The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera Author Harzhauser, Mathias 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at Author Landau, Bernard 0000-0002-7768-8494 bernardmlandau@gmail.com Author Janssen, Ronald 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at text Zootaxa 2022 2022-03-30 5123 1 1 172 http://dx.doi.org/10.11646/zootaxa.5123.1.1 journal article 10.11646/zootaxa.5123.1.1 1175-5326 6399827 036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A ‘Clavatula’ rusticula ( Mayer-Eymar, 1886 ) species inquirenda Figs 29B * Pleurotoma ( Clavatula ) rusticula Mayer-Eymar — Mayer-Eymar, 1886: 311 , pl. 16, fig. 8 Type material. Lectotype designated herein: S 1615, SL. 12.5 mm , MD : 5.0 mm, Grund ( Austria ), stored in the collection of the ETH Zürich ( Switzerland ) ( Fig. 29A ); Mayer-Eymar (1886) mentioned two specimens , but the specimen illustrated in Mayer-Eymar (1886 , pl. 16, fig. 8, SL. 24 mm , MD : 10 mm ) seems to be lost. Revised description. Small to medium-sized, slender fusiform with high spire; apical angle ~28°. Protoconch and early teleoconch not preserved. Spire whorls with faint concavity. Subsutural spiral cord bearing rounded beads in some distance to narrowly incised, slightly wavy suture. Broad spiral cords below subsutural collar. Last whorl short, conical above strongly protruding shoulder with large nodes. Two beaded peribasal spiral cords. Base strongly constricted with weak spiral cords. Fasciole indistinct. Outer lip thin, smooth within. Siphonal canal moderately long, deflected to the left. Columella slightly twisted, smooth. Columellar and parietal callus thickened forming broad rim. Discussion. This species is based on poorly preserved fragments. It is unique within the Paratethyan fauna due to its small beads on the subsutural collar, which appear at some distance from the suture, and due to its strongly angulated and protruding shoulder Mayer-Eymar (1886) discussed similarities with ‘ Clavatula ’ concatenata ( Grateloup, 1832 ) , from the Chattian and Aquitanian of France , but the French species has little in common with ‘ C. ’ rusticula and has a smooth subsutural cord, and beads only above the abapical suture (see Peyrot 1931 , pl. 8, figs 63, 84, 85; Lozouet 2017 , pl. 33, figs 18–20). Paleoenvironment. At the locality Grund fossiliferous channel fills, which formed in middle to outer neritic environments bear allochthonous assemblages uniting coastal-mudflat faunas with inner neritic ones ( Zuschin et al . 2005 ; Roetzel 2009 ). Distribution in Central Paratethys. Badenian (middle Miocene): North Alpine -Carpathian Foreland Basin : Grund ( Austria ) ( Mayer-Eymar 1886 ).