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<document id="FA3C3CC733D98F84AEF5487480C0A5F3" ID-CLB-Dataset="22501" ID-DOI="10.11646/zootaxa.4670.1.1" ID-GBIF-Dataset="8e19d5d0-be5e-4022-a16c-b3ce0e554c54" ID-ISSN="1175-5334" ID-Zenodo-Dep="3773507" ID-ZooBank="urn:lsid:zoobank.org:pub:5D56C44D-BBAE-4CE7-9184-8A6DDC8009A4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588077119377" checkinUser="plazi" docAuthor="Sinclair, Bradley J., Vajda, Élodie A., Saigusa, Toyohei, Shamshev, Igor V. &amp; Wheeler, Terry A." docDate="2019" docId="03C52A7BEB19FF90FF13FA0398E0F818" docLanguage="en" docName="zootaxa.4670.1.1.pdf.imf" docOrigin="Zootaxa 4670 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Rhamphomyia Meigen 1822" docType="key" docVersion="6" lastPageNumber="7" masterDocId="FFFC5203EB1CFF96FF84FFED9D7CFFFF" masterDocTitle="Rhamphomyia Meigen of the Canadian Arctic Archipelago, Greenland and Iceland (Diptera: Empididae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="6" updateTime="1698837924851" updateUser="ExternalLinkService">
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<mods:title id="AD13CE540CE129AB15D6FDFE8D626174">Rhamphomyia Meigen of the Canadian Arctic Archipelago, Greenland and Iceland (Diptera: Empididae)</mods:title>
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<mods:namePart id="05A7610B3B6188B43385C59E0C690548">Sinclair, Bradley J.</mods:namePart>
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<mods:namePart id="214536393521CFFDCD18D9BF42207542">Vajda, Élodie A.</mods:namePart>
<mods:affiliation id="9FAAFF142D6042345C160C9C1B898050">Department of Natural Resource Sciences, McGill University, Macdonald Campus, Ste-Anne-de-Bellevue, QC, H 9 X 3 V 9, Canada</mods:affiliation>
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<mods:namePart id="862F2FCC0C108D676308606BECF6F49B">Saigusa, Toyohei</mods:namePart>
<mods:affiliation id="D1F2B0AB696A5B8EEF7F623285FD1CC8">17 - 1 - 402 Baikoen 2 - chome, Chuo-ku, Fukuoka-shi 810 - 0035, Japan</mods:affiliation>
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<mods:affiliation id="B3EC1C2FA703397C53D0BB0996818F3A">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg, 199034, Russia</mods:affiliation>
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<heading id="D09B2C01EB19FF93FF13FA0399E5F9F7" bold="true" box="[151,1177,1517,1544]" fontSize="10" level="2" pageId="5" pageNumber="6" reason="2">
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Key to male
<taxonomicName id="4C6CE0EEEB19FF93FEB0FA009C9AF9F7" authorityName="Meigen" authorityYear="1822" box="[308,486,1517,1544]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B918477FEB19FF93FEB0FA009C9AF9F7" bold="true" box="[308,486,1517,1544]" italics="true" pageId="5" pageNumber="6">Rhamphomyia</emphasis>
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of the Canadian Arctic Islands,
<collectingCountry id="F37BDBFDEB19FF93FCFDFA009E82F9F8" box="[889,1022,1517,1543]" name="Greenland" pageId="5" pageNumber="6">Greenland</collectingCountry>
and
<collectingCountry id="F37BDBFDEB19FF93FBB8FA0399E5F9F7" box="[1084,1177,1518,1544]" name="Iceland" pageId="5" pageNumber="6">Iceland</collectingCountry>
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1 Prosternum clothed entirely with white to brownish silky, hair-like setae (
<figureCitation id="135787E8EB19FF93FC1CF9C19E98F9BC" box="[920,996,1580,1603]" captionStart="FIGURE 3" captionStartId="12.[151,250,1972,1997]" captionTargetBox="[151,1435,409,1901]" captionTargetId="figure@12.[151,1435,409,1901]" captionTargetPageId="12" captionText="FIGURE 3. Rhamphomyia (Ctenempis) albopilosa Coquillett. A. Male forelegs; B. Male hindleg, anterior view; C. Male terminalia, lateral view; D. Female head and prosternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773513" httpUri="https://zenodo.org/record/3773513/files/figure.png" pageId="5" pageNumber="6">Fig. 3D</figureCitation>
); sternite 6 with cluster of strong golden setae; sternite 7 with pair of horn-like projections ventrally (
<figureCitation id="135787E8EB19FF93FD6CF9A49E4DF9A0" box="[744,817,1609,1631]" captionStart="FIGURE 3" captionStartId="12.[151,250,1972,1997]" captionTargetBox="[151,1435,409,1901]" captionTargetId="figure@12.[151,1435,409,1901]" captionTargetPageId="12" captionText="FIGURE 3. Rhamphomyia (Ctenempis) albopilosa Coquillett. A. Male forelegs; B. Male hindleg, anterior view; C. Male terminalia, lateral view; D. Female head and prosternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773513" httpUri="https://zenodo.org/record/3773513/files/figure.png" pageId="5" pageNumber="6">Fig. 3C</figureCitation>
)....................
<taxonomicName id="4C6CE0EEEB19FF93FBB4F9A798E1F9A0" authority="Coquillett" authorityName="Coquillett" box="[1072,1437,1609,1631]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="albopilosa" subGenus="Ctenempis">
<emphasis id="B918477FEB19FF93FBB4F9A798E1F9A0" bold="true" box="[1072,1437,1609,1631]" pageId="5" pageNumber="6">
<emphasis id="B918477FEB19FF93FBB4F9A7993AF9A0" bold="true" box="[1072,1094,1610,1631]" italics="true" pageId="5" pageNumber="6">R.</emphasis>
(
<emphasis id="B918477FEB19FF93FBD0F9A499C0F9A1" bold="true" box="[1108,1212,1609,1630]" italics="true" pageId="5" pageNumber="6">Ctenempis</emphasis>
)
<emphasis id="B918477FEB19FF93FB4EF9A49853F9A0" bold="true" box="[1226,1327,1609,1631]" italics="true" pageId="5" pageNumber="6">albopilosa</emphasis>
Coquillett
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</paragraph>
<paragraph id="8BD39B6DEB19FF93FF13F98A98E0F984" blockId="5.[151,1437,1580,2035]" box="[151,1436,1638,1660]" pageId="5" pageNumber="6">
<keyLead id="309D3858EB19FF93FF13F98A98E0F984" box="[151,1436,1638,1660]" pageId="5" pageNumber="6">- Prosternum bare; sternite 6 without row of golden setae; sternite 7 unmodified or modified different than above.......... 2</keyLead>
</paragraph>
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<paragraph id="8BD39B6DEB19FF93FF13F96E98E0F967" blockId="5.[151,1437,1580,2035]" box="[151,1436,1667,1689]" pageId="5" pageNumber="6">
<keyLead id="309D3858EB19FF93FF13F96E98E0F967" box="[151,1436,1667,1689]" pageId="5" pageNumber="6">
2 Cerci directed anteriorly (
<figureCitation id="135787E8EB19FF93FE5DF96E9F55F966" box="[473,553,1667,1689]" captionStart="FIGURE 6" captionStartId="15.[151,250,1956,1981]" captionTargetBox="[151,1436,174,1935]" captionTargetId="figure@15.[151,1436,174,1936]" captionTargetPageId="15" captionText="FIGURE 6. Rhamphomyia (Dasyrhamphomyia) erinacioides Malloch, holotype. A. Male terminalia, lateral view; B. Male terminalia, dorsal view; C. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773519" httpUri="https://zenodo.org/record/3773519/files/figure.png" pageId="5" pageNumber="6">Figs 6A</figureCitation>
,
<figureCitation id="135787E8EB19FF93FDB7F96E9F2EF967" box="[563,594,1667,1688]" captionStart="FIGURE 9" captionStartId="18.[151,250,1924,1949]" captionTargetBox="[151,1435,301,1895]" captionTargetId="figure@18.[151,1435,301,1905]" captionTargetPageId="18" captionText="FIGURE 9. Rhamphomyia (Dasyrhamphomyia) hovgaardii Holmgren. A. Male hindleg, anterior view; B. Female hindleg, anterior view; C. Male terminalia, lateral view; D. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773526" httpUri="https://zenodo.org/record/3773526/files/figure.png" pageId="5" pageNumber="6">9C</figureCitation>
,
<figureCitation id="135787E8EB19FF93FDDFF96E9FF9F967" box="[603,645,1667,1688]" captionStart="FIGURE 12" captionStartId="22.[151,250,1871,1896]" captionTargetBox="[151,1435,301,1850]" captionTargetId="figure@22.[151,1435,301,1851]" captionTargetPageId="22" captionText="FIGURE 12. Rhamphomyia (Dasyrhamphomyia) leptidiformis Frey. A. Male hindleg, anterior view; B. Male terminalia, lateral view; C. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773532" httpUri="https://zenodo.org/record/3773532/files/figure.png" pageId="5" pageNumber="6">12B</figureCitation>
,
<figureCitation id="135787E8EB19FF93FD0BF96E9FC4F966" box="[655,696,1667,1689]" captionStart="FIGURE 15" captionStartId="26.[151,250,1874,1899]" captionTargetBox="[151,1435,589,1854]" captionTargetId="figure@26.[151,1435,589,1854]" captionTargetPageId="26" captionText="FIGURE 15. Rhamphomyia (Dasyrhamphomyia) nigrita Zetterstedt. A. Male hindleg, anterior view; B. Male terminalia, lateral view; C. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773538" httpUri="https://zenodo.org/record/3773538/files/figure.png" pageId="5" pageNumber="6">15B</figureCitation>
) (subgenus
<taxonomicName id="4C6CE0EEEB19FF93FCA8F96E9E9EF967" box="[812,994,1667,1688]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subGenus" subGenus="Dasyrhamphomyia">
<emphasis id="B918477FEB19FF93FCA8F96E9E9EF967" box="[812,994,1667,1688]" italics="true" pageId="5" pageNumber="6">Dasyrhamphomyia</emphasis>
</taxonomicName>
)................................... 3
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB19FF93FF13F94D98E0F949" blockId="5.[151,1437,1580,2035]" box="[151,1436,1696,1718]" pageId="5" pageNumber="6">
<keyLead id="309D3858EB19FF93FF13F94D98E0F949" box="[151,1436,1696,1718]" pageId="5" pageNumber="6">- Cerci directed posteriorly............................................................................... 6</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BD39B6DEB19FF93FF13F95098E1F8F4" blockId="5.[151,1437,1580,2035]" pageId="5" pageNumber="6">
<keyLead id="309D3858EB19FF93FF13F95098E1F8F4" pageId="5" pageNumber="6">
3 Thorax short haired (setae shorter than gaps between rows) with acrostichals and dorsocentrals 12 serial, with brown vittae beneath rows; cerci with dark apex, with left cercus overlapping right at tip (
<figureCitation id="135787E8EB19FF93FC3AF934994AF910" box="[958,1078,1753,1775]" captionStart="FIGURE 12" captionStartId="22.[151,250,1871,1896]" captionTargetBox="[151,1435,301,1850]" captionTargetId="figure@22.[151,1435,301,1851]" captionTargetPageId="22" captionText="FIGURE 12. Rhamphomyia (Dasyrhamphomyia) leptidiformis Frey. A. Male hindleg, anterior view; B. Male terminalia, lateral view; C. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773532" httpUri="https://zenodo.org/record/3773532/files/figure.png" pageId="5" pageNumber="6">Figs 12B, C</figureCitation>
); hind tibia with anteroventral row of dark, stout, spine-like setae shorter than half tibial width..................................
<taxonomicName id="4C6CE0EEEB19FF93FB24F91B98E1F8F4" authority="Frey" authorityName="Frey" box="[1184,1437,1782,1804]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="leptidiformis" subGenus="Dasyrhamphomyia">
<emphasis id="B918477FEB19FF93FB24F91B98E1F8F4" bold="true" box="[1184,1437,1782,1804]" pageId="5" pageNumber="6">
<emphasis id="B918477FEB19FF93FB24F91B99CAF8F4" bold="true" box="[1184,1206,1782,1803]" italics="true" pageId="5" pageNumber="6">R.</emphasis>
(
<emphasis id="B918477FEB19FF93FB40F91B99A9F8F4" bold="true" box="[1220,1237,1782,1803]" italics="true" pageId="5" pageNumber="6">D</emphasis>
.)
<emphasis id="B918477FEB19FF93FB6DF91B981AF8F3" bold="true" box="[1257,1382,1782,1804]" italics="true" pageId="5" pageNumber="6">leptidiformis</emphasis>
Frey
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</paragraph>
<paragraph id="8BD39B6DEB19FF93FF13F8FE98E0F89D" blockId="5.[151,1437,1580,2035]" pageId="5" pageNumber="6">
<keyLead id="309D3858EB19FF93FF13F8FE98E0F89D" pageId="5" pageNumber="6">
- Thorax long haired (setae longer than gaps between rows) with acrostichals and dorsocentrals 24 serial, without brown vittae beneath rows; cerci not darkened at apex, with right cercus overlapping left at tip (
<figureCitation id="135787E8EB19FF93FC75F8DD993DF8BA" box="[1009,1089,1840,1862]" captionStart="FIGURE 9" captionStartId="18.[151,250,1924,1949]" captionTargetBox="[151,1435,301,1895]" captionTargetId="figure@18.[151,1435,301,1905]" captionTargetPageId="18" captionText="FIGURE 9. Rhamphomyia (Dasyrhamphomyia) hovgaardii Holmgren. A. Male hindleg, anterior view; B. Female hindleg, anterior view; C. Male terminalia, lateral view; D. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773526" httpUri="https://zenodo.org/record/3773526/files/figure.png" pageId="5" pageNumber="6">Figs 9D</figureCitation>
,
<figureCitation id="135787E8EB19FF93FBCEF8DD9909F8B9" box="[1098,1141,1840,1862]" captionStart="FIGURE 15" captionStartId="26.[151,250,1874,1899]" captionTargetBox="[151,1435,589,1854]" captionTargetId="figure@26.[151,1435,589,1854]" captionTargetPageId="26" captionText="FIGURE 15. Rhamphomyia (Dasyrhamphomyia) nigrita Zetterstedt. A. Male hindleg, anterior view; B. Male terminalia, lateral view; C. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773538" httpUri="https://zenodo.org/record/3773538/files/figure.png" pageId="5" pageNumber="6">15C</figureCitation>
); hind tibia without anteroventral row of dark, stout, spine-like setae..................................................................... 4
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</keyStep>
<keyStep id="309883C8EB19FF93FF13F88498E0F846" pageId="5" pageNumber="6">
<paragraph id="8BD39B6DEB19FF93FF13F88498E0F864" blockId="5.[151,1437,1580,2035]" pageId="5" pageNumber="6">
<keyLead id="309D3858EB19FF93FF13F88498E0F864" pageId="5" pageNumber="6">
4 Cerci pale, elongate, extending to tergite 3, overlapping subapically in addition to apical overlap (
<figureCitation id="135787E8EB19FF93FB0FF884987DF880" box="[1163,1281,1897,1919]" captionStart="FIGURE 15" captionStartId="26.[151,250,1874,1899]" captionTargetBox="[151,1435,589,1854]" captionTargetId="figure@26.[151,1435,589,1854]" captionTargetPageId="26" captionText="FIGURE 15. Rhamphomyia (Dasyrhamphomyia) nigrita Zetterstedt. A. Male hindleg, anterior view; B. Male terminalia, lateral view; C. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773538" httpUri="https://zenodo.org/record/3773538/files/figure.png" pageId="5" pageNumber="6">Figs 15B, C</figureCitation>
)...........................................................................................
<taxonomicName id="4C6CE0EEEB19FF93FB1DF86B98E0F864" authority="Zetterstedt" authorityName="Zetterstedt" box="[1177,1436,1926,1948]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nigrita" subGenus="Dasyrhamphomyia">
<emphasis id="B918477FEB19FF93FB1DF86B98E0F864" bold="true" box="[1177,1436,1926,1948]" pageId="5" pageNumber="6">
<emphasis id="B918477FEB19FF93FB1DF86B99D3F864" bold="true" box="[1177,1199,1926,1947]" italics="true" pageId="5" pageNumber="6">R.</emphasis>
(
<emphasis id="B918477FEB19FF93FB39F86B99A8F864" bold="true" box="[1213,1236,1926,1947]" italics="true" pageId="5" pageNumber="6">D.</emphasis>
)
<emphasis id="B918477FEB19FF93FB66F86B9859F864" bold="true" box="[1250,1317,1926,1947]" italics="true" pageId="5" pageNumber="6">nigrita</emphasis>
Zetterstedt
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB19FF93FF13F84E98E0F846" blockId="5.[151,1437,1580,2035]" box="[151,1436,1955,1977]" pageId="5" pageNumber="6">
<keyLead id="309D3858EB19FF93FF13F84E98E0F846" box="[151,1436,1955,1977]" pageId="5" pageNumber="6">
- Cerci brownish to black, shorter, extending to tergite 5, only overlapping apically (
<figureCitation id="135787E8EB19FF93FC78F84E9937F846" box="[1020,1099,1955,1977]" captionStart="FIGURE 6" captionStartId="15.[151,250,1956,1981]" captionTargetBox="[151,1436,174,1935]" captionTargetId="figure@15.[151,1436,174,1936]" captionTargetPageId="15" captionText="FIGURE 6. Rhamphomyia (Dasyrhamphomyia) erinacioides Malloch, holotype. A. Male terminalia, lateral view; B. Male terminalia, dorsal view; C. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773519" httpUri="https://zenodo.org/record/3773519/files/figure.png" pageId="5" pageNumber="6">Figs 6A</figureCitation>
,
<figureCitation id="135787E8EB19FF93FBD2F84E990FF846" box="[1110,1139,1955,1977]" captionStart="FIGURE 9" captionStartId="18.[151,250,1924,1949]" captionTargetBox="[151,1435,301,1895]" captionTargetId="figure@18.[151,1435,301,1905]" captionTargetPageId="18" captionText="FIGURE 9. Rhamphomyia (Dasyrhamphomyia) hovgaardii Holmgren. A. Male hindleg, anterior view; B. Female hindleg, anterior view; C. Male terminalia, lateral view; D. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773526" httpUri="https://zenodo.org/record/3773526/files/figure.png" pageId="5" pageNumber="6">9C</figureCitation>
)....................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB19FF90FF13F82D98E1FF33" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<paragraph id="8BD39B6DEB19FF93FF13F82D98E0F80D" blockId="5.[151,1437,1580,2035]" pageId="5" pageNumber="6">
<keyLead id="309D3858EB19FF93FF13F82D98E0F80D" pageId="5" pageNumber="6">
5 Dichoptic; abdomen with setae longer than length of abdominal tergites; cerci blackish (
<figureCitation id="135787E8EB19FF93FBC5F82D99ECF829" box="[1089,1168,1984,2006]" captionStart="FIGURE 6" captionStartId="15.[151,250,1956,1981]" captionTargetBox="[151,1436,174,1935]" captionTargetId="figure@15.[151,1436,174,1936]" captionTargetPageId="15" captionText="FIGURE 6. Rhamphomyia (Dasyrhamphomyia) erinacioides Malloch, holotype. A. Male terminalia, lateral view; B. Male terminalia, dorsal view; C. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773519" httpUri="https://zenodo.org/record/3773519/files/figure.png" pageId="5" pageNumber="6">Fig. 6A</figureCitation>
); hind tibia and femur with pronounced fine, pale ventral pubescence (
<figureCitation id="135787E8EB19FF93FDF9F8309FBBF80C" box="[637,711,2013,2035]" captionStart="FIGURE 6" captionStartId="15.[151,250,1956,1981]" captionTargetBox="[151,1436,174,1935]" captionTargetId="figure@15.[151,1436,174,1936]" captionTargetPageId="15" captionText="FIGURE 6. Rhamphomyia (Dasyrhamphomyia) erinacioides Malloch, holotype. A. Male terminalia, lateral view; B. Male terminalia, dorsal view; C. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773519" httpUri="https://zenodo.org/record/3773519/files/figure.png" pageId="5" pageNumber="6">Fig. 6C</figureCitation>
)....................................
