treatments-xml/data/03/93/91/0393914C2A7B8722C282FDCE6B23F856.xml

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<mods:title id="16E743EA38247BBF71E048139535E8E2">Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)</mods:title>
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<treatment id="0393914C2A7B8722C282FDCE6B23F856" ID-DOI="http://doi.org/10.5281/zenodo.7705609" ID-GBIF-Taxon="209579501" ID-Zenodo-Dep="7705609" LSID="urn:lsid:plazi:treatment:0393914C2A7B8722C282FDCE6B23F856" httpUri="http://treatment.plazi.org/id/0393914C2A7B8722C282FDCE6B23F856" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
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<taxonomicName id="4C3A5BD92A7B8721C282FDCE6BE0F923" ID-CoL="BDMSZ" authority="Li &amp; Chen" authorityName="Li &amp; Chen" box="[151,696,582,609]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="nigridentata" status="sp. nov." subGenus="Steganina">
<emphasis id="B94EFC482A7B8721C282FDCE69A1F922" bold="true" box="[151,249,583,609]" italics="true" pageId="89" pageNumber="90">Stegana</emphasis>
(
<emphasis id="B94EFC482A7B8721C31FFDCE68DCF922" bold="true" box="[266,388,583,609]" italics="true" pageId="89" pageNumber="90">Steganina</emphasis>
) 
<emphasis id="B94EFC482A7B8721C380FDCF6B71F922" bold="true" box="[405,553,582,609]" italics="true" pageId="89" pageNumber="90">nigridentata</emphasis>
Li &amp; Chen
</taxonomicName>
, 
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<subSubSection id="C32073D12A7B8721C282FDE2682EF9C6" box="[151,374,618,645]" pageId="89" pageNumber="90" type="description">
<paragraph id="8B85205A2A7B8721C282FDE2682EF9C6" blockId="89.[151,803,582,645]" box="[151,374,618,645]" pageId="89" pageNumber="90">
(
<figureCitation id="13013CDF2A7B8721C28AFDE2681EF9C6" box="[159,326,619,645]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="89" pageNumber="90">Figs. 12E, 12F</figureCitation>
, 
<figureCitation id="13013CDF2A7B8721C344FDE36836F9C6" box="[337,366,618,645]" captionStart="FIGURE 55" captionStartId="89.[151,250,1663,1687]" captionTargetBox="[160,1430,820,1634]" captionTargetId="figure-241@89.[151,1436,813,1640]" captionTargetPageId="89" captionText="FIGURE 55. Stegana (Steganina) nigridentata Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705804" httpUri="https://zenodo.org/record/7705804/files/figure.png" pageId="89" pageNumber="90">55</figureCitation>
)
</paragraph>
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<subSubSection id="C32073D12A7B8721C282FD3B6DE2F856" pageId="89" pageNumber="90" type="diagnosis">
<paragraph id="8B85205A2A7B8721C282FD3B6DE2F856" blockId="89.[151,1437,690,789]" pageId="89" pageNumber="90">
<emphasis id="B94EFC482A7B8721C282FD3B684AF98F" bold="true" box="[151,274,690,716]" pageId="89" pageNumber="90">Diagnosis.</emphasis>
This species is similar to 
<emphasis id="B94EFC482A7B8721C026FD3A6B19F98E" box="[563,577,691,717]" italics="true" pageId="89" pageNumber="90">S</emphasis>
. 
<taxonomicName id="4C3A5BD92A7B8721C05AFD3A6B86F98E" box="[591,734,691,717]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="menghai" status="sp. nov." subGenus="Steganina">
(
<emphasis id="B94EFC482A7B8721C04DFD3A6B3EF98E" box="[600,614,691,717]" italics="true" pageId="89" pageNumber="90">S</emphasis>
.) 
<emphasis id="B94EFC482A7B8721C06BFD3A6B86F98E" box="[638,734,691,717]" italics="true" pageId="89" pageNumber="90">menghai</emphasis>
</taxonomicName>
<emphasis id="B94EFC482A7B8721C0F0FD3A6A67F98E" bold="true" box="[741,831,691,717]" pageId="89" pageNumber="90">
<taxonomicNameLabel id="A27D41332A7B8721C0F0FD3A6A67F98E" box="[741,831,691,717]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the shape of male terminalia (
<figureCitation id="13013CDF2A7B8721C6BBFD3A6C53F98E" box="[1198,1291,690,717]" captionStart="FIGURE 50" captionStartId="84.[151,250,970,994]" captionTargetBox="[163,1431,186,938]" captionTargetId="figure-30@84.[151,1436,181,945]" captionTargetPageId="84" captionText="FIGURE 50. Stegana (Steganina) menghai Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705786" httpUri="https://zenodo.org/record/7705786/files/figure.png" pageId="89" pageNumber="90">Figs. 50</figureCitation>
, 
<figureCitation id="13013CDF2A7B8721C702FD3B6C6EF98E" box="[1303,1334,690,717]" captionStart="FIGURE 55" captionStartId="89.[151,250,1663,1687]" captionTargetBox="[160,1430,820,1634]" captionTargetId="figure-241@89.[151,1436,813,1640]" captionTargetPageId="89" captionText="FIGURE 55. Stegana (Steganina) nigridentata Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705804" httpUri="https://zenodo.org/record/7705804/files/figure.png" pageId="89" pageNumber="90">55</figureCitation>
), but can be distinguished from the latter by having the pregonite blurry or absent (
<figureCitation id="13013CDF2A7B8721C1D7FD5E6D1FF9B2" box="[962,1095,726,753]" captionStart="FIGURE 50" captionStartId="84.[151,250,970,994]" captionTargetBox="[163,1431,186,938]" captionTargetId="figure-30@84.[151,1436,181,945]" captionTargetPageId="84" captionText="FIGURE 50. Stegana (Steganina) menghai Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705786" httpUri="https://zenodo.org/record/7705786/files/figure.png" pageId="89" pageNumber="90">Fig. 50B, C</figureCitation>
); the aedeagus laterally deeply dehiscent and with 4 small black teeth per side, distally lacking serrated process (
<figureCitation id="13013CDF2A7B8721C633FD726DF5F856" box="[1062,1197,762,789]" captionStart="FIGURE 55" captionStartId="89.[151,250,1663,1687]" captionTargetBox="[160,1430,820,1634]" captionTargetId="figure-241@89.[151,1436,813,1640]" captionTargetPageId="89" captionText="FIGURE 55. Stegana (Steganina) nigridentata Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705804" httpUri="https://zenodo.org/record/7705804/files/figure.png" pageId="89" pageNumber="90">Fig. 55B, C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4570D22A7B8721C282F9F66BEFFD9C" ID-DOI="http://doi.org/10.5281/zenodo.7705804" ID-Zenodo-Dep="7705804" httpUri="https://zenodo.org/record/7705804/files/figure.png" pageId="89" pageNumber="90" startId="89.[151,250,1663,1687]" targetBox="[160,1430,820,1634]" targetPageId="89">
<paragraph id="8B85205A2A7B8721C282F9F66BEFFD9C" blockId="89.[151,1437,1663,1759]" pageId="89" pageNumber="90">
<emphasis id="B94EFC482A7B8721C282F9F66879FDD4" bold="true" box="[151,289,1663,1687]" pageId="89" pageNumber="90">FIGURE 55.</emphasis>
<taxonomicName id="4C3A5BD92A7B8721C33CF9096BAEFDD4" authority="Li &amp; Chen" authorityName="Li &amp; Chen" box="[297,758,1663,1687]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="nigridentata" status="sp. nov." subGenus="Steganina">
<emphasis id="B94EFC482A7B8721C33CF9096823FDD4" box="[297,379,1664,1687]" italics="true" pageId="89" pageNumber="90">Stegana</emphasis>
(
<emphasis id="B94EFC482A7B8721C39FF90968B7FDD4" box="[394,495,1664,1687]" italics="true" pageId="89" pageNumber="90">Steganina</emphasis>
) 
<emphasis id="B94EFC482A7B8721C015F9F66B25FDD4" box="[512,637,1663,1687]" italics="true" pageId="89" pageNumber="90">nigridentata</emphasis>
Li &amp; Chen
</taxonomicName>
, 
<emphasis id="B94EFC482A7B8721C114F9F66A0BFDD4" bold="true" box="[769,851,1663,1687]" pageId="89" pageNumber="90">
<taxonomicNameLabel id="A27D41332A7B8721C114F9F66A0BFDD4" box="[769,851,1663,1687]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view.
</paragraph>
</caption>
<subSubSection id="C32073D12A7B8722C2D2F8846B28F95A" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90" type="description">
<paragraph id="8B85205A2A7B8722C2D2F8846B9BFBBA" blockId="89.[151,1437,1805,1940]" lastBlockId="90.[151,1437,150,789]" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
<emphasis id="B94EFC482A7B8721C2D2F8846801FC64" bold="true" box="[199,345,1805,1831]" pageId="89" pageNumber="90">Description.</emphasis>
Male and female: First flagellomere dark brown. Clypeus dark brown medially, yellow laterally. Mesonotum brown to black medially, brownish yellow to brown laterally (
<figureCitation id="13013CDF2A7B8721C1CFF8B86D18FC0F" box="[986,1088,1841,1868]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="89" pageNumber="90">Fig. 12E</figureCitation>
). Katepisternum yellow above, brownish below (
<figureCitation id="13013CDF2A7B8721C349F8DC6899FC33" box="[348,449,1877,1904]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="89" pageNumber="90">Fig. 12F</figureCitation>
). Scutellum dark brown to black (
<figureCitation id="13013CDF2A7B8721C151F8DC6AF3FC33" box="[836,939,1877,1904]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="89" pageNumber="90">Fig. 12E</figureCitation>
). Halter mostly grey (
<figureCitation id="13013CDF2A7B8721C6B1F8DC6C51FC33" box="[1188,1289,1877,1904]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="89" pageNumber="90">Fig. 12F</figureCitation>
). Fore femur with a row of 3 or 4 long setae on distal part of posteroventral surface. Abdominal tergites mostly black (
<figureCitation id="13013CDF2A7B8721C726F8F06CCFFCD7" box="[1331,1431,1913,1940]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="89" pageNumber="90">Fig. 12E</figureCitation>
, margin, lacking pubescence (
<figureCitation id="13013CDF2A788722C3F4FF1E6B13FBF2" box="[481,587,150,177]" captionStart="FIGURE 55" captionStartId="89.[151,250,1663,1687]" captionTargetBox="[160,1430,820,1634]" captionTargetId="figure-241@89.[151,1436,813,1640]" captionTargetPageId="89" captionText="FIGURE 55. Stegana (Steganina) nigridentata Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705804" httpUri="https://zenodo.org/record/7705804/files/figure.png" pageId="90" pageNumber="91">Fig. 55A</figureCitation>
). Cercus with much pubescence (
<figureCitation id="13013CDF2A788722C1D3FF1E6D68FBF2" box="[966,1072,150,177]" captionStart="FIGURE 55" captionStartId="89.[151,250,1663,1687]" captionTargetBox="[160,1430,820,1634]" captionTargetId="figure-241@89.[151,1436,813,1640]" captionTargetPageId="89" captionText="FIGURE 55. Stegana (Steganina) nigridentata Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705804" httpUri="https://zenodo.org/record/7705804/files/figure.png" pageId="90" pageNumber="91">Fig. 55A</figureCitation>
). Hypandrium expanded anteromedially (
<figureCitation id="13013CDF2A788722C31DFF3268D7FB96" box="[264,399,186,213]" captionStart="FIGURE 55" captionStartId="89.[151,250,1663,1687]" captionTargetBox="[160,1430,820,1634]" captionTargetId="figure-241@89.[151,1436,813,1640]" captionTargetPageId="89" captionText="FIGURE 55. Stegana (Steganina) nigridentata Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705804" httpUri="https://zenodo.org/record/7705804/files/figure.png" pageId="90" pageNumber="91">Fig. 55B, C</figureCitation>
). Aedeagal sheath distally dorsad curved and with numerous minute, serrated processes (
<figureCitation id="13013CDF2A788722C77AFF3269B2FBBA" captionStart="FIGURE 55" captionStartId="89.[151,250,1663,1687]" captionTargetBox="[160,1430,820,1634]" captionTargetId="figure-241@89.[151,1436,813,1640]" captionTargetPageId="89" captionText="FIGURE 55. Stegana (Steganina) nigridentata Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705804" httpUri="https://zenodo.org/record/7705804/files/figure.png" pageId="90" pageNumber="91">Fig. 55D, E</figureCitation>
). Subepandrial sclerite vaulted (
<figureCitation id="13013CDF2A788722C047FF566BEFFBBA" box="[594,695,222,249]" captionStart="FIGURE 55" captionStartId="89.[151,250,1663,1687]" captionTargetBox="[160,1430,820,1634]" captionTargetId="figure-241@89.[151,1436,813,1640]" captionTargetPageId="89" captionText="FIGURE 55. Stegana (Steganina) nigridentata Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705804" httpUri="https://zenodo.org/record/7705804/files/figure.png" pageId="90" pageNumber="91">Fig. 55F</figureCitation>
).
</paragraph>
<paragraph id="8B85205A2A788722C2D2FE8A6B28F95A" blockId="90.[151,1437,150,789]" pageId="90" pageNumber="91">
<emphasis id="B94EFC482A788722C2D2FE8A6B57FA5F" bold="true" box="[199,527,258,285]" pageId="90" pageNumber="91">Measurements and indices.</emphasis>
BL = 
<quantity id="4CC28DBF2A788722C04EFE8A6B99FA5E" box="[603,705,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.28" pageId="90" pageNumber="91" unit="mm" value="3.28">3.28 mm</quantity>
in 
<typeStatus id="54819EF82A788722C0FDFE8A6A12FA5E" box="[744,842,259,285]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
(
<quantity id="4CC28DBF2A788722C14FFE8B6AB3FA5E" box="[858,1003,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="8.128" metricValueMax="8.9408" metricValueMin="7.3152" pageId="90" pageNumber="91" unit="in" value="3.2" valueMax="3.52" valueMin="2.88">2.88–3.52 in</quantity>
5♂, 
<quantity id="4CC28DBF2A788722C633FE8A6DEDFA5E" box="[1062,1205,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="9.4488" metricValueMax="10.668" metricValueMin="8.2296" pageId="90" pageNumber="91" unit="in" value="3.72" valueMax="4.2" valueMin="3.24">3.24–4.20 in</quantity>
<specimenCount id="9D3CEBD32A788722C6A8FE8B6DB8FA5E" box="[1213,1248,258,285]" pageId="90" pageNumber="91" type="female">5♀</specimenCount>
<typeStatus id="54819EF82A788722C6FCFE8A6C01FA5E" box="[1257,1369,259,285]" pageId="90" pageNumber="91" type="paratype">paratypes</typeStatus>
), ThL = 
<quantity id="4CC28DBF2A788722C2BAFEAE684CFA02" box="[175,276,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" pageId="90" pageNumber="91" unit="mm" value="1.44">1.44 mm</quantity>
(
<quantity id="4CC28DBF2A788722C331FEAE68ECFA02" box="[292,436,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="4.1021" metricValueMax="4.572" metricValueMin="3.6322" pageId="90" pageNumber="91" unit="in" value="1.615" valueMax="1.8" valueMin="1.43">1.43–1.80 in</quantity>
♂, 
<quantity id="4CC28DBF2A788722C3F5FEAF6B36FA02" box="[480,622,294,321]" metricMagnitude="-2" metricUnit="m" metricValue="4.838699999999999" metricValueMax="5.207" metricValueMin="4.4704" pageId="90" pageNumber="91" unit="in" value="1.9049999999999998" valueMax="2.05" valueMin="1.76">1.76–2.05 in</quantity>
<specimenCount id="9D3CEBD32A788722C062FEAE6BD2FA02" box="[631,650,295,321]" pageId="90" pageNumber="91" type="female">♀</specimenCount>
), WL = 
<quantity id="4CC28DBF2A788722C0FBFEAE6A0CFA02" box="[750,852,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.44" pageId="90" pageNumber="91" unit="mm" value="2.44">2.44 mm</quantity>
(2.36–3.00 in ♂, 
<quantity id="4CC28DBF2A788722C60AFEAE6DF6FA02" box="[1055,1198,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="7.5692" metricValueMax="8.2296" metricValueMin="6.9088" pageId="90" pageNumber="91" unit="in" value="2.98" valueMax="3.24" valueMin="2.72">2.72–3.24 in</quantity>
<specimenCount id="9D3CEBD32A788722C6A3FEAE6D91FA02" box="[1206,1225,295,321]" pageId="90" pageNumber="91" type="female">♀</specimenCount>
), WW = 
<quantity id="4CC28DBF2A788722C722FEAE6CC5FA02" box="[1335,1437,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="90" pageNumber="91" unit="mm" value="1.2">1.20 mm</quantity>
(
<quantity id="4CC28DBF2A788722C28AFEC26877FA26" box="[159,303,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="3.175" metricValueMax="3.5559999999999996" metricValueMin="2.794" pageId="90" pageNumber="91" unit="in" value="1.25" valueMax="1.4" valueMin="1.1">1.10–1.40 in</quantity>
