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<document id="6AEBB50C7781992F9DB6B250C045C2F1" ID-DOI="10.1080/00222933.2024.2387798" ID-ISSN="1464-5262" ID-Zenodo-Dep="13758556" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726224281730" checkinUser="plazi" docAuthor="Volcan, Matheus Vieira, Severo-Neto, Francisco &amp; Lanés, Luis Esteban Krause" docDate="2024" docId="03AB755FFFDDFFE7FE886CC844EB2A55" docLanguage="en" docName="JNATHIST.58.37-40.1493-1508.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 58 (37 - 40)" docSource="http://dx.doi.org/10.1080/00222933.2024.2387798" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Melanorivulus terena Volcan, Severo-Neto &amp; Lanes 2024, sp. n." docType="treatment" docUuid="9C25B380-D5DA-402B-8E98-C0296777AE0B" docUuidSource="ZooBank" docVersion="2" lastPageNumber="1501" masterDocId="FF920D27FFDFFFEFFFBF6E034633296F" masterDocTitle="A new species of Melanorivulus (Cyprinodontiformes: Rivulidae) from headwaters of the Taquari River basin, Paraguay River basin, central Brazilian Cerrado" masterLastPageNumber="1508" masterPageNumber="1493" pageNumber="1495" updateTime="1726225067375" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="338B00CEC2169BBA3069D20F81723354" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="45C92FDA8956385AE22EF747C63B1035">
<mods:title id="AE8DCB3435429D18A48888A359D4CA0C">A new species of Melanorivulus (Cyprinodontiformes: Rivulidae) from headwaters of the Taquari River basin, Paraguay River basin, central Brazilian Cerrado</mods:title>
</mods:titleInfo>
<mods:name id="4C4DADDC30118847E9BC21E8A90A463E" type="personal">
<mods:role id="67227D1C63034A24EE06C178724C5D3C">
<mods:roleTerm id="3D35C26E92951DB29BB3FC12D3AB3FE7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="93E6794ED3085073861E9C42544FD3F0">Volcan, Matheus Vieira</mods:namePart>
<mods:affiliation id="4F441B95C16342D93CB48F2AEE408D49">Laboratório de Ictiologia, Instituto Pró-Pampa (IPPampa), Pelotas, Brazil;</mods:affiliation>
</mods:name>
<mods:name id="EAAE3713C857E59B649DC4A32C6E48D8" type="personal">
<mods:role id="6EBA5F9F7BACFA4E9C9DB4E7286632F5">
<mods:roleTerm id="29C0E0D1003107D82C429220921FD276">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0197EB048945A5B5E492C8B4F3B8E74A">Severo-Neto, Francisco</mods:namePart>
<mods:affiliation id="AE6B1773306C7B1965A8551C1237E9E5">Instituto de Biociências, Coleção Zoológica da Universidade Federal de Mato Grosso do Sul, Campo Grande, Brazil</mods:affiliation>
</mods:name>
<mods:name id="8FBD0784CE9F11613213A28BA3809EFB" type="personal">
<mods:role id="15EC938A0A39637295CE19D8911E6152">
<mods:roleTerm id="EBDF75997FAAE5DC6C304453A6A679AE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="25A89D3CC6679DF53D2D6A7BD9B01B0F">Lanés, Luis Esteban Krause</mods:namePart>
<mods:affiliation id="BA43611B16AEC045FFC3E2CC50CB5FCB">Laboratório de Ictiologia, Instituto Pró-Pampa (IPPampa), Pelotas, Brazil;</mods:affiliation>
</mods:name>
<mods:typeOfResource id="77BB93ED801738E30C65D913EDD6F24B">text</mods:typeOfResource>
<mods:relatedItem id="72611A37F41872A56FCB32F367A0E25F" type="host">
<mods:titleInfo id="A8DDFEBAE2E0AC50DB7B5AA2EDD38676">
<mods:title id="38492B9FCC3D2D17E6D724F1E9682E35">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="0496DEA7F16DB42030AFBC32AB45DA3C">
<mods:date id="2AE9A705480D90DE367E7590C1B42304">2024</mods:date>
<mods:detail id="ECC83B9FE98BEC7063E2BAE43CD7F730" type="series">
<mods:title id="0420C0D87DC0EE5CF1F6BF2A6150DFC8">J. Nat. Hist.</mods:title>
</mods:detail>
<mods:detail id="7995252F8AAD9159AB94880927AC0716" type="pubDate">
<mods:number id="27A51442266F2EED90B7F3296E675540">2024-08-29</mods:number>
</mods:detail>
<mods:detail id="0E87A00D43E40324A071C23EF37A93CC" type="volume">
<mods:number id="7DC7286FB5A72BA4C636650201DAAF8F">58</mods:number>
</mods:detail>
<mods:detail id="E21F39580E47EFD604762C8B3E59DEE2" type="issue">
<mods:number id="B6FC1016B7EF71F27BB4CFB526C7A7FC">37 - 40</mods:number>
</mods:detail>
<mods:extent id="B4AE38ACF8D84327DAAAAC87A54A81CE" unit="page">
<mods:start id="D5453C401D42B729191DFED4914A0FDC">1493</mods:start>
<mods:end id="3381A9E5AAD13DDC1C69587E166DC7C5">1508</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="1869DB62547A7BEB09FF040FC170259F">
<mods:url id="C567B7BD616F558199439EA061EF723A">http://dx.doi.org/10.1080/00222933.2024.2387798</mods:url>
</mods:location>
<mods:classification id="AA015DA95CB2A074AB95223C07598500">journal article</mods:classification>
<mods:identifier id="BD8710192E2FBD8ECE70CE2D1EA9D900" type="DOI">10.1080/00222933.2024.2387798</mods:identifier>
<mods:identifier id="A657F797319DEAD310E3D6ED1D99C357" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="31894B2A7A78BAE9C2D98BAA37769710" type="Zenodo-Dep">13758556</mods:identifier>
<mods:identifier id="5F1F3BDE37CEBFFC49EAAB864EBF0915" type="ZooBank">8B0027D9-8346-4ADB-AE9E-8C770823B8D9</mods:identifier>
</mods:mods>
<treatment id="03AB755FFFDDFFE7FE886CC844EB2A55" LSID="urn:lsid:zoobank.org:act:9C25B380-D5DA-402B-8E98-C0296777AE0B" httpUri="http://treatment.plazi.org/id/03AB755FFFDDFFE7FE886CC844EB2A55" lastPageId="8" lastPageNumber="1501" pageId="2" pageNumber="1495">
<subSubSection id="C31897C2FFDDFFEDFE886CC8458A2A36" pageId="2" pageNumber="1495" type="nomenclature">
<paragraph id="8BBDC449FFDDFFEDFE886CC845DF2B89" blockId="2.[311,1004,715,742]" box="[311,1004,715,742]" pageId="2" pageNumber="1495">
<heading id="D0F57325FFDDFFEDFE886CC845DF2B89" box="[311,1004,715,742]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1495" reason="5">
<taxonomicName id="4C02BFCAFFDDFFEDFE886CC845AE2B89" authority="Volcan, Severo-Neto &amp; Lanes" authorityName="Volcan, Severo-Neto &amp; Lanes" authorityYear="2024" box="[311,925,715,742]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="1495" phylum="Chordata" rank="species" species="terena" status="sp. nov.">
<emphasis id="B976185BFFDDFFEDFE886CC844012B89" box="[311,562,715,742]" italics="true" pageId="2" pageNumber="1495">Melanorivulus terena</emphasis>
Volcan, Severo-Neto &amp; Lanés
</taxonomicName>
<emphasis id="B976185BFFDDFFEDFC1A6CC845DF2B89" bold="true" box="[933,1004,715,742]" pageId="2" pageNumber="1495">
<taxonomicNameLabel id="A245A520FFDDFFEDFC1A6CC845DF2B89" box="[933,1004,715,742]" pageId="2" pageNumber="1495" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBDC449FFDDFFEDFF7F6D1D458A2A36" blockId="2.[192,966,798,857]" pageId="2" pageNumber="1495">
<uri id="FF93C84BFFDDFFEDFF7F6D1D458A2A36" pageId="2" pageNumber="1495">
urn:lsid:zoobank.org:pub:
<uuid id="FFA4FE9CFFDDFFEDFE766D1D45F52A58" box="[457,966,798,823]" pageId="2" pageNumber="1495">8B0027D9-8346-4ADB-AE9E-8C770823B8D9</uuid>
urn:lsid:zoobank.org:act:
<uuid id="FFA4FE9CFFDDFFEDFE7E6D43458A2A36" box="[449,953,832,857]" pageId="2" pageNumber="1495">9C25B380-D5DA-402B-8E98-C0296777AE0B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C31897C2FFDDFFEDFF1F6D84426D2DB4" pageId="2" pageNumber="1495" type="materials_examined">
<paragraph id="8BBDC449FFDDFFEDFF1F6D84422C2D66" blockId="2.[160,1156,903,1033]" pageId="2" pageNumber="1495">
<materialsCitation id="3B6ACE14FFDDFFEDFF1F6D84422C2D66" collectingDate="2021-04-04" collectionCode="T, R" collectorName="F. Severo-Neto &amp; F. Sinani" country="Brazil" county="Camapua" elevation="518" latitude="-19.34333" location="Rio Coxim" longLatPrecision="1" longitude="-53.89277" municipality="Ribeirao Cachoeirinha" pageId="2" pageNumber="1495" specimenCode="ZUFMS-PIS08450" specimenCount="1" stateProvince="Mato Grosso do Sul" typeStatus="holotype">
<emphasis id="B976185BFFDDFFEDFF1F6D8447A72ACE" bold="true" box="[160,404,903,929]" pageId="2" pageNumber="1495">
<emphasis id="B976185BFFDDFFEDFF1F6D8447382ACE" bold="true" box="[160,267,903,929]" italics="true" pageId="2" pageNumber="1495">
<typeStatus id="54B97AEBFFDDFFEDFF1F6D8447382ACE" box="[160,267,903,929]" pageId="2" pageNumber="1495" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="1339D8CCFFDDFFEDFEA16D8447B62ACE" box="[286,389,903,929]" captionStart="Figure 1" captionStartId="2.[160,225,1704,1727]" captionTargetBox="[162,1152,1301,1673]" captionTargetId="figure-344@2.[162,1152,1301,1673]" captionTargetPageId="2" captionText="Figure 1. Melanorivulus terena, ZUFMS-PIS08450, male, holotype, 27.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13758558" httpUri="https://zenodo.org/record/13758558/files/figure.png" pageId="2" pageNumber="1495">Figure 1</figureCitation>
).
</emphasis>
<specimenCode id="DBA46C32FFDDFFEDFE1B6D8B44562ACE" box="[420,613,904,929]" collectionCode="ZUFMS-PIS" pageId="2" pageNumber="1495">ZUFMS-PIS08450</specimenCode>
, male,
<quantity id="4CFA69ACFFDDFFEDFD066D8B452D2ACE" box="[697,798,904,929]" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="2" pageNumber="1495" unit="mm" value="27.2">27.2 mm</quantity>
SL,
<collectingCountry id="F31584D9FFDDFFEDFCED6D8B45A22ACE" box="[850,913,904,929]" name="Brazil" pageId="2" pageNumber="1495">Brazil</collectingCountry>
,
<collectingRegion id="49C60AABFFDDFFEDFC216D8B46EF2AAB" country="Brazil" name="Mato Grosso do Sul" pageId="2" pageNumber="1495">Mato Grosso do Sul State</collectingRegion>
, municipality of
<collectingCounty id="62DCBCC5FFDDFFEDFE2C6DA844332AAB" box="[403,512,939,964]" pageId="2" pageNumber="1495">Camapuã</collectingCounty>
, road MS-422,
<collectingMunicipality id="6BD95E33FFDDFFEDFD1B6DA845A82AAB" box="[676,923,939,964]" pageId="2" pageNumber="1495">Ribeirão Cachoeirinha</collectingMunicipality>
, first-order stream in the headwaters of
<location id="8EDD9292FFDDFFEDFE3E6DCE47C92A89" LSID="urn:lsid:plazi:treatment:03AB755FFFDDFFE7FE886CC844EB2A55:8EDD9292FFDDFFEDFE3E6DCE47C92A89" box="[385,506,973,998]" country="Brazil" county="Camapua" latitude="-19.34333" longLatPrecision="1" longitude="-53.89277" municipality="Ribeirao Cachoeirinha" name="Rio Coxim" pageId="2" pageNumber="1495" stateProvince="Mato Grosso do Sul">Rio Coxim</location>
,
<location id="8EDD9292FFDDFFEDFDB76DCE44B82A89" LSID="urn:lsid:plazi:treatment:03AB755FFFDDFFE7FE886CC844EB2A55:8EDD9292FFDDFFEDFDB76DCE44B82A89" box="[520,651,973,998]" country="Brazil" county="Camapua" latitude="-19.34333" longLatPrecision="1" longitude="-53.89277" municipality="Ribeirao Cachoeirinha" name="Rio Taquari" pageId="2" pageNumber="1495" stateProvince="Mato Grosso do Sul">Rio Taquari</location>
drainage,
<location id="8EDD9292FFDDFFEDFCB46DCE451D2A89" LSID="urn:lsid:plazi:treatment:03AB755FFFDDFFE7FE886CC844EB2A55:8EDD9292FFDDFFEDFCB46DCE451D2A89" box="[779,814,973,998]" country="Brazil" county="Camapua" latitude="-19.34333" longLatPrecision="1" longitude="-53.89277" municipality="Ribeirao Cachoeirinha" name="Rio" pageId="2" pageNumber="1495" stateProvince="Mato Grosso do Sul">Rio</location>
<collectingCountry id="F31584D9FFDDFFEDFC866DCE45922A89" box="[825,929,973,998]" name="Paraguay" pageId="2" pageNumber="1495">Paraguay</collectingCountry>
basin, -
<geoCoordinate id="EE36A28EFFDDFFEDFBBF6DCE42B32A89" box="[1024,1152,973,998]" degrees="19.34333" direction="south" orientation="latitude" pageId="2" pageNumber="1495" precision="1" value="-19.34333">19.34333°S</geoCoordinate>
, -
<geoCoordinate id="EE36A28EFFDDFFEDFF196DF347032D66" box="[166,304,1008,1033]" degrees="53.89277" direction="west" orientation="longitude" pageId="2" pageNumber="1495" precision="1" value="-53.89277">53.89277°W</geoCoordinate>
, altitude
<elevation id="002F237AFFDDFFEDFE286DF344262D66" box="[407,533,1008,1033]" metricMagnitude="2" metricUnit="m" metricValue="5.18" pageId="2" pageNumber="1495" unit="m" value="518.0">
<quantity id="4CFA69ACFFDDFFEDFE286DF347ED2D66" box="[407,478,1008,1033]" metricMagnitude="2" metricUnit="m" metricValue="5.18" pageId="2" pageNumber="1495" unit="m" value="518.0">518 m</quantity>
a.s.l.
</elevation>
;
<collectorName id="26F7A19FFFDDFFEDFD9E6DF344F52D66" box="[545,710,1008,1033]" pageId="2" pageNumber="1495">F. Severo-Neto</collectorName>
and
<collectionCode id="ED135C8CFFDDFFEDFD406DF345232D66" box="[767,784,1008,1033]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="1495">T</collectionCode>
.
<collectionCode id="ED135C8CFFDDFFEDFCAC6DF345172D66" box="[787,804,1008,1033]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="1495">R</collectionCode>
.
<collectorName id="26F7A19FFFDDFFEDFC986DF345B52D66" box="[807,902,1008,1033]" pageId="2" pageNumber="1495">F. Sinani</collectorName>
,
<date id="FFBCE289FFDDFFEDFC2E6DF342282D66" box="[913,1051,1008,1033]" pageId="2" pageNumber="1495" value="2021-04-04">
<collectingDate id="EFF81B61FFDDFFEDFC2E6DF342282D66" box="[913,1051,1008,1033]" pageId="2" pageNumber="1495" value="2021-04-04">4 April 2021</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBDC449FFDDFFEDFF1F6A34426D2DB4" blockId="2.[160,1156,1079,1243]" pageId="2" pageNumber="1495">
<materialsCitation id="3B6ACE14FFDDFFEDFF1F6A3447092D1C" pageId="2" pageNumber="1495" specimenCode="ZUFMS-PIS08471" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B976185BFFDDFFEDFF1F6A34472F2D3E" bold="true" box="[160,284,1079,1105]" italics="true" pageId="2" pageNumber="1495">
<typeStatus id="54B97AEBFFDDFFEDFF1F6A3447242D3E" box="[160,279,1079,1105]" pageId="2" pageNumber="1495" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="DBA46C32FFDDFFEDFE936A3447DE2D3F" box="[300,493,1079,1104]" collectionCode="ZUFMS-PIS" pageId="2" pageNumber="1495">ZUFMS-PIS08471</specimenCode>
,
<specimenCount id="9D040FC0FFDDFFEDFE4A6A34447E2D3F" box="[501,589,1079,1104]" count="2" pageId="2" pageNumber="1495" type="male">2 males</specimenCount>
,
<quantity id="4CFA69ACFFDDFFEDFDEA6A3444C22D3F" box="[597,753,1079,1104]" metricMagnitude="-2" metricUnit="m" metricValue="2.71" metricValueMax="2.81" metricValueMin="2.61" pageId="2" pageNumber="1495" unit="mm" value="27.1" valueMax="28.1" valueMin="26.1">26.1–28.1 mm</quantity>
SL (2 C&amp;S),
<specimenCount id="9D040FC0FFDDFFEDFCCE6A3445ED2D3F" box="[881,990,1079,1104]" count="2" pageId="2" pageNumber="1495" type="female">2 females</specimenCount>
,
<quantity id="4CFA69ACFFDDFFEDFC586A3442B02D3F" box="[999,1155,1079,1104]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.61" metricValueMin="2.09" pageId="2" pageNumber="1495" unit="mm" value="23.5" valueMax="26.1" valueMin="20.9">20.9–26.1 mm</quantity>
SL (2 C&amp;S); M
</materialsCitation>
.
<collectionCode id="ED135C8CFFDDFFEDFE826A5947632D1C" box="[317,336,1114,1139]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="1495" type="Museum">V</collectionCode>
. Volcan and L.E.K.
<materialsCitation id="3B6ACE14FFDDFFEDFD986A5942B32D1C" box="[551,1152,1114,1139]" collectingDate="2016-12-12" collectionCode="T, R" collectorName="F. Severo-Neto &amp; F. Sinani" country="Brazil" county="Camapua" elevation="518" latitude="-19.34333" location="Rio Coxim" longLatPrecision="1" longitude="-53.89277" municipality="Ribeirao Cachoeirinha" pageId="2" pageNumber="1495" specimenCount="1" stateProvince="Mato Grosso do Sul" typeStatus="paratype">
Lanés,
<date id="FFBCE289FFDDFFEDFDCA6A5945632D1C" box="[629,848,1114,1139]" pageId="2" pageNumber="1495" value="2016-12-12">
<collectingDate id="EFF81B61FFDDFFEDFDCA6A5945632D1C" box="[629,848,1114,1139]" pageId="2" pageNumber="1495" value="2016-12-12">12 December 2016</collectingDate>
</date>
, same locality as holotype
</materialsCitation>
.
<materialsCitation id="3B6ACE14FFDDFFEDFF1F6A7E45A42DD7" collectionCode="MZUSP" country="Brazil" pageId="2" pageNumber="1495" specimenCode="MZUSP 130257" specimenCount="27" specimenCount-female="5" specimenCount-juvenile="19" specimenCount-male="3" stateProvince="Mato Grosso do Sul" typeStatus="paratype">
<specimenCode id="DBA46C32FFDDFFEDFF1F6A7E477E2DF9" box="[160,333,1149,1174]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="1495">MZUSP 130257</specimenCode>
,
<specimenCount id="9D040FC0FFDDFFEDFEE86A7E47832DF9" box="[343,432,1149,1174]" count="3" pageId="2" pageNumber="1495" type="male">3 males</specimenCount>
,
<quantity id="4CFA69ACFFDDFFEDFE046A7E446A2DF9" box="[443,601,1149,1174]" metricMagnitude="-2" metricUnit="m" metricValue="1.945" metricValueMax="1.98" metricValueMin="1.91" pageId="2" pageNumber="1495" unit="mm" value="19.45" valueMax="19.8" valueMin="19.1">19.1–19.8 mm</quantity>
SL,
<specimenCount id="9D040FC0FFDDFFEDFD386A7E44C52DF9" box="[647,758,1149,1174]" count="5" pageId="2" pageNumber="1495" type="female">5 females</specimenCount>
,
<quantity id="4CFA69ACFFDDFFEDFCBF6A7E45AD2DF9" box="[768,926,1149,1174]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" metricValueMax="2.08" metricValueMin="1.78" pageId="2" pageNumber="1495" unit="mm" value="19.3" valueMax="20.8" valueMin="17.8">17.8–20.8 mm</quantity>
SL,
<specimenCount id="9D040FC0FFDDFFEDFC726A7E42672DF9" box="[973,1108,1149,1174]" count="19" pageId="2" pageNumber="1495" type="juvenile">19 juveniles</specimenCount>
, sex undetermined,
<quantity id="4CFA69ACFFDDFFEDFEED6A9C47C52DD7" box="[338,502,1183,1208]" metricMagnitude="-2" metricUnit="m" metricValue="1.4949999999999999" metricValueMax="1.8699999999999999" metricValueMin="1.1199999999999999" pageId="2" pageNumber="1495" unit="mm" value="14.95" valueMax="18.7" valueMin="11.2">11.2–18.7 mm</quantity>
SL, all collected with the
<typeStatus id="54B97AEBFFDDFFEDFC8F6A9C45A42DD7" box="[816,919,1183,1208]" pageId="2" pageNumber="1495" type="holotype">holotype</typeStatus>
</materialsCitation>
.
<materialsCitation id="3B6ACE14FFDDFFEDFC196A9C42682DB4" country="Brazil" pageId="2" pageNumber="1495" specimenCode="ZUFMS-PIS06575" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Mato Grosso do Sul" typeStatus="paratype">
<specimenCode id="DBA46C32FFDDFFEDFC196A9C42542DD7" box="[934,1127,1183,1208]" collectionCode="ZUFMS-PIS" pageId="2" pageNumber="1495">ZUFMS-PIS06575</specimenCode>
,
<specimenCount id="9D040FC0FFDDFFEDFBCA6A9C46D72DB4" count="5" pageId="2" pageNumber="1495" type="male">5 males</specimenCount>
,
<quantity id="4CFA69ACFFDDFFEDFF516AC147BF2DB4" box="[238,396,1218,1243]" metricMagnitude="-2" metricUnit="m" metricValue="2.3600000000000003" metricValueMax="2.6100000000000003" metricValueMin="2.1100000000000003" pageId="2" pageNumber="1495" unit="mm" value="23.6" valueMax="26.1" valueMin="21.1">21.1–26.1 mm</quantity>
SL,
<specimenCount id="9D040FC0FFDDFFEDFE046AC1441A2DB4" box="[443,553,1218,1243]" count="4" pageId="2" pageNumber="1495" type="female">4 females</specimenCount>
,
<quantity id="4CFA69ACFFDDFFEDFD8C6AC144E22DB4" box="[563,721,1218,1243]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.61" metricValueMin="2.09" pageId="2" pageNumber="1495" unit="mm" value="23.5" valueMax="26.1" valueMin="20.9">20.9–26.1 mm</quantity>
SL, all collected with the
<typeStatus id="54B97AEBFFDDFFEDFC4B6AC142682DB4" box="[1012,1115,1218,1243]" pageId="2" pageNumber="1495" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF7D94C1FFDDFFEDFF1F68AB45952FD0" ID-DOI="http://doi.org/10.5281/zenodo.13758558" ID-Zenodo-Dep="13758558" box="[160,934,1704,1727]" httpUri="https://zenodo.org/record/13758558/files/figure.png" pageId="2" pageNumber="1495" startId="2.[160,225,1704,1727]" targetBox="[162,1152,1301,1673]" targetPageId="2" targetType="figure">
<paragraph id="8BBDC449FFDDFFEDFF1F68AB45952FD0" blockId="2.[160,934,1704,1727]" box="[160,934,1704,1727]" pageId="2" pageNumber="1495">
<emphasis id="B976185BFFDDFFEDFF1F68AB46C82FD0" bold="true" box="[160,251,1704,1727]" pageId="2" pageNumber="1495">Figure 1.</emphasis>
<taxonomicName id="4C02BFCAFFDDFFEDFEBD68AB44742FD0" authority=", ZUFMS-PIS" authorityName="ZUFMS-PIS" box="[258,583,1704,1727]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="1495" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFDDFFEDFEBD68AB47F82FD0" box="[258,459,1704,1727]" italics="true" pageId="2" pageNumber="1495">Melanorivulus terena</emphasis>
, ZUFMS-PIS
</taxonomicName>
08450, male, holotype, 27.2 mm SL.
</paragraph>
</caption>
<subSubSection id="C31897C2FFDCFFECFF1F6E8D42482A77" pageId="3" pageNumber="1496" type="diagnosis">
<paragraph id="8BBDC449FFDCFFECFF1F6E8D42482A77" blockId="3.[160,1156,142,792]" pageId="3" pageNumber="1496">
<emphasis id="B976185BFFDCFFECFF1F6E8D472929C7" bold="true" box="[160,282,142,168]" italics="true" pageId="3" pageNumber="1496">Diagnosis.</emphasis>
<taxonomicName id="4C02BFCAFFDCFFECFE956E8C442329C7" box="[298,528,143,168]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFDCFFECFE956E8C442329C7" box="[298,528,143,168]" italics="true" pageId="3" pageNumber="1496">Melanorivulus terena</emphasis>
</taxonomicName>
is distinguished from all species of the
<taxonomicName id="4C02BFCAFFDCFFECFC6A6E8C420429C7" baseAuthorityName="Costa" baseAuthorityYear="1989" box="[981,1079,143,168]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B976185BFFDCFFECFC6A6E8C420429C7" box="[981,1079,143,168]" italics="true" pageId="3" pageNumber="1496">M. pictus</emphasis>
</taxonomicName>
group by the distal margin of the unpaired fins and pelvic-fin orangish-red in males (vs never with the combination of distal margin orangish-red in all unpaired fins and pelvic fin).
<taxonomicName id="4C02BFCAFFDCFFECFF1F6EF447B0287F" box="[160,387,247,272]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFDCFFECFF1F6EF447B0287F" box="[160,387,247,272]" italics="true" pageId="3" pageNumber="1496">Melanorivulus terena</emphasis>
</taxonomicName>
is similar to
<emphasis id="B976185BFFDCFFECFDB36EF444D0287F" box="[524,739,247,272]" italics="true" pageId="3" pageNumber="1496">
<taxonomicName id="4C02BFCAFFDCFFECFDB36EF4445F287F" baseAuthorityName="Costa" baseAuthorityYear="2005" box="[524,620,247,272]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="egens">M. egens</taxonomicName>
,
<taxonomicName id="4C02BFCAFFDCFFECFDC96EF444D0287F" authorityName="Costa" authorityYear="2018" box="[630,739,247,272]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="linearis">M. linearis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C02BFCAFFDCFFECFCA56EF44583287F" authorityName="Volcan, Severo-Neto, and Lanes" authorityYear="2018" box="[794,944,247,272]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="interruptus">
<emphasis id="B976185BFFDCFFECFCA56EF44583287F" box="[794,944,247,272]" italics="true" pageId="3" pageNumber="1496">M. interruptus</emphasis>
</taxonomicName>
, and distinguished from all other species of the
<taxonomicName id="4C02BFCAFFDCFFECFE5B6F194475285C" baseAuthorityName="Costa" baseAuthorityYear="1989" box="[484,582,282,307]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B976185BFFDCFFECFE5B6F194475285C" box="[484,582,282,307]" italics="true" pageId="3" pageNumber="1496">M. pictus</emphasis>
</taxonomicName>
group by the absence of distinctive dark marks on the humeral region (vs presence).
<taxonomicName id="4C02BFCAFFDCFFECFDD46F3F4550283A" box="[619,867,316,341]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFDCFFECFDD46F3F4550283A" box="[619,867,316,341]" italics="true" pageId="3" pageNumber="1496">Melanorivulus terena</emphasis>
</taxonomicName>
is distinguished from
<emphasis id="B976185BFFDCFFECFF1F6F5C47842817" box="[160,439,351,376]" italics="true" pageId="3" pageNumber="1496">
<taxonomicName id="4C02BFCAFFDCFFECFF1F6F5C47722817" authorityName="Volcan, Severo-Neto, and Lanes" authorityYear="2018" box="[160,321,351,376]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="interruptus">M. interruptus</taxonomicName>
,
<taxonomicName id="4C02BFCAFFDCFFECFEED6F5C47842817" baseAuthorityName="Costa" baseAuthorityYear="2005" box="[338,439,351,376]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="egens">M. egens</taxonomicName>
</emphasis>
and
<taxonomicName id="4C02BFCAFFDCFFECFE446F5C44432817" authorityName="Costa" authorityYear="2018" box="[507,624,351,376]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="linearis">
<emphasis id="B976185BFFDCFFECFE446F5C44432817" box="[507,624,351,376]" italics="true" pageId="3" pageNumber="1496">M. linearis</emphasis>
</taxonomicName>
by having a larger number of scales in the transversal series (10–11 vs 8–9). It is distinguished from
<taxonomicName id="4C02BFCAFFDCFFECFC966F8145F128F4" authorityName="Volcan, Severo-Neto, and Lanes" authorityYear="2018" box="[809,962,386,411]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="interruptus">
<emphasis id="B976185BFFDCFFECFC966F8145F128F4" box="[809,962,386,411]" italics="true" pageId="3" pageNumber="1496">M. interruptus</emphasis>
</taxonomicName>
by having fewer pectoral fin rays (13 vs 14), smaller pelvic fin (pelvic-fin length 10.3–11.6% SL, vs 12.1– 13.9% SL), larger head in males (head length 27.8–30.5% SL vs 26.2–27.5% SL), and larger caudal fin in females (caudal-fin length 30.6–32.1% SL vs 26.8–29.4% SL).
<taxonomicName id="4C02BFCAFFDCFFECFC546FE946D62B4A" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFDCFFECFC546FE946D62B4A" italics="true" pageId="3" pageNumber="1496">Melanorivulus terena</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C02BFCAFFDCFFECFE646C0F440A2B4A" baseAuthorityName="Costa" baseAuthorityYear="2005" box="[475,569,524,549]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="egens">
<emphasis id="B976185BFFDCFFECFE646C0F440A2B4A" box="[475,569,524,549]" italics="true" pageId="3" pageNumber="1496">M. egens</emphasis>
</taxonomicName>
by the presence of red oblique bars in the caudal fin in males (vs caudal fin without bars), smaller eye diameter in males (eye diameter 28.8– 32.3% HL vs 33.1–36.4% HL); larger body depth (22.0–23.1% SL vs 21.3–21.8% SL), and smaller pre-pelvic length (52.8–55.7% SL vs 56.7–56.8% SL).
<taxonomicName id="4C02BFCAFFDCFFECFCC86C7742552BE2" box="[887,1126,628,653]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFDCFFECFCC86C7742552BE2" box="[887,1126,628,653]" italics="true" pageId="3" pageNumber="1496">Melanorivulus terena</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C02BFCAFFDCFFECFE3B6C9447C42BDF" authorityName="Costa" authorityYear="2018" box="[388,503,663,688]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="linearis">
<emphasis id="B976185BFFDCFFECFE3B6C9447C42BDF" box="[388,503,663,688]" italics="true" pageId="3" pageNumber="1496">M. linearis</emphasis>
</taxonomicName>
by smaller caudal fin in males (caudal-fin length 29.9– 33.6% SL vs 36.6–36.9% SL), and in females (caudal-fin length 30.6–32.1% SL vs 34.2– 37.4% SL); by smaller pelvic fin in males (pelvic-fin length 10.3–11.6% SL, vs 13.1–15.4% SL); and by larger head width in females (head width 75.0–80.6% HL vs 70.5–74.0% HL).
</paragraph>
</subSubSection>
<subSubSection id="C31897C2FFDCFFE9FF1F6D50456D28F5" lastPageId="6" lastPageNumber="1499" pageId="3" pageNumber="1496" type="description">
<paragraph id="8BBDC449FFDCFFECFF1F6D5047152A02" blockId="3.[160,1156,851,981]" box="[160,294,851,877]" pageId="3" pageNumber="1496">
<emphasis id="B976185BFFDCFFECFF1F6D5047152A02" bold="true" box="[160,294,851,877]" italics="true" pageId="3" pageNumber="1496">Description</emphasis>
</paragraph>
<paragraph id="8BBDC449FFDCFFEBFF1F6D7444152982" blockId="3.[160,1156,851,981]" lastBlockId="4.[160,1156,142,1242]" lastPageId="4" lastPageNumber="1497" pageId="3" pageNumber="1496">
Morphometric data is presented in
<tableCitation id="C680F1F2FFDCFFECFDF96D7444AE2AFF" box="[582,669,887,912]" captionStart="Table 1" captionStartId="3.[220,276,1049,1072]" captionText="Table 1. Morphometric and meristic data for the holotype and paratypes of Melanorivulus terena sp. n." pageId="3" pageNumber="1496">Table 1</tableCitation>
. Males larger than females. Largest male examined
<quantity id="4CFA69ACFFDCFFECFEA76D99474E2ADC" box="[280,381,922,947]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="3" pageNumber="1496" unit="mm" value="28.1">28.1 mm</quantity>
SL, largest female
<quantity id="4CFA69ACFFDCFFECFDE26D9944F12ADC" box="[605,706,922,947]" metricMagnitude="-2" metricUnit="m" metricValue="2.61" pageId="3" pageNumber="1496" unit="mm" value="26.1">26.1 mm</quantity>
SL. Dorsal profile slightly convex from snout to end of dorsal-fin base. Ventral profile convex from lower jaw to origin of anal fin. Dorsal and ventral profiles of caudal peduncle nearly straight. Body slender, approximately cylindrical and compressed. Greatest body depth at pelvic-fin base. Snout weakly pointed in lateral view. Jaws short.
</paragraph>
<caption id="DF7D94C1FFDCFFECFF636A1A47692D22" pageId="3" pageNumber="1496" startId="3.[220,276,1049,1072]" targetType="table">
<paragraph id="8BBDC449FFDCFFECFF636A1A47692D22" blockId="3.[220,1095,1049,1101]" pageId="3" pageNumber="1496">
<emphasis id="B976185BFFDCFFECFF636A1A471F2D5F" bold="true" box="[220,300,1049,1072]" pageId="3" pageNumber="1496">Table 1.</emphasis>
Morphometric and meristic data for the holotype and paratypes of
<taxonomicName id="4C02BFCAFFDCFFECFC7D6A1A472A2D22" authorityName="Volcan &amp; Severo-Neto &amp; Lanés" authorityYear="2024" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="1496" phylum="Chordata" rank="species" species="terena" status="sp. nov.">
<emphasis id="B976185BFFDCFFECFC7D6A1A472A2D22" italics="true" pageId="3" pageNumber="1496">Melanorivulus terena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245A520FFDCFFECFE9F6A3547692D22" box="[288,346,1078,1101]" pageId="3" pageNumber="1496" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BBDC449FFDCFFECFD806A5942362FD9" pageId="3" pageNumber="1496">
<table id="F90236E9FFDC0010FF586A59420E2FD9" box="[231,1085,1114,1718]" gridcols="4" gridrows="25" pageId="3" pageNumber="1496">
<tr id="3532C60BFFDC0010FF586A59420E2D01" box="[231,1085,1114,1134]" gridrow="0" pageId="3" pageNumber="1496" rowspan-0="1">
<th id="76E3AF77FFDC0010FD806A5944B92D01" box="[575,650,1114,1134]" gridcol="1" gridrow="0" pageId="3" pageNumber="1496">Holotype</th>
<th id="76E3AF77FFDC0010FD546A5945692D01" box="[747,858,1114,1134]" gridcol="2" gridrow="0" pageId="3" pageNumber="1496">Males (n = 5)</th>
<th id="76E3AF77FFDC0010FC056A59420E2D01" box="[954,1085,1114,1134]" gridcol="3" gridrow="0" pageId="3" pageNumber="1496">Females (n = 4)</th>
</tr>
<tr id="3532C60BFFDC0010FF586A79420E2DE1" box="[231,1085,1146,1166]" gridrow="1" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586A79479A2DE1" box="[231,425,1146,1166]" gridcol="0" gridrow="1" pageId="3" pageNumber="1496">Standard length (mm)</th>
<td id="76E3AF77FFDC0010FD806A7944B92DE1" box="[575,650,1146,1166]" gridcol="1" gridrow="1" pageId="3" pageNumber="1496">27.2</td>
<td id="76E3AF77FFDC0010FD546A7945692DE1" box="[747,858,1146,1166]" gridcol="2" gridrow="1" pageId="3" pageNumber="1496">21.1–26.1</td>
<td id="76E3AF77FFDC0010FC056A79420E2DE1" box="[954,1085,1146,1166]" gridcol="3" gridrow="1" pageId="3" pageNumber="1496">20.9–26.1</td>
</tr>
<tr id="3532C60BFFDC0010FF586A91420E2DC9" box="[231,1085,1170,1190]" gridrow="2" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586A91420E2DC9" box="[231,1085,1170,1190]" colspan="4" colspanRight="3" gridcol="0" gridrow="2" pageId="3" pageNumber="1496">
<emphasis id="B976185BFFDCFFECFF586A9144312DC9" bold="true" box="[231,514,1170,1190]" pageId="3" pageNumber="1496">Percentages of standard length</emphasis>
</th>
</tr>
<tr id="3532C60BFFDC0010FF586AA9420E2DD1" box="[231,1085,1194,1214]" gridrow="3" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586AA9479A2DD1" box="[231,425,1194,1214]" gridcol="0" gridrow="3" pageId="3" pageNumber="1496">Body depth</th>
<td id="76E3AF77FFDC0010FD806AA944B92DD1" box="[575,650,1194,1214]" gridcol="1" gridrow="3" pageId="3" pageNumber="1496">25.7</td>
<td id="76E3AF77FFDC0010FD546AA945692DD1" box="[747,858,1194,1214]" gridcol="2" gridrow="3" pageId="3" pageNumber="1496">22.3–24.5</td>
<td id="76E3AF77FFDC0010FC056AA9420E2DD1" box="[954,1085,1194,1214]" gridcol="3" gridrow="3" pageId="3" pageNumber="1496">22.0–23.1</td>
</tr>
<tr id="3532C60BFFDC0010FF586AC1420E2DB9" box="[231,1085,1218,1238]" gridrow="4" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586AC1479A2DB9" box="[231,425,1218,1238]" gridcol="0" gridrow="4" pageId="3" pageNumber="1496">Caudal peduncle depth</th>
<td id="76E3AF77FFDC0010FD806AC144B92DB9" box="[575,650,1218,1238]" gridcol="1" gridrow="4" pageId="3" pageNumber="1496">15.8</td>
<td id="76E3AF77FFDC0010FD546AC145692DB9" box="[747,858,1218,1238]" gridcol="2" gridrow="4" pageId="3" pageNumber="1496">14.1–16.1</td>
<td id="76E3AF77FFDC0010FC056AC1420E2DB9" box="[954,1085,1218,1238]" gridcol="3" gridrow="4" pageId="3" pageNumber="1496">13.8–14.9</td>
</tr>
<tr id="3532C60BFFDC0010FF586AD9420E2D81" box="[231,1085,1242,1262]" gridrow="5" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586AD9479A2D81" box="[231,425,1242,1262]" gridcol="0" gridrow="5" pageId="3" pageNumber="1496">Predorsal length</th>
<td id="76E3AF77FFDC0010FD806AD944B92D81" box="[575,650,1242,1262]" gridcol="1" gridrow="5" pageId="3" pageNumber="1496">76.1</td>
<td id="76E3AF77FFDC0010FD546AD945692D81" box="[747,858,1242,1262]" gridcol="2" gridrow="5" pageId="3" pageNumber="1496">74.1–77.6</td>
<td id="76E3AF77FFDC0010FC056AD9420E2D81" box="[954,1085,1242,1262]" gridcol="3" gridrow="5" pageId="3" pageNumber="1496">75.1–78.7</td>
</tr>
<tr id="3532C60BFFDC0010FF586AF1420E2C69" box="[231,1085,1266,1286]" gridrow="6" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586AF1479A2C69" box="[231,425,1266,1286]" gridcol="0" gridrow="6" pageId="3" pageNumber="1496">Prepelvic length</th>
<td id="76E3AF77FFDC0010FD806AF144B92C69" box="[575,650,1266,1286]" gridcol="1" gridrow="6" pageId="3" pageNumber="1496">52.6</td>
<td id="76E3AF77FFDC0010FD546AF145692C69" box="[747,858,1266,1286]" gridcol="2" gridrow="6" pageId="3" pageNumber="1496">53.3–56.8</td>
<td id="76E3AF77FFDC0010FC056AF1420E2C69" box="[954,1085,1266,1286]" gridcol="3" gridrow="6" pageId="3" pageNumber="1496">52.9–55.7</td>
</tr>
<tr id="3532C60BFFDC0010FF586B09420E2C71" box="[231,1085,1290,1310]" gridrow="7" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586B09479A2C71" box="[231,425,1290,1310]" gridcol="0" gridrow="7" pageId="3" pageNumber="1496">Dorsal-fin base length</th>
<td id="76E3AF77FFDC0010FD806B0944B92C71" box="[575,650,1290,1310]" gridcol="1" gridrow="7" pageId="3" pageNumber="1496">11.8</td>
<td id="76E3AF77FFDC0010FD546B0945692C71" box="[747,858,1290,1310]" gridcol="2" gridrow="7" pageId="3" pageNumber="1496">10.7–12.3</td>
<td id="76E3AF77FFDC0010FC056B09420E2C71" box="[954,1085,1290,1310]" gridcol="3" gridrow="7" pageId="3" pageNumber="1496">10.2–12.0</td>
</tr>
<tr id="3532C60BFFDC0010FF586B21420E2C59" box="[231,1085,1314,1334]" gridrow="8" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586B21479A2C59" box="[231,425,1314,1334]" gridcol="0" gridrow="8" pageId="3" pageNumber="1496">Anal-fin base length</th>
<td id="76E3AF77FFDC0010FD806B2144B92C59" box="[575,650,1314,1334]" gridcol="1" gridrow="8" pageId="3" pageNumber="1496">19.5</td>
<td id="76E3AF77FFDC0010FD546B2145692C59" box="[747,858,1314,1334]" gridcol="2" gridrow="8" pageId="3" pageNumber="1496">19.3–20.7</td>
<td id="76E3AF77FFDC0010FC056B21420E2C59" box="[954,1085,1314,1334]" gridcol="3" gridrow="8" pageId="3" pageNumber="1496">18.4–19.6</td>
</tr>
<tr id="3532C60BFFDC0010FF586B39420E2C21" box="[231,1085,1338,1358]" gridrow="9" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586B39479A2C21" box="[231,425,1338,1358]" gridcol="0" gridrow="9" pageId="3" pageNumber="1496">Caudal-fin length</th>
<td id="76E3AF77FFDC0010FD806B3944B92C21" box="[575,650,1338,1358]" gridcol="1" gridrow="9" pageId="3" pageNumber="1496">32.7</td>
<td id="76E3AF77FFDC0010FD546B3945692C21" box="[747,858,1338,1358]" gridcol="2" gridrow="9" pageId="3" pageNumber="1496">29.9–33.6</td>
<td id="76E3AF77FFDC0010FC056B39420E2C21" box="[954,1085,1338,1358]" gridcol="3" gridrow="9" pageId="3" pageNumber="1496">30.7–32.1</td>
</tr>
<tr id="3532C60BFFDC0010FF586B51420E2C09" box="[231,1085,1362,1382]" gridrow="10" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586B51479A2C09" box="[231,425,1362,1382]" gridcol="0" gridrow="10" pageId="3" pageNumber="1496">Pectoral-fin length</th>
<td id="76E3AF77FFDC0010FD806B5144B92C09" box="[575,650,1362,1382]" gridcol="1" gridrow="10" pageId="3" pageNumber="1496">20.6</td>
<td id="76E3AF77FFDC0010FD546B5145692C09" box="[747,858,1362,1382]" gridcol="2" gridrow="10" pageId="3" pageNumber="1496">18.5–21.8</td>
<td id="76E3AF77FFDC0010FC056B51420E2C09" box="[954,1085,1362,1382]" gridcol="3" gridrow="10" pageId="3" pageNumber="1496">18.3–20.8</td>
</tr>
<tr id="3532C60BFFDC0010FF586B69420E2C11" box="[231,1085,1386,1406]" gridrow="11" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586B69479A2C11" box="[231,425,1386,1406]" gridcol="0" gridrow="11" pageId="3" pageNumber="1496">Pelvic-fin length</th>
<td id="76E3AF77FFDC0010FD806B6944B92C11" box="[575,650,1386,1406]" gridcol="1" gridrow="11" pageId="3" pageNumber="1496">10.3</td>
<td id="76E3AF77FFDC0010FD546B6945692C11" box="[747,858,1386,1406]" gridcol="2" gridrow="11" pageId="3" pageNumber="1496">10.7–11.6</td>
<td id="76E3AF77FFDC0010FC056B69420E2C11" box="[954,1085,1386,1406]" gridcol="3" gridrow="11" pageId="3" pageNumber="1496">9.4–9.6</td>
</tr>
<tr id="3532C60BFFDC0010FF586B81420E2CF9" box="[231,1085,1410,1430]" gridrow="12" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586B81479A2CF9" box="[231,425,1410,1430]" gridcol="0" gridrow="12" pageId="3" pageNumber="1496">Head length</th>
<td id="76E3AF77FFDC0010FD806B8144B92CF9" box="[575,650,1410,1430]" gridcol="1" gridrow="12" pageId="3" pageNumber="1496">28.7</td>
<td id="76E3AF77FFDC0010FD546B8145692CF9" box="[747,858,1410,1430]" gridcol="2" gridrow="12" pageId="3" pageNumber="1496">27.8–30.5</td>
<td id="76E3AF77FFDC0010FC056B81420E2CF9" box="[954,1085,1410,1430]" gridcol="3" gridrow="12" pageId="3" pageNumber="1496">27.6–28.8</td>
</tr>
<tr id="3532C60BFFDC0010FF586B99420E2CC1" box="[231,1085,1434,1454]" gridrow="13" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586B99420E2CC1" box="[231,1085,1434,1454]" colspan="4" colspanRight="3" gridcol="0" gridrow="13" pageId="3" pageNumber="1496">
<emphasis id="B976185BFFDCFFECFF586B9947EC2CC1" bold="true" box="[231,479,1434,1454]" pageId="3" pageNumber="1496">Percentages of head length</emphasis>
</th>
</tr>
<tr id="3532C60BFFDC0010FF586BB1420E2CA9" box="[231,1085,1458,1478]" gridrow="14" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586BB1479A2CA9" box="[231,425,1458,1478]" gridcol="0" gridrow="14" pageId="3" pageNumber="1496">Head depth</th>
<td id="76E3AF77FFDC0010FD806BB144B92CA9" box="[575,650,1458,1478]" gridcol="1" gridrow="14" pageId="3" pageNumber="1496">71.8</td>
<td id="76E3AF77FFDC0010FD546BB145692CA9" box="[747,858,1458,1478]" gridcol="2" gridrow="14" pageId="3" pageNumber="1496">63.1–72.2</td>
<td id="76E3AF77FFDC0010FC056BB1420E2CA9" box="[954,1085,1458,1478]" gridcol="3" gridrow="14" pageId="3" pageNumber="1496">66.7–69.4</td>
</tr>
<tr id="3532C60BFFDC0010FF586BC9420E2CB1" box="[231,1085,1482,1502]" gridrow="15" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586BC9479A2CB1" box="[231,425,1482,1502]" gridcol="0" gridrow="15" pageId="3" pageNumber="1496">Head width</th>
<td id="76E3AF77FFDC0010FD806BC944B92CB1" box="[575,650,1482,1502]" gridcol="1" gridrow="15" pageId="3" pageNumber="1496">78.2</td>
<td id="76E3AF77FFDC0010FD546BC945692CB1" box="[747,858,1482,1502]" gridcol="2" gridrow="15" pageId="3" pageNumber="1496">70.8–76.4</td>
<td id="76E3AF77FFDC0010FC056BC9420E2CB1" box="[954,1085,1482,1502]" gridcol="3" gridrow="15" pageId="3" pageNumber="1496">75.0–80.6</td>
</tr>
<tr id="3532C60BFFDC0010FF586BE1420E2C99" box="[231,1085,1506,1526]" gridrow="16" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586BE1479A2C99" box="[231,425,1506,1526]" gridcol="0" gridrow="16" pageId="3" pageNumber="1496">Snout length</th>
<td id="76E3AF77FFDC0010FD806BE144B92C99" box="[575,650,1506,1526]" gridcol="1" gridrow="16" pageId="3" pageNumber="1496">20.5</td>
<td id="76E3AF77FFDC0010FD546BE145692C99" box="[747,858,1506,1526]" gridcol="2" gridrow="16" pageId="3" pageNumber="1496">15.8–22.2</td>
<td id="76E3AF77FFDC0010FC056BE1420E2C99" box="[954,1085,1506,1526]" gridcol="3" gridrow="16" pageId="3" pageNumber="1496">14.9–22.2</td>
</tr>
<tr id="3532C60BFFDC0010FF586BF9420E2F61" box="[231,1085,1530,1550]" gridrow="17" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586BF9479A2F61" box="[231,425,1530,1550]" gridcol="0" gridrow="17" pageId="3" pageNumber="1496">Lower jaw length</th>
<td id="76E3AF77FFDC0010FD806BF944B92F61" box="[575,650,1530,1550]" gridcol="1" gridrow="17" pageId="3" pageNumber="1496">24.4</td>
<td id="76E3AF77FFDC0010FD546BF945692F61" box="[747,858,1530,1550]" gridcol="2" gridrow="17" pageId="3" pageNumber="1496">18.0–21.5</td>
<td id="76E3AF77FFDC0010FC056BF9420E2F61" box="[954,1085,1530,1550]" gridcol="3" gridrow="17" pageId="3" pageNumber="1496">17.9–23.3</td>
</tr>
<tr id="3532C60BFFDC0010FF586811420E2F49" box="[231,1085,1554,1574]" gridrow="18" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586811479A2F49" box="[231,425,1554,1574]" gridcol="0" gridrow="18" pageId="3" pageNumber="1496">Eye diameter</th>
<td id="76E3AF77FFDC0010FD80681144B92F49" box="[575,650,1554,1574]" gridcol="1" gridrow="18" pageId="3" pageNumber="1496">29.5</td>
<td id="76E3AF77FFDC0010FD54681145692F49" box="[747,858,1554,1574]" gridcol="2" gridrow="18" pageId="3" pageNumber="1496">28.8–32.3</td>
<td id="76E3AF77FFDC0010FC056811420E2F49" box="[954,1085,1554,1574]" gridcol="3" gridrow="18" pageId="3" pageNumber="1496">28.4–34.4</td>
</tr>
<tr id="3532C60BFFDC0010FF586829420E2F51" box="[231,1085,1578,1598]" gridrow="19" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586829420E2F51" box="[231,1085,1578,1598]" colspan="4" colspanRight="3" gridcol="0" gridrow="19" pageId="3" pageNumber="1496">
<emphasis id="B976185BFFDCFFECFF58682947162F51" bold="true" box="[231,293,1578,1598]" pageId="3" pageNumber="1496">Counts</emphasis>
</th>
</tr>
<tr id="3532C60BFFDC0010FF586841420E2F39" box="[231,1085,1602,1622]" gridrow="20" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586841479A2F39" box="[231,425,1602,1622]" gridcol="0" gridrow="20" pageId="3" pageNumber="1496">Dorsal fin ray</th>
<td id="76E3AF77FFDC0010FD80684144B92F39" box="[575,650,1602,1622]" gridcol="1" gridrow="20" pageId="3" pageNumber="1496">10</td>
<td id="76E3AF77FFDC0010FD54684145692F39" box="[747,858,1602,1622]" gridcol="2" gridrow="20" pageId="3" pageNumber="1496">9–10</td>
<td id="76E3AF77FFDC0010FC056841420E2F39" box="[954,1085,1602,1622]" gridcol="3" gridrow="20" pageId="3" pageNumber="1496">9–10</td>
</tr>
<tr id="3532C60BFFDC0010FF586859420E2F01" box="[231,1085,1626,1646]" gridrow="21" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586859479A2F01" box="[231,425,1626,1646]" gridcol="0" gridrow="21" pageId="3" pageNumber="1496">Caudal fin ray</th>
<td id="76E3AF77FFDC0010FD80685944B92F01" box="[575,650,1626,1646]" gridcol="1" gridrow="21" pageId="3" pageNumber="1496">28</td>
<td id="76E3AF77FFDC0010FD54685945692F01" box="[747,858,1626,1646]" gridcol="2" gridrow="21" pageId="3" pageNumber="1496">28–31</td>
<td id="76E3AF77FFDC0010FC056859420E2F01" box="[954,1085,1626,1646]" gridcol="3" gridrow="21" pageId="3" pageNumber="1496">30–32</td>
</tr>
<tr id="3532C60BFFDC0010FF586871420E2FE9" box="[231,1085,1650,1670]" gridrow="22" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586871479A2FE9" box="[231,425,1650,1670]" gridcol="0" gridrow="22" pageId="3" pageNumber="1496">Anal fin ray</th>
<td id="76E3AF77FFDC0010FD80687144B92FE9" box="[575,650,1650,1670]" gridcol="1" gridrow="22" pageId="3" pageNumber="1496">14</td>
<td id="76E3AF77FFDC0010FD54687145692FE9" box="[747,858,1650,1670]" gridcol="2" gridrow="22" pageId="3" pageNumber="1496">14–15</td>
<td id="76E3AF77FFDC0010FC056871420E2FE9" box="[954,1085,1650,1670]" gridcol="3" gridrow="22" pageId="3" pageNumber="1496">14–16</td>
</tr>
<tr id="3532C60BFFDC0010FF586889420E2FF1" box="[231,1085,1674,1694]" gridrow="23" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF586889479A2FF1" box="[231,425,1674,1694]" gridcol="0" gridrow="23" pageId="3" pageNumber="1496">Pelvic fin ray</th>
<td id="76E3AF77FFDC0010FD80688944B92FF1" box="[575,650,1674,1694]" gridcol="1" gridrow="23" pageId="3" pageNumber="1496">7</td>
<td id="76E3AF77FFDC0010FD54688945692FF1" box="[747,858,1674,1694]" gridcol="2" gridrow="23" pageId="3" pageNumber="1496">7</td>
<td id="76E3AF77FFDC0010FC056889420E2FF1" box="[954,1085,1674,1694]" gridcol="3" gridrow="23" pageId="3" pageNumber="1496">7</td>
</tr>
<tr id="3532C60BFFDC0010FF5868A1420E2FD9" box="[231,1085,1698,1718]" gridrow="24" pageId="3" pageNumber="1496">
<th id="76E3AF77FFDC0010FF5868A1479A2FD9" box="[231,425,1698,1718]" gridcol="0" gridrow="24" pageId="3" pageNumber="1496">Pectoral fin ray</th>
<td id="76E3AF77FFDC0010FD8068A144B92FD9" box="[575,650,1698,1718]" gridcol="1" gridrow="24" pageId="3" pageNumber="1496">13</td>
<td id="76E3AF77FFDC0010FD5468A145692FD9" box="[747,858,1698,1718]" gridcol="2" gridrow="24" pageId="3" pageNumber="1496">13</td>
<td id="76E3AF77FFDC0010FC0568A1420E2FD9" box="[954,1085,1698,1718]" gridcol="3" gridrow="24" pageId="3" pageNumber="1496">13</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBDC449FFDBFFEBFF7F6EF5444628B0" blockId="4.[160,1156,142,1242]" pageId="4" pageNumber="1497">Short dorsal and anal fins. Dorsal-fin rays 9–10. Dorsal fin slightly pointed in males and rounded in females. Dorsal-fin origin on vertical through between base of 8th and 10th anal-fin ray. Anal-fin rays 14–16. Anal fin slightly pointed in males and females. Caudal fin rounded 28–32 rays. Pectoral-fin rays 13. Pectoral fin rounded, with posterior margin reaching vertical at about 70–80% of length between pectoral-fin and pelvic-fin bases. Pelvic-fin rays 7. Pelvic-fin posterior tip reaching vertical at anterior to the anus to 1st anal-fin ray. Pelvic-fin bases in close proximity.</paragraph>
<paragraph id="8BBDC449FFDBFFEBFF7F6FEA42B72BC0" blockId="4.[160,1156,142,1242]" pageId="4" pageNumber="1497">Scales cycloid. Body and head entirely scaled, except anterior ventral surface of head. Body squamation extending over anterior 40% of caudal-fin base. No scales on dorsal and anal-fin bases. Frontal squamation E-patterned; E-scales not overlapping medially; scales arranged in regular circular pattern around A-scale without exposed margins; transverse row of scales anterior to H-scale. Longitudinal series of scales 31–32; transverse series of scales 10–11; scale rows around caudal peduncle 16. No contact organs on flank and fins.</paragraph>
<paragraph id="8BBDC449FFDBFFEBFF7F6CBA45712A55" blockId="4.[160,1156,142,1242]" pageId="4" pageNumber="1497">Cephalic neuromasts: supraorbital 3 + 3, parietal 1, anterior rostral 1, posterior rostral 1, infraorbital 1 + 11–14 + 1, preorbital 1–2, otic 1, postotic 1, supratemporal 1, median opercular 1, ventral opercular 1, preopercular 2 + 4, mandibular 3 + 1, lateral mandibular 1–2, paramandibular 1. Two neuromasts on caudal-fin base.</paragraph>
<paragraph id="8BBDC449FFDBFFEBFF7F6D4745962DB5" blockId="4.[160,1156,142,1242]" pageId="4" pageNumber="1497">
Six branchiostegal rays. Gill rakers on first branchial arch 1 + 7. First epibranchial approximately straight. Dorsal portion of preopercle short and pointed. Ventral process of posttemporal absent. Total number of vertebrae 30–31, 13 precaudal vertebrae in males and
<quantity id="4CFA69ACFFDBFFEBFEA86DAF47452AAA" box="[279,374,940,965]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="3.81" metricValueMin="3.556" pageId="4" pageNumber="1497" unit="in" value="14.5" valueMax="15.0" valueMin="14.0">14–15 in</quantity>
females, 17 caudal vertebrae in males and
<quantity id="4CFA69ACFFDBFFEBFCEE6DAF45832AAA" box="[849,944,940,965]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="4" pageNumber="1497" unit="in" value="16.5" valueMax="17.0" valueMin="16.0">16–17 in</quantity>
females. Dorsal-fin origin on vertical between neural spines of 19th and 20th vertebrae in males and between the 20th and 21st vertebrae in females. Origin of anal fin at vertical through pleural ribs of 13th and 14th vertebrae in males and between the 14th and 15th vertebrae in females. Ventral process of angulo-articular short, pointed. Vomerine teeth 1–3. Dermosphenotic present. Basihyal sub-triangular, greatest width 55% of length; basihyal cartilage 25% of total basihyal length. Second pharyngobranchial teeth absent. Teeth on the fourth ceratobranchial divided into two rows exhibiting a median gap. Interarcual cartilage well developed. Lateral process of ethmoid well developed and free.
</paragraph>
<paragraph id="8BBDC449FFDBFFEAFF1F6B0547A82F6E" blockId="4.[160,1157,1286,1728]" lastBlockId="5.[160,1156,1235,1537]" lastPageId="5" lastPageNumber="1498" pageId="4" pageNumber="1497">
<emphasis id="B976185BFFDBFFEBFF1F6B05445B2C4F" bold="true" box="[160,616,1286,1312]" pageId="4" pageNumber="1497">
<emphasis id="B976185BFFDBFFEBFF1F6B0547D12C4F" bold="true" box="[160,482,1286,1312]" italics="true" pageId="4" pageNumber="1497">Colouration in life – males</emphasis>
(
<figureCitation id="1339D8CCFFDBFFEBFE4C6B0544692C4F" box="[499,602,1286,1312]" captionStart="Figure 2" captionStartId="5.[160,227,1095,1118]" captionTargetBox="[163,1153,142,1062]" captionTargetId="figure-151@5.[163,1153,142,1063]" captionTargetPageId="5" captionText="Figure 2. Male ZUFMS-PIS08471 (above), and female ZUFMS-PIS06575 (below) paratypes of Melanorivulus terena, sp.n. Camapuã municipality, Mato Grosso do Sul, Brazil. Photos: Matheus Vieira Volcan (male) and Francisco Severo-Neto (female)." figureDoi="http://doi.org/10.5281/zenodo.13758562" httpUri="https://zenodo.org/record/13758562/files/figure.png" pageId="4" pageNumber="1497">Figure 2</figureCitation>
).
</emphasis>
Metallic greenish-grey flank, becoming purplish-blue close to the base of the anal fin; red dots scattered on anterior flank area, above the pectoral-fin insertion, forming 3–4 small oblique bars; 1 or 2 narrow continuous or interrupted red oblique bars, between the medial pectoral-fin portion and the pelvic-fin base; continuous bars forming chevron-like marks with an angle on the dorsal region of the flank; 7–9 continuous or interrupted oblique narrow red bars irregularly arranged between the pelvic-fin base and caudal peduncle, anteriorly directed, forming chevron-like marks, with an angle that goes from dorsal to midline in the anterior-posterior direction, no bar interruption pattern; vestigial red bars in the lateral-dorsal region of flank, from the medial pectoral fin portion to the dorsal origin of caudal fin. Dark red pigmentation in the postorbital region of the head, forming 2–3 oblique bars, sometimes contiguous with the red dots in the humeral region. Dorsum brownish-green, venter whitish-grey. Dorsal portion of the head brownish-green; ventral portion yellowish-grey; pale greenish-golden iridescence on opercular region. Dark grey jaws. Iridescent yellow iris. Dorsal fin yellow with 4–5 oblique red bars, orangish-red distal margin. Anal fin yellow, proximal portion light blue, medial posterior proximal portion with faint red marks, sometimes forming a U-shape; distal region with an orangish-red margin. Caudal fin yellow with 4–5 red oblique parallel bars, covering generally 1/3–1/2 of caudal fin on mid-dorsal portion; ventral portion light yellow without bars; distal margin of fin with an orangish-red colour, well delineated. Pectoral fin yellowish-hyaline. Pelvic fin yellow with orangish-red posterior margin.
</paragraph>
<caption id="DF7D94C1FFDAFFEAFF1F6A4444E12DF7" ID-DOI="http://doi.org/10.5281/zenodo.13758562" ID-Zenodo-Dep="13758562" httpUri="https://zenodo.org/record/13758562/files/figure.png" pageId="5" pageNumber="1498" startId="5.[160,227,1095,1118]" targetBox="[163,1153,142,1062]" targetPageId="5" targetType="figure">
<paragraph id="8BBDC449FFDAFFEAFF1F6A4444E12DF7" blockId="5.[160,1156,1095,1176]" pageId="5" pageNumber="1498">
<emphasis id="B976185BFFDAFFEAFF1F6A4447302D31" bold="true" box="[160,259,1095,1118]" pageId="5" pageNumber="1498">Figure 2.</emphasis>
Male ZUFMS-PIS08471 (above), and female ZUFMS-PIS06575 (below) paratypes of
<taxonomicName id="4C02BFCAFFDAFFEAFF1F6A67475C2D14" authorityName="Volcan &amp; Severo-Neto &amp; Lanés" authorityYear="2024" box="[160,367,1124,1147]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="1498" phylum="Chordata" rank="species" species="terena" status="sp. nov.">
<emphasis id="B976185BFFDAFFEAFF1F6A67475C2D14" box="[160,367,1124,1147]" italics="true" pageId="5" pageNumber="1498">Melanorivulus terena</emphasis>
</taxonomicName>
,
<emphasis id="B976185BFFDAFFEAFE3F6A6747872D14" bold="true" box="[384,436,1124,1147]" pageId="5" pageNumber="1498">
<taxonomicNameLabel id="A245A520FFDAFFEAFE3F6A6747872D14" box="[384,436,1124,1147]" pageId="5" pageNumber="1498" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
Camapuã municipality, Mato Grosso do Sul, Brazil. Photos: Matheus Vieira Volcan (male) and Francisco Severo-Neto (female).
</paragraph>
</caption>
<paragraph id="8BBDC449FFDAFFE9FF1F683F456D28F5" blockId="5.[160,1155,1596,1726]" lastBlockId="6.[160,1156,142,410]" lastPageId="6" lastPageNumber="1499" pageId="5" pageNumber="1498">
<emphasis id="B976185BFFDAFFEAFF1F683F47B02F39" bold="true" box="[160,387,1596,1622]" pageId="5" pageNumber="1498">
<emphasis id="B976185BFFDAFFEAFF1F683F47322F39" bold="true" box="[160,257,1596,1622]" italics="true" pageId="5" pageNumber="1498">Females</emphasis>
(
<figureCitation id="1339D8CCFFDAFFEAFEAF683F47472F39" box="[272,372,1596,1622]" captionStart="Figure 2" captionStartId="5.[160,227,1095,1118]" captionTargetBox="[163,1153,142,1062]" captionTargetId="figure-151@5.[163,1153,142,1063]" captionTargetPageId="5" captionText="Figure 2. Male ZUFMS-PIS08471 (above), and female ZUFMS-PIS06575 (below) paratypes of Melanorivulus terena, sp.n. Camapuã municipality, Mato Grosso do Sul, Brazil. Photos: Matheus Vieira Volcan (male) and Francisco Severo-Neto (female)." figureDoi="http://doi.org/10.5281/zenodo.13758562" httpUri="https://zenodo.org/record/13758562/files/figure.png" pageId="5" pageNumber="1498">Figure 2</figureCitation>
).
</emphasis>
Same bar pattern as males, with a greyish-ochre flank, becoming lighter towards the base of the caudal fin. Light ochre venter. Dorsum and dorsal portion of head brownish-ochre; ventral portion of head yellowish. Jaws brownishorange; opercular region pale ochre, with a few chromatophores scattered randomly and a conspicuous greenish-gold iridescent spot. Dorsal fin yellow, with 2 or 3 faint brown bars; distal margin faint dark brown. Anal fin yellow; basal portion light yellow becoming dark brown gradually towards distal margin; distal region with a dark brown margin. Caudal fin pale yellow, with 5 dark brown bars in the mid upper portion of the fin; faint dark brown distal margin, with mixed orange dots in the dorsal margin; conspicuous black spot on the dorso-basal portion of the fin, generally contiguous with the first bar. Pectoral fin hyaline with narrow orange margins. Pelvic fin yellowish hyaline with dark brown posterior margin with orange dots.
</paragraph>
</subSubSection>
<subSubSection id="C31897C2FFD9FFE9FF1F6FCE47042B21" pageId="6" pageNumber="1499" type="distribution">
<paragraph id="8BBDC449FFD9FFE9FF1F6FCE47042B21" blockId="6.[160,1156,461,590]" pageId="6" pageNumber="1499">
<emphasis id="B976185BFFD9FFE9FF1F6FCE479F2888" bold="true" box="[160,428,461,487]" pageId="6" pageNumber="1499">
<emphasis id="B976185BFFD9FFE9FF1F6FCE471F2888" bold="true" box="[160,300,461,487]" italics="true" pageId="6" pageNumber="1499">Distribution</emphasis>
(
<figureCitation id="1339D8CCFFD9FFE9FE856FCE47AE2888" box="[314,413,461,487]" captionStart="Figure 3" captionStartId="6.[160,225,1675,1698]" captionTargetBox="[165,1152,944,1642]" captionTargetId="figure-243@6.[163,1153,940,1644]" captionTargetPageId="6" captionText="Figure 3. Occurrence area of Melanorivulus terena (type locality) in the upper Rio Coxim basin, Rio Paraguay basin and closest related species of M. pictus group in the Rio Paraná basin." figureDoi="http://doi.org/10.5281/zenodo.13758564" httpUri="https://zenodo.org/record/13758564/files/figure.png" pageId="6" pageNumber="1499">Figure 3</figureCitation>
).
</emphasis>
<taxonomicName id="4C02BFCAFFD9FFE9FE036FCE44AC2889" authorityName="Volcan &amp; Severo-Neto &amp; Lanés" authorityYear="2024" box="[444,671,461,486]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="1499" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFD9FFE9FE036FCE44AC2889" box="[444,671,461,486]" italics="true" pageId="6" pageNumber="1499">Melanorivulus terena</emphasis>
</taxonomicName>
is only known from one location in Ribeirão Cachoeirinha, a small stream in the headwaters of the Coxim River, a tributary of the Taquari River, in the
<collectingCountry id="F31584D9FFD9FFE9FE3A6C1047DE2B43" box="[389,493,531,556]" name="Paraguay" pageId="6" pageNumber="1499">Paraguay</collectingCountry>
River basin, municipality of Camapuã, state of
<collectingRegion id="49C60AABFFD9FFE9FC486C1046D92B21" country="Brazil" name="Mato Grosso do Sul" pageId="6" pageNumber="1499">Mato Grosso do Sul</collectingRegion>
,
<collectingCountry id="F31584D9FFD9FFE9FF4B6C3647002B21" box="[244,307,565,590]" name="Brazil" pageId="6" pageNumber="1499">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31897C2FFD9FFE7FF1F6C8244EB2A55" lastPageId="8" lastPageNumber="1501" pageId="6" pageNumber="1499" type="etymology">
<paragraph id="8BBDC449FFD9FFE9FF1F6C82450F2A6C" blockId="6.[160,1155,641,771]" pageId="6" pageNumber="1499">
<emphasis id="B976185BFFD9FFE9FF1F6C8247102BF4" bold="true" box="[160,291,641,667]" italics="true" pageId="6" pageNumber="1499">Etymology.</emphasis>
The specific epithet ‘terena’ is in allusion to the Terena ethnic group. The Terena people belong to the Aruak linguistic family and are the last remaining members of the Guará nation. The Terena are currently the most visible Indigenous people in the upper
<collectingCountry id="F31584D9FFD9FFE9FF546CE947602A6C" box="[235,339,746,771]" name="Paraguay" pageId="6" pageNumber="1499">Paraguay</collectingCountry>
portion in the state of Mato Grosso do Sul.
</paragraph>
<paragraph id="8BBDC449FFD9FFE7FF1F6D3545602860" blockId="6.[160,1156,822,882]" lastBlockId="8.[160,1155,142,271]" lastPageId="8" lastPageNumber="1501" pageId="6" pageNumber="1499">
<emphasis id="B976185BFFD9FFE9FF1F6D3547332A3F" bold="true" box="[160,256,822,848]" italics="true" pageId="6" pageNumber="1499">Habitat.</emphasis>
The new species was found in a small first-order stream (locally called ‘veredas’), with a depth of about
<quantity id="4CFA69ACFFD9FFE9FE7E6D5A443C2A1D" box="[449,527,857,882]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="1499" unit="cm" value="20.0">20 cm</quantity>
, at the headwaters of the Taquari River (
<figureCitation id="1339D8CCFFD9FFE9FBAE6D5A42452A1D" box="[1041,1142,857,882]" captionStart="Figure 4" captionStartId="7.[160,225,1525,1548]" captionTargetBox="[163,1152,142,1494]" captionTargetId="figure-3@7.[163,1153,142,1494]" captionTargetPageId="7" captionText="Figure 4. Type locality of Melanorivulus terena sp. n. in Ribeirão Cachoeirinha, tributary of headwaters of Rio Coxim, Paraguay River Basin, Camapuã municipality, Mato Grosso do Sul state, Brazil (A). Photo: Matheus Volcan. Detail of the type of environment where the species was recorded (B). Photo: Francisco Severo-Neto." figureDoi="http://doi.org/10.5281/zenodo.13758566" httpUri="https://zenodo.org/record/13758566/files/figure.png" pageId="6" pageNumber="1499">Figure 4</figureCitation>
).
<taxonomicName id="4C02BFCAFFD7FFE7FF1F6E8D47B729C8" authorityName="Volcan &amp; Severo-Neto &amp; Lanés" authorityYear="2024" box="[160,388,142,167]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="1501" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFD7FFE7FF1F6E8D47B729C8" box="[160,388,142,167]" italics="true" pageId="8" pageNumber="1501">Melanorivulus terena</emphasis>
</taxonomicName>
was recorded in the stream bank and adjacent shallow pools where the water was clear, still or gently flowing, with muddy and clay substrates, within the savanna-like Cerrado domain. The species is found in areas exposed to sun, as well as in areas shaded by trees, mainly buritis palm (
<taxonomicName id="4C02BFCAFFD7FFE7FD356EF545752860" authorityName="Ulmer" authorityYear="1905" box="[650,838,246,271]" class="Gastropoda" family="Cypraeidae" genus="Mauritia" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="1501" phylum="Mollusca" rank="species" species="flexuosa">
<emphasis id="B976185BFFD7FFE7FD356EF545752860" box="[650,838,246,271]" italics="true" pageId="8" pageNumber="1501">Mauritia flexuosa</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF7D94C1FFD9FFE9FF1F688845C02FD0" ID-DOI="http://doi.org/10.5281/zenodo.13758564" ID-Zenodo-Dep="13758564" httpUri="https://zenodo.org/record/13758564/files/figure.png" pageId="6" pageNumber="1499" startId="6.[160,225,1675,1698]" targetBox="[165,1152,944,1642]" targetPageId="6" targetType="figure">
<paragraph id="8BBDC449FFD9FFE9FF1F688845C02FD0" blockId="6.[160,1155,1675,1727]" pageId="6" pageNumber="1499">
<emphasis id="B976185BFFD9FFE9FF1F688846CF2FCD" bold="true" box="[160,252,1675,1698]" pageId="6" pageNumber="1499">Figure 3.</emphasis>
Occurrence area of
<taxonomicName id="4C02BFCAFFD9FFE9FE74688844A62FCD" authorityName="Volcan &amp; Severo-Neto &amp; Lanés" authorityYear="2024" box="[459,661,1675,1698]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="1499" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFD9FFE9FE74688844A62FCD" box="[459,661,1675,1698]" italics="true" pageId="6" pageNumber="1499">Melanorivulus terena</emphasis>
</taxonomicName>
(type locality) in the upper Rio Coxim basin, Rio Paraguay basin and closest related species of
<taxonomicName id="4C02BFCAFFD9FFE9FDD868AB448D2FD0" baseAuthorityName="Costa" baseAuthorityYear="1989" box="[615,702,1704,1727]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="1499" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B976185BFFD9FFE9FDD868AB448D2FD0" box="[615,702,1704,1727]" italics="true" pageId="6" pageNumber="1499">M. pictus</emphasis>
</taxonomicName>
group in the Rio Paraná basin.
</paragraph>
</caption>
<caption id="DF7D94C1FFD8FFE8FF1F6BF647B12F0B" ID-DOI="http://doi.org/10.5281/zenodo.13758566" ID-Zenodo-Dep="13758566" httpUri="https://zenodo.org/record/13758566/files/figure.png" pageId="7" pageNumber="1500" startId="7.[160,225,1525,1548]" targetBox="[163,1152,142,1494]" targetPageId="7" targetType="figure">
<paragraph id="8BBDC449FFD8FFE8FF1F6BF647B12F0B" blockId="7.[160,1157,1525,1636]" pageId="7" pageNumber="1500">
<emphasis id="B976185BFFD8FFE8FF1F6BF646CA2F63" bold="true" box="[160,249,1525,1548]" pageId="7" pageNumber="1500">Figure 4.</emphasis>
Type locality of
<taxonomicName id="4C02BFCAFFD8FFE8FE266BF644522F63" authorityName="Volcan &amp; Severo-Neto &amp; Lanés" authorityYear="2024" box="[409,609,1525,1548]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="1500" phylum="Chordata" rank="species" species="terena" status="sp. nov.">
<emphasis id="B976185BFFD8FFE8FE266BF644522F63" box="[409,609,1525,1548]" italics="true" pageId="7" pageNumber="1500">Melanorivulus terena</emphasis>
</taxonomicName>
<emphasis id="B976185BFFD8FFE8FDD96BF644AC2F63" bold="true" box="[614,671,1525,1548]" pageId="7" pageNumber="1500">
<taxonomicNameLabel id="A245A520FFD8FFE8FDD96BF644AC2F63" box="[614,671,1525,1548]" pageId="7" pageNumber="1500" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
in Ribeirão Cachoeirinha, tributary of headwaters of Rio Coxim, Paraguay River Basin, Camapuã municipality, Mato Grosso do Sul state, Brazil (A). Photo: Matheus Volcan. Detail of the type of environment where the species was recorded (B). Photo: Francisco Severo-Neto.
</paragraph>
</caption>
<paragraph id="8BBDC449FFD7FFE7FF1F6F3844EB2A55" blockId="8.[160,1156,315,826]" pageId="8" pageNumber="1501">
<emphasis id="B976185BFFD7FFE7FF1F6F3847A4283A" bold="true" box="[160,407,315,341]" italics="true" pageId="8" pageNumber="1501">Conservation status.</emphasis>
The new species is endemic to the Coxim River, a tributary of Taquari River, and was recorded in only one location, but there are potential habitats for its occurrence along the stream and in the surrounding area. It has been recorded in an area with major impacts from intense agricultural activity. Another main activity in the region is cattle farming, which has drastically altered the original vegetation. The main threats are erosion and excessive sedimentation in the drainages of Ribeirão Cachoeirinha and its tributaries, as well as trampling by cattle in shallow wetlands (veredas) located along the stream banks. In addition, the area of occurrence of
<taxonomicName id="4C02BFCAFFD7FFE7FCE46C2C45F22B27" authorityName="Volcan &amp; Severo-Neto &amp; Lanés" authorityYear="2024" box="[859,961,559,584]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="1501" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFD7FFE7FCE46C2C45F22B27" box="[859,961,559,584]" italics="true" pageId="8" pageNumber="1501">M. terena</emphasis>
</taxonomicName>
is crossed by the MS-422 highway. Consequently, the species is also subject to transportation-based accidents with toxic loads, which pose a threat to the
<taxonomicName id="4C02BFCAFFD7FFE7FD5D6C7745792BE2" authorityName="Volcan &amp; Severo-Neto &amp; Lanés" authorityYear="2024" box="[738,842,628,653]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="1501" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFD7FFE7FD5D6C7745792BE2" box="[738,842,628,653]" italics="true" pageId="8" pageNumber="1501">M. terena</emphasis>
</taxonomicName>
population. Because of the previously mentioned threats, the high degree of endemism, and the field observations,
<taxonomicName id="4C02BFCAFFD7FFE7FF1F6CBA473B2BBD" authorityName="Volcan &amp; Severo-Neto &amp; Lanés" authorityYear="2024" box="[160,264,697,722]" family="Rivulidae" genus="Melanorivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="1501" phylum="Chordata" rank="species" species="terena">
<emphasis id="B976185BFFD7FFE7FF1F6CBA473B2BBD" box="[160,264,697,722]" italics="true" pageId="8" pageNumber="1501">M. terena</emphasis>
</taxonomicName>
is considered ‘Vulnerable’, following the criteria of the International Union for Conservation of Nature (
<bibRefCitation id="EF93B9B8FFD7FFE7FE0A6CDF441C2B9A" author="IUCN" box="[437,559,732,757]" pageId="8" pageNumber="1501" refId="ref7974" refString="IUCN. 2012. IUCN Red list categories and criteria: version 3.1. 2 nd. Gland (Switzerland): IUCN; p. 32." type="journal volume" year="2012">IUCN 2012</bibRefCitation>
). The species has an area of occupation of less than
<quantity id="4CFA69ACFFD7FFE7FF1F6CFC46CC2A77" box="[160,255,767,792]" metricMagnitude="6" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="1501" unit="km" value="2000.0">2000 km</quantity>
2, it occurs in only one locality, and it suffers from a reduction in its present area of occurrence and in its habitat quality (VU B2abii,iii).
</paragraph>
</subSubSection>
</treatment>
</document>
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