239 lines
38 KiB
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239 lines
38 KiB
XML
<document id="B0C06D61A30A5C596F4B2EC41CA9E70D" ID-CLB-Dataset="32199" ID-DOI="10.5852/ejt.2017.261" ID-GBIF-Dataset="708e152e-4450-4439-84a9-3c56148cd082" ID-Zenodo-Dep="885318" ID-ZooBank="C59239D1-0BF0-446A-AF6E-71234D01B23D" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1504621962129" checkinUser="plazi" docAuthor="Reboleira, Ana Sofa P. S., Fresneda, Javier & Salgado, José Maria" docDate="2017" docId="03FB5E336667AB51B7246466FAA03CAA" docLanguage="en" docName="ejt-261.pdf.imf" docOrigin="European Journal of Taxonomy 261" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Speonemadus Jeannel 1922" docType="treatment" docVersion="18" lastPageNumber="3" masterDocId="FFC2264B6665AB53B56C656DFFD63B4C" masterDocTitle="A new species of Speonemadus from Portugal, with the revision of the escalerai-group (Coleoptera, Leiodidae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="3" updateTime="1734103965070" updateUser="ExternalLinkService">
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<mods:title id="B6E34756D069E670C6BCC526832EE51D">A new species of Speonemadus from Portugal, with the revision of the escalerai-group (Coleoptera, Leiodidae)</mods:title>
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<mods:namePart id="4442C309A8356E8317DC835150582A15">Reboleira, Ana Sofa P. S.</mods:namePart>
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<mods:namePart id="C5E9E3BF5285A5EF8965AF1D8B5C29A4">Fresneda, Javier</mods:namePart>
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<mods:namePart id="74C74E538B222E90196E6721255BCC22">Salgado, José Maria</mods:namePart>
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<mods:title id="76F0B20051314A6168D42313140A3551">European Journal of Taxonomy</mods:title>
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<mods:date id="403A24F8CF93F92E816C90CFD8C0A677">2017</mods:date>
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<treatment id="03FB5E336667AB51B7246466FAA03CAA" ID-DOI="http://doi.org/10.5281/zenodo.5689566" ID-GBIF-Taxon="133183536" ID-Zenodo-Dep="5689566" LSID="urn:lsid:plazi:treatment:03FB5E336667AB51B7246466FAA03CAA" httpUri="http://treatment.plazi.org/id/03FB5E336667AB51B7246466FAA03CAA" lastPageNumber="3" pageId="2" pageNumber="3" scope_family="Leiodidae" scope_order="Coleoptera" scope_subFamily="Cholevinae" scope_tribe="Anemadini">
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<subSubSection id="C348BCAE6667AB51B7246466FC3D3A69" box="[584,1003,267,294]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BEDEF256667AB51B7246466FC3D3A69" blockId="2.[584,1003,267,294]" box="[584,1003,267,294]" pageId="2" pageNumber="3">
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<heading id="D0A558496667AB51B7246466FC3D3A69" box="[584,1003,267,294]" centered="true" fontSize="11" level="3" pageId="2" pageNumber="3" reason="4">
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Genus
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<treatmentCitation id="0AF3C9346667AB51B7F76466FC3D3A69" author="Jeannel" authority="Jeannel, 1922" authorityName="Jeannel" authorityYear="1922" box="[667,1003,267,294]" class="Insecta" family="Leiodidae" firstAuthor="Jeannel" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" pagination="1 - 98" phylum="Arthropoda" rank="genus" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." treatmentTaxon="Speonemadus" type="journal article" year="1922">
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<taxonomicName id="4C5294A66667AB51B7F76466FC3D3A69" ID-CoL="8X7VV" authority="Jeannel, 1922" authorityName="Jeannel" authorityYear="1922" box="[667,1003,267,294]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B92633376667AB51B7F76466FC953A6A" bold="true" box="[667,835,267,294]" italics="true" pageId="2" pageNumber="3">Speonemadus</emphasis>
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<bibRefCitation id="EFC392D46667AB51B6266466FC3D3A69" author="Jeannel" box="[842,1003,267,293]" firstAuthor="Jeannel" pageId="2" pageNumber="3" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel, 1922</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C348BCAE6667AB51B5D16420FB113A24" box="[189,1223,333,360]" pageId="2" pageNumber="3" type="type_taxon">
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<paragraph id="8BEDEF256667AB51B5D16420FB113A24" blockId="2.[189,1223,333,360]" box="[189,1223,333,360]" pageId="2" pageNumber="3">
