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<mods:title id="6DB3E647D441B3EA9814DCD78F7E790F">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="986F4911FB06FF04FAF26BBECA2A889F">Harzhauser, Mathias</mods:namePart>
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<subSubSection id="C327654CFFDDFFF7FFBAFB2D6BE1FB60" box="[151,915,1158,1185]" pageId="95" pageNumber="96" type="nomenclature">
<paragraph id="8B8236C7FFDDFFF7FFBAFB2D6BE1FB60" blockId="95.[151,915,1158,1221]" box="[151,915,1158,1185]" pageId="95" pageNumber="96">
<heading id="D0CA81ABFFDDFFF7FFBAFB2D6BE1FB60" bold="true" box="[151,915,1158,1185]" fontSize="11" level="2" pageId="95" pageNumber="96" reason="4">
<emphasis id="B949EAD5FFDDFFF7FFBAFB2D6BE1FB60" bold="true" box="[151,915,1158,1185]" pageId="95" pageNumber="96">
<emphasis id="B949EAD5FFDDFFF7FFBAFB2D696CFB61" bold="true" box="[151,286,1158,1185]" italics="true" pageId="95" pageNumber="96">Clavatula</emphasis>
<taxonomicName id="4C3D4D44FFDDFFF7FE08FB2D6AD6FB60" authority="(Mayer-Eymar, 1886)" baseAuthorityName="Mayer-Eymar" baseAuthorityYear="1886" box="[293,676,1158,1185]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="rusticula">
<emphasis id="B949EAD5FFDDFFF7FE08FB2D69E3FB61" bold="true" box="[293,401,1158,1185]" italics="true" pageId="95" pageNumber="96">rusticula</emphasis>
(
<bibRefCitation id="EFAC4B36FFDDFFF7FE8DFB2D6AEEFB60" author="Mayer-Eymar, C." box="[416,668,1158,1184]" pageId="95" pageNumber="96" pagination="302 - 312" refId="ref138634" refString="Mayer-Eymar, C. (1886) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 26, 302 - 312, pl. 16. [https: // www. biodiversitylibrary. org / item / 54516 # page / 310 / mode / 1 up]" type="journal article" year="1886">Mayer-Eymar, 1886</bibRefCitation>
)
</taxonomicName>
species inquirenda
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C327654CFFDDFFF7FFBAFB00688FFB05" box="[151,253,1195,1221]" pageId="95" pageNumber="96" type="description">
<paragraph id="8B8236C7FFDDFFF7FFBAFB00688FFB05" blockId="95.[151,915,1158,1221]" box="[151,253,1195,1221]" pageId="95" pageNumber="96">
<figureCitation id="13062A42FFDDFFF7FFBAFB00688FFB05" box="[151,253,1195,1221]" captionStart="FIGURE 29" captionStartId="96.[151,250,1397,1421]" captionTargetBox="[151,1436,236,1373]" captionTargetId="figure-45@96.[151,1436,236,1373]" captionTargetPageId="96" captionText="FIGURE 29 A. Clavatula inedita Bellardi, 1877, Letkés (Hungary), private collection of Zoltán Kovács and Zoltán Vicián. B. Clavatula rusticula (Mayer-Eymar, 1886), lectotype, S 1615, Grund (Austria). C. Clavatula orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). D1D2. Clavatula orientoromana Báldi, 1960, paratype, M60.146, Szokolya (Hungary). E1E2. Clavatula justinae (Hoernes &amp; Auinger, 1891), paralectotypes, NHMW 1849/0005/0110b, Buituri (Romania).F1F3. Clavatula justinae (Hoernes &amp;Auinger, 1891), paralectotype, NHMW 1949/0005/0110a, Buituri (Romania). G. Clavatula justinae (Hoernes &amp; Auinger, 1891), lectotype, NHMW 1849/0005/0110c, Buituri (Romania)." figureDoi="http://doi.org/10.5281/zenodo.6399893" httpUri="https://zenodo.org/record/6399893/files/figure.png" pageId="95" pageNumber="96">Figs 29B</figureCitation>
</paragraph>
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<heading id="D0CA81ABFFDDFFF7FFBAFB586CE3FACD" box="[151,1169,1267,1293]" fontSize="11" level="2" pageId="95" pageNumber="96" reason="5">
<treatmentCitationGroup id="AB2D11E9FFDDFFF7FFBAFB586CE3FACD" box="[151,1169,1267,1293]" pageId="95" pageNumber="96">
<emphasis id="B949EAD5FFDDFFF7FFBAFB5868D7FACD" box="[151,165,1267,1293]" italics="true" pageId="95" pageNumber="96">*</emphasis>
<taxonomicName id="4C3D4D44FFDDFFF7FF81FB586AB4FACD" authority="Mayer-Eymar" authorityName="Mayer-Eymar" authorityYear="1886" box="[172,710,1267,1293]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="rusticula" subGenus="Clavatula">
<emphasis id="B949EAD5FFDDFFF7FF81FB58695DFACD" box="[172,303,1267,1293]" italics="true" pageId="95" pageNumber="96">Pleurotoma</emphasis>
(
<emphasis id="B949EAD5FFDDFFF7FE6DFB5869DCFACD" box="[320,430,1267,1293]" italics="true" pageId="95" pageNumber="96">Clavatula</emphasis>
)
<emphasis id="B949EAD5FFDDFFF7FE93FB586A50FACD" box="[446,546,1267,1293]" italics="true" pageId="95" pageNumber="96">rusticula</emphasis>
Mayer-Eymar
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<treatmentCitation id="0A9C10D6FFDDFFF7FDCEFB586B8FFACD" author="Mayer-Eymar, C." box="[739,1021,1267,1293]" page="311" pageId="95" pageNumber="96" year="1886">
<bibRefCitation id="EFAC4B36FFDDFFF7FDCEFB586B8FFACD" author="Mayer-Eymar, C." box="[739,1021,1267,1293]" pageId="95" pageNumber="96" pagination="302 - 312" refId="ref138634" refString="Mayer-Eymar, C. (1886) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 26, 302 - 312, pl. 16. [https: // www. biodiversitylibrary. org / item / 54516 # page / 310 / mode / 1 up]" type="journal article" year="1886">Mayer-Eymar, 1886: 311</bibRefCitation>
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, pl. 16, fig. 8
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<subSubSection id="C327654CFFDDFFF7FFBAFA90697EF959" pageId="95" pageNumber="96" type="materials_examined">
<paragraph id="8B8236C7FFDDFFF7FFBAFA906D77FA5D" blockId="95.[151,1437,1338,2013]" pageId="95" pageNumber="96">
<emphasis id="B949EAD5FFDDFFF7FFBAFA906935FA94" bold="true" box="[151,327,1338,1365]" pageId="95" pageNumber="96">Type material.</emphasis>
<typeStatus id="54868865FFDDFFF7FE62FA9069CDFA95" box="[335,447,1339,1365]" pageId="95" pageNumber="96" type="lectotype">Lectotype</typeStatus>
designated herein: S 1615, SL.
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,
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: 5.0 mm, Grund (
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), stored in the collection of the
<collectionCode id="ED2CAE02FFDDFFF7FE76FAF469E3FAB9" box="[347,401,1375,1401]" country="Switzerland" httpUri="http://grbio.org/cool/h54f-yj1d" name="Kultursammlungen der Eidgenosische Technische Hochschule" pageId="95" pageNumber="96">ETH</collectionCode>
<collectingRegion id="49F9F825FFDDFFF7FEB6FAF46994FAB9" box="[411,486,1375,1401]" country="Switzerland" name="Zuerich" pageId="95" pageNumber="96">Zürich</collectingRegion>
(
<collectingCountry id="F32A7657FFDDFFF7FEDAFAF46AF2FABA" box="[503,640,1375,1402]" name="Switzerland" pageId="95" pageNumber="96">Switzerland</collectingCountry>
) (
<figureCitation id="13062A42FFDDFFF7FDB4FAF46B76FAB9" box="[665,772,1375,1401]" captionStart="FIGURE 29" captionStartId="96.[151,250,1397,1421]" captionTargetBox="[151,1436,236,1373]" captionTargetId="figure-45@96.[151,1436,236,1373]" captionTargetPageId="96" captionText="FIGURE 29 A. Clavatula inedita Bellardi, 1877, Letkés (Hungary), private collection of Zoltán Kovács and Zoltán Vicián. B. Clavatula rusticula (Mayer-Eymar, 1886), lectotype, S 1615, Grund (Austria). C. Clavatula orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). D1D2. Clavatula orientoromana Báldi, 1960, paratype, M60.146, Szokolya (Hungary). E1E2. Clavatula justinae (Hoernes &amp; Auinger, 1891), paralectotypes, NHMW 1849/0005/0110b, Buituri (Romania).F1F3. Clavatula justinae (Hoernes &amp;Auinger, 1891), paralectotype, NHMW 1949/0005/0110a, Buituri (Romania). G. Clavatula justinae (Hoernes &amp; Auinger, 1891), lectotype, NHMW 1849/0005/0110c, Buituri (Romania)." figureDoi="http://doi.org/10.5281/zenodo.6399893" httpUri="https://zenodo.org/record/6399893/files/figure.png" pageId="95" pageNumber="96">Fig. 29A</figureCitation>
);
<bibRefCitation id="EFAC4B36FFDDFFF7FC36FAF46C79FABA" author="Mayer-Eymar, C." box="[795,1035,1375,1402]" pageId="95" pageNumber="96" pagination="302 - 312" refId="ref138634" refString="Mayer-Eymar, C. (1886) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 26, 302 - 312, pl. 16. [https: // www. biodiversitylibrary. org / item / 54516 # page / 310 / mode / 1 up]" type="journal article" year="1886">Mayer-Eymar (1886)</bibRefCitation>
mentioned
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, but the specimen illustrated in
<bibRefCitation id="EFAC4B36FFDDFFF7FEB1FA286AF1FA5D" author="Mayer-Eymar, C." box="[412,643,1411,1437]" pageId="95" pageNumber="96" pagination="302 - 312" refId="ref138634" refString="Mayer-Eymar, C. (1886) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 26, 302 - 312, pl. 16. [https: // www. biodiversitylibrary. org / item / 54516 # page / 310 / mode / 1 up]" type="journal article" year="1886">Mayer-Eymar (1886</bibRefCitation>
, pl. 16, fig. 8, SL.
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,
<collectionCode id="ED2CAE02FFDDFFF7FC9CFA286B90FA5D" box="[945,994,1411,1437]" country="Germany" name="Museum Donaueschingen" pageId="95" pageNumber="96">MD</collectionCode>
:
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) seems to be lost.
