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<document id="B59ED2627B52E1DB2637B7C38A35CAF3" ID-DOI="10.11646/phytotaxa.500.1.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="5424389" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630391626960" checkinUser="marcus" docAuthor="Ralaiveloarisoa, Anna Berthe, Liimatainen, Kare, Ralimanana, Helene, Jeannoda, Vololoniaina, Cable, Stuart &amp; Niskanen, Tuula" docDate="2021" docId="923E7A0AB905430FFF21F8AEFE177EB3" docLanguage="en" docName="phytotaxa.500.1.4.pdf" docOrigin="Phytotaxa 500 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.500.1.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Gloeocantharellus andasibensis Ralaiv., Niskanen &amp; Liimat. 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="34" masterDocId="6E070272B906430AFFA9FFDAFFA17C02" masterDocTitle="Gloeocantharellus andasibensis sp. nov. (Gomphaceae) from Madagascar" masterLastPageNumber="36" masterPageNumber="29" pageNumber="32" updateTime="1699258330370" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8287B2EC6F5B4012BDE1263585CB6B39">Gloeocantharellus andasibensis sp. nov. (Gomphaceae) from Madagascar</mods:title>
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<mods:name id="0474CD0812101FAC79401D18F81D51AC" type="personal">
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<mods:namePart id="8A951AC547D8C98AE07E1D7405AD09EF">Ralaiveloarisoa, Anna Berthe</mods:namePart>
<mods:nameIdentifier id="0FF6A5EBAD130E744C91E9C6114FF0B6" type="ORCID">0000-0002-4046-9028</mods:nameIdentifier>
<mods:affiliation id="0FBE7BC032B1D6E44A7CE8A5B8EAEA7F">University of Antananarivo Madagascar, BP 906, Antananarivo 101, Madagascar &amp; Royal Botanic Gardens, Kew, Lot II J 131 B Ambodivoanjo, Ivandry 101 Antananarivo, Madagascar &amp; A. Ralaiveloarisoa @ kew. org; annaralaivelo @ yahoo. fr; https: // orcid. org / 0000 - 0002 - 4046 - 9028</mods:affiliation>
<mods:nameIdentifier id="3887BE0A1466D7ACBFD2CBD3BFABC7BD" type="email">alaiveloarisoa@kew.org</mods:nameIdentifier>
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<mods:namePart id="0190D7B11CEFF90399F9341DD2336CC6">Liimatainen, Kare</mods:namePart>
<mods:nameIdentifier id="CE70C7D14BCC744DBACC7E589E6FABD4" type="ORCID">0000-0002-5422-2384</mods:nameIdentifier>
<mods:affiliation id="3DE45565722C5EDB93E897EF28AA705C">Jodrell Laboratory, Royal Botanic Gardens, Kew Surrey TW 9 3 AB, United Kingdom &amp; K. Liimatainen @ kew. org; https: // orcid. org / 0000 - 0002 - 5422 - 2384</mods:affiliation>
<mods:nameIdentifier id="20FC2A521C416116BED1594FF0690862" type="email">iimatainen@kew.org</mods:nameIdentifier>
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<mods:namePart id="F889E28897C2D02CF31D0F2BDA35D223">Ralimanana, Helene</mods:namePart>
<mods:nameIdentifier id="3A8C38DE0AB9A5AE4E558F41A0D84535" type="ORCID">0000-0002-7963-5644</mods:nameIdentifier>
<mods:affiliation id="90C98D30C941DBEB540ABC4796245C57">Royal Botanic Gardens, Kew, Lot II J 131 B Ambodivoanjo, Ivandry 101 Antananarivo, Madagascar &amp; H. Ralimanana @ kew. org; https: // orcid. org / 0000 - 0002 - 7963 - 5644</mods:affiliation>
<mods:nameIdentifier id="75FBA5F07478E2EBCA31D6285CCB14B7" type="email">alimanana@kew.org</mods:nameIdentifier>
</mods:name>
<mods:name id="F4AE3880ED8B9C8FB6B8C2C5BD8CFBAF" type="personal">
<mods:role id="1766E31A186A2942F530E66A97ACFD8C">
<mods:roleTerm id="03670192CB265D05B866A008459934AA">Author</mods:roleTerm>
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<mods:namePart id="5D58781FD007FE870956DBFB6AEA0B1D">Jeannoda, Vololoniaina</mods:namePart>
<mods:nameIdentifier id="7D53186DD0B7BD245F1E0D42D5324078" type="ORCID">0000-0002-5178-6889</mods:nameIdentifier>
<mods:affiliation id="6610B6B5B900BE057DC82274154A4C7A">University of Antananarivo Madagascar, BP 906, Antananarivo 101, Madagascar &amp; vololoniaina. jeannoda @ gmail. com; https: // orcid. org / 0000 - 0002 - 5178 - 6889</mods:affiliation>
<mods:nameIdentifier id="69395D8BA004BBBEE419B8E0BB1226AE" type="email">vololoniaina.jeannoda@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B63CDF4187B05D1244BB17974D2BAFD3">Cable, Stuart</mods:namePart>
<mods:nameIdentifier id="563AA6D6D3822008E8A4DB48A1A5E79A" type="ORCID">0000-0001-7119-9844</mods:nameIdentifier>
<mods:affiliation id="21AF9B8A30AFF337A5C0CA68E2D2BF1C">Jodrell Laboratory, Royal Botanic Gardens, Kew Surrey TW 9 3 AB, United Kingdom &amp; S. Cable @ kew. org; https: // orcid. org / 0000 - 0001 - 7119 - 9844</mods:affiliation>
<mods:nameIdentifier id="9A096B260079FA816F9EDAA064551E88" type="email">able@kew.org</mods:nameIdentifier>
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<mods:namePart id="E6F5DBB4A2853FA04447EFC142929478">Niskanen, Tuula</mods:namePart>
<mods:nameIdentifier id="9BCB597F87E176EED69CC44B07A0001D" type="ORCID">0000-0003-1479-5548</mods:nameIdentifier>
<mods:affiliation id="B3F7711D480E6DE4082E8809F5F5F31C">Jodrell Laboratory, Royal Botanic Gardens, Kew Surrey TW 9 3 AB, United Kingdom &amp; T. Niskanen @ kew. org; https: // orcid. org / 0000 - 0003 - 1479 - 5548</mods:affiliation>
<mods:nameIdentifier id="164CCF341D9D47E4703645C07F5493F6" type="email">iskanen@kew.org</mods:nameIdentifier>
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<mods:titleInfo id="E1966E0CFF4174152CA3A20F46E06F5F">
<mods:title id="47F099D28E7F2FC6CB07EBD64AE8B6B8">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="D1C3A44279C53BFDAA27C75556365F65">
<mods:date id="42AA1DD7CCFCEC5BED01B9BB435B4189">2021</mods:date>
<mods:detail id="DE53582CE815E3E2637372E78D565610" type="pubDate">
<mods:number id="6C167EC2657C45AD6596DD441F473EE2">2021-05-07</mods:number>
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<mods:detail id="4AF05D2100E5CFAFE6278DCB4059BDEB" type="volume">
<mods:number id="F5CC7F37F32B48E7DA27F4A54BCF7107">500</mods:number>
</mods:detail>
<mods:detail id="96645A11406FAFB4F6038429315F2015" type="issue">
<mods:number id="0B3F143745B5C02FAAF3412C4E5589C5">1</mods:number>
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<mods:start id="29E1464243FDE611E7F245C3D5BD2B00">29</mods:start>
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<mods:url id="BC574DD8ED121EEE64C2E0937C95E205">http://dx.doi.org/10.11646/phytotaxa.500.1.4</mods:url>
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<mods:classification id="AA952D11D8E5EE618CF1ABDC154CF92B">journal article</mods:classification>
<mods:identifier id="3DF44940251EFBBB6A3E4046E65DEED1" type="DOI">10.11646/phytotaxa.500.1.4</mods:identifier>
<mods:identifier id="4D2035233852DAD9D8BDB5CF58C0211C" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="AAB13A0AF093B27006E7AF80395D5790" type="Zenodo-Dep">5424389</mods:identifier>
</mods:mods>
<treatment id="923E7A0AB905430FFF21F8AEFE177EB3" LSID="urn:lsid:plazi:treatment:923E7A0AB905430FFF21F8AEFE177EB3" httpUri="http://treatment.plazi.org/id/923E7A0AB905430FFF21F8AEFE177EB3" lastPageId="5" lastPageNumber="34" pageId="3" pageNumber="32">
<subSubSection id="528D9897B9054309FF21F8AEFE017BB1" pageId="3" pageNumber="32" type="nomenclature">
<paragraph id="1A28CB1CB9054309FF21F8AEFC4D7B8C" blockId="3.[136,1004,1908,1935]" box="[136,1004,1908,1935]" pageId="3" pageNumber="32">
<heading id="41607C70B9054309FF21F8AEFC4D7B8C" box="[136,1004,1908,1935]" fontSize="11" level="2" pageId="3" pageNumber="32" reason="3">
<taxonomicName id="DD97B09FB9054309FF21F8AEFC9A7B8C" ID-CoL="9Z7SW" authority="Ralaiv., Niskanen &amp; Liimat." authorityName="Ralaiv., Niskanen &amp; Liimat." authorityYear="2021" box="[136,827,1908,1935]" class="Agaricomycetes" family="Gomphaceae" genus="Gloeocantharellus" kingdom="Fungi" order="Gomphales" pageId="3" pageNumber="32" phylum="Basidiomycota" rank="species" species="andasibensis" status="sp. nov.">
<emphasis id="28E3170EB9054309FF21F8AEFE597B8D" bold="true" box="[136,504,1908,1935]" italics="true" pageId="3" pageNumber="32">Gloeocantharellus andasibensis</emphasis>
Ralaiv., Niskanen &amp; Liimat.
</taxonomicName>
<taxonomicNameLabel id="33D0AA75B9054309FCE8F8AFFC397B8D" box="[833,920,1909,1935]" pageId="3" pageNumber="32" rank="species">
<emphasis id="28E3170EB9054309FCE8F8AFFC317B8C" box="[833,912,1909,1934]" italics="true" pageId="3" pageNumber="32">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="82ACD799B9054309FC0EF8AEFC427B8C" box="[935,995,1908,1934]" captionStart="FIGURE 2" captionStartId="4.[136,229,1835,1857]" captionTargetBox="[318,1270,387,1802]" captionTargetId="figure-117@4.[318,1270,387,1811]" captionTargetPageId="4" captionText="FIGURE 2. a. Basidiomata, b. Basidiospores in Cotton Blue, c. Gloeocystidia with contents in KOH d. Gloeocystidia without contents of Gloeocantharellus andasibensis in KOH (Holotype MAF18-226 (TAN!), Isotype K(M) 255527(K!)). Photographs: T. Niskanen and A. Ralaiveloarisoa. Scale bars a = 1 cm, b–d = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5424393" httpUri="https://zenodo.org/record/5424393/files/figure.png" pageId="3" pageNumber="32">Fig 2</figureCitation>
)
</heading>
</paragraph>
<paragraph id="1A28CB1CB9054309FF21F842FE017BB1" blockId="3.[136,416,1944,1971]" box="[136,416,1944,1971]" pageId="3" pageNumber="32">Index fungorum: 558149</paragraph>
</subSubSection>
<subSubSection id="528D9897B905430FFF21F83EFE177EB3" lastPageId="5" lastPageNumber="34" pageId="3" pageNumber="32" type="diagnosis">
<paragraph id="1A28CB1CB905430EFF21F83EFBAB7CDB" blockId="3.[136,1452,2020,2078]" lastBlockId="4.[136,1452,159,361]" lastPageId="4" lastPageNumber="33" pageId="3" pageNumber="32">
Diagnosis:—
<taxonomicName id="DD97B09FB9054309FEAEF83EFD9F7BFB" box="[263,574,2020,2041]" class="Agaricomycetes" family="Gomphaceae" genus="Gloeocantharellus" kingdom="Fungi" order="Gomphales" pageId="3" pageNumber="32" phylum="Basidiomycota" rank="species" species="andasibensis">
<emphasis id="28E3170EB9054309FEAEF83EFD9F7BFB" box="[263,574,2020,2041]" italics="true" pageId="3" pageNumber="32">Gloeocantharellus andasibensis</emphasis>
</taxonomicName>
differs from its closest relatives,
<taxonomicName id="DD97B09FB9054309FC3AF83EFB927BFB" authorityName="Corner" authorityYear="1969" box="[915,1075,2020,2041]" class="Agaricomycetes" family="Gomphaceae" genus="Gloeocantharellus" kingdom="Fungi" order="Gomphales" pageId="3" pageNumber="32" phylum="Basidiomycota" rank="species" species="echinosporus">
<emphasis id="28E3170EB9054309FC3AF83EFB927BFB" box="[915,1075,2020,2041]" italics="true" pageId="3" pageNumber="32">G. echinosporus</emphasis>
</taxonomicName>
and
<taxonomicName id="DD97B09FB9054309FBC0F83EFA8C7BFB" authorityName="Ming Zhang &amp; T. H. Li" authorityYear="2019" box="[1129,1325,2020,2041]" class="Agaricomycetes" family="Gomphaceae" genus="Gloeocantharellus" kingdom="Fungi" order="Gomphales" pageId="3" pageNumber="32" phylum="Basidiomycota" rank="species" species="neoechinosporus">
<emphasis id="28E3170EB9054309FBC0F83EFA8C7BFB" box="[1129,1325,2020,2041]" italics="true" pageId="3" pageNumber="32">G. neoechinosporus</emphasis>
</taxonomicName>
by its small basidia and small, verrucose basidiospores. In addition,
<taxonomicName id="DD97B09FB9054309FD75F7D2FC3C741F" authorityName="Ming Zhang &amp; T. H. Li" authorityYear="2019" box="[732,925,2056,2077]" class="Agaricomycetes" family="Gomphaceae" genus="Gloeocantharellus" kingdom="Fungi" order="Gomphales" pageId="3" pageNumber="32" phylum="Basidiomycota" rank="species" species="neoechinosporus">
<emphasis id="28E3170EB9054309FD75F7D2FC3C741F" box="[732,925,2056,2077]" italics="true" pageId="3" pageNumber="32">G. neoechinosporus</emphasis>
</taxonomicName>
can be distinguished by its pale yellow/pale orange to greyish orange hymenophore turning pale brown to dark brown when bruised, and the presence of a weak superior annulus; while
<taxonomicName id="DD97B09FB902430EFA3DFF45FE9C7CDA" authorityName="Corner" authorityYear="1969" class="Agaricomycetes" family="Gomphaceae" genus="Gloeocantharellus" kingdom="Fungi" order="Gomphales" pageId="4" pageNumber="33" phylum="Basidiomycota" rank="species" species="echinosporus">
<emphasis id="28E3170EB902430EFA3DFF45FE9C7CDA" italics="true" pageId="4" pageNumber="33">G. echinosporus</emphasis>
</taxonomicName>
has a squamulose pileus and lacks or only has a few gloeoplerous hyphae.
</paragraph>
<paragraph id="1A28CB1CB902430EFF21FF3DFED37D6B" blockId="4.[136,1452,159,361]" pageId="4" pageNumber="33">
Type:—
<collectingCountry id="62808B8CB902430EFF7CFF3DFE257CFE" box="[213,388,231,252]" name="Madagascar" pageId="4" pageNumber="33">MADAGASCAR</collectingCountry>
.
<collectingRegion id="D85305FEB902430EFE24FF3DFE587CFF" box="[397,505,231,253]" country="Madagascar" name="Toamasina" pageId="4" pageNumber="33">Toamasina</collectingRegion>
: Moramanga district, Mantadia National Park, elev.,
<quantity id="DD6F66F9B902430EFC5AFF3DFBCD7CFF" box="[1011,1132,231,253]" metricMagnitude="2" metricUnit="m" metricValue="9.975" metricValueMax="10.15" metricValueMin="9.8" pageId="4" pageNumber="33" unit="m" value="997.5" valueMax="1015.0" valueMin="980.0">980–1015 m</quantity>
a.s.l.,
<geoCoordinate id="7FA3ADDBB902430EFB03FF3DFA9E7CFE" box="[1194,1343,231,252]" degrees="18" direction="south" minutes="47" orientation="latitude" pageId="4" pageNumber="33" precision="1" seconds="44.3" value="-18.795639">S 18° 47’ 44.3’’</geoCoordinate>
<geoCoordinate id="7FA3ADDBB902430EFAECFF32FF517D23" degrees="048" direction="east" minutes="25" orientation="longitude" pageId="4" pageNumber="33" precision="1" seconds="36.9" value="48.426918">E 048° 25’ 36.9</geoCoordinate>
’‘, a forest dominated by
<taxonomicName id="DD97B09FB902430EFE76FED6FD287D22" box="[479,649,267,289]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="species" species="thouarsii">
<emphasis id="28E3170EB902430EFE76FED6FD287D22" box="[479,649,267,289]" italics="true" pageId="4" pageNumber="33">Uapaca thouarsii</emphasis>
</taxonomicName>
,
<taxonomicName id="DD97B09FB902430EFD3FFED6FCDE7D22" box="[662,895,267,289]" class="Magnoliopsida" family="Cunoniaceae" genus="Weinmannia" kingdom="Plantae" order="Oxalidales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="species" species="rutenbergii">
<emphasis id="28E3170EB902430EFD3FFED6FCDE7D22" box="[662,895,267,289]" italics="true" pageId="4" pageNumber="33">Weinmannia rutenbergii</emphasis>
</taxonomicName>
,
<taxonomicName id="DD97B09FB902430EFC25FED1FBBB7D23" box="[908,1050,267,289]" class="Magnoliopsida" family="Clusiaceae" genus="Symphonia" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="28E3170EB902430EFC25FED1FBBB7D23" box="[908,1050,267,289]" italics="true" pageId="4" pageNumber="33">Symphonia sp.</emphasis>
</taxonomicName>
,
<taxonomicName id="DD97B09FB902430EFB8FFED1FB2D7D23" box="[1062,1164,267,289]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
,
<taxonomicName id="DD97B09FB902430EFB3EFED6FAAB7D23" box="[1175,1290,268,289]" class="Magnoliopsida" family="Myrtaceae" genus="Eugenia" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="28E3170EB902430EFB3EFED6FAAB7D23" box="[1175,1290,268,289]" italics="true" pageId="4" pageNumber="33">Eugenia sp.</emphasis>
</taxonomicName>
,
<taxonomicName id="DD97B09FB902430EFABEFED1FA097D23" box="[1303,1448,267,289]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
, and
<taxonomicName id="DD97B09FB902430EFF40FEF5FE1E7D46" authorityName="Thouars" authorityYear="1805" box="[233,447,303,324]" class="Magnoliopsida" family="Sarcolaenaceae" genus="Sarcolaena" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="species" species="eriophora">
<emphasis id="28E3170EB902430EFF40FEF5FE1E7D46" box="[233,447,303,324]" italics="true" pageId="4" pageNumber="33">Sarcolaena eriophora</emphasis>
</taxonomicName>
,
<date id="6E29EDDCB902430EFE67FEF5FD217D46" box="[462,640,303,325]" pageId="4" pageNumber="33" value="2018-02-24">24 February 2018</date>
,
<emphasis id="28E3170EB902430EFD24FEEAFCA07D46" box="[653,769,303,325]" italics="true" pageId="4" pageNumber="33">T. Niskanen</emphasis>
(
<typeStatus id="C52C75BEB902430EFCB8FEF5FCCD7D47" box="[785,876,303,325]" pageId="4" pageNumber="33" type="holotype">Holotype</typeStatus>
MAF18-226(TAN!), Isotype K(M)255527(K!), GenBank:
<accessionNumber id="05C456FFB902430EFF14FE89FE907D6B" box="[189,305,339,361]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MW506830" pageId="4" pageNumber="33" type="GenBank">MW506830</accessionNumber>
(ITS).
</paragraph>
<caption id="4EE89B94B902430EFF21F8F1FDD97B8B" ID-DOI="http://doi.org/10.5281/zenodo.5424393" ID-Zenodo-Dep="5424393" httpUri="https://zenodo.org/record/5424393/files/figure.png" pageId="4" pageNumber="33" startId="4.[136,229,1835,1857]" targetBox="[318,1270,387,1802]" targetPageId="4">
<paragraph id="1A28CB1CB902430EFF21F8F1FDD97B8B" blockId="4.[136,1452,1835,1930]" pageId="4" pageNumber="33">
<emphasis id="28E3170EB902430EFF21F8F1FF597B43" bold="true" box="[136,248,1835,1857]" pageId="4" pageNumber="33">FIGURE 2</emphasis>
.
<emphasis id="28E3170EB902430EFEACF8F6FEB07B43" bold="true" box="[261,273,1836,1857]" pageId="4" pageNumber="33">a</emphasis>
. Basidiomata,
<emphasis id="28E3170EB902430EFE0BF8F6FE0E7B43" bold="true" box="[418,431,1836,1857]" pageId="4" pageNumber="33">b</emphasis>
. Basidiospores in Cotton Blue,
<emphasis id="28E3170EB902430EFD4CF8F6FD517B43" bold="true" box="[741,752,1836,1857]" pageId="4" pageNumber="33">c</emphasis>
. Gloeocystidia with contents in KOH
<emphasis id="28E3170EB902430EFBCCF8F6FBD37B43" bold="true" box="[1125,1138,1836,1857]" pageId="4" pageNumber="33">d</emphasis>
. Gloeocystidia without contents of
<taxonomicName id="DD97B09FB902430EFF0BF88AFE747B67" box="[162,469,1872,1893]" class="Agaricomycetes" family="Gomphaceae" genus="Gloeocantharellus" kingdom="Fungi" order="Gomphales" pageId="4" pageNumber="33" phylum="Basidiomycota" rank="species" species="andasibensis">
<emphasis id="28E3170EB902430EFF0BF88AFE747B67" box="[162,469,1872,1893]" italics="true" pageId="4" pageNumber="33">Gloeocantharellus andasibensis</emphasis>
</taxonomicName>
in KOH (Holotype MAF18-226 (TAN!), Isotype K(M) 255527(K!)). Photographs: T. Niskanen and A. Ralaiveloarisoa. Scale bars
<emphasis id="28E3170EB902430EFE38F8AEFE3C7B8B" bold="true" box="[401,413,1908,1929]" pageId="4" pageNumber="33">a</emphasis>
= 1 cm,
<emphasis id="28E3170EB902430EFE5BF8AEFDB97B8B" bold="true" box="[498,536,1908,1929]" pageId="4" pageNumber="33">b–d</emphasis>
= 10 µm.
</paragraph>
</caption>
<paragraph id="1A28CB1CB902430EFF21F862FCD37BD0" blockId="4.[136,1452,1976,2075]" box="[136,882,1976,2003]" pageId="4" pageNumber="33">
Etymology:—The species epithet is derived from the
<typeStatus id="C52C75BEB902430EFD48F863FCB07BD1" box="[737,785,1977,2003]" pageId="4" pageNumber="33">type</typeStatus>
locality.
</paragraph>
<paragraph id="1A28CB1CB902430FFF14F806FAEC7CDE" blockId="4.[136,1452,1976,2075]" lastBlockId="5.[136,1452,158,689]" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33">
Description:—
<emphasis id="28E3170EB902430EFECDF806FE0B7BF4" box="[356,426,2012,2038]" italics="true" pageId="4" pageNumber="33">Pileus</emphasis>
<quantity id="DD6F66F9B902430EFE06F806FDA17BF5" box="[431,512,2012,2039]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="4" pageNumber="33" unit="cm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 cm</quantity>
in diameter, at first convex then plane, depressed in the centre, orange-red, somewhat innately fibrillose, viscid.
<emphasis id="28E3170EB902430EFE1DF7DAFDF37418" box="[436,594,2048,2074]" italics="true" pageId="4" pageNumber="33">Hymenophore</emphasis>
decurrent, crowded, at first white, turning blackish on handling.
<emphasis id="28E3170EB902430EFAE9F7DAFAD97418" box="[1344,1400,2048,2074]" italics="true" pageId="4" pageNumber="33">Stipe</emphasis>
5–8 ×
<quantity id="DD6F66F9B903430FFF09FF44FEAD7CBB" box="[160,268,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="5" pageNumber="34" unit="cm" value="1.25" valueMax="2.0" valueMin="0.5">0.5–2 cm</quantity>
, cylindrical, tapered at the base, orange-red to rusty red, becoming blackish rose when handled.
<emphasis id="28E3170EB903430FFAFDFF45FA0A7CBB" box="[1364,1451,159,185]" italics="true" pageId="5" pageNumber="34">Context</emphasis>
white.
<emphasis id="28E3170EB903430FFF7DFF19FEF37CDE" box="[212,338,195,221]" italics="true" pageId="5" pageNumber="34">Spore print</emphasis>
white.
<emphasis id="28E3170EB903430FFE0CFF18FE417CDE" box="[421,480,194,220]" italics="true" pageId="5" pageNumber="34">Odor</emphasis>
in hymenophore indistinct, not observed in the other parts of the basidiomata.
</paragraph>
<paragraph id="1A28CB1CB903430FFF14FF3CFF4E7DDA" blockId="5.[136,1452,158,689]" pageId="5" pageNumber="34">
<emphasis id="28E3170EB903430FFF14FF3CFEFA7D02" box="[189,347,230,256]" italics="true" pageId="5" pageNumber="34">Basidiospores</emphasis>
7.5–9 × 4.5–5.5 µm (av.=8.4 × 5 µm, Q=1.45–1.95, Q
<subScript id="8613C959B903430FFC0CFF23FC187D05" attach="left" box="[933,953,249,263]" fontSize="6" pageId="5" pageNumber="34">av.</subScript>
=1.7 µm), ellipsoid to somewhat amygdaloidobovoid, verrucose, ornamentation distinct, up to 0.5 µm high, cyanophilic with Cotton Blue.
<emphasis id="28E3170EB903430FFB6EFED0FABC7D26" box="[1223,1309,266,292]" italics="true" pageId="5" pageNumber="34">Basidia</emphasis>
25–40 × 9– 10.5 µm, clavate, hyaline, with 2–4 sterigmata (1.5–6 µm long).
<emphasis id="28E3170EB903430FFCCFFEF4FC627D4A" box="[870,963,302,328]" italics="true" pageId="5" pageNumber="34">Cystidia</emphasis>
50–60(–80) × 7–9.5 µm, lanceolate, rarely subclavate, apically attenuated, cyanophilic with Cotton Blue, with or without yellow granulose contents in KOH, with basal clamp connections.
<emphasis id="28E3170EB903430FFE06FEACFD807D92" box="[431,545,374,400]" italics="true" pageId="5" pageNumber="34">Pileipellis</emphasis>
an ixocutis composed of hyphae 2–4.5 µm diam, with yellow-green granulose contents or without and then pale green and transparent, wall hyaline; some hyphae up to 6 µm in diameter, yellowish, clamped.
</paragraph>
<paragraph id="1A28CB1CB903430FFF14FE38FA267E47" blockId="5.[136,1452,158,689]" pageId="5" pageNumber="34">
Habitat and distribution:—on soil among grasses.
<collectingCountry id="62808B8CB903430FFD41FE38FCD57DFE" box="[744,884,482,508]" name="Madagascar" pageId="5" pageNumber="34">Madagascar</collectingCountry>
: in undisturbed primary humid forest dominated by
<taxonomicName id="DD97B09FB903430FFF21FDDDFDFB7E23" authority="Baill." authorityName="Baill." box="[136,602,518,545]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="species" species="thouarsii">
<emphasis id="28E3170EB903430FFF21FDDDFEEC7E22" box="[136,333,518,544]" italics="true" pageId="5" pageNumber="34">Uapaca thouarsii</emphasis>
Baill. (Phyllanthaceae)
</taxonomicName>
,
<taxonomicName id="DD97B09FB903430FFDCCFDDDFC197E22" authority="Engl." authorityName="Engl." box="[613,952,518,544]" class="Magnoliopsida" family="Cunoniaceae" genus="Weinmannia" kingdom="Plantae" order="Oxalidales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="species" species="rutenbergii">
<emphasis id="28E3170EB903430FFDCCFDDDFCD47E22" box="[613,885,518,544]" italics="true" pageId="5" pageNumber="34">Weinmannia rutenbergii</emphasis>
Engl.
</taxonomicName>
Cunnoniaceae,
<taxonomicName id="DD97B09FB903430FFBC4FDDCFA067E23" box="[1133,1447,518,545]" class="Magnoliopsida" family="Clusiaceae" genus="Symphonia" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="28E3170EB903430FFBC4FDDCFAAB7E22" box="[1133,1290,518,544]" italics="true" pageId="5" pageNumber="34">Symphonia sp</emphasis>
. (Clusiaceae)
</taxonomicName>
,
<taxonomicName id="DD97B09FB903430FFF21FDF0FE1B7E47" box="[136,442,554,581]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="28E3170EB903430FFF21FDF0FE807E46" box="[136,289,554,580]" italics="true" pageId="5" pageNumber="34">Psychotria sp</emphasis>
. (Rubiaceae)
</taxonomicName>
,
<taxonomicName id="DD97B09FB903430FFE6CFDF1FD7D7E46" box="[453,732,554,581]" class="Magnoliopsida" family="Myrtaceae" genus="Eugenia" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="28E3170EB903430FFE6CFDF1FDE37E46" box="[453,578,555,580]" italics="true" pageId="5" pageNumber="34">Eugenia sp</emphasis>
. (Myrtaceae)
</taxonomicName>
,
<taxonomicName id="DD97B09FB903430FFD4EFDF0FC2C7E46" box="[743,909,554,580]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
and
<taxonomicName id="DD97B09FB903430FFC6AFDF0FA227E47" authorityName="Thouars" authorityYear="1805" box="[963,1411,554,581]" class="Magnoliopsida" family="Sarcolaenaceae" genus="Sarcolaena" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="species" species="eriophora">
<emphasis id="28E3170EB903430FFC6AFDF0FB187E46" box="[963,1209,554,580]" italics="true" pageId="5" pageNumber="34">Sarcolaena eriophora</emphasis>
(Sarcolaenaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="1A28CB1CB903430FFF21FD94FE177EB3" blockId="5.[136,1452,158,689]" pageId="5" pageNumber="34">
<materialsCitation id="AAFFC141B903430FFF21FD94FE127EB3" collectingDate="2018-02-24" collectionCode="T, TAN, K, M" country="Madagascar" county="Specimen" elevation="998" latitude="-18.795639" location="Mantadia National Park" longLatPrecision="1" longitude="48.426918" municipality="Moramanga district" pageId="5" pageNumber="34" specimenCode="MAF18-226" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<collectingCounty id="F349B390B903430FFF21FD94FF577E6A" box="[136,246,590,616]" pageId="5" pageNumber="34">Specimen</collectingCounty>
examined:—
<collectingCountry id="62808B8CB903430FFE27FD94FDFA7E6A" box="[398,603,590,616]" name="Madagascar" pageId="5" pageNumber="34">MADAGASCAR</collectingCountry>
.
<collectingRegion id="D85305FEB903430FFDCCFD94FD437E6A" box="[613,738,590,616]" country="Madagascar" name="Toamasina" pageId="5" pageNumber="34">Toamasina</collectingRegion>
:
<collectingMunicipality id="FA4C5166B903430FFD47FD95FC6E7E6A" box="[750,975,590,617]" pageId="5" pageNumber="34">Moramanga district</collectingMunicipality>
,
<location id="1F489DC7B903430FFC72FD94FB4D7E6A" LSID="urn:lsid:plazi:treatment:923E7A0AB905430FFF21F8AEFE177EB3:1F489DC7B903430FFC72FD94FB4D7E6A" box="[987,1260,590,616]" country="Madagascar" county="Specimen" latitude="-18.795639" longLatPrecision="1" longitude="48.426918" municipality="Moramanga district" name="Mantadia National Park" pageId="5" pageNumber="34" stateProvince="Toamasina">Mantadia National Park</location>
, elev.,
<elevation id="91BA2C2FB903430FFA92FD94FF7D7E8E" metricMagnitude="2" metricUnit="m" metricValue="9.975" metricValueMax="10.15" metricValueMin="9.8" pageId="5" pageNumber="34" unit="m" value="997.5" valueMax="1015.0" valueMin="980.0">
<quantity id="DD6F66F9B903430FFA92FD94FF3F7E8F" metricMagnitude="2" metricUnit="m" metricValue="9.975" metricValueMax="10.15" metricValueMin="9.8" pageId="5" pageNumber="34" unit="m" value="997.5" valueMax="1015.0" valueMin="980.0">980–1015 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="7FA3ADDBB903430FFF40FDA8FE3D7E8E" box="[233,412,626,652]" degrees="18" direction="south" minutes="47" orientation="latitude" pageId="5" pageNumber="34" precision="1" seconds="44.3" value="-18.795639">S 18° 47’ 44.3’’</geoCoordinate>
<geoCoordinate id="7FA3ADDBB903430FFE0DFDA9FDC37E8E" box="[420,610,626,653]" degrees="048" direction="east" minutes="25" orientation="longitude" pageId="5" pageNumber="34" precision="1" seconds="36.9" value="48.426918">E 048° 25’ 36.9</geoCoordinate>
’‘,
<date id="6E29EDDCB903430FFDD0FDA8FCA67E8E" box="[633,775,626,652]" pageId="5" pageNumber="34" value="2018-02-24">
<collectingDate id="7E6D1434B903430FFDD0FDA8FCA67E8E" box="[633,775,626,652]" pageId="5" pageNumber="34" value="2018-02-24">24 Feb 2018</collectingDate>
</date>
(rainy season),
<emphasis id="28E3170EB903430FFC17FDA9FBE27E8E" box="[958,1091,626,652]" italics="true" pageId="5" pageNumber="34">
<collectionCode id="7C8653D9B903430FFC17FDA9FC6E7E8E" box="[958,975,627,652]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="34">T</collectionCode>
. Niskanen
</emphasis>
(
<typeStatus id="C52C75BEB903430FFBFAFDA8FB1C7E8E" box="[1107,1213,626,652]" pageId="5" pageNumber="34" type="holotype">Holotype</typeStatus>
<specimenCode id="4A316367B903430FFB6CFDA8FAF77E8F" box="[1221,1366,626,653]" collectionCode="MAF" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15320" lsid="urn:lsid:biocol.org:col:15320" name="Universidad Complutense" pageId="5" pageNumber="34" type="Herbarium">MAF18-226</specimenCode>
(
<collectionCode id="7C8653D9B903430FFAF4FDA8FA3D7E8F" box="[1373,1436,626,653]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="5" pageNumber="34" type="Herbarium">TAN</collectionCode>
!), Isotype
<collectionCode id="7C8653D9B903430FFF48FD4CFF567EB3" box="[225,247,662,689]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="34" type="Herbarium">K</collectionCode>
(
<collectionCode id="7C8653D9B903430FFF56FD4CFEBB7EB3" box="[255,282,662,689]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="5" pageNumber="34" type="Herbarium">M</collectionCode>
)255527(
<collectionCode id="7C8653D9B903430FFE2EFD4CFE3C7EB3" box="[391,413,662,689]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="34" type="Herbarium">K</collectionCode>
!))
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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