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<figureCitation id="1369F3A06666AB50B7EC643FFD1F3A20" box="[640,713,338,364]" captionStart="Figs 1 5" captionStartId="3.[189,241,1892,1918]" captionTargetBox="[189,1395,1323,1850]" captionTargetId="figure@3.[189,1398,1323,1850]" captionTargetPageId="3" captionText="Figs 1 5. Speonemadus of the escalera i- group habitus. 1. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 2. S. angusticollis (Kraatz, 1870) (Rute, Spain). 3. S. bolivari (Jeannel, 1922) (Ardales, Spain). 4. S. breuili (Jeannel, 1922) (Motillas, Spain). 5. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 1 mm." httpUri="https://zenodo.org/record/889348/files/figure.png" pageId="3" pageNumber="4">Figs 1</figureCitation>
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<figureCitation id="1369F3A06666AB50B7BB643FFCD03A20" box="[727,774,338,364]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="3" pageNumber="4">67</figureCitation>
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<figureCitation id="1369F3A06666AB50B67E643FFCE43A20" box="[786,818,338,364]" captionStart="Figs 16 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="3" pageNumber="4">16</figureCitation>
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<figureCitation id="1369F3A06666AB50B652643FFC883A20" box="[830,862,338,364]" captionStart="Figs 21 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="3" pageNumber="4">21</figureCitation>
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<figureCitation id="1369F3A06666AB50B606643FFC653A20" box="[874,947,338,364]" captionStart="Figs 26 - 31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26 - 31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the frst tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 26 27 = 100 μm, 28 31 = 10 μm." httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="3" pageNumber="4">2631</figureCitation>
<figureCitation id="1369F3A06666AB50B606643FFC653A20" box="[874,947,338,364]" captionStart="Figs 26-31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26-31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the first tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="3" pageNumber="4">2631</figureCitation>
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@ -112,14 +112,14 @@ with a total length of 4.0
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<collectingMunicipality id="6B89755F6666AB50B77F6643FD8C3804" box="[531,602,814,840]" pageId="3" pageNumber="4">Loulé</collectingMunicipality>
,
<collectingCounty id="628C97A96666AB50B71C6642FC4F3804" box="[624,921,814,841]" pageId="3" pageNumber="4">Gruta do Vale Telheiro</collectingCounty>
<location id="8E8DB9FE6666AB50B71C6642FC4F3804" LSID="urn:lsid:plazi:treatment:03FB5E336666AB59B7216466FD873A20:8E8DB9FE6666AB50B71C6642FC4F3804" box="[624,921,814,841]" country="Portugal" county="Gruta do Vale Telheiro" latitude="37.170555" longLatPrecision="19" longitude="-8.035001" municipality="Loule" name="Gruta do Vale Telheiro" pageId="3" pageNumber="4" stateProvince="Algarve">Gruta do Vale Telheiro</location>
, coordinates: WGS 84:
<geoCoordinate id="EE6689E26666AB50B1B96640FAA43805" box="[1237,1394,813,841]" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="15" value="37.170555">37º1014″ N</geoCoordinate>
,
@ -132,7 +132,9 @@ with a total length of 4.0
<collectorName id="26A78AF36666AB50B776663FFD7D3820" box="[538,683,850,876]" pageId="3" pageNumber="4">S. Reboleira</collectorName>
leg. (
<collectionCode id="ED4377E06666AB50B780663FFC953820" box="[748,835,850,876]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="3" pageNumber="4">ZMUC</collectionCode>
number 00036257).
number
<specimenCode id="DBF4475E6666AB50B6C7663CFBF03820" box="[939,1062,849,876]" pageId="3" pageNumber="4">00036257</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEDEF256666AB50B58666FEFEB138E1" blockId="3.[189,1399,915,1188]" box="[234,359,915,941]" pageId="3" pageNumber="4">
@ -141,18 +143,22 @@ number 00036257).
</heading>
</paragraph>
<paragraph id="8BEDEF256666AB50B5D166DBFB333FE8" blockId="3.[189,1399,915,1188]" pageId="3" pageNumber="4">
<materialsCitation id="3B3AE5786666AB50B5D166DBFD98389D" ID-GBIF-Occurrence="1609370440" box="[189,590,948,977]" country="Portugal" location="PORTUGAL" pageId="3" pageNumber="4" specimenCount="30" specimenCount-female="30">
<materialsCitation id="3B3AE5786666AB50B5D166DBFD98389D" ID-GBIF-Occurrence="1609370440" box="[189,590,948,977]" country="Portugal" location="PORTUGAL" pageId="3" pageNumber="4" specimenCount="68" specimenCount-female="30" specimenCount-male="38">
<collectingCountry id="F345AFB56666AB50B5D166DBFE8B389C" box="[189,349,950,976]" name="Portugal" pageId="3" pageNumber="4">PORTUGAL</collectingCountry>
: 38 ♁♁ and
<specimenCount id="9D5424AC6666AB50B49666DBFD98389D" box="[506,590,950,977]" pageId="3" pageNumber="4" type="female">30 ♀♀</specimenCount>
:
<specimenCount id="9D5424AC6666AB50B40566DBFE6A389C" box="[361,444,948,976]" count="38" pageId="3" pageNumber="4" type="male">38 ♂♂</specimenCount>
and
<specimenCount id="9D5424AC6666AB50B49666DBFD98389D" box="[506,590,950,977]" count="30" pageId="3" pageNumber="4" type="female">30 ♀♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B3AE5786666AB50B73266DAFBF9389D" ID-GBIF-Occurrence="1609370451" box="[606,1071,948,977]" country="Portugal" location="PORTUGAL" pageId="3" pageNumber="4" specimenCount="18" specimenCount-female="18">
same data as holotype; 11 ♁♁,
<specimenCount id="9D5424AC6666AB50B6B766DBFBF9389D" box="[987,1071,950,977]" pageId="3" pageNumber="4" type="female">18 ♀♀</specimenCount>
<materialsCitation id="3B3AE5786666AB50B73266DAFBF9389D" ID-GBIF-Occurrence="1609370451" box="[606,1071,948,977]" country="Portugal" location="PORTUGAL" pageId="3" pageNumber="4" specimenCount="29" specimenCount-female="18" specimenCount-male="11" typeStatus="paratype">
same data as holotype;
<specimenCount id="9D5424AC6666AB50B61466DBFC1D389C" box="[888,971,948,976]" count="11" pageId="3" pageNumber="4" type="male">11 ♂♂</specimenCount>
,
<specimenCount id="9D5424AC6666AB50B6B766DBFBF9389D" box="[987,1071,950,977]" count="18" pageId="3" pageNumber="4" type="female">18 ♀♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B3AE5786666AB50B15366DAFDFB38BF" ID-GBIF-Occurrence="1609370455" collectingDate="2009-03-13" collectorName="S. Reboleira" country="Portugal" location="PORTUGAL" pageId="3" pageNumber="4" specimenCount="1">
<materialsCitation id="3B3AE5786666AB50B15366DAFDFB38BF" ID-GBIF-Occurrence="1609370455" collectingDate="2009-03-13" collectorName="S. Reboleira" country="Portugal" location="PORTUGAL" pageId="3" pageNumber="4" specimenCount="1" typeStatus="paratype">
same locality as holotype,
<date id="FFECC9E56666AB50B5D166B7FE8B38B8" box="[189,349,986,1012]" pageId="3" pageNumber="4" value="2009-03-13">
<collectingDate id="EFA8300D6666AB50B5D166B7FE8B38B8" box="[189,349,986,1012]" pageId="3" pageNumber="4" value="2009-03-13">13 Mar. 2009</collectingDate>
@ -161,18 +167,30 @@ same locality as holotype,
<collectorName id="26A78AF36666AB50B40466B7FE2E38BF" box="[360,504,985,1012]" pageId="3" pageNumber="4">S. Reboleira</collectorName>
leg.
</materialsCitation>
; 53 ♁♁, 63 ♀♀,
<date id="FFECC9E56666AB50B79B66B7FC4038B8" box="[759,918,986,1012]" pageId="3" pageNumber="4" value="2009-05-24">24 May 2009</date>
, S. Reboleira leg.;
<materialsCitation id="3B3AE5786666AB50B11E66B7FBE23F5B" ID-GBIF-Occurrence="1609370453" collectingDate="2009-03-15" collectorName="S. Reboleira" country="Portugal" county="Algarao do Remexido" location="Algarve" municipality="Sao Bras de Alportel" pageId="3" pageNumber="4" specimenCount="9" specimenCount-female="9">
9 ♁♁,
<specimenCount id="9D5424AC6666AB50B1AE66B7FAD338B8" box="[1218,1285,986,1012]" pageId="3" pageNumber="4" type="female">9 ♀♀</specimenCount>
;
<materialsCitation id="3B3AE5786666AB50B75666B4FBB038BF" box="[570,1126,984,1012]" collectingDate="2009-05-24" collectorName="S. Reboleira" pageId="3" pageNumber="4" specimenCount="116" specimenCount-female="63" specimenCount-male="53" typeStatus="paratype">
<specimenCount id="9D5424AC6666AB50B75666B4FD5D38B8" box="[570,651,984,1012]" count="53" pageId="3" pageNumber="4" type="male">53 ♂♂</specimenCount>
,
<location id="8E8DB9FE6666AB50B07D66B4FAA438BF" LSID="urn:lsid:plazi:treatment:03FB5E336666AB59B7216466FD873A20:8E8DB9FE6666AB50B07D66B4FAA438BF" box="[1297,1394,985,1011]" country="Portugal" county="Algarao do Remexido" municipality="Sao Bras de Alportel" name="Algarve" pageId="3" pageNumber="4">Algarve</location>
<specimenCount id="9D5424AC6666AB50B7F466B4FD3F38B8" box="[664,745,985,1012]" count="63" pageId="3" pageNumber="4" type="female">63 ♀♀</specimenCount>
,
<date id="FFECC9E56666AB50B79B66B7FC4038B8" box="[759,918,986,1012]" pageId="3" pageNumber="4" value="2009-05-24">
<collectingDate id="EFA8300D6666AB50B79B66B7FC4038B8" box="[759,918,986,1012]" pageId="3" pageNumber="4" value="2009-05-24">24 May 2009</collectingDate>
</date>
,
<collectorName id="26A78AF36666AB50B6CD66B7FBE738BF" box="[929,1073,985,1012]" pageId="3" pageNumber="4">S. Reboleira</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B3AE5786666AB50B11E66B7FBE23F5B" ID-GBIF-Occurrence="1609370453" collectingDate="2009-03-15" collectorName="S. Reboleira" country="Portugal" county="Algarao do Remexido" location="Algarao do Remexido" municipality="Sao Bras de Alportel" pageId="3" pageNumber="4" specimenCount="18" specimenCount-female="9" specimenCount-male="9" stateProvince="Algarve" typeStatus="paratype">
<specimenCount id="9D5424AC6666AB50B11E66B7FB6338B8" box="[1138,1205,984,1012]" count="9" pageId="3" pageNumber="4" type="male">9 ♂♂</specimenCount>
,
<specimenCount id="9D5424AC6666AB50B1AE66B7FAD338B8" box="[1218,1285,986,1012]" count="9" pageId="3" pageNumber="4" type="female">9 ♀♀</specimenCount>
,
<collectingRegion id="499621C76666AB50B07D66B4FAA438BF" box="[1297,1394,985,1011]" pageId="3" pageNumber="4">Algarve</collectingRegion>
,
<collectingMunicipality id="6B89755F6666AB50B5D16690FE663F5B" box="[189,432,1021,1047]" pageId="3" pageNumber="4">São Brás de Alportel</collectingMunicipality>
,
<collectingCounty id="628C97A96666AB50B4D56690FD6B3F5B" box="[441,701,1021,1047]" pageId="3" pageNumber="4">Algarão do Remexido</collectingCounty>
<location id="8E8DB9FE6666AB50B4D56690FD6B3F5B" LSID="urn:lsid:plazi:treatment:03FB5E336666AB59B7216466FD873A20:8E8DB9FE6666AB50B4D56690FD6B3F5B" box="[441,701,1021,1047]" country="Portugal" county="Algarao do Remexido" municipality="Sao Bras de Alportel" name="Algarao do Remexido" pageId="3" pageNumber="4" stateProvince="Algarve">Algarão do Remexido</location>
,
<date id="FFECC9E56666AB50B7AB6691FCB33F5B" box="[711,869,1020,1047]" pageId="3" pageNumber="4" value="2009-03-15">
<collectingDate id="EFA8300D6666AB50B7AB6691FCB33F5B" box="[711,869,1020,1047]" pageId="3" pageNumber="4" value="2009-03-15">15 Mar. 2009</collectingDate>
@ -182,8 +200,8 @@ leg.
leg.
</materialsCitation>
;
<materialsCitation id="3B3AE5786666AB50B1536690FE053F76" ID-GBIF-Occurrence="1609370445" collectingDate="2009-05-23" collectorName="S. Reboleira" country="Portugal" county="Algarao do Remexido" location="Algarve" municipality="Sao Bras de Alportel" pageId="3" pageNumber="4" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D5424AC6666AB50B1536690FBBC3F5B" box="[1087,1130,1021,1047]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
<materialsCitation id="3B3AE5786666AB50B1536690FE053F76" ID-GBIF-Occurrence="1609370445" collectingDate="2009-05-23" collectorName="S. Reboleira" country="Portugal" county="Algarao do Remexido" location="Algarve" municipality="Sao Bras de Alportel" pageId="3" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="Algarve" typeStatus="paratype">
<specimenCount id="9D5424AC6666AB50B1536690FBBC3F5B" box="[1087,1130,1021,1047]" count="1" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
, same locality,
<collectingDate id="EFA8300D6666AB50B0716690FF2D3F76" pageId="3" pageNumber="4" value="2009-05-23">
<date id="FFECC9E56666AB50B0716690FF2D3F76" pageId="3" pageNumber="4" value="2009-05-23">23 May 2009</date>
@ -193,9 +211,10 @@ leg.
leg.
</materialsCitation>
;
<materialsCitation id="3B3AE5786666AB50B48D614DFB073F76" ID-GBIF-Occurrence="1609370449" box="[481,1233,1054,1083]" collectingDate="2009-05-25" collectorName="S. Reboleira" country="Portugal" county="Algarao do Remexido" location="Algarve" municipality="Sao Bras de Alportel" pageId="3" pageNumber="4" specimenCount="27" specimenCount-female="27">
31 ♁♁,
<specimenCount id="9D5424AC6666AB50B729614DFD4E3F76" box="[581,664,1056,1082]" pageId="3" pageNumber="4" type="female">27 ♀♀</specimenCount>
<materialsCitation id="3B3AE5786666AB50B48D614DFB073F76" ID-GBIF-Occurrence="1609370449" box="[481,1233,1054,1083]" collectingDate="2009-05-25" collectorName="S. Reboleira" country="Portugal" county="Algarao do Remexido" location="Algarve" municipality="Sao Bras de Alportel" pageId="3" pageNumber="4" specimenCount="58" specimenCount-female="27" specimenCount-male="31" stateProvince="Algarve" typeStatus="paratype">
<specimenCount id="9D5424AC6666AB50B48D614DFDE33F76" box="[481,565,1054,1082]" count="31" pageId="3" pageNumber="4" type="male">31 ♂♂</specimenCount>
,
<specimenCount id="9D5424AC6666AB50B729614DFD4E3F76" box="[581,664,1056,1082]" count="27" pageId="3" pageNumber="4" type="female">27 ♀♀</specimenCount>
, same locality,
<date id="FFECC9E56666AB50B638614DFC2E3F76" box="[852,1016,1056,1083]" pageId="3" pageNumber="4" value="2009-05-25">
<collectingDate id="EFA8300D6666AB50B638614DFC2E3F76" box="[852,1016,1056,1083]" pageId="3" pageNumber="4" value="2009-05-25">25 May 2009</collectingDate>
@ -205,10 +224,10 @@ leg.
leg.
</materialsCitation>
;
<materialsCitation id="3B3AE5786666AB50B1B3614DFD003F11" ID-GBIF-Occurrence="1609370437" collectingDate="2009-09-05" collectorName="S. Reboleira" country="Portugal" county="Algarao do Remexido" location="Algarve" municipality="Sao Bras de Alportel" pageId="3" pageNumber="4" specimenCount="2" specimenCount-female="1" specimenCount-pupa="1">
<specimenCount id="9D5424AC6666AB50B1B3614DFAE53F76" box="[1247,1331,1056,1082]" pageId="3" pageNumber="4" type="pupa">1 pupa</specimenCount>
<materialsCitation id="3B3AE5786666AB50B1B3614DFD003F11" ID-GBIF-Occurrence="1609370437" collectingDate="2009-09-05" collectorName="S. Reboleira" country="Portugal" county="Algarao do Remexido" location="Algarve" municipality="Sao Bras de Alportel" pageId="3" pageNumber="4" specimenCount="2" specimenCount-female="1" specimenCount-pupa="1" stateProvince="Algarve" typeStatus="paratype">
<specimenCount id="9D5424AC6666AB50B1B3614DFAE53F76" box="[1247,1331,1056,1082]" count="1" pageId="3" pageNumber="4" type="pupa">1 pupa</specimenCount>
,
<specimenCount id="9D5424AC6666AB50B02D614DFAB93F76" box="[1345,1391,1056,1082]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
<specimenCount id="9D5424AC6666AB50B02D614DFAB93F76" box="[1345,1391,1056,1082]" count="1" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
, same locality,
<date id="FFECC9E56666AB50B404612EFDD73F11" box="[360,513,1091,1118]" pageId="3" pageNumber="4" value="2009-09-05">
<collectingDate id="EFA8300D6666AB50B404612EFDD73F11" box="[360,513,1091,1118]" pageId="3" pageNumber="4" value="2009-09-05">5 Sept. 2009</collectingDate>
@ -218,9 +237,10 @@ leg.
leg.
</materialsCitation>
;
<materialsCitation id="3B3AE5786666AB50B788612EFF3D3FCC" ID-GBIF-Occurrence="1609370442" collectingDate="2009-12-29" collectorName="S. Reboleira" country="Portugal" county="Algarao do Remexido" location="Algarve" municipality="Sao Bras de Alportel" pageId="3" pageNumber="4" specimenCount="3" specimenCount-female="3">
3 ♁♁,
<specimenCount id="9D5424AC6666AB50B65B612EFCAD3F12" box="[823,891,1091,1118]" pageId="3" pageNumber="4" type="female">3 ♀♀</specimenCount>
<materialsCitation id="3B3AE5786666AB50B788612EFF3D3FCC" ID-GBIF-Occurrence="1609370442" collectingDate="2009-12-29" collectorName="S. Reboleira" country="Portugal" county="Algarao do Remexido" location="Algarve" municipality="Sao Bras de Alportel" pageId="3" pageNumber="4" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Algarve" typeStatus="paratype">
<specimenCount id="9D5424AC6666AB50B788612EFCFE3F11" box="[740,808,1089,1117]" count="3" pageId="3" pageNumber="4" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D5424AC6666AB50B65B612EFCAD3F12" box="[823,891,1091,1118]" count="3" pageId="3" pageNumber="4" type="female">3 ♀♀</specimenCount>
, same locality,
<date id="FFECC9E56666AB50B159612EFB0E3F11" box="[1077,1240,1091,1118]" pageId="3" pageNumber="4" value="2009-12-29">
<collectingDate id="EFA8300D6666AB50B159612EFB0E3F11" box="[1077,1240,1091,1118]" pageId="3" pageNumber="4" value="2009-12-29">29 Dec. 2009</collectingDate>
@ -230,31 +250,34 @@ leg.
leg.
</materialsCitation>
;
<materialsCitation id="3B3AE5786666AB50B597610BFAFE3FCC" ID-GBIF-Occurrence="1609370456" box="[251,1320,1124,1153]" collectingDate="2009-12-29" collectorName="S. Reboleira" country="Portugal" county="Gruta da Senhora" location="Algarve" municipality="Moncarapacho" pageId="3" pageNumber="4" specimenCount="4" specimenCount-female="4">
<specimenCount id="9D5424AC6666AB50B597610BFE973FCD" box="[251,321,1126,1153]" pageId="3" pageNumber="4" type="female">4 ♀♀</specimenCount>
<materialsCitation id="3B3AE5786666AB50B597610BFAFE3FCC" ID-GBIF-Occurrence="1609370456" box="[251,1320,1124,1153]" collectingDate="2009-12-29" collectorName="S. Reboleira" country="Portugal" county="Gruta da Senhora" location="Gruta da Senhora" municipality="Moncarapacho" pageId="3" pageNumber="4" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Algarve" typeStatus="paratype">
<specimenCount id="9D5424AC6666AB50B597610BFE973FCD" box="[251,321,1126,1153]" count="4" pageId="3" pageNumber="4" type="female">4 ♀♀</specimenCount>
,
<location id="8E8DB9FE6666AB50B43C610BFE673FCC" LSID="urn:lsid:plazi:treatment:03FB5E336666AB59B7216466FD873A20:8E8DB9FE6666AB50B43C610BFE673FCC" box="[336,433,1126,1152]" country="Portugal" county="Gruta da Senhora" municipality="Moncarapacho" name="Algarve" pageId="3" pageNumber="4">Algarve</location>
<collectingRegion id="499621C76666AB50B43C610BFE673FCC" box="[336,433,1126,1152]" pageId="3" pageNumber="4">Algarve</collectingRegion>
,
<collectingMunicipality id="6B89755F6666AB50B4AD610BFDA43FCC" box="[449,626,1126,1152]" pageId="3" pageNumber="4">Moncarapacho</collectingMunicipality>
,
<collectingCounty id="628C97A96666AB50B7ED610BFC8E3FCC" box="[641,856,1126,1152]" pageId="3" pageNumber="4">Gruta da Senhora</collectingCounty>
<location id="8E8DB9FE6666AB50B7ED610BFC8E3FCC" LSID="urn:lsid:plazi:treatment:03FB5E336666AB59B7216466FD873A20:8E8DB9FE6666AB50B7ED610BFC8E3FCC" box="[641,856,1126,1152]" country="Portugal" county="Gruta da Senhora" municipality="Moncarapacho" name="Gruta da Senhora" pageId="3" pageNumber="4" stateProvince="Algarve">Gruta da Senhora</location>
,
<date id="FFECC9E56666AB50B60B610BFBD83FCC" box="[871,1038,1126,1153]" pageId="3" pageNumber="4" value="2009-12-29">
<collectingDate id="EFA8300D6666AB50B60B610BFBD83FCC" box="[871,1038,1126,1153]" pageId="3" pageNumber="4" value="2009-12-29">29 Dec. 2009</collectingDate>
</date>
,
<collectorName id="26A78AF36666AB50B170610BFB673FCC" box="[1052,1201,1126,1152]" pageId="3" pageNumber="4">S. Reboleira</collectorName>
leg.; 1 ♁
leg.;
<specimenCount id="9D5424AC6666AB50B195610BFAFE3FCC" box="[1273,1320,1124,1152]" count="1" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B3AE5786666AB50B056610AFB333FE8" ID-GBIF-Occurrence="1609370454" collectingDate="2009-05-24" collectionCode="CASR, CJMS, CZULE, CFL, CPB, ZMUC" collectorName="S. Reboleira" country="Portugal" county="Gruta da Senhora" location="Algarve" municipality="Moncarapacho" pageId="3" pageNumber="4" specimenCount="1">
<materialsCitation id="3B3AE5786666AB50B056610AFB333FE8" ID-GBIF-Occurrence="1609370454" collectingDate="2009-05-24" collectionCode="SR, CASR, CJMS, CZULE, CFL, CPB, ZMUC" collectorName="S. Reboleira" country="Portugal" county="Gruta da Senhora" location="Algarve" municipality="Moncarapacho" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Algarve" typeStatus="paratype">
same locality,
<date id="FFECC9E56666AB50B44E61E7FE173FE8" box="[290,449,1162,1188]" pageId="3" pageNumber="4" value="2009-05-24">
<collectingDate id="EFA8300D6666AB50B44E61E7FE173FE8" box="[290,449,1162,1188]" pageId="3" pageNumber="4" value="2009-05-24">24 May 2009</collectingDate>
</date>
,
<collectorName id="26A78AF36666AB50B4A161E7FD883FEF" box="[461,606,1161,1188]" pageId="3" pageNumber="4">S. Reboleira</collectorName>
leg. (SR,
leg. (
<collectionCode id="ED4377E06666AB50B7CC61E7FD113FE8" box="[672,711,1162,1188]" pageId="3" pageNumber="4">SR</collectionCode>
,
<collectionCode id="ED4377E06666AB50B7B861E7FCF23FE8" box="[724,804,1162,1188]" pageId="3" pageNumber="4">CASR</collectionCode>
,
<collectionCode id="ED4377E06666AB50B64361E7FCAD3FE8" box="[815,891,1162,1188]" pageId="3" pageNumber="4">CJMS</collectionCode>
@ -265,7 +288,7 @@ leg. (SR,
,
<collectionCode id="ED4377E06666AB50B15561E7FBA53FE8" box="[1081,1139,1162,1188]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13713" name="National Institute for the Control of Pharmaceutical and Biological Products" pageId="3" pageNumber="4">CPB</collectionCode>
,
<collectionCode id="ED4377E06666AB50B11361E7FB0F3FE8" box="[1151,1241,1162,1188]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="3" pageNumber="4">ZMUC</collectionCode>
<collectionCode id="ED4377E06666AB50B11361E7FB0F3FE8" box="[1151,1241,1162,1188]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="3" pageNumber="4">ZMUC</collectionCode>
).
</materialsCitation>
</paragraph>
@ -402,7 +425,7 @@ of the
</th>
</tr>
<tr id="3562ED67666154ACB5AD64BCFABE3AA7" box="[193,1384,465,491]" gridrow="2" pageId="4" pageNumber="5">
<th id="76B3841B666154ACB5AD64BCFE973AA7" box="[193,321,465,491]" gridcol="0" gridrow="2" pageId="4" pageNumber="5">holotype </th>
<th id="76B3841B666154ACB5AD64BCFE973AA7" box="[193,321,465,491]" gridcol="0" gridrow="2" pageId="4" pageNumber="5">holotype </th>
<td id="76B3841B666154ACB40D64BCFEAC3AA7" box="[353,378,465,491]" gridcol="1" gridrow="2" pageId="4" pageNumber="5">L</td>
<td id="76B3841B666154ACB4CD64BCFE0B3AA7" box="[417,477,465,491]" gridcol="2" gridrow="2" pageId="4" pageNumber="5">0.189</td>
<td id="76B3841B666154ACB49064BCFDEE3AA7" box="[508,568,465,491]" gridcol="3" gridrow="2" pageId="4" pageNumber="5">0.195</td>
@ -431,7 +454,7 @@ of the
<td id="76B3841B666154ACB0406499FABE3940" box="[1324,1384,500,524]" gridcol="12" gridrow="3" pageId="4" pageNumber="5">0.090</td>
</tr>
<tr id="3562ED67666154ACB5AD6779FABE3962" box="[193,1384,532,558]" gridrow="4" pageId="4" pageNumber="5">
<th id="76B3841B666154ACB5AD6779FE973962" box="[193,321,532,558]" gridcol="0" gridrow="4" pageId="4" pageNumber="5">paratypes </th>
<th id="76B3841B666154ACB5AD6779FE973962" box="[193,321,532,558]" gridcol="0" gridrow="4" pageId="4" pageNumber="5">paratypes </th>
<td id="76B3841B666154ACB40D6779FEAC3962" box="[353,378,532,558]" gridcol="1" gridrow="4" pageId="4" pageNumber="5">L</td>
<td id="76B3841B666154ACB4CD6779FE0B3962" box="[417,477,532,558]" gridcol="2" gridrow="4" pageId="4" pageNumber="5">0.191</td>
<td id="76B3841B666154ACB4906779FDEE3962" box="[508,568,532,558]" gridcol="3" gridrow="4" pageId="4" pageNumber="5">0.195</td>
@ -501,7 +524,7 @@ of the
</th>
</tr>
<tr id="3562ED67666154ACB5AD67A5FABE39AE" box="[193,1384,712,738]" gridrow="9" pageId="4" pageNumber="5">
<th id="76B3841B666154ACB5AD67A5FE9739AE" box="[193,321,712,738]" gridcol="0" gridrow="9" pageId="4" pageNumber="5"></th>
<th id="76B3841B666154ACB5AD67A5FE9739AE" box="[193,321,712,738]" gridcol="0" gridrow="9" pageId="4" pageNumber="5"></th>
<td id="76B3841B666154ACB40D67A5FEAC39AE" box="[353,378,712,738]" gridcol="1" gridrow="9" pageId="4" pageNumber="5">L</td>
<td id="76B3841B666154ACB4CD67A5FE0B39AE" box="[417,477,712,738]" gridcol="2" gridrow="9" pageId="4" pageNumber="5">0.182</td>
<td id="76B3841B666154ACB49067A5FDEE39AE" box="[508,568,712,738]" gridcol="3" gridrow="9" pageId="4" pageNumber="5">0.208</td>
@ -571,7 +594,7 @@ of the
</th>
</tr>
<tr id="3562ED67666154ACB5AD6611FABE38DA" box="[193,1384,892,918]" gridrow="14" pageId="4" pageNumber="5">
<th id="76B3841B666154ACB5AD6611FE9738DA" box="[193,321,892,918]" gridcol="0" gridrow="14" pageId="4" pageNumber="5"></th>
<th id="76B3841B666154ACB5AD6611FE9738DA" box="[193,321,892,918]" gridcol="0" gridrow="14" pageId="4" pageNumber="5"></th>
<td id="76B3841B666154ACB40D6611FEAC38DA" box="[353,378,892,918]" gridcol="1" gridrow="14" pageId="4" pageNumber="5">L</td>
<td id="76B3841B666154ACB4CD6611FE0B38DA" box="[417,477,892,918]" gridcol="2" gridrow="14" pageId="4" pageNumber="5">0.174</td>
<td id="76B3841B666154ACB4906611FDEE38DA" box="[508,568,892,918]" gridcol="3" gridrow="14" pageId="4" pageNumber="5">0.177</td>
@ -641,7 +664,7 @@ of the
</th>
</tr>
<tr id="3562ED67666154ACB5AD615DFABE3F06" box="[193,1384,1072,1098]" gridrow="19" pageId="4" pageNumber="5">
<th id="76B3841B666154ACB5AD615DFE973F06" box="[193,321,1072,1098]" gridcol="0" gridrow="19" pageId="4" pageNumber="5"></th>
<th id="76B3841B666154ACB5AD615DFE973F06" box="[193,321,1072,1098]" gridcol="0" gridrow="19" pageId="4" pageNumber="5"></th>
<td id="76B3841B666154ACB40D615DFEAC3F06" box="[353,378,1072,1098]" gridcol="1" gridrow="19" pageId="4" pageNumber="5">L</td>
<td id="76B3841B666154ACB4CD615DFE0B3F06" box="[417,477,1072,1098]" gridcol="2" gridrow="19" pageId="4" pageNumber="5">0.182</td>
<td id="76B3841B666154ACB490615DFDEE3F06" box="[508,568,1072,1098]" gridcol="3" gridrow="19" pageId="4" pageNumber="5">0.192</td>
@ -711,7 +734,7 @@ of the
</th>
</tr>
<tr id="3562ED67666154ACB5AD618EFABE3FB1" box="[193,1384,1251,1277]" gridrow="24" pageId="4" pageNumber="5">
<th id="76B3841B666154ACB5AD618EFE973FB1" box="[193,321,1251,1277]" gridcol="0" gridrow="24" pageId="4" pageNumber="5"></th>
<th id="76B3841B666154ACB5AD618EFE973FB1" box="[193,321,1251,1277]" gridcol="0" gridrow="24" pageId="4" pageNumber="5"></th>
<td id="76B3841B666154ACB40D618EFEAC3FB1" box="[353,378,1251,1277]" gridcol="1" gridrow="24" pageId="4" pageNumber="5">L</td>
<td id="76B3841B666154ACB4CD618EFE0B3FB1" box="[417,477,1251,1277]" gridcol="2" gridrow="24" pageId="4" pageNumber="5">0.190</td>
<td id="76B3841B666154ACB490618EFDEE3FB1" box="[508,568,1251,1277]" gridcol="3" gridrow="24" pageId="4" pageNumber="5">0.194</td>
@ -788,41 +811,41 @@ BODY. Length
(females) (
<figureCitation id="1369F3A06661AB57B486609CFDE23D40" box="[490,564,1521,1548]" captionStart="Figs 1 5" captionStartId="3.[189,241,1892,1918]" captionTargetBox="[189,1395,1323,1850]" captionTargetId="figure@3.[189,1398,1323,1850]" captionTargetPageId="3" captionText="Figs 1 5. Speonemadus of the escalera i- group habitus. 1. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 2. S. angusticollis (Kraatz, 1870) (Rute, Spain). 3. S. bolivari (Jeannel, 1922) (Ardales, Spain). 4. S. breuili (Jeannel, 1922) (Motillas, Spain). 5. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 1 mm." httpUri="https://zenodo.org/record/889348/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
,
<figureCitation id="1369F3A06661AB57B72D609CFD463D40" box="[577,656,1521,1548]" captionStart="Figs 26 - 31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26 - 31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the frst tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 26 27 = 100 μm, 28 31 = 10 μm." httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="4" pageNumber="5">2627</figureCitation>
). Color reddish brown, darker in central area of head and lighter on antennae and legs. Integument evenly covered with a yellowish pubescence, thin, short and fattened, slightly raised in forehead area and clypeus. Head retractable; head sculpture composed of a tight mesh of small dots. Eyes developed with pigmented ommatidia (
<figureCitation id="1369F3A06661AB57B60C6336FC603D39" box="[864,950,1627,1653]" captionStart="Figs 26 - 31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26 - 31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the frst tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 26 27 = 100 μm, 28 31 = 10 μm." httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="4" pageNumber="5">Fig. 27</figureCitation>
<figureCitation id="1369F3A06661AB57B72D609CFD463D40" box="[577,656,1521,1548]" captionStart="Figs 26-31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26-31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the first tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="4" pageNumber="5">2627</figureCitation>
). Color reddish brown, darker in central area of head and lighter on antennae and legs. Integument evenly covered with a yellowish pubescence, thin, short and flattened, slightly raised in forehead area and clypeus. Head retractable; head sculpture composed of a tight mesh of small dots. Eyes developed with pigmented ommatidia (
<figureCitation id="1369F3A06661AB57B60C6336FC603D39" box="[864,950,1627,1653]" captionStart="Figs 26-31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26-31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the first tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="4" pageNumber="5">Fig. 27</figureCitation>
); antennae long (
<quantity id="4CAA42C06661AB57B1166336FB033D3A" box="[1146,1237,1627,1654]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="5" unit="mm" value="1.9">1.9 mm</quantity>
) and reaching almost one third of basal area of elytra; antennomeres (
<figureCitation id="1369F3A06661AB57B6276313FBD63DD4" box="[843,1024,1662,1688]" captionStart="Figs 26 - 31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26 - 31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the frst tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 26 27 = 100 μm, 28 31 = 10 μm." httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="4" pageNumber="5">Figs 2627, 29</figureCitation>
<figureCitation id="1369F3A06661AB57B6276313FBD63DD4" box="[843,1024,1662,1688]" captionStart="Figs 26-31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26-31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the first tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="4" pageNumber="5">Figs 2627, 29</figureCitation>
) slender, all clearly longer than wide; 2th, 4th, 5th and 7th antennomeres equal, 3rd slightly longer, 6th shorter and 8th almost twice as long as wide (
<tableCitation id="C6D0DA9E6661AB57B44963A8FEAB3D93" box="[293,381,1733,1759]" captionStart="Table 1" captionStartId="4.[188,257,267,293]" captionTargetBox="[193,1384,385,1378]" captionTargetPageId="4" captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width." httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04" pageId="4" pageNumber="5" tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
); tip of antenna with sensorial organ (
<figureCitation id="1369F3A06661AB57B65163A8FC433D93" box="[829,917,1733,1759]" captionStart="Figs 26 - 31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26 - 31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the frst tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 26 27 = 100 μm, 28 31 = 10 μm." httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="4" pageNumber="5">Fig. 29</figureCitation>
<figureCitation id="1369F3A06661AB57B65163A8FC433D93" box="[829,917,1733,1759]" captionStart="Figs 26-31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26-31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the first tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="4" pageNumber="5">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="8BEDEF256661AB57B5D1626AFC673C9D" blockId="4.[189,1399,1798,2001]" pageId="4" pageNumber="5">
PRONOTUM (
<figureCitation id="1369F3A06661AB57B42B626BFE643C6D" box="[327,434,1798,1825]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="4" pageNumber="5">Figs 67</figureCitation>
). Slightly transverse, maximum width/length ratio = 1.291.43 (males) and 1.28 1.38 (females), with maximum width at middle; almost fattened; basal impressions not observed; lateral and basal margins weakly beaded; pronotal sides regularly curved anteriorly, posteriorly almost straight and converging posteriorly; basal angles obtuse with apex almost rounded. Pronotal base narrower than base of elytra; middle area gently arched, lateral areas almost straight. Pronotum surface uniformly granulate, well defned laterally and more indistinct in disk area.
<figureCitation id="1369F3A06661AB57B42B626BFE643C6D" box="[327,434,1798,1825]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="4" pageNumber="5">Figs 67</figureCitation>
). Slightly transverse, maximum width/length ratio = 1.291.43 (males) and 1.28 1.38 (females), with maximum width at middle; almost flattened; basal impressions not observed; lateral and basal margins weakly beaded; pronotal sides regularly curved anteriorly, posteriorly almost straight and converging posteriorly; basal angles obtuse with apex almost rounded. Pronotal base narrower than base of elytra; middle area gently arched, lateral areas almost straight. Pronotum surface uniformly granulate, well defined laterally and more indistinct in disk area.
</paragraph>
<paragraph id="8BEDEF256660AB56B5D16461FD2A3AFE" blockId="5.[189,1399,267,435]" pageId="5" pageNumber="6">
ELYTRA. Elliptical, very elongate and convex, except along suture where they are fattened in the central area; maximum length/width ratio =
ELYTRA. Elliptical, very elongate and convex, except along suture where they are flattened in the central area; maximum length/width ratio =
<quantity id="4CAA42C06660AB56B7A96443FC8F3A04" box="[709,857,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="4.8641" metricValueMax="5.0292" metricValueMin="4.699" pageId="5" pageNumber="6" unit="in" value="1.915" valueMax="1.98" valueMin="1.85">1.851.98 in</quantity>
males and
<quantity id="4CAA42C06660AB56B6B26443FBA43A04" box="[990,1138,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="5.0419" metricValueMax="5.1308" metricValueMin="4.953" pageId="5" pageNumber="6" unit="in" value="1.9849999999999999" valueMax="2.02" valueMin="1.95">1.952.02 in</quantity>
females; elytron apex rounded; transverse striations visible, clearly separated, shallow and perpendicular to elytral suture (
<figureCitation id="1369F3A06660AB56B5AA6418FECB3AC3" box="[198,285,373,399]" captionStart="Figs 26 - 31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26 - 31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the frst tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 26 27 = 100 μm, 28 31 = 10 μm." httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="5" pageNumber="6">Fig. 28</figureCitation>
<figureCitation id="1369F3A06660AB56B5AA6418FECB3AC3" box="[198,285,373,399]" captionStart="Figs 26-31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26-31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the first tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="5" pageNumber="6">Fig. 28</figureCitation>
); sutural groove present and well marked along entire elytron, sub-parallel to suture and clearly convergent. Metathoracic wings fully developed.
</paragraph>
<paragraph id="8BEDEF256660AB56B5D164B7FA863976" blockId="5.[189,1398,473,571]" pageId="5" pageNumber="6">
LEGS. Relatively long and thin; protibial keel (
<figureCitation id="1369F3A06660AB56B78F64B4FCEA3AB8" box="[739,828,473,500]" captionStart="Figs 16 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="5" pageNumber="6">Fig. 16</figureCitation>
), which is high and long, elevated distally, with a well marked vertex, sharp at apex; frst three protarsomeres dilated (
<figureCitation id="1369F3A06660AB56B1646490FBB6395B" box="[1032,1120,509,535]" captionStart="Figs 26 - 31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26 - 31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the frst tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 26 27 = 100 μm, 28 31 = 10 μm." httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="5" pageNumber="6">Fig. 30</figureCitation>
) in males, frst slightly narrower than apical area of protibia (tarsus/tibia ratio = 0.86); mesotibia curved, metatibia straight.
), which is high and long, elevated distally, with a well marked vertex, sharp at apex; first three protarsomeres dilated (
<figureCitation id="1369F3A06660AB56B1646490FBB6395B" box="[1032,1120,509,535]" captionStart="Figs 26-31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26-31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the first tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="5" pageNumber="6">Fig. 30</figureCitation>
) in males, first slightly narrower than apical area of protibia (tarsus/tibia ratio = 0.86); mesotibia curved, metatibia straight.
</paragraph>
<caption id="DF2DBFAD6660AB56B5D1622CFD2A3CA5" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="5" pageNumber="6" targetBox="[189,1398,686,1816]" targetPageId="5">
<caption id="DF2DBFAD6660AB56B5D1622CFD2A3CA5" ID-DOI="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="5" pageNumber="6" targetBox="[189,1398,686,1816]" targetPageId="5">
<paragraph id="8BEDEF256660AB56B5D1622CFD2A3CA5" blockId="5.[189,1399,1857,2025]" pageId="5" pageNumber="6">
<emphasis id="B92633376660AB56B5D1622CFEEA3C17" bold="true" box="[189,316,1857,1883]" pageId="5" pageNumber="6">Figs 615.</emphasis>
<taxonomicName id="4C5294A66660AB56B428622CFE333C17" authorityName="Jeannel" authorityYear="1922" box="[324,485,1857,1883]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
@ -839,7 +862,7 @@ of the
<emphasis id="B92633376660AB56B6D4622FFB8B3C17" box="[952,1117,1857,1884]" italics="true" pageId="5" pageNumber="6">S. algarvensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2158E4C6660AB56B10A622FFB173C10" box="[1126,1217,1858,1884]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(Gruta do Vale Telheiro, Portugal), respectively and ♀.
(Gruta do Vale Telheiro, Portugal), respectively and ♀.
<emphasis id="B92633376660AB56B7C46209FD023C32" bold="true" box="[680,724,1892,1918]" pageId="5" pageNumber="6">89</emphasis>
.
<treatmentCitation id="0AF3C9346660AB56B78D6208FB943C33" author="Kraatz" authority="(Kraatz, 1870)" authorityName="Kraatz" authorityYear="1870" baseAuthorityName="Kraatz" baseAuthorityYear="1870" box="[737,1090,1893,1919]" class="Insecta" family="Leiodidae" firstAuthor="Kraatz" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" pagination="98 - 99" phylum="Arthropoda" rank="species" refId="ref12372" refString="Kraatz G. 1870. Choleva angusticollis Kraatz n. sp. In: Heyden L. von (ed.) Entomologische Reise nach dem sudlichen Spanien der Sierra Guadarrama und Sierra Morena, Portugal und den Cantabrischen Gebirgen: 98 - 99. Entomologischer Verein in Berlin, Berlin." species="angusticollis" treatmentTaxon="S. angusticollis" type="book chapter" year="1870">
@ -850,7 +873,7 @@ of the
)
</taxonomicName>
</treatmentCitation>
(Rute, Spain) respectively and ♀.
(Rute, Spain) respectively and ♀.
<emphasis id="B92633376660AB56B45B62E5FEAB3CEE" bold="true" box="[311,381,1928,1954]" pageId="5" pageNumber="6">1011</emphasis>
.
<treatmentCitation id="0AF3C9346660AB56B4E362E5FD1D3CEE" author="Jeannel" authority="(Jeannel, 1922)" authorityName="Jeannel" authorityYear="1922" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[399,715,1928,1954]" class="Insecta" family="Leiodidae" firstAuthor="Jeannel" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" pagination="1 - 98" phylum="Arthropoda" rank="species" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." species="bolivari" treatmentTaxon="S. bolivari" type="journal article" year="1922">
@ -861,7 +884,7 @@ of the
)
</taxonomicName>
</treatmentCitation>
(Ardales, Spain) respectively and ♀.
(Ardales, Spain) respectively and ♀.
<emphasis id="B92633376660AB56B1DC62E5FB2C3CEE" bold="true" box="[1200,1274,1928,1954]" pageId="5" pageNumber="6">1213</emphasis>
.
<treatmentCitation id="0AF3C9346660AB56B06662E5FEA53C89" author="Jeannel" authority="(Jeannel, 1922)" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" class="Insecta" family="Leiodidae" firstAuthor="Jeannel" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" pagination="1 - 98" phylum="Arthropoda" rank="species" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." species="breuili" treatmentTaxon="S. breuili" type="journal article" year="1922">
@ -872,7 +895,7 @@ of the
)
</taxonomicName>
</treatmentCitation>
(Motillas, Spain) respectively and ♀.
(Motillas, Spain) respectively and ♀.
<emphasis id="B92633376660AB56B63262C6FC713C89" bold="true" box="[862,935,1963,1989]" pageId="5" pageNumber="6">1415</emphasis>
.
<treatmentCitation id="0AF3C9346660AB56B6DB62C6FAD03C89" author="Uhagon" authority="(Uhagon, 1898)" authorityName="Uhagon" authorityYear="1898" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[951,1286,1963,1989]" class="Insecta" family="Leiodidae" firstAuthor="Uhagon" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" pagination="117 - 126" phylum="Arthropoda" rank="species" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi &quot; Ensayo sobre las especies espanolas del grupo &quot; Cholevae &quot;. Anales de la Sociedad espanola de Historia natural, 7 (2), 27 &quot;. Actas de la Sociedad espanola de Historia natural 1: 117 - 126." species="escalerai" treatmentTaxon="S. escalerai" type="journal article" year="1898">
@ -883,7 +906,7 @@ of the
)
</taxonomicName>
</treatmentCitation>
(Jumilla, Spain) respectively and ♀. Scale bars = 1 mm.
(Jumilla, Spain) respectively and ♀. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BEDEF256663AB55B5866466FEFC3A69" blockId="6.[189,1399,267,645]" box="[234,298,267,293]" pageId="6" pageNumber="7">
@ -900,10 +923,10 @@ Some variability is observed in male
<quantity id="4CAA42C06663AB55B1746418FBA23AC3" box="[1048,1140,373,399]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="mm" value="1.2">1.2 mm</quantity>
(1.3 if parameres are included) (
<figureCitation id="1369F3A06663AB55B45364F5FE4C3AFE" box="[319,410,408,434]" captionStart="Figs 21 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="6" pageNumber="7">Fig. 21</figureCitation>
). Median lobe, lanceolate in dorsal view, with elongate and curved apex close to dorsal face; sharp apex with marked shoulders at margins; basal lamina developed, almost two times shorter than middle lobe; ventral blade of tegmen short and poorly defned. Parameres, in dorsal view, sub rectilinear, slightly curved medially on inner side, with apical areas rounded and slightly truncated on inner side; four thin setae at apex, three subequal upper setae and a larger one, close to largest setae, another seta, shorter, more robust, tooth-like and highly sclerotized. Inner sac with two long rows of spinules surrounding two long parallel rows of sclerotized teeth arranged in a zipper-shape (
). Median lobe, lanceolate in dorsal view, with elongate and curved apex close to dorsal face; sharp apex with marked shoulders at margins; basal lamina developed, almost two times shorter than middle lobe; ventral blade of tegmen short and poorly defined. Parameres, in dorsal view, sub rectilinear, slightly curved medially on inner side, with apical areas rounded and slightly truncated on inner side; four thin setae at apex, three subequal upper setae and a larger one, close to largest setae, another seta, shorter, more robust, tooth-like and highly sclerotized. Inner sac with two long rows of spinules surrounding two long parallel rows of sclerotized teeth arranged in a zipper-shape (
<figureCitation id="1369F3A06663AB55B18F6706FAEB39C9" box="[1251,1341,619,645]" captionStart="Figs 21 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="6" pageNumber="7">Figs 21</figureCitation>
,
<figureCitation id="1369F3A06663AB55B0246706FABE39C9" box="[1352,1384,619,645]" captionStart="Figs 26 - 31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26 - 31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the frst tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 26 27 = 100 μm, 28 31 = 10 μm." httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="6" pageNumber="7">31</figureCitation>
<figureCitation id="1369F3A06663AB55B0246706FABE39C9" box="[1352,1384,619,645]" captionStart="Figs 26-31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26-31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the first tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="6" pageNumber="7">31</figureCitation>
).
</paragraph>
<paragraph id="8BEDEF256663AB55B58667C1FE93398A" blockId="6.[189,1399,684,816]" box="[234,325,684,710]" pageId="6" pageNumber="7">
@ -913,8 +936,8 @@ Some variability is observed in male
</paragraph>
<paragraph id="8BEDEF256663AB54B5D167A2FC153AC3" blockId="6.[189,1399,684,816]" lastBlockId="7.[189,1398,266,399]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Generally smaller, with a less transverse pronotum (
<figureCitation id="1369F3A06663AB55B65967A2FCA839A6" box="[821,894,719,746]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
) and longer elytra, but same body width. No sexual dimorphism observed in colour, pubescence, sculpture, granularity, striation, mesotibia or metatibia. Females have a slender protarsus; antennomeres proportionally thicker and elytral apex notched and serrate. Seventh uroventrite with a narrow slit in middle of posterior margin and 8th uroventrite with posterior margin as a soft membranous arc, its ventral spine is short, wide and slightly sharp at apex. Ratio of frst tarsus to apical part of tibia: 0.840.89. Female genitalia typical of genus as described by
<figureCitation id="1369F3A06663AB55B65967A2FCA839A6" box="[821,894,719,746]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
) and longer elytra, but same body width. No sexual dimorphism observed in colour, pubescence, sculpture, granularity, striation, mesotibia or metatibia. Females have a slender protarsus; antennomeres proportionally thicker and elytral apex notched and serrate. Seventh uroventrite with a narrow slit in middle of posterior margin and 8th uroventrite with posterior margin as a soft membranous arc, its ventral spine is short, wide and slightly sharp at apex. Ratio of first tarsus to apical part of tibia: 0.840.89. Female genitalia typical of genus as described by
<bibRefCitation id="EFC392D46662AB54B4356418FD5A3AC3" author="Giachino P. M. &amp; Vailati D." box="[345,652,373,399]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="7" pageNumber="8" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati (1993)</bibRefCitation>
and
<bibRefCitation id="EFC392D46662AB54B7A96418FC693AC3" author="Salgado" box="[709,959,373,399]" etAl="et al." firstAuthor="Salgado" pageId="7" pageNumber="8" refId="ref13714" refString="Salgado J. M., Blas M. &amp; Fresneda J. 2008. Coleoptera, Cholevidae. Fauna Iberica, vol. 31, Museo Nacional de Ciencias Naturales, CSIC, Madrid." type="journal volume" year="2008">
@ -1109,7 +1132,7 @@ by the different shape of the male protibial keel, by the antennomeres ratio and
</taxonomicName>
are much more clear in the shape of the protibial keel.
</paragraph>
<caption id="DF2DBFAD666DAB5BB5D16229FD9F3CA7" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="8" pageNumber="9" targetBox="[189,1398,600,1819]" targetPageId="8">
<caption id="DF2DBFAD666DAB5BB5D16229FD9F3CA7" ID-DOI="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="8" pageNumber="9" targetBox="[189,1398,600,1819]" targetPageId="8">
<paragraph id="8BEDEF25666DAB5BB5D16229FD9F3CA7" blockId="8.[189,1399,1860,2027]" pageId="8" pageNumber="9">
<emphasis id="B9263337666DAB5BB5D16229FE9F3C12" bold="true" box="[189,329,1860,1886]" pageId="8" pageNumber="9">Figs 26-31.</emphasis>
<taxonomicName id="4C5294A6666DAB5BB4386229FD513C12" authority="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado, 2017" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" box="[340,647,1860,1886]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="algarvensis" status="sp. nov.">
@ -1133,7 +1156,7 @@ pointed out.
<emphasis id="B9263337666DAB5BB7CF62E7FD163CE8" bold="true" box="[675,704,1930,1956]" pageId="8" pageNumber="9">29</emphasis>
. Tip of the antenna.
<emphasis id="B9263337666DAB5BB6D262E7FC0D3CE8" bold="true" box="[958,987,1930,1956]" pageId="8" pageNumber="9">30</emphasis>
. Male protarsus with detail of the ventral pads of the frst tarsomere.
. Male protarsus with detail of the ventral pads of the first tarsomere.
<emphasis id="B9263337666DAB5BB70562C0FD503C8B" bold="true" box="[617,646,1965,1991]" pageId="8" pageNumber="9">31</emphasis>
. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm.
</paragraph>
@ -1156,7 +1179,7 @@ Jeannel
<emphasis id="B9263337666CAB5AB74C6752FD143914" box="[544,706,574,601]" italics="true" pageId="9" pageNumber="10">S. algarvensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2158E4C666CAB5AB7A46752FCF43915" box="[712,802,575,601]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, it had previously been identifed as
, it had previously been identified as
<taxonomicName id="4C5294A6666CAB5AB1D26752FAB93914" baseAuthorityName="Kraatz" baseAuthorityYear="1870" box="[1214,1391,574,601]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B9263337666CAB5AB1D26752FAB93914" box="[1214,1391,574,601]" italics="true" pageId="9" pageNumber="10">S. angusticollis</emphasis>
</taxonomicName>
@ -1253,8 +1276,8 @@ Thaxt., 1901
(of the order
<taxonomicName id="4C5294A6666CAB5AB49C61FBFD733FFD" authorityName="Engler" authorityYear="1898" box="[496,677,1174,1201]" class="Laboulbeniomycetes" kingdom="Fungi" order="Laboulbeniales" pageId="9" pageNumber="10" phylum="Ascomycota" rank="order">Laboulbeniales</taxonomicName>
) attached to the cuticle (Santamaria pers. com.) (
<figureCitation id="1369F3A0666CAB5AB18C61FBFAE13FFD" box="[1248,1335,1174,1201]" captionStart="Figs 26 - 31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26 - 31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the frst tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 26 27 = 100 μm, 28 31 = 10 μm." httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="9" pageNumber="10">Fig. 26</figureCitation>
); this represents the frst record of the fungal species for the Portuguese territory. This fungus species has only been previously found in
<figureCitation id="1369F3A0666CAB5AB18C61FBFAE13FFD" box="[1248,1335,1174,1201]" captionStart="Figs 26-31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26-31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the first tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="9" pageNumber="10">Fig. 26</figureCitation>
); this represents the first record of the fungal species for the Portuguese territory. This fungus species has only been previously found in
<taxonomicName id="4C5294A6666CAB5AB49861B1FD433FBA" box="[500,661,1244,1270]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
beetles of the genus
<taxonomicName id="4C5294A6666CAB5AB6F561B1FB043FBB" authority="Gistel, 1856" authorityName="Gistel" authorityYear="1856" box="[921,1234,1244,1271]" class="Insecta" family="Staphylinidae" genus="Sepedophilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
@ -1271,11 +1294,11 @@ Haelewaters
<taxonomicName id="4C5294A6666CAB5AB5D1604EFEA63E71" authorityName="Engler" authorityYear="1898" box="[189,368,1315,1341]" class="Laboulbeniomycetes" kingdom="Fungi" order="Laboulbeniales" pageId="9" pageNumber="10" phylum="Ascomycota" rank="order">Laboulbeniales</taxonomicName>
species on a new host of the family
<taxonomicName id="4C5294A6666CAB5AB677604EFC463E71" authorityName="Fleming" authorityYear="1821" box="[795,912,1315,1341]" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
is the frst case of host shifting following an ecological opportunity (
is the first case of host shifting following an ecological opportunity (
<emphasis id="B9263337666CAB5AB494602AFDEF3E2C" box="[504,569,1351,1376]" italics="true" pageId="9" pageNumber="10">sensu</emphasis>
<bibRefCitation id="EFC392D4666CAB5AB72C602AFC703E2C" author="De Kesel A. &amp; Haelewaters D." box="[576,934,1350,1377]" firstAuthor="De Kesel" journalOrPublisher="Mycologia" pageId="9" pageNumber="10" pagination="407 - 414" part="106" publicationUrl="http://dx.doi.org/10.3852/13-348" refId="ref11709" refString="De Kesel A. &amp; Haelewaters D. 2014. Laboulbenia slackensis and L. littoralis sp. nov. (Ascomycota, Laboulbeniales), two sibling species as a result of ecological speciation. Mycologia 106 (3): 407 - 414. http: // dx. doi. org / 10.3852 / 13 - 348" title="Laboulbenia slackensis and L. littoralis sp. nov. (Ascomycota, Laboulbeniales), two sibling species as a result of ecological speciation" type="journal article" year="2014">De Kesel &amp; Haelewaters 2014</bibRefCitation>
), in the subterranean environment. Also phoretic undetermined mites were observed on some specimens (
<figureCitation id="1369F3A0666CAB5AB6D66004FBC43ECF" box="[954,1042,1385,1411]" captionStart="Figs 26 - 31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26 - 31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the frst tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 26 27 = 100 μm, 28 31 = 10 μm." httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="9" pageNumber="10">Fig. 27</figureCitation>
<figureCitation id="1369F3A0666CAB5AB6D66004FBC43ECF" box="[954,1042,1385,1411]" captionStart="Figs 26-31" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1398,600,1819]" captionTargetId="figure@8.[189,1398,600,1819]" captionTargetPageId="8" captionText="Figs 26-31. Speonemadus algarvensis sp. nov. scanning electron micrograph. 26. Dorsal view with Laboulbeniales Stichomyces conosomatis pointed out. 27. Ventral view, with detail of phoretic acari. 28. Details of elytron stria and setation. 29. Tip of the antenna. 30. Male protarsus with detail of the ventral pads of the first tarsomere. 31. Posterior dorsal view of the evaginated aedeagus. Scale bars 2627 = 100 μm, 2831 = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.889356" httpUri="https://zenodo.org/record/889356/files/figure.png" pageId="9" pageNumber="10">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="8BEDEF25666CAB5AB5D160C6FC3E3C69" blockId="9.[189,1399,1451,1829]" pageId="9" pageNumber="10">
@ -1439,7 +1462,7 @@ was collected in three caves of the southernmost province of
<collectingCountry id="F345AFB5666CAB5AB0606216FAA43CD9" box="[1292,1394,1915,1941]" name="Portugal" pageId="9" pageNumber="10">Portugal</collectingCountry>
, the Algarve (
<figureCitation id="1369F3A0666CAB5AB43A62F3FE783CF4" box="[342,430,1950,1976]" captionStart="Fig. 32" captionStartId="18.[189,232,1955,1981]" captionTargetBox="[189,1398,986,1883]" captionTargetId="figure@18.[189,1398,986,1914]" captionTargetPageId="18" captionText="Fig. 32. Distribution of Speonemadus of the escalerai - group: S. algarvensis sp. nov., S. angusticollis (Kraatz, 1870), S. bolivari (Jeannel, 1922), S. breuili (Jeannel, 1922) and S. escalerai (Uhagón, 1898)." httpUri="https://zenodo.org/record/889358/files/figure.png" pageId="9" pageNumber="10">Fig. 32</figureCitation>
), where it seems to be endemic to the central and eastern part of the Algarve karst massif. The species was not found in the most western part of the Algarve massif, where feldwork was also conducted. The karst included in this region is also know as “Barrocal” and is the richest area for subterranean-adapted fauna of the country (
), where it seems to be endemic to the central and eastern part of the Algarve karst massif. The species was not found in the most western part of the Algarve massif, where fieldwork was also conducted. The karst included in this region is also know as “Barrocal” and is the richest area for subterranean-adapted fauna of the country (
<bibRefCitation id="EFC392D4666FAB59B7976443FC6E3A05" author="Reboleira" box="[763,952,302,329]" firstAuthor="Reboleira" pageId="10" pageNumber="11" refId="ref12782" refString="Reboleira A. S. P. S. 2012. Biodiversity and conservation of subterranean fauna of Portuguese karst. PhD thesis. University of Aveiro, Portugal." type="book" year="2012">Reboleira 2012</bibRefCitation>
;
<bibRefCitation id="EFC392D4666FAB59B6AB6443FB013A05" author="Reboleira" box="[967,1239,302,329]" etAl="et al." firstAuthor="Reboleira" pageId="10" pageNumber="11" pagination="23 - 37" refId="ref13205" refString="Reboleira A. S. P. S., Borges P., Goncalves F., Serrano A. R. M. &amp; Oromi P. 2011 a. The subterranean fauna of a biodiversity hotspot region - Portugal: an overview and its conservation. International Journal of Speleology 40 (1): 23 - 37. http: // dx. doi. org / 10.5038 / 1827 - 806 X. 40.1.4" type="journal article" year="2011" yearSuffix="a">

View file

@ -1,53 +1,53 @@
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@ -61,7 +61,7 @@ Genus
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<taxonomicName id="4C5294A66667AB51B5D16420FE043A2B" ID-CoL="4YHYZ" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[189,466,333,359]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="escalerai">
<emphasis id="B92633376667AB51B5D16420FE043A2B" box="[189,466,333,359]" italics="true" pageId="2" pageNumber="3">Speonemadus escalerai</emphasis>
@ -79,12 +79,12 @@ Genus
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Representatives of this group are characterized by a body length between 3.9 to 5.0 mm and have a slightly transverse pronotum (
<figureCitation id="1369F3A06667AB51B74464C7FD733A89" box="[552,677,426,453]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="2" pageNumber="3">Figs 615</figureCitation>
<figureCitation id="1369F3A06667AB51B74464C7FD733A89" box="[552,677,426,453]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="2" pageNumber="3">Figs 615</figureCitation>
); the anterior tibia of the males is compressed towards the apical half and has a keel with different shapes on the dorsal edge (
<figureCitation id="1369F3A06667AB51B6B464A0FBB43AA4" box="[984,1122,461,488]" captionStart="Figs 16 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Figs 1620</figureCitation>
); elytra have a rounded apex in males and a notched one in females. The median lobe of the aedeagus is elongated and bottleshaped in dorsal view (
<figureCitation id="1369F3A06667AB51B4B8677EFD8A3962" box="[468,604,531,558]" captionStart="Figs 21 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="2" pageNumber="3">Figs 2125</figureCitation>
) and the inner sac lacks a ventral tooth in the apical region of the middle lobe and has two long parallel rows of fne spines and teeth (
) and the inner sac lacks a ventral tooth in the apical region of the middle lobe and has two long parallel rows of fine spines and teeth (
<figureCitation id="1369F3A06667AB51B6B0675AFBB0391D" box="[988,1126,567,593]" captionStart="Figs 21 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="2" pageNumber="3">Figs 2324</figureCitation>
).
</paragraph>
@ -104,9 +104,9 @@ Key to the species of the
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE6667AB51B5D16610FAA03CAA" pageId="2" pageNumber="3" type="key">
<subSubSection id="C348BCAE6667AB51B5D16610FC173F9F" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BEDEF256667AB51B5D16610FC173F9F" blockId="2.[189,1399,845,1235]" pageId="2" pageNumber="3">
Note to the key: diversifcation of the species of the
Note to the key: diversification of the species of the
<taxonomicName id="4C5294A66667AB51B6596611FC7738DA" ID-CoL="4YHYZ" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[821,929,892,918]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="escalerai">
<emphasis id="B92633376667AB51B6596611FC7738DA" box="[821,929,892,918]" italics="true" pageId="2" pageNumber="3">escalerai</emphasis>
</taxonomicName>
@ -122,8 +122,10 @@ Herbst, 1797. Only females of
<taxonomicName id="4C5294A66667AB51B64A613DFC0E3F26" ID-CoL="4YHYV" baseAuthorityName="Kraatz" baseAuthorityYear="1870" box="[806,984,1104,1130]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B92633376667AB51B64A613DFC0E3F26" box="[806,984,1104,1130]" italics="true" pageId="2" pageNumber="3">S. angusticollis</emphasis>
</taxonomicName>
can be separated from other species by the shape of the pronotum, being the only one with a bell-shape, as in males. Therefore, in this species group only males provide distinctive characters useful for identifcation. These characters are identifed in all key items and constitute the basis of specifc differentiation.
can be separated from other species by the shape of the pronotum, being the only one with a bell-shape, as in males. Therefore, in this species group only males provide distinctive characters useful for identification. These characters are identified in all key items and constitute the basis of specific differentiation.
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE6667AB51B5D16181FAA03CAA" pageId="2" pageNumber="3" type="key">
<keyStep id="30A6F7806667AB51B5D16181FAA13E65" pageId="2" pageNumber="3">
<paragraph id="8BEDEF256667AB51B5D16181FAA03E4A" blockId="2.[189,1399,1259,2023]" box="[189,1398,1259,1286]" pageId="2" pageNumber="3">
<keyLead id="30A34C106667AB51B5D16181FAA03E4A" box="[189,1398,1259,1286]" pageId="2" pageNumber="3">
@ -146,7 +148,7 @@ can be separated from other species by the shape of the pronotum, being the only
2. Protibial keel with sharp vertex and slightly lobed from the margin to the base (
<figureCitation id="1369F3A06667AB51B1D36024FACD3E2F" box="[1215,1307,1353,1379]" captionStart="Figs 16 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Fig. 20</figureCitation>
); elytra sculpture and pronotum granularity poorly marked; pronotum sexually dimorphic (
<figureCitation id="1369F3A06667AB51B1C56001FAFB3ECB" box="[1193,1325,1388,1415]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="2" pageNumber="3">Figs 1415</figureCitation>
<figureCitation id="1369F3A06667AB51B1C56001FAFB3ECB" box="[1193,1325,1388,1415]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="2" pageNumber="3">Figs 1415</figureCitation>
) …… …………………………………………………………………………
<treatmentCitation id="0AF3C9346667AB51B14C60E2FAA03EE5" author="Uhagon" authority="(Uhagon, 1898)" authorityName="Uhagon" authorityYear="1898" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[1056,1398,1423,1450]" class="Insecta" family="Leiodidae" firstAuthor="Uhagon" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" pagination="117 - 126" phylum="Arthropoda" rank="species" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi &quot; Ensayo sobre las especies espanolas del grupo &quot; Cholevae &quot;. Anales de la Sociedad espanola de Historia natural, 7 (2), 27 &quot;. Actas de la Sociedad espanola de Historia natural 1: 117 - 126." species="escalerai" treatmentTaxon="S. escalerai" type="journal article" year="1898">
<taxonomicName id="4C5294A66667AB51B14C60E2FAA03EE5" ID-CoL="4YHYZ" authority="(Uhagon, 1898)" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[1056,1398,1423,1450]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="escalerai">
@ -163,7 +165,7 @@ can be separated from other species by the shape of the pronotum, being the only
Protibial keel with a slightly sharp vertex and linear margin to the base (
<figureCitation id="1369F3A06667AB51B14460DFFB7B3E80" box="[1064,1197,1458,1484]" captionStart="Figs 16 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="2" pageNumber="3">Figs 16, 18</figureCitation>
); elytra sculpture and pronotum with well marked granularity; pronotum not sexually dimorphic (
<figureCitation id="1369F3A06667AB51B1EC60B8FAEA3EBC" box="[1152,1340,1493,1520]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="2" pageNumber="3">Figs 67, 1011</figureCitation>
<figureCitation id="1369F3A06667AB51B1EC60B8FAEA3EBC" box="[1152,1340,1493,1520]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="2" pageNumber="3">Figs 67, 1011</figureCitation>
) …3
</keyLead>
</paragraph>

View file

@ -1,53 +1,53 @@
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<treatment id="03FB5E336669AB5DB75061A2FC11393F" ID-DOI="http://doi.org/10.5281/zenodo.5689572" ID-GBIF-Taxon="133183539" ID-Zenodo-Dep="5689572" LSID="urn:lsid:plazi:treatment:03FB5E336669AB5DB75061A2FC11393F" httpUri="http://treatment.plazi.org/id/03FB5E336669AB5DB75061A2FC11393F" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<treatment id="03FB5E336669AB5DB75061A2FC11393F" ID-DOI="http://doi.org/10.5281/zenodo.5689572" ID-GBIF-Taxon="133183539" ID-Zenodo-Dep="5689572" LSID="urn:lsid:plazi:treatment:03FB5E336669AB5DB75061A2FC11393F" httpUri="http://treatment.plazi.org/id/03FB5E336669AB5DB75061A2FC11393F" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" scope_family="Leiodidae" scope_order="Coleoptera" scope_subFamily="Cholevinae" scope_tribe="Anemadini">
<subSubSection id="C348BCAE6669AB5FB75061A2FC213FA6" box="[572,1015,1231,1258]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BEDEF256669AB5FB75061A2FC213FA6" blockId="12.[572,1015,1231,1258]" box="[572,1015,1231,1258]" pageId="12" pageNumber="13">
<heading id="D0A558496669AB5FB75061A2FC213FA6" box="[572,1015,1231,1258]" centered="true" fontSize="11" level="3" pageId="12" pageNumber="13" reason="4">
@ -77,7 +77,7 @@
.
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D1603EFCED3E21" blockId="12.[189,999,1363,1460]" box="[189,827,1363,1389]" pageId="12" pageNumber="13">
<taxonomicName id="4C5294A66669AB5FB5D1603EFD863E21" box="[189,592,1363,1389]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="verneri" subGenus="Speonemadus">
<taxonomicName id="4C5294A66669AB5FB5D1603EFD863E21" authority="Jeannel 1922: 61" box="[189,592,1363,1389]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="verneri" subGenus="Speonemadus">
<emphasis id="B92633376669AB5FB5D1603EFEE13E21" box="[189,311,1363,1389]" italics="true" pageId="12" pageNumber="13">Anemadus</emphasis>
(
<emphasis id="B92633376669AB5FB424603EFE3F3E21" box="[328,489,1363,1389]" italics="true" pageId="12" pageNumber="13">Speonemadus</emphasis>
@ -89,7 +89,7 @@
.
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D1601BFC093EDC" blockId="12.[189,999,1363,1460]" box="[189,991,1398,1424]" pageId="12" pageNumber="13">
<taxonomicName id="4C5294A66669AB5FB5D1601BFD333EDC" ID-CoL="DWR5" box="[189,741,1398,1424]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="angusticollis" subGenus="Speonemadus" subSpecies="breuili">
<taxonomicName id="4C5294A66669AB5FB5D1601BFD333EDC" ID-CoL="DWR5" authority="Jeannel, 1922" box="[189,741,1398,1424]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="angusticollis" subGenus="Speonemadus" subSpecies="breuili">
<emphasis id="B92633376669AB5FB5D1601BFEE13EDC" box="[189,311,1398,1424]" italics="true" pageId="12" pageNumber="13">Anemadus</emphasis>
(
<emphasis id="B92633376669AB5FB424601BFE3F3EDC" box="[328,489,1398,1424]" italics="true" pageId="12" pageNumber="13">Speonemadus</emphasis>
@ -101,7 +101,7 @@
.
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D160F4FC313EFF" blockId="12.[189,999,1363,1460]" box="[189,999,1433,1460]" pageId="12" pageNumber="13">
<taxonomicName id="4C5294A66669AB5FB5D160F4FD3B3EFF" box="[189,749,1433,1460]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="angusticollis" subGenus="Speonemadus" subSpecies="verneri">
<taxonomicName id="4C5294A66669AB5FB5D160F4FD3B3EFF" authority="(Jeannel 1922: 61)" box="[189,749,1433,1460]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="angusticollis" subGenus="Speonemadus" subSpecies="verneri">
<emphasis id="B92633376669AB5FB5D160F4FEE13EFF" box="[189,311,1433,1459]" italics="true" pageId="12" pageNumber="13">Anemadus</emphasis>
(
<emphasis id="B92633376669AB5FB42460F4FE3F3EFF" box="[328,489,1433,1459]" italics="true" pageId="12" pageNumber="13">Speonemadus</emphasis>
@ -113,7 +113,7 @@
.
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE6669AB5EB5D160B6FEBD39E2" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<subSubSection id="C348BCAE6669AB5EB5D160B6FDEE3922" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BEDEF256669AB5FB5D160B6FE8B3EB8" blockId="12.[189,1140,1498,1572]" box="[189,349,1498,1525]" pageId="12" pageNumber="13">
<heading id="D0A558496669AB5FB5D160B6FE8B3EB8" bold="true" box="[189,349,1498,1525]" fontSize="11" level="2" pageId="12" pageNumber="13" reason="6">
<emphasis id="B92633376669AB5FB5D160B6FE8B3EB8" bold="true" box="[189,349,1498,1525]" pageId="12" pageNumber="13">Type locality</emphasis>
@ -130,11 +130,10 @@
</heading>
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D16314FADE3CC6" blockId="12.[189,1399,1655,1930]" pageId="12" pageNumber="13">
<materialsCitation id="3B3AE5786669AB5FB5D16314FDC23DFA" ID-GBIF-Occurrence="1609370450" collectingDate="1970-07" collectionCode="CMCN" collectorName="Cadiz &amp; Villaluenga &amp; Cueva &amp; Aljibe &amp; Ribera &amp; Vinas" country="Spain" location="SPAIN" pageId="12" pageNumber="13" specimenCount="2" specimenCount-female="2">
<collectingCountry id="F345AFB56669AB5FB5D16314FEC23DDF" box="[189,276,1657,1683]" name="Spain" pageId="12" pageNumber="13">SPAIN</collectingCountry>
:
<collectorName id="26A78AF36669AB5FB44D6314FEBC3DDF" box="[289,362,1657,1683]" pageId="12" pageNumber="13">Cádiz</collectorName>
: 3 ♁♁,
: 3 ♂♂,
<specimenCount id="9D5424AC6669AB5FB4A16314FDC53DD8" box="[461,531,1657,1684]" pageId="12" pageNumber="13" type="female">2 ♀♀</specimenCount>
,
<collectorName id="26A78AF36669AB5FB74F6314FD7C3DDF" box="[547,682,1657,1683]" pageId="12" pageNumber="13">Villaluenga</collectorName>
@ -152,25 +151,23 @@ del
<collectorName id="26A78AF36669AB5FB44163F1FEA63DFA" box="[301,368,1692,1718]" pageId="12" pageNumber="13">Viñas</collectorName>
leg. (
<collectionCode id="ED4377E06669AB5FB4C363F1FDDA3DFA" box="[431,524,1692,1718]" pageId="12" pageNumber="13">CMCN</collectionCode>
)
</materialsCitation>
. — Málaga: many ♁♁ and ♀♀, Cortés de la Frontera, Cueva del Berrueco,
). — Málaga: many ♂♂ and ♀♀, Cortés de la Frontera, Cueva del Berrueco,
<date id="FFECC9E56669AB5FB5D163ADFE8B3D96" box="[189,349,1728,1754]" pageId="12" pageNumber="13" value="2009-09-12">12 Sep. 2009</date>
and
<date id="FFECC9E56669AB5FB4F063D2FDFB3D96" box="[412,557,1727,1754]" pageId="12" pageNumber="13" value="2010-06-06">6 Jun. 2010</date>
, GIEX leg. (CFL, CPB, CJMS, CZULE); 15 ♁♁, 9 ♀♀ Cortés de la Frontera, Peñón del Berrueco, Cueva del Humo,
, GIEX leg. (CFL, CPB, CJMS, CZULE); 15 ♂♂, 9 ♀♀ Cortés de la Frontera, Peñón del Berrueco, Cueva del Humo,
<date id="FFECC9E56669AB5FB662638EFC633DB1" box="[782,949,1763,1789]" pageId="12" pageNumber="13" value="2012-12-22">22 Dec. 2012</date>
, GIEX leg. (CFL, CJMS); many ♁♁ and ♀♀, Cortés de la Frontera, Complejo Motillas, Cueva Motilla fósil,
, GIEX leg. (CFL, CJMS); many ♂♂ and ♀♀, Cortés de la Frontera, Complejo Motillas, Cueva Motilla fósil,
<date id="FFECC9E56669AB5FB160626BFB713C6C" box="[1036,1191,1798,1824]" pageId="12" pageNumber="13" value="2009-09-13">13 Sep. 2009</date>
and
<date id="FFECC9E56669AB5FB1B3626BFAA43C6C" box="[1247,1394,1798,1824]" pageId="12" pageNumber="13" value="2009-11-08">8 Nov. 2009</date>
, GIEX leg. (CFL; CPB; CJMS); many ♁♁ and ♀♀, Cortés de la Frontera, Complejo Motillas, Cueva Motilla Parralejo,
, GIEX leg. (CFL; CPB; CJMS); many ♂♂ and ♀♀, Cortés de la Frontera, Complejo Motillas, Cueva Motilla Parralejo,
<date id="FFECC9E56669AB5FB4FF6221FDE63C2A" box="[403,560,1868,1894]" pageId="12" pageNumber="13" value="2009-09-13">13 Sep. 2009</date>
,
<date id="FFECC9E56669AB5FB7576221FD1D3C2A" box="[571,715,1868,1895]" pageId="12" pageNumber="13" value="2009-11-08">8 Nov. 2009</date>
and
<date id="FFECC9E56669AB5FB6686221FC593C2A" box="[772,911,1868,1895]" pageId="12" pageNumber="13" value="2010-06-05">5 Jun. 2010</date>
, GIEX leg. (CFL, CPB, CJMS, CZULE); 1 , 2 ♀♀, Montejaque, El Pozuelo, Cueva Sima Nueva,
, GIEX leg. (CFL, CPB, CJMS, CZULE); 1 , 2 ♀♀, Montejaque, El Pozuelo, Cueva Sima Nueva,
<date id="FFECC9E56669AB5FB60F621DFBD43CC5" box="[867,1026,1903,1930]" pageId="12" pageNumber="13" value="1986-03-29">29 Mar. 1986</date>
, Escolà leg. (CMCN).
</paragraph>
@ -251,13 +248,13 @@ Fresneda
<bibRefCitation id="EFC392D46668AB5EB4C8673EFDFA3922" author="Perez" box="[420,556,595,622]" firstAuthor="Perez" pageId="13" pageNumber="14" pagination="1 - 97" refId="ref12561" refString="Perez T. 2015. Catalogo de invertebrados citados en cuevas, simas y minas de Andalucia. Monografias Bioespeleologicas 9: 1 - 97." type="journal article" year="2015">Pérez 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE6668AB5EB5D167F9FAA13E87" pageId="13" pageNumber="14" type="description">
<paragraph id="8BEDEF256668AB5EB5D167F9FEBD39E2" blockId="13.[189,363,660,686]" box="[189,363,660,686]" pageId="13" pageNumber="14">
<heading id="D0A558496668AB5EB5D167F9FEBD39E2" bold="true" box="[189,363,660,686]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="B92633376668AB5EB5D167F9FEBD39E2" bold="true" box="[189,363,660,686]" pageId="13" pageNumber="14">Redescription</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE6668AB5EB5D167AEFAA13E87" pageId="13" pageNumber="14" type="description">
<paragraph id="8BEDEF256668AB5EB5D167AEFAA6384D" blockId="13.[189,1399,707,769]" pageId="13" pageNumber="14">
BODY. Length 4.04.5 (males) and
<quantity id="4CAA42C06668AB5EB73467AEFD303991" box="[600,742,707,733]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.7" metricValueMin="3.9" pageId="13" pageNumber="14" unit="mm" value="4.3" valueMax="4.7" valueMin="3.9">3.94.7 mm</quantity>
@ -279,8 +276,8 @@ ANTENNAE. Reaching the basal 4th part of elytra; the antennomeres 1
).
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D166DDFD9C3F5C" blockId="13.[189,1398,943,1040]" pageId="13" pageNumber="14">
PRONOTUM. Slightly transverse, maximum width at middle third part; the angles converge, smoothly from the middle area to the base in a straight line; posterior angles obtuse and rounded at vertex; sculpture and granularity well defned (
<figureCitation id="1369F3A06668AB5EB489669BFDEB3F5C" box="[485,573,1014,1040]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="13" pageNumber="14">Fig. 12</figureCitation>
PRONOTUM. Slightly transverse, maximum width at middle third part; the angles converge, smoothly from the middle area to the base in a straight line; posterior angles obtuse and rounded at vertex; sculpture and granularity well defined (
<figureCitation id="1369F3A06668AB5EB489669BFDEB3F5C" box="[485,573,1014,1040]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="13" pageNumber="14">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D16155FEDF3FD4" blockId="13.[189,1399,1079,1176]" pageId="13" pageNumber="14">
@ -315,7 +312,7 @@ Features of the genital segment and aedeagus are similar to those described for
The antennae are similar to those of the male, but the antennomeres are more transverse; 2nd, 4th, 5th and 7th antennomeres having similar length (
<tableCitation id="C6D0DA9E6668AB5EB7F26007FD2E3EC8" box="[670,760,1386,1412]" captionStart="Table 1" captionStartId="4.[188,257,267,293]" captionTargetBox="[193,1384,385,1378]" captionTargetPageId="4" captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width." httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04" pageId="13" pageNumber="14" tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
). Pronotum less elongated; the overall shape and the ratio of width/ length ratio similar to that described for males (
<figureCitation id="1369F3A06668AB5EB6C860E0FC2A3EEB" box="[932,1020,1421,1447]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="13" pageNumber="14">Fig. 13</figureCitation>
<figureCitation id="1369F3A06668AB5EB6C860E0FC2A3EEB" box="[932,1020,1421,1447]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="13" pageNumber="14">Fig. 13</figureCitation>
). Elytra elliptical and elongated with apical parts notched and serrated. 7th and 8th uroventrites, and genitalia typical of the species group.
</paragraph>
</subSubSection>
@ -337,7 +334,7 @@ is re-established as a valid species due to the clear morphological characters t
<treatmentCitation id="0AF3C9346668AB5EB6A16329FB8A3D12" author="Blas M." box="[973,1116,1604,1630]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="13" pageNumber="14" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" treatmentTaxon="unknown" type="journal article" year="1989">
<bibRefCitation id="EFC392D46668AB5EB6A16329FB8A3D12" author="Blas M." box="[973,1116,1604,1630]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="13" pageNumber="14" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas (1989)</bibRefCitation>
</treatmentCitation>
. This classifcation was subsequently followed by several authors (
. This classification was subsequently followed by several authors (
<treatmentCitation id="0AF3C9346668AB5EB7C2630AFC063DCD" author="Giachino P. M. &amp; Vailati D." box="[686,976,1639,1665]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="13" pageNumber="14" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" treatmentTaxon="unknown" type="journal article" year="1993">
<bibRefCitation id="EFC392D46668AB5EB7C2630AFC063DCD" author="Giachino P. M. &amp; Vailati D." box="[686,976,1639,1665]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="13" pageNumber="14" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
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@ -353,7 +350,7 @@ Salgado
2008
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). The main character used to re-evaluate its specifc status is the shape of the male protibial keel, which is shorter and with a more rounded ridge, but also because in
). The main character used to re-evaluate its specific status is the shape of the male protibial keel, which is shorter and with a more rounded ridge, but also because in
<taxonomicName id="4C5294A66668AB5EB63063C3FC123D8B" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[860,964,1709,1736]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B92633376668AB5EB63063C3FC123D8B" box="[860,964,1709,1736]" italics="true" pageId="13" pageNumber="14">S. breuili</emphasis>
</taxonomicName>

View file

@ -1,53 +1,53 @@
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Jeannel l936: 221.
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The specifc epithet proposed by
The specific epithet proposed by
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in the description was
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@ -133,32 +135,39 @@ has mostly been used in the entomological literature; since its use is prevalent
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);
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<materialsCitation id="3B3AE5786675AB43B40367B0FB2A39BB" ID-GBIF-Occurrence="1609370446" box="[367,1276,731,760]" collectingDate="2005-04-06" collectingDateMax="2005-04-06" collectingDateMin="1976-05-25" collectionCode="CZULE" collectorName="Ortuno" country="Spain" county="Alicante" location="Penas Roset" municipality="Xixona" pageId="16" pageNumber="17" specimenCount="12" specimenCount-female="7" specimenCount-male="5" stateProvince="Valencia">
<specimenCount id="9D5424AC6675AB43B40367B0FE6339BB" box="[367,437,731,760]" count="5" pageId="16" pageNumber="17" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D5424AC6675AB43B4A967B0FDDD39BB" box="[453,523,733,759]" count="7" pageId="16" pageNumber="17" type="female">7 ♀♀</specimenCount>
,
<collectingMunicipality id="6B89755F6675AB43B77067B0FDA339BB" box="[540,629,733,759]" pageId="16" pageNumber="17">Xixona</collectingMunicipality>
,
<location id="8E8DB9FE6675AB43B7E867B0FCCF39BB" LSID="urn:lsid:plazi:treatment:03FB5E33666AAB42B740621DFD293D2F:8E8DB9FE6675AB43B7E867B0FCCF39BB" box="[644,793,733,759]" country="Spain" county="Alicante" municipality="Xixona" name="Penas Roset" pageId="16" pageNumber="17" stateProvince="Valencia">Peñas Roset</location>
,
<date id="FFECC9E56675AB43B64B67B0FC6C39B4" box="[807,954,733,760]" pageId="16" pageNumber="17" value="2005-04-06">
<collectingDate id="EFA8300D6675AB43B64B67B0FC6C39B4" box="[807,954,733,760]" pageId="16" pageNumber="17" value="2005-04-06">6 Apr. 2005</collectingDate>
</date>
@ -166,20 +175,28 @@ leg. (
<collectorName id="26A78AF36675AB43B6A467B0FBCD39BB" box="[968,1051,733,759]" pageId="16" pageNumber="17">Ortuño</collectorName>
leg. (
<collectionCode id="ED4377E06675AB43B10F67B0FB1E39BB" box="[1123,1224,733,759]" pageId="16" pageNumber="17">CZULE</collectionCode>
). —
). —
</materialsCitation>
<materialsCitation id="3B3AE5786675AB43B06A67B0FC453856" ID-GBIF-Occurrence="1609370446" collectingDate="1976-05-25" collectingDateMax="2005-04-06" collectingDateMin="1976-05-25" collectionCode="CMCN" collectorName="Zaragoza" country="Spain" county="Alicante" location="Bocairent" municipality="Xixona" pageId="16" pageNumber="17" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Valencia">
<collectingRegion id="499621C76675AB43B06A67B0FAA539BB" box="[1286,1395,733,759]" country="Spain" name="Comunitat Valenciana" pageId="16" pageNumber="17">Valencia</collectingRegion>
: 3 ♁♁,
<specimenCount id="9D5424AC6675AB43B478666DFE923856" box="[276,324,768,794]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, Bocairent, Cova Sarsa,
:
<specimenCount id="9D5424AC6675AB43B5D1666DFED53856" box="[189,259,766,794]" count="3" pageId="16" pageNumber="17" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D5424AC6675AB43B478666DFE923856" box="[276,324,768,794]" count="1" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<location id="8E8DB9FE6675AB43B439666DFE1C3856" LSID="urn:lsid:plazi:treatment:03FB5E33666AAB42B740621DFD293D2F:8E8DB9FE6675AB43B439666DFE1C3856" box="[341,458,768,794]" country="Spain" county="Alicante" municipality="Xixona" name="Bocairent" pageId="16" pageNumber="17" stateProvince="Valencia">Bocairent</location>
, Cova Sarsa,
<collectorName id="26A78AF36675AB43B71F666DFD363856" box="[627,736,768,794]" pageId="16" pageNumber="17">Zaragoza</collectorName>
leg. (
<collectionCode id="ED4377E06675AB43B645666DFC503856" box="[809,902,768,794]" pageId="16" pageNumber="17">CMCN</collectionCode>
);
<specimenCount id="9D5424AC6675AB43B6F2666DFC1B3856" box="[926,973,768,794]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
);
</materialsCitation>
<materialsCitation id="3B3AE5786675AB43B6F2666DFD513871" ID-GBIF-Occurrence="1609370446" collectingDate="1993-03-08" collectingDateMax="2005-04-06" collectingDateMin="1976-05-25" collectionCode="CZULE" collectorName="Montagud" country="Spain" county="Alicante" location="Sima de la Carrasquilla" municipality="Enguera" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1" stateProvince="Valencia">
<specimenCount id="9D5424AC6675AB43B6F2666DFC1B3856" box="[926,973,768,794]" count="1" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectorName id="26A78AF36675AB43B6B3666DFB923856" box="[991,1092,768,794]" pageId="16" pageNumber="17">Enguera</collectorName>
<collectingMunicipality id="6B89755F6675AB43B6B3666DFB923856" box="[991,1092,768,794]" pageId="16" pageNumber="17">Enguera</collectingMunicipality>
,
<collectorName id="26A78AF36675AB43B13F666DFAA43856" box="[1107,1394,768,794]" pageId="16" pageNumber="17">Sima de la Carrasquilla</collectorName>
<location id="8E8DB9FE6675AB43B13F666DFAA43856" LSID="urn:lsid:plazi:treatment:03FB5E33666AAB42B740621DFD293D2F:8E8DB9FE6675AB43B13F666DFAA43856" box="[1107,1394,768,794]" country="Spain" county="Alicante" municipality="Enguera" name="Sima de la Carrasquilla" pageId="16" pageNumber="17" stateProvince="Valencia">Sima de la Carrasquilla</location>
,
<date id="FFECC9E56675AB43B5D1664EFE983871" box="[189,334,803,830]" pageId="16" pageNumber="17" value="1993-03-08">
<collectingDate id="EFA8300D6675AB43B5D1664EFE983871" box="[189,334,803,830]" pageId="16" pageNumber="17" value="1993-03-08">8 Mar. 1993</collectingDate>
@ -335,8 +352,8 @@ ANTENNAE. Long and slender, when oriented back reaching sixth basal part of elyt
</paragraph>
<paragraph id="8BEDEF256675AB43B5D163FBFAA13C55" blockId="16.[189,1399,1686,1818]" pageId="16" pageNumber="17">
PRONOTUM. Slightly transverse, maximum width / length ratio = 1.191.30 (males) and 1.13 1.20 (females); hexagonal in dorsal view, with maximum width in middle; sides arcuate and almost linear converging towards the base (
<figureCitation id="1369F3A06675AB43B74863B1FDAA3DBA" box="[548,636,1756,1782]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="16" pageNumber="17">Fig. 14</figureCitation>
); posterior angles obtuse with rounded apex. Base of pronotum almost as wide as base of elytra, gently curved sideways and fnely beaded. Median granularity sculpture.
<figureCitation id="1369F3A06675AB43B74863B1FDAA3DBA" box="[548,636,1756,1782]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="16" pageNumber="17">Fig. 14</figureCitation>
); posterior angles obtuse with rounded apex. Base of pronotum almost as wide as base of elytra, gently curved sideways and finely beaded. Median granularity sculpture.
</paragraph>
<paragraph id="8BEDEF256675AB43B5D1622FFAA13CEE" blockId="16.[189,1399,1857,1954]" pageId="16" pageNumber="17">ELYTRA. Elliptical and very elongated, maximum length / width ratio = 2.062.18 (males) and 1.96 2.06 (female), with rounded apical areas; elytral disc slightly convex and depressed along suture in its half basal part; elytral suture well-marked with thin transverse striae, distant and perpendicular to suture.</paragraph>
<paragraph id="8BEDEF256674AB42B5866466FEFC3A69" blockId="17.[189,1399,267,434]" box="[234,298,267,293]" pageId="17" pageNumber="18">
@ -369,7 +386,7 @@ GENITAL SEGMENT STRUCTURE. Similar to that described for
<paragraph id="8BEDEF256674AB42B5D16776FEA4398D" blockId="17.[189,1398,538,705]" pageId="17" pageNumber="18">
LEGS. Relatively long and slender; protibia widened in apical area and with a long keel running through the two basal thirds of the margin, showing two sharp protrusions, separated by a slight depression, anterior is sharp and much higher and the second is fuzzy (
<figureCitation id="1369F3A06674AB42B60B670CFC683937" box="[871,958,609,635]" captionStart="Figs 16 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="17" pageNumber="18">Fig. 20</figureCitation>
); frst three protarsomeres are dilated, the frst being slightly narrower than the anterior tip of protibia; mesotibiae arched and metatibiae slightly arched.
); first three protarsomeres are dilated, the first being slightly narrower than the anterior tip of protibia; mesotibiae arched and metatibiae slightly arched.
</paragraph>
<paragraph id="8BEDEF256674AB42B5866784FE93384F" blockId="17.[189,1399,745,1018]" box="[234,325,745,771]" pageId="17" pageNumber="18">
<heading id="D0A558496674AB42B5866784FE93384F" bold="true" box="[234,325,745,771]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="6">
@ -380,7 +397,7 @@ LEGS. Relatively long and slender; protibia widened in apical area and with a lo
Antennae long, reaching the basal third of elytra, with antennomeres generally slightly longer and transverse (
<tableCitation id="C6D0DA9E6674AB42B42B6642FE723805" box="[327,420,815,841]" captionStart="Table 1" captionStartId="4.[188,257,267,293]" captionTargetBox="[193,1384,385,1378]" captionTargetPageId="4" captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width." httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04" pageId="17" pageNumber="18" tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
). Pronotum not so transverse; in dorsal view subrectangular, with side margins almost straight before maximum width; which is situated towards the third part of basal area (
<figureCitation id="1369F3A06674AB42B07D663FFABE3821" box="[1297,1384,850,877]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
<figureCitation id="1369F3A06674AB42B07D663FFABE3821" box="[1297,1384,850,877]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
). Elytra elliptical and elongate, with notched and serrated apical areas. Sexual dimorphism very clear in pronotum, protibiae and apical part of elytra. Females are generally less robust and with less transverse pronotum than males. The uroventrites 7th and 8th, and spermatheca follows the overall model for the group.
</paragraph>
</subSubSection>
@ -415,7 +432,7 @@ The great variability of this species led to the description of the subspecies
<taxonomicName id="4C5294A66674AB42B5D16044FE073E0F" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[189,465,1321,1347]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="escalerai">
<emphasis id="B92633376674AB42B5D16044FE073E0F" box="[189,465,1321,1347]" italics="true" pageId="17" pageNumber="18">Speonemadus escalerai</emphasis>
</taxonomicName>
is a subterranean species and, therefore, common in caves. It is mostly found in areas with small accumulations of bat guano, but also under stones or stalagmitic foors and in deep parts of caves (
is a subterranean species and, therefore, common in caves. It is mostly found in areas with small accumulations of bat guano, but also under stones or stalagmitic floors and in deep parts of caves (
<bibRefCitation id="EFC392D46674AB42B4406002FE713EC5" author="Blas M." box="[300,423,1391,1417]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="17" pageNumber="18" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas 1989</bibRefCitation>
).
<bibRefCitation id="EFC392D46674AB42B4D66002FD903EC5" author="Blas M." box="[442,582,1391,1417]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="17" pageNumber="18" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas (1989)</bibRefCitation>
@ -427,7 +444,7 @@ Fresneda
<emphasis id="B92633376674AB42B144601DFBB43EC5" box="[1064,1122,1391,1417]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
(2007)
</bibRefCitation>
also pointed out its presence in the Superfcial Subterranean Habitat (MSS) (
also pointed out its presence in the Superficial Subterranean Habitat (MSS) (
<emphasis id="B92633376674AB42B61060FEFC6B3EE0" box="[892,957,1427,1452]" italics="true" pageId="17" pageNumber="18">sensu</emphasis>
<bibRefCitation id="EFC392D46674AB42B6A860FFFB333EE0" author="Giachino &amp; Vailati" box="[964,1253,1426,1452]" firstAuthor="Giachino" pageId="17" pageNumber="18" pagination="3" refId="ref12117" refString="Giachino P. M. &amp; Vailati D. 2010. The subterranean environment. Hypogean life, concepts and collecting techniques. WBA Handbooks 3, World Biodiversity Association, Verona." type="book chapter" year="2010">Giachino &amp; Vailati 2010</bibRefCitation>
) in the karst area of
@ -442,7 +459,6 @@ also pointed out its presence in the Superfcial Subterranean Habitat (MSS) (
</heading>
</paragraph>
<paragraph id="8BEDEF256674AB42B5D16348FD293D2F" blockId="17.[189,1398,1573,1635]" pageId="17" pageNumber="18">
<materialsCitation id="3B3AE5786674AB42B5D16348FD293D2F" ID-GBIF-Occurrence="1609370441" country="Spain" location="Murcia" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Murcia">
This species is distributed in the southeast Iberian Peninsula and colonizes caves of the provinces of Alicante,
<collectingRegion id="499621C76674AB42B4436324FE523D2F" box="[303,388,1609,1635]" country="Spain" name="Murcia" pageId="17" pageNumber="18">Murcia</collectingRegion>
and Valencia in
@ -450,7 +466,6 @@ and Valencia in
(
<figureCitation id="1369F3A06674AB42B7F66324FD273D2F" box="[666,753,1609,1635]" captionStart="Fig. 32" captionStartId="18.[189,232,1955,1981]" captionTargetBox="[189,1398,986,1883]" captionTargetId="figure@18.[189,1398,986,1914]" captionTargetPageId="18" captionText="Fig. 32. Distribution of Speonemadus of the escalerai - group: S. algarvensis sp. nov., S. angusticollis (Kraatz, 1870), S. bolivari (Jeannel, 1922), S. breuili (Jeannel, 1922) and S. escalerai (Uhagón, 1898)." httpUri="https://zenodo.org/record/889358/files/figure.png" pageId="17" pageNumber="18">Fig. 32</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>

View file

@ -1,53 +1,53 @@
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<treatment id="03FB5E33666BAB5CB75A67F7FABB3C05" ID-DOI="http://doi.org/10.5281/zenodo.5689574" ID-GBIF-Taxon="133183537" ID-Zenodo-Dep="5689574" LSID="urn:lsid:plazi:treatment:03FB5E33666BAB5CB75A67F7FABB3C05" httpUri="http://treatment.plazi.org/id/03FB5E33666BAB5CB75A67F7FABB3C05" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<treatment id="03FB5E33666BAB5CB75A67F7FABB3C05" ID-DOI="http://doi.org/10.5281/zenodo.5689574" ID-GBIF-Taxon="133183537" ID-Zenodo-Dep="5689574" LSID="urn:lsid:plazi:treatment:03FB5E33666BAB5CB75A67F7FABB3C05" httpUri="http://treatment.plazi.org/id/03FB5E33666BAB5CB75A67F7FABB3C05" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" scope_family="Leiodidae" scope_order="Coleoptera" scope_subFamily="Cholevinae" scope_tribe="Anemadini">
<subSubSection id="C348BCAE666BAB5DB75A67F7FC2839F9" box="[566,1022,666,693]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BEDEF25666BAB5DB75A67F7FC2839F9" blockId="14.[566,1022,666,693]" box="[566,1022,666,693]" pageId="14" pageNumber="15">
<heading id="D0A55849666BAB5DB75A67F7FC2839F9" box="[566,1022,666,693]" centered="true" fontSize="11" level="3" pageId="14" pageNumber="15" reason="4">
@ -108,7 +108,7 @@
<collectingCountry id="F345AFB5666BAB5DB5D1614CFEC23F77" box="[189,276,1057,1083]" name="Spain" pageId="14" pageNumber="15">SPAIN</collectingCountry>
:
<location id="8E8DB9FE666BAB5DB44E614CFE5C3F77" LSID="urn:lsid:plazi:treatment:03FB5E33666BAB5CB75A67F7FABB3C05:8E8DB9FE666BAB5DB44E614CFE5C3F77" box="[290,394,1057,1083]" country="Spain" county="Sima San Rafael" municipality="Cerro de la Mesa" name="Granada" pageId="14" pageNumber="15">Granada</location>
: 10 ♁♁,
: 10 ♂♂,
<specimenCount id="9D5424AC666BAB5DB76C614CFD833F77" box="[512,597,1057,1083]" pageId="14" pageNumber="15" type="female">10 ♀♀</specimenCount>
,
<location id="8E8DB9FE666BAB5DB70B6172FD7F3F77" LSID="urn:lsid:plazi:treatment:03FB5E33666BAB5CB75A67F7FABB3C05:8E8DB9FE666BAB5DB70B6172FD7F3F77" box="[615,681,1055,1083]" country="Spain" county="Sima San Rafael" municipality="Cerro de la Mesa" name="Illora" pageId="14" pageNumber="15">Íllora</location>
@ -129,8 +129,8 @@
)
</materialsCitation>
;
<materialsCitation id="3B3AE578666BAB5DB7116128FCE43FCD" ID-GBIF-Occurrence="1609370439" collectingDate="15 Dec. 2012 and 10 Mar. 2013" collectionCode="CFL, CPB, CJMS, CZULE" country="Spain" location="Pinar" municipality="Cueva de Pagarrecio" pageId="14" pageNumber="15" specimenCount="1">
many ♁♁ and
<materialsCitation id="3B3AE578666BAB5DB7116128FCE43FCD" ID-GBIF-Occurrence="1609370439" collectingDate="2012-12-15" collectionCode="CFL, CPB, CJMS, CZULE" country="Spain" location="Pinar" municipality="Cueva de Pagarrecio" pageId="14" pageNumber="15" specimenCount="1">
many ♂♂ and
<specimenCount id="9D5424AC666BAB5DB6446128FC833F13" box="[808,853,1093,1119]" pageId="14" pageNumber="15">♀♀</specimenCount>
,
<location id="8E8DB9FE666BAB5DB60D6129FC773F12" LSID="urn:lsid:plazi:treatment:03FB5E33666BAB5CB75A67F7FABB3C05:8E8DB9FE666BAB5DB60D6129FC773F12" box="[865,929,1092,1118]" country="Spain" municipality="Cueva de Pagarrecio" name="Pinar" pageId="14" pageNumber="15">Piñar</location>
@ -156,7 +156,7 @@ and
<materialsCitation id="3B3AE578666BAB5DB6506105FCFF3FE9" ID-GBIF-Occurrence="1609370438" collectingDate="2006-10-17" collectionCode="CFL" collectorName="Barranco &amp; Ruiz-Aviles" country="Spain" county="Cueva de Ardales" location="Malaga" municipality="Ardales" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1">
