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<document id="22B765EF102933E5F2257DF1860265F4" ID-CLB-Dataset="27116" ID-DOI="http://dx.doi.org/10.3897/zookeys.840.33326" ID-GBIF-Dataset="363f2c7c-0635-46c8-b598-9e9aa014b6d2" ID-PMC="PMC6482118" ID-Pensoft-Pub="1313-2970-840-35" ID-PubMed="31065227" ID-ZooBank="C607C2FDC17644FABBC582D561C6434C" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-840-35" ModsDocOrigin="ZooKeys 840" ModsDocTitle="Conchological and molecular analysis of the “non-scaly” Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)" checkinTime="1555526082661" checkinUser="pensoft" docAuthor="Zacaery Khalik, Mohd, P. Hendriks, Kasper, J. Vermeulen, Jaap &amp; Menno Schilthuizen," docDate="2019" docId="4A94025A23D8300206463E367CFB426A" docLanguage="en" docName="ZooKeys 840: 35-86" docOrigin="ZooKeys 840" docSource="http://dx.doi.org/10.3897/zookeys.840.33326" docTitle="Georissa filiasaulae Haase &amp; Schilthuizen 2007" docType="treatment" docVersion="6" lastPageNumber="59" masterDocId="647C1D35F570C105FFFCFFCDB169212D" masterDocTitle="Conchological and molecular analysis of the &quot; non-scaly &quot; Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)" masterLastPageNumber="86" masterPageNumber="35" pageNumber="58" updateTime="1701381596384" updateUser="ExternalLinkService">
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<mods:title id="3BD0293E49B76745888CBDE8BBCD8682">Conchological and molecular analysis of the &quot; non-scaly &quot; Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)</mods:title>
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<mods:namePart id="F88BC07EFCE8EE8F313C3AE604C813DE">Zacaery Khalik, Mohd</mods:namePart>
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<mods:namePart id="226D25C2260A26F4B6F4EC13936FC2F7">P. Hendriks, Kasper</mods:namePart>
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<mods:namePart id="508C19CDE52E77AE804D6F27E51F2EF1">J. Vermeulen, Jaap</mods:namePart>
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<mods:namePart id="EACAB59BCB03355D48EA07BDAB8DE51C">Menno Schilthuizen,</mods:namePart>
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<treatment id="4A94025A23D8300206463E367CFB426A" ID-GBIF-Taxon="156787944" LSID="urn:lsid:plazi:treatment:4A94025A23D8300206463E367CFB426A" httpUri="http://treatment.plazi.org/id/4A94025A23D8300206463E367CFB426A" lastPageId="24" lastPageNumber="59" pageId="23" pageNumber="58">
<subSubSection id="10F9BE0320EAA9D6E762060196B33F6C" pageId="23" pageNumber="58" type="nomenclature">
<paragraph id="391E26868965E5F84A4B8E2097FFF1DD" pageId="23" pageNumber="58">
<taxonomicName id="3F57E310C5D63D709C81723519931FA3" ID-CoL="7RDV3" authority="Haase &amp; Schilthuizen, 2007" authorityName="Haase &amp; Schilthuizen" authorityYear="2007" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa filiasaulae" order="Cycloneritida" pageId="23" pageNumber="58" phylum="Mollusca" rank="species" species="filiasaulae">Georissa filiasaulae Haase &amp; Schilthuizen, 2007</taxonomicName>
Figure 13
<normalizedToken id="B41BBED03454E5E579A0DD65FCB2BD45" originalValue="AK">A-K</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="F06455B12273C171670D5236702E5FF6" pageId="23" pageNumber="58" type="reference_group">
<paragraph id="5E2930D8C1F48F77786A2CB8B69513BE" pageId="23" pageNumber="58">
<taxonomicName id="08DCAC2E5CFA8C62712C4CF7F66EA71C" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa filiasaulae" order="Cycloneritida" pageId="23" pageNumber="58" phylum="Mollusca" rank="species" species="filiasaulae">Georissa filiasaulae</taxonomicName>
Haase &amp; Schilthuizen, 2007: 216, figs 2
<normalizedToken id="386B2FA612DB845E42D0BDBEA5B35E05" originalValue="AB">A-B</normalizedToken>
and 2E;
<bibRefCitation id="27B1FCB0D80AC8D4118FBAB0C30AC3BA" author="Clements, R" journalOrPublisher="Biological Conservation" pageId="39" pageNumber="74" url="https://doi.org/10.1641/0006-3568(2006)56[733:lkosai]2.0.co;2" year="2006">Clements et al. 2006</bibRefCitation>
: 736;
<bibRefCitation id="1DA04B2D3DA57DC62A6F0ACFB44409BA" author="Clements, R" journalOrPublisher="Biological Conservation" pageId="39" pageNumber="74" pagination="2751 - 2764" title="Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts." url="https://doi.org/10.1016/j.biocon.2008.08.011" volume="141" year="2008">Clements et al. 2008</bibRefCitation>
: Appendix D;
<bibRefCitation id="D63D42D4C9B1692CBA30E6E49835FD81" author="Schilthuizen, M" journalOrPublisher="Biological Journal of the Linnean Society" pageId="41" pageNumber="76" pagination="727 - 740" title="Small-scale genetic structuring in a tropical cave snail and admixture with its above-ground sister species." url="https://doi.org/10.1111/j.1095-8312.2011.01835.x" volume="105" year="2012">Schilthuizen et al. 2012</bibRefCitation>
;
<bibRefCitation id="7DE31BC4507840882338081C811EF617" author="Khalik, MZ" journalOrPublisher="ZooKeys" pageId="40" pageNumber="75" pagination="1 - 55" title="A molecular and conchological dissection of the &quot; scaly &quot; Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)." url="https://doi.org/10.3897/zookeys.773.24878.figure10" volume="773" year="2018">Khalik et al. 2018</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5271B29BFE542E099EE2FB2C54B11C7D" pageId="23" pageNumber="58" type="type locality">
<paragraph id="B4DA0F1BEF350449218712429C392DE0" pageId="23" pageNumber="58">Type locality.</paragraph>
<paragraph id="EDF019B0E9A52B3A9650FE5FE4BD7775" pageId="23" pageNumber="58">
Malaysia, Sabah, Sepulut valley, Interior province, Batu Sanaron (
<geoCoordinate id="E95FC904D183954BC0E22DE30B61C5CF" direction="north" orientation="latitude" precision="9" value="4.7008333">04°42.05'N</geoCoordinate>
,
<geoCoordinate id="BCEFAA73608D5F765FD6D1A3E3E26869" direction="east" orientation="longitude" precision="9" value="116.60017">116°36.01'E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="1A620F8D4D195B15AA2767B1789CFAC9" pageId="23" pageNumber="58" type="type material">
<paragraph id="93A1F53C109CA6B10C8500A7A65B34BD" pageId="23" pageNumber="58">Type material.</paragraph>
<paragraph id="6FC6C11B11554232D5A3FC955A49C27A" pageId="23" pageNumber="58">
Holotype (Holotype by original designation). Malaysia, Sabah, Sepulut valley, Interior province, Batu Sanaron (
<geoCoordinate id="7E759523040415C65402003189B6C250" direction="north" orientation="latitude" precision="9" value="4.7008333">04°42.05'N</geoCoordinate>
,
<geoCoordinate id="6CAA23FC70CA77E72BC523AF9BC87A23" direction="east" orientation="longitude" precision="9" value="116.60017">116°36.01'E</geoCoordinate>
): BOR/MOL 3795. Paratypes. Malaysia, Sabah, Sepulut valley, Interior province, Batu Sanaron (
<geoCoordinate id="BAC83094AE5F0685FE3930E5EF19CD03" direction="north" orientation="latitude" precision="9" value="4.7008333">04°42.05'N</geoCoordinate>
,
<geoCoordinate id="C636BE39ED1079F9D06922A0CA8808B1" direction="east" orientation="longitude" precision="9" value="116.60017">116°36.01'E</geoCoordinate>
): BOR/MOL 3491 (7); ZMB 107143-107149 (7) (not seen).
</paragraph>
</subSubSection>
<subSubSection id="97D94ABF2E6228176C0F7E45BE94DF56" pageId="23" pageNumber="58" type="materials_examined">
<paragraph id="51F28AE722689189E6CF287478B5B816" pageId="23" pageNumber="58">Other material.</paragraph>
<paragraph id="1B7554EAADBAD884897173C181CA6FEF" pageId="23" pageNumber="58">
Batu Sanaron, Interior province, Sepulut valley, Sabah: BOR/MOL 532, BOR/MOL 3405. Batu Pungiton, Interior province, Sepulut valley, Sabah Batu Pungiton, Sabah (
<geoCoordinate id="597AFDE179A27C19EC44466FECA50951" direction="north" orientation="latitude" precision="9" value="4.7068334">04°42.41'N</geoCoordinate>
,
<geoCoordinate id="D5766BFC3781B62B225A7BCCE484EBCE" direction="east" orientation="longitude" precision="9" value="116.60067">116°36.04'E</geoCoordinate>
): BOR/MOL 12768 (Fig. 13).
</paragraph>
<caption id="A27740FC10E1201E7DA61A969AEBEEC6" pageId="23" pageNumber="58">
<paragraph id="72480C20D7E91150DBE5E12C498763F4" pageId="23" pageNumber="58">
Figure 13.
<taxonomicName id="C2655B9B5391E889E61F7374C5F0064E" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa filiasaulae" order="Cycloneritida" pageId="23" pageNumber="58" phylum="Mollusca" rank="species" species="filiasaulae">Georissa filiasaulae</taxonomicName>
Haase &amp; Schilthuizen, 2007.
<normalizedToken id="FCA02757599B9556D7C6E1ACFEE292BA" originalValue="AK">A-K</normalizedToken>
BOR/MOL 12768 A, D shell apertural view B shell side view C shell rear view E, F shell cross-section from 3D model G, H operculum frontal and ventral view from 3D model I shell top view J protoconch side view K close up of protoconch from top at 1000
<normalizedToken id="63DB36E9F21D3E758B8F3ADDDC43E504" originalValue="×">x</normalizedToken>
magnification. Scale bars: 500
<normalizedToken id="E2F68B7994D937B97A065DA93C8F1273" originalValue="µm">µm</normalizedToken>
(
<normalizedToken id="FEE30978430745541BBB029CEEB1BFD7" originalValue="AI">A-I</normalizedToken>
); 200
<normalizedToken id="E455ECC929FF20ABD798456DCD45DC30" originalValue="µm">µm</normalizedToken>
(J); 10
<normalizedToken id="71A5EEC3591CE17B8B4DC24C191AE445" originalValue="µm">µm</normalizedToken>
(K).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D07B814E6485367AAC9C4A7086343423" pageId="23" pageNumber="58" type="description">
<paragraph id="3CD62EA6838587BE7A9A7DDACC5EAF63" pageId="23" pageNumber="58">Description.</paragraph>
<paragraph id="55B8AFB30A6E935BFE48777DA9E6DDF7" pageId="23" pageNumber="58">
Protoconch. Colour: white. Sculpture: a mix of rounded, ellipsoidal to irregular sculptural shape. Mesh width: 2.5-20
<normalizedToken id="D8D87CB0C4210A079CB06A227AFBF1AA" originalValue="µm">µm</normalizedToken>
. Teleoconch. Colour: white. First whorl: convex. Subsequent whorls: convex, shell shape slender to broad. Suture: clearly impressed. Shoulder: slightly extended, regularly spaced nodules. Number of whorls: 2
<normalizedToken id="34976B3164EE412F860DD55F6684F2AB" originalValue="½–">1/2-</normalizedToken>
3. SH: 1.21-1.68 mm. SW: 1.67-1.68 mm. SI: 1.08-1.27. Shell sculpture. Radial sculpture: absent, weak to strong growth lines. Spiral sculpture: present, rather weak and thin, densely spaced on the first whorl, the ribbing appears immediately after the protoconch, sometimes distorted/discontinuous by the growth lines, superficially smooth on the later whorls. Aperture. Shape: rounded to semi-elliptic, straight to slightly convex parietal side, palatal side rounded, palatal edge partially contiguous with the body whorl and parietal side, basal side convex. AH: 0.67- 0.79 mm. AW: 0.69-0.83 mm. AI: 0.93-0.97.
</paragraph>
</subSubSection>
<subSubSection id="9B8D835706214AEFCAB05AB73017428A" pageId="23" pageNumber="58" type="diagnosis">
<paragraph id="085F72B350D19B14B89638B5FE5C695D" pageId="23" pageNumber="58">Diagnosis.</paragraph>
<paragraph id="D83D866475DC5F448457AC0691BA5599" pageId="23" pageNumber="58">
<taxonomicName id="6ADC009F7F2544CFE995B186AA4A1E40" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa filiasaulae" order="Cycloneritida" pageId="23" pageNumber="58" phylum="Mollusca" rank="species" species="filiasaulae">Georissa filiasaulae</taxonomicName>
has weak, thin, and densely arranged spiral sculpture with nodular structure on the shoulder. The shell colour and thickness are most similar to
<taxonomicName id="91013E3A289039A945833918C2A3EC51" lsidName="G. corrugata" pageId="23" pageNumber="58" rank="species" species="corrugata">G. corrugata</taxonomicName>
, which has white and partially transparent shell.
</paragraph>
</subSubSection>
<subSubSection id="1EABFF490DB8F5F1D5D9B36473FA848A" pageId="24" pageNumber="59" type="distribution">
<paragraph id="1CB808D5CD90BB2D17427E84B1F9F024" pageId="24" pageNumber="59">
<pageBreakToken id="8D6648C8AC04D3AF214181C5B2784DA2" pageId="24" pageNumber="59" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph id="9B07152EC84D35BA0B6AB19F807070EB" pageId="24" pageNumber="59">
<taxonomicName id="0D9C6BE0ED2D4428C22E2AC297CD7B6C" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa filiasaulae" order="Cycloneritida" pageId="24" pageNumber="59" phylum="Mollusca" rank="species" species="filiasaulae">Georissa filiasaulae</taxonomicName>
is a cave specialist, known from the cave system of Batu Sanaron and Batu Tinahas in the Sepulut valley.
<bibRefCitation id="3737BCC25096D4B6DF94C5A78DAD1904" author="Schilthuizen, M" journalOrPublisher="Biological Journal of the Linnean Society" pageId="41" pageNumber="76" pagination="727 - 740" title="Small-scale genetic structuring in a tropical cave snail and admixture with its above-ground sister species." url="https://doi.org/10.1111/j.1095-8312.2011.01835.x" volume="105" year="2012">Schilthuizen et al. (2012)</bibRefCitation>
studied the population genetics of
<taxonomicName id="ABD0FFF9C689923C6458958ED05599B4" lsidName="G. filiasaulae" pageId="24" pageNumber="59" rank="species" species="filiasaulae">G. filiasaulae</taxonomicName>
and its sister species,
<taxonomicName id="B34A7818D050C468B8F0F338414B8523" lsidName="G. saulae" pageId="24" pageNumber="59" rank="species" species="saulae">G. saulae</taxonomicName>
. They found narrow hybrid zones between the two species in cave entrances.
</paragraph>
</subSubSection>
<subSubSection id="FE29FFDCBCA0D38BFC42E3D41F70D8A0" pageId="24" pageNumber="59" type="molecular analysis">
<paragraph id="F9B772DCEF78A9E5CCC0FC5EC3330016" pageId="24" pageNumber="59">Molecular analysis.</paragraph>
<paragraph id="F8758E802175422FA33D3FAE2C26C777" pageId="24" pageNumber="59">
ML and Bayesian analyses of
<taxonomicName id="486CC1C7F78E2DF91CD51814F115CEB0" lsidName="G. filiasaulae" pageId="24" pageNumber="59" rank="species" species="filiasaulae">G. filiasaulae</taxonomicName>
(16S: n = 3; CO1: n = 3) show that
<taxonomicName id="3271E2259B03EEF634445A0AA12F8712" lsidName="G. filiasaulae" pageId="24" pageNumber="59" rank="species" species="filiasaulae">G. filiasaulae</taxonomicName>
form one clade with 98% BS and 100% PP. The sister group is the
<taxonomicName id="764BADA8A6AD2CE5707C81150391599A" lsidName="G. saulae" pageId="24" pageNumber="59" rank="species" species="saulae">G. saulae</taxonomicName>
population from Pungiton (
<taxonomicName id="49B87F460AD0AEFD9A17419B24837D20" lsidName="G. saulae" pageId="24" pageNumber="59" rank="species" species="saulae">G. saulae</taxonomicName>
is paraphyletic).
</paragraph>
</subSubSection>
<subSubSection id="FFBF2864C8FE857DAAE1038206C4DD6E" pageId="24" pageNumber="59" type="discussion">
<paragraph id="EFC352CBDE16F4921AAF633939D49DB9" pageId="24" pageNumber="59">Discussion.</paragraph>
<paragraph id="2B720744A837A30E533A8D9BCDBEAF37" pageId="24" pageNumber="59">
<taxonomicName id="3CF4E634ADB73F6B27802A6DB2E53B2D" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa filiasaulae" order="Cycloneritida" pageId="24" pageNumber="59" phylum="Mollusca" rank="species" species="filiasaulae">Georissa filiasaulae</taxonomicName>
is one of the two known Bornean
<taxonomicName id="51542F6C65E4B8D22F397ABFBB14F0DD" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa" order="Cycloneritida" pageId="24" pageNumber="59" phylum="Mollusca" rank="genus">Georissa</taxonomicName>
that is troglobitic.
<bibRefCitation id="80E0D5DBD9AD7610BF7A85F8280B5F5E" author="Khalik, MZ" journalOrPublisher="ZooKeys" pageId="40" pageNumber="75" pagination="1 - 55" title="A molecular and conchological dissection of the &quot; scaly &quot; Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)." url="https://doi.org/10.3897/zookeys.773.24878.figure10" volume="773" year="2018">Khalik et al. (2018)</bibRefCitation>
described
<taxonomicName id="8AD9E52CF33CBC8374E686649644F7CF" lsidName="G. silaburensis" pageId="24" pageNumber="59" rank="species" species="silaburensis">G. silaburensis</taxonomicName>
, another species of Bornean
<taxonomicName id="939EB23CC28BF45D94034C5675BB081E" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa" order="Cycloneritida" pageId="24" pageNumber="59" phylum="Mollusca" rank="genus">Georissa</taxonomicName>
from the
<normalizedToken id="726465F6EA7347E301E5ADE998C4B388" originalValue="“scaly”">&quot;scaly&quot;</normalizedToken>
group as a possible troglobite from Gunung Silabur, Serian, Sarawak.
<taxonomicName id="7E712C332862425A1295736A6F98F0F7" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa filiasaulae" order="Cycloneritida" pageId="24" pageNumber="59" phylum="Mollusca" rank="species" species="filiasaulae">Georissa filiasaulae</taxonomicName>
differs from
<taxonomicName id="F9693B22EDD1D4DDC0C0DF80AD136FE2" lsidName="G. saulae" pageId="24" pageNumber="59" rank="species" species="saulae">G. saulae</taxonomicName>
by the absence of any scale-like sculpture, reduced shell pigmentation, and relatively larger shell size and broader shell shape. Population genetic studies suggest that the hybrid zone between the two is restricted to a narrow region at the cave entrances, rendering the two species as independent evolutionary units. Therefore, considering them as separate species is warranted (
<bibRefCitation id="7380C3628535B161975BCD0BC539658A" author="Schilthuizen, M" journalOrPublisher="BioEssays" pageId="41" pageNumber="76" pagination="1134 - 1141" title="Dualism and conflicts in understanding speciation." url="https://doi.org/10.1002/1521-1878(200012)22:12&lt;1134::aid-bies11&gt;3.0.co;2-5" volume="22" year="2000">Schilthuizen 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>