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<document id="11FC91E4100FD87AA30E7E96436047CB" ID-DOI="10.5852/ejt.2023.910.2365" ID-ISSN="2118-9773" ID-Zenodo-Dep="10350702" ID-ZooBank="DED6151F-FD3A-488D-9CAF-1B0DAD697BA3" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1702283260375" checkinUser="plazi" docAuthor="Kiel, Steffen, Sami, Marco &amp; Taviani, Marco" docDate="2023" docId="C80687AB9C2CFFC2FD18FD92FBF869FA" docLanguage="en" docName="EJT.2023.910.115-160.pdf" docOrigin="European Journal of Taxonomy 910" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2365/10359" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Putzeysia diversii Kiel &amp; Sami &amp; Taviani 2023, sp. nov." docType="treatment" docUuid="6184825A-5A82-44BA-891F-5E40A0E43B7C" docUuidSource="ZooBank" docVersion="1" lastPageNumber="122" masterDocId="343FFFD39C2BFFC5FF98FF85FFA06E2D" masterDocTitle="Mollusks (Gastropoda, Bivalvia) from Miocene cold-seep deposits in northern Italy: revisions and additions" masterLastPageNumber="160" masterPageNumber="115" pageNumber="122" updateTime="1702568871304" updateUser="julia" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2FD2D53436BC5C219CE87AC98473EE42">Mollusks (Gastropoda, Bivalvia) from Miocene cold-seep deposits in northern Italy: revisions and additions</mods:title>
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<mods:name id="427BED2BE1C39F26D6E6A65D6BA1A1AE" type="personal">
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<mods:namePart id="5D66415E540A11CB45644409BA84AF37">Kiel, Steffen</mods:namePart>
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<mods:affiliation id="974F0D65C6B69BA2A511D39CEC20C44C">Swedish Museum of Natural History, Department of Palaeobiology, Box 50007, 10405 Stockholm, Sweden.</mods:affiliation>
<mods:nameIdentifier id="218CCDFBA35A9A58260E039BF18B269C" type="email">steffen.kiel@nrm.se</mods:nameIdentifier>
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<mods:namePart id="EB5D44D18D453042CD27BB2E693CC8FA">Sami, Marco</mods:namePart>
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<mods:affiliation id="482D280F0C1C700F33F9CD5F526D7EA7">Museo Civico di Scienze Naturali, 51, Via Medaglie D’Oro 51, 48018 Faenza, Italy.</mods:affiliation>
<mods:nameIdentifier id="FBCFDD9B3818E2711F07CC4CF156C082" type="email">marco.sami@cheapnet.it</mods:nameIdentifier>
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<mods:namePart id="646F3DE562E723CDF1F38C5C2AA3C260">Taviani, Marco</mods:namePart>
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<mods:affiliation id="EEA8181CF1086620AC9048F2E80818EA">Institute of Marine Sciences, Italian National Research Council, Via Gobetti 101, 40129 Bologna, Italy; and Stazione Zoologica Anton Dohrn, Villa Comunale, 80121 Napoli, Italy.</mods:affiliation>
<mods:nameIdentifier id="0A1B53A6E438774C9DEEB9EE9F27EE29" type="email">marco.taviani@bo.ismar.cnr.it</mods:nameIdentifier>
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<mods:title id="F8A7BA13F37CF7F03F2E7026B0F5FECE">European Journal of Taxonomy</mods:title>
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<mods:date id="67D358BBCAB18C7C950F7006CDD037C0">2023</mods:date>
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<mods:number id="FA88D068EE8BC902D4097018FB6BA785">2023-12-11</mods:number>
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<mods:number id="FEEAF93B592049DBB44EDC9C9818854C">910</mods:number>
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<treatment id="C80687AB9C2CFFC2FD18FD92FBF869FA" LSID="urn:lsid:zoobank.org:act:6184825A-5A82-44BA-891F-5E40A0E43B7C" httpUri="http://treatment.plazi.org/id/C80687AB9C2CFFC2FD18FD92FBF869FA" lastPageNumber="122" pageId="7" pageNumber="122">
<subSubSection id="08B565369C2CFFC2FD18FD92FC146C1F" box="[640,948,534,562]" pageId="7" pageNumber="122" type="nomenclature">
<paragraph id="401036BD9C2CFFC2FD18FD92FC146C1F" blockId="7.[368,1219,534,628]" box="[640,948,534,562]" pageId="7" pageNumber="122">
<heading id="1B5881D19C2CFFC2FD18FD92FC146C1F" box="[640,948,534,562]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="122" reason="2">
<taxonomicName id="87AF4D3E9C2CFFC2FD18FD92FCF26C1C" authority="Kiel &amp; Sami &amp; Taviani, 2023" authorityName="Kiel &amp; Sami &amp; Taviani" authorityYear="2023" box="[640,850,534,561]" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="diversii" status="sp. nov.">
<emphasis id="72DBEAAF9C2CFFC2FD18FD92FCF26C1C" bold="true" box="[640,850,534,561]" italics="true" pageId="7" pageNumber="122">Putzeysia diversii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69E857D49C2CFFC2FCC1FD9DFC146C1F" box="[857,948,536,562]" pageId="7" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="343E3ABF9C2CFFC2FEE8FDBDFB636C7E" box="[368,1219,568,595]" pageId="7" pageNumber="122">
urn:lsid:zoobank.org:act:
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<figureCitation id="D8942A389C2CFFC2FD44FDDFFCF76C59" box="[732,855,602,628]" captionStart="Fig" captionStartId="8.[188,231,1928,1954]" captionTargetBox="[189,1398,893,1886]" captionTargetId="figure-150@8.[189,1398,893,1886]" captionTargetPageId="8" captionText="Fig. 3. Chilodontaidae Wenz, 1938 (Vetigastropoda) from the Miocene Calcari a Lucina seep deposits in northern Italy. A–C. Putzeysia diversii sp. nov. A–B. Holotype, Ca’ Piantè (MSF 1079). C. Paratype, Ca’ Cavalmagra (MSF 1300). D–F. Chilodontaidae indet., Le Colline (MSF 1209)." figureDoi="http://doi.org/10.5281/zenodo.10350714" httpUri="https://zenodo.org/record/10350714/files/figure.png" pageId="7" pageNumber="122">Fig. 3A–C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="08B565369C2CFFC2FF25FD1FFD206D6B" pageId="7" pageNumber="122" type="diagnosis">
<paragraph id="401036BD9C2CFFC2FF25FD1FFE966C99" blockId="7.[189,310,666,692]" box="[189,310,666,692]" pageId="7" pageNumber="122">
<heading id="1B5881D19C2CFFC2FF25FD1FFE966C99" bold="true" box="[189,310,666,692]" fontSize="11" level="3" pageId="7" pageNumber="122" reason="3">
<emphasis id="72DBEAAF9C2CFFC2FF25FD1FFE966C99" bold="true" box="[189,310,666,692]" pageId="7" pageNumber="122">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="401036BD9C2CFFC2FF25FD42FD206D6B" blockId="7.[189,1399,711,838]" pageId="7" pageNumber="122">Shell turbiniform made of at least 4 whorls; suture deep; whorls slightly convex, sculpture cancellate, made of 20–24 opisthocyrt axial ribs per whorl, crossed by 4–5 spirals, intersections tuberculate; basal margin rounded-angular, marked by spiral cord; base weakly sculptured, at least two spiral cords near umbilical slit; aperture nearly circular.</paragraph>
</subSubSection>
<subSubSection id="08B565369C2CFFC2FF25FCE9FB7F6DDB" pageId="7" pageNumber="122" type="etymology">
<paragraph id="401036BD9C2CFFC2FF25FCE9FEE46DAB" blockId="7.[189,324,876,902]" box="[189,324,876,902]" pageId="7" pageNumber="122">
<heading id="1B5881D19C2CFFC2FF25FCE9FEE46DAB" bold="true" box="[189,324,876,902]" fontSize="11" level="3" pageId="7" pageNumber="122" reason="3">
<emphasis id="72DBEAAF9C2CFFC2FF25FCE9FEE46DAB" bold="true" box="[189,324,876,902]" pageId="7" pageNumber="122">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="401036BD9C2CFFC2FF25FC1DFB7F6DDB" blockId="7.[189,1399,920,1014]" pageId="7" pageNumber="122">
The new taxon is dedicated to the amateur paleontologist Mauro Diversi (1964–2021) who provided over the years many important samples to the ‘Museo Civico di Scienze Naturali’ of Faenza, including specimens from the Calcari a
<taxonomicName id="87AF4D3E9C2CFFC2FD82FC58FDCB6DDB" authorityName="Bruguiere" authorityYear="1797" box="[538,619,989,1014]" class="Bivalvia" family="Lucinidae" genus="Lucina" kingdom="Animalia" order="Lucinida" pageId="7" pageNumber="122" phylum="Mollusca" rank="genus">
<emphasis id="72DBEAAF9C2CFFC2FD82FC58FDCB6DDB" box="[538,619,989,1014]" italics="true" pageId="7" pageNumber="122">Lucina</emphasis>
</taxonomicName>
deposits discussed in previous papers by the authors.
</paragraph>
</subSubSection>
<subSubSection id="08B565369C2CFFC2FF25FB9EFAA76ACB" pageId="7" pageNumber="122" type="materials_examined">
<paragraph id="401036BD9C2CFFC2FF25FB9EFE0A6A18" blockId="7.[189,426,1051,1077]" box="[189,426,1051,1077]" pageId="7" pageNumber="122">
<heading id="1B5881D19C2CFFC2FF25FB9EFE0A6A18" bold="true" box="[189,426,1051,1077]" fontSize="11" level="3" pageId="7" pageNumber="122" reason="3">
<emphasis id="72DBEAAF9C2CFFC2FF25FB9EFE0A6A18" bold="true" box="[189,426,1051,1077]" pageId="7" pageNumber="122">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="401036BD9C2CFFC2FF72FBCCFEFC6A4E" blockId="7.[189,976,1097,1157]" box="[234,348,1097,1123]" pageId="7" pageNumber="122">
<emphasis id="72DBEAAF9C2CFFC2FF72FBCCFEFC6A4E" bold="true" box="[234,348,1097,1123]" pageId="7" pageNumber="122">
<typeStatus id="9F14881F9C2CFFC2FF72FBCCFEFC6A4E" box="[234,348,1097,1123]" pageId="7" pageNumber="122" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="401036BD9C2CFFC2FF25FBEFFC706AA9" blockId="7.[189,976,1097,1157]" box="[189,976,1130,1157]" pageId="7" pageNumber="122">
<materialsCitation id="F0C73CE09C2CFFC2FF25FBEFFC706AA9" box="[189,976,1130,1157]" collectionCode="MSF" country="Italy" location="Ca' Piante" pageId="7" pageNumber="122" specimenCode="MSF 1079" specimenCount="1" stateProvince="Emilia-Romagna" typeStatus="holotype">
<collectingCountry id="38B8762D9C2CFFC2FF25FBEFFEAF6AA9" box="[189,271,1130,1156]" name="Italy" pageId="7" pageNumber="122">ITALY</collectingCountry>
–
<emphasis id="72DBEAAF9C2CFFC2FEB4FBEFFDA16AA9" bold="true" box="[300,513,1130,1156]" pageId="7" pageNumber="122">
<collectingRegion id="826BF85F9C2CFFC2FEB4FBEFFDA16AA9" box="[300,513,1130,1156]" country="Italy" name="Emilia-Romagna" pageId="7" pageNumber="122">Emilia-Romagna</collectingRegion>
</emphasis>
•
<location id="457060669C2CFFC2FD82FBEFFD376AA9" LSID="urn:lsid:plazi:treatment:C80687AB9C2CFFC2FD18FD92FBF869FA:457060669C2CFFC2FD82FBEFFD376AA9" box="[538,663,1130,1156]" country="Italy" name="Ca' Piante" pageId="7" pageNumber="122" stateProvince="Emilia-Romagna">Ca’ Piantè</location>
;
<specimenCode id="10099EC69C2CFFC2FD3BFBEFFC806AA9" box="[675,800,1130,1156]" collectionCode="MSF" country="Switzerland" httpUri="http://grbio.org/cool/7f85-3xtv" name="Sauriermuseum Frick" pageId="7" pageNumber="122">MSF 1079</specimenCode>
(H =
<quantity id="87579B589C2CFFC2FCFDFBEFFC646AA9" box="[869,964,1130,1156]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="122" unit="mm" value="3.3">3.3 mm</quantity>
).
</materialsCitation>
</paragraph>
<paragraph id="401036BD9C2CFFC2FF72FB2FFE066AE9" blockId="7.[189,1287,1194,1254]" box="[234,422,1194,1220]" pageId="7" pageNumber="122">
<emphasis id="72DBEAAF9C2CFFC2FF72FB2FFE066AE9" bold="true" box="[234,422,1194,1220]" pageId="7" pageNumber="122">Other material</emphasis>
</paragraph>
<paragraph id="401036BD9C2CFFC2FF25FB49FAA76ACB" blockId="7.[189,1287,1194,1254]" box="[189,1287,1227,1254]" pageId="7" pageNumber="122">
<materialsCitation id="F0C73CE09C2CFFC2FF25FB49FBAB6ACB" box="[189,1035,1227,1254]" collectionCode="MSF" country="Italy" location="Ca' Cavalmagra" pageId="7" pageNumber="122" specimenCode="MSF 1300" specimenCount="1" stateProvince="Tuscany">
<collectingCountry id="38B8762D9C2CFFC2FF25FB49FEAF6ACB" box="[189,271,1228,1254]" name="Italy" pageId="7" pageNumber="122">ITALY</collectingCountry>
–
<emphasis id="72DBEAAF9C2CFFC2FEB4FB49FE336ACB" bold="true" box="[300,403,1228,1254]" pageId="7" pageNumber="122">
<collectingRegion id="826BF85F9C2CFFC2FEB4FB49FE336ACB" box="[300,403,1228,1254]" country="Italy" name="Toscana" pageId="7" pageNumber="122">Tuscany</collectingRegion>
</emphasis>
• 3 specs;
<location id="457060669C2CFFC2FD89FB49FD736AC8" LSID="urn:lsid:plazi:treatment:C80687AB9C2CFFC2FD18FD92FBF869FA:457060669C2CFFC2FD89FB49FD736AC8" box="[529,723,1227,1254]" country="Italy" name="Ca' Cavalmagra" pageId="7" pageNumber="122" stateProvince="Tuscany">Ca’ Cavalmagra</location>
;
<specimenCode id="10099EC69C2CFFC2FD46FB49FCFB6ACB" box="[734,859,1228,1254]" pageId="7" pageNumber="122">MSF 1300</specimenCode>
(H =
<quantity id="87579B589C2CFFC2FC38FB49FC5E6ACB" box="[928,1022,1228,1254]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="122" unit="mm" value="3.2">3.2 mm</quantity>
),
</materialsCitation>
<materialsCitation id="F0C73CE09C2CFFC2FB8AFB49FAA76ACB" box="[1042,1287,1228,1254]" collectionCode="MSF" country="Italy" location="Ca' Cavalmagra" pageId="7" pageNumber="122" specimenCode="MSF 1301" specimenCount="2" stateProvince="Tuscany">
<specimenCode id="10099EC69C2CFFC2FB8AFB49FB2F6ACB" box="[1042,1167,1228,1254]" pageId="7" pageNumber="122">MSF 1301</specimenCode>
[
<specimenCount id="56A9FD349C2CFFC2FB06FB49FB5C6ACB" box="[1182,1276,1228,1254]" pageId="7" pageNumber="122">2 specs</specimenCount>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="08B565369C2CFFC2FF25FA89FD316947" pageId="7" pageNumber="122" type="discussion">
<paragraph id="401036BD9C2CFFC2FF25FA89FE8F6B0B" blockId="7.[189,303,1292,1318]" box="[189,303,1292,1318]" pageId="7" pageNumber="122">
<heading id="1B5881D19C2CFFC2FF25FA89FE8F6B0B" bold="true" box="[189,303,1292,1318]" fontSize="11" level="3" pageId="7" pageNumber="122" reason="3">
<emphasis id="72DBEAAF9C2CFFC2FF25FA89FE8F6B0B" bold="true" box="[189,303,1292,1318]" pageId="7" pageNumber="122">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="401036BD9C2CFFC2FF25FABCFBBB6BD6" blockId="7.[188,1399,1336,1531]" pageId="7" pageNumber="122">
<taxonomicName id="87AF4D3E9C2CFFC2FF25FABCFE2C6B7F" authority="Kiel &amp; Sami &amp; Taviani, 2023" authorityName="Kiel &amp; Sami &amp; Taviani" authorityYear="2023" box="[189,396,1336,1362]" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="diversii" status="sp. nov.">
<emphasis id="72DBEAAF9C2CFFC2FF25FABCFE2C6B7F" box="[189,396,1336,1362]" italics="true" pageId="7" pageNumber="122">Putzeysia diversii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69E857D49C2CFFC2FE0BFABCFE4C6B7E" box="[403,492,1337,1363]" pageId="7" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
differs from the
<typeStatus id="9F14881F9C2CFFC2FD37FABCFD426B7E" box="[687,738,1337,1363]" pageId="7" pageNumber="122">type</typeStatus>
species
<taxonomicName id="87AF4D3E9C2CFFC2FCDBFABCFB876B7F" baseAuthorityName="Aradas" baseAuthorityYear="1847" box="[835,1063,1336,1362]" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="clathrata">
<emphasis id="72DBEAAF9C2CFFC2FCDBFABCFB876B7F" box="[835,1063,1336,1362]" italics="true" pageId="7" pageNumber="122">Putzeysia clathrata</emphasis>
</taxonomicName>
by having much less convex whorl flanks, a less incised suture, and more pronounced sculpture on the base (
<bibRefCitation id="243E4B4C9C2CFFC2FBF0FADFFAF16B59" author="Reitano A. &amp; Scuderi D. &amp; Villari A." box="[1128,1361,1370,1396]" pageId="7" pageNumber="122" pagination="387 - 397" refId="ref20380" refString="Reitano A., Scuderi D. &amp; Villari A. 2022. Lectotype designation for Putzeysia clathrata (Aradas, 1847) (Gastropoda Eucyclidae) and its systematic position. Biodiversity Journal 13: 387 - 397. https: // doi. org / 10.31396 / Biodiv. Jour. 2022.13.2.387.397" type="journal article" year="2022">
Reitano
<emphasis id="72DBEAAF9C2CFFC2FB56FADEFAA96B59" box="[1230,1289,1370,1396]" italics="true" pageId="7" pageNumber="122">et al.</emphasis>
2022
</bibRefCitation>
). A specimen from uppermost Miocene (Messinian) deep-water strata in
<collectingCountry id="38B8762D9C2CFFC2FC6FFAFEFB9A6BB8" box="[1015,1082,1403,1429]" name="Spain" pageId="7" pageNumber="122">Spain</collectingCountry>
identified as
<taxonomicName id="87AF4D3E9C2CFFC2FB46FAF9FE8B6B9A" baseAuthorityName="Aradas" baseAuthorityYear="1847" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" isUncertain="true" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="clathrata">
<emphasis id="72DBEAAF9C2CFFC2FB46FAF9FAEE6BB8" box="[1246,1358,1404,1429]" italics="true" pageId="7" pageNumber="122">Putzeysia</emphasis>
cf.
<emphasis id="72DBEAAF9C2CFFC2FF27FA18FE8B6B9A" box="[191,299,1437,1463]" italics="true" pageId="7" pageNumber="122">clathrata</emphasis>
</taxonomicName>
(see
<bibRefCitation id="243E4B4C9C2CFFC2FEF0FA18FDE56B9A" author="Barrier P. &amp; Zibrowius H. &amp; Lozouet P. &amp; Montenat C. &amp; Ott d'Estevou P. &amp; Serrano F. &amp; Soudet H. - J." box="[360,581,1437,1463]" pageId="7" pageNumber="122" pagination="3 - 13" refId="ref15448" refString="Barrier P., Zibrowius H., Lozouet P., Montenat C., Ott d'Estevou P., Serrano F. &amp; Soudet H. - J. 1991. Une faune de fond dur du bathyal superieur dans le Miocene terminal des cordilleres Betiques (Carboneras, SE Espagne). Mesogee 51: 3 - 13." type="journal article" year="1991">
Barrier
<emphasis id="72DBEAAF9C2CFFC2FE5CFA1BFE5E6B9A" box="[452,510,1437,1463]" italics="true" pageId="7" pageNumber="122">et al.</emphasis>
1991
</bibRefCitation>
: fig. 4.2) differs notably from
<taxonomicName id="87AF4D3E9C2CFFC2FC33FA1BFBDC6B9A" authorityName="Kiel &amp; Sami &amp; Taviani" authorityYear="2023" box="[939,1148,1437,1463]" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="diversii">
<emphasis id="72DBEAAF9C2CFFC2FC33FA1BFBDC6B9A" box="[939,1148,1437,1463]" italics="true" pageId="7" pageNumber="122">Putzeysia diversii</emphasis>
</taxonomicName>
by its much stronger ornament, consisting of only two strong spiral cords per whorl and the axial ribs being more oblique; the ornament is also very strong on the base, in contrast to
<taxonomicName id="87AF4D3E9C2CFFC2FCDCFA64FBB46BD7" authorityName="Kiel &amp; Sami &amp; Taviani" authorityYear="2023" box="[836,1044,1504,1530]" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="diversii">
<emphasis id="72DBEAAF9C2CFFC2FCDCFA64FBB46BD7" box="[836,1044,1504,1530]" italics="true" pageId="7" pageNumber="122">Putzeysia diversii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="401036BD9C2CFFC2FF25F9A4FD316947" blockId="7.[189,1399,1568,1899]" pageId="7" pageNumber="122">
<taxonomicName id="87AF4D3E9C2CFFC2FF25F9A4FE2C6817" authority="Kiel &amp; Sami &amp; Taviani, 2023" authorityName="Kiel &amp; Sami &amp; Taviani" authorityYear="2023" box="[189,396,1568,1594]" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="diversii" status="sp. nov.">
<emphasis id="72DBEAAF9C2CFFC2FF25F9A4FE2C6817" box="[189,396,1568,1594]" italics="true" pageId="7" pageNumber="122">Putzeysia diversii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69E857D49C2CFFC2FE0BF9A4FE4D6816" box="[403,493,1569,1595]" pageId="7" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
differs from the extant
<taxonomicName id="87AF4D3E9C2CFFC2FC98F9A4FB466817" authority="Reitano &amp; Scuderi, 2021" authorityName="Reitano &amp; Scuderi" authorityYear="2021" box="[768,1254,1568,1594]" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="rickyi">
<emphasis id="72DBEAAF9C2CFFC2FC98F9A4FC1A6817" box="[768,954,1568,1594]" italics="true" pageId="7" pageNumber="122">Putzeysia rickyi</emphasis>
<bibRefCitation id="243E4B4C9C2CFFC2FC59F9A5FB466817" author="Reitano A. &amp; Scuderi D." box="[961,1254,1568,1594]" pageId="7" pageNumber="122" pagination="937 - 946" refId="ref20329" refString="Reitano A. &amp; Scuderi D. 2021. A new species of Putzeysia Sulliotti, 1889 (Gastropoda Eucyclidae) from the Ionian Sea (Italy). Biodiversity Journal 12: 937 - 946. https: // doi. org / 10.31396 / Biodiv. Jour. 2021.12.4.937.946" type="journal article" year="2021">Reitano &amp; Scuderi, 2021</bibRefCitation>
</taxonomicName>
, from
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mainly by a slightly lower spire, straighter whorl flanks, and the much weaker ornamentation on the base (
<bibRefCitation id="243E4B4C9C2CFFC2FE99F9E0FD876852" author="Reitano A. &amp; Scuderi D." box="[257,551,1637,1663]" pageId="7" pageNumber="122" pagination="937 - 946" refId="ref20329" refString="Reitano A. &amp; Scuderi D. 2021. A new species of Putzeysia Sulliotti, 1889 (Gastropoda Eucyclidae) from the Ionian Sea (Italy). Biodiversity Journal 12: 937 - 946. https: // doi. org / 10.31396 / Biodiv. Jour. 2021.12.4.937.946" type="journal article" year="2021">Reitano &amp; Scuderi 2021</bibRefCitation>
). Even more similar is the Early Pleistocene to Recent
<taxonomicName id="87AF4D3E9C2CFFC2FB2EF9E3FEDA688C" authority="(Calcara, 1842)" baseAuthorityName="Calcara" baseAuthorityYear="1842" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="wiseri">
<emphasis id="72DBEAAF9C2CFFC2FB2EF9E3FAD76852" box="[1206,1399,1638,1663]" italics="true" pageId="7" pageNumber="122">Putzeysia wiseri</emphasis>
(
<bibRefCitation id="243E4B4C9C2CFFC2FF5DF903FED1688C" author="Calcara P." box="[197,369,1670,1697]" pageId="7" pageNumber="122" pagination="1 - 14" refId="ref16033" refString="Calcara P. 1842. Nuove ricerche ed osservazioni sopra vari molluschi Siciliani. Nuove specie di Calyptrea. Il Maurolico, Giornale del Gabinetto Letterario di Messina 13: 1 - 14." type="journal article" year="1842">Calcara, 1842</bibRefCitation>
)
</taxonomicName>
, also from the Mediterranean Sea, and fossil in southern
<collectingCountry id="38B8762D9C2CFFC2FBA0F903FBCF688D" box="[1080,1135,1670,1696]" name="Italy" pageId="7" pageNumber="122">Italy</collectingCountry>
, which has a spire of similar height, but the whorls are more convex and it has a sculptured base (
<bibRefCitation id="243E4B4C9C2CFFC2FBF7F92DFAD268EF" author="Guidastri R. &amp; Melone G. &amp; Taviani M." box="[1135,1394,1704,1730]" pageId="7" pageNumber="122" pagination="125 - 136" refId="ref17693" refString="Guidastri R., Melone G. &amp; Taviani M. 1984. Systematic position of &quot; Trochus &quot; wiseri Calcara (Prosobranchia: Trochidae). Archiv fur Molluskenkunde 114: 125 - 136." type="journal article" year="1984">
Guidastri
<emphasis id="72DBEAAF9C2CFFC2FB72F92CFA8868EF" box="[1258,1320,1704,1730]" italics="true" pageId="7" pageNumber="122">et al.</emphasis>
1984
</bibRefCitation>
;
<bibRefCitation id="243E4B4C9C2CFFC2FF25F94FFE7E68C9" author="Reitano A. &amp; Scuderi D." box="[189,478,1738,1764]" pageId="7" pageNumber="122" pagination="937 - 946" refId="ref20329" refString="Reitano A. &amp; Scuderi D. 2021. A new species of Putzeysia Sulliotti, 1889 (Gastropoda Eucyclidae) from the Ionian Sea (Italy). Biodiversity Journal 12: 937 - 946. https: // doi. org / 10.31396 / Biodiv. Jour. 2021.12.4.937.946" type="journal article" year="2021">Reitano &amp; Scuderi 2021</bibRefCitation>
). The same applies to two extant species of
<taxonomicName id="87AF4D3E9C2CFFC2FC7BF94EFBF368C9" authorityName="Sulliotti" authorityYear="1889" box="[995,1107,1739,1764]" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="genus">
<emphasis id="72DBEAAF9C2CFFC2FC7BF94EFBF368C9" box="[995,1107,1739,1764]" italics="true" pageId="7" pageNumber="122">Putzeysia</emphasis>
</taxonomicName>
from the
<collectingRegion id="826BF85F9C2CFFC2FB5FF94FFAD668C9" box="[1223,1398,1738,1764]" country="Spain" name="Canarias" pageId="7" pageNumber="122">Canary Islands</collectingRegion>
(Engl &amp;
<bibRefCitation id="243E4B4C9C2CFFC2FEBDF96EFE136928" author="Rolan E." box="[293,435,1771,1797]" pageId="7" pageNumber="122" pagination="93 - 98" refId="ref16950" refString="Engl W. &amp; Rolan E. 2009. Two new species of Putzeysia (Prosobranchia, Chilodontidae) from the Canary Islands. Iberus 27: 93 - 98." type="journal article" year="2009">Rolán 2009</bibRefCitation>
).
<taxonomicName id="87AF4D3E9C2CFFC2FE50F969FD286928" baseAuthorityName="Calcara" baseAuthorityYear="1842" box="[456,648,1772,1797]" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="wiseri">
<emphasis id="72DBEAAF9C2CFFC2FE50F969FD286928" box="[456,648,1772,1797]" italics="true" pageId="7" pageNumber="122">Putzeysia wiseri</emphasis>
</taxonomicName>
is common at mud volcanoes, and carbonate and coral mounds in the deep Gulf of Cadiz (
<bibRefCitation id="243E4B4C9C2CFFC2FE6CF888FD64690A" author="Genio L. &amp; Waren A. &amp; Matos F. L. &amp; Cunha M. R." box="[500,708,1805,1831]" pageId="7" pageNumber="122" pagination="5159 - 5170" refId="ref17321" refString="Genio L., Waren A., Matos F. L. &amp; Cunha M. R. 2013. The snails' tale in deep-sea habitats in the Gulf of Cadiz (NE Atlantic). Biogeosciences 10: 5159 - 5170. https: // doi. org / 10.5194 / bg- 10 - 5159 - 2013" type="journal article" year="2013">
Génio
<emphasis id="72DBEAAF9C2CFFC2FDDBF88BFDDD690A" box="[579,637,1805,1831]" italics="true" pageId="7" pageNumber="122">et al.</emphasis>
2013
</bibRefCitation>
). The Middle Miocene (upper Langhian)
<taxonomicName id="87AF4D3E9C2CFFC2FB3FF88BFAD6690A" authorityName="Kiel &amp; Sami &amp; Taviani" authorityYear="2023" box="[1191,1398,1805,1831]" class="Gastropoda" family="Eucyclidae" genus="Putzeysia" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="122" phylum="Mollusca" rank="species" species="diversii">
<emphasis id="72DBEAAF9C2CFFC2FB3FF88BFAD6690A" box="[1191,1398,1805,1831]" italics="true" pageId="7" pageNumber="122">Putzeysia diversii</emphasis>
</taxonomicName>
appears to be the earliest fossil record of the genus, previously known from the Messinian (uppermost Miocene) of
<collectingCountry id="38B8762D9C2CFFC2FECDF8D5FE386947" box="[341,408,1872,1898]" name="Spain" pageId="7" pageNumber="122">Spain</collectingCountry>
(
<bibRefCitation id="243E4B4C9C2CFFC2FE3FF8D5FD246947" author="Barrier P. &amp; Zibrowius H. &amp; Lozouet P. &amp; Montenat C. &amp; Ott d'Estevou P. &amp; Serrano F. &amp; Soudet H. - J." box="[423,644,1872,1899]" pageId="7" pageNumber="122" pagination="3 - 13" refId="ref15448" refString="Barrier P., Zibrowius H., Lozouet P., Montenat C., Ott d'Estevou P., Serrano F. &amp; Soudet H. - J. 1991. Une faune de fond dur du bathyal superieur dans le Miocene terminal des cordilleres Betiques (Carboneras, SE Espagne). Mesogee 51: 3 - 13." type="journal article" year="1991">
Barrier
<emphasis id="72DBEAAF9C2CFFC2FD9BF8D4FD966947" box="[515,566,1872,1898]" italics="true" pageId="7" pageNumber="122">et al</emphasis>
. 1991
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="08B565369C2CFFC2FF25F815FBF869FA" pageId="7" pageNumber="122" type="distribution">
<paragraph id="401036BD9C2CFFC2FF25F815FDDD6987" blockId="7.[189,637,1936,1962]" box="[189,637,1936,1962]" pageId="7" pageNumber="122">
<heading id="1B5881D19C2CFFC2FF25F815FDDD6987" bold="true" box="[189,637,1936,1962]" fontSize="11" level="3" pageId="7" pageNumber="122" reason="3">
<emphasis id="72DBEAAF9C2CFFC2FF25F815FDDD6987" bold="true" box="[189,637,1936,1962]" pageId="7" pageNumber="122">Stratigraphic and geographic range</emphasis>
</heading>
</paragraph>
<paragraph id="401036BD9C2CFFC2FF25F838FBF869FA" blockId="7.[189,1112,1981,2007]" box="[189,1112,1981,2007]" pageId="7" pageNumber="122">
Middle to Upper Miocene (upper Langhian to upper Tortonian), northern
<collectingCountry id="38B8762D9C2CFFC2FB84F838FBF469FA" box="[1052,1108,1981,2007]" name="Italy" pageId="7" pageNumber="122">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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