</paragraph>
<paragraph id="8B8236C7FFDDFFF7FFEAFA0D697EF959" blockId="95.[151,1437,1338,2013]" pageId="95" pageNumber="96">
<emphasis id="B949EAD5FFDDFFF7FFEAFA0D69CBFA00" bold="true" box="[199,441,1446,1472]" pageId="95" pageNumber="96">Revised description.</emphasis>
Small to medium-sized, slender fusiform with high spire; apical angle ~28°. Protoconch and early teleoconch not preserved. Spire whorls with faint concavity. Subsutural spiral cord bearing rounded beads in some distance to narrowly incised, slightly wavy suture. Broad spiral cords below subsutural collar. Last whorl short, conical above strongly protruding shoulder with large nodes. Two beaded peribasal spiral cords. Base strongly constricted with weak spiral cords. Fasciole indistinct. Outer lip thin, smooth within. Siphonal canal moderately long, deflected to the left. Columella slightly twisted, smooth. Columellar and parietal callus thickened forming broad rim.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFDDFFF7FFEAF90969F1F81D" pageId="95" pageNumber="96" type="discussion">
<paragraph id="8B8236C7FFDDFFF7FFEAF9096AB7F8B1" blockId="95.[151,1437,1338,2013]" pageId="95" pageNumber="96">
<emphasis id="B949EAD5FFDDFFF7FFEAF909693EF97C" bold="true" box="[199,332,1698,1724]" pageId="95" pageNumber="96">Discussion.</emphasis>
This species is based on poorly preserved fragments. It is unique within the Paratethyan fauna due to its small beads on the subsutural collar, which appear at some distance from the suture, and due to its strongly angulated and protruding shoulder
<bibRefCitation id="EFAC4B36FFDDFFF7FD11F9406B43F8C6" author="Mayer-Eymar, C." box="[572,817,1771,1798]" pageId="95" pageNumber="96" pagination="302 - 312" refId="ref138634" refString="Mayer-Eymar, C. (1886) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 26, 302 - 312, pl. 16. [https: // www. biodiversitylibrary. org / item / 54516 # page / 310 / mode / 1 up]" type="journal article" year="1886">Mayer-Eymar (1886)</bibRefCitation>
discussed similarities with
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<emphasis id="B949EAD5FFDDFFF7FBA0F9406C89F8C5" box="[1165,1275,1771,1797]" italics="true" pageId="95" pageNumber="96">Clavatula</emphasis>
</taxonomicName>
<taxonomicName id="4C3D4D44FFDDFFF7FA3EF9406916F8E9" authority="(Grateloup, 1832)" baseAuthorityName="Grateloup" baseAuthorityYear="1832" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="concatenata">
<emphasis id="B949EAD5FFDDFFF7FA3EF9406DEEF8C4" box="[1299,1436,1771,1796]" italics="true" pageId="95" pageNumber="96">concatenata</emphasis>
(
<bibRefCitation id="EFAC4B36FFDDFFF7FFB2F8A4692EF8E9" author="Grateloup, J. P. S. de" box="[159,348,1807,1833]" pageId="95" pageNumber="96" pagination="314 - 344" refId="ref133310" refString="Grateloup, J. P. S. de (1832) Tableau des coquilles Fossiles qu'on rencontre dans les terrains calcaires tertiaries (faluns) des environs de Dax, dans le departement des Landes. Actes de la Societe Linneenne de Bordeaux, 5, 314 - 344. [https: // www. biodiversitylibrary. org / item / 102890 # page / 325 / mode / 1 up]" type="journal article" year="1832">Grateloup, 1832</bibRefCitation>
)
</taxonomicName>
, from the Chattian and Aquitanian of
<collectingCountry id="F32A7657FFDDFFF7FC3BF8A46B17F8E9" box="[790,869,1807,1833]" name="France" pageId="95" pageNumber="96">France</collectingCountry>
, but the French species has little in common with
<emphasis id="B949EAD5FFDDFFF7FF8CF89868C8F88D" box="[161,186,1843,1869]" italics="true" pageId="95" pageNumber="96">C.</emphasis>
<taxonomicName id="4C3D4D44FFDDFFF7FFE6F898695DF88D" authorityName="Mayer-Eymar" authorityYear="1886" box="[203,303,1843,1869]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="rusticula">
<emphasis id="B949EAD5FFDDFFF7FFE6F898695DF88D" box="[203,303,1843,1869]" italics="true" pageId="95" pageNumber="96">rusticula</emphasis>
</taxonomicName>
and has a smooth subsutural cord, and beads only above the abapical suture (see
<bibRefCitation id="EFAC4B36FFDDFFF7FBE6F8986D24F88D" author="Peyrot, A." box="[1227,1366,1843,1869]" pageId="95" pageNumber="96" pagination="5 - 288" refId="ref141284" refString="Peyrot, A. (1931 &amp; 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]" type="journal article" year="1931">Peyrot 1931</bibRefCitation>
, pl. 8, figs 63, 84, 85;
<bibRefCitation id="EFAC4B36FFDDFFF7FE6AF8FC6997F8B1" author="Lozouet, P." box="[327,485,1879,1905]" pageId="95" pageNumber="96" pagination="3 - 179" refId="ref137939" refString="Lozouet, P. (2017) Les Conoidea de l'Oligocene superieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France). Cossmanniana, 19, 3 - 179." type="journal article" year="2017">Lozouet 2017</bibRefCitation>
, pl. 33, figs 1820).
</paragraph>
<paragraph id="8B8236C7FFDDFFF7FFEAF8D169F1F81D" blockId="95.[151,1437,1338,2013]" pageId="95" pageNumber="96">
<emphasis id="B949EAD5FFDDFFF7FFEAF8D169DAF854" bold="true" box="[199,424,1914,1940]" pageId="95" pageNumber="96">Paleoenvironment.</emphasis>
At the locality Grund fossiliferous channel fills, which formed in middle to outer neritic environments bear allochthonous assemblages uniting coastal-mudflat faunas with inner neritic ones (
<bibRefCitation id="EFAC4B36FFDDFFF7FA2FF83468A0F81D" author="Zuschin, M. &amp; Harzhauser, M. &amp; Mandic, O." pageId="95" pageNumber="96" pagination="142 - 158" refId="ref146832" refString="Zuschin, M., Harzhauser, M. &amp; Mandic, O. (2005) Influence of Size-sorting on diversity estimates from tempestitic shell beds in the middle Miocene of Austria. Palaios, 20, 142 - 158. https: // doi. org / 10.2110 / palo. 2003. p 03 - 87" type="journal article" year="2005">
Zuschin
<emphasis id="B949EAD5FFDDFFF7FA48F8346DE7F879" box="[1381,1429,1951,1977]" italics="true" pageId="95" pageNumber="96">et al</emphasis>
. 2005
</bibRefCitation>
;
<bibRefCitation id="EFAC4B36FFDDFFF7FFF3F868690AF81D" author="Roetzel, R." box="[222,376,1987,2013]" pageId="95" pageNumber="96" refId="ref142559" refString="Roetzel, R. (2009) Erlauterungen zu Blatt 23 Hadres. Geologische Karte der Republik Osterreich 1: 50000. Geologische Bundesanstalt, Wien, 150 pp." type="book" year="2009">Roetzel 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFE2FFC8FFEAFF3D6A35FF15" pageId="96" pageNumber="97" type="distribution">
<paragraph id="8B8236C7FFE2FFC8FFEAFF3D6DEEFF71" blockId="96.[151,1436,150,213]" box="[199,1436,150,177]" pageId="96" pageNumber="97">
<emphasis id="B949EAD5FFE2FFC8FFEAFF3D6A1FFF70" bold="true" box="[199,621,150,176]" pageId="96" pageNumber="97">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="B949EAD5FFE2FFC8FCECFF3D6DE7FF70" box="[961,1429,150,177]" italics="true" pageId="96" pageNumber="97">North Alpine -Carpathian Foreland Basin</emphasis>
:
</paragraph>
<paragraph id="8B8236C7FFE2FFC8FFBAFF116A35FF15" blockId="96.[151,1436,150,213]" box="[151,583,186,213]" pageId="96" pageNumber="97">
Grund (
<collectingCountry id="F32A7657FFE2FFC8FFC0FF116931FF15" box="[237,323,186,213]" name="Austria" pageId="96" pageNumber="97">Austria</collectingCountry>
) (
<bibRefCitation id="EFAC4B36FFE2FFC8FE74FF116A49FF15" author="Mayer-Eymar, C." box="[345,571,186,213]" pageId="96" pageNumber="97" pagination="302 - 312" refId="ref138634" refString="Mayer-Eymar, C. (1886) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 26, 302 - 312, pl. 16. [https: // www. biodiversitylibrary. org / item / 54516 # page / 310 / mode / 1 up]" type="journal article" year="1886">Mayer-Eymar 1886</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>