<taxonomicName id="4C6CE0EEEB19FF93FB05F83098E0F80D" authority="Malloch" authorityName="Malloch" box="[1153,1436,2013,2035]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="erinacioides" subGenus="Dasyrhamphomyia">
<emphasis id="B918477FEB19FF93FB05F83098E0F80D" bold="true" box="[1153,1436,2013,2035]" pageId="5" pageNumber="6">
<emphasis id="B918477FEB19FF93FB05F83099EBF80D" bold="true" box="[1153,1175,2013,2034]" italics="true" pageId="5" pageNumber="6">R.</emphasis>
(
<emphasis id="B918477FEB19FF93FB21F83099C0F80D" bold="true" box="[1189,1212,2013,2034]" italics="true" pageId="5" pageNumber="6">D.</emphasis>
)
<emphasis id="B918477FEB19FF93FB4EF830983EF80C" bold="true" box="[1226,1346,2013,2035]" italics="true" pageId="5" pageNumber="6">erinacioides</emphasis>
Malloch
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</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FF7798E1FF33" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FF7798E1FF33" pageId="6" pageNumber="7">
- Holoptic; abdomen with setae shorter than length of abdominal tergites; cerci pale brown (
<figureCitation id="135787E8EB1AFF90FBCDFF7499E9FF4F" box="[1097,1173,153,176]" captionStart="FIGURE 9" captionStartId="18.[151,250,1924,1949]" captionTargetBox="[151,1435,301,1895]" captionTargetId="figure@18.[151,1435,301,1905]" captionTargetPageId="18" captionText="FIGURE 9. Rhamphomyia (Dasyrhamphomyia) hovgaardii Holmgren. A. Male hindleg, anterior view; B. Female hindleg, anterior view; C. Male terminalia, lateral view; D. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773526" httpUri="https://zenodo.org/record/3773526/files/figure.png" pageId="6" pageNumber="7">Fig. 9C</figureCitation>
); hind tibia and femur with long setae beneath (
<figureCitation id="135787E8EB1AFF90FE24FF5A9C90FF32" box="[416,492,183,205]" captionStart="FIGURE 9" captionStartId="18.[151,250,1924,1949]" captionTargetBox="[151,1435,301,1895]" captionTargetId="figure@18.[151,1435,301,1905]" captionTargetPageId="18" captionText="FIGURE 9. Rhamphomyia (Dasyrhamphomyia) hovgaardii Holmgren. A. Male hindleg, anterior view; B. Female hindleg, anterior view; C. Male terminalia, lateral view; D. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773526" httpUri="https://zenodo.org/record/3773526/files/figure.png" pageId="6" pageNumber="7">Fig. 9A</figureCitation>
)......................................................
<emphasis id="B918477FEB1AFF90FBFFFF5A98E1FF33" bold="true" box="[1147,1437,182,205]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FBFFFF5A99EDFF33" bold="true" box="[1147,1169,183,204]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
<taxonomicName id="4C6CE0EEEB1AFF90FB13FF5A98E1FF33" authority="Holmgren" authorityName="Holmgren" box="[1175,1437,182,205]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hovgaardii" subGenus="Dasyrhamphomyia">
(
<emphasis id="B918477FEB1AFF90FB1BFF5A99CCFF33" bold="true" box="[1183,1200,183,204]" italics="true" pageId="6" pageNumber="7">D</emphasis>
.)
<emphasis id="B918477FEB1AFF90FB40FF5B9853FF33" bold="true" box="[1220,1327,182,204]" italics="true" pageId="6" pageNumber="7">hovgaardii</emphasis>
Holmgren
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<keyStep id="309883C8EB1AFF90FF13FF3998E0FEBD" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FF3998E0FEF8" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FF3998E0FEF8" pageId="6" pageNumber="7">
6 Wing without dm-m crossvein (cell dm open) (
<figureCitation id="135787E8EB1AFF90FD21FF399F81FF15" box="[677,765,212,234]" captionStart="FIGURE 62" captionStartId="88.[151,250,1773,1798]" captionTargetBox="[151,1435,1165,1750]" captionTargetId="figure@88.[151,1435,1165,1753]" captionTargetPageId="88" captionText="FIGURE 62. Wings of arctic Rhamphomyia. A. R. albopilosa, female, dorsal view; B. R. diversipennis, female, ventral view; C. R. diversipennis, male, ventral view; D. R. hilariformis, female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3773637" httpUri="https://zenodo.org/record/3773637/files/figure.png" pageId="6" pageNumber="7">Fig. 62C</figureCitation>
); scutum with shiny vittae.................................................................................................
<taxonomicName id="4C6CE0EEEB1AFF90FC5BFF1F98E0FEF8" authority="Becker" authorityName="Becker" box="[991,1436,241,263]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="diversipennis" subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FC5BFF1F98E0FEF8" bold="true" box="[991,1436,241,263]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FC5BFF1F9E89FEF8" bold="true" box="[991,1013,242,263]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FB87FF1C99C2FEF8" bold="true" box="[1027,1214,241,263]" italics="true" pageId="6" pageNumber="7">Pararhamphomyia</emphasis>
)
<emphasis id="B918477FEB1AFF90FB48FF1C9833FEF8" bold="true" box="[1228,1359,241,263]" italics="true" pageId="6" pageNumber="7">diversipennis</emphasis>
Becker
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<paragraph id="8BD39B6DEB1AFF90FF13FEE298E0FEBD" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FEE298E0FEBD" pageId="6" pageNumber="7">
- Wing with dm-m crossvein (cell dm closed), sometimes dm-m crossvein or M
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base absent in
<emphasis id="B918477FEB1AFF90FB0CFEE299E2FEDB" box="[1160,1182,271,292]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
<taxonomicName id="4C6CE0EEEB1AFF90FB21FEE298E5FEDA" authority="Becker" authorityName="Becker" box="[1189,1433,271,293]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="omissinervis" subGenus="Pararhamphomyia">
(
<emphasis id="B918477FEB1AFF90FB29FEE299C0FEDB" box="[1197,1212,271,292]" italics="true" pageId="6" pageNumber="7">P</emphasis>
.)
<emphasis id="B918477FEB1AFF90FB56FEE29836FEDB" box="[1234,1354,271,292]" italics="true" pageId="6" pageNumber="7">omissinervis</emphasis>
Becker
</taxonomicName>
; scutum without shiny vittae............................................................................. 7
</keyLead>
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</keyStep>
<keyStep id="309883C8EB1AFF90FF13FEA798E0FE83" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FEA798E0FEA0" blockId="6.[151,1437,153,2023]" box="[151,1436,329,351]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FEA798E0FEA0" box="[151,1436,329,351]" pageId="6" pageNumber="7">
7 Cercus with distinctive finger-like projection dorsally near base (
<figureCitation id="135787E8EB1AFF90FCD5FEA49ED0FEA0" box="[849,940,329,351]" captionStart="FIGURE 53" captionStartId="78.[151,250,1571,1596]" captionTargetBox="[151,1415,661,1543]" captionTargetId="figure@78.[151,1435,661,1551]" captionTargetPageId="78" captionText="FIGURE 53. Rhamphomyia hirtula Zetterstedt. A. Male hindleg, anterior view; B. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773619" httpUri="https://zenodo.org/record/3773619/files/figure.png" pageId="6" pageNumber="7">Figs 53B</figureCitation>
,
<figureCitation id="135787E8EB1AFF90FC32FEA49E9CFEA0" box="[950,992,329,351]" captionStart="FIGURE 60" captionStartId="86.[151,250,1785,1810]" captionTargetBox="[151,1411,949,1765]" captionTargetId="figure@86.[151,1435,949,1765]" captionTargetPageId="86" captionText="FIGURE 60. Rhamphomyia setosa Coquillett. A. Male termimalia, lateral view; B. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773633" httpUri="https://zenodo.org/record/3773633/files/figure.png" pageId="6" pageNumber="7">60A</figureCitation>
)................................... 8
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FE8A98E0FE83" blockId="6.[151,1437,153,2023]" box="[151,1436,359,381]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FE8A98E0FE83" box="[151,1436,359,381]" pageId="6" pageNumber="7">- Upper margins of cercus without finger-like projections....................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FE6998E1FDD2" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FE6998E0FE2B" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FE6998E0FE2B" pageId="6" pageNumber="7">
8 Large species (wing length more than
<quantity id="4C943688EB1AFF90FDD9FE699FE6FE65" box="[605,666,388,410]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="mm" value="5.0">5 mm</quantity>
); phallus curvature occurring before folding into epandrium; cercal finger-like projection stout (
<figureCitation id="135787E8EB1AFF90FE01FE4C9CA1FE48" box="[389,477,417,439]" captionStart="FIGURE 53" captionStartId="78.[151,250,1571,1596]" captionTargetBox="[151,1415,661,1543]" captionTargetId="figure@78.[151,1435,661,1551]" captionTargetPageId="78" captionText="FIGURE 53. Rhamphomyia hirtula Zetterstedt. A. Male hindleg, anterior view; B. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773619" httpUri="https://zenodo.org/record/3773619/files/figure.png" pageId="6" pageNumber="7">Fig. 53B</figureCitation>
); mid tibia with at least 3 preapical anteroventral and posteroventral setae dark, stout, spine-like.....................................................................................
<taxonomicName id="4C6CE0EEEB1AFF90FB41FE5298E0FE2B" authority="Zetterstedt" authorityName="Zetterstedt" box="[1221,1436,446,468]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hirtula">
<emphasis id="B918477FEB1AFF90FB41FE5298E0FE2B" bold="true" box="[1221,1436,446,468]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB41FE529859FE2B" bold="true" box="[1221,1317,446,468]" italics="true" pageId="6" pageNumber="7">R. hirtula</emphasis>
Zetterstedt
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</taxonomicName>
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</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FE3098E1FDD2" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FE3098E1FDD2" pageId="6" pageNumber="7">
- Small species (wing length less than
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); phallus curvature hidden within epandrium; cercal finger-like projection slim, short, less than half cercal length (
<figureCitation id="135787E8EB1AFF90FDB9FE149FE6FDF0" box="[573,666,505,527]" captionStart="FIGURE 60" captionStartId="86.[151,250,1785,1810]" captionTargetBox="[151,1411,949,1765]" captionTargetId="figure@86.[151,1435,949,1765]" captionTargetPageId="86" captionText="FIGURE 60. Rhamphomyia setosa Coquillett. A. Male termimalia, lateral view; B. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773633" httpUri="https://zenodo.org/record/3773633/files/figure.png" pageId="6" pageNumber="7">Fig. 60A</figureCitation>
); mid tibia with 2 anterodorsals and 2 posterodorsals longer than twice tibial width................................................................................
<taxonomicName id="4C6CE0EEEB1AFF90FB5CFDFA98E1FDD2" authority="Coquillett" authorityName="Coquillett. The" authorityYear="1895" box="[1240,1437,535,557]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B918477FEB1AFF90FB5CFDFA98E1FDD2" bold="true" box="[1240,1437,535,557]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB5CFDFA984CFDD3" bold="true" box="[1240,1328,535,556]" italics="true" pageId="6" pageNumber="7">R. setosa</emphasis>
Coquillett
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</keyStep>
<keyStep id="309883C8EB1AFF90FF13FDD998E0FD98" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FDD998E0FDB6" blockId="6.[151,1437,153,2023]" box="[151,1436,564,586]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FDD998E0FDB6" box="[151,1436,564,586]" pageId="6" pageNumber="7">
9 Tergite 8 long, subequal to length of epandrium (
<collectingCountry id="F37BDBFDEB1AFF90FD33FDD99F83FDB5" box="[695,767,564,586]" name="Iceland" pageId="6" pageNumber="7">Iceland</collectingCountry>
only) (
<figureCitation id="135787E8EB1AFF90FCC1FDD99EE6FDB5" box="[837,922,564,586]" captionStart="FIGURE 46" captionStartId="69.[151,250,1598,1623]" captionTargetBox="[179,1361,174,1568]" captionTargetId="figure@69.[179,1408,174,1579]" captionTargetPageId="69" captionText="FIGURE 46. Rhamphomyia (Pararhamphomyia) simplex Zetterstedt. A. Male hindleg, anterior view; B. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773605" httpUri="https://zenodo.org/record/3773605/files/figure.png" pageId="6" pageNumber="7">Fig. 46B</figureCitation>
)....................
<taxonomicName id="4C6CE0EEEB1AFF90FB11FDD998E0FDB6" authority="Zetterstedt" authorityName="Zetterstedt" box="[1173,1436,564,586]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="simplex" subGenus="Pararhamphomyia">
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<emphasis id="B918477FEB1AFF90FB11FDD999D7FDB6" bold="true" box="[1173,1195,564,585]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FB3DFDD999B0FDB6" bold="true" box="[1209,1228,564,585]" italics="true" pageId="6" pageNumber="7">P.</emphasis>
)
<emphasis id="B918477FEB1AFF90FB5DFDD99859FDB5" bold="true" box="[1241,1317,564,586]" italics="true" pageId="6" pageNumber="7">simplex</emphasis>
Zetterstedt
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</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FDBF98E0FD98" blockId="6.[151,1437,153,2023]" box="[151,1436,593,615]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FDBF98E0FD98" box="[151,1436,593,615]" pageId="6" pageNumber="7">- Tergite 8 shorter than length of epandrium................................................................ 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FD8298E0FD40" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FD8298E0FD5E" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FD8298E0FD5E" pageId="6" pageNumber="7">
10 Subepandrial sclerite prolonged into pair of medioposterior lobes beneath cerci (
<figureCitation id="135787E8EB1AFF90FC62FD82993CFD7B" box="[998,1088,623,645]" captionStart="FIGURE 31" captionStartId="48.[151,250,1643,1668]" captionTargetBox="[151,1435,174,1608]" captionTargetId="figure@48.[151,1435,174,1624]" captionTargetPageId="48" captionText="FIGURE 31. Rhamphomyia (Pararhamphomyia) hilariformis Frey. A. Male foreleg; B. Male hindlegs; C. Male terminalia, dorsal view; D, E. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773572" httpUri="https://zenodo.org/record/3773572/files/figure.png" pageId="6" pageNumber="7">Figs 31C</figureCitation>
,
<figureCitation id="135787E8EB1AFF90FBCEFD82990AFD7B" box="[1098,1142,623,644]" captionStart="FIGURE 28" captionStartId="44.[151,250,1981,2006]" captionTargetBox="[158,1430,625,1962]" captionTargetId="figure@44.[158,1430,625,1962]" captionTargetPageId="44" captionText="FIGURE 28. Rhamphomyia (Pararhamphomyia) helleni Frey. A. Male terminalia, lateral view; B. Male terminalia, phallus, lateral view; C. Male hindleg, anterior view; D. Female hindlegs, posterior views." figureDoi="http://doi.org/10.5281/zenodo.3773566" httpUri="https://zenodo.org/record/3773566/files/figure.png" pageId="6" pageNumber="7">28A</figureCitation>
,
<figureCitation id="135787E8EB1AFF90FB04FD8299D6FD7A" box="[1152,1194,623,645]" captionStart="FIGURE 25" captionStartId="40.[151,250,1853,1878]" captionTargetBox="[151,1435,166,1819]" captionTargetId="figure@40.[151,1435,166,1834]" captionTargetPageId="40" captionText="FIGURE 25. Rhamphomyia (Pararhamphomyia) frigida sp. nov. A. Male terminalia, lateral view; B. Male hindleg, anterior view; C. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773560" httpUri="https://zenodo.org/record/3773560/files/figure.png" pageId="6" pageNumber="7">25C</figureCitation>
,
<figureCitation id="135787E8EB1AFF90FB30FD82999CFD7B" box="[1204,1248,623,644]" captionStart="FIGURE 34" captionStartId="52.[151,250,1143,1168]" captionTargetBox="[151,1419,229,1124]" captionTargetId="figure@52.[151,1435,229,1124]" captionTargetPageId="52" captionText="FIGURE 34. Rhamphomyia (Pararhamphomyia) hoeli Frey. A. Male terminalia, lateral view; B. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773578" httpUri="https://zenodo.org/record/3773578/files/figure.png" pageId="6" pageNumber="7">34A</figureCitation>
,
<figureCitation id="135787E8EB1AFF90FB6DFD829868FD7A" box="[1257,1300,623,645]" captionStart="FIGURE 36" captionStartId="54.[151,250,1971,1996]" captionTargetBox="[151,1430,769,1951]" captionTargetId="figure@54.[151,1435,769,1951]" captionTargetPageId="54" captionText="FIGURE 36. Rhamphomyia (Pararhamphomyia) kjellmanii Holmgren. A. uralensis Becker lectotype, lateral view; B. uralensis Becker type labels; C. Male terminalia, lateral view; D. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773582" httpUri="https://zenodo.org/record/3773582/files/figure.png" pageId="6" pageNumber="7">36C</figureCitation>
,
<figureCitation id="135787E8EB1AFF90FA99FD82983BFD7B" box="[1309,1351,623,644]" captionStart="FIGURE 38" captionStartId="57.[151,250,1788,1813]" captionTargetBox="[151,1435,553,1755]" captionTargetId="figure@57.[151,1435,553,1768]" captionTargetPageId="57" captionText="FIGURE 38. Rhamphomyia (Pararhamphomyia) lymaniana sp. nov. A. Male hindleg, anterior view; B. Male abdominal tergite 5, pointing to stout lateral view; C, D. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773586" httpUri="https://zenodo.org/record/3773586/files/figure.png" pageId="6" pageNumber="7">38C</figureCitation>
,
<figureCitation id="135787E8EB1AFF90FAD5FD829800FD7A" box="[1361,1404,623,645]" captionStart="FIGURE 40" captionStartId="60.[151,250,1618,1643]" captionTargetBox="[151,1435,336,1598]" captionTargetId="figure@60.[151,1435,336,1599]" captionTargetPageId="60" captionText="FIGURE 40. Rhamphomyia (Pararhamphomyia) omissinervis Becker. A. Male terminalia, lateral view; B. Male hindleg, anterior view C. Lectotype, lateral view; B. Type labels." figureDoi="http://doi.org/10.5281/zenodo.3773591" httpUri="https://zenodo.org/record/3773591/files/figure.png" pageId="6" pageNumber="7">40A</figureCitation>
).. .................................................................................................. 11
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FD4798E0FD40" blockId="6.[151,1437,153,2023]" box="[151,1436,681,703]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FD4798E0FD40" box="[151,1436,681,703]" pageId="6" pageNumber="7">- Cercus without pair of subepandrial lobes beneath.......................................................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FD2A98E0FD06" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FD2A98E0FD23" blockId="6.[151,1437,153,2023]" box="[151,1436,711,733]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FD2A98E0FD23" box="[151,1436,711,733]" pageId="6" pageNumber="7">11 Apex of subepandrial lobe forked....................................................................... 12</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FD0898E0FD06" blockId="6.[151,1437,153,2023]" box="[151,1436,740,762]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FD0898E0FD06" box="[151,1436,740,762]" pageId="6" pageNumber="7">- Apex of subepandrial lobe unforked..................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FCEF98E0FC90" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FCEF98E1FCCB" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FCEF98E1FCCB" pageId="6" pageNumber="7">
12 Phallus stout, looping dorsally above cercus and epandrium; apex of phallus expanded with ridge of teeth-like projections (
<figureCitation id="135787E8EB1AFF90FAF6FCEC9C54FCCB" captionStart="FIGURE 28" captionStartId="44.[151,250,1981,2006]" captionTargetBox="[158,1430,625,1962]" captionTargetId="figure@44.[158,1430,625,1962]" captionTargetPageId="44" captionText="FIGURE 28. Rhamphomyia (Pararhamphomyia) helleni Frey. A. Male terminalia, lateral view; B. Male terminalia, phallus, lateral view; C. Male hindleg, anterior view; D. Female hindlegs, posterior views." figureDoi="http://doi.org/10.5281/zenodo.3773566" httpUri="https://zenodo.org/record/3773566/files/figure.png" pageId="6" pageNumber="7">Figs 28A, B</figureCitation>
)..............................................................................
