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<document id="DB070B4BE18C0A800024516D8245D767" ID-DOI="10.11646/zootaxa.5536.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14033739" ID-ZooBank="DAE89796-6FFD-4B60-A5EA-00E391A8270A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730713116456" checkinUser="plazi" docAuthor="Trýzna, Miloš, Blažej, Lukáš &amp; Rakotonirina, Jean Claude" docDate="2024" docId="038087A3C032FF8825861A9BFEFE89F5" docLanguage="en" docName="zootaxa.5536.2.5.pdf" docOrigin="Zootaxa 5536 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.2.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diastatotropis petulae Tryzna, Blazej &amp; Rakotonirina 2024, sp. nov." docType="treatment" docUuid="215CD05F-3555-4AB7-B8E4-989EC7300906" docUuidSource="ZooBank" docVersion="2" lastPageNumber="303" masterDocId="FFB9FFDBC035FF8425111A0CFFCF8D30" masterDocTitle="Two new species of Diastatotropis Lacordaire from protected areas in central and east Madagascar (Coleoptera: Anthribidae)" masterLastPageNumber="304" masterPageNumber="291" pageNumber="298" updateTime="1730713724897" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7CF2DBCA1151EB1843387176F4D62791">Two new species of Diastatotropis Lacordaire from protected areas in central and east Madagascar (Coleoptera: Anthribidae)</mods:title>
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<mods:namePart id="54AC244AAC56189D1EA5B33FDB0A169D">Trýzna, Miloš</mods:namePart>
<mods:affiliation id="8ED3FD524172AB2AED07537B3D647D87">Mendel University in Brno, Faculty of Forestry and Wood Technology, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. &amp; Municipal Museum of Ústí nad Labem, Natural History Department, Masarykova 1000 / 3, CZ- 400 01 Ústí nad Labem, Czech Republic.</mods:affiliation>
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<mods:namePart id="C1B33A8F6C803594D562EEED41F48B9F">Blažej, Lukáš</mods:namePart>
<mods:affiliation id="FE3DF8C9A63F38C0BCF27A83E4E7BDAD">Regional Museum and Gallery in Česká Lípa, square Osvobození 297, CZ- 470 01 Česká Lípa, Czech Republic. &amp; Bohemian Switzerland National Park Administration, Research and Nature Conservation Department, Pražská 52, CZ- 407 46 Krásná Lípa, Czech Republic.</mods:affiliation>
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<mods:namePart id="D5D92288D1D302B881CD5633065DCFB9">Rakotonirina, Jean Claude</mods:namePart>
<mods:affiliation id="F46765F0F6584A1DC5F93B883AC1F491">Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar. &amp; Département dEntomologie-Cultures, Elevage et Santé, Faculté des Sciences, BP 906, Université dAntananarivo, Antananarivo, Madagascar.</mods:affiliation>
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<taxonomicName id="4C294D36C032FF8325861A9BFCE38D81" authority="Tryzna, Blazej &amp; Rakotonirina" authorityName="Tryzna, Blazej &amp; Rakotonirina" authorityYear="2024" box="[151,812,150,178]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="species" species="petulae" status="sp. nov.">
<emphasis id="B95DEAA7C032FF8325861A9BFE6F8D82" bold="true" box="[151,416,151,178]" italics="true" pageId="7" pageNumber="298">Diastatotropis petulae</emphasis>
Trýzna, Blažej &amp; Rakotonirina
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http://zoobank.org/
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<paragraph id="8B9636B5C032FF8325861AD3FEE88DC9" blockId="7.[151,1169,150,249]" box="[151,295,223,249]" pageId="7" pageNumber="298">
(
<figureCitation id="13122A30C032FF83258E1AD3FED08DC9" box="[159,287,223,249]" captionStart-0="FIGURES 1620" captionStart-1="FIGURES 2125" captionStartId-0="8.[151,264,1943,1967]" captionStartId-1="9.[151,264,1943,1967]" captionTargetBox-0="[178,1408,181,1912]" captionTargetBox-1="[178,1408,182,1918]" captionTargetId-0="figure-17@8.[178,1408,181,1918]" captionTargetId-1="figure-17@9.[178,1408,181,1918]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 1620. Diastatotropis petulae, holotype male; 16, habitus in dorsal view; 17, head; 18, pronotum; 19, left antenna; 20, habitus in lateral view. Scale bars in mm." captionText-1="FIGURES 2125. Diastatotropis petulae, 2124, paratype male, terminalia; 21, aedeagus in lateral view; 22, aedeagus in dorsal view; 23, tegmen in ventral view; 24, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm. 25, Ambohitantely Special Reserve, overall view of collecting site of D. petulae." figureDoi-0="http://doi.org/10.5281/zenodo.14033755" figureDoi-1="http://doi.org/10.5281/zenodo.14033757" httpUri-0="https://zenodo.org/record/14033755/files/figure.png" httpUri-1="https://zenodo.org/record/14033757/files/figure.png" pageId="7" pageNumber="298">Figs 1624</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C333653EC032FF8325861B2BFC008CC5" pageId="7" pageNumber="298" type="materials_examined">
<paragraph id="8B9636B5C032FF8325861B2BFD618C55" blockId="7.[151,1437,295,1978]" pageId="7" pageNumber="298">
<materialsCitation id="3B413CE8C032FF8325861B2BFF1D8C56" country="Madagascar" location="Ankazobe town" municipality="Ambohitantely Special Reserve" pageId="7" pageNumber="298" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
<emphasis id="B95DEAA7C032FF8325861B2BFEF98C71" bold="true" box="[151,310,295,321]" pageId="7" pageNumber="298">
<typeStatus id="54928817C032FF8325861B2BFF1D8C71" box="[151,210,295,321]" pageId="7" pageNumber="298">Type</typeStatus>
locality.
</emphasis>
C
<collectingCountry id="F33E7625C032FF8324461B2BFE2F8C71" box="[343,480,295,321]" name="Madagascar" pageId="7" pageNumber="298">Madagascar</collectingCountry>
,
<collectingRegion id="49EDF857C032FF8324FA1B2BFD098C72" box="[491,710,295,322]" country="Madagascar" name="Antananarivo" pageId="7" pageNumber="298">Antananarivo prov.</collectingRegion>
, natural forest in
<collectingMunicipality id="6BF2ACCFC032FF83269B1B2BFB228C72" box="[906,1261,295,322]" pageId="7" pageNumber="298">Ambohitantely Special Reserve</collectingMunicipality>
near
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</materialsCitation>
,
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;
<geoCoordinate id="EE1D5072C032FF83246C1B40FDDD8C55" box="[381,530,331,358]" degrees="47" direction="east" minutes="17" orientation="longitude" pageId="7" pageNumber="298" precision="15" seconds="16" value="47.287777">E 47°17´16´´</geoCoordinate>
,
<quantity id="4CD19B50C032FF83270F1B47FDBC8C56" box="[542,627,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.623" pageId="7" pageNumber="298" unit="m" value="1623.0">1623 m</quantity>
a.s.l.
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<paragraph id="8B9636B5C032FF8325D61B63FC008CC5" blockId="7.[151,1437,295,1978]" pageId="7" pageNumber="298">
<emphasis id="B95DEAA7C032FF8325D61B63FD888CB9" bold="true" box="[199,583,367,393]" pageId="7" pageNumber="298">
Type material.
<typeStatus id="54928817C032FF8324921B63FE3F8CB9" box="[387,496,367,393]" pageId="7" pageNumber="298" type="holotype">Holotype</typeStatus>
(male)
</emphasis>
(
<figureCitation id="13122A30C032FF8327481B63FD2F8CB9" box="[601,736,367,393]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[178,1408,181,1912]" captionTargetId="figure-17@8.[178,1408,181,1918]" captionTargetPageId="8" captionText="FIGURES 1620. Diastatotropis petulae, holotype male; 16, habitus in dorsal view; 17, head; 18, pronotum; 19, left antenna; 20, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033755" httpUri="https://zenodo.org/record/14033755/files/figure.png" pageId="7" pageNumber="298">Figs 1620</figureCitation>
):
<collectingCountry id="F33E7625C032FF8327EA1B63FC058CB9" box="[763,970,367,393]" name="Madagascar" pageId="7" pageNumber="298">MADAGASCAR</collectingCountry>
, 19.-25.xii. / AMBOHITANTELY Spec. Res., / circuit Botanique, 2019 /
<geoCoordinate id="EE1D5072C032FF8327031B9FFD668C9D" box="[530,681,403,429]" degrees="18" direction="south" minutes="11" orientation="latitude" pageId="7" pageNumber="298" precision="15" seconds="44" value="-18.195555">S 18°11´44´´</geoCoordinate>
;
<geoCoordinate id="EE1D5072C032FF8327A61B98FC808C9D" box="[695,847,403,430]" degrees="47" direction="east" minutes="17" orientation="longitude" pageId="7" pageNumber="298" precision="15" seconds="16" value="47.287777">E 47°17´16´´</geoCoordinate>
, /
<quantity id="4CD19B50C032FF83267E1B9FFC068C9E" box="[879,969,403,430]" metricMagnitude="3" metricUnit="m" metricValue="1.623" pageId="7" pageNumber="298" unit="m" value="1623.0">1623 m</quantity>
, M. Trýzna leg. [p] (
<collectionCode id="ED38AE70C032FF8321D41B9FFAEC8C9D" box="[1221,1315,403,429]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="7" pageNumber="298">MMBC</collectionCode>
).
<emphasis id="B95DEAA7C032FF8320261B9FFDD88CE1" bold="true" pageId="7" pageNumber="298">
<typeStatus id="54928817C032FF8320261B9FFA538C9D" box="[1335,1436,403,429]" pageId="7" pageNumber="298" type="allotype">Allotype</typeStatus>
(female),
<specimenCount id="9D2FFD3CC032FF8324161BBBFE7F8CE1" box="[263,432,439,465]" count="2" pageId="7" pageNumber="298" type="generic" typeStatus="paratype">two paratypes</specimenCount>
(males):
</emphasis>
the same data as holotype (
<collectionCode id="ED38AE70C032FF83265C1BBBFC6B8CE1" box="[845,932,439,465]" pageId="7" pageNumber="298">MTDC</collectionCode>
). Red label [p]
<emphasis id="B95DEAA7C032FF83215E1BBBFC008CC5" bold="true" pageId="7" pageNumber="298">
<typeStatus id="54928817C032FF83215E1BBBFB218CE1" box="[1103,1262,439,465]" pageId="7" pageNumber="298" type="holotype">HOLOTYPE</typeStatus>
/
<typeStatus id="54928817C032FF8320131BBBFA538CE1" box="[1282,1436,439,465]" pageId="7" pageNumber="298" type="allotype">ALLOTYPE</typeStatus>
/
<typeStatus id="54928817C032FF8325B71BD7FEF58CC5" box="[166,314,475,501]" pageId="7" pageNumber="298" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C294D36C032FF8324411BD7FE3D8CC5" authorityName="Lacordaire" authorityYear="1866" box="[336,498,475,501]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA7C032FF8324411BD7FE3D8CC5" bold="true" box="[336,498,475,501]" italics="true" pageId="7" pageNumber="298">Diastatotropis</emphasis>
</taxonomicName>
/
<taxonomicName id="4C294D36C032FF8327191BD7FD938CC6" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[520,604,475,502]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="species" species="petulae" status="sp. nov.">
<emphasis id="B95DEAA7C032FF8327191BD7FD938CC6" bold="true" box="[520,604,475,502]" italics="true" pageId="7" pageNumber="298">petulae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57DCC032FF8327721BD7FD738CC5" box="[611,700,475,501]" pageId="7" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
/ M. Trýzna det., 2024.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C333653EC032FF8325D61BF3FB7A8FC2" pageId="7" pageNumber="298" type="diagnosis">
<paragraph id="8B9636B5C032FF8325D61BF3FB7A8FC2" blockId="7.[151,1437,295,1978]" pageId="7" pageNumber="298">
<emphasis id="B95DEAA7C032FF8325D61BF3FE8D8F29" bold="true" box="[199,322,511,537]" pageId="7" pageNumber="298">Diagnosis.</emphasis>
Small-sized species (7.09.0 mm). Rostrum with two distinct longitudinal carinae, between these carinae longitudinal flat shiny area imitating central carina. Antennal club slender and oblong, terminal segment elongate-oval, distinctly longer than wide. Elytra convex, with indistinct sub-basal tubercles. Pronotum with one central slender and two lateral wider longitudinal dark yellow stripes. Each elytron with four longitudinal dark yellow stripes: one short in scutellar area; one in the second elytral interval reaching from posterior part of sub-basal tubercle to apex of elytron; one on the fourth elytral interval reaching from anterior margin of elytron to its half; and one on the eighth elytral interval. Pygidium with longitudinal dark yellow strip in central part.
</paragraph>
</subSubSection>
<subSubSection id="C333653EC032FF8825D618F7FD088F29" lastPageId="12" lastPageNumber="303" pageId="7" pageNumber="298" type="description">
<paragraph id="8B9636B5C032FF8325D618F7FF008EDD" blockId="7.[151,1437,295,1978]" pageId="7" pageNumber="298">
<emphasis id="B95DEAA7C032FF8325D618F7FDE88E25" bold="true" box="[199,551,763,789]" pageId="7" pageNumber="298">
Description.
<typeStatus id="54928817C032FF83247218F7FE1F8E25" box="[355,464,763,789]" pageId="7" pageNumber="298" type="holotype">Holotype</typeStatus>
(male)
</emphasis>
(
<figureCitation id="13122A30C032FF83272818F7FD0F8E25" box="[569,704,763,789]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[178,1408,181,1912]" captionTargetId="figure-17@8.[178,1408,181,1918]" captionTargetPageId="8" captionText="FIGURES 1620. Diastatotropis petulae, holotype male; 16, habitus in dorsal view; 17, head; 18, pronotum; 19, left antenna; 20, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033755" httpUri="https://zenodo.org/record/14033755/files/figure.png" pageId="7" pageNumber="298">Figs 1620</figureCitation>
).
<emphasis id="B95DEAA7C032FF8327C718F0FCB78E25" box="[726,888,764,789]" italics="true" pageId="7" pageNumber="298">Measurements</emphasis>
(in mm): Total body length—9.55. Head: total length—2.05; length of rostrum—1.18; maximum width of rostrum—1.30; length of eye—0.93; maximum width across eyes—1.68; minimum distance between eyes—0.63. Antenna: length of segments: II—0.30, III—0.41, IV—0.31, V—0.26, VI—0.21, VII—0.19, VIII—0.17, IX—0.41, X—0.21, XI—0.46, width of segment IX—0.20. Pronotum: maximum length—2.13; width at carina (= maximal width in this case)—2.58; minimum width—1.63. Elytra: maximum length—4.95; maximum width—3.15. Pygidium: maximum length—0.80; maximum width— 1.00.
</paragraph>
<paragraph id="8B9636B5C032FF8325D619FBFA8D8905" blockId="7.[151,1437,295,1978]" pageId="7" pageNumber="298">
<emphasis id="B95DEAA7C032FF8325D619FBFE878921" bold="true" box="[199,328,1015,1041]" pageId="7" pageNumber="298">Coloration</emphasis>
of the cuticle of entire body black, lower side of the body nearly black. Mandibles dark brown. Basal part of head, pronotum and elytra black. All antennomeres and labrum light brown, legs dark brown.
</paragraph>
<paragraph id="8B9636B5C032FF8325D61E33FA4688D6" blockId="7.[151,1437,295,1978]" pageId="7" pageNumber="298">
<emphasis id="B95DEAA7C032FF8325D61E33FEF58969" bold="true" box="[199,314,1087,1113]" pageId="7" pageNumber="298">Vestiture.</emphasis>
Head nearly naked, only vertex with two distinct stripes behind lateral carinae of rostrum and continued to inner margin of eyes, these stripes dark yellow with very dense appressed setae. Lateral sides of rostrum almost bare. Labrum light brown with a few longer light setae. Antennae with indistinct brown pubescence, only with very short, soft, setae. Pronotum with three distinct longitudinal dark yellow stripes posteriorly extended to transverse pronotal carina, the same on pronotal declivity. Each elytron with four longitudinal stripes: one short in scutellar area; one in the second elytral interval reaching from posterior part of sub-basal tubercle to apex of elytron; one on the fourth reaching from anterior margin of elytron to its half; and one on the eighth elytral interval. Stripes of the second and eighth elytral interval converge at the apex of elytron. Rest of elytra completely black. Scutellum with dense dark yellow setae. All legs covered with dense fine brown setae. Tibiae covered with dense indistinct brown setae appressed in proximal part and subdecumbent in distal part. All tibiae on dorsal side with spot of yellowish setae in proximal third. Tarsomeres with brownish setae. Lower side of the body covered with only indistinct dark-coloured fine setae. Pygidium with distinct longitudinal stripe of dark yellow setae in central part.
</paragraph>
<paragraph id="8B9636B5C032FF8325D61FE3FEE38A19" blockId="7.[151,1437,295,1978]" pageId="7" pageNumber="298">
<emphasis id="B95DEAA7C032FF8325D61FE3FE8E8B39" bold="true" box="[199,321,1519,1545]" pageId="7" pageNumber="298">Structure.</emphasis>
Head (
<figureCitation id="13122A30C032FF8324851FE3FE288B39" box="[404,487,1519,1545]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[178,1408,181,1912]" captionTargetId="figure-17@8.[178,1408,181,1918]" captionTargetPageId="8" captionText="FIGURES 1620. Diastatotropis petulae, holotype male; 16, habitus in dorsal view; 17, head; 18, pronotum; 19, left antenna; 20, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033755" httpUri="https://zenodo.org/record/14033755/files/figure.png" pageId="7" pageNumber="298">Fig. 17</figureCitation>
) rather shorter, rostrum wider than long, ratio of rostrum length to maximum width 0.90. Rostrum flat in lateral view. Lateral sides of rostrum weakly extended apically in dorsal view. Rostrum with two distinct longitudinal lateral carinae, lacking central carina, but between lateral carinae with longitudinal flat shiny area imitating central carina, this area short and indistinct. Lateral carinae not extended distal part of rostrum and reaching anterior margin of eyes. Dorsum of rostrum with only shallow depression. Dorsal part of head coarsely sculptured. Eyes oval, not emarginate, dorsal ocular index 1.51. Ratio of maximum width across eyes to maximum width of rostrum 1.29. Antennae (
<figureCitation id="13122A30C032FF8327001CCBFDAD8BD1" box="[529,610,1735,1761]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[178,1408,181,1912]" captionTargetId="figure-17@8.[178,1408,181,1918]" captionTargetPageId="8" captionText="FIGURES 1620. Diastatotropis petulae, holotype male; 16, habitus in dorsal view; 17, head; 18, pronotum; 19, left antenna; 20, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033755" httpUri="https://zenodo.org/record/14033755/files/figure.png" pageId="7" pageNumber="298">Fig. 19</figureCitation>
) shorter, not reaching to posterior margin of pronotum. Scape and pedicel approximately twice as thick as the funicle, club widened, antennomeres IX and XI ca. 2x longer than wide, X as long as wide.
</paragraph>
<paragraph id="8B9636B5C032FF8325D61D38FBC18A89" blockId="7.[151,1437,295,1978]" pageId="7" pageNumber="298">
Pronotum (
<figureCitation id="13122A30C032FF8324521D3FFE5C8A7D" box="[323,403,1843,1869]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[178,1408,181,1912]" captionTargetId="figure-17@8.[178,1408,181,1918]" captionTargetPageId="8" captionText="FIGURES 1620. Diastatotropis petulae, holotype male; 16, habitus in dorsal view; 17, head; 18, pronotum; 19, left antenna; 20, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033755" httpUri="https://zenodo.org/record/14033755/files/figure.png" pageId="7" pageNumber="298">Fig. 18</figureCitation>
) slightly transverse, ratio of its length to maximum width 0.83, the widest in dorsal transverse carina. Disc of pronotum convex, without any imprint. Dorsal transverse carina of pronotum sinuated, not interrupted in middle (in
<typeStatus id="54928817C032FF8324211D77FE458AA5" box="[304,394,1915,1941]" pageId="7" pageNumber="298" type="allotype">allotype</typeStatus>
only indistinctly interrupted). Lateral carinae of pronotum well developed, extending to half of length of pronotum, widely rounded at contact with dorsal transverse carina.
</paragraph>
<caption id="DF56663DC03DFF8C25861D9BFD908AE3" ID-DOI="http://doi.org/10.5281/zenodo.14033755" ID-Zenodo-Dep="14033755" httpUri="https://zenodo.org/record/14033755/files/figure.png" pageId="8" pageNumber="299" startId="8.[151,264,1943,1967]" targetBox="[178,1408,181,1912]" targetPageId="8" targetType="figure">
<paragraph id="8B9636B5C03DFF8C25861D9BFD908AE3" blockId="8.[151,1436,1943,2003]" pageId="8" pageNumber="299">
<emphasis id="B95DEAA7C03DFF8C25861D9BFE9A8A9F" bold="true" box="[151,341,1943,1967]" pageId="8" pageNumber="299">FIGURES 1620.</emphasis>
<taxonomicName id="4C294D36C03DFF8C244A1D94FDF18A9F" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[347,574,1943,1967]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="299" phylum="Arthropoda" rank="species" species="petulae">
<emphasis id="B95DEAA7C03DFF8C244A1D94FDF18A9F" box="[347,574,1943,1967]" italics="true" pageId="8" pageNumber="299">Diastatotropis petulae</emphasis>
</taxonomicName>
, holotype male; 16, habitus in dorsal view; 17, head; 18, pronotum; 19, left antenna; 20, habitus in lateral view. Scale bars in mm.
</paragraph>
</caption>
<caption id="DF56663DC03CFF8D25861D9BFC6B8AC7" ID-DOI="http://doi.org/10.5281/zenodo.14033757" ID-Zenodo-Dep="14033757" httpUri="https://zenodo.org/record/14033757/files/figure.png" pageId="9" pageNumber="300" startId="9.[151,264,1943,1967]" targetBox="[178,1408,182,1918]" targetPageId="9" targetType="figure">
<paragraph id="8B9636B5C03CFF8D25861D9BFC6B8AC7" blockId="9.[151,1436,1943,2039]" pageId="9" pageNumber="300">
<emphasis id="B95DEAA7C03CFF8D25861D9BFE9C8A9F" bold="true" box="[151,339,1943,1967]" pageId="9" pageNumber="300">FIGURES 2125.</emphasis>
<taxonomicName id="4C294D36C03CFF8D24491D94FDF58A9F" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[344,570,1943,1967]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="300" phylum="Arthropoda" rank="species" species="petulae">
<emphasis id="B95DEAA7C03CFF8D24491D94FDF58A9F" box="[344,570,1943,1967]" italics="true" pageId="9" pageNumber="300">Diastatotropis petulae</emphasis>
</taxonomicName>
, 2124, paratype male, terminalia; 21, aedeagus in lateral view; 22, aedeagus in dorsal view; 23, tegmen in ventral view; 24, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm. 25, Ambohitantely Special Reserve, overall view of collecting site of
<taxonomicName id="4C294D36C03CFF8D26221DECFC6F8AC7" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[819,928,2015,2039]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="300" phylum="Arthropoda" rank="species" species="petulae">D. petulae</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF56663DC03FFF8E25861D9BFB2E8A9F" ID-DOI="http://doi.org/10.5281/zenodo.14033759" ID-Zenodo-Dep="14033759" box="[151,1249,1943,1967]" httpUri="https://zenodo.org/record/14033759/files/figure.png" pageId="10" pageNumber="301" startId="10.[151,264,1943,1967]" targetBox="[178,1408,181,1918]" targetPageId="10" targetType="figure">
<paragraph id="8B9636B5C03FFF8E25861D9BFB2E8A9F" blockId="10.[151,1249,1943,1967]" box="[151,1249,1943,1967]" pageId="10" pageNumber="301">
<emphasis id="B95DEAA7C03FFF8E25861D9BFE9B8A9F" bold="true" box="[151,340,1943,1967]" pageId="10" pageNumber="301">FIGURES 2627.</emphasis>
Ambohitantely Special Reserve, microhabitat of collecting site of
<taxonomicName id="4C294D36C03FFF8E26E61D94FB128A9F" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[1015,1245,1943,1967]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="301" phylum="Arthropoda" rank="species" species="petulae">Diastatotropis petulae</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF56663DC03EFF8F25861D9BFC8E8A9F" ID-DOI="http://doi.org/10.5281/zenodo.14033761" ID-Zenodo-Dep="14033761" box="[151,833,1943,1968]" httpUri="https://zenodo.org/record/14033761/files/figure.png" pageId="11" pageNumber="302" startId="11.[151,250,1943,1967]" targetBox="[183,1406,188,1914]" targetPageId="11" targetType="figure">
<paragraph id="8B9636B5C03EFF8F25861D9BFC8E8A9F" blockId="11.[151,833,1943,1968]" box="[151,833,1943,1968]" pageId="11" pageNumber="302">
<emphasis id="B95DEAA7C03EFF8F25861D9BFEEF8A9F" bold="true" box="[151,288,1943,1967]" pageId="11" pageNumber="302">FIGURE 28.</emphasis>
Type localities of
<taxonomicName id="4C294D36C03EFF8F24CA1D94FD508A9F" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[475,671,1943,1967]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="302" phylum="Arthropoda" rank="species" species="lusti">
<emphasis id="B95DEAA7C03EFF8F24CA1D94FD508A9F" box="[475,671,1943,1967]" italics="true" pageId="11" pageNumber="302">Diastatotropis lusti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C294D36C03EFF8F27C11D94FCF28A9F" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[720,829,1943,1967]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="302" phylum="Arthropoda" rank="species" species="petulae">D. petulae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B9636B5C039FF8825D61A9BFDAB8DC9" blockId="12.[151,1437,151,1221]" pageId="12" pageNumber="303">
Elytra (
<figureCitation id="13122A30C039FF88240A1A9BFE548D81" box="[283,411,151,177]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[178,1408,181,1912]" captionTargetId="figure-17@8.[178,1408,181,1918]" captionTargetPageId="8" captionText="FIGURES 1620. Diastatotropis petulae, holotype male; 16, habitus in dorsal view; 17, head; 18, pronotum; 19, left antenna; 20, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033755" httpUri="https://zenodo.org/record/14033755/files/figure.png" pageId="12" pageNumber="303">Figs 16, 20</figureCitation>
) rather short, with indistinct sub-basal tubercles, ratio of the maximum length to maximum width of elytra 1.57. Elytral intervals completely flat, dorsal punctures distinct, their diameter approximately the same as distance of individual punctures.
</paragraph>
<paragraph id="8B9636B5C039FF8825D61B0FFACB8C2D" blockId="12.[151,1437,151,1221]" box="[199,1284,259,285]" pageId="12" pageNumber="303">Claws without teeth. Pygidium transverse, ratio of the maximum length to maximum width 0.80.</paragraph>
<paragraph id="8B9636B5C039FF8825D61B2BFD088F29" blockId="12.[151,1437,151,1221]" pageId="12" pageNumber="303">
<emphasis id="B95DEAA7C039FF8825D61B2BFD678C71" bold="true" box="[199,680,294,321]" pageId="12" pageNumber="303">Male genitalia and associated structures</emphasis>
(
<figureCitation id="13122A30C039FF8827A61B2BFCF58C71" box="[695,826,295,321]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[178,1408,182,1918]" captionTargetId="figure-17@9.[178,1408,181,1918]" captionTargetPageId="9" captionText="FIGURES 2125. Diastatotropis petulae, 2124, paratype male, terminalia; 21, aedeagus in lateral view; 22, aedeagus in dorsal view; 23, tegmen in ventral view; 24, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm. 25, Ambohitantely Special Reserve, overall view of collecting site of D. petulae." figureDoi="http://doi.org/10.5281/zenodo.14033757" httpUri="https://zenodo.org/record/14033757/files/figure.png" pageId="12" pageNumber="303">Figs 2124</figureCitation>
). Aedeagus (
<figureCitation id="13122A30C039FF8826DB1B2BFB858C71" box="[970,1098,295,321]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[178,1408,182,1918]" captionTargetId="figure-17@9.[178,1408,181,1918]" captionTargetPageId="9" captionText="FIGURES 2125. Diastatotropis petulae, 2124, paratype male, terminalia; 21, aedeagus in lateral view; 22, aedeagus in dorsal view; 23, tegmen in ventral view; 24, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm. 25, Ambohitantely Special Reserve, overall view of collecting site of D. petulae." figureDoi="http://doi.org/10.5281/zenodo.14033757" httpUri="https://zenodo.org/record/14033757/files/figure.png" pageId="12" pageNumber="303">Figs 2122</figureCitation>
) rather short and wide. Pedon and tectum wide. Tectum inconspicuously overlapping pedon, with pointed projection apically. Apodemes of aedeagus long, parallel. Bridge of aedeagus well developed. Tegmen (
<figureCitation id="13122A30C039FF88268E1B63FC208CB9" box="[927,1007,367,393]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[178,1408,182,1918]" captionTargetId="figure-17@9.[178,1408,181,1918]" captionTargetPageId="9" captionText="FIGURES 2125. Diastatotropis petulae, 2124, paratype male, terminalia; 21, aedeagus in lateral view; 22, aedeagus in dorsal view; 23, tegmen in ventral view; 24, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm. 25, Ambohitantely Special Reserve, overall view of collecting site of D. petulae." figureDoi="http://doi.org/10.5281/zenodo.14033757" httpUri="https://zenodo.org/record/14033757/files/figure.png" pageId="12" pageNumber="303">Fig. 23</figureCitation>
) rather long and slender, with strongly sclerotized basal piece, apodeme of tegmen straight, slightly shorter than body of tegmen. Tegminal plate rather slender with exception of basal part. Apex of tegmen broad, with distinct longer setae. Segment VIII (
<figureCitation id="13122A30C039FF88202E1BBBFA5B8CE1" box="[1343,1428,439,465]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[178,1408,182,1918]" captionTargetId="figure-17@9.[178,1408,181,1918]" captionTargetPageId="9" captionText="FIGURES 2125. Diastatotropis petulae, 2124, paratype male, terminalia; 21, aedeagus in lateral view; 22, aedeagus in dorsal view; 23, tegmen in ventral view; 24, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm. 25, Ambohitantely Special Reserve, overall view of collecting site of D. petulae." figureDoi="http://doi.org/10.5281/zenodo.14033757" httpUri="https://zenodo.org/record/14033757/files/figure.png" pageId="12" pageNumber="303">Fig. 24</figureCitation>
) moderately sclerotized, approximately as long as wide, anterior margin of tergite rounded. Sternite IX (
<figureCitation id="13122A30C039FF88201D1BD7FA938CC5" box="[1292,1372,475,501]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[178,1408,182,1918]" captionTargetId="figure-17@9.[178,1408,181,1918]" captionTargetPageId="9" captionText="FIGURES 2125. Diastatotropis petulae, 2124, paratype male, terminalia; 21, aedeagus in lateral view; 22, aedeagus in dorsal view; 23, tegmen in ventral view; 24, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm. 25, Ambohitantely Special Reserve, overall view of collecting site of D. petulae." figureDoi="http://doi.org/10.5281/zenodo.14033757" httpUri="https://zenodo.org/record/14033757/files/figure.png" pageId="12" pageNumber="303">Fig. 24</figureCitation>
) with symmetrical short arms, apodeme slightly curved.
</paragraph>
</subSubSection>
<subSubSection id="C333653EC039FF8825D6182FFCB68E95" pageId="12" pageNumber="303" type="etymology">
<paragraph id="8B9636B5C039FF8825D6182FFE958F51" blockId="12.[151,1437,151,1221]" pageId="12" pageNumber="303">
<emphasis id="B95DEAA7C039FF8825D6182FFE818F0D" bold="true" box="[199,334,547,573]" pageId="12" pageNumber="303">Etymology.</emphasis>
Matronym, dedicated to the wife of the second author, Petra (tenderly Peťula) Blažejová (Varnsdorf,
<collectingCountry id="F33E7625C039FF882586184BFE818F51" box="[151,334,583,609]" name="Czech Republic" pageId="12" pageNumber="303">Czech Republic</collectingCountry>
).
</paragraph>
<paragraph id="8B9636B5C039FF8825D61866FC748E25" blockId="12.[151,1437,151,1221]" pageId="12" pageNumber="303">
<emphasis id="B95DEAA7C039FF8825D61866FE3E8FB4" bold="true" box="[199,497,618,644]" pageId="12" pageNumber="303">Collecting circumstances</emphasis>
(
<figureCitation id="13122A30C039FF8827111867FD4B8FB5" box="[512,644,619,645]" captionStart="FIGURES 2627" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-17@10.[178,1408,181,1918]" captionTargetPageId="10" captionText="FIGURES 2627. Ambohitantely Special Reserve, microhabitat of collecting site of Diastatotropis petulae." figureDoi="http://doi.org/10.5281/zenodo.14033759" httpUri="https://zenodo.org/record/14033759/files/figure.png" pageId="12" pageNumber="303">Figs 2627</figureCitation>
).
