153 lines
13 KiB
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153 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.79.57686" ID-GBIF-Dataset="054f2212-802b-42b3-bba6-e8bca2b22198" ID-Pensoft-Pub="1314-2607-79-77" ID-Pensoft-UUID="9771F5D9B2AA51558A0A56AC96840EC0" ID-Zenodo-Dep="4255453" ID-ZooBank="ED8D2D07C0384429B5FF1011D2B48907" ModsDocID="1314-2607-79-77" checkinTime="1604097865600" checkinUser="pensoft" docAuthor="Martins, Andre L. & Melo, Gabriel A. R." docDate="2020" docId="62F4983C9C9D5282B73D66AC9FBB144C" docLanguage="en" docName="JourHymenoptRes 79: 77-88" docOrigin="Journal of Hymenoptera Research 79" docPubDate="2020-10-30" docSource="http://dx.doi.org/10.3897/jhr.79.57686" docTitle="Thaumatodryinus miocenicus Olmi 1995" docType="treatment" docVersion="4" id="9771F5D9B2AA51558A0A56AC96840EC0" lastPageNumber="77" masterDocId="9771F5D9B2AA51558A0A56AC96840EC0" masterDocTitle="Revision of the fossil species of Thaumatodryinus Perkins from Dominican amber, with a new combination and description of a new species (Hymenoptera, Dryinidae)" masterLastPageNumber="88" masterPageNumber="77" pageNumber="77" updateTime="1678754720945" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the fossil species of Thaumatodryinus Perkins from Dominican amber, with a new combination and description of a new species (Hymenoptera, Dryinidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Martins, Andre L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4794-0644</mods:nameIdentifier>
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<mods:affiliation>Laboratorio de Biologia Comparada de Hymenoptera, Departamento de Zoologia, Universidade Federal do Parana, Cx. postal 19020, 81531 - 980, Curitiba, PR, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Melo, Gabriel A. R.</mods:namePart>
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<mods:affiliation>Laboratorio de Biologia Comparada de Hymenoptera, Departamento de Zoologia, Universidade Federal do Parana, Cx. postal 19020, 81531 - 980, Curitiba, PR, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">garmelo@ufpr.br</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-10-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>79</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>77</mods:start>
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<mods:end>88</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.79.57686</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.79.57686</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-79-77</mods:identifier>
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<mods:identifier type="ZooBank">ED8D2D07C0384429B5FF1011D2B48907</mods:identifier>
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<mods:identifier type="Pensoft-UUID">9771F5D9B2AA51558A0A56AC96840EC0</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4255453</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168745944" LSID="urn:lsid:plazi:treatment:62F4983C9C9D5282B73D66AC9FBB144C" httpUri="http://treatment.plazi.org/id/62F4983C9C9D5282B73D66AC9FBB144C" lastPageNumber="77" pageId="0" pageNumber="77">
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<subSubSection pageId="0" pageNumber="77" type="nomenclature">
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<paragraph pageId="0" pageNumber="77">
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†
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<taxonomicName LSID="62F4983C-9C9D-5282-B73D-66AC9FBB144C" authority="Olmi, 1995" authorityName="Olmi" authorityYear="1995" class="Insecta" family="Dryinidae" genus="Thaumatodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thaumatodryinus miocenicus" order="Hymenoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="miocenicus">Thaumatodryinus miocenicus Olmi, 1995</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="77" type="description">
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<paragraph pageId="0" pageNumber="77">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. † Thaumatodryinus miocenicus Olmi, 1995, female holotype. A habitus, dorsal view B habitus, ventral view C habitus, lateral view D habitus, dorsal view E head and mesosoma, ventral view F habitus, lateral view. Scale bars: 1 mm (A-C), 1 mm (D, F), 1 mm (E)." figureDoi="10.3897/jhr.79.57686.figure2" httpUri="https://binary.pensoft.net/fig/469491" pageId="0" pageNumber="77">Figure 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="77" type="reference_group">
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<paragraph pageId="0" pageNumber="77">
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<taxonomicName authorityName="Olmi" authorityYear="1995" class="Insecta" family="Dryinidae" genus="Thaumatodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thaumatodryinus miocenicus" order="Hymenoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="miocenicus">Thaumatodryinus miocenicus</taxonomicName>
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Olmi, 1995: 263. Female holotype. Dominican Republic: amber from unknown mine, probably El Valle mine (AMNH, examined).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="77" type="extended diagnosis">
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<paragraph pageId="0" pageNumber="77">Extended diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="77">
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Female holotype, 5.6 mm (see fig. 1-2 in
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<bibRefCitation author="Olmi, M" journalOrPublisher="Redia" pageId="0" pageNumber="77" pagination="253 - 271" refId="B11" refString="Olmi, M, 1995. Dryinids and embolemids in amber (Hymenoptera Dryinidae et Embolemidae). Redia 78: 253 - 271" title="Dryinids and embolemids in amber (Hymenoptera Dryinidae et Embolemidae)." volume="78" year="1995">Olmi (1995)</bibRefCitation>
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and fig. 140 in
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3792.2.1" author="Olmi, M" journalOrPublisher="Zootaxa" pageId="0" pageNumber="77" pagination="1 - 534" refId="B13" refString="Olmi, M, Virla, EG, 2014. Dryinidae of the Neotropical region (Hymenoptera: Chrysidoidea). Zootaxa 3792 (1): 1 - 534, DOI: https://doi.org/10.11646/zootaxa.3792.2.1" title="Dryinidae of the Neotropical region (Hymenoptera: Chrysidoidea)." url="https://doi.org/10.11646/zootaxa.3792.2.1" volume="3792" year="2014">Olmi and Virla (2014)</bibRefCitation>
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).
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<taxonomicName authorityName="Olmi" authorityYear="1995" class="Insecta" family="Dryinidae" genus="Thaumatodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thaumatodryinus miocenicus" order="Hymenoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="miocenicus">
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<emphasis italics="true" pageId="0" pageNumber="77">Thaumatodryinus miocenicus</emphasis>
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</taxonomicName>
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is easily recognized by the following combination of characters: black head, except antenna brown to dark brown; mesosoma mostly black; legs and metasoma apparently brown. Antenna with dense and short setae. Head slightly convex, granulated; upper frons and vertex with pilosity; occiput slightly excavated and densely hairy; frontal line present but apparently incomplete in holotype (complete in paratype Do-4066-M); mid portion of clypeus with anterior margin convex. Mesoscutum reticulate rugose; notauli incomplete, reaching approximately half the length of mesoscutum; mesoscutellum reticulate rugose, hairy and shorter than mesoscutum; anterior margin of mesoscutellum with a foveate narrow groove; metanotum reticulate rugose, shorter than scutellum. Propodeum apparently reticulate rugose and short, distinctly shorter than high in lateral view; fore wing overlapped in holotype and color not visible, but in paratypes, fore wing with one dark transverse band beneath pterostigma. Ocellar ratio: POL= 7; OL= 3; OOL= 10; OPL= 2; TL= 3. Antennomeres in following proportions: 14:7:22:24:34:20:19 (flagellomere 8-10 not visible). Eye somewhat elongate, about 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide; maximum head width about 1.7
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<normalizedToken originalValue="×">x</normalizedToken>
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distance between inner margins of tegulae. Pronotum shorter than head (16:37). Mesoscutum longer than pronotum (20:16). Fore wing with 2r-rs shorter than 3Rs&4Rs. Fore leg segments in following proportions: 30 (coxa): 15 (trochanter): 40 (femur): 43 (tibia): 22 (1th tarsomere): 3 (2nd tarsomere): 7 (3rd tarsomere): 22 (4th tarsomere): 36 (5th tarsomere); 5th tarsomere longer than enlarged claw and with two rows of at least 30 lamellae; distal apex with group of approximately three long lamellae; enlarged claw with two subapical teeth and one row of 29 lamellae. Propodeum short, distinctly shorter than high in lateral view, with dorsal surface shorter than posterior surface (45:50).