<taxonomicName id="4C6CE0EEEB1AFF90FB5BFCF298E1FCCB" authority="Frey" authorityName="Frey" box="[1247,1437,798,821]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni" subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FB5BFCF298E1FCCB" bold="true" box="[1247,1437,798,821]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB5BFCF29993FCCB" bold="true" box="[1247,1263,799,820]" italics="true" pageId="6" pageNumber="7">R</emphasis>
. (
<emphasis id="B918477FEB1AFF90FA87FCF29869FCCB" bold="true" box="[1283,1301,799,820]" italics="true" pageId="6" pageNumber="7">P.</emphasis>
)
<emphasis id="B918477FEB1AFF90FAA7FCF3981BFCCB" bold="true" box="[1315,1383,798,820]" italics="true" pageId="6" pageNumber="7">helleni</emphasis>
Frey
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FCD098E0FC90" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FCD098E0FC90" pageId="6" pageNumber="7">- Phallus slender, filamentous apically, extending slightly beyond epandrium; tip of phallus slender, tapered, lacking teeth-like projections......................................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FC9A98E0FBE0" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FC9A98E1FC38" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FC9A98E1FC38" pageId="6" pageNumber="7">
13 Fore tarsomere 1 swollen, slightly broader than width of fore tibia at apex (
<figureCitation id="135787E8EB1AFF90FC4FFC9A995AFC72" box="[971,1062,887,909]" captionStart="FIGURE 31" captionStartId="48.[151,250,1643,1668]" captionTargetBox="[151,1435,174,1608]" captionTargetId="figure@48.[151,1435,174,1624]" captionTargetPageId="48" captionText="FIGURE 31. Rhamphomyia (Pararhamphomyia) hilariformis Frey. A. Male foreleg; B. Male hindlegs; C. Male terminalia, dorsal view; D, E. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773572" httpUri="https://zenodo.org/record/3773572/files/figure.png" pageId="6" pageNumber="7">Fig. 31A</figureCitation>
); tergite 5 without stout posterolateral setae; subepandrial lobe with outer finger-like process lacking apical seta, longer than inner lobe (
<figureCitation id="135787E8EB1AFF90FB08FC7999AFFC55" box="[1164,1235,916,938]" captionStart="FIGURE 31" captionStartId="48.[151,250,1643,1668]" captionTargetBox="[151,1435,174,1608]" captionTargetId="figure@48.[151,1435,174,1624]" captionTargetPageId="48" captionText="FIGURE 31. Rhamphomyia (Pararhamphomyia) hilariformis Frey. A. Male foreleg; B. Male hindlegs; C. Male terminalia, dorsal view; D, E. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773572" httpUri="https://zenodo.org/record/3773572/files/figure.png" pageId="6" pageNumber="7">Figs 31</figureCitation>
C-E)............................................................................................
<taxonomicName id="4C6CE0EEEB1AFF90FB28FC5F98E1FC38" authority="Frey" authorityName="Frey" box="[1196,1437,945,967]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hilariformis" subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FB28FC5F98E1FC38" bold="true" box="[1196,1437,945,967]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB28FC5F99BEFC38" bold="true" box="[1196,1218,946,967]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FB54FC5F999FFC38" bold="true" box="[1232,1251,946,967]" italics="true" pageId="6" pageNumber="7">P.</emphasis>
)
<emphasis id="B918477FEB1AFF90FB74FC5C981BFC38" bold="true" box="[1264,1383,945,967]" italics="true" pageId="6" pageNumber="7">hilariformis</emphasis>
Frey
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FC2298E0FBE0" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FC2298E0FBE0" pageId="6" pageNumber="7">
- Fore tarsomere 1 slender, narrower than width of fore tibia at apex; tergite 5 with several closely set stout posterolateral setae (
<figureCitation id="135787E8EB1AFF90FF6DFC019C3DFBFD" box="[233,321,1004,1026]" captionStart="FIGURE 38" captionStartId="57.[151,250,1788,1813]" captionTargetBox="[151,1435,553,1755]" captionTargetId="figure@57.[151,1435,553,1768]" captionTargetPageId="57" captionText="FIGURE 38. Rhamphomyia (Pararhamphomyia) lymaniana sp. nov. A. Male hindleg, anterior view; B. Male abdominal tergite 5, pointing to stout lateral view; C, D. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773586" httpUri="https://zenodo.org/record/3773586/files/figure.png" pageId="6" pageNumber="7">Fig. 38B</figureCitation>
); subepandrial lobe with slender, outer finger-like process bearing 1 very long, fine, black seta (
<figureCitation id="135787E8EB1AFF90FB5CFC019833FBFD" box="[1240,1359,1004,1026]" captionStart="FIGURE 38" captionStartId="57.[151,250,1788,1813]" captionTargetBox="[151,1435,553,1755]" captionTargetId="figure@57.[151,1435,553,1768]" captionTargetPageId="57" captionText="FIGURE 38. Rhamphomyia (Pararhamphomyia) lymaniana sp. nov. A. Male hindleg, anterior view; B. Male abdominal tergite 5, pointing to stout lateral view; C, D. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773586" httpUri="https://zenodo.org/record/3773586/files/figure.png" pageId="6" pageNumber="7">Figs 38C, D</figureCitation>
)....................................................
<emphasis id="B918477FEB1AFF90FC9AFBE798E0FBE0" bold="true" box="[798,1436,1033,1055]" pageId="6" pageNumber="7">
<taxonomicName id="4C6CE0EEEB1AFF90FC9AFBE79835FBE0" authority="Sinclair, Vajda, Saigusa &amp; Shamshev" authorityName="Sinclair, Vajda, Saigusa &amp; Shamshev" authorityYear="2019" box="[798,1353,1033,1055]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="lymaniana" status="sp. nov." subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FC9AFBE79E48FBE0" bold="true" box="[798,820,1034,1055]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FCC6FBE79E28FBE0" bold="true" box="[834,852,1034,1055]" italics="true" pageId="6" pageNumber="7">P.</emphasis>
)
<emphasis id="B918477FEB1AFF90FCE6FBE49EB7FBE0" bold="true" box="[866,971,1033,1055]" italics="true" pageId="6" pageNumber="7">lymaniana</emphasis>
Sinclair, Vajda, Saigusa &amp; Shamshev
</taxonomicName>
<taxonomicNameLabel id="A22BFA04EB1AFF90FACBFBE798E0FBE0" box="[1359,1436,1034,1055]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FBCA98E0FBA5" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FBCA98E0FBC2" blockId="6.[151,1437,153,2023]" box="[151,1436,1063,1085]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FBCA98E0FBC2" box="[151,1436,1063,1085]" pageId="6" pageNumber="7">14 Phallus not forming loops.............................................................................. 15</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FBA898E0FBA5" blockId="6.[151,1437,153,2023]" box="[151,1436,1092,1114]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FBA898E0FBA5" box="[151,1436,1092,1114]" pageId="6" pageNumber="7">- Phallus forming loops................................................................................. 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FB8C98E1FB30" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FB8C98E0FB6B" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FB8C98E0FB6B" pageId="6" pageNumber="7">
15 Hind tibia with some anterodorsal, posterodorsal and dorsal setae more than 2X tibial width (
<figureCitation id="135787E8EB1AFF90FBECFB8C99BDFB88" box="[1128,1217,1121,1143]" captionStart="FIGURE 25" captionStartId="40.[151,250,1853,1878]" captionTargetBox="[151,1435,166,1819]" captionTargetId="figure@40.[151,1435,166,1834]" captionTargetPageId="40" captionText="FIGURE 25. Rhamphomyia (Pararhamphomyia) frigida sp. nov. A. Male terminalia, lateral view; B. Male hindleg, anterior view; C. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773560" httpUri="https://zenodo.org/record/3773560/files/figure.png" pageId="6" pageNumber="7">Fig. 25B</figureCitation>
); apex of subepandrial lobe rounded (
<figureCitation id="135787E8EB1AFF90FEE9FB929C9FFB6A" box="[365,483,1151,1173]" captionStart="FIGURE 25" captionStartId="40.[151,250,1853,1878]" captionTargetBox="[151,1435,166,1819]" captionTargetId="figure@40.[151,1435,166,1834]" captionTargetPageId="40" captionText="FIGURE 25. Rhamphomyia (Pararhamphomyia) frigida sp. nov. A. Male terminalia, lateral view; B. Male hindleg, anterior view; C. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773560" httpUri="https://zenodo.org/record/3773560/files/figure.png" pageId="6" pageNumber="7">Figs 25A, C</figureCitation>
)............................
