156 lines
17 KiB
XML
156 lines
17 KiB
XML
<document id="3CAAE31AFFC1FBDA587FCF587A7B94DA" ID-DOI="10.11646/zootaxa.5536.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14033618" ID-ZooBank="5F1FBB59-9C69-4E66-9B0D-69A015F30EAA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730712964139" checkinUser="plazi" docAuthor="Alekseev, Vitalii & Bukejs, Andris" docDate="2024" docId="03F4DF38FFF76654FF506B2FBA6C52B8" docLanguage="en" docName="zootaxa.5536.2.1.pdf" docOrigin="Zootaxa 5536 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thanatoplagia Alekseev et Bukejs 2024, gen. nov." docType="treatment" docVersion="2" lastPageNumber="224" masterDocId="FFCDA740FFE06643FFC76A41BB4B5607" masterDocTitle="A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene ‘ amber forests’" masterLastPageNumber="247" masterPageNumber="201" pageNumber="224" updateTime="1730713620892" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="4908481822E150AD3950225BAE18124A">
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<mods:title id="11DDE901A47E9490B3049762C8F5DFBE">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene ‘ amber forests’</mods:title>
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<mods:name id="7AEAD87ACA0C12A60E0BE42CA019D408" type="personal">
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<mods:role id="BBEF47295D4DA38BE7955FB5F72529A6">
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<mods:roleTerm id="0951773D5A60FA4079CA11D5198A886D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C28167E832BBEAF86B693DC2116EED40">Alekseev, Vitalii</mods:namePart>
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<mods:affiliation id="559A0E19F60626F88C48034802F02393">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia. & Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. & Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; & Department of Ecology & Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
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</mods:name>
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<mods:name id="E665ABF621302FE8E7A5FE56D7CE2610" type="personal">
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<mods:roleTerm id="3291B2019134CDD8E7198CD69E1C4E55">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DB15A18D09B46B0DBB0C0F822A3267AB">Bukejs, Andris</mods:namePart>
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<mods:affiliation id="6FE8C16EE627ACD5D6A8177155E1CE7F">Institute of Life Sciences and Technologies, Daugavpils University, Vienības 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
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<mods:typeOfResource id="4D3BD7E1178F6E55C5F9BF422E0DD378">text</mods:typeOfResource>
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<mods:title id="749A49CE05642BF74775283082C16021">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E2211D0B922B6255F6793F14F9D39484">
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<mods:date id="F683A97BA9FEDAC1A6A4B171624D80DE">2024</mods:date>
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<mods:detail id="E9D6145D71DC8319BAE874443D479DF4" type="pubDate">
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<mods:number id="D4BFE16A27F3B679BBC2F51196FA2313">2024-11-04</mods:number>
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<mods:number id="0D117738EF920385DFB4F22C1FFBE0B4">5536</mods:number>
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<mods:detail id="33AF227A4C04F52B26CDF5EC4CF8038F" type="issue">
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<mods:number id="E5482575135EC6A1934B1AC9AE81AB0B">2</mods:number>
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<mods:start id="6977BE61551FC95E44CEF3C846A65245">201</mods:start>
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<mods:end id="391105F3C4F4A6D5EDA8ED3A9BC42E71">247</mods:end>
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<mods:url id="FA4371D159A884EA9D1749723352F17C">http://dx.doi.org/10.11646/zootaxa.5536.2.1</mods:url>
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<mods:classification id="861BA4CDDB716A55759AA0A398371D31">journal article</mods:classification>
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<mods:identifier id="5AEF1DFEDF707EE020ED43445F366B81" type="DOI">10.11646/zootaxa.5536.2.1</mods:identifier>
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<mods:identifier id="B0ED537287D346D8672FB0E8349C9E28" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F4DF38FFF76654FF506B2FBA6C52B8" LSID="urn:lsid:plazi:treatment:03F4DF38FFF76654FF506B2FBA6C52B8" httpUri="http://treatment.plazi.org/id/03F4DF38FFF76654FF506B2FBA6C52B8" lastPageNumber="224" pageId="23" pageNumber="224">
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<subSubSection id="C3473DA5FFF76654FF506B2FB84D578E" box="[151,774,366,393]" pageId="23" pageNumber="224" type="nomenclature">
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<paragraph id="8BE26E2EFFF76654FF506B2FB84D578E" blockId="23.[151,774,366,393]" box="[151,774,366,393]" pageId="23" pageNumber="224">
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<heading id="D0AAD942FFF76654FF506B2FB84D578E" bold="true" box="[151,774,366,393]" fontSize="11" level="1" pageId="23" pageNumber="224" reason="1">
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<emphasis id="B929B23CFFF76654FF506B2FB84D578E" bold="true" box="[151,774,366,393]" pageId="23" pageNumber="224">
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Genus
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<taxonomicName id="4C5D15ADFFF76654FF296B2FB9DB578F" authority="Alekseev et Bukejs" authorityName="Alekseev et Bukejs" authorityYear="2024" box="[238,656,366,393]" class="Magnoliopsida" family="Fagaceae" genus="Thanatoplagia" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="224" phylum="Tracheophyta" rank="genus" status="gen. nov.">
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<emphasis id="B929B23CFFF76654FF296B2FBAEB578E" bold="true" box="[238,416,366,393]" italics="true" pageId="23" pageNumber="224">Thanatoplagia</emphasis>
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Alekseev et Bukejs
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</taxonomicName>
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<taxonomicNameLabel id="A21A0F47FFF76654FD5F6B2EB84D578E" box="[664,774,367,393]" pageId="23" pageNumber="224" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3473DA5FFF76654FF506BF5BA1354A4" pageId="23" pageNumber="224" type="materials_examined">
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<paragraph id="8BE26E2EFFF76654FF506BF5B8BF57CC" blockId="23.[151,1012,435,460]" box="[151,1012,435,460]" pageId="23" pageNumber="224">
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<typeStatus id="54E6D08CFFF76654FF506BF5BB8257CB" box="[151,201,436,460]" pageId="23" pageNumber="224">Type</typeStatus>
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species:
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<taxonomicName id="4C5D15ADFFF76654FEE16BF2B99657CC" authority="Alekseev et Bukejs" authorityName="Alekseev et Bukejs" authorityYear="2024" box="[294,733,435,459]" class="Magnoliopsida" family="Fagaceae" genus="Thanatoplagia" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="224" phylum="Tracheophyta" rank="species" species="tamutisi" status="sp. nov.">
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<emphasis id="B929B23CFFF76654FEE16BF2B95F57CC" box="[294,532,435,459]" italics="true" pageId="23" pageNumber="224">Thanatoplagia tamutisi</emphasis>
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Alekseev et Bukejs
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</taxonomicName>
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<taxonomicNameLabel id="A21A0F47FFF76654FD236BF2B87D57CC" box="[740,822,435,459]" pageId="23" pageNumber="224" rank="species">
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<emphasis id="B929B23CFFF76654FD236BF2B86457CC" bold="true" box="[740,815,435,459]" pageId="23" pageNumber="224">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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, designated herein
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</paragraph>
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<paragraph id="8BE26E2EFFF76654FF506BB8BA1354A4" blockId="23.[151,1437,505,1216]" pageId="23" pageNumber="224">
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<emphasis id="B929B23CFFF76654FF506BB8BAE55414" bold="true" box="[151,430,505,531]" pageId="23" pageNumber="224">Taxonomic assignment.</emphasis>
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The specimen under study belongs to the tribe
