E003FA4958297" baseAuthorityName="Liu" baseAuthorityYear="1950" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="omeimontis">
<emphasis id="C712C0BBF16E620A3C4258352D0C7873" italics="true">B. omeimontis</emphasis>
</taxonomicName>
,
<taxonomicName id="7D3C3D91A976E6DC6A48F55874BB1AAD" baseAuthorityName="Bourret" baseAuthorityYear="1937" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="palpebralespinosa">
<emphasis id="C724760791D862FD951DA64439F110F7" italics="true">B. palpebralespinosa</emphasis>
</taxonomicName>
,
<taxonomicName id="CD6087187B76C70D0B9F50D1A192A4DB" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="qianbeiensis">
<emphasis id="1E7112EE361D0AD2E79AE7D5D4254145" italics="true">B. qianbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="1DFBBC6AECF3801047A98AD60D4954B0" baseAuthorityName="Jiang, Ye &amp; Fei" baseAuthorityYear="2008" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sangzhiensis">
<emphasis id="B71B97889110930F8F48C390F928DB7D" italics="true">B. sangzhiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="84E22F7AA6EDD8E1DF138E8737393DAD" baseAuthorityName="Lyu &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sanmingensis">
<emphasis id="1D047C74F368AC7C14CB9CDD799FA181" italics="true">B. sanmingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="73F788D240F316F547CEE46A9644E6B9" baseAuthorityName="Lyu, Liu &amp; Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shimentaina">
<emphasis id="F80FFF8C5774C4550570B17967A347A8" italics="true">B. shimentaina</emphasis>
</taxonomicName>
,
<taxonomicName id="14DF84D175CD51B4F327F0E666A8769E" baseAuthorityName="Wang, Deng, Liu, Wu &amp; Liu" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shunhuangensis">
<emphasis id="5EB869893A0FB3E686F944DAB437E128" italics="true">B. shunhuangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="206F6D74F5852422419E9E3A0719C489" baseAuthorityName="Liu &amp; Hu" baseAuthorityYear="1973" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="spinata">
<emphasis id="93EDA55C909B72C3D76A6D76C3CA404C" italics="true">B. spinata</emphasis>
</taxonomicName>
,
<taxonomicName id="E34B86F7307E94E91B335B1BCA5FEC58" baseAuthorityName="Lyu &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sanmingensis">
<emphasis id="1F8BC3B6EFD840112F4FFB3B78740CF9" italics="true">B. sanmingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="B652BE8388D418787066CCEE74616804" baseAuthorityName="Wang &amp; Lyu" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="tongboensis">
<emphasis id="397FA37E6D0657AD58FA56F8C458451E" italics="true">B. tongboensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6866E07DF5AC6CD45574DDEB3124933D" baseAuthorityName="Shen, Mo &amp; Li" baseAuthorityYear="2010" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="tuberogranulatus">
<emphasis id="B9A439E1FD4881EC73FF546D404F6A2E" italics="true">B. tuberogranulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="FE2C4F6784481548AA8FB25A1266D3D8" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="wuliangshanensis">
<emphasis id="F68B21C24DCBBF69EC2E223526EBD196" italics="true">B. wuliangshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="B1AB0DFD5D5152EF03B924EA6A634E57" baseAuthorityName="Wang, Wu, Peng, Shi, Lu &amp; Wu" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xianjuensis">
<emphasis id="A45C2780216F9084E90B90B553FBE4E4" italics="true">B. xianjuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="76975B23A432CDF7D6078C3DFF161D3D" baseAuthorityName="Lyu, Zeng &amp; Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yangmingensis">
<emphasis id="B1F3D471C98B10AA6C5FF02458360088" italics="true">B. yangmingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="1C0A3A12E7159B553319F3106B7FD6A2" authorityName="Qi, Mo, Lyu, Wang &amp; Wang" authorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yaoshanensis">
<emphasis id="AC91795EEC4F34C0DC4A42F572D49C93" italics="true">B. yaoshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="DE35F143E855FD23C572153EF34AB52F" authorityName="Qi, Lyu, Wang &amp; Wang" authorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yingdeensis">
<emphasis id="807D84A015638AC26577C6AD5D2100E3" italics="true">B. yingdeensis</emphasis>
</taxonomicName>
(vs. heels overlapping), from
<taxonomicName id="9202F481B0A9F28C289193194958911D" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="binchuanensis">
<emphasis id="5E4E2113D1C84A18A1BBFCEABE643B11" italics="true">B. binchuanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="07128C59E7681CDC8A79C4CEE8135883" authorityName="Zeng, Wang, Chen, Xiao, Zhan, Li &amp; Lin" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="elongata">
<emphasis id="1EB938B26E504534504F523C6A943017" italics="true">B. elongata</emphasis>
</taxonomicName>
,
<taxonomicName id="71FDC347E7C15092AF1AED184860896E" baseAuthorityName="Wang, Liu &amp; Jiang" baseAuthorityYear="2017" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lishuiensis">
<emphasis id="63DF000FEC90555ED20882F0ABFC14BF" italics="true">B. lishuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="1D0E02B7F84875E9F78B0A3A64DBD771" baseAuthorityName="Stejneger" baseAuthorityYear="1926" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="minor">
<emphasis id="4DE23E4F826F44067DAB9FC30A569676" italics="true">B. minor</emphasis>
</taxonomicName>
,
<taxonomicName id="342EA315ECF2CBBD86CEF336E997E616" baseAuthorityName="Lyu, Zeng &amp; Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xiangnanensis">
<emphasis id="2CF27856E9B4A7AB55457081B5EF7989" italics="true">B. xiangnanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="2001AC8866CF1E5B3D4D094F1C6F0297" authorityName="Lyu &amp; Wang" authorityYear="2023" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xuefengmontis">
<emphasis id="B4FC2119858431BB13BCE9EA172B3A10" italics="true">B. xuefengmontis</emphasis>
</taxonomicName>
(vs. heels just meeting), and from
<taxonomicName id="E3E5776EA98348CFCE78BAD8B4975415" baseAuthorityName="Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy &amp; Che" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="angka">
<emphasis id="B8704DE6695727D995064393CAD3E4FA" italics="true">B. angka</emphasis>
</taxonomicName>
,
<taxonomicName id="111AAA49A0B2075171561A6C224EE91C" baseAuthorityName="Lyu, Wang &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="daiyunensis">
<emphasis id="F474135F640EA4BA5E54DBC8EEF22E26" italics="true">B. daiyunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="01171BBFB09401CFB1B25551A400ECEE" baseAuthorityName="Ye, Fei &amp; Xie" baseAuthorityYear="2007" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="baolongensis">
<emphasis id="10EA953861E9084143EB6EB9A346EA2E" italics="true">B. baolongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="A9B80A2497A0FDD0F455B2775DB84B2E" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="wushanensis">
<emphasis id="331E16E99C71CA01F6BCA9ED8D02DF44" italics="true">B. wushanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="72E21FDD0071D72C987D83A144601A5C" authorityName="Qi, Wang, Lyu &amp; Wang" authorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yunkaiensis">
<emphasis id="E760581733EBCE98A223E5FC2259AFB7" italics="true">B. yunkaiensis</emphasis>
</taxonomicName>
(vs. heels just meeting or slightly overlapping).
</paragraph>
<paragraph id="17C657E13DDC77522A03382FC364D2AA">
<taxonomicName id="E30C43F588CFDB4414B418257EA4BF73" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="69CAA55C486EAC2A901EE0962BE559DE" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="785D093D2A8FC3147026BA22E1DBA76E" rank="species">sp. nov.</taxonomicNameLabel>
can easily be distinguished from the following congeners by its tongue not notched distally:
<taxonomicName id="101CFBEE877C4C94A3C8C60976154AAB" baseAuthorityName="Inger &amp; Romer" baseAuthorityYear="1961" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="brachykolos">
<emphasis id="31EC3ABCCB69C0954CD396357BC0378D" italics="true">B. brachykolos</emphasis>
</taxonomicName>
,
<taxonomicName id="B5250C18C47EEF3E3F8C0598BAF2DD7F" baseAuthorityName="Wang, Liu, Lyu, Zeng &amp; Wang" baseAuthorityYear="2017" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="insularis">
<emphasis id="E223C53E64471E8BF4CC42F2809A2163" italics="true">B. insularis</emphasis>
</taxonomicName>
,
<taxonomicName id="F0D64AE1A50E2EAA85501BD092A207A4" baseAuthorityName="Wang &amp; Zeng" baseAuthorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="pepe">
<emphasis id="6E26BA6DA4AD4B7860C39EC35D39C7B4" italics="true">B. pepe</emphasis>
</taxonomicName>
(vs. tongue notched distally).
<taxonomicName id="BF37737B0F952B1A61709DCF224D79F0" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="EFF88ADDCEB335B28D8F35E34F207BF7" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="552D32E5750602D119EE88F9FBAD11E6" rank="species">sp. nov.</taxonomicNameLabel>
can easily be distinguished from the following congeners by its presence of vomerine teeth:
<taxonomicName id="142BBB7FCEEDB8B5C5954DA28A917DB5" baseAuthorityName="Wang, Li &amp; Jin" baseAuthorityYear="2014" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="acuta">
<emphasis id="0C74A7D9E9B65D6C4ED32A39FAD81BF2" italics="true">B. acuta</emphasis>
</taxonomicName>
,
<taxonomicName id="FCD16EE464725D139455A8B8158425E4" baseAuthorityName="Boulenger" baseAuthorityYear="1899" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="0E1EFCF3CCD07949498A525A3830A1AF" italics="true">B. boettgeri</emphasis>
</taxonomicName>
,
<taxonomicName id="65DE6A3EB264C178624A1138B85563E6" baseAuthorityName="Nguyen, Pham, Nguyen, Luong &amp; Ziegler" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="caobangensis">
<emphasis id="053B8F644D573530D913532D1CC3219D" italics="true">B. caobangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="EC9FFF6A6B66F9510DBB0E53B0A8D93D" baseAuthorityName="Lyu, Zeng, Wang &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="daoji">
<emphasis id="677AE47755B8DED4A0BBD7B4CB6FC56A" italics="true">B. daoji</emphasis>
</taxonomicName>
,
<taxonomicName id="28C0A60B961EF28738E3370E097B7298" authorityName="Wang, Zeng, Lyu, Xiao &amp; Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hungtai">
<emphasis id="254C7680A45062108AE0735DB344D760" italics="true">B. hungtai</emphasis>
</taxonomicName>
,
<taxonomicName id="7124B97207AF3BAB4BF2107678259160" authorityName="Qian, Hu, Mo, Gao, Zhang &amp; Yang" authorityYear="2023" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hengshanensis">
<emphasis id="A60DE18E62ECCB925CF6A304404E1BBE" italics="true">B. hengshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="2862DA29E43734D9EA8C5E1E94612842" baseAuthorityName="Pope" baseAuthorityYear="1929" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="CED8C3860F4DBFDB01142EF323383E79" italics="true">B. kuatunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="5E56EA7928F5BDE7A6FFFB41812D2687" baseAuthorityName="Messenger &amp; Dahn" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ombrophila">
<emphasis id="64E79D0D5F12892B7973357F1A5EEE90" italics="true">B. ombrophila</emphasis>
</taxonomicName>
,
<taxonomicName id="09201E7967E82E6588F5C64420225A4D" baseAuthorityName="Wang, Li &amp; Zhao" baseAuthorityYear="2014" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="obesa">
<emphasis id="2711C568D0A531BDCBE79C0CD832CAD0" italics="true">B. obesa</emphasis>
</taxonomicName>
,
<taxonomicName id="55F521B8AE119FDDBDBB57CFFFD24F1A" baseAuthorityName="Tian &amp; Sun" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shuichengensis">
<emphasis id="BC0A5DA1650A29BBB73594727D224591" italics="true">B. shuichengensis</emphasis>
</taxonomicName>
,
<taxonomicName id="1FBF22B5F89B1256CE58F93B85E8ED0E" baseAuthorityName="Wang, Lyu &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="wugongensis">
<emphasis id="47A79CFE7209ACD75BAE9CFF62CD3713" italics="true">B. wugongensis</emphasis>
</taxonomicName>
(vs. vomerine teeth absent).
</paragraph>
<paragraph id="C2448FFC024885178C202432797A89B7">
<taxonomicName id="FFAE7CEEA6F21DCC32066BEF3B8803CD" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="235B2E6B3D6AB00FCBAA9A046D740D5B" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9E751C4FDE68D5F12E17D38C9FFEFEB6" rank="species">sp. nov.</taxonomicNameLabel>
can easily be distinguished from the following congeners by its absence of lateral fringes on webbing on toes:
<taxonomicName id="03AA7123829FA53E468F85151C124348" baseAuthorityName="Wang &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="dongguanensis">
<emphasis id="F58085F19BCC1CB8189E8F866F74BEEB" italics="true">B. dongguanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="79E874B00651E705F3ADC47C0BEC3040" authorityName="Wang, Zeng, Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="fengshunensis">
<emphasis id="763A7A58D40355ED36F5B0AE1043EC95" italics="true">B. fengshunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="09802D9FCF59BDFEA3084CFD61BFD506" baseAuthorityName="Wang, Zeng &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nankunensis">
<emphasis id="6D94D9F8B13BB107A7605B198F65FD9F" italics="true">B. nankunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="0D11DA042BA41DECFB8AC102ADA9D9DC" authorityName="Wang, Zeng, Lyu, Xiao &amp; Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="puningensis">
<emphasis id="E27EA79349C947CDA2CDD1F730181563" italics="true">B. puningensis</emphasis>
</taxonomicName>
(vs. toes with rudimentary webbing), and from
<taxonomicName id="92D463A73D2E38606C912C4B2B3CE143" baseAuthorityName="Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong &amp; Rowley" baseAuthorityYear="2017" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rubrimera">
<emphasis id="2A4A7530615DEBD25BC51B9518E4CB13" italics="true">B. rubrimera</emphasis>
</taxonomicName>
(vs. toes with narrow lateral fringes).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFEAI" type="distribution">
<paragraph id="DBE0BDFE48A1A74ACC7DFF4FDF624C66">
<heading id="893C6222C352E659C09A3073EAEE9676" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="CBE97A313C6BB2BA511D81F4B8BA1FA9">
Currently,
<taxonomicName id="9AB26459ABE07966F7D4B14C24B9B491" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="B005AAA343905782967722A00A7B41DE" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44D9B16CC3637B160882B5D1EA738336" rank="species">sp. nov.</taxonomicNameLabel>
is only known from the coastal hills of Ningde City, eastern
<collectingRegion id="EA3E3E471C223FB70450FD45BB169646" country="China" name="Fujian">Fujian Province</collectingRegion>
,
<collectingCountry id="B7F545A70E114ABDFCF3A7E6AC0EF83C" name="China">China</collectingCountry>
. It inhabits flowing montane seeps and the nearby forest floor and leaf litter. The habitat is surrounded by secondary forest mixed with bamboo groves at elevations between
<quantity id="C3A6A9743B8888C3012887403DAAB7E4" metricMagnitude="2" metricUnit="m" metricValue="3.3" metricValueMax="5.1" metricValueMin="1.5" unit="m" value="330.0" valueMax="510.0" valueMin="150.0">
150
<specimenCount id="7F72168ACBCF1329AC55E71405C70DA5" count="510" type="generic">510 m.</specimenCount>
</quantity>
Advertisement calls of males were heard from February to May. Males were found calling in rock crevices.
</paragraph>
</subSubSection>
</treatment>
</document>