<emphasis id="B918477FEB1AFF90FCC1FB9298E0FB6B" bold="true" box="[837,1436,1150,1173]" pageId="6" pageNumber="7">
<taxonomicName id="4C6CE0EEEB1AFF90FCC1FB929835FB6A" authority="Sinclair, Vajda, Saigusa &amp; Shamshev" authorityName="Sinclair, Vajda, Saigusa &amp; Shamshev" authorityYear="2019" box="[837,1353,1150,1173]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="frigida" status="sp. nov." subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FCC1FB929E27FB6B" bold="true" box="[837,859,1151,1172]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FCEDFB929E07FB6B" bold="true" box="[873,891,1151,1172]" italics="true" pageId="6" pageNumber="7">P.</emphasis>
)
<emphasis id="B918477FEB1AFF90FC0CFB939EB7FB6B" bold="true" box="[904,971,1150,1172]" italics="true" pageId="6" pageNumber="7">frigida</emphasis>
Sinclair, Vajda, Saigusa &amp; Shamshev
</taxonomicName>
<taxonomicNameLabel id="A22BFA04EB1AFF90FAD4FB9298E0FB6B" box="[1360,1436,1151,1172]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FB7098E1FB30" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FB7098E1FB30" pageId="6" pageNumber="7">
- Hind tibia with anterodorsal, posterodorsal and dorsal setae shorter than 2X tibial width (
<figureCitation id="135787E8EB1AFF90FBBFFB7199EFFB4D" box="[1083,1171,1180,1202]" captionStart="FIGURE 34" captionStartId="52.[151,250,1143,1168]" captionTargetBox="[151,1419,229,1124]" captionTargetId="figure@52.[151,1435,229,1124]" captionTargetPageId="52" captionText="FIGURE 34. Rhamphomyia (Pararhamphomyia) hoeli Frey. A. Male terminalia, lateral view; B. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773578" httpUri="https://zenodo.org/record/3773578/files/figure.png" pageId="6" pageNumber="7">Fig. 34B</figureCitation>
); apex of subepandrial lobe pointed (
<figureCitation id="135787E8EB1AFF90FEB3FB549CF2FB2F" box="[311,398,1209,1232]" captionStart="FIGURE 34" captionStartId="52.[151,250,1143,1168]" captionTargetBox="[151,1419,229,1124]" captionTargetId="figure@52.[151,1435,229,1124]" captionTargetPageId="52" captionText="FIGURE 34. Rhamphomyia (Pararhamphomyia) hoeli Frey. A. Male terminalia, lateral view; B. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773578" httpUri="https://zenodo.org/record/3773578/files/figure.png" pageId="6" pageNumber="7">Fig. 34A</figureCitation>
).......................................................................
<taxonomicName id="4C6CE0EEEB1AFF90FB76FB5798E1FB30" authority="Frey" authorityName=": Frey" authorityYear="1955" box="[1266,1437,1209,1231]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hoeli" subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FB76FB5798E1FB30" bold="true" box="[1266,1437,1209,1231]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB76FB579874FB30" bold="true" box="[1266,1288,1210,1231]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FA92FB579854FB30" bold="true" box="[1302,1320,1210,1231]" italics="true" pageId="6" pageNumber="7">P.</emphasis>
)
<emphasis id="B918477FEB1AFF90FAB1FB54981AFB30" bold="true" box="[1333,1382,1209,1231]" italics="true" pageId="6" pageNumber="7">hoeli</emphasis>
Frey
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FB3A98E0FABB" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FB3A98E1FAF6" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FB3A98E1FAF6" pageId="6" pageNumber="7">
16 Hind tibia clavate (
<figureCitation id="135787E8EB1AFF90FE1EFB3A9C88FB12" box="[410,500,1239,1261]" captionStart="FIGURE 36" captionStartId="54.[151,250,1971,1996]" captionTargetBox="[151,1430,769,1951]" captionTargetId="figure@54.[151,1435,769,1951]" captionTargetPageId="54" captionText="FIGURE 36. Rhamphomyia (Pararhamphomyia) kjellmanii Holmgren. A. uralensis Becker lectotype, lateral view; B. uralensis Becker type labels; C. Male terminalia, lateral view; D. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773582" httpUri="https://zenodo.org/record/3773582/files/figure.png" pageId="6" pageNumber="7">Fig. 36D</figureCitation>
); hind tarsomere 1 shorter than length of tarsomeres 24 combined (
<figureCitation id="135787E8EB1AFF90FB00FB3A99A7FB12" box="[1156,1243,1239,1261]" captionStart="FIGURE 36" captionStartId="54.[151,250,1971,1996]" captionTargetBox="[151,1430,769,1951]" captionTargetId="figure@54.[151,1435,769,1951]" captionTargetPageId="54" captionText="FIGURE 36. Rhamphomyia (Pararhamphomyia) kjellmanii Holmgren. A. uralensis Becker lectotype, lateral view; B. uralensis Becker type labels; C. Male terminalia, lateral view; D. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773582" httpUri="https://zenodo.org/record/3773582/files/figure.png" pageId="6" pageNumber="7">Fig. 36D</figureCitation>
)............................................................................................
<taxonomicName id="4C6CE0EEEB1AFF90FB00FB1998E1FAF6" authority="Holmgren" authorityName="Holmgren" box="[1156,1437,1268,1290]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="8" phylum="Arthropoda" rank="species" species="kjellmanii" subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FB00FB1998E1FAF6" bold="true" box="[1156,1437,1268,1290]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB00FB1999E6FAF6" bold="true" box="[1156,1178,1268,1289]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FB2CFB1999CBFAF6" bold="true" box="[1192,1207,1268,1289]" italics="true" pageId="6" pageNumber="7">P</emphasis>
.)
<emphasis id="B918477FEB1AFF90FB4FFB199853FAF5" bold="true" box="[1227,1327,1268,1290]" italics="true" pageId="6" pageNumber="7">kjellmanii</emphasis>
Holmgren
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FAFF98E0FABB" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FAFF98E0FABB" pageId="6" pageNumber="7">
- Hind tibia not clavate (
<figureCitation id="135787E8EB1AFF90FE3AFAFC9F6AFAD7" box="[446,534,1297,1320]" captionStart="FIGURE 40" captionStartId="60.[151,250,1618,1643]" captionTargetBox="[151,1435,336,1598]" captionTargetId="figure@60.[151,1435,336,1599]" captionTargetPageId="60" captionText="FIGURE 40. Rhamphomyia (Pararhamphomyia) omissinervis Becker. A. Male terminalia, lateral view; B. Male hindleg, anterior view C. Lectotype, lateral view; B. Type labels." figureDoi="http://doi.org/10.5281/zenodo.3773591" httpUri="https://zenodo.org/record/3773591/files/figure.png" pageId="6" pageNumber="7">Fig. 40B</figureCitation>
); hind tarsomere 1 longer than length of tarsomeres 24 combined (
<figureCitation id="135787E8EB1AFF90FB26FAFC998BFAD7" box="[1186,1271,1297,1320]" captionStart="FIGURE 40" captionStartId="60.[151,250,1618,1643]" captionTargetBox="[151,1435,336,1598]" captionTargetId="figure@60.[151,1435,336,1599]" captionTargetPageId="60" captionText="FIGURE 40. Rhamphomyia (Pararhamphomyia) omissinervis Becker. A. Male terminalia, lateral view; B. Male hindleg, anterior view C. Lectotype, lateral view; B. Type labels." figureDoi="http://doi.org/10.5281/zenodo.3773591" httpUri="https://zenodo.org/record/3773591/files/figure.png" pageId="6" pageNumber="7">Fig. 40B</figureCitation>
)...........................................................................................
<emphasis id="B918477FEB1AFF90FB09FAC298E0FABB" bold="true" box="[1165,1436,1327,1349]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB09FAC299DFFABB" bold="true" box="[1165,1187,1327,1348]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
<taxonomicName id="4C6CE0EEEB1AFF90FB2DFAC298E0FABB" authority="Becker" authorityName="Becker" box="[1193,1436,1327,1349]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="omissinervis" subGenus="Pararhamphomyia">
(
<emphasis id="B918477FEB1AFF90FB35FAC299BCFABB" bold="true" box="[1201,1216,1327,1348]" italics="true" pageId="6" pageNumber="7">P</emphasis>
.)
<emphasis id="B918477FEB1AFF90FB50FAC29833FABB" bold="true" box="[1236,1359,1327,1348]" italics="true" pageId="6" pageNumber="7">omissinervis</emphasis>
Becker
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FAA198E0FA63" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FAA198E0FA80" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FAA198E0FA80" pageId="6" pageNumber="7">
17 Phallus hidden within epandrium, leaving only base visible (
<figureCitation id="135787E8EB1AFF90FCA3FAA19E01FA9D" box="[807,893,1356,1378]" captionStart="FIGURE 17" captionStartId="30.[151,250,1971,1996]" captionTargetBox="[161,1426,166,1951]" captionTargetId="figure@30.[161,1426,166,1951]" captionTargetPageId="30" captionText="FIGURE 17. Rhamphomyia (Eorhamphomyia) shewelli sp. nov. A. Male hindleg, anterior view; B, C. Male terminalia, lateral view; D. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3773544" httpUri="https://zenodo.org/record/3773544/files/figure.png" pageId="6" pageNumber="7">Fig. 17B</figureCitation>
).................................................................................
<emphasis id="B918477FEB1AFF90FD26FA8798E0FA80" bold="true" box="[674,1436,1385,1407]" pageId="6" pageNumber="7">
<taxonomicName id="4C6CE0EEEB1AFF90FD26FA879835FA80" authority="Sinclair, Vajda, Saigusa &amp; Shamshev" authorityName="Sinclair, Vajda, Saigusa &amp; Shamshev" authorityYear="2019" box="[674,1353,1385,1407]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="shewelli" status="sp. nov." subGenus="Eorhamphomyia">
<emphasis id="B918477FEB1AFF90FD26FA879FCEFA80" bold="true" box="[674,690,1386,1407]" italics="true" pageId="6" pageNumber="7">R</emphasis>
. (
<emphasis id="B918477FEB1AFF90FD42FA849E11FA80" bold="true" box="[710,877,1385,1407]" italics="true" pageId="6" pageNumber="7">Eorhamphomyia</emphasis>
)
<emphasis id="B918477FEB1AFF90FCFFFA849EB7FA80" bold="true" box="[891,971,1385,1407]" italics="true" pageId="6" pageNumber="7">shewelli</emphasis>
Sinclair, Vajda, Saigusa &amp; Shamshev
</taxonomicName>
<taxonomicNameLabel id="A22BFA04EB1AFF90FACBFA8798E0FA80" box="[1359,1436,1386,1407]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FA6A98E0FA63" blockId="6.[151,1437,153,2023]" box="[151,1436,1415,1437]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FA6A98E0FA63" box="[151,1436,1415,1437]" pageId="6" pageNumber="7">- Phallus slim to hair-like, usually extending farther than length of epandrium, and often with curvatures................ 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FA4998E0FA28" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FA4998E0FA46" blockId="6.[151,1437,153,2023]" box="[151,1436,1444,1466]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FA4998E0FA46" box="[151,1436,1444,1466]" pageId="6" pageNumber="7">
18 Phallus with distinct loop on apical half around apex of epandrium (
<figureCitation id="135787E8EB1AFF90FCEAFA499EB5FA45" box="[878,969,1444,1466]" captionStart="FIGURE 51" captionStartId="75.[151,250,1877,1902]" captionTargetBox="[151,1435,481,1854]" captionTargetId="figure@75.[151,1435,481,1857]" captionTargetPageId="75" captionText="FIGURE 51. Rhamphomyia herschelli Malloch. A. Female hindleg, posterior view; B. Male hindleg, anterior view; C. Male terminalia, lateral view; D. Male terminalia, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3773615" httpUri="https://zenodo.org/record/3773615/files/figure.png" pageId="6" pageNumber="7">Figs 51C</figureCitation>
,
<figureCitation id="135787E8EB1AFF90FC57FA499E81FA45" box="[979,1021,1444,1466]" captionStart="FIGURE 57" captionStartId="82.[151,250,1970,1995]" captionTargetBox="[187,1386,166,1950]" captionTargetId="figure@82.[187,1400,166,1950]" captionTargetPageId="82" captionText="FIGURE 57. Rhamphomyia laevigata Loew, male terminalia, lateral view. A. Tuktayktuk (NWT); B. Blairemore (Alberta); C. (Pt. Pelee, Ontario); D. (Ottawa, Ontario); E. (Old Chelsea, Quebec); F. Lac La Hache (British Columbia)." figureDoi="http://doi.org/10.5281/zenodo.3773627" httpUri="https://zenodo.org/record/3773627/files/figure.png" pageId="6" pageNumber="7">57A</figureCitation>
)............................... 19
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13FA2F98E0FA28" blockId="6.[151,1437,153,2023]" box="[151,1436,1473,1495]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FA2F98E0FA28" box="[151,1436,1473,1495]" pageId="6" pageNumber="7">- Phallus without loop, gradually curved................................................................... 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13FA3298E0F9B3" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13FA3298E0F9EE" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13FA3298E0F9EE" pageId="6" pageNumber="7">
19 Sternite 8 with stout, “horn-like” lateral projections (
<figureCitation id="135787E8EB1AFF90FD65FA329E46FA0A" box="[737,826,1503,1525]" captionStart="FIGURE 51" captionStartId="75.[151,250,1877,1902]" captionTargetBox="[151,1435,481,1854]" captionTargetId="figure@75.[151,1435,481,1857]" captionTargetPageId="75" captionText="FIGURE 51. Rhamphomyia herschelli Malloch. A. Female hindleg, posterior view; B. Male hindleg, anterior view; C. Male terminalia, lateral view; D. Male terminalia, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3773615" httpUri="https://zenodo.org/record/3773615/files/figure.png" pageId="6" pageNumber="7">Fig. 51D</figureCitation>
); phallus with loop forming acute angle (
<figureCitation id="135787E8EB1AFF90FB4EFA32985EFA0A" box="[1226,1314,1503,1525]" captionStart="FIGURE 51" captionStartId="75.[151,250,1877,1902]" captionTargetBox="[151,1435,481,1854]" captionTargetId="figure@75.[151,1435,481,1857]" captionTargetPageId="75" captionText="FIGURE 51. Rhamphomyia herschelli Malloch. A. Female hindleg, posterior view; B. Male hindleg, anterior view; C. Male terminalia, lateral view; D. Male terminalia, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3773615" httpUri="https://zenodo.org/record/3773615/files/figure.png" pageId="6" pageNumber="7">Fig. 51C</figureCitation>
); hind tarsomere 1 swollen, wider than hind tibial width at apex (
<figureCitation id="135787E8EB1AFF90FD67FA119E44F9ED" box="[739,824,1532,1554]" captionStart="FIGURE 51" captionStartId="75.[151,250,1877,1902]" captionTargetBox="[151,1435,481,1854]" captionTargetId="figure@75.[151,1435,481,1857]" captionTargetPageId="75" captionText="FIGURE 51. Rhamphomyia herschelli Malloch. A. Female hindleg, posterior view; B. Male hindleg, anterior view; C. Male terminalia, lateral view; D. Male terminalia, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3773615" httpUri="https://zenodo.org/record/3773615/files/figure.png" pageId="6" pageNumber="7">Fig. 51B</figureCitation>
)................................
<taxonomicName id="4C6CE0EEEB1AFF90FB41FA1198E0F9EE" authority="Malloch" authorityName="Malloch" box="[1221,1436,1532,1554]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="herschelli">
<emphasis id="B918477FEB1AFF90FB41FA1198E0F9EE" bold="true" box="[1221,1436,1532,1554]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB41FA11983EF9ED" bold="true" box="[1221,1346,1532,1554]" italics="true" pageId="6" pageNumber="7">R. herschelli</emphasis>
Malloch
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13F9F798E0F9B3" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13F9F798E0F9B3" pageId="6" pageNumber="7">
- Sternite 8 without projections; phallus with small in-ward U-shaped loop (
<figureCitation id="135787E8EB1AFF90FC28F9F49979F9D0" box="[940,1029,1561,1583]" captionStart="FIGURE 57" captionStartId="82.[151,250,1970,1995]" captionTargetBox="[187,1386,166,1950]" captionTargetId="figure@82.[187,1400,166,1950]" captionTargetPageId="82" captionText="FIGURE 57. Rhamphomyia laevigata Loew, male terminalia, lateral view. A. Tuktayktuk (NWT); B. Blairemore (Alberta); C. (Pt. Pelee, Ontario); D. (Ottawa, Ontario); E. (Old Chelsea, Quebec); F. Lac La Hache (British Columbia)." figureDoi="http://doi.org/10.5281/zenodo.3773627" httpUri="https://zenodo.org/record/3773627/files/figure.png" pageId="6" pageNumber="7">Fig. 57A</figureCitation>
); hind tarsomere 1 slender (
<figureCitation id="135787E8EB1AFF90FA94F9F4981BF9D0" box="[1296,1383,1561,1583]" captionStart="FIGURE 56" captionStartId="81.[151,250,1903,1928]" captionTargetBox="[156,1435,1201,1883]" captionTargetId="figure@81.[151,1435,1201,1884]" captionTargetPageId="81" captionText="FIGURE 56. Rhamphomyia laevigata Loew. A. Female hindleg, posterior view; B. Male hindleg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3773625" httpUri="https://zenodo.org/record/3773625/files/figure.png" pageId="6" pageNumber="7">Fig. 56B</figureCitation>
)........................................................................................
<taxonomicName id="4C6CE0EEEB1AFF90FB6DF9DA98E0F9B3" authority="Loew" authorityName="Loew" box="[1257,1436,1590,1612]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="laevigata">
<emphasis id="B918477FEB1AFF90FB6DF9DA98E0F9B3" bold="true" box="[1257,1436,1590,1612]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB6DF9DA9822F9B3" bold="true" box="[1257,1374,1590,1612]" italics="true" pageId="6" pageNumber="7">R. laevigata</emphasis>
Loew
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13F9B998E0F93E" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13F9B998E0F95B" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13F9B998E0F95B" pageId="6" pageNumber="7">
20 Midleg distinctly modified: femur strongly curved upwards; tibia curved inwards, with 35 dark, stout mid-posterodorsal setae longer than 2X tibial width; tarsomere 1 spindle-shaped, covered with dense, erect and moderately long setae (
<figureCitation id="135787E8EB1AFF90FA83F99C9802F978" box="[1287,1406,1649,1671]" captionStart="FIGURE 42" captionStartId="63.[151,250,1860,1885]" captionTargetBox="[151,1435,174,1825]" captionTargetId="figure@63.[151,1435,174,1841]" captionTargetPageId="63" captionText="FIGURE 42. Rhamphomyia (Pararhamphomyia) petervajdai sp. nov. A. Male midleg, posterior view; B. Male midleg (dorsal view) and hindleg, lateral view; C, D. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773595" httpUri="https://zenodo.org/record/3773595/files/figure.png" pageId="6" pageNumber="7">Figs 42A, B</figureCitation>
)................................................