<location id="8E8DB9FE666BAB5DB60C610AFC6B3FCD" LSID="urn:lsid:plazi:treatment:03FB5E33666BAB5CB75A67F7FABB3C05:8E8DB9FE666BAB5DB60C610AFC6B3FCD" box="[864,957,1127,1153]" country="Spain" county="Cueva de Ardales" municipality="Ardales" name="Malaga" pageId="14" pageNumber="15">Málaga</location>
: 1 ,
: 1 ,
<specimenCount id="9D5424AC666BAB5DB16D610AFBFA3FCE" box="[1025,1068,1127,1154]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="6B89755F666BAB5DB154610AFB403FCD" box="[1080,1174,1127,1153]" pageId="14" pageNumber="15">Ardales</collectingMunicipality>
@ -176,7 +176,7 @@ leg. (
</materialsCitation>
;
<materialsCitation id="3B3AE578666BAB5DB65A61E7FCBC3F84" ID-GBIF-Occurrence="1609370457" collectingDate="2007-01-17" collectionCode="CPB, CJMS" collectorName="Barranco &amp; Ruiz-Aviles" country="Trinidad and Tobago" location="Trinidad" pageId="14" pageNumber="15" specimenCount="8" specimenCount-female="8">
6 ♁♁,
6 ♂♂,
<specimenCount id="9D5424AC666BAB5DB6E761E6FC063FE9" box="[907,976,1163,1189]" pageId="14" pageNumber="15" type="female">8 ♀♀</specimenCount>
, Ardales, Cueva de Doña
<collectingCountry id="F345AFB5666BAB5DB06761E7FAA43FE8" box="[1291,1394,1162,1188]" name="Trinidad and Tobago" pageId="14" pageNumber="15">Trinidad</collectingCountry>
@ -195,8 +195,8 @@ leg. (
)
</materialsCitation>
;
<materialsCitation id="3B3AE578666BAB5DB61961C3FE7E3E42" ID-GBIF-Occurrence="1609370448" collectingDate="28 Nov. 2009 and 30 Sep. 2010" collectionCode="CFL, CPB, CJMS" country="Trinidad and Tobago" location="Higuerones" municipality="Gobantes-Meliones" pageId="14" pageNumber="15" specimenCount="1">
many ♁♁ and
<materialsCitation id="3B3AE578666BAB5DB61961C3FE7E3E42" ID-GBIF-Occurrence="1609370448" collectingDate="2009-11-28" collectionCode="CFL, CPB, CJMS" country="Trinidad and Tobago" location="Higuerones" municipality="Gobantes-Meliones" pageId="14" pageNumber="15" specimenCount="1">
many ♂♂ and
<specimenCount id="9D5424AC666BAB5DB14C61C3FB9B3F84" box="[1056,1101,1198,1224]" pageId="14" pageNumber="15">♀♀</specimenCount>
, Antequera, Cueva de los
<location id="8E8DB9FE666BAB5DB5D161BCFE923FA7" LSID="urn:lsid:plazi:treatment:03FB5E33666BAB5CB75A67F7FABB3C05:8E8DB9FE666BAB5DB5D161BCFE923FA7" box="[189,324,1233,1259]" country="Trinidad and Tobago" municipality="Gobantes-Meliones" name="Higuerones" pageId="14" pageNumber="15">Higuerones</location>
@ -218,7 +218,7 @@ and
</materialsCitation>
;
<materialsCitation id="3B3AE578666BAB5DB4D96199FE863E7D" ID-GBIF-Occurrence="1609370436" collectingDate="2007-01-17" collectionCode="CJMS" collectorName="Barranco" country="Trinidad and Tobago" county="Cueva de la Pileta" location="Benaojan" municipality="Sierra de Grazalema" pageId="14" pageNumber="15" specimenCount="1">
4 ♁♁,
4 ♂♂,
<location id="8E8DB9FE666BAB5DB76A6199FDAE3E42" LSID="urn:lsid:plazi:treatment:03FB5E33666BAB5CB75A67F7FABB3C05:8E8DB9FE666BAB5DB76A6199FDAE3E42" box="[518,632,1268,1294]" country="Trinidad and Tobago" county="Cueva de la Pileta" municipality="Sierra de Grazalema" name="Benaojan" pageId="14" pageNumber="15">Benaoján</location>
,
<collectingMunicipality id="6B89755F666BAB5DB7EF6199FCA03E42" box="[643,886,1268,1294]" pageId="14" pageNumber="15">Sierra de Grazalema</collectingMunicipality>
@ -236,7 +236,7 @@ leg. (
</materialsCitation>
;
<materialsCitation id="3B3AE578666BAB5DB437607AFB723E7D" ID-GBIF-Occurrence="1609370443" box="[347,1188,1301,1329]" collectingDate="1977-07-31" collectionCode="CFL" collectorName="Abad" country="Trinidad and Tobago" location="Botijos" pageId="14" pageNumber="15" specimenCount="1">
1 , Benalmádena, Cueva de los
1 , Benalmádena, Cueva de los
<location id="8E8DB9FE666BAB5DB78C607AFCEE3E7D" LSID="urn:lsid:plazi:treatment:03FB5E33666BAB5CB75A67F7FABB3C05:8E8DB9FE666BAB5DB78C607AFCEE3E7D" box="[736,824,1303,1329]" country="Trinidad and Tobago" name="Botijos" pageId="14" pageNumber="15">Botijos</location>
,
<date id="FFECC9E5666BAB5DB62E607AFC023E7D" box="[834,980,1303,1329]" pageId="14" pageNumber="15" value="1977-07-31">
@ -255,7 +255,7 @@ leg. (
<date id="FFECC9E5666BAB5DB4376056FE2E3E19" box="[347,504,1339,1365]" pageId="14" pageNumber="15" value="2009-09-28">
<collectingDate id="EFA8300D666BAB5DB4376056FE2E3E19" box="[347,504,1339,1365]" pageId="14" pageNumber="15" value="2009-09-28">28 Sep. 2009</collectingDate>
</date>
; 1 ♁♁ and
; 1 ♂♂ and
<specimenCount id="9D5424AC666BAB5DB7136056FD143E19" box="[639,706,1339,1365]" pageId="14" pageNumber="15" type="female">3 ♀♀</specimenCount>
, same locality,
<date id="FFECC9E5666BAB5DB61B6057FBDC3E19" box="[887,1034,1338,1365]" pageId="14" pageNumber="15" value="2008-07-25">
@ -398,13 +398,13 @@ ANTENNAE. Long, slender, light brown and extended beyond the basal fourth of ely
PRONOTUM. Slightly transverse, maximum width/ length ratio = 1.231.39 (males) and
<quantity id="4CAA42C0666AAB5CB1A664FEFAA13AE2" box="[1226,1399,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" metricValueMax="1.36" metricValueMin="1.28" pageId="15" pageNumber="16" unit="mm" value="1.32" valueMax="1.36" valueMin="1.28">1.281.36 mm</quantity>
(females); with a maximum width towards the middle; basal impressions are not hardly noticeable; side edges regularly arched forward, almost straight and towards the posterior angles that are obtuse and rounded in the vertex (
<figureCitation id="1369F3A0666AAB5CB4AB6490FDC8395B" box="[455,542,509,535]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
<figureCitation id="1369F3A0666AAB5CB4AB6490FDC8395B" box="[455,542,509,535]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
); pronotal base as wide as elytral base; sculpture has a coarse granularity with obvious microsculpture.
</paragraph>
<paragraph id="8BEDEF25666AAB5CB5D1670FFC27398E" blockId="15.[189,1399,609,706]" pageId="15" pageNumber="16">
ELYTRA. Elliptical and elongated with rounded apical areas, maximum length / width ratio = 1.95 2.04 (males) and
<quantity id="4CAA42C0666AAB5CB4E167E8FDEF39D3" box="[397,569,645,671]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" metricValueMax="1.98" metricValueMin="1.88" pageId="15" pageNumber="16" unit="mm" value="1.93" valueMax="1.98" valueMin="1.88">1.881.98 mm</quantity>
(females); elytral disc is slightly convex, and briefy depressed along suture at half of basal part. Sutural stria and discal striae well marked.
(females); elytral disc is slightly convex, and briefly depressed along suture at half of basal part. Sutural stria and discal striae well marked.
</paragraph>
<paragraph id="8BEDEF25666AAB5CB5866784FEFC384F" blockId="15.[189,1399,745,1124]" box="[234,298,745,771]" pageId="15" pageNumber="16">
<heading id="D0A55849666AAB5CB5866784FEFC384F" bold="true" box="[234,298,745,771]" fontSize="11" level="2" pageId="15" pageNumber="16" reason="6">
@ -423,12 +423,12 @@ Aedeagus long,
<quantity id="4CAA42C0666AAB5CB6D0665DFBCC3806" box="[956,1050,816,842]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="16" unit="mm" value="1.1">1.1 mm</quantity>
; basal lamina of median lobe well developed and more than half of the median lobe length; ventral blade of tegmen short, inconsistent and forms the widest part of the aedeagus. In dorsal view, median lobe is bottle-shaped, with converging sides towards the base and narrowed distally; apex tapers and ends in a triangular tip with the sides slightly beaded dorsally (
<figureCitation id="1369F3A0666AAB5CB49A66D1FD82389B" box="[502,596,956,983]" captionStart="Figs 21 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="15" pageNumber="16">Fig. 23</figureCitation>
). Parameres robust, only curved forwards; apical parts obliquely truncated on the inner side, where four setae are inserted, three thin, subequal, and a larger one, as well as a thick and well sclerotized small tooth. Inner sac with two longitudinal chains formed by small sclerotized parts and surrounded by two longer chains of fne and sharp spines that come together in the apical region, but do not form a typical sclerotized tooth as in other species.
). Parameres robust, only curved forwards; apical parts obliquely truncated on the inner side, where four setae are inserted, three thin, subequal, and a larger one, as well as a thick and well sclerotized small tooth. Inner sac with two longitudinal chains formed by small sclerotized parts and surrounded by two longer chains of fine and sharp spines that come together in the apical region, but do not form a typical sclerotized tooth as in other species.
</paragraph>
<paragraph id="8BEDEF25666AAB5CB5D161E6FBF53FA7" blockId="15.[189,1399,1163,1259]" pageId="15" pageNumber="16">
LEGS. Quite long; the protibial keel is elevated in the anterior middle third, showing a single, well marked, projection angle (
<figureCitation id="1369F3A0666AAB5CB48361C3FD933F84" box="[495,581,1198,1224]" captionStart="Figs 16 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="15" pageNumber="16">Fig. 18</figureCitation>
); frst three protarsomeres dilated, being the frst almost as wide as the apical tip of the protibia; mesotibia strongly arched and metatibia straight.
); first three protarsomeres dilated, being the first almost as wide as the apical tip of the protibia; mesotibia strongly arched and metatibia straight.
</paragraph>
<paragraph id="8BEDEF25666AAB5CB586607EFE933E61" blockId="15.[189,1399,1299,1501]" box="[234,325,1299,1325]" pageId="15" pageNumber="16">
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@ -439,7 +439,7 @@ LEGS. Quite long; the protibial keel is elevated in the anterior middle third, s
Antennae slightly shorter and more robust than in the male, reaching third basal part of elytra; 2nd antennomere is slightly longer than the 3rd one and 5th one longer than 4th and 6th ones; the 8th being clearly longer than wide (
<tableCitation id="C6D0DA9E666AAB5CB4FE6011FE3C3EDA" box="[402,490,1404,1430]" captionStart="Table 1" captionStartId="4.[188,257,267,293]" captionTargetBox="[193,1384,385,1378]" captionTargetPageId="4" captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width." httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04" pageId="15" pageNumber="16" tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
). Pronotum less transverse, but appears longer (
<figureCitation id="1369F3A0666AAB5CB14E6011FBAF3EDA" box="[1058,1145,1404,1430]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="15" pageNumber="16">Fig. 11</figureCitation>
<figureCitation id="1369F3A0666AAB5CB14E6011FBAF3EDA" box="[1058,1145,1404,1430]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="15" pageNumber="16">Fig. 11</figureCitation>
). Elytra elliptical and elongated with notched and serrated apical areas. The sexual dimorphism is clear for all species of this group, as well as the structures in uroventrites 7th and 8th, and spermatheca model.
</paragraph>
</subSubSection>

View file

@ -1,53 +1,53 @@
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.
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<taxonomicName id="4C5294A6666FAB59B5D1677AFE7B397C" box="[189,429,534,561]" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="angusticollis">
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<emphasis id="B9263337666FAB59B5D1677AFE7B397C" box="[189,429,534,561]" italics="true" pageId="10" pageNumber="11">Catops angusticollis</emphasis>
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<paragraph id="8BEDEF25666FAB59B5D1662DFCD03F91" blockId="10.[189,1399,830,1245]" pageId="10" pageNumber="11">
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<materialsCitation id="3B3AE578666FAB59B5D1662DFCF03831" ID-GBIF-Occurrence="2432507016" collectingDate="2009-11-22" collectionCode="CFL, CPB, CJMS, CZULE" country="Spain" county="Espino" location="Cordoba" municipality="Sima de Fuente" pageId="10" pageNumber="11" specimenCount="1">
<collectingCountry id="F345AFB5666FAB59B5D1662DFEC23816" box="[189,276,832,858]" name="Spain" pageId="10" pageNumber="11">SPAIN</collectingCountry>
:
<location id="8E8DB9FE666FAB59B44C6652FE5C3815" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B44C6652FE5C3815" box="[288,394,831,857]" country="Spain" county="Espino" municipality="Sima de Fuente" name="Cordoba" pageId="10" pageNumber="11">Córdoba</location>
: many ♁♁ and
: many ♂♂ and
<specimenCount id="9D5424AC666FAB59B726662DFDA13816" box="[586,631,832,858]" pageId="10" pageNumber="11">♀♀</specimenCount>
,
<location id="8E8DB9FE666FAB59B7EA662DFD023816" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B7EA662DFD023816" box="[646,724,832,858]" country="Spain" county="Espino" municipality="Sima de Fuente" name="Luque" pageId="10" pageNumber="11">Luque</location>
@ -142,7 +142,7 @@ and
</materialsCitation>
;
<materialsCitation id="3B3AE578666FAB59B65E660EFE0238EC" ID-GBIF-Occurrence="2432507017" collectingDate="2010-04-03" collectionCode="CFL" country="Spain" location="Sima Fuente de Alhama" pageId="10" pageNumber="11" specimenCount="2" specimenCount-female="2">
1 ,
1 ,
<specimenCount id="9D5424AC666FAB59B601660EFC663831" box="[877,944,867,893]" pageId="10" pageNumber="11" type="female">2 ♀♀</specimenCount>
,
<location id="8E8DB9FE666FAB59B6D2660EFB0C3831" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B6D2660EFB0C3831" box="[958,1242,867,893]" country="Spain" name="Sima Fuente de Alhama" pageId="10" pageNumber="11">Sima Fuente de Alhama</location>
@ -155,8 +155,8 @@ and
)
</materialsCitation>
;
<materialsCitation id="3B3AE578666FAB59B48D66EAFCFF388F" ID-GBIF-Occurrence="2432507010" collectingDate="6 Dec. 2009 and 7 Feb. 2010" collectionCode="CFL, CPB, CJMS" collectorName="Cueva de la Solana" country="Spain" location="Priego de Cordoba" pageId="10" pageNumber="11" specimenCount="1">
many ♁♁ and
<materialsCitation id="3B3AE578666FAB59B48D66EAFCFF388F" ID-GBIF-Occurrence="2432507010" collectingDate="2009-12-06" collectionCode="CFL, CPB, CJMS" collectorName="Cueva de la Solana" country="Spain" location="Priego de Cordoba" pageId="10" pageNumber="11" specimenCount="1">
many ♂♂ and
<specimenCount id="9D5424AC666FAB59B7FF66EAFD1638ED" box="[659,704,903,929]" pageId="10" pageNumber="11">♀♀</specimenCount>
,
<location id="8E8DB9FE666FAB59B7A266EBFC6638EC" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B7A266EBFC6638EC" box="[718,944,902,928]" country="Spain" name="Priego de Cordoba" pageId="10" pageNumber="11">Priego de Córdoba</location>
@ -180,7 +180,7 @@ and
<materialsCitation id="3B3AE578666FAB59B65566C7FD2238AA" ID-GBIF-Occurrence="2432507013" collectingDate="1994-04-10" collectionCode="CZULE" collectorName="Carabajal" country="Spain" county="Cueva de Remigin" location="Guadalajara" municipality="Tamajon" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1">
<location id="8E8DB9FE666FAB59B60E66C4FC22388F" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B60E66C4FC22388F" box="[866,1012,937,963]" country="Spain" county="Cueva de Remigin" municipality="Tamajon" name="Guadalajara" pageId="10" pageNumber="11">Guadalajara</location>
: 1 ,
: 1 ,
<specimenCount id="9D5424AC666FAB59B12B66C4FBAE3888" box="[1095,1144,937,964]" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="6B89755F666FAB59B1E766C4FB22388F" box="[1163,1268,937,963]" pageId="10" pageNumber="11">Tamajón</collectingMunicipality>
@ -216,7 +216,7 @@ CDP leg. (
</materialsCitation>
;
<materialsCitation id="3B3AE578666FAB59B7C2669DFEC53F61" ID-GBIF-Occurrence="2432507020" collectingDate="2013-06-05" collectionCode="CPB, CJMS" country="Spain" location="Huelva" municipality="Guerrero" pageId="10" pageNumber="11" specimenCount="4" specimenCount-female="4">
3 ♁♁,
3 ♂♂,
<specimenCount id="9D5424AC666FAB59B66B669DFC983F46" box="[775,846,1008,1034]" pageId="10" pageNumber="11" type="female">4 ♀♀</specimenCount>
, same locality,
<date id="FFECC9E5666FAB59B17D6682FB753F46" box="[1041,1187,1007,1034]" pageId="10" pageNumber="11" value="2013-06-05">
@ -232,7 +232,7 @@ CDP leg. (
<materialsCitation id="3B3AE578666FAB59B44C617EFEC53F1C" ID-GBIF-Occurrence="2432507011" collectingDate="2009-11-15" collectionCode="CFL, CJMS" country="Spain" county="Cueva de la Murcielaguina" location="Jaen" municipality="Hornos" pageId="10" pageNumber="11" specimenCount="28" specimenCount-female="28">
<location id="8E8DB9FE666FAB59B42A617EFEA93F61" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B42A617EFEA93F61" box="[326,383,1043,1069]" country="Spain" county="Cueva de la Murcielaguina" municipality="Hornos" name="Jaen" pageId="10" pageNumber="11">Jaén</location>
: 11 ♁♁,
: 11 ♂♂,
<specimenCount id="9D5424AC666FAB59B480617EFDE93F61" box="[492,575,1043,1069]" pageId="10" pageNumber="11" type="female">28 ♀♀</specimenCount>
,
<collectingMunicipality id="6B89755F666FAB59B722617EFD713F61" box="[590,679,1043,1069]" pageId="10" pageNumber="11">Hornos</collectingMunicipality>
@ -250,7 +250,7 @@ CDP leg. (
</materialsCitation>
;
<materialsCitation id="3B3AE578666FAB59B44D615BFE643F3F" ID-GBIF-Occurrence="2432507012" collectingDate="1953-05-30" collectionCode="CMCN" collectorName="Mateu &amp; Cobos" country="Spain" county="Sagreo" location="La Iruela" municipality="Cueva Secreta" pageId="10" pageNumber="11" specimenCount="4" specimenCount-female="4">
