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<document id="A64CD7632CF8FEF3B8E5714D8374168E" ID-CLB-Dataset="304586" ID-DOI="10.3897/zookeys.1216.128080" ID-GBIF-Dataset="6379ab15-2e99-467e-b16a-328ab30bb6b2" ID-ZooBank="9FAD26B7-8734-4CFC-94A6-66211BECF33F" ID-publisher-id="128080" URI-arpha="684B2E7F-2598-5003-B982-6CA9AE1761FF" URI-zoobank="http://zoobank.org/9FAD26B7-8734-4CFC-94A6-66211BECF33F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="rapid-communication" checkinTime="1729263425651" checkinUser="pensoft" docAuthor="Wu, Zhenfei, Zhang, Sihang, Qin, Fuxue &amp; Cong, Peiyun" docDate="2024" docId="55752B89A0455C0DB3777486D658CA67" docLanguage="en" docName="ZooKeys 1216: 17-26" docOrigin="ZooKeys 1216" docSource="https://zookeys.pensoft.net/article/128080/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Trichopeltis jiyue Wu, Zhang, Qin &amp; Cong, 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="26" masterDocId="684B2E7F25985003B9826CA9AE1761FF" masterDocTitle="A new epigean species of Trichopeltis Pocock, 1894 from southwest China (Diplopoda, Polydesmida, Cryptodesmidae)" masterLastPageNumber="26" masterPageNumber="17" pageNumber="17" updateTime="1729275551043" updateUser="ExternalLinkService">
<mods:mods id="373E6E376905F604D153A1F8A7F21301" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="522721108F397090F978FF6F1435977F">A new epigean species of Trichopeltis Pocock, 1894 from southwest China (Diplopoda, Polydesmida, Cryptodesmidae)</mods:title>
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<mods:name id="E47FBEF53052DFF91D9BC1E70A3F5F58" type="personal">
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<mods:namePart id="1B235E16B4B08AE68F6DB2E24CE0D909">Wu, Zhenfei</mods:namePart>
<mods:nameIdentifier id="9AE38725777041AED52F366381CE2E2F" type="ORCID">https://orcid.org/0009-0004-5586-6511</mods:nameIdentifier>
<mods:affiliation id="A7A02130557BAB4A27A9E1586BD56D4F">Yunnan Key Laboratory for Palaeobiology &amp; MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming 650500, China</mods:affiliation>
</mods:name>
<mods:name id="609B81348DBAB820C486BE1EAF36CA4C" type="personal">
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</mods:role>
<mods:namePart id="EFBA41F1BCB72AEA3BB1879825BD9F3D">Zhang, Sihang</mods:namePart>
<mods:nameIdentifier id="218AB94ABD940BA75D364A240CCA5017" type="ORCID">0000-0002-8230-6892</mods:nameIdentifier>
<mods:affiliation id="8C5612F627EB2F691041D56776F851BC">Yunnan Key Laboratory for Palaeobiology &amp; MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming 650500, China</mods:affiliation>
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<mods:namePart id="5B8D9F3E8433F51093CCA8CE1F8862FC">Qin, Fuxue</mods:namePart>
<mods:nameIdentifier id="2F864BD34D40D89EF073638085C241E8" type="ORCID">https://orcid.org/0009-0001-1140-3927</mods:nameIdentifier>
<mods:affiliation id="69455397EAF0372FDEBA752999304E35">Earthquake Prevention and Disaster Reduction Bureau of Zhaoyang, Zhaotong 657099, China</mods:affiliation>
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<mods:namePart id="16B6217C12387C3BD4E71E8E4DC8A6B5">Cong, Peiyun</mods:namePart>
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<mods:affiliation id="894A2D5656D741D5C3B79A03B4BCF893">Yunnan Key Laboratory for Palaeobiology &amp; MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming 650500, China</mods:affiliation>
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<mods:date id="9801218C8533B092994FEB2C920C966D">2024</mods:date>
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<mods:number id="52B21558DA7EEAD18CCFF164090DAD33">2024-10-18</mods:number>
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<mods:number id="DD839BAB8BE2BE05E715B3B3CAE6EF2E">1216</mods:number>
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<treatment id="55752B89A0455C0DB3777486D658CA67" ID-DOI="http://doi.org/10.5281/zenodo.13952662" ID-Zenodo-Dep="13952662" ID-arpha="55752B89-A045-5C0D-B377-7486D658CA67" ID-zoobank="https://zoobank.org/77367C20-B420-457F-A095-AB493A6BB2F7" LSID="urn:lsid:plazi:treatment:55752B89A0455C0DB3777486D658CA67" LSID-ZBK="urn:lsid:zoobank.org:act:77367C20-B420-457F-A095-AB493A6BB2F7" httpUri="http://treatment.plazi.org/id/55752B89A0455C0DB3777486D658CA67">
<subSubSection id="8608866FFFC26BF3E9E2A160DAC12108" type="nomenclature">
<paragraph id="6BCB1CA9E9549844FC1DC5C2780AE019">
<taxonomicName id="5A6C46ED3B32E4676C58A7C7EB1B99B4" authorityName="Wu &amp; Zhang &amp; Qin &amp; Cong" authorityYear="2024" class="Diplopoda" family="Cryptodesmidae" genus="Trichopeltis" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="jiyue" status="sp. nov.">
<emphasis id="CCEFF376759775CA6B510A4BD50A9B4B" italics="true">Trichopeltis jiyue</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DF22190D2B6275E82890475A303DA75D" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERGAC" type="type material">
<paragraph id="E06B9B7B5945EC696290107A46811619">
<heading id="1F64232D8AC7E3E97D290A8032AB295F" reason="title">Type material.</heading>
</paragraph>
<paragraph id="89D8ACF2E382C99DF57CE85BECB19A8F">
<materialsCitation id="3F8A731A3C6F08A500397C47C5D73B40" collectingDate="2021-10-04" collectorName="Peiyun Cong &amp; Sihang Zhang &amp; Zhenfei Wu &amp; Fuxue Qin" country="China" county="Jingdong Yi Autonomous County" elevation="2450" location="Pu'er City" longitude="101.06612" specimenCode="YNU-MD 0151" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
<emphasis id="087167AF630570DDED404840C731790E" bold="true" italics="true">
<typeStatus id="AF04ADB5BA18A0E9CDC97D763FD81F68" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<specimenCount id="4D41FE9083A6882A2337ACE7607F1B5C" count="1" type="male"></specimenCount>
(
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),
<collectingCountry id="DB66AD2E38289F17F0448406F00C4F1F" name="China">China</collectingCountry>
,
<collectingRegion id="19B93EC95FEBAF6D09FE3565509D2CCC" country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<location id="51741E067C69468F991B7C0DF51AF918" LSID="urn:lsid:plazi:treatment:55752B89A0455C0DB3777486D658CA67:51741E067C69468F991B7C0DF51AF918" country="China" county="Jingdong Yi Autonomous County" longitude="101.06612" name="Pu'er City" stateProvince="Yunnan Province">Puer City</location>
,
<collectingCounty id="D954C23F15D19A2B088C4BC3DE4919DD">Jingdong Yi Autonomous County</collectingCounty>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6GAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.066111,24.921667]}">
24 ° 54 ' 78 &quot; N,
<geoCoordinate id="0C8BA8B539C431D6408ACC36350C77AB" degrees="101" direction="east" minutes="03" orientation="longitude" precision="15" seconds="58" value="101.06612">101 ° 03 ' 58 &quot; E</geoCoordinate>
</named-content>
,
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<elevation id="A699A36B442F805944BA879FF88B97BD" metricMagnitude="3" metricUnit="m" metricValue="2.45" unit="m" value="2450.0">2450 m</elevation>
</quantity>
elev.,
<date id="8CE4EBEA7216A810373B183437B18A2C" value="2021-10-04">
<collectingDate id="21586175EF8E5EB220051891E9633F73" value="2021-10-04">4. X. 2021</collectingDate>
</date>
, leg.
<collectorName id="A8F5E868997168947405DA96ABF21C57">Peiyun Cong</collectorName>
,
<collectorName id="1A1E6FD68949CEDAA9AEBF68663F5135">Sihang Zhang</collectorName>
,
<collectorName id="6246667E082F05B13216F336D93AC566">Zhenfei Wu</collectorName>
&amp;
<collectorName id="D617C656AB972507789FE1F06638908B">Fuxue Qin</collectorName>
</materialsCitation>
.
<materialsCitation id="69B89459E947AC05484D8E9BDECC2D22" collectingDate="2021-10-04" collectorName="Peiyun Cong &amp; Sihang Zhang &amp; Zhenfei Wu &amp; Fuxue Qin." country="China" county="Jingdong Yi Autonomous County" elevation="2450" location="Pu'er City" longitude="101.06612" specimenCode="YNU-MD 0152-165" specimenCount="13" specimenCount-female="9" specimenCount-male="4" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="BC841002DF361083A24375C6B0D4244F" bold="true" italics="true">
<typeStatus id="06239273DD03DDB9EFA83A8A698A6BD6" type="paratype">Paratypes</typeStatus>
</emphasis>
: •
<specimenCount id="82BD42BE6AC00FCEBBB8240DEEF1D588" count="4" type="male">4 ♂</specimenCount>
,
<specimenCount id="4B11F753C1FF7B10C54730C8042818B5" count="9" type="female">9 ♀</specimenCount>
(
<specimenCode id="CFF305E1506A65D3682DB0ACC529E6BF">YNU-MD 0152-165</specimenCode>
) same location as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHHAC" type="etymology">
<paragraph id="B02F8FDD1F0EF667640B711E20055B25">
<heading id="372B329631EA6697FE7EE5FCD4238E7D" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="CA7E0844B8958B4700C974DB9FA37A4E">Jiyue (Chinese spelling) alludes to the bright white appearance when the animal emerges from the leaf mold, like the moon appearing from behind a dark rain cloud.</paragraph>
</subSubSection>
<subSubSection id="SECID0EMHAC" type="diagnosis">
<paragraph id="6118BD008677075CD8492544CCB52C20">
<heading id="C234AA6CBBC6A79C6830CB4B6CE29F6B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="70F64C48E4FB2A477D33ABFC4C6F7BB9">
<taxonomicName id="43EA5C6C6A5C6D3B4502AAD11F80D53B" authorityName="Pocock" authorityYear="1894" class="Diplopoda" family="Cryptodesmidae" genus="Trichopeltis" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
<emphasis id="0B04384EA575E66B5501F6AC0A466308" italics="true">Trichopeltis</emphasis>
</taxonomicName>
is characterized by the relatively long and stout setae on the gonopodal coxae, with the posterior part having two conspicuous wing-like processes (cxp); gonopodal telopodites glabrous and four-branched; and the acropodite curved caudolaterad. The living animal is uniformly bright white.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZHAC" type="description">
<paragraph id="D9326C0EE257C2C9FB3EA014518EBF46">
<heading id="81FF7AA17DF484CFAE4FE92BA286694C" reason="title">Description.</heading>
</paragraph>
<paragraph id="BA6D7380BFDA4BDC3F7D89B7458FC179">
Length of
<specimenCount id="0B22669F2803E910743709D34D50918F" count="1" type="male"></specimenCount>
ca
<quantity id="0568F464691A151AB2AA6ECC17887782" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.78" metricValueMin="1.72" unit="mm" value="17.5" valueMax="17.8" valueMin="17.2">17.217.8 mm</quantity>
,
<typeStatus id="CCAF493A3B2A96CB7FBC81B2840085CE" type="paratype">paratype</typeStatus>
<specimenCount id="5C1BA237FFEF9A99E4152F397277C61B" count="1" type="female"></specimenCount>
ca 17.0
<quantity id="5ED8AFEDE3B25B2F8397DC5B743A2BEC" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" unit="mm" value="17.4">17.4 mm</quantity>
, width of midbody pro- and metazonae
<quantity id="33846338653EE374A8580DB8995D9D0A" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.22.4 mm</quantity>
and
<quantity id="B2189554E371F4071A7074AD3288A2DE" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.4" metricValueMin="5.3" unit="mm" value="5.35" valueMax="5.4" valueMin="5.3">5.35.4 mm</quantity>
(
<specimenCount id="282FBB0969C9B94F96C0908253EFAB33" count="1" type="male"></specimenCount>
),
<quantity id="6A07AC8B46EF22994D6B7D9A30ACD7A2" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.22.5 mm</quantity>
and
<quantity id="424BE6BBBEDCC5D61F82E911B05EE4BC" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.4" metricValueMin="5.1" unit="mm" value="5.25" valueMax="5.4" valueMin="5.1">5.15.4 mm</quantity>
(
<specimenCount id="AC4AE4C57EB95939037310DB8F3BFAE2" count="1" type="female"></specimenCount>
), respectively.
</paragraph>
<paragraph id="7A7B01F5225B2657E9907C3FEDE83AE3">
<emphasis id="7EEB57AF5686390623623CEB8E19A34C" bold="true" italics="true">Coloration</emphasis>
of tergites uniformly bright white (Fig.
<figureCitation id="2849E909E6D4D4A72542EFAD7B7A3009" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. External morphology and colouration of Trichopeltis jiyue sp. nov. A ♂ holotype in habitus and live B fed 12 months in laboratory C, D ♀ paratype, after 3 months storage in 75 % alcohol. Scale bars: 10 mm (A); 5 mm (B D)." figureDoi="10.3897/zookeys.1216.128080.figure1" httpUri="https://binary.pensoft.net/fig/1156162">1 A</figureCitation>
); fed 12 months with local mor and leaves, yellow (Fig.
<figureCitation id="54ABE5DE07E69F8319F1B3FBDEFA3553" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. External morphology and colouration of Trichopeltis jiyue sp. nov. A ♂ holotype in habitus and live B fed 12 months in laboratory C, D ♀ paratype, after 3 months storage in 75 % alcohol. Scale bars: 10 mm (A); 5 mm (B D)." figureDoi="10.3897/zookeys.1216.128080.figure1" httpUri="https://binary.pensoft.net/fig/1156162">1 B</figureCitation>
); in alcohol, after months of preservation, whitish-yellow to yellow (Fig.
<figureCitation id="4941AE318722FCCAD69E0B5DB6B3833D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. External morphology and colouration of Trichopeltis jiyue sp. nov. A ♂ holotype in habitus and live B fed 12 months in laboratory C, D ♀ paratype, after 3 months storage in 75 % alcohol. Scale bars: 10 mm (A); 5 mm (B D)." figureDoi="10.3897/zookeys.1216.128080.figure1" httpUri="https://binary.pensoft.net/fig/1156162">1 C, D</figureCitation>
). Antenna whitish-yellow (proximal) to reddish-purple (distal).
</paragraph>
<caption id="21BB8D7C46B8C2C0EBE1AB071AD9813D" ID-DOI="10.3897/zookeys.1216.128080.figure1" ID-arpha="B4318DBC-127F-5371-8F6D-B4DB7D07BFD0" httpUri="https://binary.pensoft.net/fig/1156162" startId="F1">
<paragraph id="99F30F6A2B305581E7743DADBC990769">
<label id="52B33D927878404FBFD12F452ACF409D">Figure 1.</label>
</paragraph>
<paragraph id="40E4EE1623181C58DCC3F0476FB015D3">
External morphology and colouration of
<taxonomicName id="1CDD0ED5F23C202C84C8F9D37586BD3D" authorityName="Wu &amp; Zhang &amp; Qin &amp; Cong" authorityYear="2024" class="Diplopoda" family="Cryptodesmidae" genus="Trichopeltis" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="jiyue" status="sp. nov.">
<emphasis id="8E132D67C9A0DF95D1BB5CE2DAA57640" italics="true">Trichopeltis jiyue</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3298C10728F6C89F5A6339DA432AC726" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="0A1E0063613225BBE58A972F60D36C27" bold="true">A</emphasis>
♂ holotype in habitus and live
<emphasis id="DEB9B1D2A06C4D5A8C98A6CDC296094A" bold="true">B</emphasis>
fed 12 months in laboratory
<emphasis id="6FBA2471070C3FFCFDE7186BD9923481" bold="true">C, D</emphasis>
♀ paratype, after 3 months storage in 75 % alcohol. Scale bars: 10 mm (
<emphasis id="20C8EFEA73DDDED97DE8F230536FF360" bold="true">A</emphasis>
); 5 mm (
<emphasis id="115C00C205BFDD2BEC7E184AAC1519D8" bold="true">B D</emphasis>
).
</paragraph>
</caption>
<paragraph id="3CE836DCFAD36EDFEBB693617091A113">
<emphasis id="0F627ACF80FAE2E4ED243F4B1A6C6F29" bold="true" italics="true">Adults body</emphasis>
with 20 segments, collum plus 17 podous and 1 apodous tergites, plus 1 telson. In width, head &lt;&lt; collum &lt;segment 2 &lt;3 &lt;4 &lt;5 &lt;6 &lt;7-17, thereafter body tapered towards telson.
</paragraph>
<paragraph id="16617EE3DD794D5A09B6A778E142F2FD">
<emphasis id="C3F764CF6D6EE5ED6524016F3DB6149D" bold="true" italics="true">Head</emphasis>
sparsely pilose, epicranial suture present (Fig.
<figureCitation id="611AD69A9DEE868C26594E993E7CE5DC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 A</figureCitation>
). Antennae short and clavate, reaching tergite 4 when stretched ventrally; in length, antennomere 6&gt; 3&gt; 2 = 4&gt; 5&gt; 1&gt; 7 (Fig.
<figureCitation id="B61A5A9D01E38AF26B2C367B1E08D49D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 A</figureCitation>
); antennomeres 57 each with a bacilliform sensilla field apico-laterally, the numbers of bacilliform sensilla are 100, 67, and 34, respectively.
</paragraph>
<caption id="DC26C2360BCACD4EF0020312CFD16D00" ID-DOI="10.3897/zookeys.1216.128080.figure2" ID-arpha="E7821E50-8057-51D5-8F67-55269D44AA09" httpUri="https://binary.pensoft.net/fig/1156163" startId="F2">
<paragraph id="E70FC53A3946CCAF95619471199D40D5">
<label id="62FE3CBAE31B834E20F2DBDB3376D59D">Figure 2.</label>
</paragraph>
<paragraph id="442D225DF0DDF721BFD26FF537687FB9">
<abbrev id="ABBRID0ERKAC" xlink_title="Scanning electron microscope">SEM</abbrev>
images of
<taxonomicName id="71712BE5DB8231A3A7092D11B3A72E68" authorityName="Wu &amp; Zhang &amp; Qin &amp; Cong" authorityYear="2024" class="Diplopoda" family="Cryptodesmidae" genus="Trichopeltis" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="jiyue" status="sp. nov.">
<emphasis id="DB2F234B24E29F6A4016E9260E5F6959" italics="true">Trichopeltis jiyue</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70FE5C11683E705811BAB9A224BC1E17" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ holotype
<emphasis id="1303D56B59412E8F8495E860832DAEC1" bold="true">A</emphasis>
head, dorsal view
<emphasis id="72CA13FA915084FEC892E58C56302E09" bold="true">B</emphasis>
collum and the second segment, dorsal view
<emphasis id="328320212EA7C0A482D82CBB0B1BA5C2" bold="true">C</emphasis>
segments 69, dorsal view
<emphasis id="59C0E911A2E88560D0E65E53F917A88E" bold="true">D</emphasis>
cross-section of segment 5, caudal view
<emphasis id="07CC9A558EB30D6CA30282DDBE3C2916" bold="true">E</emphasis>
antenna disc coeloconic sensilla, plan view
<emphasis id="FC73C0C4A3AAAAC1381B4CD59A62E9DA" bold="true">F</emphasis>
telson and anal, ventral view
<emphasis id="C59DBD1590E1A54E36CEB81A402C788A" bold="true">G</emphasis>
limbus of segment 5, subventral view
<emphasis id="BEBD24D8D070B3F4C9F5CA25E555BE96" bold="true">H</emphasis>
claw of leg, subventral view
<emphasis id="C3617931C62990A7D7204E6706102A43" bold="true">I</emphasis>
paraterga of segment 6, ventral view. Scale bars: 1 mm (
<emphasis id="806D66D323E11EC4ED860669D04229CF" bold="true">A D</emphasis>
); 50 μm (
<emphasis id="1EACF1F0E00B9107B8E96533590AF465" bold="true">E, H</emphasis>
); 20 μm (
<emphasis id="51A39D670A66D3D8880A5C01BCDD0D4F" bold="true">G</emphasis>
); 500 μm (
<emphasis id="5A7F33C1B513778F922412A5FBACE04C" bold="true">F, I</emphasis>
).
</paragraph>
</caption>
<paragraph id="3F9AF4B3562245A3C928A9159F0786ED">
<emphasis id="766475415B16F55E0DDAD0B33D012CCA" bold="true" italics="true">Collum</emphasis>
completely covering the head from above, inverted subtrapeziform, regularly convex at peripheric margin, caudal margin slightly concave (Fig.
<figureCitation id="FC54B4BE5B60F738324F7B4E0E76FD2E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 B</figureCitation>
); arranged with 12 or 13 regular, transverse rows of small, spherical, setigerous tubercles on the surface, tubercles 8-13 + 8 - 13 per row, surrounded with spherical granulations, seta on each tubercle directed caudad (Fig.
<figureCitation id="3ECBA89BB944521D75C4912D1828B575" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 B</figureCitation>
).
</paragraph>
<paragraph id="30DEC13C2BA849DE5672A54E17478F68">
<emphasis id="9DE296B42D386716248ADC8894E1CE7A" bold="true" italics="true">Prozona</emphasis>
of segments following collum finely shagreened, metazona densely tuberculate and setose; fore and caudolateral margins of collum, anterolateral, lateral and caudal margins of following paraterga of segments besides telson with obvious dentiform-lobulate lobules, smallest at mid-dorsal region and slightly larger bidirectionally at caudal margins of paraterga.
</paragraph>
<paragraph id="CC72F4B4612DFE861AC58CA8AFE01F8D">
<emphasis id="AED9ECA4B2F83D4B64163DFD0130F0D7" bold="true" italics="true">Dorsum</emphasis>
convex, postcollum paraterga flat, very broad and long, narrowly rounded laterally, axial line absent. Metatergal segments 216 with four or five irregular transverse rows of similarly small, spherical, setigerous tubercles. Tubercles decreasingly extend to paraterga, but each of the latter only with three or four irregular rows of similar tubercles (Fig.
<figureCitation id="46A6F87627B083CB0CF50336D21999DA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 C</figureCitation>
), surrounded by spherical granulations, same to collum; following metatergal segments 17 and 18 with 68 rows of smaller tubercles.
</paragraph>
<paragraph id="AA6C01A244D7AFF14627ACA25A61497A">
<emphasis id="B4535EDEDE591B993E6F478CA0CE2704" bold="true" italics="true">Paraterga</emphasis>
very strongly developed (Fig.
<figureCitation id="6286ECB83DE292ABB91F6DD9491DE42E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 C</figureCitation>
), regularly declivous, the tips extending down below level of venter (Fig.
<figureCitation id="610BF75D65239DE53C18EFB19E1B1800" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 D</figureCitation>
). Segments 215 slightly projecting forward, each with 69 small, crown-like dentiform, lateral lobules (Fig.
<figureCitation id="8426019C2ABAD4DBA9878E2C13A693C5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 I</figureCitation>
) and 79 tongue-shaped to squarish caudolateral lobules; all evident, setigerous and microvillose segments 1619 projecting caudally, each with 57 small, crown-like dentiform, lateral lobules and 913 tongue-shaped to squarish, caudolateral lobules; all evident, setigerous, and microvillose.
</paragraph>
<paragraph id="F505BE38545D02A4DE7F8ADCBF850F88">
<emphasis id="25A22276B9DEC801D65DB03E110EA0DA" bold="true" italics="true">Sterna</emphasis>
sparsely setose; axial line present; tergite stricture divided into pro- and metazone parts. Limbus, with a row of tongue-shaped lobules, microdenticulate apically (Fig.
<figureCitation id="AE4492BA2F9A5BD50789634DAA84E068" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 G</figureCitation>
). Pore visible, lying on the ventral paraterga of segment 5, ozopores formula not discernable.
</paragraph>
<paragraph id="9B03820CAA0646DB129E1E9655E4A0DB">
<emphasis id="6BCDAD46D1842E443FD72BC3B159566D" bold="true" italics="true">Telson</emphasis>
(Fig.
<figureCitation id="9132E2518761F0CF56D6E00396485BD0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 F</figureCitation>
) conical, with numerous spherical granulations; epiproct flattened dorsoventrally, microtuberculate, with four strong apical papillae. Hypoproct roundly subtrapeziform, 1 + 1 caudal setae separated, surface rugged.
</paragraph>
<paragraph id="B774DC6713F4F0DCAF279DEC5EBA5265">
<emphasis id="82373CCE4EB2F13BAEE3CBF92F2BF881" bold="true" italics="true">Legs</emphasis>
(Fig.
<figureCitation id="1A7EF5F58AD9E44BE8D65B0E91A6019C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of Trichopeltis jiyue sp. nov., ♂ holotype A head, dorsal view B collum and the second segment, dorsal view C segments 69, dorsal view D cross-section of segment 5, caudal view E antenna disc coeloconic sensilla, plan view F telson and anal, ventral view G limbus of segment 5, subventral view H claw of leg, subventral view I paraterga of segment 6, ventral view. Scale bars: 1 mm (A D); 50 μm (E, H); 20 μm (G); 500 μm (F, I)." figureDoi="10.3897/zookeys.1216.128080.figure2" httpUri="https://binary.pensoft.net/fig/1156163">2 D, H</figureCitation>
) long and slender, without modifications, longer than paraterga when stretched straight, about 1.2 times as long as the width of paraterga. In length, femur ≈ tarsus &gt;&gt; prefemur&gt; coxa = tibia&gt; postfemur&gt; claw.
</paragraph>
<paragraph id="AAC7CCACF2DA00320F07C1DC148DBF97">
<emphasis id="EA32921B10FBDD0436161A44FD6D0418" bold="true" italics="true">Gonopods</emphasis>
complex (Figs
<figureCitation id="9CC786A3E5C728BBFB52F168008EB24B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gonopodal characters of Trichopeltis jiyue sp. nov., ♂ holotype, paratype. A, C right gonopod, sublateral and subfrontal views B left gonopod, subcaudal view D right gonopod, ventral view, coxite and telopodite divided by a notch E enlargement of box-E of A, prefemur sheet micro-setose on the surface F gonopod, ventral view from above G enlargement of box-G of A, corolliform solenomere H seminal groove I box-I of C, tripartite apical process. Abbreviations: p 1, p 2, p 3, p 4 = processes of telopodite; cp = corolliform pulvillus; cxp = coxa process; ms = microsetae; sg = seminal groove; tap = tripartite apical process; ca = cannula; cx = coxa; te = telopodite; n = notch. Scale bars: 500 μm (A D, F); 50 μm (E, G); 100 μm (H, I)." figureDoi="10.3897/zookeys.1216.128080.figure3" httpUri="https://binary.pensoft.net/fig/1156164">3</figureCitation>
,
<figureCitation id="8072CF7C0D8C721431316216BDD37E27" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trichopeltis jiyue sp. nov., ♂ holotype, paratype. A subfrontal view B subcaudal view C cadual view. Abbreviations: p 1, p 2, p 3, p 4 = processes of telopodite; cxp = coxa process; sg = seminal groove; ca = cannula. Scale bar: 200 μm." figureDoi="10.3897/zookeys.1216.128080.figure4" httpUri="https://binary.pensoft.net/fig/1156165">4</figureCitation>
). Coxae with relatively long stout setae; with two conspicuous wing-like processes (cxp). Telopodite complex, with four-branched process (
<emphasis id="DD3BAC28A8953ACE4A7E1239EA496188" bold="true">p</emphasis>
), clearly curved (Figs
<figureCitation id="D9E4A6C69E5863EE318ED0EADD6AAD82" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gonopodal characters of Trichopeltis jiyue sp. nov., ♂ holotype, paratype. A, C right gonopod, sublateral and subfrontal views B left gonopod, subcaudal view D right gonopod, ventral view, coxite and telopodite divided by a notch E enlargement of box-E of A, prefemur sheet micro-setose on the surface F gonopod, ventral view from above G enlargement of box-G of A, corolliform solenomere H seminal groove I box-I of C, tripartite apical process. Abbreviations: p 1, p 2, p 3, p 4 = processes of telopodite; cp = corolliform pulvillus; cxp = coxa process; ms = microsetae; sg = seminal groove; tap = tripartite apical process; ca = cannula; cx = coxa; te = telopodite; n = notch. Scale bars: 500 μm (A D, F); 50 μm (E, G); 100 μm (H, I)." figureDoi="10.3897/zookeys.1216.128080.figure3" httpUri="https://binary.pensoft.net/fig/1156164">3</figureCitation>
,
<figureCitation id="E8F9CC6A4753F52F9D7D8D951956205B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trichopeltis jiyue sp. nov., ♂ holotype, paratype. A subfrontal view B subcaudal view C cadual view. Abbreviations: p 1, p 2, p 3, p 4 = processes of telopodite; cxp = coxa process; sg = seminal groove; ca = cannula. Scale bar: 200 μm." figureDoi="10.3897/zookeys.1216.128080.figure4" httpUri="https://binary.pensoft.net/fig/1156165">4</figureCitation>
), approximately as long as coxa, divided by a notch (Fig.
<figureCitation id="0CA5B3DBFC539FD86965EC5A62DD03CD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gonopodal characters of Trichopeltis jiyue sp. nov., ♂ holotype, paratype. A, C right gonopod, sublateral and subfrontal views B left gonopod, subcaudal view D right gonopod, ventral view, coxite and telopodite divided by a notch E enlargement of box-E of A, prefemur sheet micro-setose on the surface F gonopod, ventral view from above G enlargement of box-G of A, corolliform solenomere H seminal groove I box-I of C, tripartite apical process. Abbreviations: p 1, p 2, p 3, p 4 = processes of telopodite; cp = corolliform pulvillus; cxp = coxa process; ms = microsetae; sg = seminal groove; tap = tripartite apical process; ca = cannula; cx = coxa; te = telopodite; n = notch. Scale bars: 500 μm (A D, F); 50 μm (E, G); 100 μm (H, I)." figureDoi="10.3897/zookeys.1216.128080.figure3" httpUri="https://binary.pensoft.net/fig/1156164">3 D</figureCitation>
); prefemur glabrous; femorite (
<emphasis id="3EE583FD09A93952E156CB3FACE1D993" bold="true">p 1</emphasis>
) one leaf-shaped lobe on the inner side mesally; branch
<emphasis id="E8AE3135706D206D5B7A411DAD9EA082" bold="true">p 2</emphasis>
leaf-shaped, three times as long as
<emphasis id="369438B367EE891C4302BD8FDE0E8CD5" bold="true">p 1</emphasis>
, rather thick, curved caudolaterally, with dense micro-setae on surface, distal margin with serrate process;
<emphasis id="E56366BF5F31D3B0DF70EC5109EA0AEF" bold="true">p 3</emphasis>
subconical, with three apical processes;
<emphasis id="EE61F4911A55E8D937F58F9D8841CBFA" bold="true">p 4</emphasis>
leaf-shaped, close to the
<emphasis id="53413C7FB363A8AE8AA0016DAD3B447B" bold="true">p 2</emphasis>
, the distal margin consists of numerous conical processes, forming a corolliform pulvillus; with no distinct solenomere.
</paragraph>
<caption id="A6A8081B26A8B4E09EB895F6E307D9B2" ID-DOI="10.3897/zookeys.1216.128080.figure3" ID-arpha="EB870C49-86B2-5BE8-A26E-00781117566C" httpUri="https://binary.pensoft.net/fig/1156164" startId="F3">
<paragraph id="15D046E9F6AECDBF908C4619F2405699">
<label id="15AA38E2AF4A547932ED02C3F5FCEC34">Figure 3.</label>
</paragraph>
<paragraph id="CD86B3A5463E5BF0CA95748212F3D064">
Gonopodal characters of
<taxonomicName id="299CE3E1FA9F9EAE5B74CC2C172362E0" authorityName="Wu &amp; Zhang &amp; Qin &amp; Cong" authorityYear="2024" class="Diplopoda" family="Cryptodesmidae" genus="Trichopeltis" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="jiyue" status="sp. nov.">
<emphasis id="1BFC9A904A54404F1056FA263B076116" italics="true">Trichopeltis jiyue</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2C9EA4DD5B736753A7DF150FDE98B81" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ holotype, paratype.
<emphasis id="D2C66343256B16FE88E0957E97B796F9" bold="true">A, C</emphasis>
right gonopod, sublateral and subfrontal views
<emphasis id="E5A4A087D7B5F6A1C5B31C57BA3F0BC4" bold="true">B</emphasis>
left gonopod, subcaudal view
<emphasis id="A384D5DF665D957470FC7A9FD931B3EB" bold="true">D</emphasis>
right gonopod, ventral view, coxite and telopodite divided by a notch
<emphasis id="B6DC7C00C172CCF74A241B4450BCA6B4" bold="true">E</emphasis>
enlargement of box-E of A, prefemur sheet micro-setose on the surface
<emphasis id="C96312946A5095ECC4EC26021B7F8206" bold="true">F</emphasis>
gonopod, ventral view from above
<emphasis id="82722A98A0E1FAE7FCCF67CB056F5D71" bold="true">G</emphasis>
enlargement of box-G of A, corolliform solenomere
<emphasis id="74AF7ACD125EF14CEFDE13F1007D8664" bold="true">H</emphasis>
seminal groove
<emphasis id="23F34861DA182BBE8B38DF918525AB80" bold="true">I</emphasis>
box-I of C, tripartite apical process. Abbreviations: p 1, p 2, p 3, p 4 = processes of telopodite; cp = corolliform pulvillus; cxp = coxa process; ms = microsetae; sg = seminal groove; tap = tripartite apical process; ca = cannula; cx = coxa; te = telopodite; n = notch. Scale bars: 500 μm (
<emphasis id="895A93E816936018EDFFFDC93D9D0C31" bold="true">A D, F</emphasis>
); 50 μm (
<emphasis id="5FA242AF199E672F50E81C09CF485BA7" bold="true">E, G</emphasis>
); 100 μm (
<emphasis id="8ABD8FB543DDE0ACF5222A41AD83B7ED" bold="true">H, I</emphasis>
).
</paragraph>
</caption>
<caption id="01F21A23D496F3D614F1AD209170492D" ID-DOI="10.3897/zookeys.1216.128080.figure4" ID-arpha="E16F36C1-5E62-5428-8AF0-DAAC28E6B4FD" httpUri="https://binary.pensoft.net/fig/1156165" startId="F4">
<paragraph id="9F8A6276F7C1C72DA11FC4743B83CF40">
<label id="FABEA40109DFAD15DA62E572CC9DD629">Figure 4.</label>
</paragraph>
<paragraph id="D4F1030F6530040619281B9B07C81D98">
<taxonomicName id="E843333196A6948C0A60F8A78DE9D18E" authorityName="Wu &amp; Zhang &amp; Qin &amp; Cong" authorityYear="2024" class="Diplopoda" family="Cryptodesmidae" genus="Trichopeltis" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="jiyue" status="sp. nov.">
<emphasis id="74E33A784CA4FDF17F8B4CD9E880DD4E" italics="true">Trichopeltis jiyue</emphasis>
</taxonomicName>
<taxonomicNameLabel id="162AC3714DEEA451C6927C24FA1425D7" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ holotype, paratype.
<emphasis id="FCFDA01253E1FAE2B07CC3B43DE53EB4" bold="true">A</emphasis>
subfrontal view
<emphasis id="3E6E5D2E131929C6AEBB272903A9CB34" bold="true">B</emphasis>
subcaudal view
<emphasis id="6C997D40D07B192311E5C123791FE00D" bold="true">C</emphasis>
cadual view. Abbreviations: p 1, p 2, p 3, p 4 = processes of telopodite; cxp = coxa process; sg = seminal groove; ca = cannula. Scale bar: 200 μm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EKCAE" type="remarks">
<paragraph id="5DEA946331A871B7C02A3E862290D30C">
<heading id="1B9DFEB439DDB96BBEBA51562446A331" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="6F8BC0EEB18EEB80F4888B9FB00FDF3A">The specimens were found on a stoney roadside, which some researchers usually walk around. As compared with virgin forests, the surroundings were relatively densely populated. However, the environment is undeveloped, and it the new species seemed abundant.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,876 @@
<document id="AA6EABE5EA134236BB56190BA4EA19A3" ID-DOI="10.3897/zookeys.1216.130017" ID-ZooBank="A6D95900-0A19-4778-9AA5-6D87622B419F" ID-publisher-id="130017" URI-arpha="F535348F-BBA1-5E6C-B72A-6E5D85153D3B" URI-zoobank="http://zoobank.org/A6D95900-0A19-4778-9AA5-6D87622B419F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1729263426519" checkinUser="pensoft" docAuthor="Lin, Shi-Shi, Chen, Hong-Hui, Li, Yuan-Hang, Peng, Zhao-Ning, Zeng, Zhao-Chi &amp; Wang, Jian" docDate="2024" docId="AF6296E2B1615E10819274EAB15B0D8F" docLanguage="en" docName="ZooKeys 1216: 1-15" docOrigin="ZooKeys 1216" docSource="https://zookeys.pensoft.net/article/130017/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Boulenophrys lichun Lin, Chen, Li, Peng, Zeng &amp; Wang, 2024, sp. nov." docType="treatment" docUuid="02303B1E-ED1A-482B-8BD3-B32BC0DB9D6B" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="15" masterDocId="F535348FBBA15E6CB72A6E5D85153D3B" masterDocTitle="A new Boulenophrys species (Anura, Megophryidae) from the coastal hills of eastern Fujian Province, China" masterLastPageNumber="15" masterPageNumber="1" pageNumber="1" updateTime="1729275296647" updateUser="admin">
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<mods:title id="1776DFFB4084B876CE038DD1B504B179">A new Boulenophrys species (Anura, Megophryidae) from the coastal hills of eastern Fujian Province, China</mods:title>
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<mods:namePart id="0D444D80D12A233C29DAE001742E121C">Lin, Shi-Shi</mods:namePart>
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<mods:namePart id="5C5CDEE41ED381E14DB9D58CA70250D4">Chen, Hong-Hui</mods:namePart>
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<mods:namePart id="453780905578CABCCEF23CB41034E590">Li, Yuan-Hang</mods:namePart>
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<mods:date id="CC2A3594DCD12D4B9C3FA82C3AB60B72">2024</mods:date>
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<subSubSection id="96814DDC7AA93331346FF9933D1F0E71" type="nomenclature">
<paragraph id="69D4D87F05E4895B0794FFD3E5A0E8C3">
<taxonomicName id="031EEE677BC30E14167330E78F7415C6" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="A63568DF62BF8FF9D553CE15D72CB620" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AB90265A369E1B92AA371A987A520840" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="07B553921D823682B689CBC32F7EB794" type="description">
<paragraph id="735D68D77958EDBEB4ACA38809D44937">
<figureCitation id="F599BCED89E4BE8DE6D3B6529700F5B4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The male holotype (GEP a 214, A C) and the female paratype (GEP a 215, D F) of Boulenophrys lichun sp. nov. in life." figureDoi="10.3897/zookeys.1216.130017.figure3" httpUri="https://binary.pensoft.net/fig/1156127">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQWAG" type="material">
<paragraph id="21D4438E06133DD48BA415328542F275">
<heading id="568EEE22023837BC329BC2931F8A4E62" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="10CE74763BDA39EE1E025021F1F955B8">
<materialsCitation id="B6FBF24E33008A8740FF24CF314E1F38" collectingDate="2024-02-04" collectionCode="GEP" collectorName="Jian Wang &amp; Zhao-Chi Zeng &amp; Shi-Shi Lin &amp; Yuan-Hang Li" country="China" county="Ningde City" elevation="230" latitude="26.645775" location="Mt Nanji" longLatPrecision="1" longitude="119.519936" municipality="Jiaocheng District" specimenCount="1" specimenCount-male="1" stateProvince="Fujian Province" typeStatus="holotype">
<emphasis id="E2FB06A0ADA9E9B26D38B43C2E5E50E3" bold="true">
<emphasis id="76BD50B47D05FC388B9F6345C69110B7" bold="true" italics="true">
<typeStatus id="3305849CE846A782158678C629740D00" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="866218AF02769AFBB9A9DACA21F62C16" name="China">China</collectingCountry>
<specimenCount id="1C4F50874D03D620299222F7E511D6C8" count="1" type="male"></specimenCount>
;
<collectingRegion id="57C18B1DC8A182645A89589508C30801" country="China" name="Fujian">Fujian Province</collectingRegion>
,
<collectingCounty id="18B5719013D71DBA244D1CD520C1ABD8">Ningde City</collectingCounty>
,
<collectingMunicipality id="03FBAAB78139765F50EA5CA35A047F86">Jiaocheng District</collectingMunicipality>
,
<location id="DE6207C5BB6B7362B58F832C12364DDE" LSID="urn:lsid:plazi:treatment:AF6296E2B1615E10819274EAB15B0D8F:DE6207C5BB6B7362B58F832C12364DDE" country="China" county="Ningde City" latitude="26.645775" longLatPrecision="1" longitude="119.519936" municipality="Jiaocheng District" name="Mt Nanji" stateProvince="Fujian Province">Mt Nanji</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5WAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[119.519939,26.645774]}">
<geoCoordinate id="967A3B1B95D18BED9E947E10D946CD2C" degrees="26.645774" direction="north" orientation="latitude" precision="1" value="26.645775">26.645774 ° N</geoCoordinate>
,
<geoCoordinate id="B5852F7616BD278F00E3BD11444801B1" degrees="119.519939" direction="east" orientation="longitude" precision="1" value="119.519936">119.519939 ° E</geoCoordinate>
</named-content>
, ca.
<quantity id="FC66C952E3B80EB39AA032196B9736F9" metricMagnitude="2" metricUnit="m" metricValue="2.3" unit="m" value="230.0">
<elevation id="7F73EED5053DBABA8D574A6918FE80CE" metricMagnitude="2" metricUnit="m" metricValue="2.3" unit="m" value="230.0">230 m</elevation>
</quantity>
elev.;
<date id="37DFA94075086614C564684DE0F96D3C" value="2024-02-04">
<collectingDate id="D418BFA3AADC142ED963ED7E5274AD0F" value="2024-02-04">4 Feb. 2024</collectingDate>
</date>
;
<collectorName id="6592F6C511E42837DC501ED1AB3D863B">Jian Wang</collectorName>
,
<collectorName id="CB056EF3341E2EC4DEA8B8DE93A8ED9B">Zhao-Chi Zeng</collectorName>
,
<collectorName id="7862A6FC298FB98D24E057FE0E01FE3D">Shi-Shi Lin</collectorName>
and
<collectorName id="BC4564DB01F86975C5B25DD869221513">Yuan-Hang Li</collectorName>
leg.;
<abbrev id="ABBRID0EDXAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 214.
</materialsCitation>
</paragraph>
<caption id="D53DEA791A1D5574A1C7A2BD10DBDCB4" ID-DOI="10.3897/zookeys.1216.130017.figure3" ID-arpha="E7F83773-006E-5EB9-B15F-D9DEA63D520C" httpUri="https://binary.pensoft.net/fig/1156127" startId="F3">
<paragraph id="E48D97F35FD56BD62620E1DBECCDD6F8">
<label id="3BC83A9A52342898546438B577318BBE">Figure 3.</label>
</paragraph>
<paragraph id="12BAE93FD80B0CC445431A6B78B14A4A">
The male holotype (
<abbrev id="ABBRID0EPXAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 214,
<emphasis id="94AA5D543ABF3AF99FF99077798264DB" bold="true">A C</emphasis>
) and the female paratype (
<abbrev id="ABBRID0EVXAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 215,
<emphasis id="5A5EC990F6C6CAD258E0ECAE10B84E9F" bold="true">D F</emphasis>
) of
<taxonomicName id="FC53F6FE3F61AD931B21F4D494900294" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="03B7DAB662232C3C2B124D596F711A39" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AA4A5091AD29037F53D5DD0709140BF8" rank="species">sp. nov.</taxonomicNameLabel>
in life.
</paragraph>
</caption>
<paragraph id="C01E78590826CF741F64B2928A73A06F">
<materialsCitation id="F4229613C249F24AD2BA4425C5521C01" collectingDate="2024-02-04" collectionCode="CIB" collectorName="Jian Wang &amp; Zhao-Chi Zeng &amp; Shi-Shi Lin &amp; Yuan-Hang Li" country="China" county="Ningde City" elevation="230" latitude="26.645775" location="China" longLatPrecision="1" longitude="119.519936" municipality="Jiaocheng District" specimenCode="CIB 121428" specimenCount="4" specimenCount-male="4" stateProvince="Fujian" typeStatus="paratype">
<emphasis id="C807152715572C9EF308E5C33CFCC1C6" bold="true">
<emphasis id="46674839CCE3335187CFBCD3D48A3DDD" bold="true" italics="true">
<typeStatus id="5F162DE408EEC8397BE136C6DEA2C344" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="ECF24295A223CCBDA791656BB3C028F4" name="China">China</collectingCountry>
<specimenCount id="6389891E63B50A3285CC3E37C7F0BED2" count="4" type="male">4 ♂♂</specimenCount>
; same data as for holotype;
<specimenCode id="00EB43A2FF2439BCB54C08326EF8D9DA" collectionCode="CIB" country="China" httpUri="http://grbio.org/cool/vxkr-andy" name="Chengdu Institute of Biology" type="">
<named-content id="06FBE4BF760F565539D1073BF7E837C2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chengdu-institute-biology" xlink_title="Herpetological Museum, Chengdu Institute of Biology, the Chinese Academy of Sciences">CIB</named-content>
121428
</specimenCode>
[field number
<abbrev id="ABBRID0EWYAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 210],
<abbrev id="ABBRID0E1YAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 211213
</materialsCitation>
<materialsCitation id="A93161AD17461AE4A2530F73F0EDBBC8" collectingDate="2024-02-04" collectionCode="GEP" collectorName="Jian Wang &amp; Zhao-Chi Zeng &amp; Shi-Shi Lin &amp; Yuan-Hang Li" country="China" county="Ningde City" elevation="230" latitude="26.645775" location="Mt Nanji" longLatPrecision="1" longitude="119.519936" municipality="Jiaocheng District" specimenCount="1" specimenCount-female="1" stateProvince="Fujian" typeStatus="paratype">
<specimenCount id="8BB68CCCB1C31E4F9C04B8DDC299ADBA" count="1" type="female">1 ♀</specimenCount>
; same data as for holotype;
<abbrev id="ABBRID0E5YAG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 215
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECZAG" type="etymology">
<paragraph id="92C180E593FEFBD7F072E258C6F2DB68">
<heading id="7ED79F399FBC8B64B52731ABE3D3991C" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="7CC4B3701B803939B310AD9505099C42">
The specific name
<taxonomicName id="03CB938BCBB375066139BB4B4CF96AF6" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun">
<emphasis id="0428019D5072279D5B7F1620B9528080" italics="true">lichun</emphasis>
</taxonomicName>
is derived from Chinese Pinyin Lì Chūn, i. e. 立春 in Chinese, which means the beginning of spring, the first of the 24 solar terms (24 节气) of
<collectingCountry id="2A711593687B42B6046BA10E20108634" name="China">China</collectingCountry>
. The specific name refers to the breeding season of the new species which begins around this period. The song of the new species heralds the spring of a year. The type specimens of the new species were also collected on “ Lichun ” of the Year 2024.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESZAG" type="diagnosis">
<paragraph id="C97CD7C6EB1230B85E3F8236CB3A244C">
<heading id="05A00138F2B06019A7FBCD3BC1E7A59B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D861EC12CD81CCA7509EAEC1C14206A1">
(1) small size (
<abbrev id="ABBRID0EYZAG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
33.537.0 mm in
<specimenCount id="B3FA198ABAE1F41F3359528EF1B1D817" count="5" type="adult">five adult</specimenCount>
males,
<abbrev id="ABBRID0E3ZAG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
<quantity id="8C2478430DFFD9D633B6AEFD8C0E8B2B" metricMagnitude="-2" metricUnit="m" metricValue="4.71" unit="mm" value="47.1">47.1 mm</quantity>
in a single adult female); (2) canthus rostralis well developed, tongue not notched posteriorly; (3) tympanum distinct; (4) vomerine ridges and vomerine teeth present; (5) dorsal skin rough and highly granular, discontinuous X-shaped ridge on center of dorsum, discontinuous dorsolateral ridges present, sparse large tubercles on flanks, dorsal limbs with discontinuous transverse ridges and tubercles, ventral skin with dense raised tubercles; (6) outer margin of upper eyelid with a small horn-like prominent tubercle, supratympanic fold distinct and narrow, curving posteroventrally to above arm; (7) two metacarpal tubercles distinct, inner one observably enlarged; relative finger lengths I &lt;
<collectionCode id="770B80BC54A6923ECE90FC29A8A6C3E3">II</collectionCode>
&lt;
<collectionCode id="472A53BC938581808472CAE4BAC9167C">IV</collectionCode>
&lt;
<collectionCode id="A0FFC6EAF11600DDC886766AE3037587">III</collectionCode>
; distinct subarticular tubercle at base of each finger; (8) heels not meeting when hindlimbs folded; tibio-tarsal articulation reaching shoulder to posterior corner of eye; (9) toes without webbing and lateral fringes, inner metatarsal tubercle long ovoid, outer one absent, relative toe length I &lt;
<collectionCode id="9DBF51B57765AFD268B53DD853F24268">II</collectionCode>
&lt;
<collectionCode id="48E44799D9EAF7E9336D7A9952500841" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
&lt;
<collectionCode id="289E46F80D7AC294F457DB3C85829EC5">III</collectionCode>
&lt;
<collectionCode id="7154D65FADFBB31C6602659F5226972E">IV</collectionCode>
; (10) dorsal surface yellowish-brown with irregular dark-brown patches, and dark-brown triangular marking between eyes, dorsal limbs and digits light brown with dark-brown transverse bands; and (11) dense nuptial spines on dorsal bases of fingers I and II in breeding adult males, subgular vocal sac present in males.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EA1AG" type="description">
<paragraph id="38647783CC98549DD21E6929F48569A4">
<heading id="7EE3B7352624C3E305EB223EB5A1AA44" reason="title">
Description of
<typeStatus id="68D46B223EBEDDD3688110C16F42D8F2" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="963E5247AF99523288BE7FD7EC43E54D">
Adult male. Body size small,
<abbrev id="ABBRID0EG1AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
37.0 mm. Head width larger than head length, HWD /
<abbrev id="ABBRID0EK1AG" xlink_title="head length, from tip of snout to the articulation of the jaw">HDL</abbrev>
1.04; snout rounded in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eyes moderate in size,
<abbrev id="ABBRID0EO1AG" xlink_title="eye diameter, from the anterior corner of the eye to posterior corner of the eye">ED</abbrev>
0.34 of
<abbrev id="ABBRID0ES1AG" xlink_title="head length, from tip of snout to the articulation of the jaw">HDL</abbrev>
, pupil vertical, near diamond-shaped; nostril obliquely ovoid; canthus rostralis well developed; loreal region slightly oblique; internasal distance slightly larger than interorbital distance; tympanic region oblique, tympanum distinct and visible in dorsal view; tympanum moderate in size, margin clear, upper margin in contact with supratympanic fold, lower margin in contact with upper lip,
<abbrev id="ABBRID0EW1AG" xlink_title="tympanum diameter, horizontal diameter of tympanum">TD</abbrev>
/
<abbrev id="ABBRID0E11AG" xlink_title="eye diameter, from the anterior corner of the eye to posterior corner of the eye">ED</abbrev>
0.55; large ovoid choanae at base of maxilla; vomerine ridge and vomerine teeth present, maxillary teeth present; margin of tongue rounded, not notched distally; presence of single subgular vocal sac.
</paragraph>
<paragraph id="73BBD0DAA8EECF113AA572B7D0BF29AA">
Forearm length 0.23 of
<abbrev id="ABBRID0EA2AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
, hand 0.24 of
<abbrev id="ABBRID0EE2AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
; webbing absent between fingers, lateral fringes absent, relative finger length I &lt;
<collectionCode id="B5A70161DD3E82EF72B3ADD5C48CF042">II</collectionCode>
&lt;
<collectionCode id="41ABB0E53CC8BCB5BECC4222F53F70A1">IV</collectionCode>
&lt;
<collectionCode id="7023D05580953F3F7FFC0C5B7DE5E5B2">III</collectionCode>
; tips of fingers slightly dilated, round; subarticular tubercles on base of fingers present, distinct; inner metacarpal tubercle observably enlarged, outer one slightly smaller; single nuptial pad bearing nuptial spines present on dorsal surface of first and second fingers, respectively. Hindlimbs short, tibio-tarsal articulation reaching forward to posterior conner of eye when hindlimb stretched along body; heels not meeting when flexed hindlimbs held at right angles to body axis; crus length 0.40 of
<abbrev id="ABBRID0EI2AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
and foot length 0.58 of
<abbrev id="ABBRID0EM2AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">SVL</abbrev>
; relative toe length I &lt;
<collectionCode id="7A7EE6D2EFBD63E7BD08A123412242C9">II</collectionCode>
&lt;
<collectionCode id="61DBC4FE9B23DBD317B27BBB5CDDDB57" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
&lt;
<collectionCode id="5DE27FC7CACE106DBCB5B0F21AF36BF8">III</collectionCode>
&lt;
<collectionCode id="901BCF7C4017A075DC4F1CA9E46A51AF">IV</collectionCode>
; tips of toes round and slightly dilated; toes without lateral fringes and webbing; subarticular tubercles on base of toes present and distinct; inner metatarsal tubercle long ovoid and lacking outer metatarsal tubercle.
</paragraph>
<paragraph id="4734D26911AA43CFC8B6127EC5A2F0AB">Dorsal skin rough and highly granular; dense large tubercles on flanks; single horn-like prominent tubercle on edge of upper eyelid; obvious supratympanic fold curving posteroventrally from posterior corner of eye to level above insertion of arm; upper lip, mandibular articulation, loreal, temporal region excluding tympanum, upper eyelid and surface around cloaca with conical tubercles; discontinuous X-shaped ridge on center of dorsum, discontinuous dorsolateral ridges present; ventral surface with dense raised tubercles; tubercles on ventral hindlimbs and around cloaca bearing tiny spines on their tips; small and distinct pectoral gland closer to axilla; single femoral gland positioned on posterior surface of thigh at midpoint between knee and cloaca.</paragraph>
</subSubSection>
<subSubSection id="SECID0ER2AG" type="Coloration of holotype">
<paragraph id="B6271C329E968C040FD7A209914A3CA2">
<heading id="D52D074E86447DB558C5C6BA45BD5C27" reason="title">
Coloration of
<typeStatus id="1471056AE28271617DBBAFED457CBB9A" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="801916717A651819C67249A8BD85D970">In life, dorsal surface of body yellowish-brown with irregular dark-brown patches, dark-brown X-shaped marking on center of dorsum, dark-brown triangular marking between eyes. A vertical dark-brown band present below eye. Dorsal surface of limbs with dark-brown transverse bands. Tubercles on edge of upper eyelids orange. Supratympanic fold light brown. Surface of throat and chest yellowish-brown with irregular dark brown and white patches and white and orange dots. Center of throat with black longitudinal band. Surface of abdomen white, mottled with orange dots and black patches. Surface of ventral limbs purple brown, with white mottling and dark-brown patches. Spines on tips of tubercles on ventral hindlimbs and area around cloaca black. Digits gray white; subarticular tubercles, inner and outer metacarpal tubercles and inner metatarsal tubercle grayish-brown. Pectoral glands and femoral glands white. Iris yellowish-brown with range mottling.</paragraph>
<paragraph id="F1239DB3829FDD7CE831755D55F2B2C9">In preservative, the dorsal surface of the body is dark brown, with markings and patches more distinct. Surface of chest, throat and limbs are dark brown, with dark-brown markings and patches more distinct, white patches and dots faded and orange dots absent. Color of pectoral glands and femoral glands faded.</paragraph>
</subSubSection>
<subSubSection id="SECID0EX2AG" type="variation">
<paragraph id="A05EE43468A52F5E89D8E035470F520A">
<heading id="24A90567E5E25DB12CD4257C7BBD9320" reason="title">Variation.</heading>
</paragraph>
<paragraph id="1CAB12BD573BE69772DAEAC1F0644AB8">
Morphometric variation is listed in Table
<tableCitation id="58EB5F81A86768F92996A51BE55598DE" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements (in mm) of voucher specimens of Boulenophrys lichun sp. nov.; * holotype." httpUri="http://table.plazi.org/id/D7A01796E20787A9657B8242C75A0570" tableUuid="D7A01796E20787A9657B8242C75A0570">2</tableCitation>
. Most of the
<typeStatus id="D6734D31C7CE3F4A0B4CD4DB892F682E" type="paratype">paratypes</typeStatus>
are similar to the
<typeStatus id="399D572516854E98B39EB3B11A099B0F" type="holotype">holotype</typeStatus>
in morphology and color pattern, except for the following: tibio-tarsal articulation reaching forward to posterior corner of eye when hindlimb stretched along body in the
<typeStatus id="94E92210034E424CD44299F6EDA9D9C7" type="holotype">holotype</typeStatus>
<abbrev id="ABBRID0EB3AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 214, while reaching to shoulder in female
<typeStatus id="79F10013B192518984DB5BD07B2D0039" type="paratype">paratype</typeStatus>
<abbrev id="ABBRID0EF3AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 215; absence of nuptial pads and spines in the female
<typeStatus id="F2E552D0D89127E1E74A1C3431F29F98" type="paratype">paratype</typeStatus>
; larger body size in the female
<typeStatus id="866F3D355B5033C4436C299E4493472A" type="paratype">paratype</typeStatus>
.
</paragraph>
<caption id="D7A01796E20787A9657B8242C75A0570" ID-Table-UUID="D7A01796E20787A9657B8242C75A0570" httpUri="http://table.plazi.org/id/D7A01796E20787A9657B8242C75A0570" startId="T2" targetIsTable="true">
<paragraph id="6CA7C23C57EC05ACCF370E0EFB2F8706">
<label id="67B894C971DF5AB609BA0172C86E490B">Table 2.</label>
</paragraph>
<paragraph id="805032C2F6223927388B992DB159BE75">
Measurements (in mm) of voucher specimens of
<taxonomicName id="E76D9F8CE35830C7FEE2E253D9182869" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="67AD94F17D3693F0D769D4E2FB0751CA" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="514146FED6A315B9242D5D6C95CFA703" rank="species">sp. nov.</taxonomicNameLabel>
; * holotype.
</paragraph>
</caption>
<paragraph id="A6C52B2D2377C1964F7CC05202432890">
<table id="TID0EMFBI" rules="all">
<tbody id="36E3B7183B13F1276238D087B8E649B8">
<tr id="F51F397C98409923553C8BA2D17A3C28">
<th id="049887F96C8FF5D6897CFA64A5C0B654" colspan="1" rowspan="1">Voucher</th>
<th id="DB8DC50175D957C19DCA3A7E692CC27F" colspan="1" rowspan="1">
<named-content id="8D527AA7A55CBB37457ADEE52051DCD2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chengdu-institute-biology" xlink_title="Herpetological Museum, Chengdu Institute of Biology, the Chinese Academy of Sciences">CIB</named-content>
121428
</th>
<th id="0CA4E6B5F1CF2906F196C049163B2365" colspan="1" rowspan="1">
<abbrev id="ABBRID0ER4AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 211
</th>
<th id="DDC60DC5F600A676CAF3F09830C5456D" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZ4AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 212
</th>
<th id="B44F3FD5CC718314C296736B2745EE22" colspan="1" rowspan="1">
<abbrev id="ABBRID0EB5AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 213
</th>
<th id="31B362FEEF17A1B10142106D4931C82D" colspan="1" rowspan="1">
<abbrev id="ABBRID0EJ5AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 214 *
</th>
<th id="5397D18DC1FAA4D6BDBB0CD662A57880" colspan="1" rowspan="1">
<abbrev id="ABBRID0ER5AG" xlink_title="Guangdong Polytechnic of Environmental Protection Engineering">GEP</abbrev>
a 215
</th>
</tr>
<tr id="C93855150C4F8713157657F22D11AED2">
<td id="E8EBC8994251B0808D08F12BAE518F40" colspan="1" rowspan="1">
<emphasis id="CE3EAEAFC816C38EC2841B687B901552" bold="true">Sex</emphasis>
</td>
<td id="72A185572C78C8A4DFA64AC659CD6D5F" colspan="1" rowspan="1">male</td>
<td id="26D17BEF9E42DB7C61A35A33203C12E0" colspan="1" rowspan="1">male</td>
<td id="9B5BC789F04E9028D8F638F1BF22B024" colspan="1" rowspan="1">male</td>
<td id="63F422351A8CB4296A15FC23F918F6CE" colspan="1" rowspan="1">male</td>
<td id="3D276470CE88E549A3BB40CA75EE3ADA" colspan="1" rowspan="1">male</td>
<td id="1FAD1EAFB39C9E20C34C70666F96724A" colspan="1" rowspan="1">female</td>
</tr>
<tr id="837206366F948802BB8652B46F2D8482">
<td id="0EA5A4DC01394BB6A47F650C3CABD6D5" colspan="1" rowspan="1">
<abbrev id="ABBRID0EU6AG" xlink_title="snoutvent length, from tip of snout to posterior margin of vent">
<emphasis id="836BC9227213F70705FCAB622914323A" bold="true">SVL</emphasis>
</abbrev>
</td>
<td id="E6E77903A0E468DDB8446C91C6A4EE46" colspan="1" rowspan="1">36.7</td>
<td id="1E988A9285124D1F36A35DC8643EF954" colspan="1" rowspan="1">33.5</td>
<td id="5BC78BF7ACAEDAF5BA8846C52782AE72" colspan="1" rowspan="1">35.7</td>
<td id="01EDEA2094248502067A4C76EBFB3835" colspan="1" rowspan="1">34.6</td>
<td id="2C528598EFBD687E8B18FB9E221585E4" colspan="1" rowspan="1">37.0</td>
<td id="6DF0424C0454D1221495A78F8DB7DF51" colspan="1" rowspan="1">47.1</td>
</tr>
<tr id="75040B7A321018737F7F57E04C8B8C2A">
<td id="FDA8154537EB3E9739900077B83A8B82" colspan="1" rowspan="1">
<abbrev id="ABBRID0EQABG" xlink_title="head length, from tip of snout to the articulation of the jaw">
<emphasis id="542561CB99529E1956A708C77F33FFA5" bold="true">HDL</emphasis>
</abbrev>
</td>
<td id="C17C2254E2DA8E44C75600E2DA1BF5EA" colspan="1" rowspan="1">13.1</td>
<td id="098CD03A141193C5CCE97F89D2A3C4DB" colspan="1" rowspan="1">12.7</td>
<td id="86858C9A684984C08F3326BADFF7CE9C" colspan="1" rowspan="1">13.5</td>
<td id="9E8746D54D5DD09C535E869DD94321E1" colspan="1" rowspan="1">12.9</td>
<td id="327043E8B1B9DB37DA08D9F43E9DABA1" colspan="1" rowspan="1">13.3</td>
<td id="D398C48E384A5BF7921DCCA6C65613AF" colspan="1" rowspan="1">15.3</td>
</tr>
<tr id="4881AD5ED959143B2F629D0AD811F818">
<td id="FDC4C9E3DC04CE8FD83F93A0D782540B" colspan="1" rowspan="1">
<abbrev id="ABBRID0EMBBG" xlink_title="head width, head width at the commissure of the jaws">
<emphasis id="991DADDEF5EDC9129CBD39E218919338" bold="true">HDW</emphasis>
</abbrev>
</td>
<td id="F26CE8FC60C4807AA6DF6CB6EBF06916" colspan="1" rowspan="1">13.6</td>
<td id="2114F11BCD04FDBDDEAA1FDE10193939" colspan="1" rowspan="1">13.3</td>
<td id="DF4D73B960699E6397BE850D025F67F2" colspan="1" rowspan="1">13.8</td>
<td id="9EF3D3E4DDD716E8B4B9F3EF6858B17F" colspan="1" rowspan="1">13.6</td>
<td id="B7ECD76DE5F61A9DE8CC37BCE97CE6A2" colspan="1" rowspan="1">13.9</td>
<td id="0AEEABB8B490A8E567008C95306969C1" colspan="1" rowspan="1">16.5</td>
</tr>
<tr id="7FCB56D4F0089B5E93DF821E7E411AE5">
<td id="A2F06853CEE82191B6AB68D670C2537E" colspan="1" rowspan="1">
<abbrev id="ABBRID0EICBG" xlink_title="eye diameter, from the anterior corner of the eye to posterior corner of the eye">
<emphasis id="4C293E554603FEC204996F0FE9C36402" bold="true">ED</emphasis>
</abbrev>
</td>
<td id="895F6A2509CBFDF8688C910B66C6F3C3" colspan="1" rowspan="1">4.5</td>
<td id="4A98977A01EA7BD8248402DDCF7241EF" colspan="1" rowspan="1">4.0</td>
<td id="9371D4F35B0DFDEC61FBF8055AB5CE66" colspan="1" rowspan="1">4.5</td>
<td id="F168E37842191EF6BF89EA4CEC5C56C3" colspan="1" rowspan="1">4.5</td>
<td id="E37F63BBB44165519C3F2C0F37C041A7" colspan="1" rowspan="1">4.8</td>
<td id="5E848F9895BBA3F944C663DB6BBD9F64" colspan="1" rowspan="1">5.5</td>
</tr>
<tr id="DFA07F122FB37ADD221C553E7F04BF7E">
<td id="36DCF704884B0F227EDECA6007B94EEE" colspan="1" rowspan="1">
<abbrev id="ABBRID0EEDBG" xlink_title="tympanum diameter, horizontal diameter of tympanum">
<emphasis id="B9EF7459A0B992C8D0FD4C6243110E56" bold="true">TD</emphasis>
</abbrev>
</td>
<td id="9AEE1C580EA8877D13768E47BEBCF7CB" colspan="1" rowspan="1">2.3</td>
<td id="8E075FA5B85383DA471D5E01D003A313" colspan="1" rowspan="1">2.6</td>
<td id="6F829CFE7AB9EF9C4679628E60F44AAA" colspan="1" rowspan="1">2.5</td>
<td id="AFEB03FE6E9FF4BB16314642D5A283B6" colspan="1" rowspan="1">2.5</td>
<td id="5BCF0497C339322372C754DD8AF2FD36" colspan="1" rowspan="1">2.6</td>
<td id="EA68076DDF2CCEAD933F4E5712B47FFD" colspan="1" rowspan="1">2.8</td>
</tr>
<tr id="62E1EA674F983185FEC2B930B47F61F7">
<td id="B7C1B46FCADA13D8143CC42749DBD8A5" colspan="1" rowspan="1">
<abbrev id="ABBRID0EAEBG" xlink_title="tympanumeye distance, from anterior edge of tympanum to posterior corner of the eye">
<emphasis id="8F2278A59BA90DBFD5053590EE507221" bold="true">TED</emphasis>
</abbrev>
</td>
<td id="DC83CA011294231E55E87FCCC843C09B" colspan="1" rowspan="1">1.7</td>
<td id="1FD511F64C860832241AEB92B6864294" colspan="1" rowspan="1">1.7</td>
<td id="3DF903E38F468B722CAFE760DAC50849" colspan="1" rowspan="1">1.8</td>
<td id="C4A34397B0D15C6E4CD3746C77C0993D" colspan="1" rowspan="1">1.8</td>
<td id="D646DCA0A2286CFF1BFBBC0180A5D1E2" colspan="1" rowspan="1">1.7</td>
<td id="FDDDBC69FE2035E818EF1D19364C3E61" colspan="1" rowspan="1">2.4</td>
</tr>
<tr id="CEE70C15D1F6F47109B336F9006817C6">
<td id="A4C790331A905078FAB53957F2B2976E" colspan="1" rowspan="1">
<abbrev id="ABBRID0E3EBG" xlink_title="snout length, from tip of snout to the anterior corner of the eye">
<emphasis id="2370E57CA98AE47F32FF329618C8CE67" bold="true">SNT</emphasis>
</abbrev>
</td>
<td id="6858B6CBE3641A1C40D27D8A4FFC9EAE" colspan="1" rowspan="1">4.5</td>
<td id="F4B1F48E4D4E44055223C24A0DDAEFDF" colspan="1" rowspan="1">4.4</td>
<td id="76B55FEF75ECA495FD47452A8C5907B8" colspan="1" rowspan="1">4.5</td>
<td id="3DDEC2E3474C1FAAA1EDB4465233672A" colspan="1" rowspan="1">4.5</td>
<td id="A8B3AFBA28CB97B436968FC7F376B40B" colspan="1" rowspan="1">4.6</td>
<td id="97BC150C73E30FDE53F64D7041E550B1" colspan="1" rowspan="1">4.9</td>
</tr>
<tr id="FC4C843E00E54AD1782541ECB85B8B01">
<td id="CDDE20A5786956A1E1F609CC71AED899" colspan="1" rowspan="1">
<abbrev id="ABBRID0EYFBG" xlink_title="internasal distance, distance between nares">
<emphasis id="9F94FDE8684335AEF85B76AB29E82BEB" bold="true">IND</emphasis>
</abbrev>
</td>
<td id="34330699A17B440F3D918DFE135FEB16" colspan="1" rowspan="1">3.6</td>
<td id="E3AEBF782E4EBB2B9BC589AAD9031228" colspan="1" rowspan="1">3.7</td>
<td id="CE04950AC81AF8B94A52A8E03F8A108B" colspan="1" rowspan="1">3.8</td>
<td id="D5FA6308A105D4DD78AC199AB8C7EC99" colspan="1" rowspan="1">3.7</td>
<td id="68F11F770A1E7DB9383D16DF5606B650" colspan="1" rowspan="1">3.9</td>
<td id="8CD0BCCCE0802570410CDE3715D32810" colspan="1" rowspan="1">4.3</td>
</tr>
<tr id="ABA95591697C706BDFA6221857661B1C">
<td id="099301FB9BA322A17148FB7C8ABC38AF" colspan="1" rowspan="1">
<abbrev id="ABBRID0EUGBG" xlink_title="interorbital distance, minimum distance between upper eyelids">
<emphasis id="30F493A5A76AE8B3E65F400774021CF4" bold="true">IOD</emphasis>
</abbrev>
</td>
<td id="4466D45C2232AACA74FF5148DFB31C3A" colspan="1" rowspan="1">3.5</td>
<td id="5D4FCCB81CE9AA4D23B14293CB045461" colspan="1" rowspan="1">3.3</td>
<td id="D124B416B9D9BD8569BD86795AD69A3E" colspan="1" rowspan="1">3.4</td>
<td id="8130E6AA8A6CC9E5A985D3407CA8878A" colspan="1" rowspan="1">3.5</td>
<td id="D41D4374C0DD317EFAAE2FB3136677DD" colspan="1" rowspan="1">3.7</td>
<td id="E71BE03E633733E9906F22BE6892AD3C" colspan="1" rowspan="1">4.3</td>
</tr>
<tr id="336B68D02418E26419DE10C8CFD1483B">
<td id="E775385E74FD8C793800BDB0BE18E3D3" colspan="1" rowspan="1">
<emphasis id="12301992514B37BCD0C86BB595C2058E" bold="true">HDN</emphasis>
</td>
<td id="DEDDDE40B833D9603A3B3AB644E605AF" colspan="1" rowspan="1">8.7</td>
<td id="21F0A68966074FF0FEF070ECD68FB82C" colspan="1" rowspan="1">7.8</td>
<td id="5DA0C61BEBDB75FE9D28DB7456EF344B" colspan="1" rowspan="1">8.7</td>
<td id="7559EA8BA19814B5DC2E45390BBF28A9" colspan="1" rowspan="1">9.1</td>
<td id="4C1274E021587BA9D5D77CCAADFF9C8D" colspan="1" rowspan="1">9.2</td>
<td id="57EC47621EE5734CDBEFD8F6A8DA878E" colspan="1" rowspan="1">10.2</td>
</tr>
<tr id="918FFCA1FE96E3C58DEE294E981B81D1">
<td id="5EC2BC172CAFBD35CA53229B3983502E" colspan="1" rowspan="1">
<abbrev id="ABBRID0EJIBG" xlink_title="forearm or radiulna length, from the flexed elbow to the proximal border of the outer palmar tubercle">
<emphasis id="6066D2CAB027DA10A07B99B25BB43209" bold="true">RAD</emphasis>
</abbrev>
</td>
<td id="7AC0B388EC756766A2D6795467B1C77F" colspan="1" rowspan="1">8.6</td>
<td id="FA7C181065AD18C3A9613A34BE4A02CA" colspan="1" rowspan="1">7.0</td>
<td id="461EBCD62B7E08C4DA78827B2FBCCC7E" colspan="1" rowspan="1">8.5</td>
<td id="A4645527919747E813A8AD132A8A9A3A" colspan="1" rowspan="1">8.1</td>
<td id="40433DB7EF899C0A6764B9CE5FC0FA5D" colspan="1" rowspan="1">8.7</td>
<td id="D09CB509751A7E7713DBB2B7EDA3A337" colspan="1" rowspan="1">9.0</td>
</tr>
<tr id="12F9B569A327495B6762F32E1B89C8CA">
<td id="74A1A4509C09C0A0B5C6B1D6F191CB01" colspan="1" rowspan="1">
<abbrev id="ABBRID0EFJBG" xlink_title="foot length, from distal end of shank to the tip of digit IV">
<emphasis id="D1105FED716839ED0C96F79AC1DA206D" bold="true">FTL</emphasis>
</abbrev>
</td>
<td id="D90F4074B2AC6D12C3738D556D3A6749" colspan="1" rowspan="1">20.7</td>
<td id="31EFE29ADDA1FCDA5BDB07FE7DF37FCF" colspan="1" rowspan="1">18.9</td>
<td id="D91089EA322F744A56C24B1677DDD767" colspan="1" rowspan="1">20.7</td>
<td id="99ADC8395DAD6E4920B04E48E264C99E" colspan="1" rowspan="1">20.9</td>
<td id="3F65D824066F1FDE0D8DE065B40C984C" colspan="1" rowspan="1">21.4</td>
<td id="8EA6C31A474B8CE73B728BB77BFE6D8F" colspan="1" rowspan="1">24.1</td>
</tr>
<tr id="A674B4931E9612DB95BCA6A42EAC6810">
<td id="7F106DAA7E452DAC075FC199228C1D51" colspan="1" rowspan="1">
<abbrev id="ABBRID0EBKBG" xlink_title="crus or tibiofibula length, from the outer surface of the flexed knee to the heel">
<emphasis id="06732C834DDD5C076E6C87ED3E806710" bold="true">TIB</emphasis>
</abbrev>
</td>
<td id="0944694D1DD0B27046D3855EB6B05E84" colspan="1" rowspan="1">14.3</td>
<td id="3A71C93BE3D94444AA564639AE9B0D9F" colspan="1" rowspan="1">12.5</td>