<emphasis id="B918477FEB1AFF90FC9EF96298E0F95B" bold="true" box="[794,1436,1678,1701]" pageId="6" pageNumber="7">
<taxonomicName id="4C6CE0EEEB1AFF90FC9EF9629835F95A" authority="Sinclair, Vajda, Saigusa &amp; Shamshev" authorityName="Sinclair, Vajda, Saigusa &amp; Shamshev" authorityYear="2019" box="[794,1353,1678,1701]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="petervajdai" status="sp. nov." subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FC9EF9629E4CF95B" bold="true" box="[794,816,1679,1700]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FCBAF9629E2CF95B" bold="true" box="[830,848,1679,1700]" italics="true" pageId="6" pageNumber="7">P.</emphasis>
)
<emphasis id="B918477FEB1AFF90FCDAF9639EB7F95B" bold="true" box="[862,971,1678,1700]" italics="true" pageId="6" pageNumber="7">petervajdai</emphasis>
Sinclair, Vajda, Saigusa &amp; Shamshev
</taxonomicName>
<taxonomicNameLabel id="A22BFA04EB1AFF90FACBF96298E0F95B" box="[1359,1436,1679,1700]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13F94098E0F93E" blockId="6.[151,1437,153,2023]" box="[151,1436,1708,1730]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13F94098E0F93E" box="[151,1436,1708,1730]" pageId="6" pageNumber="7">- Midleg simple....................................................................................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13F92798E0F8C8" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13F92798E0F8E6" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13F92798E0F8E6" pageId="6" pageNumber="7">
21 Apex of epandrium truncate (
<figureCitation id="135787E8EB1AFF90FD86F9249F01F91F" box="[514,637,1737,1760]" captionStart="FIGURE 44" captionStartId="66.[151,250,1980,2005]" captionTargetBox="[184,1404,166,1957]" captionTargetId="figure@66.[184,1404,166,1960]" captionTargetPageId="66" captionText="FIGURE 44. Rhamphomyia (Pararhamphomyia) septentrionalis sp. nov. A. Male terminalia, lateral view; B. Male hindlegs; C. Male terminalia, lateral view; D. Female habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773599" httpUri="https://zenodo.org/record/3773599/files/figure.png" pageId="6" pageNumber="7">Figs 44A, C</figureCitation>
); hind tibia with distinctive inward bend on apical part; hind tarsomere 1 spindleshaped, convex dorsally but straight ventrally (
<figureCitation id="135787E8EB1AFF90FD20F90A9F86F902" box="[676,762,1767,1789]" captionStart="FIGURE 44" captionStartId="66.[151,250,1980,2005]" captionTargetBox="[184,1404,166,1957]" captionTargetId="figure@66.[184,1404,166,1960]" captionTargetPageId="66" captionText="FIGURE 44. Rhamphomyia (Pararhamphomyia) septentrionalis sp. nov. A. Male terminalia, lateral view; B. Male hindlegs; C. Male terminalia, lateral view; D. Female habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773599" httpUri="https://zenodo.org/record/3773599/files/figure.png" pageId="6" pageNumber="7">Fig. 44B</figureCitation>
)...................................................................................................
<emphasis id="B918477FEB1AFF90FD72F8E998E0F8E6" bold="true" box="[758,1436,1796,1818]" pageId="6" pageNumber="7">
<taxonomicName id="4C6CE0EEEB1AFF90FD72F8E99835F8E5" authority="Sinclair, Vajda, Saigusa &amp; Shamshev" authorityName="Sinclair, Vajda, Saigusa &amp; Shamshev" authorityYear="2019" box="[758,1353,1796,1818]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="septentrionalis" status="sp. nov." subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FD72F8E99E70F8E6" bold="true" box="[758,780,1796,1817]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FC9EF8E99E50F8E6" bold="true" box="[794,812,1796,1817]" italics="true" pageId="6" pageNumber="7">P.</emphasis>
)
<emphasis id="B918477FEB1AFF90FCBEF8E99EB7F8E5" bold="true" box="[826,971,1796,1818]" italics="true" pageId="6" pageNumber="7">septentrionalis</emphasis>
Sinclair, Vajda, Saigusa &amp; Shamshev
</taxonomicName>
<taxonomicNameLabel id="A22BFA04EB1AFF90FACBF8E998E0F8E6" box="[1359,1436,1796,1817]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13F8CF98E0F8C8" blockId="6.[151,1437,153,2023]" box="[151,1436,1825,1847]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13F8CF98E0F8C8" box="[151,1436,1825,1847]" pageId="6" pageNumber="7">- Apex of epandrium rounded; hind tibia straight; hind tarsomere 1 clavate or uniformly slender....................... 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="309883C8EB1AFF90FF13F8D298E0F818" pageId="6" pageNumber="7">
<paragraph id="8BD39B6DEB1AFF90FF13F8D298E0F870" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13F8D298E0F870" pageId="6" pageNumber="7">
22 Anepisternum shiny and scutum with 12 pairs of shiny vittae; hind tarsomere 1 clavate, increasing in size apically such that apical width broader than hind tibia (
<figureCitation id="135787E8EB1AFF90FDCEF8B19FD8F88D" box="[586,676,1884,1906]" captionStart="FIGURE 48" captionStartId="71.[151,250,1508,1533]" captionTargetBox="[151,1434,174,1488]" captionTargetId="figure@71.[151,1435,174,1489]" captionTargetPageId="71" captionText="FIGURE 48. Rhamphomyia (Pararhamphomyia) ursinella Melander. A. Male terminalia, lateral view; B. Male hindleg, anterior view; C. ursina Malloch holotype, lateral view; D. ursina Malloch type labels." figureDoi="http://doi.org/10.5281/zenodo.3773609" httpUri="https://zenodo.org/record/3773609/files/figure.png" pageId="6" pageNumber="7">Fig. 48B</figureCitation>
); cerci weakly constricted in middle, posterior half parallel to epandrium (
<figureCitation id="135787E8EB1AFF90FAF1F8B19C70F86F" captionStart="FIGURE 48" captionStartId="71.[151,250,1508,1533]" captionTargetBox="[151,1434,174,1488]" captionTargetId="figure@71.[151,1435,174,1489]" captionTargetPageId="71" captionText="FIGURE 48. Rhamphomyia (Pararhamphomyia) ursinella Melander. A. Male terminalia, lateral view; B. Male hindleg, anterior view; C. ursina Malloch holotype, lateral view; D. ursina Malloch type labels." figureDoi="http://doi.org/10.5281/zenodo.3773609" httpUri="https://zenodo.org/record/3773609/files/figure.png" pageId="6" pageNumber="7">Fig. 48A</figureCitation>
)..........................................................................
<taxonomicName id="4C6CE0EEEB1AFF90FB13F89798E0F870" authority="Melander" authorityName="Melander" box="[1175,1436,1913,1935]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ursinella" subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FB13F89798E0F870" bold="true" box="[1175,1436,1913,1935]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB13F89799D1F870" bold="true" box="[1175,1197,1914,1935]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FB3FF89799B1F870" bold="true" box="[1211,1229,1914,1935]" italics="true" pageId="6" pageNumber="7">P.</emphasis>
)
<emphasis id="B918477FEB1AFF90FB5EF894984EF870" bold="true" box="[1242,1330,1913,1935]" italics="true" pageId="6" pageNumber="7">ursinella</emphasis>
Melander
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD39B6DEB1AFF90FF13F87A98E0F818" blockId="6.[151,1437,153,2023]" pageId="6" pageNumber="7">
<keyLead id="309D3858EB1AFF90FF13F87A98E0F818" pageId="6" pageNumber="7">
- Anepisternum pruinose and scutum with pruinescent vittae; hind tarsomere 1 evenly slender, nearly as broad as apical width of hind tibia, (
<figureCitation id="135787E8EB1AFF90FED4F8599CD5F835" box="[336,425,1972,1994]" captionStart="FIGURE 25" captionStartId="40.[151,250,1853,1878]" captionTargetBox="[151,1435,166,1819]" captionTargetId="figure@40.[151,1435,166,1834]" captionTargetPageId="40" captionText="FIGURE 25. Rhamphomyia (Pararhamphomyia) frigida sp. nov. A. Male terminalia, lateral view; B. Male hindleg, anterior view; C. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773560" httpUri="https://zenodo.org/record/3773560/files/figure.png" pageId="6" pageNumber="7">Fig. 25B</figureCitation>
); cerci strongly constricted in middle, posterior half strongly divergent from epandrium (
<figureCitation id="135787E8EB1AFF90FB7AF8599829F835" box="[1278,1365,1972,1994]" captionStart="FIGURE 25" captionStartId="40.[151,250,1853,1878]" captionTargetBox="[151,1435,166,1819]" captionTargetId="figure@40.[151,1435,166,1834]" captionTargetPageId="40" captionText="FIGURE 25. Rhamphomyia (Pararhamphomyia) frigida sp. nov. A. Male terminalia, lateral view; B. Male hindleg, anterior view; C. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3773560" httpUri="https://zenodo.org/record/3773560/files/figure.png" pageId="6" pageNumber="7">Fig. 25A</figureCitation>
).......................................................................
<taxonomicName id="4C6CE0EEEB1AFF90FB8CF83F98E0F818" authority="Henriksen &amp; Lundbeck" authorityName="Henriksen &amp; Lundbeck" box="[1032,1436,2001,2023]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="filicauda" subGenus="Pararhamphomyia">
<emphasis id="B918477FEB1AFF90FB8CF83F98E0F818" bold="true" box="[1032,1436,2001,2023]" pageId="6" pageNumber="7">
<emphasis id="B918477FEB1AFF90FB8CF83F9962F818" bold="true" box="[1032,1054,2002,2023]" italics="true" pageId="6" pageNumber="7">R.</emphasis>
(
<emphasis id="B918477FEB1AFF90FBA8F83F9942F818" bold="true" box="[1068,1086,2002,2023]" italics="true" pageId="6" pageNumber="7">P.</emphasis>
)
<emphasis id="B918477FEB1AFF90FBC8F83C99D8F818" bold="true" box="[1100,1188,2001,2023]" italics="true" pageId="6" pageNumber="7">filicauda</emphasis>
Henriksen &amp; Lundbeck
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>