5 ♁♁,
5 ♂♂,
<specimenCount id="9D5424AC666FAB59B41F615BFE613F1D" box="[371,439,1078,1105]" pageId="10" pageNumber="11" type="female">4 ♀♀</specimenCount>
,
<location id="8E8DB9FE666FAB59B4AA615AFDE23F1C" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B4AA615AFDE23F1C" box="[454,564,1078,1105]" country="Spain" county="Sagreo" municipality="Cueva Secreta" name="La Iruela" pageId="10" pageNumber="11">La Iruela</location>
@ -274,7 +274,7 @@ leg. (
</materialsCitation>
;
<materialsCitation id="3B3AE578666FAB59B4AC6134FDD13FDA" ID-GBIF-Occurrence="2432507014" collectingDate="1964-08-08" collectionCode="CMCN" collectorName="Castells" country="Spain" county="Nacimiento de Guadalquivir" location="Sierra de Cazorla" municipality="Cueva" pageId="10" pageNumber="11" specimenCount="2" specimenCount-female="2">
4 ♁♁,
4 ♂♂,
<specimenCount id="9D5424AC666FAB59B7786134FD8E3F38" box="[532,600,1113,1140]" pageId="10" pageNumber="11" type="female">2 ♀♀</specimenCount>
,
<location id="8E8DB9FE666FAB59B70B6134FCEF3F3F" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B70B6134FCEF3F3F" box="[615,825,1113,1139]" country="Spain" county="Nacimiento de Guadalquivir" municipality="Cueva" name="Sierra de Cazorla" pageId="10" pageNumber="11">Sierra de Cazorla</location>
@ -293,8 +293,8 @@ leg. (
)
</materialsCitation>
;
<materialsCitation id="3B3AE578666FAB59B77E6110FCEE3FF6" ID-GBIF-Occurrence="2432507015" collectingDate="8 Sep. 2008 and 16 Jan. 2010" collectionCode="CFL, CPB, CJMS" country="Spain" county="Sima" location="Siles" municipality="Cerro de Bucentaina" pageId="10" pageNumber="11" specimenCount="1">
many ♁♁ and
<materialsCitation id="3B3AE578666FAB59B77E6110FCEE3FF6" ID-GBIF-Occurrence="2432507015" collectingDate="2008-09-08" collectionCode="CFL, CPB, CJMS" country="Spain" county="Sima" location="Siles" municipality="Cerro de Bucentaina" pageId="10" pageNumber="11" specimenCount="1">
many ♂♂ and
<specimenCount id="9D5424AC666FAB59B7D16110FD3C3FDB" box="[701,746,1149,1175]" pageId="10" pageNumber="11">♀♀</specimenCount>
,
<location id="8E8DB9FE666FAB59B79A6111FCE43FDA" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B79A6111FCE43FDA" box="[758,818,1148,1174]" country="Spain" county="Sima" municipality="Cerro de Bucentaina" name="Siles" pageId="10" pageNumber="11">Siles</location>
@ -320,7 +320,7 @@ and
<materialsCitation id="3B3AE578666FAB59B62861CDFCD43F91" ID-GBIF-Occurrence="2432507021" collectingDate="2013-01-23" collectionCode="CFL, CPB, CJMS" country="Spain" location="Sevilla" municipality="Paro" pageId="10" pageNumber="11" specimenCount="1">
<location id="8E8DB9FE666FAB59B60561F2FC693FF5" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B60561F2FC693FF5" box="[873,959,1183,1209]" country="Spain" municipality="Paro" name="Sevilla" pageId="10" pageNumber="11">Sevilla</location>
: many ♁♁ and
: many ♂♂ and
<specimenCount id="9D5424AC666FAB59B11561CDFB703FF6" box="[1145,1190,1184,1210]" pageId="10" pageNumber="11">♀♀</specimenCount>
, Alanís, Sima del
<collectingMunicipality id="6B89755F666FAB59B5D161AEFF203F91" box="[189,246,1219,1245]" pageId="10" pageNumber="11">Paro</collectingMunicipality>
@ -341,7 +341,7 @@ and
<paragraph id="8BEDEF25666FAB59B5D16069FE7E3E0E" blockId="10.[189,1398,1282,1346]" pageId="10" pageNumber="11">
<materialsCitation id="3B3AE578666FAB59B5D16069FE7E3E0E" ID-GBIF-Occurrence="2432507018" collectingDate="2009-03-30" collectorName="A. S. Reboleira" country="Portugal" county="Algar de Santo Antonio" location="Alandroal" municipality="Estremoz-Cano" pageId="10" pageNumber="11" specimenCount="1">
<collectingCountry id="F345AFB5666FAB59B5D16069FE8B3E52" box="[189,349,1284,1310]" name="Portugal" pageId="10" pageNumber="11">PORTUGAL</collectingCountry>
: 1 ,
: 1 ,
<location id="8E8DB9FE666FAB59B4C16069FDF33E52" LSID="urn:lsid:plazi:treatment:03FB5E33666FAB5FB77064FEFC693FE5:8E8DB9FE666FAB59B4C16069FDF33E52" box="[429,549,1284,1310]" country="Portugal" county="Algar de Santo Antonio" municipality="Estremoz-Cano" name="Alandroal" pageId="10" pageNumber="11">Alandroal</location>
,
<collectingMunicipality id="6B89755F666FAB59B7596069FD3D3E52" box="[565,747,1284,1310]" pageId="10" pageNumber="11">Estremoz-Cano</collectingMunicipality>
@ -651,13 +651,13 @@ BODY. Length 4.0
</paragraph>
<paragraph id="8BEDEF25666EAB58B5D166C7FCB93F61" blockId="11.[189,1399,937,1070]" pageId="11" pageNumber="12">
PRONOTUM. Slightly transverse with surface slightly punctured (
<figureCitation id="1369F3A0666EAB58B6F566C4FBD73888" box="[921,1025,937,964]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="11" pageNumber="12">Figs 89</figureCitation>
<figureCitation id="1369F3A0666EAB58B6F566C4FBD73888" box="[921,1025,937,964]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="11" pageNumber="12">Figs 89</figureCitation>
), maximum width/length ratio = 1.201.32 (males) and 1.181.25 (females), with maximum width in the middle third and the lateral edges regularly arcuate, converging towards the base; posterior angles obtuse and slightly rounded at apex. Pronotal base sinuate and narrower than elytral base.
</paragraph>
<paragraph id="8BEDEF25666EAB58B5D16138FCFD3F95" blockId="11.[189,1399,1108,1241]" pageId="11" pageNumber="12">
ELYTRA. Elliptical and very elongated, maximum length / width ratio = 2.042.19 (males) and
<quantity id="4CAA42C0666EAB58B0586138FEF53FDE" metricMagnitude="-3" metricUnit="m" metricValue="2.04" metricValueMax="2.11" metricValueMin="1.97" pageId="11" pageNumber="12" unit="mm" value="2.04" valueMax="2.11" valueMin="1.97">1.97 2.11 mm</quantity>
(females); apex of each elytron rounded; elytral disk slightly convex and fattened in the middle. Elytral sculpture formed by very strong sutural striae and transverse strias with regular arrangement, sunken, well marked and perpendicular to the suture.
(females); apex of each elytron rounded; elytral disk slightly convex and flattened in the middle. Elytral sculpture formed by very strong sutural striae and transverse strias with regular arrangement, sunken, well marked and perpendicular to the suture.
</paragraph>
<paragraph id="8BEDEF25666EAB58B586606DFEFC3E56" blockId="11.[189,1399,1280,1764]" box="[234,298,1280,1306]" pageId="11" pageNumber="12">
<heading id="D0A55849666EAB58B586606DFEFC3E56" bold="true" box="[234,298,1280,1306]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="6">
@ -669,9 +669,9 @@ The structure of genital segment corresponds to the typical pattern of the genus
<quantity id="4CAA42C0666EAB58B6976004FB8E3EC8" box="[1019,1112,1385,1412]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="12" unit="mm" value="1.3">1.3 mm</quantity>
(
<figureCitation id="1369F3A0666EAB58B1076004FB113ECF" box="[1131,1223,1385,1411]" captionStart="Figs 21 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="11" pageNumber="12">Fig. 22</figureCitation>
); basal lamina almost 2 × shorter than middle lobe; ventral blade of tegmen short and poorly defned. Median lobe in dorsal view, similar to other species of the group, although it has the triangular apex with less protruding edge and narrower at its distal third part. Parameres robust and wide with a marked hump on the outside of distal and the apical area is beveled with a blunt tip, in this area four setae are inserted, of which three are short and thin, and the fourth is longer with a small tooth, coarse and fnely sclerotized. Inner sac showing two longitudinal chains of small sclerotized pieces, surrounded by a longer structure formed by numerous fne spines that are joined at the apex, but not tooth-shaped. Legs long with a peculiar structure of the keel protibial (
); basal lamina almost 2 × shorter than middle lobe; ventral blade of tegmen short and poorly defined. Median lobe in dorsal view, similar to other species of the group, although it has the triangular apex with less protruding edge and narrower at its distal third part. Parameres robust and wide with a marked hump on the outside of distal and the apical area is beveled with a blunt tip, in this area four setae are inserted, of which three are short and thin, and the fourth is longer with a small tooth, coarse and finely sclerotized. Inner sac showing two longitudinal chains of small sclerotized pieces, surrounded by a longer structure formed by numerous fine spines that are joined at the apex, but not tooth-shaped. Legs long with a peculiar structure of the keel protibial (
<figureCitation id="1369F3A0666EAB58B74B63EEFDAB3DD1" box="[551,637,1667,1693]" captionStart="Figs 16 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="11" pageNumber="12">Fig. 17</figureCitation>
) which is long, slightly elevated, with a straight area, and fuzzy projections; in addition, the frst three protarsomeres are dilated, although frst protarsomere is narrower than anterior area of protibia; mesotibia arched and metatibia straight.
) which is long, slightly elevated, with a straight area, and fuzzy projections; in addition, the first three protarsomeres are dilated, although first protarsomere is narrower than anterior area of protibia; mesotibia arched and metatibia straight.
</paragraph>
<paragraph id="8BEDEF25666EAB58B5866266FE933C69" blockId="11.[189,1399,1803,2005]" box="[234,325,1803,1829]" pageId="11" pageNumber="12">
<heading id="D0A55849666EAB58B5866266FE933C69" bold="true" box="[234,325,1803,1829]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="6">
@ -679,10 +679,10 @@ The structure of genital segment corresponds to the typical pattern of the genus
</heading>
</paragraph>
<paragraph id="8BEDEF25666EAB5FB5D16243FAD93A20" blockId="11.[189,1399,1803,2005]" lastBlockId="12.[189,1399,267,364]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
The most signifcant external differences are observed on the antennae, pronotum, elytra, protibia and protarsomeres. Antennae very similar to those observed in males (
The most significant external differences are observed on the antennae, pronotum, elytra, protibia and protarsomeres. Antennae very similar to those observed in males (
<tableCitation id="C6D0DA9E666EAB58B6D7623CFBC43C27" box="[955,1042,1873,1899]" captionStart="Table 1" captionStartId="4.[188,257,267,293]" captionTargetBox="[193,1384,385,1378]" captionTargetPageId="4" captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width." httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04" pageId="11" pageNumber="12" tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
); the 2nd antennomere equal to the 5th one, and the 8th almost 1.5 × longer than wide. Pronotum slightly transverse and clearly narrower than in males (
<figureCitation id="1369F3A0666EAB58B40662FAFE643CFE" box="[362,434,1943,1970]" captionStart="Figs 6 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 15. Speonemadus of the escalerai - group pronotum. 6 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
<figureCitation id="1369F3A0666EAB58B40662FAFE643CFE" box="[362,434,1943,1970]" captionStart="Figs 615" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 615. Speonemadus of the escalerai-group pronotum. 67. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 89. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 1011. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 1213. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 1415. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
), its sides almost straight and divergent on the posterior part; the maximum width is near the base and bell-like in form; posterior angles are almost straight with rounded angles; pronotal base is almost as wide as base of elytra. Elytra are elliptical and elongated and proportionally slightly shorter than in males. Apex of elytra is notched and toothed. Protibiae lacking keel and protarsi slender. 7th and 8th uroventrites and genital segment typical of the genus (see
<bibRefCitation id="EFC392D46669AB5FB6B3643FFAD43A20" author="Giachino P. M. &amp; Vailati D." box="[991,1282,338,364]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="12" pageNumber="13" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
).
@ -693,7 +693,7 @@ The most signifcant external differences are observed on the antennae, pronotum,
</heading>
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D164AFFA9C3925" blockId="12.[189,1399,450,617]" pageId="12" pageNumber="13">
The most signifcant variation was observed in the samples from the two caves of Huelva (Gruta de la Maravillas and Cueva del Guerrero), specimens from both caves show intermediate characteristics between
The most significant variation was observed in the samples from the two caves of Huelva (Gruta de la Maravillas and Cueva del Guerrero), specimens from both caves show intermediate characteristics between
<taxonomicName id="4C5294A66669AB5FB4446764FE08396E" baseAuthorityName="Kraatz" baseAuthorityYear="1870" box="[296,478,520,547]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B92633376669AB5FB4446764FE08396E" box="[296,478,520,547]" italics="true" pageId="12" pageNumber="13">S. angusticollis</emphasis>
</taxonomicName>
@ -701,7 +701,7 @@ and
<taxonomicName id="4C5294A66669AB5FB7706764FD5F396E" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[540,649,520,547]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B92633376669AB5FB7706764FD5F396E" box="[540,649,520,547]" italics="true" pageId="12" pageNumber="13">S. breuili</emphasis>
</taxonomicName>
, with the protibial keel more close to the frst one and female pronotum most similar to the second species. Systematic position of the specimens collected from these two caves will be solved only by further study of specimens from caves near the Sierra de Aracena.
, with the protibial keel more close to the first one and female pronotum most similar to the second species. Systematic position of the specimens collected from these two caves will be solved only by further study of specimens from caves near the Sierra de Aracena.
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE6669AB5FB5D167FDFC103829" pageId="12" pageNumber="13" type="biology_ecology">
@ -717,7 +717,7 @@ and now also in the Portuguese region of Alentejo. In less dry regions it is fou
<bibRefCitation id="EFC392D46669AB5FB48C6645FD8A380E" author="Blas M." box="[480,604,808,834]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="12" pageNumber="13" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">Blas 1977</bibRefCitation>
). It has been also been reported (
<bibRefCitation id="EFC392D46669AB5FB6B36645FAD4380E" author="Giachino P. M. &amp; Vailati D." box="[991,1282,808,834]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="12" pageNumber="13" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
) from the Superfcial Subterranean Habitat (
) from the Superficial Subterranean Habitat (
<emphasis id="B92633376669AB5FB7216621FD583829" box="[589,654,844,869]" italics="true" pageId="12" pageNumber="13">sensu</emphasis>
<bibRefCitation id="EFC392D46669AB5FB7F96626FC6C3829" author="Giachino &amp; Vailati" box="[661,954,843,869]" firstAuthor="Giachino" pageId="12" pageNumber="13" pagination="3" refId="ref12117" refString="Giachino P. M. &amp; Vailati D. 2010. The subterranean environment. Hypogean life, concepts and collecting techniques. WBA Handbooks 3, World Biodiversity Association, Verona." type="book chapter" year="2010">Giachino &amp; Vailati 2010</bibRefCitation>
).
@ -738,7 +738,7 @@ and Sevilla and now also from southeastern
<collectingCountry id="F345AFB56669AB5FB65E6148FC433F73" box="[818,917,1061,1087]" name="Portugal" pageId="12" pageNumber="13">Portugal</collectingCountry>
(
<figureCitation id="1369F3A06669AB5FB6C46148FBD43F73" box="[936,1026,1061,1087]" captionStart="Fig. 32" captionStartId="18.[189,232,1955,1981]" captionTargetBox="[189,1398,986,1883]" captionTargetId="figure@18.[189,1398,986,1914]" captionTargetPageId="18" captionText="Fig. 32. Distribution of Speonemadus of the escalerai - group: S. algarvensis sp. nov., S. angusticollis (Kraatz, 1870), S. bolivari (Jeannel, 1922), S. breuili (Jeannel, 1922) and S. escalerai (Uhagón, 1898)." httpUri="https://zenodo.org/record/889358/files/figure.png" pageId="12" pageNumber="13">Fig. 32</figureCitation>
). The present fnding in a cave of Alandroal, in the Alentejo province, is the frst confrmed record of this species for the Portuguese territory as previous records for
). The present finding in a cave of Alandroal, in the Alentejo province, is the first confirmed record of this species for the Portuguese territory as previous records for
<collectingCountry id="F345AFB56669AB5FB7286106FD713FC9" box="[580,679,1131,1157]" name="Portugal" pageId="12" pageNumber="13">Portugal</collectingCountry>
(
<bibRefCitation id="EFC392D46669AB5FB7D46106FC8F3FC9" author="Jeannel" box="[696,857,1131,1157]" firstAuthor="Jeannel" pageId="12" pageNumber="13" pagination="5 - 15" refId="ref12333" refString="Jeannel R. 1941. Premieres explorations des grottes du Portugal par M. A. de B. Machado, Coleopteres. Publicacoes do Instituto de Zoologia &quot; Augusto Nobre &quot; 4: 5 - 15." type="journal article" year="1941">Jeannel 1941</bibRefCitation>