treatments-xml/data/DA/4A/24/DA4A247D2C9955F28788813AF6E97F17.xml

<document id="BCFF1CCAEF63DD4AD3E5BA72540A315A" ID-DOI="10.3897/contrib.entomol.74.e121614" ID-ZooBank="5AA51752-2447-410B-A249-7B421D573CE4" ID-publisher-id="121614" URI-arpha="763A6B14-4A48-516F-A3A4-FABB0D86C17F" URI-zoobank="http://zoobank.org/5AA51752-2447-410B-A249-7B421D573CE4" article-type="research-article" checkinTime="1722264656219" checkinUser="pensoft" docAuthor="Alexiou, Sotiris, Bakolitsas, Kostas, Di Russo, Claudio &amp; Rampini, Mauro" docDate="2024" docId="DA4A247D2C9955F28788813AF6E97F17" docLanguage="en" docName="Contributions to Entomology 74(1): 103-111" docOrigin="Contributions to Entomology 74 (1)" docSource="https://contributions-to-entomology.arphahub.com/article/121614/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Dolichopoda karoutsosi Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini 2024, sp. nov." docType="treatment" docUuid="B8FB5B44-87C3-492F-82BB-3587D83A304E" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="111" masterDocId="763A6B144A48516FA3A4FABB0D86C17F" masterDocTitle="Four new Dolichopoda species from Greece, one Troglophilus new to Greece and new locality records (Orthoptera, Rhaphidophoridae)" masterLastPageNumber="111" masterPageNumber="103" pageNumber="103" updateTime="1722265077055" updateUser="ExternalLinkService">
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<mods:title id="6460FFF7ECF4D322ACC5993BB8AFAE61">Four new Dolichopoda species from Greece, one Troglophilus new to Greece and new locality records (Orthoptera, Rhaphidophoridae)</mods:title>
</mods:titleInfo>
<mods:name id="C62AF1C55D4D8AA9CF3C09B5E29FE0C0" type="personal">
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<mods:namePart id="843A3D25E4AA9CE8A0C181C94443522F">Alexiou, Sotiris</mods:namePart>
<mods:nameIdentifier id="2B600A22EAD3EBC1A7A9AB925CBF5C7C" type="ORCID">0000-0003-1031-3003</mods:nameIdentifier>
<mods:affiliation id="049E609BFEF00034A0019A8169652DFA">Korinthian Museum of Natural History, Isthmion 201, Korinthos 20100, Greece</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="04A2F9C16DD2DF3CC27EE81EF9E1DBD6">Bakolitsas, Kostas</mods:namePart>
<mods:nameIdentifier id="95347C9AD139EA068AA342F216BAC22F" type="ORCID">0000-0003-3137-4024</mods:nameIdentifier>
<mods:affiliation id="F03504CC97ACA314F3930E95FEF3EBD1">Laboratory of Natural Sciences 3 rd General Lyceum, 301 31 Agrinion, Greece</mods:affiliation>
</mods:name>
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<mods:namePart id="18028049C5B8C7746FEAA0CE453BFBA9">Di Russo, Claudio</mods:namePart>
<mods:nameIdentifier id="8DC63DAA5EA1EA8D05A151D580A86F97" type="ORCID">0000-0002-1620-9592</mods:nameIdentifier>
<mods:affiliation id="13293D15FA76BA6B4849824BA4D06A4A">Dipartimento di Biologia e Biotecnologie “ C. Darwin ”, Università di Roma “ La Sapienza ”, Roma, Italy</mods:affiliation>
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<mods:namePart id="B2B88687256B17EF29DBE97AACB6AF84">Rampini, Mauro</mods:namePart>
<mods:affiliation id="EF15F9F536A70EF36BD88637539AE1A4">Dipartimento di Biologia e Biotecnologie “ C. Darwin ”, Università di Roma “ La Sapienza ”, Roma, Italy</mods:affiliation>
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<mods:title id="C218EC6EB509981029B16F049E185CED">Contributions to Entomology</mods:title>
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<mods:date id="EB8E3CA94B450127510B02946D11992A">2024</mods:date>
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<mods:number id="0771B8FACFC8FFA7AE817ABA291E0414">74</mods:number>
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<subSubSection id="898E6B539779131BC70F0BA25F8BC917" type="nomenclature">
<paragraph id="3BE5D90FE91688E8EA5A81AAD4ECBCAF">
<taxonomicName id="2C9ABD1F374078790768CFB10CB0EEEA" authority="Alexiou" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="karoutsosi" status="sp. nov.">
<emphasis id="908E0FADF17315D0BE52B7EE5FB2EE9B" italics="true">Dolichopoda karoutsosi</emphasis>
Alexiou
</taxonomicName>
<taxonomicNameLabel id="E45977E6CFB82A874C1AC838E30E8FFE" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="1ED8D8F36D10D7097A2C595DE04B88D3" type="description">
<paragraph id="A9A86F1C3993357D9FFC1CC8F80F7D4C">
<figureCitation id="95A0C24E0F2A2B0011883031C64F0507" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographic distribution of D. athosensis sp. nov., D. dirussoi sp. nov., D. karoutsosi sp. nov. and D. kotsabasi sp. nov." figureDoi="10.3897/contrib.entomol.74.e121614.figure1" httpUri="https://binary.pensoft.net/fig/1103848">Figs 1</figureCitation>
, 
<figureCitation id="09D4F5D8730758140C4EA3AEEAA30C83" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Geographic distribution of Dolichopoda species at W Sterea Ellas and S Ionian Islands." figureDoi="10.3897/contrib.entomol.74.e121614.figure4" httpUri="https://binary.pensoft.net/fig/1103851">4</figureCitation>
, 
<figureCitation id="FF801DA002D381EC148B888AD0B5766E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dolichopoda karoutsosi sp. nov., male: A. X tergum, dorsal view; B. Epiphallus, dorsal view; C. Epiphallus, lateral view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure6" httpUri="https://binary.pensoft.net/fig/1103853">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENHAE" type="type material">
<paragraph id="82DFDD2020CFAD06916EFBCD990DC966">
<heading id="66B92AD8830FEF402BCAE2CE556536F0" reason="title">Type material.</heading>
</paragraph>
<paragraph id="95FCFE650F75EBA74CBC2E171D9CE649">
<materialsCitation id="369975D24C4C76ECD4ECBF6EE1D3B6C0" collectingDate="2017-08-07" collectorName="K. Bakolitsas" country="Greece" county="cave Siko" elevation="344" latitude="38.723915" location="Sterea Ellas" longLatPrecision="1" longitude="21.116283" municipality="Agios Nikolaos" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="0556FCEC0D5F404B577CFC6EF366C7B1" bold="true" italics="true">
<typeStatus id="893FF442153798D6A2C30EDB1A8DDBC6" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<collectingCountry id="3B004BA50D78D4968DD68B8432969BD7" name="Greece">GREECE</collectingCountry>
• 
<specimenCount id="3CA40C1D25DAD8298CDE70ABA82C4EFD" count="1" type="male">♂</specimenCount>
; 
<location id="D55B6DAE1A4E9EE2309E863C935324E2" LSID="urn:lsid:plazi:treatment:DA4A247D2C9955F28788813AF6E97F17:D55B6DAE1A4E9EE2309E863C935324E2" country="Greece" county="cave Siko" latitude="38.723915" longLatPrecision="1" longitude="21.116283" municipality="Agios Nikolaos" name="Sterea Ellas">Sterea Ellas</location>
, 
<location id="4529271D92BBA4B83DE48F420C43C5BB" LSID="urn:lsid:plazi:treatment:DA4A247D2C9955F28788813AF6E97F17:4529271D92BBA4B83DE48F420C43C5BB" country="Greece" county="cave Siko" latitude="38.723915" longLatPrecision="1" longitude="21.116283" municipality="Agios Nikolaos" name="Nomos Etolias-Akarnanias">Nomos Etolias-Akarnanias</location>
, 
<location id="64834A829A9AF17ED20307E3333F7FD1" LSID="urn:lsid:plazi:treatment:DA4A247D2C9955F28788813AF6E97F17:64834A829A9AF17ED20307E3333F7FD1" country="Greece" county="cave Siko" latitude="38.723915" longLatPrecision="1" longitude="21.116283" municipality="Agios Nikolaos" name="Aetos">Aetos</location>
, 
<collectingMunicipality id="8789517FEB56149FA68BDB8C71DE8DA4">Agios Nikolaos</collectingMunicipality>
, 
<collectingCounty id="99771C6DA7CC9913F61B483A8D81A049">cave Siko</collectingCounty>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2HAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[21.116283,38.723917]}">
<geoCoordinate id="C9C644DD94CC19FABC5DF9CB41756E13" degrees="38" direction="north" minutes="43.435" orientation="latitude" precision="1" value="38.723915">38 ° 43.435 ' N</geoCoordinate>
, 
<geoCoordinate id="4FE27D71F945F8A6E14C4C60712C32B6" degrees="21" direction="east" minutes="6.977" orientation="longitude" precision="1" value="21.116283">21 ° 6.977 ' E</geoCoordinate>
</named-content>
; 
<elevation id="E274C9F7DEE9682E638BD50C0D3A16A5" metricMagnitude="2" metricUnit="m" metricValue="3.44" unit="m" value="344.0">
<quantity id="AD808AB674E33B15FA6CF24D4BBF608A" metricMagnitude="2" metricUnit="m" metricValue="3.44" unit="m" value="344.0">344 m</quantity>
a. s. l.
</elevation>
; 
<date id="F8ECD91AC11946DBC555C6937AE2869C" value="2017-08-07">
<collectingDate id="99E28897AAB8A241C2E71A40DD79C4CE" value="2017-08-07">7 Aug. 2017</collectingDate>
</date>
; 
<collectorName id="15DD81089F1FBA4025E1FAAB0A1E4AAD">K. Bakolitsas</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0EAIAE" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
. 
<materialsCitation id="1C70BC713693215661D2F7912313407B" collectingDate="2017-08-07" collectorName="K. Bakolitsas" country="Greece" county="cave Siko" elevation="344" latitude="38.723915" location="Sterea Ellas" longLatPrecision="1" longitude="21.116283" municipality="Agios Nikolaos" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="5E44DDEC821E658EEC970CBE0E57B1F1" bold="true" italics="true">
<typeStatus id="6FEAD42ED3C585A52FF5F3FCE3F43D78" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<collectingCountry id="3B19610AA3A394E4E515237EECD2BA04" name="Greece">GREECE</collectingCountry>
• 
<specimenCount id="2F7E491092B338FEC210AAE4ADCD5DCC" count="2" type="male">2 ♂</specimenCount>
, 1 nymph, same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIIAE" type="diagnosis">
<paragraph id="7929F7B8BDBA58BFAB7F0375109727C4">
<heading id="F2F247A79E623D44260DD88132ABBD27" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="19E60C23F57347B1C6F66EB6FDA54206">
The new taxon presents one morphological character unique for the genus: the presence of a notch at the median process of the epiphallus (present at both dissected males). The shape of the lateral lobes of X tergum of the males shows resemblance to 
<taxonomicName id="C6321ADCAB20CBACE363FC912D8E03BE" authorityName="Rampini &amp; Di Russo" authorityYear="2017" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="bakolitsasi">
<emphasis id="86C218635926A1BA17D7AABCDA4D866B" italics="true">D. bakolitsasi</emphasis>
</taxonomicName>
, a species known from a cave a few km away. The elevated ridge at the posterior margin of the concavity of the X tergite and the rather slender median process of epiphallus connects the new species with the species of the south Ionian region, e. g. 
<taxonomicName id="89632443E8116AD7A9C53E052C6781F1" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="garparoi">
<emphasis id="2855861BE773481155D5C2D6FB66B30D" italics="true">D. garparoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="28FFF187A42F9D293BAC3E0C8D1BAEAF" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="ithakii">
<emphasis id="AE292D36795FEFEFA0E2582A7FC1C669" italics="true">D. ithakii</emphasis>
</taxonomicName>
, 
<taxonomicName id="6CB1AA62D6C7C338209D1940AE2F0CC3" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="giachinoi">
<emphasis id="EB6F2DAD611FE5DAC20DE7F7833E8B34" italics="true">D. giachinoi</emphasis>
</taxonomicName>
and 
<taxonomicName id="783C9A02BC00746CAC92B8EE8A4F567C" authorityName="Chopard" authorityYear="1964" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="patrizii">
<emphasis id="EED2CE83640F8A463F48A65B5F27C215" italics="true">D. patrizii</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="062460714ACC276F95FC209A4846385A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Geographic distribution of Dolichopoda species at W Sterea Ellas and S Ionian Islands." figureDoi="10.3897/contrib.entomol.74.e121614.figure4" httpUri="https://binary.pensoft.net/fig/1103851">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJKAE" type="description">
<paragraph id="9B7FC9EB5C7418D6FF164F0B0C6E3DC7">
<heading id="80C77066FF2141F8080D114D125D67B6" reason="title">Description.</heading>
</paragraph>
<paragraph id="535EECA276741B8337383AB79D0B271B">Body pale brownish-yellow, legs elongated, hind femora unarmed. Fore tibiae with 4 spines on both sides of the ventral side, mid tibiae with 5 spines on both sides of the ventral side and hind tibiae with 19 spines on both sides of the dorsal side.</paragraph>
<paragraph id="556CFB0BB3A2B23C067CCCC52699EE7D">
X tergum (last abdominal tergite) (Fig. 
<figureCitation id="127BE2985E4253BF638543BCB2B39D60" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dolichopoda karoutsosi sp. nov., male: A. X tergum, dorsal view; B. Epiphallus, dorsal view; C. Epiphallus, lateral view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure6" httpUri="https://binary.pensoft.net/fig/1103853">6 A</figureCitation>
) with two lateral lobes, shallowly bilobed at apex. Bilobed apex very unequal, the external side elongated, cylindrical, pointing dorso-posteriorly. The two lobes are separated by a large concavity showing an elevated ridge on the posterior margin.
</paragraph>
<caption id="9B5E093D36AFF30F28F7CD5AC6689DBE" ID-DOI="10.3897/contrib.entomol.74.e121614.figure6" ID-arpha="289EE310-3C14-5E94-99DC-2262648604E4" httpUri="https://binary.pensoft.net/fig/1103853" startId="F6">
<paragraph id="3055F48FDB05D1F3110D557C91E59563">
<label id="AB9D84C9FB1D80CAA55682B81815DDA6">Figure 6.</label>
</paragraph>
<paragraph id="7F63E763078A297B48B1A718A7399421">
<taxonomicName id="653D0029885598CFE0597F128B902F7E" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="karoutsosi" status="sp. nov.">
<emphasis id="4499BA87D5419E25AC342F9FAD8E8499" italics="true">Dolichopoda karoutsosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC3666B6ACF3FC68FFBE67A3B81FE768" rank="species">sp. nov.</taxonomicNameLabel>
, male: 
<emphasis id="C71ED4297E88A1D655C667CE1BCCBF06" bold="true">A.</emphasis>
X tergum, dorsal view; 
<emphasis id="B71C829295572A737712F9C496E09FDC" bold="true">B.</emphasis>
Epiphallus, dorsal view; 
<emphasis id="3DDC2E10B81A61D6A2DC931B9D388191" bold="true">C.</emphasis>
Epiphallus, lateral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="D709175F5BD25F7BF2D1456CE1DBC5ED">Subgenital plate convex, with a wide median incision that runs for 1 / 3 of the total length; lateral lobes rounded, styli short, inserted at the side of the lateral lobes.</paragraph>
<paragraph id="2E124FB89F6DC31615F65DD2197BB911">
Epiphallus sclerotized, median process rather slender and acute, the last ¼ curved inwards, an obvious notch is present at the beginning of the curve (Fig. 
<figureCitation id="E95660A046B620CFEC856A554D1F6F40" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dolichopoda karoutsosi sp. nov., male: A. X tergum, dorsal view; B. Epiphallus, dorsal view; C. Epiphallus, lateral view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure6" httpUri="https://binary.pensoft.net/fig/1103853">6 B, C</figureCitation>
).
</paragraph>
<paragraph id="8F93F48FEEC81930FB94B624E912F8CE">Measurements (length in mm): pronotum 3.6, fore femora 17, middle femora 18, hind femora 26, body 16.</paragraph>
<paragraph id="0E44C3601BF79B3964FEE5A34ADE5D18">
<emphasis id="A1564ABFA6B4AA4032BC3A337A0B31FB" bold="true">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5LAE" type="etymology">
<paragraph id="DB71086C5E40D411DF7CA65AB6B18F3E">
<heading id="C89F960FFD9FDB16BDE261AF5F87439E" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="5BCC2DC7EBC46606523B2D575CA00B8B">
The new species is named after our friend, geologist Panos Karoutsos, a dear companion of the second author who visited the 
<typeStatus id="81AEB76D0CD0AEEE4BF9B53C2628AC79">type</typeStatus>
locality and many other caves together.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDMAE" type="distribution">
<paragraph id="FDBD532FD03B5E3394BBFF42F248E3F1">
<heading id="8497B8A4D295DBE434CF1520EF808B47" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="27993E720DDC949B65BCB76761C63F8A">
Known only from the 
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locality. Siko is a pit fall 
<quantity id="EFC11DAEF9D15B9256C33C7BFD562456" metricMagnitude="0" metricUnit="m" metricValue="7.0" unit="m" value="7.0">7 m</quantity>
deep, with a spacious 8 × 
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room at its base, without decoration and with high humidity. The entrance is about 0.5 × 
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, facing east and on the floor there are many bones belonging to animals that have fallen from the cave entrance.
</paragraph>
</subSubSection>
</treatment>
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