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<taxonomicName id="4C5294A66667AB51B5D16420FE043A2B" ID-CoL="4YHYZ" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[189,466,333,359]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B92633376667AB51B5D16420FE043A2B" box="[189,466,333,359]" italics="true" pageId="2" pageNumber="3">Speonemadus escalerai</emphasis>
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</taxonomicName>
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-group; “Gruppo
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<taxonomicName id="4C5294A66667AB51B7F16420FCDF3A2B" ID-CoL="4YHYZ" authority="Giachino" authorityName="Giachino" authorityPageNumber="205" authorityYear="1993" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[669,777,333,359]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B92633376667AB51B7F16420FCDF3A2B" box="[669,777,333,359]" italics="true" pageId="2" pageNumber="3">escalerai</emphasis>
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</taxonomicName>
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”
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<emphasis id="B92633376667AB51B6716423FC883A2B" box="[797,862,334,359]" italics="true" pageId="2" pageNumber="3">sensu</emphasis>
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<bibRefCitation id="EFC392D46667AB51B6096420FB123A24" author="Giachino P. M. & Vailati D." box="[869,1220,333,360]" firstAuthor="Giachino" journalOrPublisher="Museo Cívico di Scienze Naturali di Brescia. Monografe di “ Natura Bresciana ”" pageId="2" pageNumber="3" pagination="1 – 314" part="18" refId="ref12035" refString="Fuente J. M. 1925. Catalogo sistematico-geografco de los Coleopteros observados en la Peninsula Iberica, Pirineos propiamente dichos y Baleares. Boletin de la Real Sociedad Entomologica de Espana vii (2 - 3) [1924]: 93 - 117. Giachino P. M. & Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana " 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino & Vailati 1993: 205</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C348BCAE6667AB51B5D164EAFB933862" pageId="2" pageNumber="3" type="description">
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<paragraph id="8BEDEF256667AB51B5D164EAFBA4391D" blockId="2.[189,1399,390,593]" pageId="2" pageNumber="3">
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Representatives of this group are characterized by a body length between 3.9 to 5.0 mm and have a slightly transverse pronotum (
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<figureCitation id="1369F3A06667AB51B74464C7FD733A89" box="[552,677,426,453]" captionStart="Figs 6–15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6–15. Speonemadus of the escalerai-group pronotum. 6–7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 8–9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 10–11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 12–13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 14–15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="2" pageNumber="3">Figs 6–15</figureCitation>
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); the anterior tibia of the males is compressed towards the apical half and has a keel with different shapes on the dorsal edge (
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<figureCitation id="1369F3A06667AB51B6B464A0FBB43AA4" box="[984,1122,461,488]" captionStart="Figs 16 – 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 – 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Figs 16–20</figureCitation>
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); elytra have a rounded apex in males and a notched one in females. The median lobe of the aedeagus is elongated and bottleshaped in dorsal view (
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<figureCitation id="1369F3A06667AB51B4B8677EFD8A3962" box="[468,604,531,558]" captionStart="Figs 21 – 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 – 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="2" pageNumber="3">Figs 21–25</figureCitation>
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) and the inner sac lacks a ventral tooth in the apical region of the middle lobe and has two long parallel rows of fine spines and teeth (
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<figureCitation id="1369F3A06667AB51B6B0675AFBB0391D" box="[988,1126,567,593]" captionStart="Figs 21 – 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 – 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="2" pageNumber="3">Figs 23–24</figureCitation>
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).
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</paragraph>
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<paragraph id="8BEDEF256667AB51B5D1671DFCC2399D" blockId="2.[189,1399,624,721]" pageId="2" pageNumber="3">All species of the group have similar color patterns, the same antennal shape and development of the membranous wings in both sexes, as well as similar genital segments and aedeagi in males, and similar 7th and 8th urosternites and spermatheca of females.</paragraph>
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<paragraph id="8BEDEF256667AB51B5D1679CFB933862" blockId="2.[189,1398,753,815]" pageId="2" pageNumber="3">The life-style of this group is mostly subterranean, they are frequently found under stones or in leaf litter in forests and in caves, walking in bat guano or in the subterranean substrate.</paragraph>
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</subSubSection>
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<subSubSection id="C348BCAE6667AB51B5D16623FCB63824" box="[189,864,845,872]" pageId="2" pageNumber="3" type="multiple">
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<paragraph id="8BEDEF256667AB51B5D16623FCB63824" blockId="2.[189,1399,845,1235]" box="[189,864,845,872]" pageId="2" pageNumber="3">
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<heading id="D0A558496667AB51B5D16623FCB63824" bold="true" box="[189,864,845,872]" fontSize="11" level="2" pageId="2" pageNumber="3" reason="6">
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<emphasis id="B92633376667AB51B5D16623FCB63824" bold="true" box="[189,864,845,872]" pageId="2" pageNumber="3">
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Key to the species of the
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<taxonomicName id="4C5294A66667AB51B49D6620FCDA3824" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[497,780,845,872]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B92633376667AB51B49D6620FCDA3824" bold="true" box="[497,780,845,872]" italics="true" pageId="2" pageNumber="3">Speonemadus escalerai</emphasis>
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</taxonomicName>
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-group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C348BCAE6667AB51B5D16610FC173F9F" pageId="2" pageNumber="3" type="discussion">
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<paragraph id="8BEDEF256667AB51B5D16610FC173F9F" blockId="2.[189,1399,845,1235]" pageId="2" pageNumber="3">
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Note to the key: diversification of the species of the
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<taxonomicName id="4C5294A66667AB51B6596611FC7738DA" ID-CoL="4YHYZ" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[821,929,892,918]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B92633376667AB51B6596611FC7738DA" box="[821,929,892,918]" italics="true" pageId="2" pageNumber="3">escalerai</emphasis>
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</taxonomicName>
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-group is likely to be recent (see below “Speciation and biogeography”). Therefore, morphological differences are not particularly visible and a combination of characters should be used for species recognition. All species and both sexes of the “
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<taxonomicName id="4C5294A66667AB51B5A6668BFEE03F4C" ID-CoL="4YHYZ" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[202,310,998,1024]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B92633376667AB51B5A6668BFEE03F4C" box="[202,310,998,1024]" italics="true" pageId="2" pageNumber="3">escalerai</emphasis>
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</taxonomicName>
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” group show a similar coloration, shape of the antenna, development of membranous wings, genital segment in the male aedeagus and 7th and 8th uroventrites, as well as spermatheca of the females. Females have no characters to distinguish between species, which also occurs in other genera of the
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<taxonomicName id="4C5294A66667AB51B5D1613DFEE33F26" ID-CoL="8VR3J" authorityName="Fleming" authorityYear="1821" box="[189,309,1104,1130]" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
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, e.g.,
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<emphasis id="B92633376667AB51B41B613DFE693F26" box="[375,447,1104,1130]" italics="true" pageId="2" pageNumber="3">Colon</emphasis>
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Herbst, 1797. Only females of
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<taxonomicName id="4C5294A66667AB51B64A613DFC0E3F26" ID-CoL="4YHYV" baseAuthorityName="Kraatz" baseAuthorityYear="1870" box="[806,984,1104,1130]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="angusticollis">
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<emphasis id="B92633376667AB51B64A613DFC0E3F26" box="[806,984,1104,1130]" italics="true" pageId="2" pageNumber="3">S. angusticollis</emphasis>
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</taxonomicName>
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can be separated from other species by the shape of the pronotum, being the only one with a bell-shape, as in males. Therefore, in this species group only males provide distinctive characters useful for identification. These characters are identified in all key items and constitute the basis of specific differentiation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C348BCAE6667AB51B5D16181FAA03CAA" pageId="2" pageNumber="3" type="key">
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<keyStep id="30A6F7806667AB51B5D16181FAA13E65" pageId="2" pageNumber="3">
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<paragraph id="8BEDEF256667AB51B5D16181FAA03E4A" blockId="2.[189,1399,1259,2023]" box="[189,1398,1259,1286]" pageId="2" pageNumber="3">
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<keyLead id="30A34C106667AB51B5D16181FAA03E4A" box="[189,1398,1259,1286]" pageId="2" pageNumber="3">
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1. Male with a high protibial keel, with the highest point around middle of the tibia (
|
||
<figureCitation id="1369F3A06667AB51B1FC6186FAEB3E4A" box="[1168,1341,1259,1286]" captionStart="Figs 16 – 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 – 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Figs 16, 18, 20</figureCitation>
|
||
) …2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256667AB51B5AD6062FAA13E65" blockId="2.[189,1399,1259,2023]" box="[193,1399,1295,1321]" pageId="2" pageNumber="3">
|
||
<keyLead id="30A34C106667AB51B5AD6062FAA13E65" box="[193,1399,1295,1321]" pageId="2" pageNumber="3">
|
||
– Male with a lower protibial keel, with the highest point at the basal third of the tibia (
|
||
<figureCitation id="1369F3A06667AB51B1D56062FAED3E65" box="[1209,1339,1295,1321]" captionStart="Figs 16 – 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 – 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Figs 17, 19</figureCitation>
|
||
) …4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A6F7806667AB51B5D16024FAA03EBC" pageId="2" pageNumber="3">
|
||
<paragraph id="8BEDEF256667AB51B5D16024FAA03EE5" blockId="2.[189,1399,1259,2023]" pageId="2" pageNumber="3">
|
||
<keyLead id="30A34C106667AB51B5D16024FAA03EE5" pageId="2" pageNumber="3">
|
||
2. Protibial keel with sharp vertex and slightly lobed from the margin to the base (
|
||
<figureCitation id="1369F3A06667AB51B1D36024FACD3E2F" box="[1215,1307,1353,1379]" captionStart="Figs 16 – 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 – 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Fig. 20</figureCitation>
|
||
); elytra sculpture and pronotum granularity poorly marked; pronotum sexually dimorphic (
|
||
<figureCitation id="1369F3A06667AB51B1C56001FAFB3ECB" box="[1193,1325,1388,1415]" captionStart="Figs 6–15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6–15. Speonemadus of the escalerai-group pronotum. 6–7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 8–9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 10–11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 12–13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 14–15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="2" pageNumber="3">Figs 14–15</figureCitation>
|
||
) …… …………………………………………………………………………
|
||
<treatmentCitation id="0AF3C9346667AB51B14C60E2FAA03EE5" author="Uhagon" authority="(Uhagon, 1898)" authorityName="Uhagon" authorityYear="1898" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[1056,1398,1423,1450]" class="Insecta" family="Leiodidae" firstAuthor="Uhagon" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" pagination="117 - 126" phylum="Arthropoda" rank="species" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." species="escalerai" treatmentTaxon="S. escalerai" type="journal article" year="1898">
|
||
<taxonomicName id="4C5294A66667AB51B14C60E2FAA03EE5" ID-CoL="4YHYZ" authority="(Uhagon, 1898)" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[1056,1398,1423,1450]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="escalerai">
|
||
<emphasis id="B92633376667AB51B14C60E2FB783EE6" bold="true" box="[1056,1198,1423,1450]" italics="true" pageId="2" pageNumber="3">S. escalerai</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC392D46667AB51B1AD60E2FAB83EE5" author="Uhagon" box="[1217,1390,1423,1449]" firstAuthor="Uhagon" pageId="2" pageNumber="3" pagination="117 - 126" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." type="journal article" year="1898">Uhagón, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256667AB51B5AD60DEFAA03EBC" blockId="2.[189,1399,1259,2023]" pageId="2" pageNumber="3">
|
||
<keyLead id="30A34C106667AB51B5AD60DEFAA03EBC" pageId="2" pageNumber="3">
|
||
– Protibial keel with a slightly sharp vertex and linear margin to the base (
|
||
<figureCitation id="1369F3A06667AB51B14460DFFB7B3E80" box="[1064,1197,1458,1484]" captionStart="Figs 16 – 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 – 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Figs 16, 18</figureCitation>
|
||
); elytra sculpture and pronotum with well marked granularity; pronotum not sexually dimorphic (
|
||
<figureCitation id="1369F3A06667AB51B1EC60B8FAEA3EBC" box="[1152,1340,1493,1520]" captionStart="Figs 6–15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6–15. Speonemadus of the escalerai-group pronotum. 6–7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 8–9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 10–11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 12–13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 14–15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="2" pageNumber="3">Figs 6–7, 10–11</figureCitation>
|
||
) …3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A6F7806667AB51B5D16362FAA13DB1" pageId="2" pageNumber="3">
|
||
<paragraph id="8BEDEF256667AB51B5D16362FAA03D3C" blockId="2.[189,1399,1259,2023]" pageId="2" pageNumber="3">
|
||
<keyLead id="30A34C106667AB51B5D16362FAA03D3C" pageId="2" pageNumber="3">
|
||
3. Body size generally small (male length around
|
||
<quantity id="4CAA42C06667AB51B64E6362FC553D66" box="[802,899,1551,1578]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="3" unit="mm" value="4.2">4.2 mm</quantity>
|
||
); protibial keel with sharp apex and basal part of the protibia with a wide keel (
|
||
<figureCitation id="1369F3A06667AB51B7CB635FFD293D01" box="[679,767,1586,1613]" captionStart="Figs 16 – 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 – 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Fig. 18</figureCitation>
|
||
); 8th antennomere around 1.5 × longer than wide and 4th one length different from 5th and 7th antennomeres in both sexes …
|
||
<treatmentCitation id="0AF3C9346667AB51B154633BFAA03D3C" author="Jeannel" authority="(Jeannel, 1922)" authorityName="Jeannel" authorityYear="1922" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[1080,1398,1621,1648]" class="Insecta" family="Leiodidae" firstAuthor="Jeannel" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" pagination="1 - 98" phylum="Arthropoda" rank="species" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." species="bolivari" treatmentTaxon="S. bolivari" type="journal article" year="1922">
|
||
<taxonomicName id="4C5294A66667AB51B154633BFAA03D3C" ID-CoL="4YHYW" authority="(Jeannel, 1922)" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[1080,1398,1621,1648]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bolivari">
|
||
<emphasis id="B92633376667AB51B154633BFB603D3C" bold="true" box="[1080,1206,1621,1648]" italics="true" pageId="2" pageNumber="3">S. bolivari</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC392D46667AB51B1AB633BFAB83D3C" author="Jeannel" box="[1223,1390,1622,1648]" firstAuthor="Jeannel" pageId="2" pageNumber="3" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel, 1922</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256667AB51B5AD6314FAA13DB1" blockId="2.[189,1399,1259,2023]" pageId="2" pageNumber="3">
|
||
<keyLead id="30A34C106667AB51B5AD6314FAA13DB1" pageId="2" pageNumber="3">
|
||
– Body size generally larger (mean male length around
|
||
<quantity id="4CAA42C06667AB51B61A6314FC0E3DDF" box="[886,984,1657,1683]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="2" pageNumber="3" unit="mm" value="4.8">4.8 mm</quantity>
|
||
); protibial keel with rounded apex and the basal area of the protibia with a wider keel (
|
||
<figureCitation id="1369F3A06667AB51B61F63F1FC183DFB" box="[883,974,1692,1719]" captionStart="Figs 16 – 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 – 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Fig. 16</figureCitation>
|
||
); 8th antennomere more than 1.4 × longer than wide and 4th antennomere equal or almost equal in length to 5th and 7th antennomeres in both sexes …………………………………………………………………
|
||
<taxonomicName id="4C5294A66667AB51B107638FFAC53DB1" ID-CoL="6YX6R" authority="Ana Sofa P.S. Reboleira & Javier Fresneda & José Maria Salgado, 2017" authorityName="Ana Sofa P.S. Reboleira & Javier Fresneda & José Maria Salgado" authorityYear="2017" box="[1131,1299,1762,1789]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="algarvensis" status="sp. nov.">
|
||
<emphasis id="B92633376667AB51B107638FFAC53DB1" bold="true" box="[1131,1299,1762,1789]" italics="true" pageId="2" pageNumber="3">S. algarvensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2158E4C6667AB51B077638EFAA13DB1" box="[1307,1399,1763,1789]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A6F7806667AB51B5D16271FAA03CAA" pageId="2" pageNumber="3">
|
||
<paragraph id="8BEDEF256667AB51B5D16271FAA03C31" blockId="2.[189,1399,1259,2023]" pageId="2" pageNumber="3">
|
||
<keyLead id="30A34C106667AB51B5D16271FAA03C31" pageId="2" pageNumber="3">
|
||
4. Upper margin of protibial keel long, linear medially (
|
||
<figureCitation id="1369F3A06667AB51B61D6271FC1D3C7A" box="[881,971,1820,1846]" captionStart="Figs 16 – 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 – 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Fig. 17</figureCitation>
|
||
); 8th antennomere slightly less than 1.5 × longer than wide, and 4th, 5th and 7th antennomeres of unequal length in both sexes …… ………………………………………………………………………
|
||
<treatmentCitation id="0AF3C9346667AB51B16E620EFAA03C31" author="Kraatz" authority="(Kraatz, 1870)" authorityName="Kraatz" authorityYear="1870" baseAuthorityName="Kraatz" baseAuthorityYear="1870" box="[1026,1398,1890,1917]" class="Insecta" family="Leiodidae" firstAuthor="Kraatz" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" pagination="98 - 99" phylum="Arthropoda" rank="species" refId="ref12372" refString="Kraatz G. 1870. Choleva angusticollis Kraatz n. sp. In: Heyden L. von (ed.) Entomologische Reise nach dem sudlichen Spanien der Sierra Guadarrama und Sierra Morena, Portugal und den Cantabrischen Gebirgen: 98 - 99. Entomologischer Verein in Berlin, Berlin." species="angusticollis" treatmentTaxon="S. angusticollis" type="book chapter" year="1870">
|
||
<taxonomicName id="4C5294A66667AB51B16E620EFAA03C31" ID-CoL="4YHYV" authority="(Kraatz, 1870)" baseAuthorityName="Kraatz" baseAuthorityYear="1870" box="[1026,1398,1890,1917]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="angusticollis">
|
||
<emphasis id="B92633376667AB51B16E620EFB6B3C31" bold="true" box="[1026,1213,1890,1917]" italics="true" pageId="2" pageNumber="3">S. angusticollis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC392D46667AB51B1BC620EFAB83C31" author="Kraatz" box="[1232,1390,1891,1917]" firstAuthor="Kraatz" pageId="2" pageNumber="3" pagination="98 - 99" refId="ref12372" refString="Kraatz G. 1870. Choleva angusticollis Kraatz n. sp. In: Heyden L. von (ed.) Entomologische Reise nach dem sudlichen Spanien der Sierra Guadarrama und Sierra Morena, Portugal und den Cantabrischen Gebirgen: 98 - 99. Entomologischer Verein in Berlin, Berlin." type="book chapter" year="1870">Kraatz, 1870</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256667AB51B5AD62EBFAA03CAA" blockId="2.[189,1399,1259,2023]" pageId="2" pageNumber="3">
|
||
<keyLead id="30A34C106667AB51B5AD62EBFAA03CAA" pageId="2" pageNumber="3">
|
||
– Upper margin of protibial keel short, curved medially (
|
||
<figureCitation id="1369F3A06667AB51B61362EBFC0E3CEC" box="[895,984,1926,1952]" captionStart="Figs 16 – 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 – 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Fig. 19</figureCitation>
|
||
); 8th antennomere more than 1.5 × longer than wide and 4th, 5th and 7th antennomeres of almost equal length in both sexes …………… ……………………………………………………………………………
|
||
<treatmentCitation id="0AF3C9346667AB51B15162A1FAA03CAA" author="Jeannel" authority="Jeannel, 1922" authorityName="Ana Sofa P.S. Reboleira & Javier Fresneda & José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[1085,1398,1996,2023]" class="Insecta" family="Leiodidae" firstAuthor="Jeannel" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" pagination="1 - 98" phylum="Arthropoda" rank="species" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." species="breuili" status="sp. nov." treatmentTaxon="S. breuili" type="journal article" year="1922">
|
||
<taxonomicName id="4C5294A66667AB51B15162A1FAA03CAA" ID-CoL="4YHYX" authority="Jeannel, 1922" authorityName="Ana Sofa P.S. Reboleira & Javier Fresneda & José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[1085,1398,1996,2023]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="breuili" status="sp. nov.">
|
||
<emphasis id="B92633376667AB51B15162A1FB673CAB" bold="true" box="[1085,1201,1996,2023]" italics="true" pageId="2" pageNumber="3">S. breuili</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC392D46667AB51B1A962A1FAB83CAA" author="Jeannel" box="[1221,1390,1996,2022]" firstAuthor="Jeannel" pageId="2" pageNumber="3" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel, 1922</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |