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<document id="C1FBA77D9EF62203D5B6835828107ADB" ID-DOI="10.17109/AZH.63.2.143.2017" ID-ISSN="2064-2474" ID-Zenodo-Dep="12625425" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638279146406" checkinUser="felipe" docAuthor="Makranczy, György" docDate="2017" docId="03B24C0FFFA9FFE89991FF7DFC87F23B" docLanguage="en" docName="ActaZoolAcadSciH.63.2.143-262.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 63 (2)" docSource="http://dx.doi.org/10.17109/azh.63.2.143.2017" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.6:ActaZoolAcadSciH.2015-2023.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-2023.journal_article" docStyleVersion="6" docTitle="Anotylus schawalleri Makranczy, 2017, sp. n." docType="treatment" docUuid="F6CA242C-828A-4787-8037-61B2FFD281E1" docUuidSource="ZooBank" docVersion="2" lastPageNumber="250" masterDocId="FF8B3477FFC0FF839847FFC2FFBCF364" masterDocTitle="Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="262" masterPageNumber="143" pageNumber="248" updateTime="1719925380098" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="7F763A937EC25E389B884A6B63CB5C5A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="76F191946E9460F95E54E8F38521DD93">Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:namePart id="B3748051EAE8C4DCD7EE7AA17B32F30D">Makranczy, György</mods:namePart>
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<mods:title id="6F97F687B2BCB69B470E9B8467823F29">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
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<mods:date id="7E79B8926C9614599DC0B53B15D08DB2">2017</mods:date>
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<mods:number id="8285DC361537A872C56C1AD684DEE2F9">2017-06-19</mods:number>
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<mods:identifier id="52C9475C9E85CFE50E8A2DEB506A418D" type="DOI">10.17109/AZH.63.2.143.2017</mods:identifier>
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</mods:mods>
<treatment id="03B24C0FFFA9FFE89991FF7DFC87F23B" LSID="urn:lsid:zoobank.org:act:F6CA242C-828A-4787-8037-61B2FFD281E1" httpUri="http://treatment.plazi.org/id/03B24C0FFFA9FFE89991FF7DFC87F23B" lastPageId="107" lastPageNumber="250" pageId="105" pageNumber="248">
<subSubSection id="C301AE92FFA9FFEA9991FF7DFC81F3BD" box="[470,829,191,217]" pageId="105" pageNumber="248" type="nomenclature">
<paragraph id="8BA4FD19FFA9FFEA9991FF7DFC81F3BD" blockId="105.[470,829,191,217]" box="[470,829,191,217]" pageId="105" pageNumber="248">
<heading id="D0EC4A75FFA9FFEA9991FF7DFC81F3BD" box="[470,829,191,217]" centered="true" fontSize="11" level="2" pageId="105" pageNumber="248" reason="2">
<taxonomicName id="4C1B869AFFA9FFEA9991FF7DFD4CF3BD" ID-CoL="66YDL" authorityName="Makranczy" authorityYear="2017" box="[470,752,191,217]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="248" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov.">
<emphasis id="B96F210BFFA9FFEA9991FF7DFD4CF3BD" bold="true" box="[470,752,191,217]" pageId="105" pageNumber="248">Anotylus schawalleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C9C70FFA9FFEA9ABFFF7DFC81F3BD" box="[760,829,191,217]" pageId="105" pageNumber="248" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C301AE92FFA9FFE899BCFF20FC87F23B" lastPageId="107" lastPageNumber="250" pageId="105" pageNumber="248" type="description">