♂, 
<quantity id="4CC28DBF2A788722C349FEC268A0FA26" box="[348,504,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="3.56616" metricValueMax="4.064" metricValueMin="3.06832" pageId="90" pageNumber="91" unit="in" value="1.404" valueMax="1.6" valueMin="1.208">1.208–1.60 in</quantity>
<specimenCount id="9D3CEBD32A788722C014FEC26B4CFA26" box="[513,532,331,357]" pageId="90" pageNumber="91" type="female">♀</specimenCount>
), arb = 14/5 (8–14/4–6), avd = 0.60 (0.44–0.91), adf = 1.43 (1.57–2.25), flw = 1.57 (1.50–2.60), FW/HW = 0.38 (0.27–0.38), ch/o = 0.22 (0.13–0.27), prorb = 1.25 (0.75–1.14), rcorb = 0.92 (0.58–1.17), vb = 0.75 (0.29–0.67), dcl = 0.75 (0.27–0.75), presctl = 0.63 (0.50–0.93), sctl = 1.50 (1.24–1.93), sterno = 0.92 (0.59–0.82), orbito = 2.60 (1.67–2.50), dcp = 0.21 (0.18–0.26), sctlp = 1.73 (1.50–2.27), C = 2.62 (1.73–2.25), 4c = 0.81 (1.09–1.26), 4v = 1.63 (1.63–2.19), 5x = 1.60 (1.36–2.00), ac = 13.00 (10.00–21.25), M = 0.50 (0.50–0.81), C3F = 0.57 (0.69–0.82).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A788722C2D2FDAA6D68F9EA" pageId="90" pageNumber="91" type="materials_examined">
<paragraph id="8B85205A2A788722C2D2FDAA6D68F9EA" blockId="90.[151,1437,150,789]" pageId="90" pageNumber="91">
<emphasis id="B94EFC482A788722C2D2FDAA68DEF97F" bold="true" box="[199,390,546,573]" pageId="90" pageNumber="91">Type materials.</emphasis>
<materialsCitation id="3B522A072A788722C387FDAA6A1FF922" ID-GBIF-Occurrence="4097216429" collectingDate="2004-10-23" collectionCode="SCAU" collectorName="H. W. Chen" country="China" elevation="700" latitude="25.857779" location="Xing'an" longLatPrecision="20" longitude="110.41694" municipality="Maoershan" pageId="90" pageNumber="91" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="54819EF82A788722C387FDAA68A2F97E" box="[402,506,547,573]" pageId="90" pageNumber="91" type="holotype">Holotype</typeStatus>
♂ (
<collectionCode id="ED2BB89F2A788722C025FDAA6B27F97E" box="[560,639,547,573]" pageId="90" pageNumber="91">SCAU</collectionCode>
, No. 121132), 
<collectingCountry id="F32D60CA2A788722C123FDAA6ACBF97E" box="[822,915,547,573]" name="China" pageId="90" pageNumber="91">CHINA</collectingCountry>
: 
<collectingMunicipality id="6BE1BA202A788722C1B6FDAA6D7AF97E" box="[931,1058,547,573]" pageId="90" pageNumber="91">Maoershan</collectingMunicipality>
, 
<location id="8EE576812A788722C626FDAA6DD5F97E" LSID="urn:lsid:plazi:treatment:0393914C2A7B8722C282FDCE6B23F856:8EE576812A788722C626FDAA6DD5F97E" box="[1075,1165,547,573]" country="China" latitude="25.857779" longLatPrecision="20" longitude="110.41694" municipality="Maoershan" name="Xing'an" pageId="90" pageNumber="91" stateProvince="Guangxi">Xing'an</location>
, 
<collectingRegion id="49FEEEB82A788722C68AFDAA6C5AF97E" box="[1183,1282,547,573]" country="China" name="Guangxi" pageId="90" pageNumber="91">Guangxi</collectingRegion>
, 
<geoCoordinate id="EE0E469D2A788722C706FDAB6CC0F97E" box="[1299,1432,546,573]" degrees="25" direction="north" minutes="51" orientation="latitude" pageId="90" pageNumber="91" precision="15" seconds="28" value="25.857779">25°51'28''N</geoCoordinate>
, 
<geoCoordinate id="EE0E469D2A788722C282FDCF687EF922" box="[151,294,582,609]" degrees="110" direction="east" minutes="25" orientation="longitude" pageId="90" pageNumber="91" precision="15" seconds="01" value="110.41694">110°25'01''E</geoCoordinate>
, alt. 
<quantity id="4CC28DBF2A788722C348FDCE68C7F922" box="[349,415,583,609]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="90" pageNumber="91" unit="m" value="700.0">
<elevation id="0017C7692A788722C348FDCE68C7F922" box="[349,415,583,609]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="90" pageNumber="91" unit="m" value="700.0">700m</elevation>
</quantity>
, 
<date id="FF84069A2A788722C3BEFDCE6B45F922" box="[427,541,583,609]" pageId="90" pageNumber="91" value="2004-10-23">
<collectingDate id="EFC0FF722A788722C3BEFDCE6B45F922" box="[427,541,583,609]" pageId="90" pageNumber="91" value="2004-10-23">23.x.2004</collectingDate>
</date>
, ex tree trunk, 
<collectorName id="26CF458C2A788722C0D3FDCE6A1FF922" box="[710,839,583,609]" pageId="90" pageNumber="91">H.W. Chen</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B522A072A788722C144FDCE6BAFF9C6" ID-GBIF-Occurrence="4097216412" collectingDate="2006-10-19" collectionCode="SCAU" collectorName="H. W. Chen" country="China" elevation="700" latitude="25.857779" location="Xing'an" longLatPrecision="20" longitude="110.41694" municipality="Maoershan" pageId="90" pageNumber="91" specimenCount="18" specimenCount-female="18" stateProvince="Guangxi" typeStatus="paratype">
<typeStatus id="54819EF82A788722C144FDCE6A9AF922" box="[849,962,583,609]" pageId="90" pageNumber="91" type="paratype">Paratypes</typeStatus>
: 6♂, 
<specimenCount id="9D3CEBD32A788722C1EAFDCE6D68F922" box="[1023,1072,583,609]" pageId="90" pageNumber="91" type="female">18♀</specimenCount>
(
<collectionCode id="ED2BB89F2A788722C62AFDCE6DD6F922" box="[1087,1166,583,609]" pageId="90" pageNumber="91">SCAU</collectionCode>
, Nos. 121133–56, DNA #1461, 1462), 
<date id="FF84069A2A788722C32EFDE268F6F9C6" box="[315,430,619,645]" pageId="90" pageNumber="91" value="2006-10-19">
<collectingDate id="EFC0FF722A788722C32EFDE268F6F9C6" box="[315,430,619,645]" pageId="90" pageNumber="91" value="2006-10-19">19.x.2006</collectingDate>
</date>
, other data same as holotype
</materialsCitation>
; 
<materialsCitation id="3B522A072A788722C117FDE26D68F9EA" ID-GBIF-Occurrence="4097216318" collectingDate="2012-05-13" collectionCode="SCAU" collectorName="J. J. Gao" country="China" elevation="1400" latitude="22.738611" location="Pu'er" longLatPrecision="21" longitude="101.11694" municipality="Yixiang" pageId="90" pageNumber="91" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D3CEBD32A788722C117FDE26A7DF9C6" box="[770,805,619,645]" pageId="90" pageNumber="91" type="female">2♀</specimenCount>
(
<collectionCode id="ED2BB89F2A788722C121FDE26ADBF9C6" box="[820,899,619,645]" pageId="90" pageNumber="91">SCAU</collectionCode>
, Nos. 125606, 07, DNA #2190), 
<collectingMunicipality id="6BE1BA202A788722C6E2FDE26C0BF9C6" box="[1271,1363,619,645]" pageId="90" pageNumber="91">Yixiang</collectingMunicipality>
, 
<location id="8EE576812A788722C74BFDE26CC0F9C6" LSID="urn:lsid:plazi:treatment:0393914C2A7B8722C282FDCE6B23F856:8EE576812A788722C74BFDE26CC0F9C6" box="[1374,1432,619,645]" country="China" latitude="22.738611" longLatPrecision="21" longitude="101.11694" municipality="Yixiang" name="Pu'er" pageId="90" pageNumber="91" stateProvince="Yunnan">Pu'er</location>
, 
<collectingRegion id="49FEEEB82A788722C282FD0669A8F9EA" box="[151,240,655,681]" country="China" name="Yunnan" pageId="90" pageNumber="91">Yunnan</collectingRegion>
, 
<geoCoordinate id="EE0E469D2A788722C2EEFD0668D8F9EA" box="[251,384,655,681]" degrees="22" direction="north" minutes="44" orientation="latitude" pageId="90" pageNumber="91" precision="15" seconds="19" value="22.738611">22°44'19''N</geoCoordinate>
, 
<geoCoordinate id="EE0E469D2A788722C39EFD066B43F9EA" box="[395,539,655,681]" degrees="101" direction="east" minutes="07" orientation="longitude" pageId="90" pageNumber="91" precision="15" seconds="01" value="101.11694">101°07'01''E</geoCoordinate>
, alt. 
<quantity id="4CC28DBF2A788722C044FD066BF9F9EA" box="[593,673,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="90" pageNumber="91" unit="m" value="1400.0">
<elevation id="0017C7692A788722C044FD066BF9F9EA" box="[593,673,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="90" pageNumber="91" unit="m" value="1400.0">1400m</elevation>
</quantity>
, 
<date id="FF84069A2A788722C0B9FD066A46F9EA" box="[684,798,655,681]" pageId="90" pageNumber="91" value="2012-05-13">
<collectingDate id="EFC0FF722A788722C0B9FD066A46F9EA" box="[684,798,655,681]" pageId="90" pageNumber="91" value="2012-05-13">13.v.2012</collectingDate>
</date>
, ex tree trunks, 
<collectorName id="26CF458C2A788722C1DAFD066D74F9EA" box="[975,1068,655,681]" pageId="90" pageNumber="91">J.J. Gao</collectorName>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A788722C2D2FD3B6BD7F98E" box="[199,655,690,717]" pageId="90" pageNumber="91" type="distribution">
<paragraph id="8B85205A2A788722C2D2FD3B6BD7F98E" blockId="90.[151,1437,150,789]" box="[199,655,690,717]" pageId="90" pageNumber="91">
<emphasis id="B94EFC482A788722C2D2FD3B6839F98F" bold="true" box="[199,353,690,716]" pageId="90" pageNumber="91">Distribution.</emphasis>
<collectingCountry id="F32D60CA2A788722C372FD3A68F2F98E" box="[359,426,691,717]" name="China" pageId="90" pageNumber="91">China</collectingCountry>
(
<collectingRegion id="49FEEEB82A788722C3ACFD3A6B46F98E" box="[441,542,691,717]" country="China" name="Guangxi" pageId="90" pageNumber="91">Guangxi</collectingRegion>
, 
<collectingRegion id="49FEEEB82A788722C03DFD3A6BDBF98E" box="[552,643,691,717]" country="China" name="Yunnan" pageId="90" pageNumber="91">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A788722C2D2FD5E6B23F856" pageId="90" pageNumber="91" type="etymology">
<paragraph id="8B85205A2A788722C2D2FD5E6B23F856" blockId="90.[151,1437,150,789]" pageId="90" pageNumber="91">
<emphasis id="B94EFC482A788722C2D2FD5E6816F9B2" bold="true" box="[199,334,727,753]" pageId="90" pageNumber="91">Etymology.</emphasis>
A combination of the Latin words &quot; 
<emphasis id="B94EFC482A788722C0E5FD5E6A73F9B3" box="[752,811,727,752]" italics="true" pageId="90" pageNumber="91">niger</emphasis>
&quot; (= black) and &quot; 
<emphasis id="B94EFC482A788722C1FAFD5E6D16F9B2" box="[1007,1102,727,753]" italics="true" pageId="90" pageNumber="91">dentatus</emphasis>
&quot; (= dentate), referring to the aedeagus laterally with 4 small black teeth.
</paragraph>
</subSubSection>
</treatment>
</document>