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<taxonomicName id="4C5D15ADFFF76654FC796BB8BF7A5414" authorityName="Billberg" authorityYear="1820" box="[958,1073,505,531]" class="Insecta" family="Zopheridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="tribe" tribe="Sarrotriini">Sarrotriini</taxonomicName>
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based on the combination of the following characters: (1) dorsally concealed antennal insertions; (2) antennae setose, stout, 10- or 11-segmeneted, without clearly separated antennal club; (3) antennal insertions distant from eyes; (4) procoxal cavities open posteriorly; (5) all tarsi tetramerous, not dilated; and (6) metacoxae separated by intercoxal abdominal process rounded apically.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3473DA5FFF76654FF0068ECBEC9520C" pageId="23" pageNumber="224" type="diagnosis">
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<paragraph id="8BE26E2EFFF76654FF0068ECBEC9520C" blockId="23.[151,1437,505,1216]" pageId="23" pageNumber="224">
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<emphasis id="B929B23CFFF76654FF0068ECBA8654C0" bold="true" box="[199,461,685,711]" pageId="23" pageNumber="224">Differential diagnosis.</emphasis>
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The tribe
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<taxonomicName id="4C5D15ADFFF76654FDF868ECB9F954C0" authorityName="Billberg" authorityYear="1820" box="[575,690,685,711]" class="Insecta" family="Zopheridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="tribe" tribe="Sarrotriini">Sarrotriini</taxonomicName>
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includes three extant genera from the Western Palearctic:
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<taxonomicName id="4C5D15ADFFF76654FAFE68ECBA7A54EC" authority="Reitter, 1882" authorityName="Reitter" authorityYear="1882" class="Insecta" family="Zopheridae" genus="Diplagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="B929B23CFFF76654FAFE68ECBED654C0" box="[1337,1437,685,711]" italics="true" pageId="23" pageNumber="224">Diplagia</emphasis>
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<bibRefCitation id="EFCC13DFFFF76654FF506890BA7A54EC" author="Reitter, E." box="[151,305,721,747]" pageId="23" pageNumber="224" pagination="113 - 149" refId="ref27502" refString="Reitter, E. (1882) Bestimmungs-Tabellen der europaischen Coleopteren. VI. Enhaltend die Familien: Colydiidae, Rhysodidae, Trogositidae. Verhandlungen des Naturforschenden Vereines in Brunn, 22, 113 - 149." type="journal article" year="1882">Reitter, 1882</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C5D15ADFFF76654FE846890B93654EC" authority="Latreille, 1796" authorityName="Latreille" authorityYear="1796" box="[323,637,721,747]" class="Insecta" family="Zopheridae" genus="Orthocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="B929B23CFFF76654FE846890BA8854EC" box="[323,451,721,747]" italics="true" pageId="23" pageNumber="224">Orthocerus</emphasis>
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Latreille, 1796
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</taxonomicName>
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, and
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<taxonomicName id="4C5D15ADFFF76654FD036890BE7D54EC" authority="Schauffuss (Slipinski & Schuh 2008)" authorityName="Schauffuss (Slipinski & Schuh" authorityYear="2008" box="[708,1334,721,750]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="B929B23CFFF76654FD036890B82754EC" box="[708,876,721,747]" italics="true" pageId="23" pageNumber="224">Helioctamenus</emphasis>
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Schauffuss (
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<bibRefCitation id="EFCC13DFFFF76654FBCF6892BE6554EC" author="Slipinski, S. A. & Schuh, R." box="[1032,1326,721,750]" pageId="23" pageNumber="224" refId="ref28217" refString="Slipinski, S. A. & Schuh, R. (2008) Zopheridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Apollo Books, Stenstrup, pp. 78 - 87." type="book" year="2008">Ślipiński & Schuh 2008</bibRefCitation>
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)
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</taxonomicName>
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. Despite the apparent similarity, we conclude that the studied fossil cannot be attributed to any of these genera, and thus we propose a new genus for this fossil beetle.
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<taxonomicName id="4C5D15ADFFF76654FD756958B81C5534" authorityName="Alekseev & Bukejs" authorityYear="2024" box="[690,855,793,819]" class="Magnoliopsida" family="Fagaceae" genus="Thanatoplagia" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="224" phylum="Tracheophyta" rank="genus" status="gen. nov.">
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<emphasis id="B929B23CFFF76654FD756958B81C5534" box="[690,855,793,819]" italics="true" pageId="23" pageNumber="224">Thanatoplagia</emphasis>
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</taxonomicName>
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<emphasis id="B929B23CFFF76654FCA66958B8875534" bold="true" box="[865,972,793,819]" pageId="23" pageNumber="224">
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<taxonomicNameLabel id="A21A0F47FFF76654FCA66958B8875534" box="[865,972,793,819]" pageId="23" pageNumber="224" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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clearly differs from
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<taxonomicName id="4C5D15ADFFF76654FB7A6958BE6A5534" authorityName="Reitter" authorityYear="1882" box="[1213,1313,793,819]" class="Insecta" family="Zopheridae" genus="Diplagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="B929B23CFFF76654FB7A6958BE6A5534" box="[1213,1313,793,819]" italics="true" pageId="23" pageNumber="224">Diplagia</emphasis>
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</taxonomicName>
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in lacking interfacetal setae, but possessing sparsely setose antennae, and pronotum distinctly narrower than elytral base; differs from
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<taxonomicName id="4C5D15ADFFF76654FEE16920BAED557C" authorityName="Latreille" authorityYear="1796" box="[294,422,865,891]" class="Insecta" family="Zopheridae" genus="Orthocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="B929B23CFFF76654FEE16920BAED557C" box="[294,422,865,891]" italics="true" pageId="23" pageNumber="224">Orthocerus</emphasis>
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</taxonomicName>
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in antennal structure (gradually widened towards apex, not fusiform, sparsely and shortly setose in the new genus); and it differs from
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<taxonomicName id="4C5D15ADFFF76654FD5469C4B8705598" authorityName="Schauffuss (Slipinski & Schuh" authorityYear="2008" box="[659,827,901,927]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="B929B23CFFF76654FD5469C4B8705598" box="[659,827,901,927]" italics="true" pageId="23" pageNumber="224">Helioctamenus</emphasis>
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</taxonomicName>
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in having stout and strongly transverse antennomeres 4–9, and pronotum with three distinct impressions (two oval transverse, shallow impressions laterobasally and one large, rounded impression anteromedially). The new fossil genus differs from other similar sarrotriines in a well-developed antennomere 11 (as long as antennomere 10), pronounced humeri and developed metathoracic wings.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3473DA5FFF76654FF006E54B9EF5270" pageId="23" pageNumber="224" type="etymology">
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<paragraph id="8BE26E2EFFF76654FF006E54B9EF5270" blockId="23.[151,1437,505,1216]" pageId="23" pageNumber="224">
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<emphasis id="B929B23CFFF76654FF006E54BA055228" bold="true" box="[199,334,1045,1071]" pageId="23" pageNumber="224">Etymology.</emphasis>
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The name of the new genus is a compound word and combines the old Greek word θάνατος (
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<emphasis id="B929B23CFFF76654FF666E78BA4A5254" box="[161,257,1081,1107]" italics="true" pageId="23" pageNumber="224">thánatos</emphasis>
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, death) and the old Greek word πλαγία (
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<emphasis id="B929B23CFFF76654FD086E78B85C5254" box="[719,791,1081,1107]" italics="true" pageId="23" pageNumber="224">plagía</emphasis>
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, slope or hillside) referring to
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<taxonomicName id="4C5D15ADFFF76654FBB56E78BF9D5254" authorityName="Reitter" authorityYear="1882" box="[1138,1238,1081,1107]" class="Insecta" family="Zopheridae" genus="Diplagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="B929B23CFFF76654FBB56E78BF9D5254" box="[1138,1238,1081,1107]" italics="true" pageId="23" pageNumber="224">Diplagia</emphasis>
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</taxonomicName>
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, the extant genus of the tribe
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<taxonomicName id="4C5D15ADFFF76654FEDE6E1CBADB5270" authorityName="Billberg" authorityYear="1820" box="[281,400,1117,1143]" class="Insecta" family="Zopheridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="tribe" tribe="Sarrotriini">Sarrotriini</taxonomicName>
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. The gender is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3473DA5FFF76654FF006EC0BA6C52B8" pageId="23" pageNumber="224" type="description">
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<paragraph id="8BE26E2EFFF76654FF006EC0BA6C52B8" blockId="23.[151,1437,505,1216]" pageId="23" pageNumber="224">
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<emphasis id="B929B23CFFF76654FF006EC0BA12529C" bold="true" box="[199,345,1153,1179]" pageId="23" pageNumber="224">Description.</emphasis>
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The new genus is monotypic. Therefore, the generic description considerably overlaps that of the
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<typeStatus id="54E6D08CFFF76654FF506EE7BB8C52C7" box="[151,199,1190,1216]" pageId="23" pageNumber="224">type</typeStatus>
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species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |