Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae) Author Makranczy, György text Acta Zoologica Academiae Scientiarum Hungaricae 2017 2017-06-19 63 2 143 262 http://dx.doi.org/10.17109/azh.63.2.143.2017 journal article 10.17109/AZH.63.2.143.2017 2064-2474 12625425 671ADB82-5AA9-418A-BB5C-DB3B912D3BD2 Anotylus schawalleri sp. n. ( Figs 73 , 230–236 , 274 ) Typematerial – Holotype (m): MALAYSIA /BORNEO: Sabah , CrockerRangeN. P., NWKeningau, 900–1200 m [ 05°26‘N , 116°05‘E ], 16-20.XI.1996 , leg. W. Schawaller (SMNS); Paratypes (4): samedataasholotype ( 1 m , HNHM); Sabah , CrockerRange, 1550–1650 m , àproximitéducol (routeKotaKinabalu-Tambunan) [ 05°47‘55“N , 116°20‘06“E ], forêt de Lithocarpus - Castanopsis , 16.V.1996 , leg. D. Burckhardt & I. Löbl (27a), tamisagedebois pourri, feuillesmortesetmousses ( 1 m , MHNG); Sabah , CrockerRange, faceouest (W), 1600 m env., km 51-52 - route de Kota Kinabalu - Tambunan [ 05°50‘04“N , 116°20‘08“E ], foréthumideà Lithocarpus , Castanopsis etfougèresarborescentes, 18.V.1996 , leg. D. Burck- hardt & I. Löbl (30a), tamisagededébrisvegetauxdansunravinhumideetàlalisièrede laforêt (1 f, MHNG); Sabah , BanjaranCrockerMts., GunungAlabpeak, 30.IV.-27.V.1996 , 1650– 1800 m [ 05°50‘N , 116°20‘E ], leg. M. Štrba & R. Hergovits ( 1 m , NHMW). Description – Measurements (in mm, n=5): HW = 0.60 (0.58–0.61); TW = 0.57 (0.56–0.59); PW = 0.82 (0.80–0.85); EW = 0.94 (0.92–0.96); AW = 0.92 (0.91–0.93); HL = 0.43 (0.42–0.44); EL = 0.14 (0.13–0.15); TL = 0.18 (0.17–0.19); PL = 0.51 (0.50–0.52); SC = 0.60 (0.59–0.61); FB = 1.61 (1.58–1.63); BL = 3.05 (2.91–3.25). HabitusasinFig. 73 . Forebodyweaklyshiningdespite sculpture, abdomenwithpuncturesofvaryingsizes, ratherdensebutshallow, indistinctly microsculptured, yetmorelustrousthanforebody. Headblackishdarkbrown, pronotum, elyraandabdomenreddishdarkbrown, occasionallylighter, mediumbrownstripealong posteriorelytralmargin, lateralpronotalmarginandanteriormarginofhead. Legs, mouthpartsandantennaereddishmediumbrown. Headwithanteriormarginpossessingrather strongrim, almosttruncateinfront, laterallystraighttosupraantennalprominences (moderatelyelevatedobliqueridges), discslightlyimpressedneareyes. Epistomalsutureformingsubcircularimpression (muchlesspunctateinside), uppersurfacewithweaktactilesetae (intratemporalis) situatedpostero-mediallyofeacheye; eyesmediumsizedandslightly bulging, indorsalviewtemples (1.3× lengthofeye) alatiform, laterallyevenlyroundedand posteriorlyconstrictedtoformdistinctneckseparatedbyweakoccipitalfurrow. Anten- nomere 1 flattenedclub-shaped, stronglysculptured, segment 2 basallybroadsubtriangular, segment 3 incrassate, segment 4 transversediscoidwithrim, segments 5–11 withbasal dishes, articles 5–10 transverse (about 1.5–1.7× broaderthanlong). Headandpronotum withmostlyalveolatesculpture. Pronotumwithlateralportionexplanate, edgealatiform withsparseshortsetae; lateralmarginunevenlyarchedwithanumberofsmall, irregularly distributeddenticles, posteriorcornersnotincised, discmediallywithtwoweakparallel longitudinalridges, pronotalsidesweaklyimpressedaroundmiddle. Elytrawithoutmembranousposteriormargin. Elytralepipleuralridgeapparentasstrongersculpturelinesin lateralalatiformportions, discwithveryslighttonoimpressionincentre; shoulderswell developed, surfaceobscurelyfoveolateandinbetweenpuncturestorulose. Legsshort, pro- andmesotibiawithseveralspinuloserows, metatibiawithlongitudinalctenidiumofspi- nulesindistalhalf. Abdomenwithsidesmoderately/ratherarcuate, secondsegmentwith paratergitesbroadeningposteriorly (abdomenappearingconstrictedatbase), thoseofseg- mentsIII–VIIthin, mesalparatergitesbroad, posterioredgeoftergiteVIIwiththinpalisade fringe, medianpartofsterniteVIIasinFig. 230 , sterniteVIII ( Fig. 231 ), tergitesIX ( Fig. 232 ), X ( Fig. 233 ) andaedeagusasinFigs 234–236 , spermathecaasinFig. 274 . Etymology – Thespeciesisnamedafterthecollectoroftheholotype, WolfgangScha- waller (SMNS). Figs 230–236. Anotylus schawalleri sp. n. , male. 230 = median part of sternite VII, 231 = ster- nite VIII, 232 = tergite IX, 233 = tergite X, 234 = aedeagus, frontal view, 235 = median lobe, lateral view, 236 = paramere, lateral view. Scales: 0.10 mm for Figs 234–236, 0.115 mm for Fig. 233, 0.15 mm for Figs 230–232 Distribution – Thespeciesappearstobeendemictothehighestareasof theCrockerRangeinSabah (MalaysianBorneo). Remarks – Thelownumberofspecimensexamineddoesnotallowagood judgementofintraspecificvariability, butexternallythespeciescanbedifficult to separate from the closely related A. borneensis .