<paragraph id="8BA4FD19FFA9FFEA99BCFF20FCA4F398" blockId="105.[507,792,226,253]" box="[507,792,226,253]" pageId="105" pageNumber="248">
<heading id="D0EC4A75FFA9FFEA99BCFF20FCA4F398" box="[507,792,226,253]" centered="true" fontSize="11" level="3" pageId="105" pageNumber="248" reason="3">
(
<figureCitation id="1320E19CFFA9FFEA9A44FF20FDDDF398" box="[515,609,226,253]" captionStart="Figs 73–75" captionStartId="40.[136,181,1526,1548]" captionTargetBox="[140,1082,206,1495]" captionTargetId="figure-19@40.[135,1083,197,1501]" captionTargetPageId="40" captionText="Figs 73–75. Habitus of Anotylus species. 73 = A. schawalleri sp. n., 74 = A. borneensis (Cam- eron, 1928), 75 = A. gunung sp. n." figureDoi="http://doi.org/10.5281/zenodo.12625455" httpUri="https://zenodo.org/record/12625455/files/figure.png" pageId="105" pageNumber="248">Figs 73</figureCitation>
,
<figureCitation id="1320E19CFFA9FFEA9A2AFF20FD6BF398" box="[621,727,226,252]" captionStart="Figs 230–236" captionStartId="106.[136,181,1490,1512]" captionTargetBox="[148,1076,191,1464]" captionTargetId="figure-19@106.[147,1076,189,1464]" captionTargetPageId="106" captionText="Figs 230–236. Anotylus schawalleri sp. n., male. 230 = median part of sternite VII, 231 = ster- nite VIII, 232 = tergite IX, 233 = tergite X, 234 = aedeagus, frontal view, 235 = median lobe, lateral view, 236 = paramere, lateral view. Scales: 0.10 mm for Figs 234–236, 0.115 mm for Fig. 233, 0.15 mm for Figs 230–232" figureDoi="http://doi.org/10.5281/zenodo.12625527" httpUri="https://zenodo.org/record/12625527/files/figure.png" pageId="105" pageNumber="248">230–236</figureCitation>
,
<figureCitation id="1320E19CFFA9FFEA9AA4FF20FCADF398" box="[739,785,226,252]" captionStart="Figs 265–278" captionStartId="116.[136,181,1475,1497]" captionTargetBox="[138,1088,607,1439]" captionTargetId="figure-539@116.[137,1089,605,1454]" captionTargetPageId="116" captionText="Figs 265–278. Spermathecae of Anotylus species: 265 = A. coonoor sp. n., 266 = A. ganapati sp. n., 267 = A. hartmanni sp. n., 268 = A. ijen sp. n., 269 = A. jambi sp. n., 270 = A. kabasi sp. n., 271 = A. linaxi sp. n., 272 = A. riedeli sp. n., 273 = A. rurukan sp. n., 274 = A. schawalleri sp. n., 275 = A. schillhammeri sp. n., 276 = A. schuelkei sp. n., 277 = A. tanator sp. n, 278 = A. topali" figureDoi="http://doi.org/10.5281/zenodo.12625539" httpUri="https://zenodo.org/record/12625539/files/figure.png" pageId="105" pageNumber="248">274</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BA4FD19FFA9FFEA98ADFED8FC63F12B" blockId="105.[173,1126,282,593]" pageId="105" pageNumber="248">
<emphasis id="B96F210BFFA9FFEA98ADFED8FECFF24B" box="[234,371,282,303]" italics="true" pageId="105" pageNumber="248">Typematerial</emphasis>
–
<typeStatus id="54A043BBFFA9FFEA99D5FED8FE4AF24B" box="[402,502,282,304]" pageId="105" pageNumber="248" type="holotype">Holotype</typeStatus>
(m):
<collectingCountry id="F30CBD89FFA9FFEA9A76FED8FD09F24B" box="[561,693,282,303]" name="Malaysia" pageId="105" pageNumber="248">MALAYSIA</collectingCountry>
/BORNEO:
<collectingRegion id="49DF33FBFFA9FFEA9B68FED8FCD3F254" box="[815,879,282,304]" country="Malaysia" name="Sabah" pageId="105" pageNumber="248">Sabah</collectingRegion>
, CrockerRangeN. P., NWKeningau,
<quantity id="4CE350FCFFA9FFEA9914FEF5FE72F228" box="[339,462,311,332]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="105" pageNumber="248" unit="m" value="1050.0" valueMax="1200.0" valueMin="900.0">900–1200 m</quantity>
[