<td id="B350CEE68CB969FEF88CC53FBFD58CDA" colspan="1" rowspan="1">14.3</td>
<td id="B4E17C505916B6D6944C5FD0B7C275F9" colspan="1" rowspan="1">14.5</td>
<td id="D449D805881DAB681CEBC614426DD1AA" colspan="1" rowspan="1">14.7</td>
<td id="A6FFF6F5BADB81EDD0BE2D14EEFF1B8D" colspan="1" rowspan="1">15.7</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXKBG" type="comparisons">
<paragraph id="E2D10ACCFAE6CB2A9EFF30D146914556">
<heading id="77B7C96CB03F25E56DA833B49ABC1E7F" reason="title">Comparisons.</heading>
</paragraph>
<paragraph id="9C7199A408517EE176B9C32B9FC26E86">
<taxonomicName id="2F96F083B9A1CA835D02B2C357A0E14D" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="D3CF4FF24269E5F976E881F7106F1499" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F21AFA049080D8E24DC92668B5C4ED35" rank="species">sp. nov.</taxonomicNameLabel>
can easily be distinguished from the following congeners by its heels not meeting when flexed hindlimbs held at right angles to body axis:
<taxonomicName id="24155DAE73375F7DB73E283FA5704FFD" baseAuthorityName="Li, Lu, Liu &amp; Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="anlongensis">
<emphasis id="D791F609DA31DB398A4DC915611DCD7E" italics="true">B. anlongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="A260512E2FCD102E46469FED8840F9A8" baseAuthorityName="Wu, Li, Liu, Wang &amp; Wu" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="baishanzuensis">
<emphasis id="F375BD0535FC10B8DD2D01B9A4237F9B" italics="true">B. baishanzuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="F90362AB635CA49ED4CD10A8921233C8" baseAuthorityName="Jiang, Fei &amp; Ye" baseAuthorityYear="2009" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="binlingensis">
<emphasis id="74F5F6E771ACAC72226A15F8298C857F" italics="true">B. binlingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6688A3C8E36AD7A6616BD9258705A2C0" baseAuthorityName="Shen" baseAuthorityYear="1994" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="caudoprocta">
<emphasis id="A00FA677354BBF047351254AE2341935" italics="true">B. caudoprocta</emphasis>
</taxonomicName>
,
<taxonomicName id="C37E60217285E6539A484F8A88382D96" baseAuthorityName="Wang &amp; Liu" baseAuthorityYear="2014" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cheni">
<emphasis id="24357EE2235C4A3229CB2A9F6211B644" italics="true">B. cheni</emphasis>
</taxonomicName>
,
<taxonomicName id="441143FBD9992F34D0AA3C7814CE9D8A" baseAuthorityName="Xu, Li, Liu, Wei &amp; Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="chishuiensis">
<emphasis id="69A7D4CA45A16C53783631761C8FB4E3" italics="true">B. chishuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="3C371B7DAAC3CAE8425AE343103CCBDB" baseAuthorityName="Luo, Wang, Wang, Lu, Wang, Deng &amp; Zhou" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="congjiangensis">
<emphasis id="932BF2820894C46DEC1C7D9D8B53FEE9" italics="true">B. congjiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="C2CDF35B4609D28429D1ED44775F7B7D" baseAuthorityName="Rao &amp; Yang" baseAuthorityYear="1997" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="daweimontis">
<emphasis id="7E7A9AD3D8287937CC06E1DA9C489F1A" italics="true">B. daweimontis</emphasis>
</taxonomicName>
,
<taxonomicName id="2CCD4CBBC50AB9405922F7C23F0D2109" baseAuthorityName="Zhang, Liang, Ran &amp; Shen" baseAuthorityYear="2012" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="fanjingmontis">
<emphasis id="BD44715D31CA96880B5D3B9019531AFB" italics="true">B. fanjingmontis</emphasis>
</taxonomicName>
,
<taxonomicName id="9DE4F65C33248229AF37875D9045EDD2" baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong &amp; Rowley" baseAuthorityYear="2018" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="fansipanensis">
<emphasis id="8E0C4281EEA4317734FA7CAE7CCE9025" italics="true">B. fansipanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="43137B08DB5B0C605B6216694A3C34E4" baseAuthorityName="Tapley, Cutaja, Nguyen, Portway, Mahony, Nguyen, Harding, Luong &amp; Rowley" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="frigida">
<emphasis id="8E02E1D6021ABE0C352BC42F0DDD0C66" italics="true">B. frigida</emphasis>
</taxonomicName>
,
<taxonomicName id="5C42BB3EF910F8EC74EC9249E0A19694" baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong &amp; Rowley" baseAuthorityYear="2018" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hoanglienensis">
<emphasis id="7B4A28A8C83CF3BAD350DD817F44002F" italics="true">B. hoanglienensis</emphasis>
</taxonomicName>
,
<taxonomicName id="A8F881E9AD2D7FD4FFD81E1654BE795C" baseAuthorityName="Liu, Li, Wei, Xu, Cheng, Wang &amp; Wu" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jiangi">
<emphasis id="A848CFD5C6A4E583DF722F5A0A1368B9" italics="true">B. jiangi</emphasis>
</taxonomicName>
,
<taxonomicName id="960CBF0D10D4E83F56F0D36D7CE07F1E" baseAuthorityName="Fei &amp; Ye" baseAuthorityYear="1983" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="0D227A7B98D1E3599A848B2CA95FCF6D" italics="true">B. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="3FB32EFE95D6B448943441155F021B1E" baseAuthorityName="Wang" baseAuthorityYear="2012" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jinggangensis">
<emphasis id="B0261B66457C902DE73CBFAFA820D8E0" italics="true">B. jinggangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="371645458C4617C6A9CFF29CC384607A" baseAuthorityName="Wang, Zeng, Lyu &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jiulianensis">
<emphasis id="527BE408C17010EDFAF105F230DBF444" italics="true">B. jiulianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="829653CC5551636E754EAAF9DA1EAFE7" baseAuthorityName="Li, Xu, Liu, Jiang, Wei &amp; Wang" baseAuthorityYear="2018" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="leishanensis">
<emphasis id="EC7666E852A285FFE1E7D6E35EBF502D" italics="true">B. leishanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="C78930BE650F7D6984BB566ABC92639F" baseAuthorityName="Zhang, Li, Xiao, Li, Pan, Wang, Zhang &amp; Zhou" baseAuthorityYear="2017" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="liboensis">
<emphasis id="09CD2B5EAC669EBF695605C4B72973A5" italics="true">B. liboensis</emphasis>
</taxonomicName>
,
<taxonomicName id="AF8D7268FBD6FA1C20392EA5D5FFE6E6" baseAuthorityName="Wang &amp; Yang" baseAuthorityYear="2014" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lini">
<emphasis id="D157C214704ADC6C87CD0A84F4C5251F" italics="true">B. lini</emphasis>
</taxonomicName>
,
<taxonomicName id="8733BC4ADDF7479D10DBEDC3D41556D0" baseAuthorityName="Shi, Li, Zhu, Jiang, Jiang &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lushuiensis">
<emphasis id="63960180AB95EC85A77FF4D8599D7258" italics="true">B. lushuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="15AA48C10E28FE7082945706B012B8D9" baseAuthorityName="Lyu, Wang &amp; Zhao" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="mirabilis">
<emphasis id="1DE5911DDDECC7AA2729A8B787661462" italics="true">B mirabilis</emphasis>
</taxonomicName>
,
<taxonomicName id="3455D116D283C8FBB700DD2D2C0C3F2F" baseAuthorityName="Wang, Lyu &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="mufumontana">
<emphasis id="AA613C0EA26D76017D11BFDF2E51C4B2" italics="true">B. mufumontana</emphasis>
</taxonomicName>
,
<taxonomicName id="1A7C94B076A33991E64AA5712EEC8CF9" baseAuthorityName="Lyu, Wang, Liu &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nanlingensis">
<emphasis id="5442AC70817C956A99B89115187AAFE3" italics="true">B. nanlingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="EB6306A121B5024F8B3E003FA4958297" baseAuthorityName="Liu" baseAuthorityYear="1950" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="omeimontis">
<emphasis id="C712C0BBF16E620A3C4258352D0C7873" italics="true">B. omeimontis</emphasis>
</taxonomicName>
,
<taxonomicName id="7D3C3D91A976E6DC6A48F55874BB1AAD" baseAuthorityName="Bourret" baseAuthorityYear="1937" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="palpebralespinosa">
<emphasis id="C724760791D862FD951DA64439F110F7" italics="true">B. palpebralespinosa</emphasis>
</taxonomicName>
,
<taxonomicName id="CD6087187B76C70D0B9F50D1A192A4DB" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="qianbeiensis">
<emphasis id="1E7112EE361D0AD2E79AE7D5D4254145" italics="true">B. qianbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="1DFBBC6AECF3801047A98AD60D4954B0" baseAuthorityName="Jiang, Ye &amp; Fei" baseAuthorityYear="2008" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sangzhiensis">
<emphasis id="B71B97889110930F8F48C390F928DB7D" italics="true">B. sangzhiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="84E22F7AA6EDD8E1DF138E8737393DAD" baseAuthorityName="Lyu &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sanmingensis">
<emphasis id="1D047C74F368AC7C14CB9CDD799FA181" italics="true">B. sanmingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="73F788D240F316F547CEE46A9644E6B9" baseAuthorityName="Lyu, Liu &amp; Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shimentaina">
<emphasis id="F80FFF8C5774C4550570B17967A347A8" italics="true">B. shimentaina</emphasis>
</taxonomicName>
,
<taxonomicName id="14DF84D175CD51B4F327F0E666A8769E" baseAuthorityName="Wang, Deng, Liu, Wu &amp; Liu" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shunhuangensis">
<emphasis id="5EB869893A0FB3E686F944DAB437E128" italics="true">B. shunhuangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="206F6D74F5852422419E9E3A0719C489" baseAuthorityName="Liu &amp; Hu" baseAuthorityYear="1973" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="spinata">
<emphasis id="93EDA55C909B72C3D76A6D76C3CA404C" italics="true">B. spinata</emphasis>
</taxonomicName>
,
<taxonomicName id="E34B86F7307E94E91B335B1BCA5FEC58" baseAuthorityName="Lyu &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sanmingensis">
<emphasis id="1F8BC3B6EFD840112F4FFB3B78740CF9" italics="true">B. sanmingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="B652BE8388D418787066CCEE74616804" baseAuthorityName="Wang &amp; Lyu" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="tongboensis">
<emphasis id="397FA37E6D0657AD58FA56F8C458451E" italics="true">B. tongboensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6866E07DF5AC6CD45574DDEB3124933D" baseAuthorityName="Shen, Mo &amp; Li" baseAuthorityYear="2010" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="tuberogranulatus">
<emphasis id="B9A439E1FD4881EC73FF546D404F6A2E" italics="true">B. tuberogranulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="FE2C4F6784481548AA8FB25A1266D3D8" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="wuliangshanensis">
<emphasis id="F68B21C24DCBBF69EC2E223526EBD196" italics="true">B. wuliangshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="B1AB0DFD5D5152EF03B924EA6A634E57" baseAuthorityName="Wang, Wu, Peng, Shi, Lu &amp; Wu" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xianjuensis">
<emphasis id="A45C2780216F9084E90B90B553FBE4E4" italics="true">B. xianjuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="76975B23A432CDF7D6078C3DFF161D3D" baseAuthorityName="Lyu, Zeng &amp; Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yangmingensis">
<emphasis id="B1F3D471C98B10AA6C5FF02458360088" italics="true">B. yangmingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="1C0A3A12E7159B553319F3106B7FD6A2" authorityName="Qi, Mo, Lyu, Wang &amp; Wang" authorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yaoshanensis">
<emphasis id="AC91795EEC4F34C0DC4A42F572D49C93" italics="true">B. yaoshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="DE35F143E855FD23C572153EF34AB52F" authorityName="Qi, Lyu, Wang &amp; Wang" authorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yingdeensis">
<emphasis id="807D84A015638AC26577C6AD5D2100E3" italics="true">B. yingdeensis</emphasis>
</taxonomicName>
(vs. heels overlapping), from
<taxonomicName id="9202F481B0A9F28C289193194958911D" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="binchuanensis">
<emphasis id="5E4E2113D1C84A18A1BBFCEABE643B11" italics="true">B. binchuanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="07128C59E7681CDC8A79C4CEE8135883" authorityName="Zeng, Wang, Chen, Xiao, Zhan, Li &amp; Lin" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="elongata">
<emphasis id="1EB938B26E504534504F523C6A943017" italics="true">B. elongata</emphasis>
</taxonomicName>
,
<taxonomicName id="71FDC347E7C15092AF1AED184860896E" baseAuthorityName="Wang, Liu &amp; Jiang" baseAuthorityYear="2017" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lishuiensis">
<emphasis id="63DF000FEC90555ED20882F0ABFC14BF" italics="true">B. lishuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="1D0E02B7F84875E9F78B0A3A64DBD771" baseAuthorityName="Stejneger" baseAuthorityYear="1926" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="minor">
<emphasis id="4DE23E4F826F44067DAB9FC30A569676" italics="true">B. minor</emphasis>
</taxonomicName>
,
<taxonomicName id="342EA315ECF2CBBD86CEF336E997E616" baseAuthorityName="Lyu, Zeng &amp; Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xiangnanensis">
<emphasis id="2CF27856E9B4A7AB55457081B5EF7989" italics="true">B. xiangnanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="2001AC8866CF1E5B3D4D094F1C6F0297" authorityName="Lyu &amp; Wang" authorityYear="2023" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xuefengmontis">
<emphasis id="B4FC2119858431BB13BCE9EA172B3A10" italics="true">B. xuefengmontis</emphasis>
</taxonomicName>
(vs. heels just meeting), and from
<taxonomicName id="E3E5776EA98348CFCE78BAD8B4975415" baseAuthorityName="Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy &amp; Che" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="angka">
<emphasis id="B8704DE6695727D995064393CAD3E4FA" italics="true">B. angka</emphasis>
</taxonomicName>
,
<taxonomicName id="111AAA49A0B2075171561A6C224EE91C" baseAuthorityName="Lyu, Wang &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="daiyunensis">
<emphasis id="F474135F640EA4BA5E54DBC8EEF22E26" italics="true">B. daiyunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="01171BBFB09401CFB1B25551A400ECEE" baseAuthorityName="Ye, Fei &amp; Xie" baseAuthorityYear="2007" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="baolongensis">
<emphasis id="10EA953861E9084143EB6EB9A346EA2E" italics="true">B. baolongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="A9B80A2497A0FDD0F455B2775DB84B2E" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="wushanensis">
<emphasis id="331E16E99C71CA01F6BCA9ED8D02DF44" italics="true">B. wushanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="72E21FDD0071D72C987D83A144601A5C" authorityName="Qi, Wang, Lyu &amp; Wang" authorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yunkaiensis">
<emphasis id="E760581733EBCE98A223E5FC2259AFB7" italics="true">B. yunkaiensis</emphasis>
</taxonomicName>
(vs. heels just meeting or slightly overlapping).
</paragraph>
<paragraph id="17C657E13DDC77522A03382FC364D2AA">
<taxonomicName id="E30C43F588CFDB4414B418257EA4BF73" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="69CAA55C486EAC2A901EE0962BE559DE" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="785D093D2A8FC3147026BA22E1DBA76E" rank="species">sp. nov.</taxonomicNameLabel>
can easily be distinguished from the following congeners by its tongue not notched distally:
<taxonomicName id="101CFBEE877C4C94A3C8C60976154AAB" baseAuthorityName="Inger &amp; Romer" baseAuthorityYear="1961" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="brachykolos">
<emphasis id="31EC3ABCCB69C0954CD396357BC0378D" italics="true">B. brachykolos</emphasis>
</taxonomicName>
,
<taxonomicName id="B5250C18C47EEF3E3F8C0598BAF2DD7F" baseAuthorityName="Wang, Liu, Lyu, Zeng &amp; Wang" baseAuthorityYear="2017" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="insularis">
<emphasis id="E223C53E64471E8BF4CC42F2809A2163" italics="true">B. insularis</emphasis>
</taxonomicName>
,
<taxonomicName id="F0D64AE1A50E2EAA85501BD092A207A4" baseAuthorityName="Wang &amp; Zeng" baseAuthorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="pepe">
<emphasis id="6E26BA6DA4AD4B7860C39EC35D39C7B4" italics="true">B. pepe</emphasis>
</taxonomicName>
(vs. tongue notched distally).
<taxonomicName id="BF37737B0F952B1A61709DCF224D79F0" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="EFF88ADDCEB335B28D8F35E34F207BF7" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="552D32E5750602D119EE88F9FBAD11E6" rank="species">sp. nov.</taxonomicNameLabel>
can easily be distinguished from the following congeners by its presence of vomerine teeth:
<taxonomicName id="142BBB7FCEEDB8B5C5954DA28A917DB5" baseAuthorityName="Wang, Li &amp; Jin" baseAuthorityYear="2014" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="acuta">
<emphasis id="0C74A7D9E9B65D6C4ED32A39FAD81BF2" italics="true">B. acuta</emphasis>
</taxonomicName>
,
<taxonomicName id="FCD16EE464725D139455A8B8158425E4" baseAuthorityName="Boulenger" baseAuthorityYear="1899" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="0E1EFCF3CCD07949498A525A3830A1AF" italics="true">B. boettgeri</emphasis>
</taxonomicName>
,
<taxonomicName id="65DE6A3EB264C178624A1138B85563E6" baseAuthorityName="Nguyen, Pham, Nguyen, Luong &amp; Ziegler" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="caobangensis">
<emphasis id="053B8F644D573530D913532D1CC3219D" italics="true">B. caobangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="EC9FFF6A6B66F9510DBB0E53B0A8D93D" baseAuthorityName="Lyu, Zeng, Wang &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="daoji">
<emphasis id="677AE47755B8DED4A0BBD7B4CB6FC56A" italics="true">B. daoji</emphasis>
</taxonomicName>
,
<taxonomicName id="28C0A60B961EF28738E3370E097B7298" authorityName="Wang, Zeng, Lyu, Xiao &amp; Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hungtai">
<emphasis id="254C7680A45062108AE0735DB344D760" italics="true">B. hungtai</emphasis>
</taxonomicName>
,
<taxonomicName id="7124B97207AF3BAB4BF2107678259160" authorityName="Qian, Hu, Mo, Gao, Zhang &amp; Yang" authorityYear="2023" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hengshanensis">
<emphasis id="A60DE18E62ECCB925CF6A304404E1BBE" italics="true">B. hengshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="2862DA29E43734D9EA8C5E1E94612842" baseAuthorityName="Pope" baseAuthorityYear="1929" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="CED8C3860F4DBFDB01142EF323383E79" italics="true">B. kuatunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="5E56EA7928F5BDE7A6FFFB41812D2687" baseAuthorityName="Messenger &amp; Dahn" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ombrophila">
<emphasis id="64E79D0D5F12892B7973357F1A5EEE90" italics="true">B. ombrophila</emphasis>
</taxonomicName>
,
<taxonomicName id="09201E7967E82E6588F5C64420225A4D" baseAuthorityName="Wang, Li &amp; Zhao" baseAuthorityYear="2014" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="obesa">
<emphasis id="2711C568D0A531BDCBE79C0CD832CAD0" italics="true">B. obesa</emphasis>
</taxonomicName>
,
<taxonomicName id="55F521B8AE119FDDBDBB57CFFFD24F1A" baseAuthorityName="Tian &amp; Sun" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shuichengensis">
<emphasis id="BC0A5DA1650A29BBB73594727D224591" italics="true">B. shuichengensis</emphasis>
</taxonomicName>
,
<taxonomicName id="1FBF22B5F89B1256CE58F93B85E8ED0E" baseAuthorityName="Wang, Lyu &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="wugongensis">
<emphasis id="47A79CFE7209ACD75BAE9CFF62CD3713" italics="true">B. wugongensis</emphasis>
</taxonomicName>
(vs. vomerine teeth absent).
</paragraph>
<paragraph id="C2448FFC024885178C202432797A89B7">
<taxonomicName id="FFAE7CEEA6F21DCC32066BEF3B8803CD" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="235B2E6B3D6AB00FCBAA9A046D740D5B" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9E751C4FDE68D5F12E17D38C9FFEFEB6" rank="species">sp. nov.</taxonomicNameLabel>
can easily be distinguished from the following congeners by its absence of lateral fringes on webbing on toes:
<taxonomicName id="03AA7123829FA53E468F85151C124348" baseAuthorityName="Wang &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="dongguanensis">
<emphasis id="F58085F19BCC1CB8189E8F866F74BEEB" italics="true">B. dongguanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="79E874B00651E705F3ADC47C0BEC3040" authorityName="Wang, Zeng, Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="fengshunensis">
<emphasis id="763A7A58D40355ED36F5B0AE1043EC95" italics="true">B. fengshunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="09802D9FCF59BDFEA3084CFD61BFD506" baseAuthorityName="Wang, Zeng &amp; Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nankunensis">
<emphasis id="6D94D9F8B13BB107A7605B198F65FD9F" italics="true">B. nankunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="0D11DA042BA41DECFB8AC102ADA9D9DC" authorityName="Wang, Zeng, Lyu, Xiao &amp; Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="puningensis">
<emphasis id="E27EA79349C947CDA2CDD1F730181563" italics="true">B. puningensis</emphasis>
</taxonomicName>
(vs. toes with rudimentary webbing), and from
<taxonomicName id="92D463A73D2E38606C912C4B2B3CE143" baseAuthorityName="Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong &amp; Rowley" baseAuthorityYear="2017" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rubrimera">
<emphasis id="2A4A7530615DEBD25BC51B9518E4CB13" italics="true">B. rubrimera</emphasis>
</taxonomicName>
(vs. toes with narrow lateral fringes).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFEAI" type="distribution">
<paragraph id="DBE0BDFE48A1A74ACC7DFF4FDF624C66">
<heading id="893C6222C352E659C09A3073EAEE9676" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="CBE97A313C6BB2BA511D81F4B8BA1FA9">
Currently,
<taxonomicName id="9AB26459ABE07966F7D4B14C24B9B491" authorityName="Lin &amp; Chen &amp; Li &amp; Peng &amp; Zeng &amp; Wang" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichun" status="sp. nov.">
<emphasis id="B005AAA343905782967722A00A7B41DE" italics="true">Boulenophrys lichun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44D9B16CC3637B160882B5D1EA738336" rank="species">sp. nov.</taxonomicNameLabel>
is only known from the coastal hills of Ningde City, eastern
<collectingRegion id="EA3E3E471C223FB70450FD45BB169646" country="China" name="Fujian">Fujian Province</collectingRegion>
,
<collectingCountry id="B7F545A70E114ABDFCF3A7E6AC0EF83C" name="China">China</collectingCountry>
. It inhabits flowing montane seeps and the nearby forest floor and leaf litter. The habitat is surrounded by secondary forest mixed with bamboo groves at elevations between
<quantity id="C3A6A9743B8888C3012887403DAAB7E4" metricMagnitude="2" metricUnit="m" metricValue="3.3" metricValueMax="5.1" metricValueMin="1.5" unit="m" value="330.0" valueMax="510.0" valueMin="150.0">
150
<specimenCount id="7F72168ACBCF1329AC55E71405C70DA5" count="510" type="generic">510 m.</specimenCount>
</quantity>
Advertisement calls of males were heard from February to May. Males were found calling in rock crevices.
</paragraph>
</subSubSection>
</treatment>
</document>