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</paragraph>
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<caption ID-Zenodo-Dep="4255457" doi="10.3897/jhr.79.57686.figure2" httpUri="https://binary.pensoft.net/fig/469491" pageId="0" pageNumber="77" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="77">
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<emphasis bold="true" pageId="0" pageNumber="77">Figure 2.</emphasis>
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†
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<taxonomicName authorityName="Olmi" authorityYear="1995" class="Insecta" family="Dryinidae" genus="Thaumatodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thaumatodryinus miocenicus" order="Hymenoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="miocenicus">
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<emphasis italics="true" pageId="0" pageNumber="77">Thaumatodryinus miocenicus</emphasis>
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</taxonomicName>
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Olmi, 1995, female holotype.
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<emphasis bold="true" pageId="0" pageNumber="77">A</emphasis>
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habitus, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="77">B</emphasis>
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habitus, ventral view
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<emphasis bold="true" pageId="0" pageNumber="77">C</emphasis>
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habitus, lateral view
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<emphasis bold="true" pageId="0" pageNumber="77">D</emphasis>
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habitus, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="77">E</emphasis>
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head and mesosoma, ventral view
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<emphasis bold="true" pageId="0" pageNumber="77">F</emphasis>
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habitus, lateral view. Scale bars: 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="77">A-C</emphasis>
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), 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="77">D, F</emphasis>
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), 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="77">E</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="77" type="remarks">
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<paragraph pageId="0" pageNumber="77">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="77">
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While in the original description
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<bibRefCitation author="Olmi, M" journalOrPublisher="Redia" pageId="0" pageNumber="77" pagination="253 - 271" refId="B11" refString="Olmi, M, 1995. Dryinids and embolemids in amber (Hymenoptera Dryinidae et Embolemidae). Redia 78: 253 - 271" title="Dryinids and embolemids in amber (Hymenoptera Dryinidae et Embolemidae)." volume="78" year="1995">Olmi (1995)</bibRefCitation>
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mentioned that the mandibles are "not distinctly visible", Olmi and Virla (2004) stated that the mandibles are
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<normalizedToken originalValue="“tridentate”">"tridentate"</normalizedToken>
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. We could not clearly see the condition in the type material but consider more likely that
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<taxonomicName lsidName="T. miocenicus" pageId="0" pageNumber="77" rank="species" species="miocenicus">
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<emphasis italics="true" pageId="0" pageNumber="77">T. miocenicus</emphasis>
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</taxonomicName>
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has quadridentate mandibles as in all other
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<taxonomicName authorityName="Perkins" authorityYear="1905" class="Insecta" family="Dryinidae" genus="Thaumatodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thaumatodryinus" order="Hymenoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="77">Thaumatodryinus</emphasis>
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</taxonomicName>
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. When the mandibles are closed, it is quite hard to observe the tooth closer to the clypeus, giving the impression that the mandibles are tridentate.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="77" type="materials_examined">
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<paragraph pageId="0" pageNumber="77">Examined material.</paragraph>
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<paragraph pageId="0" pageNumber="77">
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<materialsCitation collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectorName="El Valle" country="the Dominican" location="the Dominican" specimenCount="1" typeStatus="holotype">
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Female
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<typeStatus>holotype</typeStatus>
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, in amber from
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<collectingCountry name="Dominican Republic">the Dominican</collectingCountry>
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Republic: amber from an unknown mine (probably
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<collectorName>El Valle</collectorName>
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mine) DR-99-146 HT (AMNH). Two
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<typeStatus>paratype</typeStatus>
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females: same locality label as
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<typeStatus>holotype</typeStatus>
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, DR-99-147 PT and DR-99-148 PT (AMNH)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |