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<document id="25C99DF052C4327E39553A09C4C93303" ID-DOI="10.5281/zenodo.5391440" ID-ISSN="1638-9387" ID-Zenodo-Dep="5391440" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630315127817" checkinUser="marcus" docAuthor="Monniot, Françoise &amp; Monniot, Claude" docDate="2001" docId="F57D87A3FF9C317EE873FB01FF781443" docLanguage="en" docName="Zoosystema.2001.23.2.201-383.pdf" docOrigin="Zoosystema 23 (2)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Didemnum madeleinae Monniot &amp; Monniot 2001, n. sp." docType="treatment" docVersion="3" lastPageNumber="270" masterDocId="0944FFDBFFDF313BEB47FF93FFBE1602" masterDocTitle="Ascidians from the tropical western Pacific" masterLastPageNumber="383" masterPageNumber="201" pageNumber="268" updateTime="1699223418854" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="ABFF5299F0894B251B34593A1A66DB3F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4BE119824BDE3CF013DE217CCC425F04">
<mods:title id="35DA68F65EFB27A4B9F5BC78B48CFF36">Ascidians from the tropical western Pacific</mods:title>
</mods:titleInfo>
<mods:name id="761938B9DC83209E342A15AB99EB1E32" type="personal">
<mods:role id="BE9AC19EB073538D36BC850BBA203E24">
<mods:roleTerm id="69198328EA8C56B463C65A65A1868F90">Author</mods:roleTerm>
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<mods:namePart id="BEFA1EAA91D0FC86C1E71E0AFD95F3E3">Monniot, Françoise</mods:namePart>
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<mods:namePart id="3A8F516C3453D684042E85C70A7CE889">Monniot, Claude</mods:namePart>
<mods:affiliation id="500BFEFA3BC35143593AAE94C69E5182">UPESA 8044, Laboratoire de Biologie des Invertébrés marins et Malacologie, Muséum national d’Histoire naturelle, 55 rue Buffon, F- 75005 Paris (France) monniot @ mnhn. fr.</mods:affiliation>
<mods:nameIdentifier id="E19CD7570B1AA9B7ACB700AC0B88C9EB" type="email">monniot@mnhn.fr</mods:nameIdentifier>
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<mods:typeOfResource id="353AC46DAA693D25AD7C643B90B5A501">text</mods:typeOfResource>
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<mods:titleInfo id="FB5877B9D7F745E8F804EBECB030606E">
<mods:title id="A2407945559F4829AD982AB945C9413E">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="4BD8733133CE719107B5D8B10D77F840">
<mods:date id="2AC00E7ADCE5A4197621984133CBB5F3">2001</mods:date>
<mods:detail id="8D27E4FB809076B2022B16CCBB84E3B7" type="volume">
<mods:number id="8F268B383194AE1844DBD38975E942CF">23</mods:number>
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<mods:number id="96F2AA82D546B5948E30D68FFAB4997D">2</mods:number>
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<mods:start id="8208D0D175FAA7CFCBFFE9DBF8C02B49">201</mods:start>
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<mods:classification id="46389D77A292BEF836137AD0505235CA">journal article</mods:classification>
<mods:identifier id="3268E7F2C1875DE28B5D97F2DE5EA155" type="DOI">10.5281/zenodo.5391440</mods:identifier>
<mods:identifier id="8760E50ECDFC6BE7CA736743F3029071" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="AD5E510206699DA929661F89EA6E046D" type="Zenodo-Dep">5391440</mods:identifier>
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<treatment id="F57D87A3FF9C317EE873FB01FF781443" LSID="urn:lsid:plazi:treatment:F57D87A3FF9C317EE873FB01FF781443" httpUri="http://treatment.plazi.org/id/F57D87A3FF9C317EE873FB01FF781443" lastPageId="69" lastPageNumber="270" pageId="67" pageNumber="268">
<subSubSection id="35CE653EFF9C3178E873FB01FBCB12A9" box="[820,1141,1169,1196]" pageId="67" pageNumber="268" type="nomenclature">
<paragraph id="7D6B36B5FF9C3178E873FB01FBCB12A9" blockId="67.[820,1141,1169,1227]" box="[820,1141,1169,1196]" pageId="67" pageNumber="268">
<heading id="262381D9FF9C3178E873FB01FBCB12A9" box="[820,1141,1169,1196]" centered="true" fontSize="11" level="2" pageId="67" pageNumber="268" reason="2">
<taxonomicName id="BAD44D36FF9C3178E873FB01FB8D12AE" ID-CoL="35WR9" authority="Monniot &amp; Monniot, 2001" authorityName="Monniot &amp; Monniot" authorityYear="2001" box="[820,1075,1170,1196]" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Aplousobranchia" pageId="67" pageNumber="268" phylum="Chordata" rank="species" species="madeleinae" status="sp. nov.">
<emphasis id="4FA0EAA7FF9C3178E873FB01FB8D12AE" bold="true" box="[820,1075,1170,1196]" italics="true" pageId="67" pageNumber="268">Didemnum madeleinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="549357DCFF9C3178EF7DFB02FBCB12A9" box="[1082,1141,1169,1195]" pageId="67" pageNumber="268" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="35CE653EFF9C3178E818FB22FBF712C9" box="[863,1097,1201,1227]" pageId="67" pageNumber="268" type="description">
<paragraph id="7D6B36B5FF9C3178E818FB22FBF712C9" blockId="67.[820,1141,1169,1227]" box="[863,1097,1201,1227]" pageId="67" pageNumber="268">
(
<figureCitation id="E5EF2A30FF9C3178E820FB22FC0712C9" box="[871,953,1201,1227]" captionStart="FIG" captionStartId="67.[154,165,978,995]" captionTargetBox="[154,687,166,940]" captionTargetId="figure-439@67.[142,705,143,966]" captionTargetPageId="67" captionText="FIG. 53. — Didemnum madeleinae n. sp.; A, thorax; B, abdomen; C, larva. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391547" httpUri="https://zenodo.org/record/5391547/files/figure.png" pageId="67" pageNumber="268">Figs 53</figureCitation>
;
<figureCitation id="E5EF2A30FF9C3178E882FB22FC4B12C9" box="[965,1013,1201,1227]" captionStart="FIG" captionStartId="68.[268,279,979,996]" captionTargetBox="[151,1247,166,942]" captionTargetId="figure-306@68.[151,700,166,942]" captionTargetPageId="68" captionText="FIG. 54. — Spicules; A, Didemnum madeleinae n. sp.; B, Polysyncraton adelon n. sp. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5391549" httpUri="https://zenodo.org/record/5391549/files/figure.png" pageId="67" pageNumber="268">54A</figureCitation>
;
<figureCitation id="E5EF2A30FF9C3178EF46FB22FBFF12C9" box="[1025,1089,1201,1227]" captionStart="FIG" captionStartId="161.[151,162,1565,1582]" captionTargetBox="[162,1240,227,1527]" captionTargetId="figure-68@161.[162,1261,227,1550]" captionTargetPageId="161" captionText="FIG. 120. — A, Didemnum algasedens n. sp.; B, Didemnum cuculliferum (Sluiter, 1909); C, Didemnum lacertosum Monniot F., 1995; D, Didemnum madeleinae n. sp.; E, Didemnum obscurum Monniot F., 1969; F, Didemnum rodriguesi Rocha &amp; Monniot F., 1993." figureDoi="http://doi.org/10.5281/zenodo.5391669" httpUri="https://zenodo.org/record/5391669/files/figure.png" pageId="67" pageNumber="268">120D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="35CE653EFF9C3178E98DFB7AFC6C134C" pageId="67" pageNumber="268" type="materials_examined">
<paragraph id="7D6B36B5FF9C3178E98DFB7AFC6C134C" blockId="67.[714,1247,1257,1424]" pageId="67" pageNumber="268">
<collectionCode id="1BC5AE70FF9C3178E98DFB7AFCD712FD" box="[714,873,1257,1280]" pageId="67" pageNumber="268">
<typeStatus id="A26F8817FF9C3178E98DFB7AFD4312FC" box="[714,765,1257,1280]" pageId="67" pageNumber="268">TYPE</typeStatus>
MATERIAL
</collectionCode>
. —
<emphasis id="4FA0EAA7FF9C3178E8DCFB7AFBCC1300" bold="true" box="[923,1138,1257,1282]" pageId="67" pageNumber="268">
<collectingCountry id="05C37625FF9C3178E8DCFB7AFBD11300" box="[923,1135,1257,1282]" name="Papua New Guinea" pageId="67" pageNumber="268">Papua New Guinea</collectingCountry>
.
</emphasis>
Louisiade Archipelago, Deboyne Lagoon, Nivani Island, overhang,
<geoCoordinate id="18E05072FF9C3178E84BFA8EFC3B1337" box="[780,901,1309,1333]" degrees="10" direction="south" minutes="47.46" orientation="latitude" pageId="67" pageNumber="268" precision="9" value="-10.791">10°47.46’S</geoCoordinate>
,
<geoCoordinate id="18E05072FF9C3178E8D5FA8EFBA71337" box="[914,1049,1309,1333]" degrees="152" direction="east" minutes="23.08" orientation="longitude" pageId="67" pageNumber="268" precision="9" value="152.38467">152°23.08’E</geoCoordinate>
,
<quantity id="BA2C9B50FF9C3178EF60FA8EFBDF1331" box="[1063,1121,1308,1333]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="67" pageNumber="268" unit="m" value="10.0">10 m</quantity>
,
<date id="096A1075FF9C3178EF29FA8EFB611337" box="[1134,1247,1309,1333]" pageId="67" pageNumber="268" value="1998-05-30">
30.
<collectionCode id="1BC5AE70FF9C3178EFC9FA8EFB1D1337" box="[1166,1187,1309,1333]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="67" pageNumber="268" type="Museum">V</collectionCode>
.1998
</date>
(
<collectionCode id="1BC5AE70FF9C3178E996FAA5FC9B134C" box="[721,805,1334,1358]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="67" pageNumber="268">MNHN</collectionCode>
A2
<collectionCode id="1BC5AE70FF9C3178E809FAA5FC3F134C" box="[846,897,1334,1358]" pageId="67" pageNumber="268">DID</collectionCode>
.C 470).
</paragraph>
</subSubSection>
<subSubSection id="35CE653EFF9C3178E98DFACDFBF7138C" pageId="67" pageNumber="268" type="etymology">
<paragraph id="7D6B36B5FF9C3178E98DFACDFBF7138C" blockId="67.[714,1247,1257,1424]" pageId="67" pageNumber="268">ETYMOLOGY. — Named after Madeleine Martin, who helped in the preparation of this paper.</paragraph>
</subSubSection>
<subSubSection id="35CE653EFF9C317FE98DFA54FCDE13A3" lastPageId="68" lastPageNumber="269" pageId="67" pageNumber="268" type="description">
<paragraph id="7D6B36B5FF9C3178E98DFA54FCE513DD" blockId="67.[714,1247,1479,1666]" box="[714,859,1479,1505]" pageId="67" pageNumber="268">DESCRIPTION</paragraph>
<paragraph id="7D6B36B5FF9C317FE98DFA7BFF6413C0" blockId="67.[714,1247,1479,1666]" lastBlockId="68.[151,685,1064,1666]" lastPageId="68" lastPageNumber="269" pageId="67" pageNumber="268">
The orange colonies (
<figureCitation id="E5EF2A30FF9C3178E897FA7BFBEE1000" box="[976,1104,1512,1538]" captionStart="FIG" captionStartId="161.[151,162,1565,1582]" captionTargetBox="[162,1240,227,1527]" captionTargetId="figure-68@161.[162,1261,227,1550]" captionTargetPageId="161" captionText="FIG. 120. — A, Didemnum algasedens n. sp.; B, Didemnum cuculliferum (Sluiter, 1909); C, Didemnum lacertosum Monniot F., 1995; D, Didemnum madeleinae n. sp.; E, Didemnum obscurum Monniot F., 1969; F, Didemnum rodriguesi Rocha &amp; Monniot F., 1993." figureDoi="http://doi.org/10.5281/zenodo.5391669" httpUri="https://zenodo.org/record/5391669/files/figure.png" pageId="67" pageNumber="268">Fig. 120D</figureCitation>
) form large crusts
<quantity id="BA2C9B50FF9C3178E856F99BFCEA1023" box="[785,852,1543,1570]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="268" unit="mm" value="2.0">2 mm</quantity>
thick that are easy to remove from the substrate. The surface is smooth with many small common cloacal apertures, which appear white in life: the orange pigment is superficial and disappears in formalin. The consistency is hard, as spicules are dense throughout the colony. The cloacal channels are wide but limited to the thoracic layer of the colony. Some accumulations of orange inclusions lie in the basal layer of the colony. The oral siphon with six sharp lobes, has the shape of a funnel with a basal sphincter (
<figureCitation id="E5EF2A30FF9B317FEBD8FA9BFEBD1320" box="[159,259,1288,1314]" captionStart="FIG" captionStartId="67.[154,165,978,995]" captionTargetBox="[154,687,166,940]" captionTargetId="figure-439@67.[142,705,143,966]" captionTargetPageId="67" captionText="FIG. 53. — Didemnum madeleinae n. sp.; A, thorax; B, abdomen; C, larva. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391547" httpUri="https://zenodo.org/record/5391547/files/figure.png" pageId="68" pageNumber="269">Fig. 53A</figureCitation>
). There is no cloacal languet. The cloacal aperture is wide and uncovers most of the four stigmata rows (
<figureCitation id="E5EF2A30FF9B317FEA06FADBFE171360" box="[321,425,1352,1378]" captionStart="FIG" captionStartId="67.[154,165,978,995]" captionTargetBox="[154,687,166,940]" captionTargetId="figure-439@67.[142,705,143,966]" captionTargetPageId="67" captionText="FIG. 53. — Didemnum madeleinae n. sp.; A, thorax; B, abdomen; C, larva. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391547" httpUri="https://zenodo.org/record/5391547/files/figure.png" pageId="68" pageNumber="269">Fig. 53A</figureCitation>
). The round and small lateral thoracic organs are near the edge of the cloacal aperture at the level of the third transverse vessel.
</paragraph>
<caption id="29AB663DFF9B317FEA4BFC40FBD215E6" ID-DOI="http://doi.org/10.5281/zenodo.5391549" ID-Zenodo-Dep="5391549" box="[268,1132,979,996]" httpUri="https://zenodo.org/record/5391549/files/figure.png" pageId="68" pageNumber="269" startId="68.[268,279,979,996]" targetBox="[151,1247,166,942]" targetPageId="68">
<paragraph id="7D6B36B5FF9B317FEA4BFC40FBD215E6" blockId="68.[268,1132,979,996]" box="[268,1132,979,996]" pageId="68" pageNumber="269">
FIG. 54. — Spicules;
<emphasis id="4FA0EAA7FF9B317FEAFFFC40FE7B15E6" bold="true" box="[440,453,979,996]" pageId="68" pageNumber="269">A</emphasis>
,
<taxonomicName id="BAD44D36FF9B317FEA88FC40FD3015E6" authority="Monniot &amp; Monniot, 2001" authorityName="Monniot &amp; Monniot" authorityYear="2001" box="[463,654,979,996]" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Aplousobranchia" pageId="68" pageNumber="269" phylum="Chordata" rank="species" species="madeleinae" status="sp. nov.">
<emphasis id="4FA0EAA7FF9B317FEA88FC40FD3015E6" box="[463,654,979,996]" italics="true" pageId="68" pageNumber="269">Didemnum madeleinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="549357DCFF9B317FE9D4FC40FD7B15E6" box="[659,709,979,996]" pageId="68" pageNumber="269" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="4FA0EAA7FF9B317FE98BFC40FD6715E6" bold="true" box="[716,729,979,996]" pageId="68" pageNumber="269">B</emphasis>
,
<taxonomicName id="BAD44D36FF9B317FE9A4FC40FC2815E6" authority="Monniot &amp; Monniot, 2001" authorityName="Monniot &amp; Monniot" authorityYear="2001" box="[739,918,979,996]" class="Ascidiacea" family="Didemnidae" genus="Polysyncraton" kingdom="Animalia" order="Aplousobranchia" pageId="68" pageNumber="269" phylum="Chordata" rank="species" species="adelon" status="sp. nov.">
<emphasis id="4FA0EAA7FF9B317FE9A4FC40FC2815E6" box="[739,918,979,996]" italics="true" pageId="68" pageNumber="269">Polysyncraton adelon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="549357DCFF9B317FE8DCFC40FC7515E6" box="[923,971,979,996]" pageId="68" pageNumber="269" rank="species">n. sp.</taxonomicNameLabel>
Scale bars: 10 µm.
</paragraph>
</caption>
<paragraph id="7D6B36B5FF9B317FEBD0FA5BFEC41020" blockId="68.[151,685,1064,1666]" pageId="68" pageNumber="269">
The oesophageal peduncle is long and the retractor muscle that parts from it lies against the abdomen (
<figureCitation id="E5EF2A30FF9B317FEA4CF99BFED11020" box="[267,367,1544,1570]" captionStart="FIG" captionStartId="67.[154,165,978,995]" captionTargetBox="[154,687,166,940]" captionTargetId="figure-439@67.[142,705,143,966]" captionTargetPageId="67" captionText="FIG. 53. — Didemnum madeleinae n. sp.; A, thorax; B, abdomen; C, larva. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391547" httpUri="https://zenodo.org/record/5391547/files/figure.png" pageId="68" pageNumber="269">Fig. 53A</figureCitation>
).
</paragraph>
<paragraph id="7D6B36B5FF9B317FEBD0F9BBFC421260" blockId="68.[151,685,1064,1666]" lastBlockId="68.[714,1247,1063,1442]" pageId="68" pageNumber="269">
The abdomen (
<figureCitation id="E5EF2A30FF9B317FEA78F9BBFE1E1040" box="[319,416,1576,1602]" captionStart="FIG" captionStartId="67.[154,165,978,995]" captionTargetBox="[154,687,166,940]" captionTargetId="figure-439@67.[142,705,143,966]" captionTargetPageId="67" captionText="FIG. 53. — Didemnum madeleinae n. sp.; A, thorax; B, abdomen; C, larva. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391547" httpUri="https://zenodo.org/record/5391547/files/figure.png" pageId="68" pageNumber="269">Fig. 53B</figureCitation>
) has about the same size as the thorax. The gut loop is slightly twisted. The single testis follicle with seven turns of the sperm duct is located inside the gut loop, beside a very large oocyte (
<figureCitation id="E5EF2A30FF9B317FE8CAFBDBFC4E1260" box="[909,1008,1096,1122]" captionStart="FIG" captionStartId="67.[154,165,978,995]" captionTargetBox="[154,687,166,940]" captionTargetId="figure-439@67.[142,705,143,966]" captionTargetPageId="67" captionText="FIG. 53. — Didemnum madeleinae n. sp.; A, thorax; B, abdomen; C, larva. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391547" httpUri="https://zenodo.org/record/5391547/files/figure.png" pageId="68" pageNumber="269">Fig. 53B</figureCitation>
).
</paragraph>
<paragraph id="7D6B36B5FF9B317FE98DFBFBFBFB1340" blockId="68.[714,1247,1063,1442]" pageId="68" pageNumber="269">
The rather spherical larvae (
<figureCitation id="E5EF2A30FF9B317FEF42FBFBFBD51280" box="[1029,1131,1128,1154]" captionStart="FIG" captionStartId="67.[154,165,978,995]" captionTargetBox="[154,687,166,940]" captionTargetId="figure-439@67.[142,705,143,966]" captionTargetPageId="67" captionText="FIG. 53. — Didemnum madeleinae n. sp.; A, thorax; B, abdomen; C, larva. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391547" httpUri="https://zenodo.org/record/5391547/files/figure.png" pageId="68" pageNumber="269">Fig. 53C</figureCitation>
) are incubated in the densely spiculated basal layer of the colony. Their trunk measures
<quantity id="BA2C9B50FF9B317FEF51FB3BFBE712C3" box="[1046,1113,1191,1218]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="68" pageNumber="269" unit="mm" value="1.0">1 mm</quantity>
but the larvae are not gemmiparous. They have three adhesive papillae, and about 12 short digitiform ampullae on each side. The tail is wound in three quarter of a turn around the trunk.
</paragraph>
<paragraph id="7D6B36B5FF9B317FE98DFADBFCDE13A3" blockId="68.[714,1247,1063,1442]" pageId="68" pageNumber="269">
The spicules (
<figureCitation id="E5EF2A30FF9B317FE82EFADBFC711360" box="[873,975,1352,1378]" captionStart="FIG" captionStartId="68.[268,279,979,996]" captionTargetBox="[151,1247,166,942]" captionTargetId="figure-306@68.[151,700,166,942]" captionTargetPageId="68" captionText="FIG. 54. — Spicules; A, Didemnum madeleinae n. sp.; B, Polysyncraton adelon n. sp. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5391549" httpUri="https://zenodo.org/record/5391549/files/figure.png" pageId="68" pageNumber="269">Fig. 54A</figureCitation>
) are stellate with rather few conical pointed rays: they measure up to 50 µm across.
</paragraph>
</subSubSection>
<subSubSection id="35CE653EFF9B317EE98DFA54FF781443" lastPageId="69" lastPageNumber="270" pageId="68" pageNumber="269" type="discussion">
<paragraph id="7D6B36B5FF9B317FE98DFA54FC9013DD" blockId="68.[714,1248,1479,1666]" box="[714,814,1479,1505]" pageId="68" pageNumber="269">REMARKS</paragraph>
<paragraph id="7D6B36B5FF9B317EE98DFA7BFEB31783" blockId="68.[714,1248,1479,1666]" lastBlockId="69.[151,684,167,577]" lastPageId="69" lastPageNumber="270" pageId="68" pageNumber="269">
<taxonomicName id="BAD44D36FF9B317FE98DFA7BFCE31000" authority="Monniot &amp; Monniot, 2001" authorityName="Monniot &amp; Monniot" authorityYear="2001" box="[714,861,1512,1538]" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Aplousobranchia" pageId="68" pageNumber="269" phylum="Chordata" rank="species" species="madeleinae" status="sp. nov.">
<emphasis id="4FA0EAA7FF9B317FE98DFA7BFCE31000" box="[714,861,1512,1538]" italics="true" pageId="68" pageNumber="269">D. madeleinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="549357DCFF9B317FE824FA74FC231003" box="[867,925,1511,1537]" pageId="68" pageNumber="269" rank="species">n. sp.</taxonomicNameLabel>
is some characters in common with
<taxonomicName id="BAD44D36FF9B317FE857F99BFC251040" authority="Monniot F. &amp; Monniot C., 1999" authorityName="Monniot F. &amp; Monniot C." authorityYear="1999" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Aplousobranchia" pageId="68" pageNumber="269" phylum="Chordata" rank="species" species="captivum">
<emphasis id="4FA0EAA7FF9B317FE857F99BFBAB1023" box="[784,1045,1544,1570]" italics="true" pageId="68" pageNumber="269">Didemnum captivum</emphasis>
Monniot F. &amp; Monniot C., 1999
</taxonomicName>
known only from
<collectingCountry id="05C37625FF9B317FEF35F9BBFB651040" box="[1138,1243,1576,1602]" name="Tanzania" pageId="68" pageNumber="269">Tanzania</collectingCountry>
: the orange colour, the shape and size of the spicules, the position of the lateral thoracic organs and the retractor muscle, and the structure of the abdomen. But the shape of the oral siphon and the cloacal aperture are very different, and the larvae are very different in size. These characters and the geographical distance of this record from
<collectingCountry id="05C37625FF9A317EEB9BFEDBFEF21760" box="[220,332,328,354]" name="Tanzania" pageId="69" pageNumber="270">Tanzania</collectingCountry>
argue persuasively against a synonymy.
</paragraph>
<paragraph id="7D6B36B5FF9A317EEBD0FE1BFF781443" blockId="69.[151,684,167,577]" pageId="69" pageNumber="270">
All other
<taxonomicName id="BAD44D36FF9A317EEA45FE1BFDCC17A0" box="[258,626,392,418]" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Aplousobranchia" pageId="69" pageNumber="270" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="4FA0EAA7FF9A317EEA45FE1BFEC417A0" box="[258,378,392,418]" italics="true" pageId="69" pageNumber="270">Didemnum</emphasis>
<bibRefCitation id="19454B44FF9A317EEAC4FE1BFDA117A0" author="SAVIGNY J. C." box="[387,543,392,418]" pageId="69" pageNumber="270" refId="ref88549" refString="SAVIGNY J. C. 1816. - Recherches anatomiques sur les ascidies composees et les ascidies simples. Systeme de la classe des Ascidies, in Memoires sur les animaux sans vertebres. Part 2. G. Dufour, Paris, 239 p." type="book" year="1816">Savigny, 1816</bibRefCitation>
species
</taxonomicName>
with only one testis follicle, without cloacal languet, and with a larva that has numerous anterior ampullae – all other such species have a very dark pigment and different spicules from this new one.
</paragraph>
</subSubSection>
</treatment>
</document>
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