<geoCoordinate id="EE2F9BDEFFA9FFEA999DFEF5FD8FF229" box="[474,563,311,333]" degrees="05" direction="north" minutes="26" orientation="latitude" pageId="105" pageNumber="248" precision="925" value="5.4333334">05°26‘N</geoCoordinate>
,
<geoCoordinate id="EE2F9BDEFFA9FFEA9A7BFEF4FD27F228" box="[572,667,310,332]" degrees="116" direction="east" minutes="05" orientation="longitude" pageId="105" pageNumber="248" precision="925" value="116.083336">116°05‘E</geoCoordinate>
],
<date id="FFA5DBD9FFA9FFEA9AECFEF5FC85F229" box="[683,825,311,333]" pageId="105" pageNumber="248" value="1996-11-16" valueMax="1996-11-20" valueMin="1996-11-16">16-20.XI.1996</date>
, leg. W. Schawaller (SMNS);
<typeStatus id="54A043BBFFA9FFEA98EAFE96FEAAF20D" box="[173,278,340,361]" pageId="105" pageNumber="248" type="paratype">Paratypes</typeStatus>
(4): samedataasholotype (
<quantity id="4CE350FCFFA9FFEA9A01FE96FDD6F20D" box="[582,618,339,361]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="105" pageNumber="248" unit="m" value="1.0">1 m</quantity>
, HNHM);
<collectingRegion id="49DF33FBFFA9FFEA9A98FE96FC9CF20D" box="[735,800,339,361]" country="Malaysia" name="Sabah" pageId="105" pageNumber="248">Sabah</collectingRegion>
, CrockerRange,
<quantity id="4CE350FCFFA9FFEA9B90FE91FBDEF20D" box="[983,1122,339,361]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.65" metricValueMin="1.55" pageId="105" pageNumber="248" unit="m" value="1600.0" valueMax="1650.0" valueMin="1550.0">1550–1650 m</quantity>
, àproximitéducol (routeKotaKinabalu-Tambunan) [
<geoCoordinate id="EE2F9BDEFFA9FFEA9B53FEB2FC2CF2E2" box="[788,912,368,390]" degrees="05" direction="north" minutes="47" orientation="latitude" pageId="105" pageNumber="248" precision="15" seconds="55" value="5.798611">05°47‘55“N</geoCoordinate>
,
<geoCoordinate id="EE2F9BDEFFA9FFEA9BD9FEB2FB9EF2E2" box="[926,1058,368,390]" degrees="116" direction="east" minutes="20" orientation="longitude" pageId="105" pageNumber="248" precision="15" seconds="06" value="116.335">116°20‘06“E</geoCoordinate>
], forêt de
<emphasis id="B96F210BFFA9FFEA9888FE4FFE02F2C6" box="[207,446,397,419]" italics="true" pageId="105" pageNumber="248">
<taxonomicName id="4C1B869AFFA9FFEA9888FE4FFEFEF2C6" box="[207,322,397,418]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" pageId="105" pageNumber="248" phylum="Tracheophyta" rank="genus">Lithocarpus</taxonomicName>
-
<taxonomicName id="4C1B869AFFA9FFEA990DFE4FFE02F2C6" box="[330,446,397,418]" class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" kingdom="Plantae" order="Fagales" pageId="105" pageNumber="248" phylum="Tracheophyta" rank="genus">Castanopsis</taxonomicName>
</emphasis>
,
<date id="FFA5DBD9FFA9FFEA998CFE4FFD8DF2C7" box="[459,561,397,419]" pageId="105" pageNumber="248" value="1996-05-16">16.V.1996</date>
, leg. D. Burckhardt &amp; I. Löbl (27a), tamisagedebois pourri, feuillesmortesetmousses (
<quantity id="4CE350FCFFA9FFEA9A69FE68FDEFF2A4" box="[558,595,426,448]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="105" pageNumber="248" unit="m" value="1.0">1 m</quantity>
, MHNG);
<collectingRegion id="49DF33FBFFA9FFEA9A8FFE68FCB5F2A4" box="[712,777,426,448]" country="Malaysia" name="Sabah" pageId="105" pageNumber="248">Sabah</collectingRegion>
, CrockerRange, faceouest (W),
<quantity id="4CE350FCFFA9FFEA98EAFE05FF47F2B8" box="[173,251,455,477]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="105" pageNumber="248" unit="m" value="1600.0">1600 m</quantity>
env., km 51-52 - route de Kota Kinabalu - Tambunan [
<geoCoordinate id="EE2F9BDEFFA9FFEA9B17FE05FC70F2B9" box="[848,972,455,477]" degrees="05" direction="north" minutes="50" orientation="latitude" pageId="105" pageNumber="248" precision="15" seconds="04" value="5.8344445">05°50‘04“N</geoCoordinate>
,
<geoCoordinate id="EE2F9BDEFFA9FFEA9B9EFE04FBE0F2B8" box="[985,1116,454,476]" degrees="116" direction="east" minutes="20" orientation="longitude" pageId="105" pageNumber="248" precision="15" seconds="08" value="116.335556">116°20‘08“E</geoCoordinate>
], foréthumideà
<taxonomicName id="4C1B869AFFA9FFEA9916FE21FE78F29C" box="[337,452,483,504]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" pageId="105" pageNumber="248" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F210BFFA9FFEA9916FE21FE78F29C" box="[337,452,483,504]" italics="true" pageId="105" pageNumber="248">Lithocarpus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1B869AFFA9FFEA9996FE26FDF9F29D" box="[465,581,484,505]" class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" kingdom="Plantae" order="Fagales" pageId="105" pageNumber="248" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F210BFFA9FFEA9996FE26FDF9F29D" box="[465,581,484,505]" italics="true" pageId="105" pageNumber="248">Castanopsis</emphasis>
</taxonomicName>
etfougèresarborescentes,
<date id="FFA5DBD9FFA9FFEA9B23FE21FC76F29D" box="[868,970,483,505]" pageId="105" pageNumber="248" value="1996-05-18">18.V.1996</date>
, leg. D. Burck- hardt &amp; I. Löbl (30a), tamisagededébrisvegetauxdansunravinhumideetàlalisièrede laforêt (1 f, MHNG);
<collectingRegion id="49DF33FBFFA9FFEA99DCFDDFFE60F157" box="[411,476,541,563]" country="Malaysia" name="Sabah" pageId="105" pageNumber="248">Sabah</collectingRegion>
, BanjaranCrockerMts., GunungAlabpeak,
<date id="FFA5DBD9FFA9FFEA9BFFFDDFFBDEF157" box="[952,1122,541,563]" pageId="105" pageNumber="248" value="1996-04-30" valueMax="1996-05-27" valueMin="1996-04-30">30.IV.-27.V.1996</date>
, 1650–
<quantity id="4CE350FCFFA9FFEA98ADFDF8FE88F12B" box="[234,308,570,592]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="105" pageNumber="248" unit="m" value="1800.0">1800 m</quantity>
[
<geoCoordinate id="EE2F9BDEFFA9FFEA9906FDF8FE25F134" box="[321,409,570,592]" degrees="05" direction="north" minutes="50" orientation="latitude" pageId="105" pageNumber="248" precision="925" value="5.8333335">05°50‘N</geoCoordinate>
,
<geoCoordinate id="EE2F9BDEFFA9FFEA99E4FDFBFDBEF12B" box="[419,514,569,591]" degrees="116" direction="east" minutes="20" orientation="longitude" pageId="105" pageNumber="248" precision="925" value="116.333336">116°20‘E</geoCoordinate>
], leg. M. Štrba &amp; R. Hergovits (
<quantity id="4CE350FCFFA9FFEA9B0CFDF8FCD2F134" box="[843,878,570,592]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="105" pageNumber="248" unit="m" value="1.0">1 m</quantity>
, NHMW).
</paragraph>
<paragraph id="8BA4FD19FFA9FFEA98ADFDB6FC1CF68D" blockId="105.[173,1126,627,1514]" pageId="105" pageNumber="248">
<emphasis id="B96F210BFFA9FFEA98ADFDB6FEE7F1ED" box="[234,347,628,649]" italics="true" pageId="105" pageNumber="248">Description</emphasis>
– Measurements (in mm, n=5): HW = 0.60 (0.58–0.61); TW = 0.57 (0.56–0.59); PW = 0.82 (0.80–0.85); EW = 0.94 (0.92–0.96); AW = 0.92 (0.91–0.93); HL = 0.43 (0.42–0.44); EL = 0.14 (0.13–0.15); TL = 0.18 (0.17–0.19); PL = 0.51 (0.50–0.52); SC = 0.60 (0.59–0.61); FB = 1.61 (1.58–1.63); BL = 3.05 (2.91–3.25).
<figureCitation id="1320E19CFFA9FFEA9A54FD08FD4AF1BB" box="[531,758,714,736]" captionStart="Figs 73–75" captionStartId="40.[136,181,1526,1548]" captionTargetBox="[140,1082,206,1495]" captionTargetId="figure-19@40.[135,1083,197,1501]" captionTargetPageId="40" captionText="Figs 73–75. Habitus of Anotylus species. 73 = A. schawalleri sp. n., 74 = A. borneensis (Cam- eron, 1928), 75 = A. gunung sp. n." figureDoi="http://doi.org/10.5281/zenodo.12625455" httpUri="https://zenodo.org/record/12625455/files/figure.png" pageId="105" pageNumber="248">HabitusasinFig. 73</figureCitation>
. Forebodyweaklyshiningdespite sculpture, abdomenwithpuncturesofvaryingsizes, ratherdensebutshallow, indistinctly microsculptured, yetmorelustrousthanforebody. Headblackishdarkbrown, pronotum, elyraandabdomenreddishdarkbrown, occasionallylighter, mediumbrownstripealong posteriorelytralmargin, lateralpronotalmarginandanteriormarginofhead. Legs, mouthpartsandantennaereddishmediumbrown. Headwithanteriormarginpossessingrather strongrim, almosttruncateinfront, laterallystraighttosupraantennalprominences (moderatelyelevatedobliqueridges), discslightlyimpressedneareyes. Epistomalsutureformingsubcircularimpression (muchlesspunctateinside), uppersurfacewithweaktactilesetae (intratemporalis) situatedpostero-mediallyofeacheye; eyesmediumsizedandslightly bulging, indorsalviewtemples (1.3× lengthofeye) alatiform, laterallyevenlyroundedand posteriorlyconstrictedtoformdistinctneckseparatedbyweakoccipitalfurrow. Anten- nomere 1 flattenedclub-shaped, stronglysculptured, segment 2 basallybroadsubtriangular, segment 3 incrassate, segment 4 transversediscoidwithrim, segments 5–11 withbasal dishes, articles 5–10 transverse (about 1.5–1.7× broaderthanlong). Headandpronotum withmostlyalveolatesculpture. Pronotumwithlateralportionexplanate, edgealatiform withsparseshortsetae; lateralmarginunevenlyarchedwithanumberofsmall, irregularly distributeddenticles, posteriorcornersnotincised, discmediallywithtwoweakparallel longitudinalridges, pronotalsidesweaklyimpressedaroundmiddle. Elytrawithoutmembranousposteriormargin. Elytralepipleuralridgeapparentasstrongersculpturelinesin lateralalatiformportions, discwithveryslighttonoimpressionincentre; shoulderswell developed, surfaceobscurelyfoveolateandinbetweenpuncturestorulose. Legsshort, pro- andmesotibiawithseveralspinuloserows, metatibiawithlongitudinalctenidiumofspi- nulesindistalhalf. Abdomenwithsidesmoderately/ratherarcuate, secondsegmentwith paratergitesbroadeningposteriorly (abdomenappearingconstrictedatbase), thoseofseg- mentsIII–VIIthin, mesalparatergitesbroad, posterioredgeoftergiteVIIwiththinpalisade fringe,
<figureCitation id="1320E19CFFA9FFEA98B0FA75FD25F6A8" box="[247,665,1463,1485]" captionStart="Figs 230–236" captionStartId="106.[136,181,1490,1512]" captionTargetBox="[148,1076,191,1464]" captionTargetId="figure-19@106.[147,1076,189,1464]" captionTargetPageId="106" captionText="Figs 230–236. Anotylus schawalleri sp. n., male. 230 = median part of sternite VII, 231 = ster- nite VIII, 232 = tergite IX, 233 = tergite X, 234 = aedeagus, frontal view, 235 = median lobe, lateral view, 236 = paramere, lateral view. Scales: 0.10 mm for Figs 234–236, 0.115 mm for Fig. 233, 0.15 mm for Figs 230–232" figureDoi="http://doi.org/10.5281/zenodo.12625527" httpUri="https://zenodo.org/record/12625527/files/figure.png" pageId="105" pageNumber="248">medianpartofsterniteVIIasinFig. 230</figureCitation>
, sterniteVIII (
<figureCitation id="1320E19CFFA9FFEA9B6CFA75FCC3F6A9" box="[811,895,1463,1485]" captionStart="Figs 230–236" captionStartId="106.[136,181,1490,1512]" captionTargetBox="[148,1076,191,1464]" captionTargetId="figure-19@106.[147,1076,189,1464]" captionTargetPageId="106" captionText="Figs 230–236. Anotylus schawalleri sp. n., male. 230 = median part of sternite VII, 231 = ster- nite VIII, 232 = tergite IX, 233 = tergite X, 234 = aedeagus, frontal view, 235 = median lobe, lateral view, 236 = paramere, lateral view. Scales: 0.10 mm for Figs 234–236, 0.115 mm for Fig. 233, 0.15 mm for Figs 230–232" figureDoi="http://doi.org/10.5281/zenodo.12625527" httpUri="https://zenodo.org/record/12625527/files/figure.png" pageId="105" pageNumber="248">Fig. 231</figureCitation>
), tergitesIX (
<figureCitation id="1320E19CFFA9FFEA9C40FA75FBE7F6A9" box="[1031,1115,1463,1485]" captionStart="Figs 230–236" captionStartId="106.[136,181,1490,1512]" captionTargetBox="[148,1076,191,1464]" captionTargetId="figure-19@106.[147,1076,189,1464]" captionTargetPageId="106" captionText="Figs 230–236. Anotylus schawalleri sp. n., male. 230 = median part of sternite VII, 231 = ster- nite VIII, 232 = tergite IX, 233 = tergite X, 234 = aedeagus, frontal view, 235 = median lobe, lateral view, 236 = paramere, lateral view. Scales: 0.10 mm for Figs 234–236, 0.115 mm for Fig. 233, 0.15 mm for Figs 230–232" figureDoi="http://doi.org/10.5281/zenodo.12625527" httpUri="https://zenodo.org/record/12625527/files/figure.png" pageId="105" pageNumber="248">Fig. 232</figureCitation>
), X (
<figureCitation id="1320E19CFFA9FFEA988DFA11FEA1F68E" box="[202,285,1491,1514]" captionStart="Figs 230–236" captionStartId="106.[136,181,1490,1512]" captionTargetBox="[148,1076,191,1464]" captionTargetId="figure-19@106.[147,1076,189,1464]" captionTargetPageId="106" captionText="Figs 230–236. Anotylus schawalleri sp. n., male. 230 = median part of sternite VII, 231 = ster- nite VIII, 232 = tergite IX, 233 = tergite X, 234 = aedeagus, frontal view, 235 = median lobe, lateral view, 236 = paramere, lateral view. Scales: 0.10 mm for Figs 234–236, 0.115 mm for Fig. 233, 0.15 mm for Figs 230–232" figureDoi="http://doi.org/10.5281/zenodo.12625527" httpUri="https://zenodo.org/record/12625527/files/figure.png" pageId="105" pageNumber="248">Fig. 233</figureCitation>
)
<figureCitation id="1320E19CFFA9FFEA996EFA16FDC1F68D" box="[297,637,1491,1513]" captionStart="Figs 230–236" captionStartId="106.[136,181,1490,1512]" captionTargetBox="[148,1076,191,1464]" captionTargetId="figure-19@106.[147,1076,189,1464]" captionTargetPageId="106" captionText="Figs 230–236. Anotylus schawalleri sp. n., male. 230 = median part of sternite VII, 231 = ster- nite VIII, 232 = tergite IX, 233 = tergite X, 234 = aedeagus, frontal view, 235 = median lobe, lateral view, 236 = paramere, lateral view. Scales: 0.10 mm for Figs 234–236, 0.115 mm for Fig. 233, 0.15 mm for Figs 230–232" figureDoi="http://doi.org/10.5281/zenodo.12625527" httpUri="https://zenodo.org/record/12625527/files/figure.png" pageId="105" pageNumber="248">andaedeagusasinFigs 234–236</figureCitation>
,
<figureCitation id="1320E19CFFA9FFEA9AC0FA16FC20F68D" box="[647,924,1491,1513]" captionStart="Figs 265–278" captionStartId="116.[136,181,1475,1497]" captionTargetBox="[138,1088,607,1439]" captionTargetId="figure-539@116.[137,1089,605,1454]" captionTargetPageId="116" captionText="Figs 265–278. Spermathecae of Anotylus species: 265 = A. coonoor sp. n., 266 = A. ganapati sp. n., 267 = A. hartmanni sp. n., 268 = A. ijen sp. n., 269 = A. jambi sp. n., 270 = A. kabasi sp. n., 271 = A. linaxi sp. n., 272 = A. riedeli sp. n., 273 = A. rurukan sp. n., 274 = A. schawalleri sp. n., 275 = A. schillhammeri sp. n., 276 = A. schuelkei sp. n., 277 = A. tanator sp. n, 278 = A. topali" figureDoi="http://doi.org/10.5281/zenodo.12625539" httpUri="https://zenodo.org/record/12625539/files/figure.png" pageId="105" pageNumber="248">spermathecaasinFig. 274</figureCitation>
.
</paragraph>
<paragraph id="8BA4FD19FFA9FFEA98ADF9CFFBDAF547" blockId="105.[173,1126,1549,1600]" box="[234,1126,1549,1571]" pageId="105" pageNumber="248">
<emphasis id="B96F210BFFA9FFEA98ADF9CFFEEFF546" box="[234,339,1549,1570]" italics="true" pageId="105" pageNumber="248">Etymology</emphasis>
– Thespeciesisnamedafterthecollectoroftheholotype, WolfgangScha-
</paragraph>
<paragraph id="8BA4FD19FFA9FFEA98EAF9E8FEECF524" blockId="105.[173,1126,1549,1600]" box="[173,336,1578,1600]" pageId="105" pageNumber="248">waller (SMNS).</paragraph>
<caption id="DF64AD91FFAAFFE998CFFA10FCA4F55A" ID-DOI="http://doi.org/10.5281/zenodo.12625527" ID-Zenodo-Dep="12625527" httpUri="https://zenodo.org/record/12625527/files/figure.png" pageId="106" pageNumber="249" startId="106.[136,181,1490,1512]" targetBox="[148,1076,191,1464]" targetPageId="106">
<paragraph id="8BA4FD19FFAAFFE998CFFA10FB83F546" blockId="106.[136,1088,1490,1599]" pageId="106" pageNumber="249">
<emphasis id="B96F210BFFAAFFE998CFFA10FEA9F68C" bold="true" box="[136,277,1490,1512]" pageId="106" pageNumber="249">Figs 230–236.</emphasis>
<taxonomicName id="4C1B869AFFAAFFE9995CFA11FE54F68C" authorityName="Makranczy" authorityYear="2017" box="[283,488,1491,1512]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="249" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov.">
<emphasis id="B96F210BFFAAFFE9995CFA11FE54F68C" box="[283,488,1491,1512]" italics="true" pageId="106" pageNumber="249">Anotylus schawalleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C9C70FFAAFFE999AAFA11FD99F68C" box="[493,549,1491,1512]" pageId="106" pageNumber="249" rank="species">sp. n.</taxonomicNameLabel>
, male. 230 = median part of sternite VII, 231 = ster- nite VIII, 232 = tergite IX, 233 = tergite X, 234 = aedeagus, frontal view, 235 = median lobe, lateral view, 236 = paramere, lateral view. Scales: 0.10 mm for Figs 234–236, 0.115 mm for
</paragraph>
<paragraph id="8BA4FD19FFAAFFE999F7F9EBFCA4F55A" blockId="106.[136,1088,1490,1599]" box="[432,792,1577,1599]" pageId="106" pageNumber="249">Fig. 233, 0.15 mm for Figs 230–232</paragraph>
</caption>
<paragraph id="8BA4FD19FFABFFE898ADFF7CFCB8F39E" blockId="107.[173,1126,190,351]" pageId="107" pageNumber="250">
<emphasis id="B96F210BFFABFFE898ADFF7CFEC4F3BC" box="[234,376,190,216]" italics="true" pageId="107" pageNumber="250">Distribution</emphasis>
– Thespeciesappearstobeendemictothehighestareasof theCrockerRangeinSabah (MalaysianBorneo).
</paragraph>
<paragraph id="8BA4FD19FFABFFE898ADFEC3FC87F23B" blockId="107.[173,1126,190,351]" pageId="107" pageNumber="250">
<emphasis id="B96F210BFFABFFE898ADFEC3FEF0F27F" box="[234,332,257,283]" italics="true" pageId="107" pageNumber="250">Remarks</emphasis>
– Thelownumberofspecimensexamineddoesnotallowagood judgementofintraspecificvariability, butexternallythespeciescanbedifficult to separate from the closely related
<taxonomicName id="4C1B869AFFABFFE89ADCFE87FC89F23A" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[667,821,324,351]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="107" pageNumber="250" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="B96F210BFFABFFE89ADCFE87FC89F23A" box="[667,821,324,351]" italics="true" pageId="107" pageNumber="250">A. borneensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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