<taxonomicName id="4C294D36C039FF8827891867FC5D8FB5" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[664,914,619,645]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="303" phylum="Arthropoda" rank="species" species="petulae">
<emphasis id="B95DEAA7C039FF8827891867FC5D8FB5" box="[664,914,619,645]" italics="true" pageId="12" pageNumber="303">Diastatotropis petulae</emphasis>
</taxonomicName>
was found in Ambohitantely Special Reserve, north-west of the town of
<collectingRegion id="49EDF857C039FF8824AB1883FD9D8F99" box="[442,594,655,681]" country="Madagascar" name="Antananarivo" pageId="12" pageNumber="303">Antananarivo</collectingRegion>
in the district of Ankazobe (Analamanga region, central
<collectingCountry id="F33E7625C039FF8821C31883FA908F99" box="[1234,1375,655,681]" name="Madagascar" pageId="12" pageNumber="303">Madagascar</collectingCountry>
). The reserve is characterized by a primary rainforest, surrounded by grassland savannas. The site of the new species discovery represents one of the last primary rainforest patches on the high plateau in central
<collectingCountry id="F33E7625C039FF8821CB18DBFAAC8FC1" box="[1242,1379,727,753]" name="Madagascar" pageId="12" pageNumber="303">Madagascar</collectingCountry>
. The detailed description of the area is presented by
<bibRefCitation id="EFB84B44C039FF8827B718F7FC788E25" author="Goodman, S. M. &amp; Raherilalao, M. J. &amp; Wohlhauser, S. &amp; Tome III" box="[678,951,763,789]" pageId="12" pageNumber="303" pagination="1233 - 1716" refId="ref5435" refString="Goodman, S. M., Raherilalao, M. J. &amp; Wohlhauser, S. (Eds.) (2018 b) s. n. In: Les aires protegees terrestres de Madagascar: Leur histoire, description et biote. Tome III: L´ouest et le sud de Madagascar - synthese / The terrestrial protected areas of Madagascar: Their history, description, and biota. Vol. III. Western and southwestern Madagascar - synthesis. Association Vahatra, Antananarivo, pp. 1233 - 1716. [the correct surname of the third editor is Wohlhauser, not Wohlauser as given on the book cover]" type="book chapter" year="2018">
Goodman
<emphasis id="B95DEAA7C039FF88260D18F7FC9B8E25" box="[796,852,763,789]" italics="true" pageId="12" pageNumber="303">et al.</emphasis>
(2018b)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B9636B5C039FF8825D61913FCB68E95" blockId="12.[151,1437,151,1221]" pageId="12" pageNumber="303">
All specimens of the
<typeStatus id="54928817C039FF8824AB1913FE258E09" box="[442,490,799,825]" pageId="12" pageNumber="303">type</typeStatus>
series were collected inside low primary forest with a canopy of 6 to 12 (max. 20) metres and one or two strata, with dense coppices of small diameter trees in elevation of
<quantity id="4CD19B50C039FF88219F194FFB2A8E6D" box="[1166,1253,835,861]" metricMagnitude="3" metricUnit="m" metricValue="1.623" pageId="12" pageNumber="303" unit="m" value="1623.0">1623 m</quantity>
a.s.l. The dying wood was already infected with an unspecified fungus. All
<specimenCount id="9D2FFD3CC039FF88263E196BFC168EB1" box="[815,985,871,897]" count="4" pageId="12" pageNumber="303" type="generic">four specimens</specimenCount>
were collected during one day, repeated collections on subsequent days did not yield any more specimens.
</paragraph>
</subSubSection>
<subSubSection id="C333653EC039FF8825D619A2FDA58EDD" pageId="12" pageNumber="303" type="distribution">
<paragraph id="8B9636B5C039FF8825D619A2FDA58EDD" blockId="12.[151,1437,151,1221]" pageId="12" pageNumber="303">
<emphasis id="B95DEAA7C039FF8825D619A2FEAE8EF8" bold="true" box="[199,353,942,968]" pageId="12" pageNumber="303">Distribution.</emphasis>
C
<collectingCountry id="F33E7625C039FF88249E19A3FDD78EF9" box="[399,536,943,969]" name="Madagascar" pageId="12" pageNumber="303">Madagascar</collectingCountry>
,
<collectingRegion id="49EDF857C039FF88273919A3FCFB8EF9" box="[552,820,943,969]" country="Madagascar" name="Antananarivo" pageId="12" pageNumber="303">Antananarivo province</collectingRegion>
. The species is only known from the
<typeStatus id="54928817C039FF88201B19A3FAF58EF9" box="[1290,1338,943,969]" pageId="12" pageNumber="303">type</typeStatus>
locality Ambohitantely Special Reserve (
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).
</paragraph>
</subSubSection>
<subSubSection id="C333653EC039FF8825D619FAFEFE89F5" pageId="12" pageNumber="303" type="diagnosis">
<paragraph id="8B9636B5C039FF8825D619FAFEFE89F5" blockId="12.[151,1437,151,1221]" pageId="12" pageNumber="303">
<emphasis id="B95DEAA7C039FF8825D619FAFE1F8920" bold="true" box="[199,464,1014,1040]" pageId="12" pageNumber="303">Differential diagnosis.</emphasis>
<taxonomicName id="4C294D36C039FF8824C819FBFD1A8921" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[473,725,1015,1041]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="303" phylum="Arthropoda" rank="species" species="petulae">
<emphasis id="B95DEAA7C039FF8824C819FBFD1A8921" box="[473,725,1015,1041]" italics="true" pageId="12" pageNumber="303">Diastatotropis petulae</emphasis>
</taxonomicName>
also belongs to a group of species with antennal club slender and cylindrical, antennomere X as long as wide, and terminal segment elongate-oval, always distinctly longer than wide. Rostrum with two lateral carinae, but between them longitudinal flat shiny area resembling central carina. Longitudinal stripes on the elytra prominent. The species characterized by well-developed lateral carinae of pronotum extending to half of length of pronotum. Pygidium with distinct longitudinal stripe of light-coloured setae in both sexes.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,338 @@
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<mods:title id="32BB394DCAD8D0261D50831B1125F53A">Two new species of Diastatotropis Lacordaire from protected areas in central and east Madagascar (Coleoptera: Anthribidae)</mods:title>
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<taxonomicName id="4C294D36C034FF8525861CCBFCC68BD0" authority="Tryzna, Blazej &amp; Rakotonirina" authorityName="Tryzna, Blazej &amp; Rakotonirina" authorityYear="2024" box="[151,777,1734,1761]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="lusti" status="sp. nov.">
<emphasis id="B95DEAA7C034FF8525861CCBFEB38BD1" bold="true" box="[151,380,1734,1761]" italics="true" pageId="1" pageNumber="292">Diastatotropis lusti</emphasis>
Trýzna, Blažej &amp; Rakotonirina
</taxonomicName>
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http://zoobank.org/
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(
<figureCitation id="13122A30C034FF85258E1D03FEDF8A19" box="[159,272,1807,1833]" captionStart-0="FIGURES 15" captionStart-1="FIGURES 69" captionStart-2="FIGURES 1013" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1274,1298]" captionStartId-2="5.[151,264,1206,1230]" captionTargetBox-0="[179,1408,191,1914]" captionTargetBox-1="[157,1429,181,1246]" captionTargetBox-2="[151,1433,182,1178]" captionTargetId-0="figure-17@2.[178,1408,181,1918]" captionTargetId-1="figure-17@3.[151,1435,181,1250]" captionTargetId-2="figure-17@5.[151,1436,181,1183]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURES 15. Diastatotropis lusti, holotype male; 1, habitus in dorsal view; 2, head; 3, pronotum; 4, right antenna; 5, habitus in lateral view. Scale bars in mm." captionText-1="FIGURES 69. Diastatotropis lusti, paratype male, terminalia; 6, aedeagus in dorsal view; 7, aedeagus in lateral view; 8, tegmen in ventral view; 9, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm." captionText-2="FIGURES 1013. Diastatotropis lusti, allotype female, terminalia; 10, ovipositor in ventral view; 11, apex of ovipositor (hemisternites) in ventral view; 12, sternite VIII (left) and tergite VIII (right) in ventral view; 13, spermatheca. Scale bars in mm." figureDoi-0="http://doi.org/10.5281/zenodo.14033741" figureDoi-1="http://doi.org/10.5281/zenodo.14033745" figureDoi-2="http://doi.org/10.5281/zenodo.14033747" httpUri-0="https://zenodo.org/record/14033741/files/figure.png" httpUri-1="https://zenodo.org/record/14033745/files/figure.png" httpUri-2="https://zenodo.org/record/14033747/files/figure.png" pageId="1" pageNumber="292">Figs 113</figureCitation>
)
</paragraph>
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<paragraph id="8B9636B5C034FF8525861D5BFE0A8AA5" blockId="1.[151,1436,1878,1941]" pageId="1" pageNumber="292">
<materialsCitation id="3B413CE8C034FF8525861D5BFB0A8A41" box="[151,1221,1878,1905]" country="Madagascar" location="Ranomafana National Park" pageId="1" pageNumber="292" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B95DEAA7C034FF8525861D5BFEF88A40" bold="true" box="[151,311,1878,1905]" pageId="1" pageNumber="292">
<typeStatus id="54928817C034FF8525861D5BFF1D8A41" box="[151,210,1879,1905]" pageId="1" pageNumber="292">Type</typeStatus>
locality.
</emphasis>
E
<collectingCountry id="F33E7625C034FF85244B1D5BFE2C8A41" box="[346,483,1879,1905]" name="Madagascar" pageId="1" pageNumber="292">Madagascar</collectingCountry>
,
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, natural forest in
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</materialsCitation>
,
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;
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,
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a.s.l.
</paragraph>
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<paragraph id="8B9636B5C037FF8625861D9BFE258AE3" blockId="2.[151,1436,1943,2003]" pageId="2" pageNumber="293">
<emphasis id="B95DEAA7C037FF8625861D9BFEF58A9F" bold="true" box="[151,314,1943,1967]" pageId="2" pageNumber="293">FIGURES 15.</emphasis>
<taxonomicName id="4C294D36C037FF8624511D94FDCB8A9F" authorityName="Tryzna, Blazej &amp; Rakotonirina" authorityYear="2024" box="[320,516,1943,1967]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="lusti">
<emphasis id="B95DEAA7C037FF8624511D94FDCB8A9F" box="[320,516,1943,1967]" italics="true" pageId="2" pageNumber="293">Diastatotropis lusti</emphasis>
</taxonomicName>
, holotype male; 1, habitus in dorsal view; 2, head; 3, pronotum; 4, right antenna; 5, habitus in lateral view. Scale bars in mm.
</paragraph>
</caption>
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<emphasis id="B95DEAA7C036FF8725861EF6FEF18822" bold="true" box="[151,318,1274,1298]" pageId="3" pageNumber="294">FIGURES 69.</emphasis>
<taxonomicName id="4C294D36C036FF8724561EF7FDC18822" authorityName="Tryzna, Blazej &amp; Rakotonirina" authorityYear="2024" box="[327,526,1274,1298]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="lusti">
<emphasis id="B95DEAA7C036FF8724561EF7FDC18822" box="[327,526,1274,1298]" italics="true" pageId="3" pageNumber="294">Diastatotropis lusti</emphasis>
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, paratype male, terminalia; 6, aedeagus in dorsal view; 7, aedeagus in lateral view; 8, tegmen in ventral view; 9, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm.
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Type material.
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(male)
</emphasis>
(
<figureCitation id="13122A30C036FF8727721F85FD008893" box="[611,719,1416,1443]" captionStart="FIGURES 15" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[179,1408,191,1914]" captionTargetId="figure-17@2.[178,1408,181,1918]" captionTargetPageId="2" captionText="FIGURES 15. Diastatotropis lusti, holotype male; 1, habitus in dorsal view; 2, head; 3, pronotum; 4, right antenna; 5, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033741" httpUri="https://zenodo.org/record/14033741/files/figure.png" pageId="3" pageNumber="294">Figs 15</figureCitation>
): E
<collectingCountry id="F33E7625C036FF8726031F85FC138893" box="[786,988,1417,1443]" name="Madagascar" pageId="3" pageNumber="294">MADAGASCAR</collectingCountry>
,
<date id="FF971075C036FF8726FC1F84FBAE8892" box="[1005,1121,1416,1442]" pageId="3" pageNumber="294" value="2010-02-16">16.ii.2010</date>
, / Ranomafana N.P.,
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, /
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;
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, / M. Trýzna leg. [p] (
<collectionCode id="ED38AE70C036FF87264D1FA1FC7588F7" box="[860,954,1453,1479]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="294">MMBC</collectionCode>
).
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<typeStatus id="54928817C036FF8726C11FA0FBFA88F6" box="[976,1077,1452,1478]" pageId="3" pageNumber="294" type="allotype">Allotype</typeStatus>
(female):
</emphasis>
<collectingCountry id="F33E7625C036FF8721AD1FA1FA4788F7" box="[1212,1416,1453,1479]" name="Madagascar" pageId="3" pageNumber="294">MADAGASCAR</collectingCountry>
/
<collectingRegion id="49EDF857C036FF8725861FDDFEA988DB" box="[151,358,1489,1515]" country="Madagascar" name="Fianarantsoa" pageId="3" pageNumber="294">Fianarantsoa prov.</collectingRegion>
/ Ranomafana env.
<emphasis id="B95DEAA7C036FF87275F1FDCFD9988DA" bold="true" box="[590,598,1488,1514]" pageId="3" pageNumber="294">/</emphasis>
28.1.-
<date id="FF971075C036FF87278A1FDCFCCD88DB" box="[667,770,1488,1515]" pageId="3" pageNumber="294" value="1995-02-06">6.2.1995</date>
/ Ivo Jeniš leg. [p] (
<collectionCode id="ED38AE70C036FF8726E91FDDFB8088DB" box="[1016,1103,1489,1515]" pageId="3" pageNumber="294">MTDC</collectionCode>
).
<emphasis id="B95DEAA7C036FF8721751FDDFAE988DA" bold="true" box="[1124,1318,1488,1515]" pageId="3" pageNumber="294">
<typeStatus id="54928817C036FF8721751FDDFB1E88DB" box="[1124,1233,1489,1515]" pageId="3" pageNumber="294" type="paratype">Paratype</typeStatus>
(male)
</emphasis>
: the same data as holotype (
<collectionCode id="ED38AE70C036FF8724731FF9FE768B3F" box="[354,441,1525,1551]" pageId="3" pageNumber="294">MTDC</collectionCode>
). Red label [p]
<emphasis id="B95DEAA7C036FF87277D1FF8FE148B02" bold="true" pageId="3" pageNumber="294">
<typeStatus id="54928817C036FF87277D1FF8FCC48B3E" box="[620,779,1524,1550]" pageId="3" pageNumber="294" type="holotype">HOLOTYPE</typeStatus>
/
<typeStatus id="54928817C036FF8726321FF8FC728B3E" box="[803,957,1524,1550]" pageId="3" pageNumber="294" type="allotype">ALLOTYPE</typeStatus>
/
<typeStatus id="54928817C036FF8726C61FF8FBA48B3E" box="[983,1131,1524,1550]" pageId="3" pageNumber="294" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C294D36C036FF8721941FF8FAE88B3E" authorityName="Lacordaire" authorityYear="1866" box="[1157,1319,1524,1550]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA7C036FF8721941FF8FAE88B3E" bold="true" box="[1157,1319,1524,1550]" italics="true" pageId="3" pageNumber="294">Diastatotropis</emphasis>
</taxonomicName>
/
<taxonomicName id="4C294D36C036FF8720511FF8FABD8B3F" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[1344,1394,1524,1551]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="lusti" status="sp. nov.">
<emphasis id="B95DEAA7C036FF8720511FF8FABD8B3F" bold="true" box="[1344,1394,1524,1551]" italics="true" pageId="3" pageNumber="294">lusti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57DCC036FF87206A1FF9FF078B03" pageId="3" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
/ M. Trýzna det., 2024.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C333653EC036FF8725D61C30FDFD8A3B" pageId="3" pageNumber="294" type="diagnosis">
<paragraph id="8B9636B5C036FF8725D61C30FDFD8A3B" blockId="3.[151,1437,1416,2019]" pageId="3" pageNumber="294">
<emphasis id="B95DEAA7C036FF8725D61C30FE8D8B66" bold="true" box="[199,322,1596,1622]" pageId="3" pageNumber="294">Diagnosis.</emphasis>
Small-sized species (about 6.0 mm). Rostrum with only two distinct longitudinal carinae, lacking central carina. Dorsum of rostrum flat, with coarse sculpture. Antennal club slender and oblong, terminal segment elongate-oval, distinctly longer than wide. Elytra convex, without distinct sub-basal tubercles. Pronotum with one central and two lateral longitudinal yellow stripes; each elytron with one longitudinal yellow stripe interrupted in the central part. This stripe is in the basal half of the fourth elytral interval, in the apical half obliquely forming from the fourth to the first elytral interval.
</paragraph>
</subSubSection>
<subSubSection id="C333653EC036FF8025D61D18FE748A34" lastPageId="4" lastPageNumber="295" pageId="3" pageNumber="294" type="description">
<paragraph id="8B9636B5C036FF8725D61D18FAA78AD3" blockId="3.[151,1437,1416,2019]" pageId="3" pageNumber="294">
<emphasis id="B95DEAA7C036FF8725D61D18FDD58A1E" bold="true" box="[199,538,1812,1839]" pageId="3" pageNumber="294">
Description.
<typeStatus id="54928817C036FF87244C1D19FE058A1F" box="[349,458,1813,1839]" pageId="3" pageNumber="294" type="holotype">Holotype</typeStatus>
(male)
</emphasis>
(
<figureCitation id="13122A30C036FF8727371D19FD468A1F" box="[550,649,1812,1839]" captionStart="FIGURES 15" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[179,1408,191,1914]" captionTargetId="figure-17@2.[178,1408,181,1918]" captionTargetPageId="2" captionText="FIGURES 15. Diastatotropis lusti, holotype male; 1, habitus in dorsal view; 2, head; 3, pronotum; 4, right antenna; 5, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033741" httpUri="https://zenodo.org/record/14033741/files/figure.png" pageId="3" pageNumber="294">Figs 15</figureCitation>
).
<emphasis id="B95DEAA7C036FF87278A1D19FCF28A1E" box="[667,829,1813,1838]" italics="true" pageId="3" pageNumber="294">Measurements</emphasis>
(in mm):Total body length—6.05.Head:total length— 1.30; length of rostrum—0.75; maximum width of rostrum—0.98; length of eye—0.65; maximum width across eyes—1.28; minimum distance between eyes—0.55. Antenna: length of segments: II—0.20, III—0.21, IV—0.19, V—0.14, VI—0.13, VII—0.10, VIII—0.09, IX—0.30, X—0.16, XI—0.31, width of segment IX—0.14. Pronotum: maximum length—1.33; width at carina (= maximal width in this case)—1.85; minimum width—1.13. Elytra: maximum length—3.25; maximum width—2.05. Pygidium: maximum length—0.50; maximum width—0.74.
</paragraph>
<paragraph id="8B9636B5C031FF8025D61A9AFE868DC8" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="295">
<emphasis id="B95DEAA7C031FF8025D61A9AFE878D80" bold="true" box="[199,328,150,176]" pageId="4" pageNumber="295">Coloration</emphasis>
of the cuticle of entire body dark, nearly black, rostrum including scrobes and mandibles, and abdominal ventrites brown. Basal part of head, pronotum and elytra black. All antennomeres and legs dark brown. Labrum yellow.
</paragraph>
<paragraph id="8B9636B5C031FF8025D61B0EFD2B8E25" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="295">
<emphasis id="B95DEAA7C031FF8025D61B0EFEF58C2C" bold="true" box="[199,314,258,284]" pageId="4" pageNumber="295">Vestiture.</emphasis>
Head with only indistinct sparse yellowish appressed setae in central part of rostrum. Frons with only very short and almost invisible setae in central part. Labrum yellowish with a few longer light setae. Vertex with two stripes behind lateral carinae of rostrum and continued to inner margin of eyes, these stripes yellow with very dense appressed setae. Lateral sides of rostrum almost bare, with yellow stripes under lower edge of eyes. Antennae with indistinct light pubescence, only with very short, soft, setae. Pronotum with one slender and two wider distinct longitudinal stripes posteriorly extended to transverse pronotal carina, the same on pronotal declivity. All stripes covered with dense yellow setae. Each elytron with one longitudinal yellow stripe interrupted in the central part. This stripe is in basal half on the fourth elytral interval, in the apical half obliquely forming from the fourth to the first elytral interval. Next yellow stripe on ninth elytral interval, also widely interrupted in middle. Rest of elytra, mainly disc and lateral part, with only fine and sparse yellow pubescence. Humeral angle lacking yellow colouration, with only sparse short yellow setae. Scutellum with dense yellow setae. All legs covered with dense fine brownish setae. Tibiae covered with dense indistinct brown setae appressed in proximal part and subdecumbent in distal part. All tibiae on dorsal side with only indistinct small spot of light-coloured setae in proximal third. Tarsomeres with brownish setae. Lower side of the body with sparse pubescence of short yellow setae. Pygidium covered with only short and indistinct dense yellowish appressed setae.
</paragraph>
<paragraph id="8B9636B5C031FF8025D61912FD828904" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="295">
<emphasis id="B95DEAA7C031FF8025D61912FE8E8E08" bold="true" box="[199,321,798,824]" pageId="4" pageNumber="295">Structure.</emphasis>
Head (
<figureCitation id="13122A30C031FF8024871912FE148E08" box="[406,475,798,824]" captionStart="FIGURES 15" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[179,1408,191,1914]" captionTargetId="figure-17@2.[178,1408,181,1918]" captionTargetPageId="2" captionText="FIGURES 15. Diastatotropis lusti, holotype male; 1, habitus in dorsal view; 2, head; 3, pronotum; 4, right antenna; 5, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033741" httpUri="https://zenodo.org/record/14033741/files/figure.png" pageId="4" pageNumber="295">Fig. 2</figureCitation>
) rather shorter, rostrum wider than long, ratio of rostrum length to maximum width 0.77. Rostrum almost flat in lateral view. Lateral sides of rostrum weakly extended apically. Rostrum with only two lateral longitudinal carinae, lacking central carina. Lateral carinae not extended distal part of rostrum and reaching anterior margin of eyes. Dorsum of rostrum with only shallow depression. Whole dorsal part of head coarsely sculptured. Eyes oval, not emarginate, dorsal ocular index 1.51. Ratio of maximum width across eyes to maximum width of rostrum 1.31. Antennae (
<figureCitation id="13122A30C031FF80270919DEFDB18EDC" box="[536,638,978,1004]" captionStart="FIGURES 15" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[179,1408,191,1914]" captionTargetId="figure-17@2.[178,1408,181,1918]" captionTargetPageId="2" captionText="FIGURES 15. Diastatotropis lusti, holotype male; 1, habitus in dorsal view; 2, head; 3, pronotum; 4, right antenna; 5, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033741" httpUri="https://zenodo.org/record/14033741/files/figure.png" pageId="4" pageNumber="295">Figs 1, 4</figureCitation>
) shorter, reaching only to dorsal transverse carina of pronotum. Scape and pedicel approximately twice as thick as the funicle, club widened, antennomeres IX and XI about twice longer than wide, X slightly longer than wide.
</paragraph>
<paragraph id="8B9636B5C031FF8025D61E33FB6789F4" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="295">
Pronotum (
<figureCitation id="13122A30C031FF8024551E32FE498968" box="[324,390,1086,1112]" captionStart="FIGURES 15" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[179,1408,191,1914]" captionTargetId="figure-17@2.[178,1408,181,1918]" captionTargetPageId="2" captionText="FIGURES 15. Diastatotropis lusti, holotype male; 1, habitus in dorsal view; 2, head; 3, pronotum; 4, right antenna; 5, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033741" httpUri="https://zenodo.org/record/14033741/files/figure.png" pageId="4" pageNumber="295">Fig. 3</figureCitation>
) slightly transverse, ratio of its length to maximum width 0.72, the widest in dorsal transverse carina, then narrowed to distal part. Disc of pronotum convex, without any imprint. Dorsal transverse carina of pronotum sinuated, not interrupted in middle. Lateral carinae of pronotum very short, not strongly indicated, not reaching half of length of pronotum, widely rounded at contact with dorsal transverse carina.
</paragraph>
<paragraph id="8B9636B5C031FF8025D61EC2FD848801" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="295">
Elytra (
<figureCitation id="13122A30C031FF80240A1EC2FEB189D9" box="[283,382,1230,1257]" captionStart="FIGURES 15" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[179,1408,191,1914]" captionTargetId="figure-17@2.[178,1408,181,1918]" captionTargetPageId="2" captionText="FIGURES 15. Diastatotropis lusti, holotype male; 1, habitus in dorsal view; 2, head; 3, pronotum; 4, right antenna; 5, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033741" httpUri="https://zenodo.org/record/14033741/files/figure.png" pageId="4" pageNumber="295">Figs 1, 5</figureCitation>
) rather short, without sub-basal tubercles, ratio of the maximum length to maximum width of elytra 1.59. Elytral intervals completely flat, dorsal punctures distinct, their diameter approximately the same as distance between individual punctures.
</paragraph>
<paragraph id="8B9636B5C031FF8025D61F36FACB8864" blockId="4.[151,1437,150,1977]" box="[199,1284,1338,1365]" pageId="4" pageNumber="295">Claws without teeth. Pygidium transverse, ratio of the maximum length to maximum width 0.68.</paragraph>
<paragraph id="8B9636B5C031FF8025D61F52FC068B60" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="295">
<emphasis id="B95DEAA7C031FF8025D61F52FD098848" bold="true" box="[199,710,1374,1400]" pageId="4" pageNumber="295">Genitalia and associated structures. Male</emphasis>
(
<figureCitation id="13122A30C031FF8027C81F52FC8B8848" box="[729,836,1374,1400]" captionStart="FIGURES 69" captionStartId="3.[151,264,1274,1298]" captionTargetBox="[157,1429,181,1246]" captionTargetId="figure-17@3.[151,1435,181,1250]" captionTargetPageId="3" captionText="FIGURES 69. Diastatotropis lusti, paratype male, terminalia; 6, aedeagus in dorsal view; 7, aedeagus in lateral view; 8, tegmen in ventral view; 9, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033745" httpUri="https://zenodo.org/record/14033745/files/figure.png" pageId="4" pageNumber="295">Figs 69</figureCitation>
). Aedeagus (
<figureCitation id="13122A30C031FF8026CD1F52FB8A8848" box="[988,1093,1374,1400]" captionStart="FIGURES 69" captionStartId="3.[151,264,1274,1298]" captionTargetBox="[157,1429,181,1246]" captionTargetId="figure-17@3.[151,1435,181,1250]" captionTargetPageId="3" captionText="FIGURES 69. Diastatotropis lusti, paratype male, terminalia; 6, aedeagus in dorsal view; 7, aedeagus in lateral view; 8, tegmen in ventral view; 9, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033745" httpUri="https://zenodo.org/record/14033745/files/figure.png" pageId="4" pageNumber="295">Figs 67</figureCitation>
) rather elongated, pedon and tectum slender. Tectum with pointed projection apically, pedon broadly rounded apically. Apodemes of aedeagus long, parallel. Bridge of aedeagus weakly developed. Tegmen (
<figureCitation id="13122A30C031FF80264A1FAAFC5188F0" box="[859,926,1446,1472]" captionStart="FIGURES 69" captionStartId="3.[151,264,1274,1298]" captionTargetBox="[157,1429,181,1246]" captionTargetId="figure-17@3.[151,1435,181,1250]" captionTargetPageId="3" captionText="FIGURES 69. Diastatotropis lusti, paratype male, terminalia; 6, aedeagus in dorsal view; 7, aedeagus in lateral view; 8, tegmen in ventral view; 9, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033745" httpUri="https://zenodo.org/record/14033745/files/figure.png" pageId="4" pageNumber="295">Fig. 8</figureCitation>
) rather long and slender, basal piece strongly sclerotized. Apodeme of tegmen straight, wide, distinctly shorter than body of tegmen. Tegminal plate slender, almost triangular, with narrow apical part. Apex of tegmen slender, with distinct longer setae. Segment VIII (
<figureCitation id="13122A30C031FF80207E1FE2FF698B1C" captionStart="FIGURES 69" captionStartId="3.[151,264,1274,1298]" captionTargetBox="[157,1429,181,1246]" captionTargetId="figure-17@3.[151,1435,181,1250]" captionTargetPageId="3" captionText="FIGURES 69. Diastatotropis lusti, paratype male, terminalia; 6, aedeagus in dorsal view; 7, aedeagus in lateral view; 8, tegmen in ventral view; 9, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033745" httpUri="https://zenodo.org/record/14033745/files/figure.png" pageId="4" pageNumber="295">Fig. 9</figureCitation>
) moderately sclerotized, approximately as long as wide, anterior margin of tergite widely rounded. Sternite IX (= spiculum gastrale,
<figureCitation id="13122A30C031FF80247B1C3AFE638B60" box="[362,428,1590,1616]" captionStart="FIGURES 69" captionStartId="3.[151,264,1274,1298]" captionTargetBox="[157,1429,181,1246]" captionTargetId="figure-17@3.[151,1435,181,1250]" captionTargetPageId="3" captionText="FIGURES 69. Diastatotropis lusti, paratype male, terminalia; 6, aedeagus in dorsal view; 7, aedeagus in lateral view; 8, tegmen in ventral view; 9, segment VIII and sternite IX (= spiculum gastrale) in dorsal view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033745" httpUri="https://zenodo.org/record/14033745/files/figure.png" pageId="4" pageNumber="295">Fig. 9</figureCitation>
) with symmetrical short arms, apodeme curved.
</paragraph>
<paragraph id="8B9636B5C031FF8025D61C56FE748A34" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="295">
<emphasis id="B95DEAA7C031FF8025D61C56FED18B44" bold="true" box="[199,286,1626,1652]" pageId="4" pageNumber="295">Female</emphasis>
(
<figureCitation id="13122A30C031FF8024201C56FE768B44" box="[305,441,1626,1652]" captionStart="FIGURES 1013" captionStartId="5.[151,264,1206,1230]" captionTargetBox="[151,1433,182,1178]" captionTargetId="figure-17@5.[151,1436,181,1183]" captionTargetPageId="5" captionText="FIGURES 1013. Diastatotropis lusti, allotype female, terminalia; 10, ovipositor in ventral view; 11, apex of ovipositor (hemisternites) in ventral view; 12, sternite VIII (left) and tergite VIII (right) in ventral view; 13, spermatheca. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033747" httpUri="https://zenodo.org/record/14033747/files/figure.png" pageId="4" pageNumber="295">Figs 1013</figureCitation>
). Ovipositor (
<figureCitation id="13122A30C031FF80274D1C56FD7D8B44" box="[604,690,1626,1652]" captionStart="FIGURES 1013" captionStartId="5.[151,264,1206,1230]" captionTargetBox="[151,1433,182,1178]" captionTargetId="figure-17@5.[151,1436,181,1183]" captionTargetPageId="5" captionText="FIGURES 1013. Diastatotropis lusti, allotype female, terminalia; 10, ovipositor in ventral view; 11, apex of ovipositor (hemisternites) in ventral view; 12, sternite VIII (left) and tergite VIII (right) in ventral view; 13, spermatheca. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033747" httpUri="https://zenodo.org/record/14033747/files/figure.png" pageId="4" pageNumber="295">Fig. 10</figureCitation>
) wide and short, lateral rods diverging. Hemisternites (
<figureCitation id="13122A30C031FF80202E1C56FA5C8B44" box="[1343,1427,1626,1652]" captionStart="FIGURES 1013" captionStartId="5.[151,264,1206,1230]" captionTargetBox="[151,1433,182,1178]" captionTargetId="figure-17@5.[151,1436,181,1183]" captionTargetPageId="5" captionText="FIGURES 1013. Diastatotropis lusti, allotype female, terminalia; 10, ovipositor in ventral view; 11, apex of ovipositor (hemisternites) in ventral view; 12, sternite VIII (left) and tergite VIII (right) in ventral view; 13, spermatheca. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033747" httpUri="https://zenodo.org/record/14033747/files/figure.png" pageId="4" pageNumber="295">Fig. 11</figureCitation>
) unarmed, without distinct toothed plates, with numerous setae apically. Sternite VIII (
<figureCitation id="13122A30C031FF8021461C72FB668BA8" box="[1111,1193,1662,1688]" captionStart="FIGURES 1013" captionStartId="5.[151,264,1206,1230]" captionTargetBox="[151,1433,182,1178]" captionTargetId="figure-17@5.[151,1436,181,1183]" captionTargetPageId="5" captionText="FIGURES 1013. Diastatotropis lusti, allotype female, terminalia; 10, ovipositor in ventral view; 11, apex of ovipositor (hemisternites) in ventral view; 12, sternite VIII (left) and tergite VIII (right) in ventral view; 13, spermatheca. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033747" httpUri="https://zenodo.org/record/14033747/files/figure.png" pageId="4" pageNumber="295">Fig. 12</figureCitation>
, left) transverse, with anterior margin almost straight, only slightly convex in middle, and with sclerotized apodeme. Tergite VIII (
<figureCitation id="13122A30C031FF8020571CAEFA588B8C" box="[1350,1431,1698,1724]" captionStart="FIGURES 1013" captionStartId="5.[151,264,1206,1230]" captionTargetBox="[151,1433,182,1178]" captionTargetId="figure-17@5.[151,1436,181,1183]" captionTargetPageId="5" captionText="FIGURES 1013. Diastatotropis lusti, allotype female, terminalia; 10, ovipositor in ventral view; 11, apex of ovipositor (hemisternites) in ventral view; 12, sternite VIII (left) and tergite VIII (right) in ventral view; 13, spermatheca. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033747" httpUri="https://zenodo.org/record/14033747/files/figure.png" pageId="4" pageNumber="295">Fig. 12</figureCitation>
, right) as wide as long, its anterior margin distinctly trilobed. Both segments with numerous setae apically. Shape of spermatheca as in
<figureCitation id="13122A30C031FF8024741CE6FE778A34" box="[357,440,1770,1796]" captionStart="FIGURES 1013" captionStartId="5.[151,264,1206,1230]" captionTargetBox="[151,1433,182,1178]" captionTargetId="figure-17@5.[151,1436,181,1183]" captionTargetPageId="5" captionText="FIGURES 1013. Diastatotropis lusti, allotype female, terminalia; 10, ovipositor in ventral view; 11, apex of ovipositor (hemisternites) in ventral view; 12, sternite VIII (left) and tergite VIII (right) in ventral view; 13, spermatheca. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14033747" httpUri="https://zenodo.org/record/14033747/files/figure.png" pageId="4" pageNumber="295">Fig. 13</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C333653EC031FF8125D61D02FF2F8B6B" lastPageId="5" lastPageNumber="296" pageId="4" pageNumber="295" type="etymology">
<paragraph id="8B9636B5C031FF8025D61D02FAF78A18" blockId="4.[151,1437,150,1977]" box="[199,1336,1806,1833]" pageId="4" pageNumber="295">
<emphasis id="B95DEAA7C031FF8025D61D02FE818A18" bold="true" box="[199,334,1806,1832]" pageId="4" pageNumber="295">Etymology.</emphasis>
Patronym, dedicated to our colleague and friend Zdeněk Lust (Rožany,
<collectingCountry id="F33E7625C031FF8021641D02FAE38A18" box="[1141,1324,1806,1832]" name="Czech Republic" pageId="4" pageNumber="295">Czech Republic</collectingCountry>
).
</paragraph>
<paragraph id="8B9636B5C031FF8025D61D3EFD198A88" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="295">
<emphasis id="B95DEAA7C031FF8025D61D3EFE3E8A7C" bold="true" box="[199,497,1842,1868]" pageId="4" pageNumber="295">Collecting circumstances</emphasis>
(
<figureCitation id="13122A30C031FF8027101D3EFD4B8A7D" box="[513,644,1842,1869]" captionStart="FIGURES 1415" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-17@6.[178,1408,181,1918]" captionTargetPageId="6" captionText="FIGURES 1415. Ranomafana National Park, overall view of collecting site of Diastatotropis lusti." figureDoi="http://doi.org/10.5281/zenodo.14033753" httpUri="https://zenodo.org/record/14033753/files/figure.png" pageId="4" pageNumber="295">Figs 1415</figureCitation>
).
<taxonomicName id="4C294D36C031FF8027891D3FFCBE8A7C" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[664,881,1842,1868]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="lusti">
<emphasis id="B95DEAA7C031FF8027891D3FFCBE8A7C" box="[664,881,1842,1868]" italics="true" pageId="4" pageNumber="295">Diastatotropis lusti</emphasis>
</taxonomicName>
was found in Ranomafana National Park located ca.
<quantity id="4CD19B50C031FF8025D01D5AFEC58A40" box="[193,266,1878,1904]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="295" unit="km" value="40.0">40 km</quantity>
(by air) NW of
<collectingRegion id="49EDF857C031FF8024DA1D5AFD958A40" box="[459,602,1878,1904]" country="Madagascar" name="Fianarantsoa" pageId="4" pageNumber="295">Fianarantsoa</collectingRegion>
town,
<collectingRegion id="49EDF857C031FF8027BD1D5AFC648A40" box="[684,939,1878,1904]" country="Madagascar" name="Fianarantsoa" pageId="4" pageNumber="295">Fianarantsoa province</collectingRegion>
. Ranomafana N.P. is covered with medium altitude moist evergreen forest, dominated by the humid climate with no typical dry season. The detailed description of the area is presented by
<bibRefCitation id="EFB84B44C031FF8024D21D92FD1D8A88" author="Goodman, S. M. &amp; Raherilalao, M. J. &amp; Wohlhauser, S. &amp; Tome II" box="[451,722,1950,1976]" pageId="4" pageNumber="295" pagination="425 - 1231" refId="ref5324" refString="Goodman, S. M., Raherilalao, M. J. &amp; Wohlhauser, S. (Eds.) (2018 a) s. n. In: Les aires protegees terrestres de Madagascar: Leur histoire, description et biote. Tome II: Le nord et l´est de Madagascar / The terrestrial protected areas of Madagascar: Their history, description, and biota. Vol. II. Northern and eastern Madagascar. Association Vahatra, Antananarivo, pp. 425 - 1231. [the correct surname of the third editor is Wohlhauser, not Wohlauser as given on the book cover]" type="book chapter" year="2018">
Goodman
<emphasis id="B95DEAA7C031FF8027281D93FDBE8A88" box="[569,625,1950,1976]" italics="true" pageId="4" pageNumber="295">et al.</emphasis>
(2018a)
</bibRefCitation>
.
</paragraph>
<caption id="DF56663DC030FF8125861EBAFF098826" ID-DOI="http://doi.org/10.5281/zenodo.14033747" ID-Zenodo-Dep="14033747" httpUri="https://zenodo.org/record/14033747/files/figure.png" pageId="5" pageNumber="296" startId="5.[151,264,1206,1230]" targetBox="[151,1433,182,1178]" targetPageId="5" targetType="figure">
<paragraph id="8B9636B5C030FF8125861EBAFF098826" blockId="5.[151,1436,1206,1302]" pageId="5" pageNumber="296">
<emphasis id="B95DEAA7C030FF8125861EBAFE9789FE" bold="true" box="[151,344,1206,1230]" pageId="5" pageNumber="296">FIGURES 1013.</emphasis>
<taxonomicName id="4C294D36C030FF8124731EBBFDE589FF" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[354,554,1207,1231]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="296" phylum="Arthropoda" rank="species" species="lusti">
<emphasis id="B95DEAA7C030FF8124731EBBFDE589FF" box="[354,554,1207,1231]" italics="true" pageId="5" pageNumber="296">Diastatotropis lusti</emphasis>
</taxonomicName>
, allotype female, terminalia; 10, ovipositor in ventral view; 11, apex of ovipositor (hemisternites) in ventral view; 12, sternite VIII (left) and tergite VIII (right) in ventral view; 13, spermatheca. Scale bars in mm.
</paragraph>
</caption>
<paragraph id="8B9636B5C030FF8125D61F65FF2F8B6B" blockId="5.[151,1437,1385,1951]" pageId="5" pageNumber="296">
<specimenCount id="9D2FFD3CC030FF8125D61F65FEF888B3" box="[199,311,1385,1411]" count="2" pageId="5" pageNumber="296" type="male">Two male</specimenCount>
specimens of
<taxonomicName id="4C294D36C030FF8124CD1F66FDE088B3" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[476,559,1385,1411]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="296" phylum="Arthropoda" rank="species" species="lusti">
<emphasis id="B95DEAA7C030FF8124CD1F66FDE088B3" box="[476,559,1385,1411]" italics="true" pageId="5" pageNumber="296">D. lusti</emphasis>
</taxonomicName>
were collected in a slightly disturbed place on the edge of the primary forest at elevation
<quantity id="4CD19B50C030FF8124311F81FEBA8897" box="[288,373,1421,1447]" metricMagnitude="3" metricUnit="m" metricValue="1.189" pageId="5" pageNumber="296" unit="m" value="1189.0">1189 m</quantity>
a.s.l. These males were caught climbing on the central wall of an installed Malaise trap during the day. No other specimens of this species were found in the collection container of the trap or in the immediate vicinity. The last specimen, a female, was collected in Ranomafana env.. According to the collector, the specimen was obtained by collecting along National Road (RN) 25, which forms the boundary of the National Park. A single specimen was obtained by beating of vegetation, mainly bushes and branches of low trees (Ivo Jeniš, pers. comm., 2000).
</paragraph>
</subSubSection>
<subSubSection id="C333653EC030FF8125D61C68FDF88B93" pageId="5" pageNumber="296" type="distribution">
<paragraph id="8B9636B5C030FF8125D61C68FDF88B93" blockId="5.[151,1437,1385,1951]" pageId="5" pageNumber="296">
<emphasis id="B95DEAA7C030FF8125D61C68FEAE8B4E" bold="true" box="[199,353,1636,1662]" pageId="5" pageNumber="296">Distribution.</emphasis>
E
<collectingCountry id="F33E7625C030FF81249F1C69FDD88B4F" box="[398,535,1637,1663]" name="Madagascar" pageId="5" pageNumber="296">Madagascar</collectingCountry>
,
<collectingRegion id="49EDF857C030FF81273B1C69FCE18B4F" box="[554,814,1637,1663]" country="Madagascar" name="Fianarantsoa" pageId="5" pageNumber="296">Fianarantsoa province</collectingRegion>
. The species is only known from the
<typeStatus id="54928817C030FF81201B1C69FAF58B4F" box="[1290,1338,1637,1663]" pageId="5" pageNumber="296">type</typeStatus>
locality Ranomafana National Park (
<figureCitation id="13122A30C030FF8124C71C85FDE58B93" box="[470,554,1673,1699]" captionStart="FIGURE 28" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[183,1406,188,1914]" captionTargetId="figure-17@11.[179,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 28. Type localities of Diastatotropis lusti and D. petulae." figureDoi="http://doi.org/10.5281/zenodo.14033761" httpUri="https://zenodo.org/record/14033761/files/figure.png" pageId="5" pageNumber="296">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C333653EC030FF8125D61CA0FDA18AAF" pageId="5" pageNumber="296" type="diagnosis">
<paragraph id="8B9636B5C030FF8125D61CA0FDA18AAF" blockId="5.[151,1437,1385,1951]" pageId="5" pageNumber="296">
<emphasis id="B95DEAA7C030FF8125D61CA0FE1C8BF6" bold="true" box="[199,467,1708,1734]" pageId="5" pageNumber="296">Differential diagnosis.</emphasis>
<taxonomicName id="4C294D36C030FF8124CE1CA2FD748BF7" authorityName="Trýzna &amp; Blažej &amp; Rakotonirina" authorityYear="2024" box="[479,699,1709,1735]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="296" phylum="Arthropoda" rank="species" species="lusti">
<emphasis id="B95DEAA7C030FF8124CE1CA2FD748BF7" box="[479,699,1709,1735]" italics="true" pageId="5" pageNumber="296">Diastatotropis lusti</emphasis>
</taxonomicName>
belongs to a group of species with antennal club slender and cylindrical, antennomere X as long as wide, and terminal segment elongate-oval, always distinctly longer than wide. Rostrum with only two lateral carinae. Oblique stripes are prominent on the elytra, that converge apically. The species is distinguished from other species by the shape of the pronotum (in dorsal view), which is very protruding laterally in the area of the dorsal transverse carina. It is also characterized by very short lateral carinae of pronotum, which are not strongly indicated and not reaching half of length of pronotum. Pygidium in both sexes lacks a longitudinal stripe of light-coloured setae.
</paragraph>
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@ -0,0 +1,155 @@
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<mods:title id="958B2EEF4125379CAD472BD5F80C5328">Two new species of Diastatotropis Lacordaire from protected areas in central and east Madagascar (Coleoptera: Anthribidae)</mods:title>
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Genus
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<emphasis id="B95DEAA7C034FF8525FF1E17FE578905" bold="true" box="[238,408,1051,1077]" italics="true" pageId="1" pageNumber="292">Diastatotropis</emphasis>
<bibRefCitation id="EFB84B44C034FF85248E1E16FDBB8904" author="Lacordaire, J. T." box="[415,628,1050,1076]" pageId="1" pageNumber="292" refId="ref5586" refString="Lacordaire, J. T. (1865) Histoire naturelle des insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu´ici dans cet ordre d´insectes. Tome septieme contenant les familles des curculionides (suite), scolytides, brenthides, anthribides et bruchides. Librairie Roret, Paris, 7 + [2] + 620 pp. [date of publication by 12. XII. 1865 (Bousquet 2016: 314), although on title page 1866]" type="book" year="1865">Lacordaire, 1865</bibRefCitation>
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species:
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<emphasis id="B95DEAA7C034FF85242D1E6FFDE0894C" box="[316,559,1123,1148]" italics="true" pageId="1" pageNumber="292">Diastatotropis tigrina</emphasis>
<bibRefCitation id="EFB84B44C034FF8527261E6FFD39894D" author="Lacordaire, J. T." box="[567,758,1122,1149]" pageId="1" pageNumber="292" refId="ref5586" refString="Lacordaire, J. T. (1865) Histoire naturelle des insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu´ici dans cet ordre d´insectes. Tome septieme contenant les familles des curculionides (suite), scolytides, brenthides, anthribides et bruchides. Librairie Roret, Paris, 7 + [2] + 620 pp. [date of publication by 12. XII. 1865 (Bousquet 2016: 314), although on title page 1866]" type="book" year="1865">Lacordaire, 1865</bibRefCitation>
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(by original designation),
<typeStatus id="54928817C034FF8521311E6FFB9F894D" box="[1056,1104,1123,1149]" pageId="1" pageNumber="292">type</typeStatus>
locality Madagascar
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<emphasis id="B95DEAA7C034FF8525D61E8BFE848991" bold="true" box="[199,331,1159,1185]" pageId="1" pageNumber="292">Placement.</emphasis>
<taxonomicName id="4C294D36C034FF8524431E8BFE138991" authorityName="Billberg" authorityYear="1820" box="[338,476,1159,1185]" class="Insecta" family="Anthribidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="292" phylum="Arthropoda" rank="subFamily" subFamily="Anthribinae">Anthribinae</taxonomicName>
:
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(see Discussion in
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).
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Diagnosis of
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<emphasis id="B95DEAA7C034FF8524481EA7FE3489F5" bold="true" box="[345,507,1195,1221]" italics="true" pageId="1" pageNumber="292">Diastatotropis</emphasis>
</taxonomicName>
.
</emphasis>
Body oval to elongate.Rostrum extended apically, with one, two or three longitudinal carinae or lacking carinae. Eyes spherical or elliptical, not emarginate. Antennae of males usually reaching posterior margin of pronotum or beyond, antennae of females shorter, not reaching posterior margin of pronotum. Antennal funicle slender, then antennal club 3-segmented (most species) or 4-segmented, or antennomeres continuously extended. Dorsal transverse carina of pronotum distinct and antebasal. Medium- or large-sized species, from
<quantity id="4CD19B50C034FF85204D1F37FA528865" box="[1372,1437,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="292" unit="mm" value="6.0">6 mm</quantity>
(
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<emphasis id="B95DEAA7C034FF8525B01F53FE8F8849" box="[161,320,1375,1401]" italics="true" pageId="1" pageNumber="292">D. arofaniana</emphasis>
<bibRefCitation id="EFB84B44C034FF8524581F53FD898849" author="Tryzna, M. &amp; Banar, P." box="[329,582,1375,1401]" pageId="1" pageNumber="292" pagination="30 - 52" refId="ref6040" refString="Tryzna, M. &amp; Banar, P. (2022) Diastatotropis arofaniana sp. nov. from eastern Madagascar, a new species close to D. rubra Frieser, with a redescription of the type of the latter, and notes on morphology (Coleoptera: Anthribidae). Zootaxa, 5120 (1), 30 - 52. https: // doi. org / 10.11646 / zootaxa. 5120.1.2" type="journal article" year="2022">Trýzna &amp; Baňař 2022</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C294D36C034FF8527421F53FBDF8849" authority="Tryzna, Blazej &amp; Rakotonirina" authorityName="Tryzna, Blazej &amp; Rakotonirina" authorityYear="2024" box="[595,1040,1375,1401]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="lusti" status="sp. nov.">
<emphasis id="B95DEAA7C034FF8527421F53FD688849" box="[595,679,1375,1401]" italics="true" pageId="1" pageNumber="292">D. lusti</emphasis>
Trýzna, Blažej &amp; Rakotonirina
</taxonomicName>
<emphasis id="B95DEAA7C034FF8521081F53FBBB8849" bold="true" box="[1049,1140,1375,1401]" pageId="1" pageNumber="292">
<taxonomicNameLabel id="A26E57DCC034FF8521081F53FBBB8849" box="[1049,1140,1375,1401]" pageId="1" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
or small specimens of
<taxonomicName id="4C294D36C034FF8520901F53FE2C88AD" authority="(Fairmaire, 1893))" baseAuthorityName="Fairmaire" baseAuthorityYear="1893" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="rubricollis">
<emphasis id="B95DEAA7C034FF8520901F53FEC288AD" italics="true" pageId="1" pageNumber="292">D. rubricollis</emphasis>
(Fairmaire, 1893))
</taxonomicName>
to more than
<quantity id="4CD19B50C034FF85276E1F8FFD0188AD" box="[639,718,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="292" unit="mm" value="18.0">18 mm</quantity>
(
<taxonomicName id="4C294D36C034FF8527CF1F8FFBFC88AD" authority="Waterhouse, 1877" authorityName="Waterhouse" authorityYear="1877" box="[734,1075,1411,1437]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B95DEAA7C034FF8527CF1F8FFC9088AD" box="[734,863,1411,1437]" italics="true" pageId="1" pageNumber="292">D. olivacea</emphasis>
Waterhouse, 1877
</taxonomicName>
).
</paragraph>
<paragraph id="8B9636B5C034FF8525D61FABFDCE8B45" blockId="1.[151,1437,1122,1653]" pageId="1" pageNumber="292">
Male and female genitalia conspicuously small compared to overall size of body (see Discussion in
<bibRefCitation id="EFB84B44C034FF8520211FABFEEE88D5" author="Tryzna, M. &amp; Banar, P." pageId="1" pageNumber="292" pagination="30 - 52" refId="ref6040" refString="Tryzna, M. &amp; Banar, P. (2022) Diastatotropis arofaniana sp. nov. from eastern Madagascar, a new species close to D. rubra Frieser, with a redescription of the type of the latter, and notes on morphology (Coleoptera: Anthribidae). Zootaxa, 5120 (1), 30 - 52. https: // doi. org / 10.11646 / zootaxa. 5120.1.2" type="journal article" year="2022">Trýzna &amp; Baňař 2022</bibRefCitation>
).
<emphasis id="B95DEAA7C034FF8524261FC7FE3688D5" box="[311,505,1483,1509]" italics="true" pageId="1" pageNumber="292">Female genitalia</emphasis>
: ovipositor short and wide, hemisternites strikingly short and delicate, unarmed, without distinct toothed plates, with only numerous setae or small preapical stylus. Lateral and median rods very short, median rod reaching about half of the length of lateral rod. Transverse bar invisible.
<emphasis id="B95DEAA7C034FF8521A21C1FFA958B1D" box="[1203,1370,1555,1581]" italics="true" pageId="1" pageNumber="292">Male genitalia</emphasis>
more diversely shaped. For depicted genitalia of
<taxonomicName id="4C294D36C034FF85277F1C3BFCC08B60" authorityName="Lacordaire" authorityYear="1866" box="[622,783,1591,1616]" class="Insecta" family="Anthribidae" genus="Diastatotropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA7C034FF85277F1C3BFCC08B60" box="[622,783,1591,1616]" italics="true" pageId="1" pageNumber="292">Diastatotropis</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFB84B44C034FF85262D1C3BFBE28B61" author="Tryzna, M. &amp; Banar, P." box="[828,1069,1591,1617]" pageId="1" pageNumber="292" pagination="537 - 544" refId="ref5921" refString="Tryzna, M. &amp; Banar, P. (2017) A new species of Diastatotropis Lacordaire (Coleoptera: Anthribidae) from Montagne d´Ambre National Park, northern Madagascar. Zootaxa, 4221 (5), 537 - 544. https: // doi. org / 10.11646 / zootaxa. 4221.5.3" type="journal article" year="2017">Trýzna &amp; Baňař 2017</bibRefCitation>
,
<bibRefCitation id="EFB84B44C034FF8521241C3BFF008B45" author="Tryzna, M. &amp; Andrianomenjanahary, M. N." pageId="1" pageNumber="292" pagination="444 - 450" refId="ref5802" refString="Tryzna, M. &amp; Andrianomenjanahary, M. N. (2019) Description of a new species of the genus Diastatotropis Lacordaire (Coleoptera: Anthribidae) with strikingly elongated elytral apices from north-eastern Madagascar. Zootaxa, 4563 (3), 444 - 450. https: // doi. org / 10.11646 / zootaxa. 4563.3.2" type="journal article" year="2019">Trýzna &amp;Andrianomenjanahary 2019</bibRefCitation>
and
<bibRefCitation id="EFB84B44C034FF8524141C57FE328B45" author="Tryzna, M. &amp; Banar, P." box="[261,509,1627,1653]" pageId="1" pageNumber="292" pagination="30 - 52" refId="ref6040" refString="Tryzna, M. &amp; Banar, P. (2022) Diastatotropis arofaniana sp. nov. from eastern Madagascar, a new species close to D. rubra Frieser, with a redescription of the type of the latter, and notes on morphology (Coleoptera: Anthribidae). Zootaxa, 5120 (1), 30 - 52. https: // doi. org / 10.11646 / zootaxa. 5120.1.2" type="journal article" year="2022">Trýzna &amp; Baňař 2022</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,619 @@
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<mods:title id="EB9E59C51550020896EFD4AB22AB84FD">A new species of Eudemopsis Falkovitsh (Lepidoptera: Tortricidae: Olethreutinae: Olethreutini) from Thailand and Laos, with the proposed synonymy of Fansipaniana Razowski</mods:title>
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Pinkaew &amp; Muadsub
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<paragraph id="8B9036CCFFD9FFAD7CFE8488FEB4B1E6" blockId="1.[151,1069,1425,1524]" box="[151,288,1498,1524]" pageId="1" pageNumber="337">
(
<figureCitation id="13142A49FFD9FFAD7CF68488FE83B1E6" box="[159,279,1498,1524]" captionStart-0="FIGURES 12" captionStart-1="FIGURES 34" captionStart-2="FIGURES 56" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStart-5="FIGURE 9" captionStart-6="FIGURES 1012" captionStart-7="FIGURE 13" captionStartId-0="2.[151,264,1365,1389]" captionStartId-1="2.[151,264,2016,2040]" captionStartId-2="3.[151,264,901,925]" captionStartId-3="3.[151,250,1655,1679]" captionStartId-4="4.[151,250,869,893]" captionStartId-5="4.[151,250,1734,1758]" captionStartId-6="5.[151,264,1546,1570]" captionStartId-7="6.[151,250,1920,1944]" captionTargetBox-0="[333,1252,319,1326]" captionTargetBox-1="[151,1435,1486,1992]" captionTargetBox-2="[166,1421,193,860]" captionTargetBox-3="[185,1403,991,1628]" captionTargetBox-4="[183,1409,191,835]" captionTargetBox-5="[579,1026,939,1709]" captionTargetBox-6="[222,1374,181,1509]" captionTargetBox-7="[535,1047,645,1882]" captionTargetId-0="figure-73@2.[322,1265,306,1338]" captionTargetId-1="figure-97@2.[151,1435,1486,1992]" captionTargetId-2="figure-29@3.[154,1434,181,877]" captionTargetId-3="figure-53@3.[172,1416,986,1632]" captionTargetId-4="figure-18@4.[175,1412,181,846]" captionTargetId-5="figure-39@4.[562,1026,918,1710]" captionTargetId-6="figure-29@5.[212,1375,181,1521]" captionTargetId-7="figure-259@6.[520,1066,633,1897]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionTargetPageId-4="4" captionTargetPageId-5="4" captionTargetPageId-6="5" captionTargetPageId-7="6" captionText-0="FIGURES 12. Wing pattern of Eudemopsis squamapanna sp. nov. 1. Male (holotype); 2. Female (paratype, np7971). Scale bars = 2 mm." captionText-1="FIGURES 34. Head of Eudemopsis squamapanna sp. nov. 3. Male (holotype); 4. Female(paratype, np7971)." captionText-2="FIGURES 56. Wing venation on the right wings of Eudemopsis squamapanna sp. nov. 5. Male (paratype, np6618); 6. Female (paratype, np7971)." captionText-3="FIGURE 7. Male abdomen of Eudemopsis squamapanna sp. nov. with two small lobes of dense scale sockets on the abdominal segment IV anterolaterally (paratype, NP4167). Scale bars = 1 mm." captionText-4="FIGURE 8. Morphological features of male genitalia of Eudemopsis squamapanna sp. nov. (holotype). Scale bars = 0.5 mm." captionText-5="FIGURE 9. Extended part of uncus, tuba analis, gnathos, phallus and anellus in male genitalia of Eudemopsis squamapanna sp. nov. (holotype). Scale bars = 0.5 mm." captionText-6="FIGURES 1012. Modified scales on the anterior surface on male valva of Eudemopsis squamapanna sp. nov. 10. Exposed valva from the abdominal segment VIII (pinned specimen, paratype, np13911); 11. Valva after dyed in EosinY (holotype); 12. Mounted valva in Euparal (holotype) (A. Modified short scales B. Modified very long scales on membranous lobe)." captionText-7="FIGURE 13. Morphological features of female genitalia of Eudemopsis squamapanna sp. nov. (paratype, NP3302). Scale bars = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14033805" figureDoi-1="http://doi.org/10.5281/zenodo.14033807" figureDoi-2="http://doi.org/10.5281/zenodo.14033809" figureDoi-3="http://doi.org/10.5281/zenodo.14033811" figureDoi-4="http://doi.org/10.5281/zenodo.14033815" figureDoi-5="http://doi.org/10.5281/zenodo.14033817" figureDoi-6="http://doi.org/10.5281/zenodo.14033819" figureDoi-7="http://doi.org/10.5281/zenodo.14033821" httpUri-0="https://zenodo.org/record/14033805/files/figure.png" httpUri-1="https://zenodo.org/record/14033807/files/figure.png" httpUri-2="https://zenodo.org/record/14033809/files/figure.png" httpUri-3="https://zenodo.org/record/14033811/files/figure.png" httpUri-4="https://zenodo.org/record/14033815/files/figure.png" httpUri-5="https://zenodo.org/record/14033817/files/figure.png" httpUri-6="https://zenodo.org/record/14033819/files/figure.png" httpUri-7="https://zenodo.org/record/14033821/files/figure.png" pageId="1" pageNumber="337">Figs. 113</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3356547FFD9FFAE7CFE8770FE08B50F" lastPageId="2" lastPageNumber="338" pageId="1" pageNumber="337" type="materials_examined">
<paragraph id="8B9036CCFFD9FFAE7CFE8770FE08B50F" blockId="1.[151,1437,1569,2029]" lastBlockId="2.[151,1437,151,285]" lastPageId="2" lastPageNumber="338" pageId="1" pageNumber="337">
<emphasis id="B95BEADEFFD9FFAD7CFE8770FED3B229" bold="true" box="[151,327,1569,1596]" pageId="1" pageNumber="337">Type Material.</emphasis>
<materialsCitation id="3B473C91FFD9FFAD7D248770FA86B272" collectingDate="2017-01-04" collectingDateMax="2017-01-07" collectingDateMin="2017-01-04" collectionCode="KKIC" collectorName="Khao Yai N. P. &amp; N. Pinkaew &amp; Deposited" country="Thailand" elevation="700" latitude="14.431945" location="Nakhon Ratchasima Prov." longLatPrecision="21" longitude="101.40139" pageId="1" pageNumber="337" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Ratchasima" typeStatus="holotype">
<emphasis id="B95BEADEFFD9FFAD7D248770FE57B22E" bold="true" box="[333,451,1570,1596]" pageId="1" pageNumber="337">
<typeStatus id="5494886EFFD9FFAD7D248770FE2AB22E" box="[333,446,1570,1596]" pageId="1" pageNumber="337" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D29FD45FFD9FFAD7DA08770FE74B22E" box="[457,480,1570,1596]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
, [
<collectingCountry id="F338765CFFD9FFAD7D988770FD18B22E" box="[497,652,1570,1596]" name="Thailand" pageId="1" pageNumber="337">THAILAND</collectingCountry>
]:
<collectingRegion id="49EBF82EFFD9FFAD7EF28770FC2BB22E" box="[667,959,1570,1596]" country="Thailand" name="Nakhon Ratchasima" pageId="1" pageNumber="337">Nakhon Ratchasima Prov.</collectingRegion>
,
<collectorName id="26DA531AFFD9FFAD7FA08770FBF3B22E" box="[969,1127,1570,1596]" pageId="1" pageNumber="337">Khao Yai N.P.</collectorName>
,
<geoCoordinate id="EE1B500BFFD9FFAD78188770FB6EB22F" box="[1137,1274,1570,1597]" degrees="14" direction="north" minutes="25" orientation="latitude" pageId="1" pageNumber="337" precision="15" seconds="55" value="14.431945">14°25ˊ55˝N</geoCoordinate>
,
<geoCoordinate id="EE1B500BFFD9FFAD796D8770FA0CB22F" box="[1284,1432,1570,1597]" degrees="101" direction="east" minutes="24" orientation="longitude" pageId="1" pageNumber="337" precision="15" seconds="05" value="101.40139">101°24ˊ05˝E</geoCoordinate>
, alt.
<quantity id="4CD79B29FFD9FFAD7CA98714FE9CB272" box="[192,264,1606,1632]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="337" unit="m" value="700.0">
<elevation id="0002D1FFFFD9FFAD7CA98714FE9CB272" box="[192,264,1606,1632]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="337" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="FF91100CFFD9FFAD7D7A8714FE24B272" box="[275,432,1606,1632]" pageId="1" pageNumber="337" value="2017-01-04" valueMax="2017-01-07" valueMin="2017-01-04">
<collectingDate id="EFD5E9E4FFD9FFAD7D7A8714FE24B272" box="[275,432,1606,1632]" pageId="1" pageNumber="337" value="2017-01-04" valueMax="2017-01-07" valueMin="2017-01-04">47 Jan. 2017</collectingDate>
</date>
,
<collectorName id="26DA531AFFD9FFAD7DD38714FDAFB272" box="[442,571,1606,1632]" pageId="1" pageNumber="337">N. Pinkaew</collectorName>
<emphasis id="B95BEADEFFD9FFAD7E2B8715FDE6B272" box="[578,626,1606,1632]" italics="true" pageId="1" pageNumber="337">et al</emphasis>
. leg., np11115 (genitalia slide NP3301).
<collectorName id="26DA531AFFD9FFAD785F8714FB3CB272" box="[1078,1192,1606,1632]" pageId="1" pageNumber="337">Deposited</collectorName>
in
<collectionCode id="ED3EAE09FFD9FFAD78A38714FA86B272" box="[1226,1298,1606,1632]" pageId="1" pageNumber="337">KKIC</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B473C91FFD9FFAE79728714FD08B4C7" collectingDate="2009-05-15" collectingDateMax="2023-09-19" collectingDateMin="2009-05-15" collectionCode="KKIC" collectorName="Khao Yai N. P. &amp; Pang Sida N. P. &amp; These &amp; N. Pinkaew" country="Thailand" county="Bueng Khong Long Non-Hunting Area" elevation="170" lastPageId="2" lastPageNumber="338" latitude="12.392777" location="Agroforestry Research Station" longLatPrecision="21" longitude="102.666664" municipality="Khao Glaur" pageId="1" pageNumber="337" specimenCount="38" specimenCount-female="2" specimenCount-male="36" stateProvince="Bueng Kan" typeStatus="paratype">
<emphasis id="B95BEADEFFD9FFAD79728714FA08B272" bold="true" box="[1307,1436,1606,1632]" pageId="1" pageNumber="337">
<typeStatus id="5494886EFFD9FFAD79728714FA03B272" box="[1307,1431,1606,1632]" pageId="1" pageNumber="337" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D29FD45FFD9FFAD7CFE8738FF5EB296" box="[151,202,1642,1668]" count="18" pageId="1" pageNumber="337" type="male">18♂</specimenCount>
,
<specimenCount id="9D29FD45FFD9FFAD7CBF8738FF6DB296" box="[214,249,1642,1668]" count="1" pageId="1" pageNumber="337" type="female">1♀</specimenCount>
, [
<collectingCountry id="F338765CFFD9FFAD7D678738FE31B296" box="[270,421,1642,1668]" name="Thailand" pageId="1" pageNumber="337">THAILAND</collectingCountry>
]
<collectingRegion id="49EBF82EFFD9FFAD7DDB8738FDE1B296" box="[434,629,1642,1668]" country="Thailand" name="Bueng Kan" pageId="1" pageNumber="337">Bueng Kan Prov.</collectingRegion>
,
<collectingCounty id="62F14E40FFD9FFAD7EE98738FBAEB296" box="[640,1082,1642,1668]" pageId="1" pageNumber="337">Bueng Khong Long Non-Hunting Area</collectingCounty>
,
<geoCoordinate id="EE1B500BFFD9FFAD782F8738FB5AB296" box="[1094,1230,1642,1668]" degrees="18" direction="north" minutes="01" orientation="latitude" pageId="1" pageNumber="337" precision="15" seconds="20" value="18.022223">18°01ˊ20˝N</geoCoordinate>
,
<geoCoordinate id="EE1B500BFFD9FFAD78B38738FAFAB297" box="[1242,1390,1642,1669]" degrees="104" direction="east" minutes="00" orientation="longitude" pageId="1" pageNumber="337" precision="15" seconds="52" value="104.01444">104°00ˊ52˝E</geoCoordinate>
, alt.
<quantity id="4CD79B29FFD9FFAD7CFE87DCFF4BB2BA" box="[151,223,1678,1704]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="337" unit="m" value="170.0">
<elevation id="0002D1FFFFD9FFAD7CFE87DCFF4BB2BA" box="[151,223,1678,1704]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="337" unit="m" value="170.0">170 m</elevation>
</quantity>
,
<date id="FF91100CFFD9FFAD7C8087DCFEF9B2BA" box="[233,365,1678,1704]" pageId="1" pageNumber="337" value="2023-04-06">
<collectingDate id="EFD5E9E4FFD9FFAD7C8087DCFEF9B2BA" box="[233,365,1678,1704]" pageId="1" pageNumber="337" value="2023-04-06">6 Apr. 2023</collectingDate>
</date>
, np13905 (
<specimenCount id="9D29FD45FFD9FFAD7D8187DCFE6BB2BA" box="[488,511,1678,1704]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
, genitalia slide NP4169); same collection data as preceding,
<date id="FF91100CFFD9FFAD78F487DCFAA4B2BA" box="[1181,1328,1678,1704]" pageId="1" pageNumber="337" value="2023-09-19">
<collectingDate id="EFD5E9E4FFD9FFAD78F487DCFAA4B2BA" box="[1181,1328,1678,1704]" pageId="1" pageNumber="337" value="2023-09-19">19 Sep. 2023</collectingDate>
</date>
, np13910 (
<specimenCount id="9D29FD45FFD9FFAD7CC887E0FF2CB2DE" box="[161,184,1714,1740]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
), np13911 (
<specimenCount id="9D29FD45FFD9FFAD7D2D87E0FECFB2DE" box="[324,347,1714,1740]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
), np13912 (
<specimenCount id="9D29FD45FFD9FFAD7D8087E0FD94B2DE" box="[489,512,1714,1740]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
), np13913 (
<specimenCount id="9D29FD45FFD9FFAD7EE787E0FD31B2DE" box="[654,677,1714,1740]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
).
<collectingRegion id="49EBF82EFFD9FFAD7ED487E0FC7EB2DE" box="[701,1002,1714,1740]" country="Thailand" name="Nakhon Ratchasima" pageId="1" pageNumber="337">Nakhon Ratchasima Prov.</collectingRegion>
, same collection data as
<typeStatus id="5494886EFFD9FFAD796487E0FAE6B2DE" box="[1293,1394,1714,1740]" pageId="1" pageNumber="337" type="holotype">holotype</typeStatus>
,
<date id="FF91100CFFD9FFAD79E987E0FE92B2E2" pageId="1" pageNumber="337" value="2016-01-14">
<collectingDate id="EFD5E9E4FFD9FFAD79E987E0FE92B2E2" pageId="1" pageNumber="337" value="2016-01-14">14 Jan. 2016</collectingDate>
</date>
, np9802 (
<specimenCount id="9D29FD45FFD9FFAD7D1C8784FE18B2E2" box="[373,396,1750,1776]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
, genitalia slide NP4167); same collection data as
<typeStatus id="5494886EFFD9FFAD7FD38784FB8BB2E2" box="[954,1055,1750,1776]" pageId="1" pageNumber="337" type="holotype">holotype</typeStatus>
,
<date id="FF91100CFFD9FFAD78438784FB3DB2E2" box="[1066,1193,1750,1776]" pageId="1" pageNumber="337" value="2016-07-07">
<collectingDate id="EFD5E9E4FFD9FFAD78438784FB3DB2E2" box="[1066,1193,1750,1776]" pageId="1" pageNumber="337" value="2016-07-07">7 Jul. 2016</collectingDate>
</date>
, np8887 (
<specimenCount id="9D29FD45FFD9FFAD79708784FAA4B2E2" box="[1305,1328,1750,1776]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
, genitalia slide NP4166), np7971 (
<specimenCount id="9D29FD45FFD9FFAD7DC787A8FE55B306" box="[430,449,1786,1812]" count="1" pageId="1" pageNumber="337" type="female"></specimenCount>
, genitalia slide NP3302, wing slide).
<collectingRegion id="49EBF82EFFD9FFAD7F0187A8FBC3B306" box="[872,1111,1786,1812]" country="Thailand" name="Nakhon Nayok" pageId="1" pageNumber="337">Nakhon Nayok Prov.</collectingRegion>
,
<collectorName id="26DA531AFFD9FFAD780B87A8FA97B306" box="[1122,1283,1786,1812]" pageId="1" pageNumber="337">Khao Yai N.P.</collectorName>
,
<geoCoordinate id="EE1B500BFFD9FFAD796687A8FA0CB307" box="[1295,1432,1786,1813]" degrees="14" direction="north" minutes="23" orientation="latitude" pageId="1" pageNumber="337" precision="15" seconds="56" value="14.398889">14°23ˊ56˝N</geoCoordinate>
,
<geoCoordinate id="EE1B500BFFD9FFAD7CFE864CFEBEB32A" box="[151,298,1822,1848]" degrees="101" direction="east" minutes="22" orientation="longitude" pageId="1" pageNumber="337" precision="15" seconds="16" value="101.37112">101°22ˊ16˝E</geoCoordinate>
, alt.
<quantity id="4CD79B29FFD9FFAD7D0D864CFE3BB32A" box="[356,431,1822,1848]" metricMagnitude="2" metricUnit="m" metricValue="7.859999999999999" pageId="1" pageNumber="337" unit="m" value="786.0">
<elevation id="0002D1FFFFD9FFAD7D0D864CFE3BB32A" box="[356,431,1822,1848]" metricMagnitude="2" metricUnit="m" metricValue="7.859999999999999" pageId="1" pageNumber="337" unit="m" value="786.0">786 m</elevation>
</quantity>
,
<date id="FF91100CFFD9FFAD7DD5864CFDCFB32A" box="[444,603,1822,1848]" pageId="1" pageNumber="337" value="2010-08-13">
<collectingDate id="EFD5E9E4FFD9FFAD7DD5864CFDCFB32A" box="[444,603,1822,1848]" pageId="1" pageNumber="337" value="2010-08-13">13 Aug. 2010</collectingDate>
</date>
, np4656 (
<specimenCount id="9D29FD45FFD9FFAD7EA7864CFD71B32A" box="[718,741,1822,1848]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
, genitalia slide NP1503);
<geoCoordinate id="EE1B500BFFD9FFAD7879864CFB0CB32A" box="[1040,1176,1822,1848]" degrees="14" direction="north" minutes="26" orientation="latitude" pageId="1" pageNumber="337" precision="15" seconds="18" value="14.4383335">14°26ˊ18˝N</geoCoordinate>
,
<geoCoordinate id="EE1B500BFFD9FFAD78CF864CFAADB32A" box="[1190,1337,1822,1848]" degrees="101" direction="east" minutes="22" orientation="longitude" pageId="1" pageNumber="337" precision="15" seconds="24" value="101.37334">101°22ˊ24˝E</geoCoordinate>
, alt.
<quantity id="4CD79B29FFD9FFAD791B864CFF3BB34E" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="337" unit="m" value="740.0">
<elevation id="0002D1FFFFD9FFAD791B864CFF3BB34E" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="337" unit="m" value="740.0">740 m</elevation>
</quantity>
,
<date id="FF91100CFFD9FFAD7CD58610FEE9B34E" box="[188,381,1858,1884]" pageId="1" pageNumber="337" value="2017-10-21" valueMax="2017-10-22" valueMin="2017-10-21">
<collectingDate id="EFD5E9E4FFD9FFAD7CD58610FEE9B34E" box="[188,381,1858,1884]" pageId="1" pageNumber="337" value="2017-10-21" valueMax="2017-10-22" valueMin="2017-10-21">2122 Oct. 2017</collectingDate>
</date>
, np11279 (
<specimenCount id="9D29FD45FFD9FFAD7D958610FD87B34E" box="[508,531,1858,1884]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
).
<collectingRegion id="49EBF82EFFD9FFAD7E408610FD59B34E" box="[553,717,1858,1884]" country="Thailand" name="Sa Kaeo" pageId="1" pageNumber="337">Sa Kaeo Prov.</collectingRegion>
,
<collectorName id="26DA531AFFD9FFAD7EB08610FC10B34E" box="[729,900,1858,1884]" pageId="1" pageNumber="337">Pang Sida N.P.</collectorName>
,
<geoCoordinate id="EE1B500BFFD9FFAD7FF88610FB8EB34F" box="[913,1050,1858,1885]" degrees="14" direction="north" minutes="02" orientation="latitude" pageId="1" pageNumber="337" precision="15" seconds="25" value="14.040278">14°02ˊ25˝N</geoCoordinate>
,
<geoCoordinate id="EE1B500BFFD9FFAD784F8610FB2EB34F" box="[1062,1210,1858,1885]" degrees="102" direction="east" minutes="15" orientation="longitude" pageId="1" pageNumber="337" precision="15" seconds="56" value="102.26556">102°15ˊ56˝E</geoCoordinate>
, alt.
<quantity id="4CD79B29FFD9FFAD78988610FAA8B34E" box="[1265,1340,1858,1884]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="337" unit="m" value="310.0">
<elevation id="0002D1FFFFD9FFAD78988610FAA8B34E" box="[1265,1340,1858,1884]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="337" unit="m" value="310.0">310 m</elevation>
</quantity>
,
<date id="FF91100CFFD9FFAD79218610FF46B392" pageId="1" pageNumber="337" value="2017-10-18">
<collectingDate id="EFD5E9E4FFD9FFAD79218610FF46B392" pageId="1" pageNumber="337" value="2017-10-18">18 Oct. 2017</collectingDate>
</date>
, np9811 (
<specimenCount id="9D29FD45FFD9FFAD7D2B8634FECDB392" box="[322,345,1894,1920]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
, wing slide); same collection data as preceding,
<date id="FF91100CFFD9FFAD7F168634FB81B392" box="[895,1045,1894,1920]" pageId="1" pageNumber="337" value="2017-04-26">
<collectingDate id="EFD5E9E4FFD9FFAD7F168634FB81B392" box="[895,1045,1894,1920]" pageId="1" pageNumber="337" value="2017-04-26">26 Apr. 2017</collectingDate>
</date>
, np10020 (
<specimenCount id="9D29FD45FFD9FFAD78FD8634FB3FB392" box="[1172,1195,1894,1920]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
), np10034 (
<specimenCount id="9D29FD45FFD9FFAD795C8634FAD8B392" box="[1333,1356,1894,1920]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
); same collection data as preceding,
<date id="FF91100CFFD9FFAD7D8486D8FD1CB3B6" box="[493,648,1930,1956]" pageId="1" pageNumber="337" value="2017-06-21">
<collectingDate id="EFD5E9E4FFD9FFAD7D8486D8FD1CB3B6" box="[493,648,1930,1956]" pageId="1" pageNumber="337" value="2017-06-21">21 Jun. 2017</collectingDate>
</date>
, np10458 (
<specimenCount id="9D29FD45FFD9FFAD7F6786D8FCB1B3B6" box="[782,805,1930,1956]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
), np10471 (
<specimenCount id="9D29FD45FFD9FFAD7FDC86D8FC58B3B6" box="[949,972,1930,1956]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
); same collection data as preceding,
<date id="FF91100CFFD9FFAD79E986D8FE98B3DA" pageId="1" pageNumber="337" value="2017-10-18">
<collectingDate id="EFD5E9E4FFD9FFAD79E986D8FE98B3DA" pageId="1" pageNumber="337" value="2017-10-18">18 Oct. 2017</collectingDate>
</date>
, np11175 (
<specimenCount id="9D29FD45FFD9FFAD7DE686FCFE32B3DA" box="[399,422,1966,1992]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
);
<geoCoordinate id="EE1B500BFFD9FFAD7DA986FCFDDCB3DB" box="[448,584,1966,1993]" degrees="14" direction="north" minutes="00" orientation="latitude" pageId="1" pageNumber="337" precision="15" seconds="54" value="14.015">14°00ˊ54˝N</geoCoordinate>
,
<geoCoordinate id="EE1B500BFFD9FFAD7E3186FCFD7EB3DB" box="[600,746,1966,1993]" degrees="101" direction="east" minutes="11" orientation="longitude" pageId="1" pageNumber="337" precision="15" seconds="45" value="101.19583">101°11ˊ45˝E</geoCoordinate>
, alt.
<quantity id="4CD79B29FFD9FFAD7F4E86FCFCE0B3DA" box="[807,884,1966,1992]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="337" unit="m" value="260.0">
<elevation id="0002D1FFFFD9FFAD7F4E86FCFCE0B3DA" box="[807,884,1966,1992]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="337" unit="m" value="260.0">260 m</elevation>
</quantity>
,
<date id="FF91100CFFD9FFAD7FEA86FCFB8BB3DA" box="[899,1055,1966,1992]" pageId="1" pageNumber="337" value="2017-04-24">
<collectingDate id="EFD5E9E4FFD9FFAD7FEA86FCFB8BB3DA" box="[899,1055,1966,1992]" pageId="1" pageNumber="337" value="2017-04-24">24 Apr. 2017</collectingDate>
</date>
, np9906 (
<specimenCount id="9D29FD45FFD9FFAD78FF86FCFB39B3DA" box="[1174,1197,1966,1992]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
).
<collectingRegion id="49EBF82EFFD9FFAD78AF86FCFA0DB3DA" box="[1222,1433,1966,1992]" country="Thailand" name="Chanthaburi" pageId="1" pageNumber="337">Chanthaburi Prov.</collectingRegion>
,
<collectingMunicipality id="6BF4ACB6FFD9FFAD7CFE8680FEB4B3FE" box="[151,288,2002,2028]" pageId="1" pageNumber="337">Khao Glaur</collectingMunicipality>
,
<geoCoordinate id="EE1B500BFFD9FFAD7D478680FE22B3FF" box="[302,438,2002,2029]" degrees="13" direction="north" minutes="05" orientation="latitude" pageId="1" pageNumber="337" precision="15" seconds="49" value="13.096944">13°05ˊ49˝N</geoCoordinate>
,
<geoCoordinate id="EE1B500BFFD9FFAD7DAC8680FDCCB3FE" box="[453,600,2002,2028]" degrees="102" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="337" precision="15" seconds="21" value="102.1725">102°10ˊ21˝E</geoCoordinate>
, alt.
<quantity id="4CD79B29FFD9FFAD7EFD8680FD74B3FE" box="[660,736,2002,2028]" metricMagnitude="2" metricUnit="m" metricValue="1.66" pageId="1" pageNumber="337" unit="m" value="166.0">
<elevation id="0002D1FFFFD9FFAD7EFD8680FD74B3FE" box="[660,736,2002,2028]" metricMagnitude="2" metricUnit="m" metricValue="1.66" pageId="1" pageNumber="337" unit="m" value="166.0">166 m</elevation>
</quantity>
,
<date id="FF91100CFFD9FFAD7E868680FC19B3FE" box="[751,909,2002,2029]" pageId="1" pageNumber="337" value="2009-05-15">
<collectingDate id="EFD5E9E4FFD9FFAD7E868680FC19B3FE" box="[751,909,2002,2029]" pageId="1" pageNumber="337" value="2009-05-15">15 May 2009</collectingDate>
</date>
, np3107 (
<specimenCount id="9D29FD45FFD9FFAD786A8680FB8EB3FE" box="[1027,1050,2002,2028]" count="1" pageId="1" pageNumber="337" type="male"></specimenCount>
, genitalia slide NP2976), np3105 (
<specimenCount id="9D29FD45FFDAFFAE7CC881C5FF2CB4A3" box="[161,184,151,177]" count="1" pageId="2" pageNumber="338" type="male"></specimenCount>
).
<collectingRegion id="49EBF82EFFDAFFAE7CA481C5FED4B4A3" box="[205,320,151,177]" country="Thailand" name="Trat" pageId="2" pageNumber="338">Trat Prov.</collectingRegion>
,
<collectingRegion id="49EBF82EFFDAFFAE7D2281C5FEEDB4A3" box="[331,377,151,177]" country="Thailand" name="Trat" pageId="2" pageNumber="338">Trat</collectingRegion>
<location id="8EF06017FFDAFFAE7D1681C5FD4CB4A3" LSID="urn:lsid:plazi:treatment:038687DAFFD9FFAB7CFE84C3FBC1B027:8EF06017FFDAFFAE7D1681C5FD4CB4A3" box="[383,728,151,177]" country="Thailand" county="Bueng Khong Long Non-Hunting Area" latitude="12.392777" longLatPrecision="21" longitude="102.666664" municipality="Khao Glaur" name="Agroforestry Research Station" pageId="2" pageNumber="338" stateProvince="Bueng Kan">Agroforestry Research Station</location>
,
<geoCoordinate id="EE1B500BFFDAFFAE7E8D81C5FCF9B4A3" box="[740,877,151,177]" degrees="12" direction="north" minutes="23" orientation="latitude" pageId="2" pageNumber="338" precision="15" seconds="34" value="12.392777">12°23ˊ34˝N</geoCoordinate>
,
<geoCoordinate id="EE1B500BFFDAFFAE7F1081C5FB98B4A3" box="[889,1036,151,177]" degrees="102" direction="east" minutes="40" orientation="longitude" pageId="2" pageNumber="338" precision="15" seconds="00" value="102.666664">102°40ˊ00˝E</geoCoordinate>
, alt.
<quantity id="4CD79B29FFDAFFAE782D81C5FB14B4A3" box="[1092,1152,151,177]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="338" unit="m" value="46.0">
<elevation id="0002D1FFFFDAFFAE782D81C5FB14B4A3" box="[1092,1152,151,177]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="338" unit="m" value="46.0">46 m</elevation>
</quantity>
,
<date id="FF91100CFFDAFFAE78E581C5FADBB4A3" box="[1164,1359,151,177]" pageId="2" pageNumber="338" value="2010-09-16" valueMax="2010-09-19" valueMin="2010-09-16">
<collectingDate id="EFD5E9E4FFDAFFAE78E581C5FADBB4A3" box="[1164,1359,151,177]" pageId="2" pageNumber="338" value="2010-09-16" valueMax="2010-09-19" valueMin="2010-09-16">1619 Sep. 2010</collectingDate>
</date>
.
<collectorName id="26DA531AFFDAFFAE793081C5FA08B4A3" box="[1369,1436,151,177]" pageId="2" pageNumber="338">These</collectorName>
specimens were collected by
<collectorName id="26DA531AFFDAFFAE7D8981E9FDF7B4C7" box="[480,611,187,213]" pageId="2" pageNumber="338">N. Pinkaew</collectorName>
<emphasis id="B95BEADEFFDAFFAE7E0381E9FD08B4C7" box="[618,668,187,213]" italics="true" pageId="2" pageNumber="338">et al</emphasis>
</materialsCitation>
.
<materialsCitation id="3B473C91FFDAFFAE7EC381E9FE03B50F" collectingDate="2017-01-26" collectionCode="KKIC" collectorName="Phou Khao Khouay N. P. &amp; P. Phewphanh &amp; All" country="Laos" elevation="785" latitude="18.35" location="Vientiane Prov." pageId="2" pageNumber="338" specimenCount="2" specimenCount-male="2" stateProvince="Viangchan" typeStatus="paratype">
<specimenCount id="9D29FD45FFDAFFAE7EC381E9FD5AB4C7" box="[682,718,187,213]" count="1" pageId="2" pageNumber="338" type="male">1♂</specimenCount>
,
<collectingCountry id="F338765CFFDAFFAE7EB281E9FCBCB4C7" box="[731,808,187,213]" name="Laos" pageId="2" pageNumber="338">LAOS</collectingCountry>
:
<collectingRegion id="49EBF82EFFDAFFAE7F5A81E9FC77B4C7" box="[819,995,187,213]" country="Laos" name="Viangchan" pageId="2" pageNumber="338">Vientiane Prov.</collectingRegion>
,
<collectorName id="26DA531AFFDAFFAE7F8781E9FA97B4C7" box="[1006,1283,187,213]" pageId="2" pageNumber="338">Phou Khao Khouay N.P.</collectorName>
,
<geoCoordinate id="EE1B500BFFDAFFAE796681E9FA0CB4C7" box="[1295,1432,187,213]" degrees="18" direction="north" minutes="20" orientation="latitude" pageId="2" pageNumber="338" precision="15" seconds="60" value="18.35">18°20ˊ60˝N</geoCoordinate>
, 102°47ˊ92˝E, alt.
<quantity id="4CD79B29FFDAFFAE7D0A818CFE39B4EB" box="[355,429,222,249]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="2" pageNumber="338" unit="m" value="785.0">
<elevation id="0002D1FFFFDAFFAE7D0A818CFE39B4EB" box="[355,429,222,249]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="2" pageNumber="338" unit="m" value="785.0">785 m</elevation>
</quantity>
,
<date id="FF91100CFFDAFFAE7DD3818DFDDAB4EB" box="[442,590,223,249]" pageId="2" pageNumber="338" value="2017-01-26">
<collectingDate id="EFD5E9E4FFDAFFAE7DD3818DFDDAB4EB" box="[442,590,223,249]" pageId="2" pageNumber="338" value="2017-01-26">26 Jan. 2017</collectingDate>
</date>
,
<collectorName id="26DA531AFFDAFFAE7E32818DFD68B4EB" box="[603,764,223,249]" pageId="2" pageNumber="338">P. Phewphanh</collectorName>
, leg., pp00393 (
<specimenCount id="9D29FD45FFDAFFAE7FDC818DFC58B4EB" box="[949,972,223,249]" count="1" pageId="2" pageNumber="338" type="male"></specimenCount>
, genitalia slide PP00207).
<collectorName id="26DA531AFFDAFFAE7892818DFA8BB4EB" box="[1275,1311,223,249]" pageId="2" pageNumber="338">All</collectorName>
specimens are deposited in
<collectionCode id="ED3EAE09FFDAFFAE7D268051FE03B50F" box="[335,407,259,285]" pageId="2" pageNumber="338">KKIC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF506644FFDAFFAE7CFE8407FE8AB183" ID-DOI="http://doi.org/10.5281/zenodo.14033805" ID-Zenodo-Dep="14033805" httpUri="https://zenodo.org/record/14033805/files/figure.png" pageId="2" pageNumber="338" startId="2.[151,264,1365,1389]" targetBox="[333,1252,319,1326]" targetPageId="2" targetType="figure">
<paragraph id="8B9036CCFFDAFFAE7CFE8407FE8AB183" blockId="2.[151,1436,1365,1426]" pageId="2" pageNumber="338">
<emphasis id="B95BEADEFFDAFFAE7CFE8407FEA8B17F" bold="true" box="[151,316,1365,1389]" pageId="2" pageNumber="338">FIGURES 12.</emphasis>
Wing pattern of
<taxonomicName id="4C2F4D4FFFDAFFAE7D828407FD6FB17F" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" box="[491,763,1365,1389]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="338" phylum="Arthropoda" rank="species" species="squamapanna" status="sp. nov.">
<emphasis id="B95BEADEFFDAFFAE7D828407FD6FB17F" box="[491,763,1365,1389]" italics="true" pageId="2" pageNumber="338">Eudemopsis squamapanna</emphasis>
</taxonomicName>
<emphasis id="B95BEADEFFDAFFAE7F6A8404FCC0B17C" bold="true" box="[771,852,1366,1390]" pageId="2" pageNumber="338">
<taxonomicNameLabel id="A26857A5FFDAFFAE7F6A8404FCC0B17C" box="[771,852,1366,1390]" pageId="2" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1. Male (holotype); 2. Female (paratype, np7971). Scale bars = 2 mm.
</paragraph>
</caption>
<caption id="DF506644FFDAFFAE7CFE86B2FB6FB3EA" ID-DOI="http://doi.org/10.5281/zenodo.14033807" ID-Zenodo-Dep="14033807" box="[151,1275,2016,2040]" httpUri="https://zenodo.org/record/14033807/files/figure.png" pageId="2" pageNumber="338" startId="2.[151,264,2016,2040]" targetBox="[151,1435,1486,1992]" targetPageId="2" targetType="figure">
<paragraph id="8B9036CCFFDAFFAE7CFE86B2FB6FB3EA" blockId="2.[151,1275,2016,2040]" box="[151,1275,2016,2040]" pageId="2" pageNumber="338">
<emphasis id="B95BEADEFFDAFFAE7CFE86B2FEAFB3EA" bold="true" box="[151,315,2016,2040]" pageId="2" pageNumber="338">FIGURES 34.</emphasis>
Head of
<taxonomicName id="4C2F4D4FFFDAFFAE7DF186B2FD33B3EA" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" box="[408,679,2016,2040]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="338" phylum="Arthropoda" rank="species" species="squamapanna" status="sp. nov.">
<emphasis id="B95BEADEFFDAFFAE7DF186B2FD33B3EA" box="[408,679,2016,2040]" italics="true" pageId="2" pageNumber="338">Eudemopsis squamapanna</emphasis>
</taxonomicName>
<emphasis id="B95BEADEFFDAFFAE7EC486B2FD6AB3EA" bold="true" box="[685,766,2016,2040]" pageId="2" pageNumber="338">
<taxonomicNameLabel id="A26857A5FFDAFFAE7EC486B2FD6AB3EA" box="[685,766,2016,2040]" pageId="2" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
3. Male (holotype); 4. Female(paratype, np7971).
</paragraph>
</caption>
<caption id="DF506644FFDBFFAF7CFE82D7FECBB7D3" ID-DOI="http://doi.org/10.5281/zenodo.14033809" ID-Zenodo-Dep="14033809" httpUri="https://zenodo.org/record/14033809/files/figure.png" pageId="3" pageNumber="339" startId="3.[151,264,901,925]" targetBox="[166,1421,193,860]" targetPageId="3" targetType="figure">
<paragraph id="8B9036CCFFDBFFAF7CFE82D7FECBB7D3" blockId="3.[151,1437,901,961]" pageId="3" pageNumber="339">
<emphasis id="B95BEADEFFDBFFAF7CFE82D7FEAEB78F" bold="true" box="[151,314,901,925]" pageId="3" pageNumber="339">FIGURES 56.</emphasis>
Wing venation on the right wings of
<taxonomicName id="4C2F4D4FFFDBFFAF7EC682D7FC2AB78F" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" box="[687,958,901,925]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="species" species="squamapanna" status="sp. nov.">
<emphasis id="B95BEADEFFDBFFAF7EC682D7FC2AB78F" box="[687,958,901,925]" italics="true" pageId="3" pageNumber="339">Eudemopsis squamapanna</emphasis>
</taxonomicName>
<emphasis id="B95BEADEFFDBFFAF7FAA82D7FB87B78F" bold="true" box="[963,1043,901,925]" pageId="3" pageNumber="339">
<taxonomicNameLabel id="A26857A5FFDBFFAF7FAA82D7FB87B78F" box="[963,1043,901,925]" pageId="3" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
5. Male (paratype, np6618); 6. Female (paratype, np7971).
</paragraph>
</caption>
<caption id="DF506644FFDBFFAF7CFE8725FCD0B2A1" ID-DOI="http://doi.org/10.5281/zenodo.14033811" ID-Zenodo-Dep="14033811" httpUri="https://zenodo.org/record/14033811/files/figure.png" pageId="3" pageNumber="339" startId="3.[151,250,1655,1679]" targetBox="[185,1403,991,1628]" targetPageId="3" targetType="figure">
<paragraph id="8B9036CCFFDBFFAF7CFE8725FCD0B2A1" blockId="3.[151,1436,1655,1716]" pageId="3" pageNumber="339">
<emphasis id="B95BEADEFFDBFFAF7CFE8725FE86B282" bold="true" box="[151,274,1655,1680]" pageId="3" pageNumber="339">FIGURE 7.</emphasis>
Male abdomen of
<taxonomicName id="4C2F4D4FFFDBFFAF7DA7872AFD48B29D" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" box="[462,732,1656,1680]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="species" species="squamapanna" status="sp. nov.">
<emphasis id="B95BEADEFFDBFFAF7DA7872AFD48B29D" box="[462,732,1656,1680]" italics="true" pageId="3" pageNumber="339">Eudemopsis squamapanna</emphasis>
</taxonomicName>
<emphasis id="B95BEADEFFDBFFAF7E8B872AFCA5B282" bold="true" box="[738,817,1656,1680]" pageId="3" pageNumber="339">
<taxonomicNameLabel id="A26857A5FFDBFFAF7E8B872AFCA5B282" box="[738,817,1656,1680]" pageId="3" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with two small lobes of dense scale sockets on the abdominal segment IV anterolaterally (paratype, NP4167). Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="C3356547FFDBFFAF7CAE8654FBDAB3EA" pageId="3" pageNumber="339" type="diagnosis">
<paragraph id="8B9036CCFFDBFFAF7CAE8654FBDAB3EA" blockId="3.[151,1437,1798,2040]" pageId="3" pageNumber="339">
<emphasis id="B95BEADEFFDBFFAF7CAE8654FED6B332" bold="true" box="[199,322,1798,1824]" pageId="3" pageNumber="339">Diagnosis.</emphasis>
The forewing pattern of
<taxonomicName id="4C2F4D4FFFDBFFAF7E0A8655FCB5B332" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" box="[611,801,1798,1824]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="species" species="squamapanna">
<emphasis id="B95BEADEFFDBFFAF7E0A8655FDE0B332" box="[611,628,1799,1824]" italics="true" pageId="3" pageNumber="339">E</emphasis>
.
<emphasis id="B95BEADEFFDBFFAF7EED8655FCB5B332" box="[644,801,1799,1824]" italics="true" pageId="3" pageNumber="339">squamapanna</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C2F4D4FFFDBFFAF78648654FEBBB356" authority="(Heppner &amp; Bae, 2016)" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" baseAuthorityName="Pinkaew &amp; Muadsub &amp; Jaikla" baseAuthorityYear="2024" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="species" species="tamdaoensis" status="comb. nov.">
<emphasis id="B95BEADEFFDBFFAF78648654FABEB332" box="[1037,1322,1798,1824]" italics="true" pageId="3" pageNumber="339">Eudemopsis tamdaoensis</emphasis>
(
<bibRefCitation id="EFBE4B3DFFDBFFAF79538654FEBCB356" author="Heppner, J. B. &amp; Bae, Y. S." pageId="3" pageNumber="339" pagination="135 - 138" refId="ref4168" refString="Heppner, J. B. &amp; Bae, Y. S. (2016) A new species of Fansipaniana from Northern Vietnam (Lepidoptera: Tortricidae: Olethreutinae: Olethreutini). Zootaxa, 4097 (1), 135 - 138. https: // doi. org / 10.11646 / zootaxa. 4097.1.10" type="journal article" year="2016">Heppner &amp; Bae, 2016</bibRefCitation>
)
</taxonomicName>
<emphasis id="B95BEADEFFDBFFAF7D5F8678FE23B356" bold="true" box="[310,439,1834,1860]" pageId="3" pageNumber="339">
<taxonomicNameLabel id="A26857A5FFDBFFAF7D5F8678FE23B356" box="[310,439,1834,1860]" pageId="3" pageNumber="339" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, but it can be recognized by the brown mixed with dark brown scales on the frons, labial palp, tegulae and mesonotum, whereas the scales are greyish brown in
<taxonomicName id="4C2F4D4FFFDBFFAF7FBA861DFA09B37A" authority="(Heppner &amp; Bae, 2016)" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" baseAuthorityName="Pinkaew &amp; Muadsub &amp; Jaikla" baseAuthorityYear="2024" box="[979,1437,1870,1896]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="species" species="tamdaoensis" status="comb. nov.">
<emphasis id="B95BEADEFFDBFFAF7FBA861DFB15B37A" box="[979,1153,1870,1896]" italics="true" pageId="3" pageNumber="339">E. tamdaoensis</emphasis>
(
<bibRefCitation id="EFBE4B3DFFDBFFAF78FB861CFA01B37A" author="Heppner, J. B. &amp; Bae, Y. S." box="[1170,1429,1870,1896]" pageId="3" pageNumber="339" pagination="135 - 138" refId="ref4168" refString="Heppner, J. B. &amp; Bae, Y. S. (2016) A new species of Fansipaniana from Northern Vietnam (Lepidoptera: Tortricidae: Olethreutinae: Olethreutini). Zootaxa, 4097 (1), 135 - 138. https: // doi. org / 10.11646 / zootaxa. 4097.1.10" type="journal article" year="2016">Heppner &amp; Bae, 2016</bibRefCitation>
)
</taxonomicName>
<emphasis id="B95BEADEFFDBFFAF7CFE8620FE82B39E" bold="true" box="[151,278,1906,1932]" pageId="3" pageNumber="339">
<taxonomicNameLabel id="A26857A5FFDBFFAF7CFE8620FE82B39E" box="[151,278,1906,1932]" pageId="3" pageNumber="339" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
Male genitalia of this new species include a large patch of modified short scales and a large membranous lobe of very long modified scales on the anterior surface of the valva, which distinguish it from other
<taxonomicName id="4C2F4D4FFFDBFFAF797C86C4FF64B3C6" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95BEADEFFDBFFAF797C86C4FA08B3A2" box="[1301,1436,1942,1968]" italics="true" pageId="3" pageNumber="339">Eudemopsis</emphasis>
species.
</taxonomicName>
<taxonomicName id="4C2F4D4FFFDBFFAF7C9F86E8FD54B3C6" authority="(Razowski, 2009)" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" baseAuthorityName="Pinkaew &amp; Muadsub &amp; Jaikla" baseAuthorityYear="2024" box="[246,704,1978,2004]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="species" species="fansipana" status="comb. nov.">
<emphasis id="B95BEADEFFDBFFAF7C9F86E8FE66B3C6" box="[246,498,1978,2004]" italics="true" pageId="3" pageNumber="339">Eudemopsis fansipana</emphasis>
(
<bibRefCitation id="EFBE4B3DFFDBFFAF7E6886E8FD2CB3C6" author="Razowski, J." box="[513,696,1978,2004]" pageId="3" pageNumber="339" pagination="115 - 143" refId="ref4923" refString="Razowski, J. (2009) Tortricidae from Vietnam in the collection of the Berlin Museum. 6. Olethreutinae (Lepidoptera: Tortricidae). SHILAP Revista De Lepidopterologia, 37 (145), 115 - 143." type="journal article" year="2009">Razowski, 2009</bibRefCitation>
)
</taxonomicName>
<emphasis id="B95BEADEFFDBFFAF7EAE86E8FCD3B3C6" bold="true" box="[711,839,1978,2004]" pageId="3" pageNumber="339">
<taxonomicNameLabel id="A26857A5FFDBFFAF7EAE86E8FCD3B3C6" box="[711,839,1978,2004]" pageId="3" pageNumber="339" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
has only a patch of modified short scales, whereas
<taxonomicName id="4C2F4D4FFFDBFFAF79ED86E9FDA7B3EA" authority="(Heppner &amp; Bae, 2016)" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" baseAuthorityName="Pinkaew &amp; Muadsub &amp; Jaikla" baseAuthorityYear="2024" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="species" species="tamdaoensis" status="comb. nov.">
<emphasis id="B95BEADEFFDBFFAF79ED86E9FEB7B3EA" italics="true" pageId="3" pageNumber="339">E. tamdaoensis</emphasis>
(
<bibRefCitation id="EFBE4B3DFFDBFFAF7D5B868CFDBFB3EA" author="Heppner, J. B. &amp; Bae, Y. S." box="[306,555,2014,2040]" pageId="3" pageNumber="339" pagination="135 - 138" refId="ref4168" refString="Heppner, J. B. &amp; Bae, Y. S. (2016) A new species of Fansipaniana from Northern Vietnam (Lepidoptera: Tortricidae: Olethreutinae: Olethreutini). Zootaxa, 4097 (1), 135 - 138. https: // doi. org / 10.11646 / zootaxa. 4097.1.10" type="journal article" year="2016">Heppner &amp; Bae, 2016</bibRefCitation>
)
</taxonomicName>
<emphasis id="B95BEADEFFDBFFAF7E53868CFD2EB3EA" bold="true" box="[570,698,2014,2040]" pageId="3" pageNumber="339">
<taxonomicNameLabel id="A26857A5FFDBFFAF7E53868CFD2EB3EA" box="[570,698,2014,2040]" pageId="3" pageNumber="339" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
has only very long modified scales.
</paragraph>
</subSubSection>
<caption id="DF506644FFDCFFA87CFE8237FA07B76F" ID-DOI="http://doi.org/10.5281/zenodo.14033815" ID-Zenodo-Dep="14033815" box="[151,1427,869,894]" httpUri="https://zenodo.org/record/14033815/files/figure.png" pageId="4" pageNumber="340" startId="4.[151,250,869,893]" targetBox="[183,1409,191,835]" targetPageId="4" targetType="figure">
<paragraph id="8B9036CCFFDCFFA87CFE8237FA07B76F" blockId="4.[151,1427,869,894]" box="[151,1427,869,894]" pageId="4" pageNumber="340">
<emphasis id="B95BEADEFFDCFFA87CFE8237FE87B76C" bold="true" box="[151,275,869,894]" pageId="4" pageNumber="340">FIGURE 8.</emphasis>
Morphological features of male genitalia of
<taxonomicName id="4C2F4D4FFFDCFFA87EB08234FC7CB76F" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" box="[729,1000,870,894]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="340" phylum="Arthropoda" rank="species" species="squamapanna" status="sp. nov.">
<emphasis id="B95BEADEFFDCFFA87EB08234FC7CB76F" box="[729,1000,870,894]" italics="true" pageId="4" pageNumber="340">Eudemopsis squamapanna</emphasis>
</taxonomicName>
<emphasis id="B95BEADEFFDCFFA87F878234FBABB76C" bold="true" box="[1006,1087,870,894]" pageId="4" pageNumber="340">
<taxonomicNameLabel id="A26857A5FFDCFFA87F878234FBABB76C" box="[1006,1087,870,894]" pageId="4" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype). Scale bars = 0.5 mm.
</paragraph>
</caption>
<caption id="DF506644FFDCFFA87CFE8794FDA8B310" ID-DOI="http://doi.org/10.5281/zenodo.14033817" ID-Zenodo-Dep="14033817" httpUri="https://zenodo.org/record/14033817/files/figure.png" pageId="4" pageNumber="340" startId="4.[151,250,1734,1758]" targetBox="[579,1026,939,1709]" targetPageId="4" targetType="figure">
<paragraph id="8B9036CCFFDCFFA87CFE8794FDA8B310" blockId="4.[151,1436,1734,1794]" pageId="4" pageNumber="340">
<emphasis id="B95BEADEFFDCFFA87CFE8794FE80B2CC" bold="true" box="[151,276,1734,1758]" pageId="4" pageNumber="340">FIGURE 9.</emphasis>
Extended part of uncus, tuba analis, gnathos, phallus and anellus in male genitalia of
<taxonomicName id="4C2F4D4FFFDCFFA878E58794FA08B2CC" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" box="[1164,1436,1734,1758]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="340" phylum="Arthropoda" rank="species" species="squamapanna" status="sp. nov.">
<emphasis id="B95BEADEFFDCFFA878E58794FA08B2CC" box="[1164,1436,1734,1758]" italics="true" pageId="4" pageNumber="340">Eudemopsis squamapanna</emphasis>
</taxonomicName>
<emphasis id="B95BEADEFFDCFFA87CFE87B8FF7CB310" bold="true" box="[151,232,1770,1794]" pageId="4" pageNumber="340">
<taxonomicNameLabel id="A26857A5FFDCFFA87CFE87B8FF7CB310" box="[151,232,1770,1794]" pageId="4" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype). Scale bars = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3356547FFDCFFAB7CAE8606FA10B7B7" lastPageId="7" lastPageNumber="343" pageId="4" pageNumber="340" type="description">
<paragraph id="8B9036CCFFDCFFA87CAE8606FEB5B3C8" blockId="4.[151,1436,1876,2010]" pageId="4" pageNumber="340">
<emphasis id="B95BEADEFFDCFFA87CAE8606FECDB37C" bold="true" box="[199,345,1876,1902]" pageId="4" pageNumber="340">Description.</emphasis>
Head (
<figureCitation id="13142A49FFDCFFA87DC48606FD88B37D" box="[429,540,1876,1903]" captionStart="FIGURES 34" captionStartId="2.[151,264,2016,2040]" captionTargetBox="[151,1435,1486,1992]" captionTargetId="figure-97@2.[151,1435,1486,1992]" captionTargetPageId="2" captionText="FIGURES 34. Head of Eudemopsis squamapanna sp. nov. 3. Male (holotype); 4. Female(paratype, np7971)." figureDoi="http://doi.org/10.5281/zenodo.14033807" httpUri="https://zenodo.org/record/14033807/files/figure.png" pageId="4" pageNumber="340">Figs. 34</figureCitation>
): Lower frons brown mixed with dark purple, upper frons and vertex brown to dark brown mixed with dark purple; antenna light brown; labial palpus porrect, first segment dark purple, second segment curved upward, brown mixed with purple, with dark brown along ventral margin, apical segment short, light brown.
</paragraph>
<caption id="DF506644FFDDFFA97CFE8758FAB9B278" ID-DOI="http://doi.org/10.5281/zenodo.14033819" ID-Zenodo-Dep="14033819" httpUri="https://zenodo.org/record/14033819/files/figure.png" pageId="5" pageNumber="341" startId="5.[151,264,1546,1570]" targetBox="[222,1374,181,1509]" targetPageId="5" targetType="figure">
<paragraph id="8B9036CCFFDDFFA97CFE8758FAB9B278" blockId="5.[151,1437,1546,1642]" pageId="5" pageNumber="341">
<emphasis id="B95BEADEFFDDFFA97CFE8758FEC1B230" bold="true" box="[151,341,1546,1570]" pageId="5" pageNumber="341">FIGURES 1012.</emphasis>
Modified scales on the anterior surface on male valva of
<taxonomicName id="4C2F4D4FFFDDFFA97FC28758FB28B230" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" box="[939,1212,1546,1570]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="341" phylum="Arthropoda" rank="species" species="squamapanna" status="sp. nov.">
<emphasis id="B95BEADEFFDDFFA97FC28758FB28B230" box="[939,1212,1546,1570]" italics="true" pageId="5" pageNumber="341">Eudemopsis squamapanna</emphasis>
</taxonomicName>
<emphasis id="B95BEADEFFDDFFA978AD8758FA81B230" bold="true" box="[1220,1301,1546,1570]" pageId="5" pageNumber="341">
<taxonomicNameLabel id="A26857A5FFDDFFA978AD8758FA81B230" box="[1220,1301,1546,1570]" pageId="5" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
10. Exposed valva from the abdominal segment VIII (pinned specimen, paratype, np13911); 11. Valva after dyed in EosinY (holotype); 12. Mounted valva in Euparal (holotype) (A. Modified short scales B. Modified very long scales on membranous lobe).
</paragraph>
</caption>
<paragraph id="8B9036CCFFDDFFAA7CAE87EEFB51B673" blockId="5.[151,1437,1724,2039]" lastBlockId="6.[151,1437,151,615]" lastPageId="6" lastPageNumber="342" pageId="5" pageNumber="341">
Thorax: Pronotal collar dark purple; mesonotum light brown mixed with purple marks mediolaterally and posterolaterally; tegulae light brown with purple mixed with dark brown basal 1/3; posterior crest light brown. Forewing subtriangular, length 5.76.9 mm in males (n = 20) (
<figureCitation id="13142A49FFDDFFA97F018656FC3FB30C" box="[872,939,1796,1822]" captionStart="FIGURES 12" captionStartId="2.[151,264,1365,1389]" captionTargetBox="[333,1252,319,1326]" captionTargetId="figure-73@2.[322,1265,306,1338]" captionTargetPageId="2" captionText="FIGURES 12. Wing pattern of Eudemopsis squamapanna sp. nov. 1. Male (holotype); 2. Female (paratype, np7971). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14033805" httpUri="https://zenodo.org/record/14033805/files/figure.png" pageId="5" pageNumber="341">Fig. 1</figureCitation>
), 6.6 mm in female (n = 1) (
<figureCitation id="13142A49FFDDFFA978958656FAD7B30C" box="[1276,1347,1796,1822]" captionStart="FIGURES 12" captionStartId="2.[151,264,1365,1389]" captionTargetBox="[333,1252,319,1326]" captionTargetId="figure-73@2.[322,1265,306,1338]" captionTargetPageId="2" captionText="FIGURES 12. Wing pattern of Eudemopsis squamapanna sp. nov. 1. Male (holotype); 2. Female (paratype, np7971). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14033805" httpUri="https://zenodo.org/record/14033805/files/figure.png" pageId="5" pageNumber="341">Fig. 2</figureCitation>
); costal margin evenly curved, with a rather small pterostigma on costa between Sc and R
<quantity id="4CD79B29FFDDFFA978498668FBD0B350" box="[1056,1092,1832,1864]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="341" unit="in" value="1.0">
<subScript id="17AB3489FFDDFFA978498668FBBCB35A" attach="left" box="[1056,1064,1850,1864]" fontSize="6" pageId="5" pageNumber="341">1</subScript>
in
</quantity>
both male and female (distinct after removed scales) (
<figureCitation id="13142A49FFDDFFA97DF4861EFD98B375" box="[413,524,1868,1895]" captionStart="FIGURES 56" captionStartId="3.[151,264,901,925]" captionTargetBox="[166,1421,193,860]" captionTargetId="figure-29@3.[154,1434,181,877]" captionTargetPageId="3" captionText="FIGURES 56. Wing venation on the right wings of Eudemopsis squamapanna sp. nov. 5. Male (paratype, np6618); 6. Female (paratype, np7971)." figureDoi="http://doi.org/10.5281/zenodo.14033809" httpUri="https://zenodo.org/record/14033809/files/figure.png" pageId="5" pageNumber="341">Figs. 56</figureCitation>
); termen round, base of M
<subScript id="17AB3489FFDDFFA97F29860CFCDCB37E" attach="left" box="[832,840,1886,1900]" fontSize="6" pageId="5" pageNumber="341">2</subScript>
close to M
<subScript id="17AB3489FFDDFFA97FA5860CFC40B37E" attach="left" box="[972,980,1886,1900]" fontSize="6" pageId="5" pageNumber="341">3</subScript>
than M
<subScript id="17AB3489FFDDFFA97859860CFBACB37E" attach="left" box="[1072,1080,1886,1900]" fontSize="6" pageId="5" pageNumber="341">1</subScript>
, M
<subScript id="17AB3489FFDDFFA97808860CFBFDB37E" attach="left" box="[1121,1129,1886,1900]" fontSize="6" pageId="5" pageNumber="341">3</subScript>
curved downward basally, CuA
<subScript id="17AB3489FFDDFFA97CA586D0FF40B382" attach="left" box="[204,212,1922,1936]" fontSize="6" pageId="5" pageNumber="341">1</subScript>
curved downward medially, distal end of M
<subScript id="17AB3489FFDDFFA97EA986D0FD5CB382" attach="left" box="[704,712,1922,1936]" fontSize="6" pageId="5" pageNumber="341">1</subScript>
, M
<subScript id="17AB3489FFDDFFA97E8686D0FD63B382" attach="left" box="[751,759,1922,1936]" fontSize="6" pageId="5" pageNumber="341">2</subScript>
, M
<subScript id="17AB3489FFDDFFA97F7786D0FCB2B382" attach="left" box="[798,806,1922,1936]" fontSize="6" pageId="5" pageNumber="341">3</subScript>
, and CuA
<subScript id="17AB3489FFDDFFA97FFE86D0FC0BB382" attach="left" box="[919,927,1922,1936]" fontSize="6" pageId="5" pageNumber="341">1</subScript>
equidistant; costal strigulae brown, separated by dark brown streaks, strigulae 1 6 distinct, strigulae 7 9 indistinct; ground color yellowish brown to light brown, with scattered brown to dark brown spots; basal patch between wing base and strigula 1, extending downward as irregular band to near middle of dorsum, brown mixed with dark brown; subbasal fascia between strigulae 2 and 3, narrowed, extending from costa curve to dorsum, brown mixed with dark brown; median fascia between strigulae 4 and 5, moderately widened band, extending from costa beyond to middle of wing, distinct on costa, slightly narrowed beyond to middle of wing; postmedian fascia narrow, sinuate, extending from costa to near tornus, brown mixed with dark brown; preterminal fascia narrow, sinuate, extending slightly obliquely from costa to termen at M
<subScript id="17AB3489FFDEFFAA79F98047FA09B531" attach="left" box="[1424,1437,277,291]" fontSize="6" pageId="6" pageNumber="342">3,</subScript>
dark brown, with a short, longitudinal, narrowed band fused with postmedian and preterminal fascia at M
<subScript id="17AB3489FFDEFFAA7949806BFABAB555" attach="both" box="[1312,1326,313,327]" fontSize="6" pageId="6" pageNumber="342">2,</subScript>
apical 1/3 with a large, subcircular patch, extending from costa to M
<subScript id="17AB3489FFDEFFAA7F4B800FFCBEB579" attach="left" box="[802,810,349,363]" fontSize="6" pageId="6" pageNumber="342">2</subScript>
, pale brown, more bright anteriorly; tornal patch, large, subelliptic, between M
<subScript id="17AB3489FFDEFFAA7DF180D3FE34B59D" attach="left" box="[408,416,385,399]" fontSize="6" pageId="6" pageNumber="342">2</subScript>
to tornus, light brown, slightly paler anteriorly, beyond preterminal fascia with an oblique band, moderately wide, extending from costa to termen between M
<subScript id="17AB3489FFDEFFAA7FEB80F7FC1EB5A1" attach="left" box="[898,906,421,435]" fontSize="6" pageId="6" pageNumber="342">2</subScript>
and M
<subScript id="17AB3489FFDEFFAA7FB180F7FC74B5A1" attach="left" box="[984,992,421,435]" fontSize="6" pageId="6" pageNumber="342">3</subScript>
; underside with pale trace of upper side markings; fringe scales dark brown, except light brown between CuA
<subScript id="17AB3489FFDEFFAA7FD1809BFC54B5C5" attach="left" box="[952,960,457,471]" fontSize="6" pageId="6" pageNumber="342">1</subScript>
to tornus. Hindwing subtriangular, costal margin in male more convex than in female (
<figureCitation id="13142A49FFDEFFAA7EEC8089FD65B5E7" box="[645,753,474,501]" captionStart="FIGURES 56" captionStartId="3.[151,264,901,925]" captionTargetBox="[166,1421,193,860]" captionTargetId="figure-29@3.[154,1434,181,877]" captionTargetPageId="3" captionText="FIGURES 56. Wing venation on the right wings of Eudemopsis squamapanna sp. nov. 5. Male (paratype, np6618); 6. Female (paratype, np7971)." figureDoi="http://doi.org/10.5281/zenodo.14033809" httpUri="https://zenodo.org/record/14033809/files/figure.png" pageId="6" pageNumber="342">Figs. 56</figureCitation>
), greyish brown, slightly paler to wing base, wing apex with a distinct mark between costa and Sc+R
<subScript id="17AB3489FFDEFFAA7E298343FDDCB60D" attach="left" box="[576,584,529,543]" fontSize="6" pageId="6" pageNumber="342">1</subScript>
, pale brown, fringe scales greyish brown, slightly paler to anal angle, except brown at apex; underside light brown, slightly paler to wing base, with large longitudinal patch extending from wing base to apex and from costa to M
<subScript id="17AB3489FFDEFFAA7E65830BFD80B675" attach="left" box="[524,532,601,615]" fontSize="6" pageId="6" pageNumber="342">1</subScript>
yellowish white with patch of light purple scales median 1/3.
</paragraph>
<caption id="DF506644FFDEFFAA7CFE86D2FE94B3AE" ID-DOI="http://doi.org/10.5281/zenodo.14033821" ID-Zenodo-Dep="14033821" httpUri="https://zenodo.org/record/14033821/files/figure.png" pageId="6" pageNumber="342" startId="6.[151,250,1920,1944]" targetBox="[535,1047,645,1882]" targetPageId="6" targetType="figure">
<paragraph id="8B9036CCFFDEFFAA7CFE86D2FE94B3AE" blockId="6.[151,1436,1920,1980]" pageId="6" pageNumber="342">
<emphasis id="B95BEADEFFDEFFAA7CFE86D2FE8BB38A" bold="true" box="[151,287,1920,1944]" pageId="6" pageNumber="342">FIGURE 13.</emphasis>
Morphological features of female genitalia of
<taxonomicName id="4C2F4D4FFFDEFFAA7E9F86D2FB91B38A" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" box="[758,1029,1920,1944]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="342" phylum="Arthropoda" rank="species" species="squamapanna" status="sp. nov.">
<emphasis id="B95BEADEFFDEFFAA7E9F86D2FB91B38A" box="[758,1029,1920,1944]" italics="true" pageId="6" pageNumber="342">Eudemopsis squamapanna</emphasis>
</taxonomicName>
<emphasis id="B95BEADEFFDEFFAA786286D3FBCFB38B" bold="true" box="[1035,1115,1921,1945]" pageId="6" pageNumber="342">
<taxonomicNameLabel id="A26857A5FFDEFFAA786286D3FBCFB38B" box="[1035,1115,1921,1945]" pageId="6" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(paratype, NP3302). Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9036CCFFDFFFAB7CAE81C5FA10B7B7" blockId="7.[151,1437,151,1077]" pageId="7" pageNumber="343">
Abdomen (
<figureCitation id="13142A49FFDFFFAB7D2E81C5FE1BB4A3" box="[327,399,151,177]" captionStart="FIGURE 7" captionStartId="3.[151,250,1655,1679]" captionTargetBox="[185,1403,991,1628]" captionTargetId="figure-53@3.[172,1416,986,1632]" captionTargetPageId="3" captionText="FIGURE 7. Male abdomen of Eudemopsis squamapanna sp. nov. with two small lobes of dense scale sockets on the abdominal segment IV anterolaterally (paratype, NP4167). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033811" httpUri="https://zenodo.org/record/14033811/files/figure.png" pageId="7" pageNumber="343">Fig. 7</figureCitation>
): Segment IV with two small membranous lobes anterolaterally, with dense scale sockets. Membrane beyond abdominal segment VIII enlarged. Male genitalia (
<figureCitation id="13142A49FFDFFFAB7FC181E9FBB4B4C7" box="[936,1056,187,213]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURES 1012" captionStartId-0="4.[151,250,869,893]" captionStartId-1="4.[151,250,1734,1758]" captionStartId-2="5.[151,264,1546,1570]" captionTargetBox-0="[183,1409,191,835]" captionTargetBox-1="[579,1026,939,1709]" captionTargetBox-2="[222,1374,181,1509]" captionTargetId-0="figure-18@4.[175,1412,181,846]" captionTargetId-1="figure-39@4.[562,1026,918,1710]" captionTargetId-2="figure-29@5.[212,1375,181,1521]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 8. Morphological features of male genitalia of Eudemopsis squamapanna sp. nov. (holotype). Scale bars = 0.5 mm." captionText-1="FIGURE 9. Extended part of uncus, tuba analis, gnathos, phallus and anellus in male genitalia of Eudemopsis squamapanna sp. nov. (holotype). Scale bars = 0.5 mm." captionText-2="FIGURES 1012. Modified scales on the anterior surface on male valva of Eudemopsis squamapanna sp. nov. 10. Exposed valva from the abdominal segment VIII (pinned specimen, paratype, np13911); 11. Valva after dyed in EosinY (holotype); 12. Mounted valva in Euparal (holotype) (A. Modified short scales B. Modified very long scales on membranous lobe)." figureDoi-0="http://doi.org/10.5281/zenodo.14033815" figureDoi-1="http://doi.org/10.5281/zenodo.14033817" figureDoi-2="http://doi.org/10.5281/zenodo.14033819" httpUri-0="https://zenodo.org/record/14033815/files/figure.png" httpUri-1="https://zenodo.org/record/14033817/files/figure.png" httpUri-2="https://zenodo.org/record/14033819/files/figure.png" pageId="7" pageNumber="343">Figs. 812</figureCitation>
) with tegumen moderately broad, oblong, bilobed apex, with dense scale sockets laterally, apodeme of muscle m4 rather small, subtriangular lobe; uncus ovate, rather small, with sparse setae; socii reduced; gnathos arising from near lateral midpoint of tegumen, completely fused medially as strongly sclerotized band, densely microtrichiate, dorsomedially with an extended sclerotized, rather narrow, tubular median prominence to tuba analis, long, slightly narrowed to apex, densely covered with microtrichia; vinculum moderately wide; juxta subtriangular, rather small; caulis moderately long; anellus surrounding basal 1/2 of phallus, with two groups of dense microtrichia laterally; phallus subtubular, rather short and moderately wide, slightly narrowed towards apex; cornuti absent; valva simple, without neck, dorsal margin rather straight, basal 2/3 broadened, margins rather parallel, apical 1/3 with ventral margin slightly narrowed towards apex, apex rather wide and round, posterior surface with a small group of dense setae basally, medially with a widened group of short, sparsely setose, apical 2/3 clothed with short setae, ventromedially with a cluster of moderately dense and long, strong spiniform setae, anterior surface at basal 1/4 with a large membranous lobe of dense, very long setae (
<figureCitation id="13142A49FFDFFFAB7DF58339FDDDB697" box="[412,585,619,645]" captionStart="FIGURES 1012" captionStartId="5.[151,264,1546,1570]" captionTargetBox="[222,1374,181,1509]" captionTargetId="figure-29@5.[212,1375,181,1521]" captionTargetPageId="5" captionText="FIGURES 1012. Modified scales on the anterior surface on male valva of Eudemopsis squamapanna sp. nov. 10. Exposed valva from the abdominal segment VIII (pinned specimen, paratype, np13911); 11. Valva after dyed in EosinY (holotype); 12. Mounted valva in Euparal (holotype) (A. Modified short scales B. Modified very long scales on membranous lobe)." figureDoi="http://doi.org/10.5281/zenodo.14033819" httpUri="https://zenodo.org/record/14033819/files/figure.png" pageId="7" pageNumber="343">Figs. 10B12B</figureCitation>
), white in pinned specimens (
<figureCitation id="13142A49FFDFFFAB7FFD8339FC6DB697" box="[916,1017,619,645]" captionStart="FIGURES 1012" captionStartId="5.[151,264,1546,1570]" captionTargetBox="[222,1374,181,1509]" captionTargetId="figure-29@5.[212,1375,181,1521]" captionTargetPageId="5" captionText="FIGURES 1012. Modified scales on the anterior surface on male valva of Eudemopsis squamapanna sp. nov. 10. Exposed valva from the abdominal segment VIII (pinned specimen, paratype, np13911); 11. Valva after dyed in EosinY (holotype); 12. Mounted valva in Euparal (holotype) (A. Modified short scales B. Modified very long scales on membranous lobe)." figureDoi="http://doi.org/10.5281/zenodo.14033819" httpUri="https://zenodo.org/record/14033819/files/figure.png" pageId="7" pageNumber="343">Fig. 10B</figureCitation>
), apical 3/4 with a large patch of very dense, rather short, modified scales (
<figureCitation id="13142A49FFDFFFAB7E4783DDFD4BB6BB" box="[558,735,655,681]" captionStart="FIGURES 1012" captionStartId="5.[151,264,1546,1570]" captionTargetBox="[222,1374,181,1509]" captionTargetId="figure-29@5.[212,1375,181,1521]" captionTargetPageId="5" captionText="FIGURES 1012. Modified scales on the anterior surface on male valva of Eudemopsis squamapanna sp. nov. 10. Exposed valva from the abdominal segment VIII (pinned specimen, paratype, np13911); 11. Valva after dyed in EosinY (holotype); 12. Mounted valva in Euparal (holotype) (A. Modified short scales B. Modified very long scales on membranous lobe)." figureDoi="http://doi.org/10.5281/zenodo.14033819" httpUri="https://zenodo.org/record/14033819/files/figure.png" pageId="7" pageNumber="343">Figs. 10A12A</figureCitation>
), dark brown in pinned specimens (
<figureCitation id="13142A49FFDFFFAB780083DDFB44B6BB" box="[1129,1232,655,681]" captionStart="FIGURES 1012" captionStartId="5.[151,264,1546,1570]" captionTargetBox="[222,1374,181,1509]" captionTargetId="figure-29@5.[212,1375,181,1521]" captionTargetPageId="5" captionText="FIGURES 1012. Modified scales on the anterior surface on male valva of Eudemopsis squamapanna sp. nov. 10. Exposed valva from the abdominal segment VIII (pinned specimen, paratype, np13911); 11. Valva after dyed in EosinY (holotype); 12. Mounted valva in Euparal (holotype) (A. Modified short scales B. Modified very long scales on membranous lobe)." figureDoi="http://doi.org/10.5281/zenodo.14033819" httpUri="https://zenodo.org/record/14033819/files/figure.png" pageId="7" pageNumber="343">Fig. 10A</figureCitation>
). Female genitalia (
<figureCitation id="13142A49FFDFFFAB7CF683E1FF64B6DF" box="[159,240,691,717]" captionStart="FIGURE 13" captionStartId="6.[151,250,1920,1944]" captionTargetBox="[535,1047,645,1882]" captionTargetId="figure-259@6.[520,1066,633,1897]" captionTargetPageId="6" captionText="FIGURE 13. Morphological features of female genitalia of Eudemopsis squamapanna sp. nov. (paratype, NP3302). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14033821" httpUri="https://zenodo.org/record/14033821/files/figure.png" pageId="7" pageNumber="343">Fig. 13</figureCitation>
) with papillae anales moderately broad, densely setose; tergum VIII moderately sclerotized, without scale socket, subtriangular lateral extension densely microtrichiate; sternum VII moderately sclerotized, with dense scale sockets, more dense posteriorly, posterior margin shallowly concave, slightly sinuate; ostium bursae elongate, beyond posterior margin of sternum VII, surrounded with large, sclerotized, subcircular sterigma, covered with dense microtrichia, posteromedially with a moderately large patch, densely microtrichiate; ductus bursae 1/2 length of corpus bursae, anterior 1/3 narrowed, posterior 2/3 widened, colliculum ill-defined; ductus seminalis arising from anterior 1/3 of ductus bursae; corpus bursae large, ovate, with two moderately large, curved, blade-shaped signa.
</paragraph>
</subSubSection>
<subSubSection id="C3356547FFDFFFAB7CAE82FCFDE7B7FF" pageId="7" pageNumber="343" type="distribution">
<paragraph id="8B9036CCFFDFFFAB7CAE82FCFDE7B7FF" blockId="7.[151,1437,151,1077]" pageId="7" pageNumber="343">
<emphasis id="B95BEADEFFDFFFAB7CAE82FCFECDB7DA" bold="true" box="[199,345,942,968]" pageId="7" pageNumber="343">Distribution</emphasis>
.
<collectingCountry id="F338765CFFDFFFAB7D0282FDFE5BB7DB" box="[363,463,943,969]" name="Thailand" pageId="7" pageNumber="343">Thailand</collectingCountry>
(
<collectingRegion id="49EBF82EFFDFFFAB7D8982FDFDF3B7DB" box="[480,615,943,969]" country="Thailand" name="Bueng Kan" pageId="7" pageNumber="343">Bueng Kan</collectingRegion>
,
<collectingRegion id="49EBF82EFFDFFFAB7E1F82FDFCCBB7DB" box="[630,863,943,969]" country="Thailand" name="Nakhon Ratchasima" pageId="7" pageNumber="343">Nakhon Ratchasima</collectingRegion>
,
<collectingRegion id="49EBF82EFFDFFFAB7F0482FDFB8BB7DB" box="[877,1055,943,969]" country="Thailand" name="Nakhon Nayok" pageId="7" pageNumber="343">Nakhon Nayok</collectingRegion>
,
<collectingRegion id="49EBF82EFFDFFFAB784482FDFB05B7DB" box="[1069,1169,943,969]" country="Thailand" name="Sa Kaeo" pageId="7" pageNumber="343">Sa Kaeo</collectingRegion>
,
<collectingRegion id="49EBF82EFFDFFFAB78C982FDFAB9B7DB" box="[1184,1325,943,969]" country="Thailand" name="Chanthaburi" pageId="7" pageNumber="343">Chanthaburi</collectingRegion>
, and
<collectingRegion id="49EBF82EFFDFFFAB790682FDFA09B7DB" box="[1391,1437,943,969]" country="Thailand" name="Trat" pageId="7" pageNumber="343">Trat</collectingRegion>
Provinces) and
<collectingCountry id="F338765CFFDFFFAB7D2C8281FEEFB7FF" box="[325,379,979,1005]" name="Laos" pageId="7" pageNumber="343">Laos</collectingCountry>
(
<collectingRegion id="49EBF82EFFDFFFAB7DE38281FDF3B7FF" box="[394,615,979,1005]" country="Laos" name="Viangchan" pageId="7" pageNumber="343">Vientiane Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3356547FFDFFFAB7CAE82A5FBC1B027" pageId="7" pageNumber="343" type="etymology">
<paragraph id="8B9036CCFFDFFFAB7CAE82A5FBC1B027" blockId="7.[151,1437,151,1077]" pageId="7" pageNumber="343">
<emphasis id="B95BEADEFFDFFFAB7CAE82A5FEDCB003" bold="true" box="[199,328,1015,1041]" pageId="7" pageNumber="343">Etymology</emphasis>
. The specific epithet
<taxonomicName id="4C2F4D4FFFDFFFAB7E5082A5FD42B002" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" box="[569,726,1015,1040]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="343" phylum="Arthropoda" rank="species" species="squamapanna">
<emphasis id="B95BEADEFFDFFFAB7E5082A5FD42B002" box="[569,726,1015,1040]" italics="true" pageId="7" pageNumber="343">squamapanna</emphasis>
</taxonomicName>
comes from two Latinized words, squama = scale, and panna = patch, and refers to the large patch of short scales on the anterior surface of the valva.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,110 @@
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<mods:title id="930CA944A6C9D21AF2B9643FDE73BE85">A new species of Eudemopsis Falkovitsh (Lepidoptera: Tortricidae: Olethreutinae: Olethreutini) from Thailand and Laos, with the proposed synonymy of Fansipaniana Razowski</mods:title>
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<mods:namePart id="4EBCFD89A8FC1A97D4275D496C9AD8FC">Muadsub, Sopita</mods:namePart>
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<mods:nameIdentifier id="1B758E50046B68741BEF13C4E24D3628" type="email">muadsub_s@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="98CB07464B0C077C0ED09F9E5E7A7E5C">Jaikla, Soraya</mods:namePart>
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<heading id="D0D881A0FFD9FFAD7CFE8596FC11B0CD" bold="true" box="[151,901,1220,1247]" fontSize="11" level="1" pageId="1" pageNumber="337" reason="1">
<emphasis id="B95BEADEFFD9FFAD7CFE8596FC11B0CD" bold="true" box="[151,901,1220,1247]" pageId="1" pageNumber="337">
<taxonomicName id="4C2F4D4FFFD9FFAD7CFE8596FD65B0CC" authority="(Heppner &amp; Bae, 2016)" authorityName="Pinkaew &amp; Muadsub &amp; Jaikla" authorityYear="2024" baseAuthorityName="Pinkaew &amp; Muadsub &amp; Jaikla" baseAuthorityYear="2024" box="[151,753,1220,1247]" class="Insecta" family="Tortricidae" genus="Eudemopsis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="337" phylum="Arthropoda" rank="species" species="tamdaoensis" status="comb. nov.">
<emphasis id="B95BEADEFFD9FFAD7CFE8596FE53B0CD" bold="true" box="[151,455,1220,1247]" italics="true" pageId="1" pageNumber="337">Eudemopsis tamdaoensis</emphasis>
(
<bibRefCitation id="EFBE4B3DFFD9FFAD7DBF8596FD7EB0CC" author="Heppner, J. B. &amp; Bae, Y. S." box="[470,746,1220,1247]" pageId="1" pageNumber="337" pagination="135 - 138" refId="ref4168" refString="Heppner, J. B. &amp; Bae, Y. S. (2016) A new species of Fansipaniana from Northern Vietnam (Lepidoptera: Tortricidae: Olethreutinae: Olethreutini). Zootaxa, 4097 (1), 135 - 138. https: // doi. org / 10.11646 / zootaxa. 4097.1.10" type="journal article" year="2016">Heppner &amp; Bae, 2016</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A26857A5FFD9FFAD7E968597FC11B0CD" box="[767,901,1221,1247]" pageId="1" pageNumber="337" rank="species">comb. nov.</taxonomicNameLabel>
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<emphasis id="B95BEADEFFD9FFAD7CFE8458FE3EB133" box="[151,426,1289,1313]" italics="true" pageId="1" pageNumber="337">Fansipaniana tamdaoensis</emphasis>
</taxonomicName>
:
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<bibRefCitation id="EFBE4B3DFFD9FFAD7DD0845BFD25B133" author="Heppner, J. B. &amp; Bae, Y. S." box="[441,689,1289,1313]" pageId="1" pageNumber="337" pagination="135 - 138" refId="ref4168" refString="Heppner, J. B. &amp; Bae, Y. S. (2016) A new species of Fansipaniana from Northern Vietnam (Lepidoptera: Tortricidae: Olethreutinae: Olethreutini). Zootaxa, 4097 (1), 135 - 138. https: // doi. org / 10.11646 / zootaxa. 4097.1.10" type="journal article" year="2016">Heppner &amp; Bae 2016: 1</bibRefCitation>
</treatmentCitation>
. Type locality:
<collectingCountry id="F338765CFFD9FFAD7F38845BFC33B133" box="[849,935,1289,1313]" name="Vietnam" pageId="1" pageNumber="337">Vietnam</collectingCountry>
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,
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”.
<typeStatus id="5494886EFFD9FFAD7962845BFAFDB133" box="[1291,1385,1289,1313]" pageId="1" pageNumber="337" type="holotype">Holotype</typeStatus>
(
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):
<collectionCode id="ED3EAE09FFD9FFAD7CAE847AFE93B152" box="[199,263,1320,1344]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="337" type="Museum">FSCA</collectionCode>
/
<collectionCode id="ED3EAE09FFD9FFAD7D66847AFECEB152" box="[271,346,1320,1344]" pageId="1" pageNumber="337">MGCL</collectionCode>
.
</materialsCitation>
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View file

@ -0,0 +1,429 @@
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<mods:title id="F758D83E8565F92F67E9E4F282057E30">A new genus for the Blackfin Sucker, Thoburnia atripinnis (Cypriniformes: Catostomidae)</mods:title>
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<taxonomicName id="4C07D3F8465BC276FF64F9BFFE1EF9A9" authority="Armbruster" authorityName="Armbruster" authorityYear="2024" box="[151,463,1571,1598]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="327" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B9737469465BC276FF64F9BFFEE3F9A9" bold="true" box="[151,306,1571,1598]" italics="true" pageId="2" pageNumber="327">Vexillichthys</emphasis>
Armbruster
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,
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<paragraph id="8BB8A87B465BC276FF64F9F0FF29F911" blockId="2.[151,967,1571,1670]" box="[151,248,1644,1670]" pageId="2" pageNumber="327">
<figureCitation id="133CB4FE465BC276FF64F9F0FF29F911" box="[151,248,1644,1670]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,846,870]" captionStartId-1="4.[151,250,1277,1301]" captionTargetBox-0="[340,1247,179,820]" captionTargetBox-1="[177,1411,181,1253]" captionTargetId-0="figure-15@3.[340,1247,179,820]" captionTargetId-1="figure-23@4.[177,1411,181,1253]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Live specimens of Vexillichthys n. gen. atripinnis. A. young in typical coloration (90 mm SL, Salt Lick Creek at Bugtussle Road crossing, Monroe Co., KY, 36.65654°, -85.92124°) and B. nuptial specimen, probably female (147 mm SL, Long Hungry Creek, at KY 1333 crossing, Allen Co., KY, 36.67412°, -86.04397°). Specimens uncataloged. Photos by M. Thomas." captionText-1="FIGURE 2. Preserved nuptial male specimen of Vexillichthys n. gen. atripinnis in dorsal, lateral, and ventral views. AUM 12812, 71.6 mm SL. Photos by J.W. Armbruster. Scale = 1 cm." figureDoi-0="http://doi.org/10.5281/zenodo.14033793" figureDoi-1="http://doi.org/10.5281/zenodo.14033795" httpUri-0="https://zenodo.org/record/14033793/files/figure.png" httpUri-1="https://zenodo.org/record/14033795/files/figure.png" pageId="2" pageNumber="327">Figs 12</figureCitation>
</paragraph>
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<paragraph id="8BB8A87B465BC276FF64F928FC81F958" blockId="2.[151,1437,1716,1958]" box="[151,848,1716,1743]" pageId="2" pageNumber="327">
<emphasis id="B9737469465BC276FF64F928FEE3F959" bold="true" box="[151,306,1716,1742]" pageId="2" pageNumber="327">
<typeStatus id="54BC16D9465BC276FF64F928FF03F959" box="[151,210,1716,1742]" pageId="2" pageNumber="327">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C07D3F8465BC276FECAF929FC9DF958" ID-CoL="44GTS" authority="Bailey 1959" authorityName="Bailey" authorityYear="1959" box="[313,844,1716,1743]" family="Catostomidae" genus="Moxostoma" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="327" phylum="Chordata" rank="species" species="atripinne" subGenus="Thoburnia">
<emphasis id="B9737469465BC276FECAF929FE6DF959" box="[313,444,1717,1742]" italics="true" pageId="2" pageNumber="327">Moxostoma</emphasis>
(
<emphasis id="B9737469465BC276FE3FF928FD93F959" box="[460,578,1716,1742]" italics="true" pageId="2" pageNumber="327">Thoburnia</emphasis>
)
<emphasis id="B9737469465BC276FDA0F929FD6BF959" box="[595,698,1717,1742]" italics="true" pageId="2" pageNumber="327">atripinne</emphasis>
<bibRefCitation id="EF96D58A465BC276FD33F928FC9DF958" author="Bailey, R. M." box="[704,844,1716,1743]" pageId="2" pageNumber="327" pagination="1 - 19" refId="ref5656" refString="Bailey, R. M. (1959) A new catostomid fish, Moxostoma (Thoburnia) atripinne, from the Green River drainage, Kentucky and Tennessee. Occasional Papers of the Museum of Zoology, The University of Michigan, 559, 1 - 19." type="journal article" year="1959">Bailey 1959</bibRefCitation>
</taxonomicName>
.
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<paragraph id="8BB8A87B465BC276FF34F944FABDF8AC" blockId="2.[151,1437,1716,1958]" pageId="2" pageNumber="327">
<emphasis id="B9737469465BC276FF34F944FE43F965" bold="true" box="[199,402,1752,1778]" pageId="2" pageNumber="327">Included species.</emphasis>
<taxonomicName id="4C07D3F8465BC276FE6AF945FC11F964" ID-CoL="44GTS" authority="Bailey 1959: 7" authorityName="Bailey" authorityPageNumber="7" authorityYear="1959" box="[409,960,1752,1779]" family="Catostomidae" genus="Moxostoma" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="327" phylum="Chordata" rank="species" species="atripinne" subGenus="Thoburnia">
<emphasis id="B9737469465BC276FE6AF945FDCDF965" box="[409,540,1753,1778]" italics="true" pageId="2" pageNumber="327">Moxostoma</emphasis>
(
<emphasis id="B9737469465BC276FDDFF944FD73F965" box="[556,674,1752,1778]" italics="true" pageId="2" pageNumber="327">Thoburnia</emphasis>
)
<emphasis id="B9737469465BC276FD41F945FCC8F965" box="[690,793,1753,1778]" italics="true" pageId="2" pageNumber="327">atripinne</emphasis>
<materialsCitation id="3B6FA226465BC276FCD3F944FC11F964" box="[800,960,1752,1779]" collectionCode="UMMZ" country="United States of America" county="Macon County" elevation="233" location="Salt Lick Creek" municipality="Red Boiling Springs" pageId="2" pageNumber="327" specimenCode="UMMZ 169506, UMMZ 165370" specimenCount="2" stateProvince="Tennessee" typeStatus="holotype">
<bibRefCitation id="EF96D58A465BC276FCD3F944FC11F964" author="Bailey, R. M." box="[800,960,1752,1779]" pageId="2" pageNumber="327" pagination="1 - 19" refId="ref5656" refString="Bailey, R. M. (1959) A new catostomid fish, Moxostoma (Thoburnia) atripinne, from the Green River drainage, Kentucky and Tennessee. Occasional Papers of the Museum of Zoology, The University of Michigan, 559, 1 - 19." type="journal article" year="1959">Bailey 1959:7</bibRefCitation>
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, Pls. 12;
<figureCitation id="133CB4FE465BC276FBC6F944FB63F965" box="[1077,1202,1752,1778]" captionStart="FIGURE 1" captionStartId="3.[151,250,846,870]" captionTargetBox="[340,1247,179,820]" captionTargetId="figure-15@3.[340,1247,179,820]" captionTargetPageId="3" captionText="FIGURE 1. Live specimens of Vexillichthys n. gen. atripinnis. A. young in typical coloration (90 mm SL, Salt Lick Creek at Bugtussle Road crossing, Monroe Co., KY, 36.65654°, -85.92124°) and B. nuptial specimen, probably female (147 mm SL, Long Hungry Creek, at KY 1333 crossing, Allen Co., KY, 36.67412°, -86.04397°). Specimens uncataloged. Photos by M. Thomas." figureDoi="http://doi.org/10.5281/zenodo.14033793" httpUri="https://zenodo.org/record/14033793/files/figure.png" pageId="2" pageNumber="327">Figs.1BD</figureCitation>
,
<figureCitation id="133CB4FE465BC276FB4DF944FB31F965" box="[1214,1248,1752,1778]" captionStart="FIGURE 2" captionStartId="4.[151,250,1277,1301]" captionTargetBox="[177,1411,181,1253]" captionTargetId="figure-23@4.[177,1411,181,1253]" captionTargetPageId="4" captionText="FIGURE 2. Preserved nuptial male specimen of Vexillichthys n. gen. atripinnis in dorsal, lateral, and ventral views. AUM 12812, 71.6 mm SL. Photos by J.W. Armbruster. Scale = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14033795" httpUri="https://zenodo.org/record/14033795/files/figure.png" pageId="2" pageNumber="327">2C</figureCitation>
,
<figureCitation id="133CB4FE465BC276FB1FF944FAFEF965" box="[1260,1327,1752,1778]" captionStart="FIGURE 3" captionStartId="5.[151,250,1009,1033]" captionTargetBox="[168,1423,181,981]" captionTargetId="figure-15@5.[164,1423,181,985]" captionTargetPageId="5" captionText="FIGURE 3. Distribution of Vexillichthys n. gen. atripinnis." figureDoi="http://doi.org/10.5281/zenodo.14033797" httpUri="https://zenodo.org/record/14033797/files/figure.png" pageId="2" pageNumber="327">3AB</figureCitation>
<location id="8ED8FEA0465BC276FAC5F944FF0CF881" LSID="urn:lsid:plazi:treatment:03AE196D465BC272FF64F9BFFEBBF879:8ED8FEA0465BC276FAC5F944FF0CF881" country="United States of America" county="Macon County" municipality="Red Boiling Springs" name="Salt Lick Creek" pageId="2" pageNumber="327" stateProvince="Tennessee">Salt Lick Creek</location>
, at state highway 52 bridge,
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,
<collectingCounty id="62D9D0F7465BC276FCDFF961FC09F881" box="[812,984,1788,1815]" pageId="2" pageNumber="327">Macon County</collectingCounty>
,
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,
<collectingCountry id="F310E8EB465BC276FB9BF960FB66F881" box="[1128,1207,1788,1814]" name="United States of America" pageId="2" pageNumber="327">U.S.A.</collectingCountry>
,
<elevation id="002A4F48465BC276FB34F960FA48F881" box="[1223,1433,1788,1815]" metricMagnitude="2" metricUnit="m" metricValue="2.33172" pageId="2" pageNumber="327" unit="ft" value="765.0">
elevation
<quantity id="4CFF059E465BC276FAC4F960FA48F881" box="[1335,1433,1788,1815]" metricMagnitude="2" metricUnit="m" metricValue="2.33172" pageId="2" pageNumber="327" unit="ft" value="765.0">765 feet</quantity>
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.
<typeStatus id="54BC16D9465BC276FF64F8BCFED5F8AD" box="[151,260,1824,1850]" pageId="2" pageNumber="327" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBA10000465BC276FEFDF8BDFE11F8AC" box="[270,448,1824,1851]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="2" pageNumber="327" type="University or college">UMMZ 169506</specimenCode>
(image examined).
<typeStatus id="54BC16D9465BC276FD53F8BDFCC0F8AC" box="[672,785,1825,1851]" pageId="2" pageNumber="327" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="DBA10000465BC276FCEFF8BDFC1FF8AC" box="[796,974,1824,1851]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="2" pageNumber="327" type="University or college">UMMZ 165370</specimenCode>
(6). Valid as
<taxonomicName id="4C07D3F8465BC276FB97F8BCFAB4F8AD" box="[1124,1381,1824,1850]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="327" phylum="Chordata" rank="species" species="atripinnis">
<emphasis id="B9737469465BC276FB97F8BCFAB4F8AD" box="[1124,1381,1824,1850]" italics="true" pageId="2" pageNumber="327">Vexillichthys atripinnis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB8A87B465BC276FF34F8D8FC10F831" blockId="2.[151,1437,1716,1958]" pageId="2" pageNumber="327">
<emphasis id="B9737469465BC276FF34F8D8FE1BF8C9" bold="true" box="[199,458,1860,1886]" pageId="2" pageNumber="327">Specimens examined.</emphasis>
Specimens used for morphometrics in supplementary table 1 and those for the map (
<figureCitation id="133CB4FE465BC276FF6CF8F4FF37F815" box="[159,230,1896,1922]" captionStart="FIGURE 3" captionStartId="5.[151,250,1009,1033]" captionTargetBox="[168,1423,181,981]" captionTargetId="figure-15@5.[164,1423,181,985]" captionTargetPageId="5" captionText="FIGURE 3. Distribution of Vexillichthys n. gen. atripinnis." figureDoi="http://doi.org/10.5281/zenodo.14033797" httpUri="https://zenodo.org/record/14033797/files/figure.png" pageId="2" pageNumber="327">Fig. 3</figureCitation>
) in supplementary table 2 available at
<bibRefCitation id="EF96D58A465BC276FD47F8F4FC5AF815" author="Armbruster, J. W." box="[692,907,1896,1922]" pageId="2" pageNumber="327" refId="ref5469" refString="Armbruster, J. W. (2024) Supplementary tables: Blackfin Sucker mensural and locality data. Available from: https: // aurora. auburn. edu / handle / 11200 / 50650 (Accessed 7 Oct. 2024) https: // doi. org / 10.35099 / nfcp- 8538" type="book" year="2024">Armbruster (2024)</bibRefCitation>
(AUM specimens were additionally used for morphological comparisons). Skeletal specimen examined AUM 86412.
</paragraph>
</subSubSection>
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<emphasis id="B9737469465AC277FF64FCD2FEC5FCF1" bold="true" box="[151,276,846,870]" pageId="3" pageNumber="328">FIGURE 1.</emphasis>
Live specimens of
<taxonomicName id="4C07D3F8465AC277FE2DFCD2FD8FFCF1" authorityName="Armbruster" authorityYear="2024" box="[478,606,846,870]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B9737469465AC277FE2DFCD2FD8FFCF1" box="[478,606,846,870]" italics="true" pageId="3" pageNumber="328">Vexillichthys</emphasis>
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<emphasis id="B9737469465AC277FD96FCD2FD7FFCF1" bold="true" box="[613,686,846,870]" pageId="3" pageNumber="328">
<taxonomicNameLabel id="A240C912465AC277FD96FCD2FD7FFCF1" box="[613,686,846,870]" pageId="3" pageNumber="328" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C07D3F8465AC277FD46FCD3FCC9FCF1" box="[693,792,847,870]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="species" species="atripinnis">
<emphasis id="B9737469465AC277FD46FCD3FCC9FCF1" box="[693,792,847,870]" italics="true" pageId="3" pageNumber="328">atripinnis</emphasis>
</taxonomicName>
. A. young in typical coloration (90 mm SL, Salt Lick Creek at Bugtussle Road crossing, Monroe Co., KY, 36.65654°, -85.92124°) and B. nuptial specimen, probably female (147 mm SL, Long Hungry Creek, at KY 1333 crossing, Allen Co., KY, 36.67412°, -86.04397°). Specimens uncataloged. Photos by M. Thomas.
</paragraph>
</caption>
<subSubSection id="C31DFBF0465AC277FF34FBB8FED7F9E8" pageId="3" pageNumber="328" type="diagnosis">
<paragraph id="8BB8A87B465AC277FF34FBB8FB88FA5C" blockId="3.[151,1437,1060,2022]" pageId="3" pageNumber="328">
<emphasis id="B9737469465AC277FF34FBB8FEEBFBA9" bold="true" box="[199,314,1060,1086]" pageId="3" pageNumber="328">Diagnosis</emphasis>
.
<taxonomicName id="4C07D3F8465AC277FEBDFBB8FE0DFBA9" authorityName="Armbruster" authorityYear="2024" box="[334,476,1060,1086]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FEBDFBB8FE0DFBA9" box="[334,476,1060,1086]" italics="true" pageId="3" pageNumber="328">Vexillichthys</emphasis>
</taxonomicName>
can be separated from all other Catostomids by the presence of well-developed, thin, black stripes on the body and a large, black mark on the anterodistal portion of the dorsal fin. Members of
<taxonomicName id="4C07D3F8465AC277FF64FBF0FEF7FB11" authorityName="Rafinesque" authorityYear="1818" box="[151,294,1132,1158]" family="Catostomidae" genus="Hypentelium" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FF64FBF0FEF7FB11" box="[151,294,1132,1158]" italics="true" pageId="3" pageNumber="328">Hypentelium</emphasis>
</taxonomicName>
and
<taxonomicName id="4C07D3F8465AC277FE93FBF0FE07FB11" box="[352,470,1132,1158]" family="Catostomidae" genus="Thoburnia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FE93FBF0FE07FB11" box="[352,470,1132,1158]" italics="true" pageId="3" pageNumber="328">Thoburnia</emphasis>
</taxonomicName>
may have stripes, but the stripes are faint, and the dorsal saddles are darker than the stripes (vs. the stripes darker than the saddles in
<taxonomicName id="4C07D3F8465AC277FD40FB0CFC90FB3D" authorityName="Armbruster" authorityYear="2024" box="[691,833,1168,1194]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FD40FB0CFC90FB3D" box="[691,833,1168,1194]" italics="true" pageId="3" pageNumber="328">Vexillichthys</emphasis>
</taxonomicName>
).
<taxonomicName id="4C07D3F8465AC277FCA4FB0CFC37FB3D" authorityName="Rafinesque" authorityYear="1818" box="[855,998,1168,1194]" family="Catostomidae" genus="Hypentelium" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FCA4FB0CFC37FB3D" box="[855,998,1168,1194]" italics="true" pageId="3" pageNumber="328">Hypentelium</emphasis>
</taxonomicName>
does have dark marks in the dorsal fin, but the fin has bands or blotches instead of a single, large, black area at the anterodistal portion of the fin and the remainder of the fin hyaline.
<taxonomicName id="4C07D3F8465AC277FE2DFB44FDBDFB65" authorityName="Armbruster" authorityYear="2024" box="[478,620,1240,1266]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FE2DFB44FDBDFB65" box="[478,620,1240,1266]" italics="true" pageId="3" pageNumber="328">Vexillichthys</emphasis>
</taxonomicName>
can be further separated from
<taxonomicName id="4C07D3F8465AC277FC30FB44FBE8FB65" box="[963,1081,1240,1266]" family="Catostomidae" genus="Thoburnia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FC30FB44FBE8FB65" box="[963,1081,1240,1266]" italics="true" pageId="3" pageNumber="328">Thoburnia</emphasis>
</taxonomicName>
by lacking a rust-colored, wide lateral stripe.
<taxonomicName id="4C07D3F8465AC277FEDCFB61FE78FA81" authorityName="Jordan" authorityYear="1878" box="[303,425,1277,1302]" family="Catostomidae" genus="Minytrema" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FEDCFB61FE78FA81" box="[303,425,1277,1302]" italics="true" pageId="3" pageNumber="328">Minytrema</emphasis>
</taxonomicName>
also has dark stripes, but these are composed of distinct spots (vs. continuous lines). Some of the species of
<taxonomicName id="4C07D3F8465AC277FEAAFABDFE0DFAAD" authorityName="Rafinesque" authorityYear="1820" box="[345,476,1313,1338]" family="Catostomidae" genus="Moxostoma" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FEAAFABDFE0DFAAD" box="[345,476,1313,1338]" italics="true" pageId="3" pageNumber="328">Moxostoma</emphasis>
</taxonomicName>
formerly ascribed to
<taxonomicName id="4C07D3F8465AC277FD22FABDFCB3FAAC" authorityName="Fowler" authorityYear="1913" box="[721,866,1313,1339]" family="Catostomidae" genus="Scartomyzon" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FD22FABDFCB3FAAC" box="[721,866,1313,1339]" italics="true" pageId="3" pageNumber="328">Scartomyzon</emphasis>
</taxonomicName>
, also may have stripes, but they are not as intense as that of
<taxonomicName id="4C07D3F8465AC277FEF5FAD8FE45FAC9" authorityName="Armbruster" authorityYear="2024" box="[262,404,1348,1374]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FEF5FAD8FE45FAC9" box="[262,404,1348,1374]" italics="true" pageId="3" pageNumber="328">Vexillichthys</emphasis>
</taxonomicName>
and there is maximally some black just at the anterodistal edge of the dorsal fin (vs. a large area).
<taxonomicName id="4C07D3F8465AC277FF2FFAF4FEBBFA15" authorityName="Armbruster" authorityYear="2024" box="[220,362,1384,1410]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FF2FFAF4FEBBFA15" box="[220,362,1384,1410]" italics="true" pageId="3" pageNumber="328">Vexillichthys</emphasis>
</taxonomicName>
may also be unique in the development of pads on the distal ends of the anterior unbranched rays of the pectoral fin as well as the anterior portion of the first branched ray and occasionally the second branched ray (other suckers may have thickened skin in these regions, but not distinct pads;
<figureCitation id="133CB4FE465AC277FBFBFA2CFB9AFA5C" box="[1032,1099,1456,1483]" captionStart="FIGURE 5" captionStartId="6.[151,250,1214,1238]" captionTargetBox="[177,1369,237,1191]" captionTargetId="figure-46@6.[177,1411,237,1191]" captionTargetPageId="6" captionText="FIGURE 5. Ventral view of right pectoral fins of A. Thoburnia rhothoeca, AUM 61433, 94.7 mm SL and B. Vexillichthys n. gen. atripinnis, AUM 12812, 133.7 mm SL. Unbranched pectoral fin rays numbered with lowercase Roman numerals and first two branched rays numbered with Arabic numerals. The specimen in B is a large, nuptial female and pads are present on all four of the numbered rays with even some thickening on the third branched ray (numbers are approximately in the center of the pads). Photos by J.W. Armbruster. Scale = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14033801" httpUri="https://zenodo.org/record/14033801/files/figure.png" pageId="3" pageNumber="328">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BB8A87B465AC277FF34FA48FED7F9E8" blockId="3.[151,1437,1060,2022]" pageId="3" pageNumber="328">
Osteologically,
<taxonomicName id="4C07D3F8465AC277FE8EFA48FDDAFA79" authorityName="Armbruster" authorityYear="2024" box="[381,523,1492,1518]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FE8EFA48FDDAFA79" box="[381,523,1492,1518]" italics="true" pageId="3" pageNumber="328">Vexillichthys</emphasis>
</taxonomicName>
can be diagnosed by two characteristics of the skull: lateral laminae of the dermethmoid angled anteriorly (
<figureCitation id="133CB4FE465AC277FDE7FA64FDA2F984" box="[532,627,1528,1555]" captionStart="FIGURE 4" captionStartId="5.[151,250,1672,1696]" captionTargetBox="[164,1423,1087,1648]" captionTargetId="figure-27@5.[164,1423,1087,1648]" captionTargetPageId="5" captionText="FIGURE 4. Dorsal view of crania of A. Hypentelium nigricans, AUM 86410, 259.5 mm SL, B. Thoburnia rhothoecum, AUM 86411, and C. Vexillichthys n. gen. atripinnis, AUM 86412. 109.7 mm SL. DE—dermethmoid (lines are to a lateral lamina), F—frontal, FPF—frontoparietal fenestra, LE—lateral ethmoid (lines are to the base of the bone), and SOC—supraorbital crest (lines are to a lateral lamina). Photos by J.W. Armbruster. Scale = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14033799" httpUri="https://zenodo.org/record/14033799/files/figure.png" pageId="3" pageNumber="328">Fig. 4C</figureCitation>
; vs. laterally in
<taxonomicName id="4C07D3F8465AC277FCCCFA64FBE2F985" box="[831,1075,1528,1554]" family="Catostomidae" genus="Thoburnia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="species" species="rhothoeca">
<emphasis id="B9737469465AC277FCCCFA64FBE2F985" box="[831,1075,1528,1554]" italics="true" pageId="3" pageNumber="328">Thoburnia rhothoeca</emphasis>
</taxonomicName>
,
<figureCitation id="133CB4FE465AC277FBBBFA64FB75F984" box="[1096,1188,1528,1555]" captionStart="FIGURE 4" captionStartId="5.[151,250,1672,1696]" captionTargetBox="[164,1423,1087,1648]" captionTargetId="figure-27@5.[164,1423,1087,1648]" captionTargetPageId="5" captionText="FIGURE 4. Dorsal view of crania of A. Hypentelium nigricans, AUM 86410, 259.5 mm SL, B. Thoburnia rhothoecum, AUM 86411, and C. Vexillichthys n. gen. atripinnis, AUM 86412. 109.7 mm SL. DE—dermethmoid (lines are to a lateral lamina), F—frontal, FPF—frontoparietal fenestra, LE—lateral ethmoid (lines are to the base of the bone), and SOC—supraorbital crest (lines are to a lateral lamina). Photos by J.W. Armbruster. Scale = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14033799" httpUri="https://zenodo.org/record/14033799/files/figure.png" pageId="3" pageNumber="328">Fig. 4B</figureCitation>
, or posteriorly as in
<taxonomicName id="4C07D3F8465AC277FF64F980FEF7F9A1" authorityName="Rafinesque" authorityYear="1818" box="[151,294,1564,1590]" family="Catostomidae" genus="Hypentelium" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FF64F980FEF7F9A1" box="[151,294,1564,1590]" italics="true" pageId="3" pageNumber="328">Hypentelium</emphasis>
</taxonomicName>
and most
<taxonomicName id="4C07D3F8465AC277FE6BF981FDCAF9A1" authorityName="Rafinesque" authorityYear="1820" box="[408,539,1565,1590]" family="Catostomidae" genus="Moxostoma" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FE6BF981FDCAF9A1" box="[408,539,1565,1590]" italics="true" pageId="3" pageNumber="328">Moxostoma</emphasis>
</taxonomicName>
,
<figureCitation id="133CB4FE465AC277FDD4F980FD51F9A0" box="[551,640,1564,1591]" captionStart="FIGURE 4" captionStartId="5.[151,250,1672,1696]" captionTargetBox="[164,1423,1087,1648]" captionTargetId="figure-27@5.[164,1423,1087,1648]" captionTargetPageId="5" captionText="FIGURE 4. Dorsal view of crania of A. Hypentelium nigricans, AUM 86410, 259.5 mm SL, B. Thoburnia rhothoecum, AUM 86411, and C. Vexillichthys n. gen. atripinnis, AUM 86412. 109.7 mm SL. DE—dermethmoid (lines are to a lateral lamina), F—frontal, FPF—frontoparietal fenestra, LE—lateral ethmoid (lines are to the base of the bone), and SOC—supraorbital crest (lines are to a lateral lamina). Photos by J.W. Armbruster. Scale = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14033799" httpUri="https://zenodo.org/record/14033799/files/figure.png" pageId="3" pageNumber="328">Fig. 4A</figureCitation>
); narrowest portion of lateral ethmoid when viewed ventrally much less than length of anterolateral processes (vs. approximately equal in
<taxonomicName id="4C07D3F8465AC277FC9DF9DDFC2BF9CD" box="[878,1018,1600,1626]" family="Catostomidae" genus="Thoburnia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="species" species="rhothoeca">
<emphasis id="B9737469465AC277FC9DF9DDFC2BF9CD" box="[878,1018,1600,1626]" italics="true" pageId="3" pageNumber="328">T. rhothoeca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C07D3F8465AC277FBDDF9DCFB6CF9CD" authorityName="Rafinesque" authorityYear="1818" box="[1070,1213,1600,1626]" family="Catostomidae" genus="Hypentelium" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="328" phylum="Chordata" rank="genus">
<emphasis id="B9737469465AC277FBDDF9DCFB6CF9CD" box="[1070,1213,1600,1626]" italics="true" pageId="3" pageNumber="328">Hypentelium</emphasis>
</taxonomicName>
, area of base shown in
<figureCitation id="133CB4FE465AC277FF47F9F8FF29F9E8" box="[180,248,1636,1663]" captionStart="FIGURE 4" captionStartId="5.[151,250,1672,1696]" captionTargetBox="[164,1423,1087,1648]" captionTargetId="figure-27@5.[164,1423,1087,1648]" captionTargetPageId="5" captionText="FIGURE 4. Dorsal view of crania of A. Hypentelium nigricans, AUM 86410, 259.5 mm SL, B. Thoburnia rhothoecum, AUM 86411, and C. Vexillichthys n. gen. atripinnis, AUM 86412. 109.7 mm SL. DE—dermethmoid (lines are to a lateral lamina), F—frontal, FPF—frontoparietal fenestra, LE—lateral ethmoid (lines are to the base of the bone), and SOC—supraorbital crest (lines are to a lateral lamina). Photos by J.W. Armbruster. Scale = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14033799" httpUri="https://zenodo.org/record/14033799/files/figure.png" pageId="3" pageNumber="328">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31DFBF0465AC272FF34F914FE20FF42" lastPageId="6" lastPageNumber="331" pageId="3" pageNumber="328" type="description">
<paragraph id="8BB8A87B465AC277FF34F914FF0BF8A5" blockId="3.[151,1437,1060,2022]" pageId="3" pageNumber="328">
<emphasis id="B9737469465AC277FF34F914FE88F935" bold="true" box="[199,345,1672,1698]" pageId="3" pageNumber="328">Description.</emphasis>
Morphometrics in table 1. Body long and low, flattened ventrally to anal fin and then rising to caudal peduncle. Dorsal surface forms gentle arc from snout to dorsal fin and then slowly lowering to caudal peduncle. Head small and roughly conical. Eye large. Mouth inferior. Upper lip plicate, about half width of lower lip; lower lip plicate proximally and with cross furrows distally; lower lip with distal split between left and right lobes.
</paragraph>
<paragraph id="8BB8A87B465AC277FF34F8A0FD4CF871" blockId="3.[151,1437,1060,2022]" pageId="3" pageNumber="328">Dorsal fin roughly triangular in juveniles and females, becoming almost rectangular in nuptial males; located approximately mid body; ii,810 (usually 9). Anal fin rounded, broader in nuptial males; ii,67 (usually 6). Caudal fin emarginate with rounded lobes, i,8,8,i. Pectoral fin short and rounded with leading unbranched rays and anterior margin of first branched ray with fleshy pad; located at ventral margin of body; ii,12,i. Pelvic fins rounded, originating below middle of dorsal fin; i,68 (usually 7),i.</paragraph>
<caption id="DF78F8F3465DC270FF64FB61FCC4FAAE" ID-DOI="http://doi.org/10.5281/zenodo.14033795" ID-Zenodo-Dep="14033795" httpUri="https://zenodo.org/record/14033795/files/figure.png" pageId="4" pageNumber="329" startId="4.[151,250,1277,1301]" targetBox="[177,1411,181,1253]" targetPageId="4" targetType="figure">
<paragraph id="8BB8A87B465DC270FF64FB61FCC4FAAE" blockId="4.[151,1436,1277,1337]" pageId="4" pageNumber="329">
<emphasis id="B9737469465DC270FF64FB61FEC7FA82" bold="true" box="[151,278,1277,1301]" pageId="4" pageNumber="329">FIGURE 2.</emphasis>
Preserved nuptial male specimen of
<taxonomicName id="4C07D3F8465DC270FD6FFB61FCCDFA82" authorityName="Armbruster" authorityYear="2024" box="[668,796,1277,1301]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="329" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B9737469465DC270FD6FFB61FCCDFA82" box="[668,796,1277,1301]" italics="true" pageId="4" pageNumber="329">Vexillichthys</emphasis>
</taxonomicName>
<emphasis id="B9737469465DC270FCD6FB61FCBEFA82" bold="true" box="[805,879,1277,1301]" pageId="4" pageNumber="329">
<taxonomicNameLabel id="A240C912465DC270FCD6FB61FCBEFA82" box="[805,879,1277,1301]" pageId="4" pageNumber="329" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C07D3F8465DC270FC84FB62FC0BFA82" box="[887,986,1278,1301]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="329" phylum="Chordata" rank="species" species="atripinnis">
<emphasis id="B9737469465DC270FC84FB62FC0BFA82" box="[887,986,1278,1301]" italics="true" pageId="4" pageNumber="329">atripinnis</emphasis>
</taxonomicName>
in dorsal, lateral, and ventral views. AUM 12812, 71.6 mm SL. Photos by J.W. Armbruster. Scale = 1 cm.
</paragraph>
</caption>
<paragraph id="8BB8A87B465DC270FF34FA17FBB0F9A2" blockId="4.[151,1437,1419,2021]" pageId="4" pageNumber="329">
Lateral line complete. Scales small,
<quantity id="4CFF059E465DC270FDACFA17FD13FA32" box="[607,706,1419,1446]" metricMagnitude="0" metricUnit="m" metricValue="1.2192" metricValueMax="1.27" metricValueMin="1.1684" pageId="4" pageNumber="329" unit="in" value="48.0" valueMax="50.0" valueMin="46.0">4650 in</quantity>
lateral line, 1620 predorsal scale rows, 9 caudal peduncle scale rows, 16 circumpeduncular scale rows, 3132 circumferential scales. Scales of breast and nape deeply embedded. Vertebrae 4345. 35 right and 39 left pharyngeal teeth; ventral most teeth with
<emphasis id="B9737469465DC270FBCEFA48FB80FA7A" box="[1085,1105,1492,1517]" italics="true" pageId="4" pageNumber="329">U</emphasis>
-shaped cusps (shallowest in middle, longest edge medial) and dorsal most teeth thin and filelike; pharyngeal arch narrow, not much wider than teeth; single tooth present on neck of arch (portion of arch ventral to widened section).
</paragraph>
<paragraph id="8BB8A87B465DC270FF34F9A3FEFFF872" blockId="4.[151,1437,1419,2021]" pageId="4" pageNumber="329">
<emphasis id="B9737469465DC270FF34F9A3FE9EF9CE" bold="true" box="[199,335,1599,1625]" pageId="4" pageNumber="329">Coloration.</emphasis>
Body distinctly dark above (base color tan with overlying dark markings) and white below. Head with thick, diffuse stripe from tip of snout, through eye, to dorsal margin of opercle. Body with approximately seven black stripes running entire length separated by tan to cream interspaces. Faint dorsal saddles present below stripes, first behind head, second below origin of dorsal fin (slanted anteroventrally), third behind dorsal fin (slanted posteroventrally), fourth at middle of caudal peduncle (wedge-shaped, narrowest dorsally), and fifth at base of caudal fin; second and fourth saddles darkest. Dorsal fin clear to hyaline with large, black spot anterodistally on first five to six rays with spot continuing only on extreme distal portion of remaining rays. Other fins hyaline with caudal darker; caudal also with broad, dark stripe down middle and sometimes with dark edge. Nuptial specimens darker often with a dark mark over the pectoral girdle that extends medially behind opercular opening and interradial membranes of pectoral and/or anal fins pigmented (mainly in center third of length of fin). Color in life similar, but pectoral fins and dorsal and ventral margins of caudal fin rust-colored with lighter yellow to rust coloring on pelvic and anal fins.
</paragraph>
<caption id="DF78F8F3465CC271FF64FC6DFD23FB9E" ID-DOI="http://doi.org/10.5281/zenodo.14033797" ID-Zenodo-Dep="14033797" box="[151,754,1009,1034]" httpUri="https://zenodo.org/record/14033797/files/figure.png" pageId="5" pageNumber="330" startId="5.[151,250,1009,1033]" targetBox="[168,1423,181,981]" targetPageId="5" targetType="figure">
<paragraph id="8BB8A87B465CC271FF64FC6DFD23FB9E" blockId="5.[151,754,1009,1034]" box="[151,754,1009,1034]" pageId="5" pageNumber="330">
<emphasis id="B9737469465CC271FF64FC6DFEC2FB9D" bold="true" box="[151,275,1009,1034]" pageId="5" pageNumber="330">FIGURE 3.</emphasis>
Distribution of
<taxonomicName id="4C07D3F8465CC271FE45FC6EFDE7FB9D" authorityName="Armbruster" authorityYear="2024" box="[438,566,1010,1034]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="330" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B9737469465CC271FE45FC6EFDE7FB9D" box="[438,566,1010,1034]" italics="true" pageId="5" pageNumber="330">Vexillichthys</emphasis>
</taxonomicName>
<emphasis id="B9737469465CC271FDCFFC6EFD52FB9D" bold="true" box="[572,643,1010,1034]" pageId="5" pageNumber="330">
<taxonomicNameLabel id="A240C912465CC271FDCFFC6EFD52FB9D" box="[572,643,1010,1034]" pageId="5" pageNumber="330" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C07D3F8465CC271FD7AFC6EFD3DFB9E" box="[649,748,1010,1033]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="330" phylum="Chordata" rank="species" species="atripinnis">
<emphasis id="B9737469465CC271FD7AFC6EFD3DFB9E" box="[649,748,1010,1033]" italics="true" pageId="5" pageNumber="330">atripinnis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF78F8F3465CC271FF64F914FCBCF89B" ID-DOI="http://doi.org/10.5281/zenodo.14033799" ID-Zenodo-Dep="14033799" httpUri="https://zenodo.org/record/14033799/files/figure.png" pageId="5" pageNumber="330" startId="5.[151,250,1672,1696]" targetBox="[164,1423,1087,1648]" targetPageId="5" targetType="figure">
<paragraph id="8BB8A87B465CC271FF64F914FCBCF89B" blockId="5.[151,1436,1672,1804]" pageId="5" pageNumber="330">
<emphasis id="B9737469465CC271FF64F914FEC2F937" bold="true" box="[151,275,1672,1696]" pageId="5" pageNumber="330">FIGURE 4.</emphasis>
Dorsal view of crania of A.
<taxonomicName id="4C07D3F8465CC271FDC7F914FCB5F937" authority=", AUM" authorityName="AUM" box="[564,868,1672,1696]" family="Catostomidae" genus="Hypentelium" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="330" phylum="Chordata" rank="species" species="nigricans">
<emphasis id="B9737469465CC271FDC7F914FCCDF937" box="[564,796,1672,1696]" italics="true" pageId="5" pageNumber="330">Hypentelium nigricans</emphasis>
, AUM
</taxonomicName>
86410, 259.5 mm SL, B.
<taxonomicName id="4C07D3F8465CC271FB9FF914FA4DF937" authority=", AUM" authorityName="AUM" box="[1132,1436,1672,1696]" family="Catostomidae" genus="Thoburnia" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="330" phylum="Chordata" rank="species" species="rhothoecum">
<emphasis id="B9737469465CC271FB9FF914FA87F937" box="[1132,1366,1672,1696]" italics="true" pageId="5" pageNumber="330">Thoburnia rhothoecum</emphasis>
, AUM
</taxonomicName>
86411, and C.
<taxonomicName id="4C07D3F8465CC271FEDCF930FE7EF953" authorityName="Armbruster" authorityYear="2024" box="[303,431,1708,1732]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="330" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B9737469465CC271FEDCF930FE7EF953" box="[303,431,1708,1732]" italics="true" pageId="5" pageNumber="330">Vexillichthys</emphasis>
</taxonomicName>
<emphasis id="B9737469465CC271FE44F930FDD1F953" bold="true" box="[439,512,1708,1732]" pageId="5" pageNumber="330">
<taxonomicNameLabel id="A240C912465CC271FE44F930FDD1F953" box="[439,512,1708,1732]" pageId="5" pageNumber="330" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C07D3F8465CC271FDF4F931FD60F953" authority=", AUM" authorityName="AUM" box="[519,689,1708,1732]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="330" phylum="Chordata" rank="species" species="atripinnis">
<emphasis id="B9737469465CC271FDF4F931FDBBF953" box="[519,618,1709,1732]" italics="true" pageId="5" pageNumber="330">atripinnis</emphasis>
, AUM
</taxonomicName>
86412. 109.7 mm SL. DE—dermethmoid (lines are to a lateral lamina), F—frontal, FPF—frontoparietal fenestra, LE—lateral ethmoid (lines are to the base of the bone), and SOC—supraorbital crest (lines are to a lateral lamina). Photos by J.W. Armbruster. Scale = 1 cm.
</paragraph>
</caption>
<paragraph id="8BB8A87B465CC272FF34F8CDFE20FF42" blockId="5.[151,1437,1873,2043]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="331" pageId="5" pageNumber="330">
<emphasis id="B9737469465CC271FF34F8CDFE6CF8FC" bold="true" box="[199,445,1873,1899]" pageId="5" pageNumber="330">Sexual Dimorphism.</emphasis>
Males much smaller than females with larger anal fin (anal fin height to SL ratio 21.4 33.7% vs. 7.917.2%) and a more rectangular dorsal fin with a greater depressed dorsal fin length to SL ratio 22.333.4% (vs. 20.022.3%). Tubercles found in both males and females. Tubercles in females small and present on anal fin and caudal peduncle ventral to lateral line. Nuptial males with large tubercles on the anal and caudal fins and small tubercles on much of body (particularly caudal peduncle head and throat). Males also with some small tubercles occasionally on ventral side of pectoral fin and dorsal side of pelvic fin. Body tubercles of males best developed on caudal peduncle.
</paragraph>
</subSubSection>
<caption id="DF78F8F3465FC272FF64FB22FDEFFAF1" ID-DOI="http://doi.org/10.5281/zenodo.14033801" ID-Zenodo-Dep="14033801" httpUri="https://zenodo.org/record/14033801/files/figure.png" pageId="6" pageNumber="331" startId="6.[151,250,1214,1238]" targetBox="[177,1369,237,1191]" targetPageId="6" targetType="figure">
<paragraph id="8BB8A87B465FC272FF64FB22FDEFFAF1" blockId="6.[151,1437,1214,1382]" pageId="6" pageNumber="331">
<emphasis id="B9737469465FC272FF64FB22FEC5FB41" bold="true" box="[151,276,1214,1238]" pageId="6" pageNumber="331">FIGURE 5.</emphasis>
Ventral view of right pectoral fins of A.
<taxonomicName id="4C07D3F8465FC272FD49FB22FC0BFB40" authority=", AUM" authorityName="AUM" box="[698,986,1214,1239]" family="Catostomidae" genus="Thoburnia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="331" phylum="Chordata" rank="species" species="rhothoeca">
<emphasis id="B9737469465FC272FD49FB22FC42FB41" box="[698,915,1214,1238]" italics="true" pageId="6" pageNumber="331">Thoburnia rhothoeca</emphasis>
, AUM
</taxonomicName>
61433, 94.7 mm SL and B.
<taxonomicName id="4C07D3F8465FC272FAF3FB22FA51FB41" authorityName="Armbruster" authorityYear="2024" box="[1280,1408,1214,1238]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="331" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B9737469465FC272FAF3FB22FA51FB41" box="[1280,1408,1214,1238]" italics="true" pageId="6" pageNumber="331">Vexillichthys</emphasis>
</taxonomicName>
<emphasis id="B9737469465FC272FA7BFB22FF15FB6D" bold="true" pageId="6" pageNumber="331">
<taxonomicNameLabel id="A240C912465FC272FA7BFB22FF15FB6D" pageId="6" pageNumber="331" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C07D3F8465FC272FF39FB7FFEA2FB6C" authority=", AUM" authorityName="AUM" box="[202,371,1251,1275]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="331" phylum="Chordata" rank="species" species="atripinnis">
<emphasis id="B9737469465FC272FF39FB7FFEFCFB6D" box="[202,301,1251,1274]" italics="true" pageId="6" pageNumber="331">atripinnis</emphasis>
, AUM
</taxonomicName>
12812, 133.7 mm SL. Unbranched pectoral fin rays numbered with lowercase Roman numerals and first two branched rays numbered with Arabic numerals. The specimen in B is a large, nuptial female and pads are present on all four of the numbered rays with even some thickening on the third branched ray (numbers are approximately in the center of the pads). Photos by J.W. Armbruster. Scale = 1 cm.
</paragraph>
</caption>
<subSubSection id="C31DFBF0465FC272FF34FA24FA9AFA44" box="[199,1355,1464,1491]" pageId="6" pageNumber="331" type="distribution">
<paragraph id="8BB8A87B465FC272FF34FA24FA9AFA44" blockId="6.[151,1437,1464,2030]" box="[199,1355,1464,1491]" pageId="6" pageNumber="331">
<emphasis id="B9737469465FC272FF34FA24FEB0FA45" bold="true" box="[199,353,1464,1490]" pageId="6" pageNumber="331">Distribution.</emphasis>
Found only in headwaters of the Barren River System of
<collectingRegion id="49C36699465FC272FC1BFA24FB84FA45" box="[1000,1109,1464,1490]" country="United States of America" name="Kentucky" pageId="6" pageNumber="331">Kentucky</collectingRegion>
and
<collectingRegion id="49C36699465FC272FB78FA25FAD0FA44" box="[1163,1281,1465,1491]" country="United States of America" name="Tennessee" pageId="6" pageNumber="331">Tennessee</collectingRegion>
,
<collectingCountry id="F310E8EB465FC272FAFFFA25FA99FA44" box="[1292,1352,1465,1491]" name="United States of America" pageId="6" pageNumber="331">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31DFBF0465FC272FF34FA41FC87F814" pageId="6" pageNumber="331" type="biology_ecology">
<paragraph id="8BB8A87B465FC272FF34FA41FC87F814" blockId="6.[151,1437,1464,2030]" pageId="6" pageNumber="331">
<emphasis id="B9737469465FC272FF34FA41FEFDFA60" bold="true" box="[199,300,1501,1527]" pageId="6" pageNumber="331">Ecology.</emphasis>
The trematode,
<taxonomicName id="4C07D3F8465FC272FE19FA40FD28FA61" authorityName="Timmons &amp; Rogers" authorityYear="1977" box="[490,761,1500,1526]" class="Monogenea" family="Dactylogyridae" genus="Dactylogyrus" kingdom="Animalia" order="Dactylogyridea" pageId="6" pageNumber="331" phylum="Platyhelminthes" rank="species" species="atripinnei">
<emphasis id="B9737469465FC272FE19FA40FD28FA61" box="[490,761,1500,1526]" italics="true" pageId="6" pageNumber="331">Dactylogyrus atripinnei</emphasis>
</taxonomicName>
was described from the gills of
<taxonomicName id="4C07D3F8465FC272FB81FA40FEE2F98C" authority="(Timmons &amp; Rogers 1977)" baseAuthorityName="Timmons &amp; Rogers" baseAuthorityYear="1977" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="331" phylum="Chordata" rank="genus">
<emphasis id="B9737469465FC272FB81FA40FAD1FA61" box="[1138,1280,1500,1526]" italics="true" pageId="6" pageNumber="331">Vexillichthys</emphasis>
(
<bibRefCitation id="EF96D58A465FC272FAE2FA40FEFAF98C" author="Timmons, T. J. &amp; Rogers, W. A." pageId="6" pageNumber="331" pagination="238 - 239" refId="ref7644" refString="Timmons, T. J. &amp; Rogers, W. A. (1977) Dactylogyrus atripinnei sp. n. from the Blackfin Sucker in Tennessee. The Journal of Parasitology, 63, 238 - 239. https: // doi. org / 10.2307 / 3280049" type="journal article" year="1977">Timmons &amp; Rogers 1977</bibRefCitation>
)
</taxonomicName>
. Life history was described in
<bibRefCitation id="EF96D58A465FC272FD62F99CFC41F98C" author="Timmons, T. J. &amp; Ramsey, J. S. &amp; Bauer, B. H." box="[657,912,1536,1563]" pageId="6" pageNumber="331" pagination="538 - 541" refId="ref7694" refString="Timmons, T. J., Ramsey, J. S. &amp; Bauer, B. H. (1983) Life history and habitat of the blackfin sucker, Moxostoma atripinne (Osteichthyes: Catostomidae). Copeia, 1983, 538 - 541. https: // doi. org / 10.2307 / 1444405" type="journal article" year="1983">
Timmons
<emphasis id="B9737469465FC272FCF7F99DFCECF98D" box="[772,829,1536,1562]" italics="true" pageId="6" pageNumber="331">et al.</emphasis>
(1983)
</bibRefCitation>
with
<taxonomicName id="4C07D3F8465FC272FC21F99DFB8DF98D" box="[978,1116,1537,1562]" family="Catostomidae" genus="Vexillichthys" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="331" phylum="Chordata" rank="species" species="atripinnis">
<emphasis id="B9737469465FC272FC21F99DFB8DF98D" box="[978,1116,1537,1562]" italics="true" pageId="6" pageNumber="331">V. atripinnis</emphasis>
</taxonomicName>
found in headwater streams (
<quantity id="4CFF059E465FC272FF6CF9B8FED1F9A8" box="[159,256,1572,1599]" metricMagnitude="0" metricUnit="m" metricValue="5.25" metricValueMax="9.0" metricValueMin="1.5" pageId="6" pageNumber="331" unit="m" value="5.25" valueMax="9.0" valueMin="1.5">1.59 m</quantity>
wide) with alternating pools and riffles. Streams were clear with a substrate of limestone, shale, and siltstone. Young were found in pools (
<quantity id="4CFF059E465FC272FDB4F9D5FD4CF9F4" box="[583,669,1609,1635]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="6" pageNumber="331" unit="m" value="0.65" valueMax="1.0" valueMin="0.3">0.31m</quantity>
deep) often in schools over gravel. Adults were collected along the shore or in pools (1.0
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deep) and would retreat into crevices in the bedrock.Adults are associated with bedrock ledges, boulders, slab rock, and rootwads (see also
<bibRefCitation id="EF96D58A465FC272FD1BF90CFC75F93C" author="Stringfield, C." box="[744,932,1680,1707]" pageId="6" pageNumber="331" refId="ref7533" refString="Stringfield, C. (2013) Population distribution and abundance of the blackfin sucker (Thoburnia atripinnis) in the upper Barren River System. Unpublished Master Thesis, Eastern Kentucky University, Richmond, Kentucky. [unknown pagination]" type="book" year="2013">Stringfield 2013</bibRefCitation>
;
<bibRefCitation id="EF96D58A465FC272FC41F90CFF05F958" author="Kentucky Ecological Services Field Office" pageId="6" pageNumber="331" refId="ref6771" refString="Kentucky Ecological Services Field Office (2017) Blackfin Sucker (Thoburnia atripinnis) species status assessment. Report to the United States Fish and Wildlife Service. Kentucky Ecological Services Field Office, Frankfort, Kentucky, 38 pp." type="book" year="2017">
<collectingRegion id="49C36699465FC272FC41F90CFBCEF93D" box="[946,1055,1680,1706]" country="United States of America" name="Kentucky" pageId="6" pageNumber="331">Kentucky</collectingRegion>
Ecological Services Field Office 2017
</bibRefCitation>
). They feed mostly on chironomids but also on cladocerans, copepods, ostracods, simuliids, melgalopterans, and trichopterans. Males are tuberculate in year two, but
<bibRefCitation id="EF96D58A465FC272FCEBF944FBC2F964" author="Timmons, T. J. &amp; Ramsey, J. S. &amp; Bauer, B. H." box="[792,1043,1752,1779]" pageId="6" pageNumber="331" pagination="538 - 541" refId="ref7694" refString="Timmons, T. J., Ramsey, J. S. &amp; Bauer, B. H. (1983) Life history and habitat of the blackfin sucker, Moxostoma atripinne (Osteichthyes: Catostomidae). Copeia, 1983, 538 - 541. https: // doi. org / 10.2307 / 1444405" type="journal article" year="1983">
Timmons
<emphasis id="B9737469465FC272FC7AF945FC13F965" box="[905,962,1752,1778]" italics="true" pageId="6" pageNumber="331">et al.</emphasis>
(1983)
</bibRefCitation>
reported a young-of-the-year male that was emitting milt in December.
<bibRefCitation id="EF96D58A465FC272FDC5F960FD00F880" author="Bailey, R. M." box="[566,721,1788,1815]" pageId="6" pageNumber="331" pagination="1 - 19" refId="ref5656" refString="Bailey, R. M. (1959) A new catostomid fish, Moxostoma (Thoburnia) atripinne, from the Green River drainage, Kentucky and Tennessee. Occasional Papers of the Museum of Zoology, The University of Michigan, 559, 1 - 19." type="journal article" year="1959">Bailey (1959)</bibRefCitation>
reported spawning males in water eight centimeters deep from 610 April while
<bibRefCitation id="EF96D58A465FC272FEAEF8BCFD88F8AC" author="Timmons, T. J. &amp; Ramsey, J. S. &amp; Bauer, B. H." box="[349,601,1824,1851]" pageId="6" pageNumber="331" pagination="538 - 541" refId="ref7694" refString="Timmons, T. J., Ramsey, J. S. &amp; Bauer, B. H. (1983) Life history and habitat of the blackfin sucker, Moxostoma atripinne (Osteichthyes: Catostomidae). Copeia, 1983, 538 - 541. https: // doi. org / 10.2307 / 1444405" type="journal article" year="1983">
Timmons
<emphasis id="B9737469465FC272FE3CF8BDFDD6F8AD" box="[463,519,1824,1850]" italics="true" pageId="6" pageNumber="331">et al.</emphasis>
(1983)
</bibRefCitation>
reported tuberculate males in March. Males gather in riffles weeks before spawning females arrive (females were normally in pools). Egg size is
<quantity id="4CFF059E465FC272FC55F8D8FBFAF8C8" box="[934,1067,1860,1887]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="6" pageNumber="331" unit="mm" value="2.1" valueMax="2.5" valueMin="1.7">1.72.5 mm</quantity>
and fecundity ranged from 1,070 to 1,755 eggs per female. Maximum reported age is five years.
</paragraph>
</subSubSection>
<subSubSection id="C31DFBF0465FC272FF34F811FEBBF879" pageId="6" pageNumber="331" type="etymology">
<paragraph id="8BB8A87B465FC272FF34F811FEBBF879" blockId="6.[151,1437,1464,2030]" pageId="6" pageNumber="331">
<emphasis id="B9737469465FC272FF34F811FE9FF830" bold="true" box="[199,334,1933,1959]" pageId="6" pageNumber="331">Etymology.</emphasis>
From the Latin
<emphasis id="B9737469465FC272FDF2F810FDB1F831" box="[513,608,1932,1958]" italics="true" pageId="6" pageNumber="331">vexillum</emphasis>
for flag and the Greek
<emphasis id="B9737469465FC272FC93F810FC7EF831" box="[864,943,1932,1958]" italics="true" pageId="6" pageNumber="331">ichthys</emphasis>
for fish. The dorsal fin has a flaglike pattern and the approximately thirteen alternating dark and light stripes are like the flag of the
<collectingCountry id="F310E8EB465FC272FB84F82CFA48F85D" box="[1143,1433,1968,1995]" name="United States of America" pageId="6" pageNumber="331">United States of America</collectingCountry>
. Gender masculine.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,771 @@
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<mods:start id="2EC4ED8AD1B45B577AE2B83E269893A7">248</mods:start>
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<mods:location id="80EDF981A7B150BF8507B3B959B96F9E">
<mods:url id="97E22DBB2B6FF3EDDA7EAAAB5DAC789E">http://dx.doi.org/10.11646/zootaxa.5536.2.2</mods:url>
</mods:location>
<mods:classification id="83862220CDA0FA1BB42D2CC719149902">journal article</mods:classification>
<mods:identifier id="75E9352BFF2AA775F3C24687C63F28CD" type="DOI">10.11646/zootaxa.5536.2.2</mods:identifier>
<mods:identifier id="16324723D66BE580544DF8372B47E6CB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8EEFA96EB066CF6E966BBB9FCF7D94F1" type="Zenodo-Dep">14033544</mods:identifier>
<mods:identifier id="F7EF4BFC46E7975678BD4895F82CC08F" type="ZooBank">E1AAF32C-6B89-4704-A408-3D7545283171</mods:identifier>
</mods:mods>
<treatment id="03BFF8153011FF9AEAE5FC6EFC5A683E" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E" httpUri="http://treatment.plazi.org/id/03BFF8153011FF9AEAE5FC6EFC5A683E" lastPageId="10" lastPageNumber="258" pageId="4" pageNumber="252">
<subSubSection id="C30C1A883011FF94EAE5FC6EFD0068C7" box="[151,714,934,961]" pageId="4" pageNumber="252" type="nomenclature">
<paragraph id="8BA949033011FF94EAE5FC6EFD0068C7" blockId="4.[151,714,934,961]" box="[151,714,934,961]" pageId="4" pageNumber="252">
<heading id="D0E1FE6F3011FF94EAE5FC6EFD0068C7" bold="true" box="[151,714,934,961]" fontSize="11" level="1" pageId="4" pageNumber="252" reason="1">
<taxonomicName id="4C1632803011FF94EAE5FC6EFD0068C7" authority="(Meyrick 1932)" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[151,714,934,961]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="252" phylum="Arthropoda" rank="species" species="stichocentra">
<emphasis id="B96295113011FF94EAE5FC6EFD0068C7" bold="true" box="[151,714,934,961]" pageId="4" pageNumber="252">
<emphasis id="B96295113011FF94EAE5FC6EFDC968C6" bold="true" box="[151,515,934,961]" italics="true" pageId="4" pageNumber="252">Pararhigognostis stichocentra</emphasis>
(
<bibRefCitation id="EF8734F23011FF94E861FC6EFD0868C7" author="Meyrick, E." box="[531,706,934,960]" pageId="4" pageNumber="252" pagination="107 - 120" refId="ref5877" refString="Meyrick, E. (1932) Entomological expedition to Abyssinia, 1926 - 27 Microlepidoptera. The Transaction of the Entomological Society of London, 80 (1), 107 - 120. https: // doi. org / 10.1111 / j. 1365 - 2311.1932. tb 00085. x" type="journal article" year="1932">Meyrick 1932</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30C1A883011FF94EAE5FC2FFC826F5C" pageId="4" pageNumber="252" type="reference_group">
<paragraph id="8BA949033011FF94EAE5FC2FFD8A68F8" blockId="4.[151,854,999,1115]" box="[151,576,999,1023]" pageId="4" pageNumber="252">
<treatmentCitationGroup id="AB066E2D3011FF94EAE5FC2FFD8A68F8" box="[151,576,999,1023]" pageId="4" pageNumber="252">
<treatmentCitation id="0AB76F123011FF94EAE5FC2FFDF668F8" author="Meyrick, E." box="[151,572,999,1023]" page="118" pageId="4" pageNumber="252" year="1932">
<taxonomicName id="4C1632803011FF94EAE5FC2FFDF668F8" ID-CoL="4KJYY" authority="Meyrick, 1932: 118" authorityName="Meyrick" authorityPageNumber="118" authorityYear="1932" box="[151,572,999,1023]" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="252" phylum="Arthropoda" rank="species" species="stichocentra">
<emphasis id="B96295113011FF94EAE5FC2FFEA168F8" box="[151,363,999,1023]" italics="true" pageId="4" pageNumber="252">Plutella stichocentra</emphasis>
<bibRefCitation id="EF8734F23011FF94EB03FC2FFDF668F8" author="Meyrick, E." box="[369,572,999,1023]" pageId="4" pageNumber="252" pagination="107 - 120" refId="ref5877" refString="Meyrick, E. (1932) Entomological expedition to Abyssinia, 1926 - 27 Microlepidoptera. The Transaction of the Entomological Society of London, 80 (1), 107 - 120. https: // doi. org / 10.1111 / j. 1365 - 2311.1932. tb 00085. x" type="journal article" year="1932">Meyrick, 1932: 118</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA949033011FF94EAE5FBCEFC9C6F19" blockId="4.[151,854,999,1115]" box="[151,854,1030,1054]" pageId="4" pageNumber="252">
<taxonomicName id="4C1632803011FF94EAE5FBCEFDC06F19" ID-CoL="4KJYY" authority="Meyrick, 1932" authorityName="Meyrick" authorityYear="1932" box="[151,522,1030,1054]" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="252" phylum="Arthropoda" rank="species" species="stichocentra">
<emphasis id="B96295113011FF94EAE5FBCEFEA16F19" box="[151,363,1030,1054]" italics="true" pageId="4" pageNumber="252">Plutella stichocentra</emphasis>
<bibRefCitation id="EF8734F23011FF94EB03FBCEFDC06F19" author="Meyrick, E." box="[369,522,1030,1054]" pageId="4" pageNumber="252" pagination="107 - 120" refId="ref5877" refString="Meyrick, E. (1932) Entomological expedition to Abyssinia, 1926 - 27 Microlepidoptera. The Transaction of the Entomological Society of London, 80 (1), 107 - 120. https: // doi. org / 10.1111 / j. 1365 - 2311.1932. tb 00085. x" type="journal article" year="1932">Meyrick, 1932</bibRefCitation>
</taxonomicName>
: Afromoths.net online data base.
</paragraph>
<paragraph id="8BA949033011FF94EAE5FBECFD5C6F3B" blockId="4.[151,854,999,1115]" box="[151,662,1060,1085]" pageId="4" pageNumber="252">
<treatmentCitationGroup id="AB066E2D3011FF94EAE5FBECFD5C6F3B" box="[151,662,1060,1085]" pageId="4" pageNumber="252">
<treatmentCitation id="0AB76F123011FF94EAE5FBECFDE76F3A" author="Bradley J. D." box="[151,557,1060,1085]" page="113" pageId="4" pageNumber="252" year="1965">
<taxonomicName id="4C1632803011FF94EAE5FBECFDE76F3A" ID-CoL="4KJYZ" authority="Bradley, 1965: 113" authorityName="Bradley" authorityPageNumber="113" authorityYear="1965" box="[151,557,1060,1085]" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="252" phylum="Arthropoda" rank="species" species="symmorpha" status="syn. nov.">
<emphasis id="B96295113011FF94EAE5FBECFEAE6F3B" box="[151,356,1060,1084]" italics="true" pageId="4" pageNumber="252">Plutella symmorpha</emphasis>
<bibRefCitation id="EF8734F23011FF94EB18FBECFDE76F3A" author="Bradley J. D." box="[362,557,1060,1085]" pageId="4" pageNumber="252" pagination="81 - 148" refId="ref5179" refString="Bradley J. D. (1965) Ruwenzori Expedition 1952. Microlepidoptera. Ruwenzori Expedition. 2 (12). Trustees of the British Museum (Natural History), London, pp. 81 - 148." type="book chapter" year="1965">Bradley, 1965: 113</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<taxonomicNameLabel id="A251286A3011FF94E845FBEDFD5C6F3B" box="[567,662,1061,1085]" pageId="4" pageNumber="252">
<emphasis id="B96295113011FF94E845FBEDFD5A6F3A" bold="true" box="[567,656,1061,1085]" pageId="4" pageNumber="252">syn. nov</emphasis>
.
</taxonomicNameLabel>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA949033011FF94EAE5FB8BFC826F5C" blockId="4.[151,854,999,1115]" box="[151,840,1091,1115]" pageId="4" pageNumber="252">
<taxonomicName id="4C1632803011FF94EAE5FB8BFE316F5C" ID-CoL="4KJYZ" authority="Bradley, 1965" authorityName="Bradley" authorityYear="1965" box="[151,507,1091,1115]" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="252" phylum="Arthropoda" rank="species" species="symmorpha">
<emphasis id="B96295113011FF94EAE5FB8BFEAE6F5C" box="[151,356,1091,1115]" italics="true" pageId="4" pageNumber="252">Plutella symmorpha</emphasis>
<bibRefCitation id="EF8734F23011FF94EB18FB8BFE316F5C" author="Bradley J. D." box="[362,507,1091,1115]" pageId="4" pageNumber="252" pagination="81 - 148" refId="ref5179" refString="Bradley J. D. (1965) Ruwenzori Expedition 1952. Microlepidoptera. Ruwenzori Expedition. 2 (12). Trustees of the British Museum (Natural History), London, pp. 81 - 148." type="book chapter" year="1965">Bradley, 1965</bibRefCitation>
</taxonomicName>
: Afromoths.net online data base.
</paragraph>
</subSubSection>
<subSubSection id="C30C1A883011FF94EAE5FB4DFE6B6FC4" pageId="4" pageNumber="252" type="diagnosis">
<paragraph id="8BA949033011FF94EAE5FB4DFE6B6FC4" blockId="4.[151,1437,1157,2041]" pageId="4" pageNumber="252">
<emphasis id="B96295113011FF94EAE5FB4DFED86F98" bold="true" box="[151,274,1157,1183]" pageId="4" pageNumber="252">Diagnosis.</emphasis>
Because the genus is monotypic, features detailed in the diagnosis of the genus above will serve to distinguish this species.
</paragraph>
</subSubSection>
<subSubSection id="C30C1A883011FF94EAB5FB04FBCA6DDC" pageId="4" pageNumber="252" type="description">
<paragraph id="8BA949033011FF94EAB5FB04FCE36D03" blockId="4.[151,1437,1157,2041]" pageId="4" pageNumber="252">
<emphasis id="B96295113011FF94EAB5FB04FEA76FE1" bold="true" box="[199,365,1228,1254]" pageId="4" pageNumber="252">Redescription</emphasis>
(
<figureCitation id="132D55863011FF94EBF2FB04FE776FE1" box="[384,445,1228,1254]" captionStart="FIGURES 12" captionStartId="5.[152,270,1531,1555]" captionTargetBox="[218,1370,179,1506]" captionTargetId="figure-26@5.[218,1370,179,1506]" captionTargetPageId="5" captionText="FIGURES 12. 1. Pararhigognostis stichocentra: adult female, paralectotype, preserved in coll. Meyrick, in NHMUK; 2. paralectotype labels: A, type, female, B, label of type (paralectotype), C, accession number in NHMUK, D, localities, E, note about of host plant." figureDoi="http://doi.org/10.5281/zenodo.14033546" httpUri="https://zenodo.org/record/14033546/files/figure.png" pageId="4" pageNumber="252">Fig.1</figureCitation>
). The forewing span
<quantity id="4CEEE4E63011FF94E8C0FB04FCED6FE1" box="[690,807,1228,1254]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="4" pageNumber="252" unit="mm" value="19.0" valueMax="22.0" valueMin="16.0">1622mm</quantity>
. Forewing mostly grey-brown, with a small proportion of single white and brown scales; costal margin grey-brown, with a small proportion of white-yellowish scales apically; basal area grey-brown, near costal margin in central part with admixture of yellowish ones, and near dorsal margin with conspicuous, large proportion of white scales; inconspicuous dark band composed of black scales at wing base, interrupted, reaching 2/3 of dorsal margin length; a narrow band basal band extending from its base to the same point, composed of yellow scales; in central parts of wings, above this a bicoloured band, a narrow yellow band starts, extending to its outer margin; wing apex grey-brown, with scattered yellowish scales; cilia coloured similarly, with a markedly larger proportion of white scales; dorsal margin white-grey from its base to 2/3 its length, with single, irregularly scattered black scales at its margin.
</paragraph>
<paragraph id="8BA949033011FF94EAB5F9C6FC286D2F" blockId="4.[151,1437,1157,2041]" box="[199,994,1550,1576]" pageId="4" pageNumber="252">Hindwing lanceolate, mostly white-grey; cilia long, coloured similarly.</paragraph>
<paragraph id="8BA949033011FF94EAB5F9FAFEB06D77" blockId="4.[151,1437,1157,2041]" pageId="4" pageNumber="252">Abdomen: white-grey, scales slightly shiny. Male genitalia as described from genus. Female genitalia as described for genus.</paragraph>
<paragraph id="8BA949033011FF94EAB5F9B1FBCA6DDC" blockId="4.[151,1437,1157,2041]" pageId="4" pageNumber="252">
<emphasis id="B96295113011FF94EAB5F9B1FECC6D94" bold="true" box="[199,262,1657,1683]" pageId="4" pageNumber="252">Note.</emphasis>
We designate
<specimenCount id="9D10828A3011FF94EBC2F9B1FE196D94" box="[432,467,1657,1683]" count="1" pageId="4" pageNumber="252" type="male">1♂</specimenCount>
as a
<typeStatus id="54ADF7A13011FF94E860F9B1FDB36D94" box="[530,633,1657,1683]" pageId="4" pageNumber="252" type="lectotype">lectotype</typeStatus>
<materialsCitation id="3B7E435E3011FF94E8F0F9B1FBCA6DDC" collectingDate="1926-12-18" collectorName="H. Scott &amp; Britt. Mus. &amp; Natural History Museum &amp; London" country="United Kingdom" elevation="3810" location="Meyrick" municipality="Mt. Chilalo" pageId="4" pageNumber="252" specimenCount="8" specimenCount-female="4" specimenCount-male="4" typeStatus="paralectotype">
(marked by
<location id="8EC91FD83011FF94E97FF9B1FCA66D94" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83011FF94E97FF9B1FCA66D94" box="[781,876,1657,1683]" country="United Kingdom" municipality="Mt. Chilalo" name="Meyrick" pageId="4" pageNumber="252">Meyrick</location>
as a type (red label), (
<collectingMunicipality id="6BCDD3793011FF94EE01F9B1FB306D94" box="[1139,1274,1657,1683]" pageId="4" pageNumber="252">Mt. Chilalo</collectingMunicipality>
, moorland, ca
<quantity id="4CEEE4E63011FF94EAE5F955FE9B6DB0" box="[151,337,1693,1719]" metricMagnitude="3" metricUnit="m" metricValue="3.81" metricValueMax="3.9624" metricValueMin="3.6576" pageId="4" pageNumber="252" unit="ft" value="12500.0" valueMax="13000.0" valueMin="12000.0">
<elevation id="003BAE303011FF94EAE5F955FE9B6DB0" box="[151,337,1693,1719]" metricMagnitude="3" metricUnit="m" metricValue="3.81" metricValueMax="3.9624" metricValueMin="3.6576" pageId="4" pageNumber="252" unit="ft" value="12500.0" valueMax="13000.0" valueMin="12000.0">1200013000 ft.</elevation>
</quantity>
<date id="FFA86FC33011FF94EB2AF955FE106DB0" box="[344,474,1693,1719]" pageId="4" pageNumber="252" value="1926-12-18">
<collectingDate id="EFEC962B3011FF94EB2AF955FE106DB0" box="[344,474,1693,1719]" pageId="4" pageNumber="252" value="1926-12-18">18.12.1926</collectingDate>
</date>
.
<collectorName id="26E32CD53011FF94EB97F956FD8B6DB0" box="[485,577,1693,1720]" pageId="4" pageNumber="252">H. Scott</collectorName>
leg.
<collectorName id="26E32CD53011FF94E80BF955FD3E6DBF" box="[633,756,1693,1720]" pageId="4" pageNumber="252">Britt. Mus.</collectorName>
1927-127, labelled as type, preserved in the
<collectorName id="26E32CD53011FF94EE9FF955FF366DDC" pageId="4" pageNumber="252">Natural History Museum</collectorName>
,
<collectorName id="26E32CD53011FF94EB75F909FEAB6DDC" box="[263,353,1729,1755]" pageId="4" pageNumber="252">London</collectorName>
,
<collectingCountry id="F30109933011FF94EB1EF909FE016DDC" box="[364,459,1729,1755]" name="United Kingdom" pageId="4" pageNumber="252">England</collectingCountry>
) and all the others (
<specimenCount id="9D10828A3011FF94E8D8F909FD2F6DDC" box="[682,741,1729,1755]" count="4" pageId="4" pageNumber="252" type="male">4♂♂</specimenCount>
,
<specimenCount id="9D10828A3011FF94E882F909FCE26DDC" box="[752,808,1729,1755]" count="4" pageId="4" pageNumber="252" type="female">4♀♀</specimenCount>
) as
<typeStatus id="54ADF7A13011FF94E925F909FC366DDC" box="[855,1020,1729,1755]" pageId="4" pageNumber="252" type="paralectotype">paralectotypes</typeStatus>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30C1A883011FF9AEAB5F92DFC456AF2" lastPageId="10" lastPageNumber="258" pageId="4" pageNumber="252" type="materials_examined">
<paragraph id="8BA949033011FF94EAB5F92DFA826C6D" blockId="4.[151,1437,1157,2041]" pageId="4" pageNumber="252">
<emphasis id="B96295113011FF94EAB5F92DFEA26DF9" bold="true" box="[199,360,1764,1791]" pageId="4" pageNumber="252">
<typeStatus id="54ADF7A13011FF94EAB5F92DFEC86DF8" box="[199,258,1765,1791]" pageId="4" pageNumber="252">Type</typeStatus>
locality.
</emphasis>
Mount Chilalo,
<collectingCountry id="F30109933011FF94E855F92CFD5D6DF9" box="[551,663,1764,1790]" name="Ethiopia" pageId="4" pageNumber="252">Abyssinia</collectingCountry>
(
<bibRefCitation id="EF8734F23011FF94E8DAF92CFC846DF8" author="Meyrick, E." box="[680,846,1764,1791]" pageId="4" pageNumber="252" pagination="107 - 120" refId="ref5877" refString="Meyrick, E. (1932) Entomological expedition to Abyssinia, 1926 - 27 Microlepidoptera. The Transaction of the Entomological Society of London, 80 (1), 107 - 120. https: // doi. org / 10.1111 / j. 1365 - 2311.1932. tb 00085. x" type="journal article" year="1932">Meyrick 1932</bibRefCitation>
), is an isolated, volcanic mountain in south-eastern
<collectingCountry id="F30109933011FF94EAE5F8C0FF306C25" box="[151,250,1800,1826]" name="Ethiopia" pageId="4" pageNumber="252">Ethiopia</collectingCountry>
. The highest point (
<quantity id="4CEEE4E63011FF94EBA1F8C0FDE06C24" box="[467,554,1800,1827]" metricMagnitude="3" metricUnit="m" metricValue="4.036" pageId="4" pageNumber="252" unit="m" value="4036.0">4036 m</quantity>
a.s.l.), in the Arsi Zone of the
<collectingRegion id="49D287E13011FF94E908F8C0FBE06C25" box="[890,1066,1800,1826]" country="Ethiopia" name="Oromiya" pageId="4" pageNumber="252">Oromia Region</collectingRegion>
, is located on the border between the Hitosa and Tiyo districts. The volcano last erupted in the Pleistocene (
<bibRefCitation id="EF8734F23011FF94E995F8E4FB3C6C40" author="Gashaw, M. &amp; Didita, M." box="[999,1270,1836,1863]" pageId="4" pageNumber="252" pagination="1 - 201" refId="ref5670" refString="Gashaw, M. &amp; Didita, M. (2015) Ethiopian panel on climate change, In: First Assessment Report, Working group II. Biodiversity and Ecosystems. Ethiopian Academy of Sciences, Addis Abeba, pp. 1 - 201." type="book chapter" year="2015">Gashaw &amp; Didita 2015</bibRefCitation>
). Its Ethiopian afro-alpine and subafroalpine ecosystems are unique habitats (
<bibRefCitation id="EF8734F23011FF94E922F898FB836C6D" author="Schweiger, A. H. &amp; Otieno, D. O. &amp; Kulunge, D. O. &amp; Reineking, B. &amp; Tenhunen, J." box="[848,1097,1871,1898]" pageId="4" pageNumber="252" pagination="293 - 303" refId="ref6366" refString="Schweiger, A. H., Otieno, D. O., Kulunge, D. O., Reineking, B. &amp; Tenhunen, J. (2015) The Afro-alpine dwarf shrub Helichrysum citrispinum favours undersorey plants through microclimate amelioration. Plant Ecology and Diversity, 8 (3), 293 - 303. https: // doi. org / 10.1080 / 17550874.2015.1014207" type="journal article" year="2015">
Schweiger
<emphasis id="B96295113011FF94E9BDF898FBC26C6D" box="[975,1032,1872,1898]" italics="true" pageId="4" pageNumber="252">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="EF8734F23011FF94EE27F898FAF16C6D" author="Uhlig, S. K. &amp; Uhlig, K." box="[1109,1339,1872,1898]" pageId="4" pageNumber="252" pagination="153 - 156" refId="ref6648" refString="Uhlig, S. K. &amp; Uhlig, K. (1991) Studies on the Altitudinal Zonation of Forests and Alpine Plants in the Central Bale Mountains, Ethiopia. Mountain Research and Development, 11 (2), 153 - 156. https: // doi. org / 10.2307 / 3673574" type="journal article" year="1991">Uhlig &amp; Uhlig 1991</bibRefCitation>
).
</paragraph>
<paragraph id="8BA949033011FF94EAB5F8BBFD1D6C89" blockId="4.[151,1437,1157,2041]" box="[199,727,1907,1934]" pageId="4" pageNumber="252">
<emphasis id="B96295113011FF94EAB5F8BBFDFC6C8A" bold="true" box="[199,566,1907,1933]" pageId="4" pageNumber="252">Additional material examined:</emphasis>
<specimenCount id="9D10828A3011FF94E84FF8BCFD4C6C89" box="[573,646,1908,1934]" count="24" pageId="4" pageNumber="252" type="male">24♂♂</specimenCount>
,
<specimenCount id="9D10828A3011FF94E8E3F8BCFD1D6C89" box="[657,727,1908,1934]" count="21" pageId="4" pageNumber="252" type="female">21♀♀</specimenCount>
</paragraph>
<paragraph id="8BA949033011FF94EAB5F850FEE06CFE" blockId="4.[151,1437,1157,2041]" pageId="4" pageNumber="252">
<materialsCitation id="3B7E435E3011FF94EAB5F850FA5D6CB6" box="[199,1431,1943,1970]" collectingDate="1926-12-18" collectionCode="NHMUK" collectorName="H. Scott" country="Ethiopia" elevation="3810" location="Mt. Chilalo" pageId="4" pageNumber="252" specimenCount="9" specimenCount-female="4" specimenCount-male="5">
<collectionCode id="ED07D1C63011FF94EAB5F850FEFF6CB5" box="[199,309,1944,1970]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="252" type="Museum">NHMUK</collectionCode>
:
<specimenCount id="9D10828A3011FF94EB37F85FFEB46CB5" box="[325,382,1943,1970]" count="5" pageId="4" pageNumber="252" type="male">5♂♂</specimenCount>
,
<specimenCount id="9D10828A3011FF94EBFDF85FFE026CB6" box="[399,456,1943,1969]" count="4" pageId="4" pageNumber="252" type="female">4♀♀</specimenCount>
,
<collectingCountry id="F30109933011FF94EBABF85FFD876CB6" box="[473,589,1943,1969]" name="Ethiopia" pageId="4" pageNumber="252">Abyssinia</collectingCountry>
:
<location id="8EC91FD83011FF94E82EF850FD2F6CB6" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83011FF94E82EF850FD2F6CB6" box="[604,741,1943,1970]" country="Ethiopia" name="Mt. Chilalo" pageId="4" pageNumber="252">Mt. Chilalo</location>
, moorland, ca
<quantity id="4CEEE4E63011FF94E9EAF85FFBA86CB6" box="[920,1122,1943,1969]" metricMagnitude="3" metricUnit="m" metricValue="3.81" metricValueMax="3.9624" metricValueMin="3.6576" pageId="4" pageNumber="252" unit="ft" value="12500.0" valueMax="13000.0" valueMin="12000.0">
<elevation id="003BAE303011FF94E9EAF85FFBA86CB6" box="[920,1122,1943,1969]" metricMagnitude="3" metricUnit="m" metricValue="3.81" metricValueMax="3.9624" metricValueMin="3.6576" pageId="4" pageNumber="252" unit="ft" value="12500.0" valueMax="13000.0" valueMin="12000.0">12000 13000 ft.</elevation>
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<date id="FFA86FC33011FF94EE1CF85FFB3A6CB6" box="[1134,1264,1943,1969]" pageId="4" pageNumber="252" value="1926-12-18">
<collectingDate id="EFEC962B3011FF94EE1CF85FFB3A6CB6" box="[1134,1264,1943,1969]" pageId="4" pageNumber="252" value="1926-12-18">18.12.1926</collectingDate>
</date>
.
<collectorName id="26E32CD53011FF94EE8DF850FAAA6CB6" box="[1279,1376,1943,1970]" pageId="4" pageNumber="252">H. Scott</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E3011FF94EAE5F873FB806CD2" box="[151,1098,1979,2005]" collectionCode="NHMUK" collectorName="F. W. Edwards" country="Kenya" elevation="2438" location="Aberdare Range" municipality="Mt. Kinango" pageId="4" pageNumber="252" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D10828A3011FF94EAE5F873FF766CD2" box="[151,188,1979,2005]" count="1" pageId="4" pageNumber="252" type="male">1♂</specimenCount>
,
<collectingCountry id="F30109933011FF94EABFF873FED06CD2" box="[205,282,1979,2005]" name="Kenya" pageId="4" pageNumber="252">Kenya</collectingCountry>
:
<location id="8EC91FD83011FF94EB5AF873FE2C6CD2" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83011FF94EB5AF873FE2C6CD2" box="[296,486,1979,2005]" country="Kenya" municipality="Mt. Kinango" name="Aberdare Range" pageId="4" pageNumber="252">Aberdare Range</location>
,
<collectingMunicipality id="6BCDD3793011FF94EB85F873FD476CD2" box="[503,653,1979,2005]" pageId="4" pageNumber="252">Mt. Kinango</collectingMunicipality>
,
<quantity id="4CEEE4E63011FF94E8EFF873FD306CD2" box="[669,762,1979,2005]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="4" pageNumber="252" unit="ft" value="8000.0">
<elevation id="003BAE303011FF94E8EFF873FD306CD2" box="[669,762,1979,2005]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="4" pageNumber="252" unit="ft" value="8000.0">8000 ft.</elevation>
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10.1934.
<collectorName id="26E32CD53011FF94E901F873FBDE6CD2" box="[883,1044,1979,2005]" pageId="4" pageNumber="252">F.W. Edwards</collectorName>
leg.
</materialsCitation>
(
<bibRefCitation id="EF8734F23011FF94EE2FF873FAC56CD1" author="Bradley J. D." box="[1117,1295,1979,2006]" pageId="4" pageNumber="252" pagination="81 - 148" refId="ref5179" refString="Bradley J. D. (1965) Ruwenzori Expedition 1952. Microlepidoptera. Ruwenzori Expedition. 2 (12). Trustees of the British Museum (Natural History), London, pp. 81 - 148." type="book chapter" year="1965">Bradley (1965)</bibRefCitation>
, as
<taxonomicName id="4C1632803011FF94EF36F873FED06CFE" authorityName="Bradley" authorityYear="1965" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="252" phylum="Arthropoda" rank="species" species="symmorpha">
<emphasis id="B96295113011FF94EF36F873FED06CFE" italics="true" pageId="4" pageNumber="252">Plutella symmorpha</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF69198B3010FF95EAEAFA33FE966D5C" ID-DOI="http://doi.org/10.5281/zenodo.14033546" ID-Zenodo-Dep="14033546" httpUri="https://zenodo.org/record/14033546/files/figure.png" pageId="5" pageNumber="253" startId="5.[152,270,1531,1555]" targetBox="[218,1370,179,1506]" targetPageId="5" targetType="figure">
<paragraph id="8BA949033010FF95EAEAFA33FE966D5C" blockId="5.[151,1437,1531,1627]" pageId="5" pageNumber="253">
<emphasis id="B96295113010FF95EAEAFA33FE8E6D14" bold="true" box="[152,324,1531,1555]" pageId="5" pageNumber="253">FIGURES 12.</emphasis>
1.
<taxonomicName id="4C1632803010FF95EB1BFA33FD6A6D14" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[361,672,1531,1555]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="stichocentra">
<emphasis id="B96295113010FF95EB1BFA33FD6A6D14" box="[361,672,1531,1555]" italics="true" pageId="5" pageNumber="253">Pararhigognostis stichocentra</emphasis>
</taxonomicName>
: adult female, paralectotype, preserved in coll. Meyrick, in NHMUK; 2. paralectotype labels: A, type, female, B, label of type (paralectotype), C, accession number in NHMUK, D, localities, E, note about of host plant.
</paragraph>
</caption>
<paragraph id="8BA949033010FF95EAB5F966FCDE6CA7" blockId="5.[151,1437,1710,1952]" pageId="5" pageNumber="253">
DA;
<collectingCountry id="F30109933010FF95EA8DF966FE866DCF" box="[255,332,1710,1736]" name="Kenya" pageId="5" pageNumber="253">Kenya</collectingCountry>
:
<specimenCount id="9D10828A3010FF95EB25F966FE5A6DCF" box="[343,400,1710,1736]" count="3" pageId="5" pageNumber="253" type="female">3♀♀</specimenCount>
,
<date id="FFA86FC33010FF95EBEFF966FDD46DCF" box="[413,542,1710,1736]" pageId="5" pageNumber="253" value="1998-10-16">16.10.1998</date>
,
<date id="FFA86FC33010FF95E858F966FD616DCF" box="[554,683,1710,1736]" pageId="5" pageNumber="253" value="1998-10-31">31.10.1998</date>
,
<date id="FFA86FC33010FF95E8C4F966FCE36DCE" box="[694,809,1710,1737]" pageId="5" pageNumber="253" value="1999-05-02">2.05.1999</date>
,
<materialsCitation id="3B7E435E3010FF95E946F966FEE96DEB" collectingDate="1999-05-13" collectorName="D. J. L. Agassiz" elevation="2438" location="Turi" municipality="Rift Valley Prov." pageId="5" pageNumber="253" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D10828A3010FF95E946F966FC936DCF" box="[820,857,1710,1736]" count="1" pageId="5" pageNumber="253" type="male">1♂</specimenCount>
,
<date id="FFA86FC33010FF95E914F966FC2D6DCE" box="[870,999,1710,1737]" pageId="5" pageNumber="253" value="1999-05-13">
<collectingDate id="EFEC962B3010FF95E914F966FC2D6DCE" box="[870,999,1710,1737]" pageId="5" pageNumber="253" value="1999-05-13">13.05.1999</collectingDate>
</date>
,
<collectingMunicipality id="6BCDD3793010FF95E980F966FB7B6DCF" box="[1010,1201,1710,1736]" pageId="5" pageNumber="253">Rift Valley Prov.</collectingMunicipality>
,
<location id="8EC91FD83010FF95EEC9F966FB276DCF" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83010FF95EEC9F966FB276DCF" box="[1211,1261,1710,1736]" municipality="Rift Valley Prov." name="Turi" pageId="5" pageNumber="253">Turi</location>
,
<quantity id="4CEEE4E63010FF95EE8BF966FA866DCF" box="[1273,1356,1710,1736]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="5" pageNumber="253" unit="ft" value="8000.0">
<elevation id="003BAE303010FF95EE8BF966FA866DCF" box="[1273,1356,1710,1736]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="5" pageNumber="253" unit="ft" value="8000.0">8000ft.</elevation>
</quantity>
,
<collectorName id="26E32CD53010FF95EF25F966FF3A6DEB" pageId="5" pageNumber="253">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
;
<specimenCount id="9D10828A3010FF95EB5DF91AFE986DEB" box="[303,338,1746,1772]" count="1" pageId="5" pageNumber="253" type="female">1♀</specimenCount>
,
<date id="FFA86FC33010FF95EB12F91AFE2B6DEB" box="[352,481,1746,1772]" pageId="5" pageNumber="253" value="1998-10-31">31.10.1998</date>
,
<materialsCitation id="3B7E435E3010FF95EB9FF91AFB106DEB" box="[493,1242,1746,1773]" collectingDate="1999-12-15" collectorName="D. J. L. Agassiz" location="Lake Elementeita" municipality="Rift Valley" pageId="5" pageNumber="253" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D10828A3010FF95EB9FF91AFDDA6DEB" box="[493,528,1746,1772]" count="1" pageId="5" pageNumber="253" type="female">1♀</specimenCount>
,
<date id="FFA86FC33010FF95E86CF91AFD556DEA" box="[542,671,1746,1773]" pageId="5" pageNumber="253" value="1999-12-15">
<collectingDate id="EFEC962B3010FF95E86CF91AFD556DEA" box="[542,671,1746,1773]" pageId="5" pageNumber="253" value="1999-12-15">15.12.1999</collectingDate>
</date>
,
<collectingMunicipality id="6BCDD3793010FF95E8D8F91AFCEF6DEB" box="[682,805,1746,1772]" pageId="5" pageNumber="253">Rift Valley</collectingMunicipality>
,
<location id="8EC91FD83010FF95E942F91AFC326DEB" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83010FF95E942F91AFC326DEB" box="[816,1016,1746,1772]" municipality="Rift Valley" name="Lake Elementeita" pageId="5" pageNumber="253">Lake Elementeita</location>
,
<collectorName id="26E32CD53010FF95EE71F91AFB6D6DEB" box="[1027,1191,1746,1772]" pageId="5" pageNumber="253">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E3010FF95EE94F91AFD406C17" collectingDate="2003-02-05" collectorName="D. J. L. Agassiz" location="Lake" municipality="Rift Valley" pageId="5" pageNumber="253" specimenCount="1" specimenCount-male="1" stateProvince="Baringo">
<specimenCount id="9D10828A3010FF95EE94F91AFAC06DEB" box="[1254,1290,1746,1772]" count="1" pageId="5" pageNumber="253" type="male">1♂</specimenCount>
,
<date id="FFA86FC33010FF95EF65F91AFA526DEA" box="[1303,1432,1746,1773]" pageId="5" pageNumber="253" value="2003-02-05">
<collectingDate id="EFEC962B3010FF95EF65F91AFA526DEA" box="[1303,1432,1746,1773]" pageId="5" pageNumber="253" value="2003-02-05">02.05.2003</collectingDate>
</date>
,
<collectingMunicipality id="6BCDD3793010FF95EAE5F93EFEC56C17" box="[151,271,1782,1808]" pageId="5" pageNumber="253">Rift Valley</collectingMunicipality>
,
<location id="8EC91FD83010FF95EB65F93EFE856C17" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83010FF95EB65F93EFE856C17" box="[279,335,1782,1808]" municipality="Rift Valley" name="Lake" pageId="5" pageNumber="253" stateProvince="Baringo">Lake</location>
<collectingRegion id="49D287E13010FF95EB26F93EFE7B6C17" box="[340,433,1782,1808]" country="Kenya" name="Baringo" pageId="5" pageNumber="253">Baringo</collectingRegion>
,
<collectorName id="26E32CD53010FF95EBCBF93EFD906C17" box="[441,602,1782,1808]" pageId="5" pageNumber="253">D.J.L.Agassiz</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E3010FF95E8E1F93EFEA46C33" collectingDate="20.10.1998, 02.10.2004, 05.12.2011" collectorName="D. J. L. Agassiz" location="Lake Naivasha" municipality="Rift Valley" pageId="5" pageNumber="253" specimenCount="3" specimenCount-male="3" stateProvince="Baringo">
<specimenCount id="9D10828A3010FF95E8E1F93EFD076C17" box="[659,717,1782,1808]" count="3" pageId="5" pageNumber="253" type="male">3♂♂</specimenCount>
,
<collectingDate id="EFEC962B3010FF95E8A4F93EFBA26C16" box="[726,1128,1782,1809]" isEnumeration="true" pageId="5" pageNumber="253">
<date id="FFA86FC33010FF95E8A4F93EFC9D6C17" box="[726,855,1782,1808]" pageId="5" pageNumber="253" value="1998-10-20">20.10.1998</date>
,
<date id="FFA86FC33010FF95E92DF93EFC2A6C17" box="[863,992,1782,1808]" pageId="5" pageNumber="253" value="2004-02-10">02.10.2004</date>
,
<date id="FFA86FC33010FF95E99AF93EFBA26C16" box="[1000,1128,1782,1809]" pageId="5" pageNumber="253" value="2011-05-12">05.12.2011</date>
</collectingDate>
,
<collectingMunicipality id="6BCDD3793010FF95EE03F93EFB236C17" box="[1137,1257,1782,1808]" pageId="5" pageNumber="253">Rift Valley</collectingMunicipality>
,
<location id="8EC91FD83010FF95EE83F93EFA526C17" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83010FF95EE83F93EFA526C17" box="[1265,1432,1782,1808]" municipality="Rift Valley" name="Lake Naivasha" pageId="5" pageNumber="253" stateProvince="Baringo">Lake Naivasha</location>
,
<collectorName id="26E32CD53010FF95EAE5F8D2FEF16C33" box="[151,315,1818,1844]" pageId="5" pageNumber="253">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E3010FF95EB0BF8D2FC2A6C33" box="[377,992,1818,1845]" collectingDate="2005-11-27" collectorName="D. J. L. Agassiz" location="Gilgil" municipality="Rift Valley" pageId="5" pageNumber="253" specimenCount="1" specimenCount-male="1" stateProvince="Baringo">
<specimenCount id="9D10828A3010FF95EB0BF8D2FE546C33" box="[377,414,1818,1844]" count="1" pageId="5" pageNumber="253" type="male">1♂</specimenCount>
,
<date id="FFA86FC33010FF95EBD8F8D2FDE06C32" box="[426,554,1818,1845]" pageId="5" pageNumber="253" value="2005-11-27">
<collectingDate id="EFEC962B3010FF95EBD8F8D2FDE06C32" box="[426,554,1818,1845]" pageId="5" pageNumber="253" value="2005-11-27">27.11.2005</collectingDate>
</date>
,
<collectingMunicipality id="6BCDD3793010FF95E847F8D2FD7A6C33" box="[565,688,1818,1844]" pageId="5" pageNumber="253">Rift Valley</collectingMunicipality>
,
<location id="8EC91FD83010FF95E8C9F8D2FD356C33" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83010FF95E8C9F8D2FD356C33" box="[699,767,1818,1844]" municipality="Rift Valley" name="Gilgil" pageId="5" pageNumber="253" stateProvince="Baringo">Gilgil</location>
,
<collectorName id="26E32CD53010FF95E978F8D2FC646C33" box="[778,942,1818,1844]" pageId="5" pageNumber="253">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E3010FF95E99EF8D2FE006C5F" collectingDate="2013-10-24" collectorName="Lodge, D. J. L. &amp; Agassiz" location="Castle Forest" municipality="Central" pageId="5" pageNumber="253" specimenCount="1" specimenCount-female="1" stateProvince="Baringo">
<specimenCount id="9D10828A3010FF95E99EF8D2FBC56C33" box="[1004,1039,1818,1844]" count="1" pageId="5" pageNumber="253" type="female">1♀</specimenCount>
,
<date id="FFA86FC33010FF95EE6FF8D2FB546C33" box="[1053,1182,1818,1844]" pageId="5" pageNumber="253" value="2013-10-24">
<collectingDate id="EFEC962B3010FF95EE6FF8D2FB546C33" box="[1053,1182,1818,1844]" pageId="5" pageNumber="253" value="2013-10-24">24.10.2013</collectingDate>
</date>
,
<collectingMunicipality id="6BCDD3793010FF95EEDBF8D2FB346C33" box="[1193,1278,1818,1844]" pageId="5" pageNumber="253">Central</collectingMunicipality>
,
<location id="8EC91FD83010FF95EF7BF8D2FA566C33" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83010FF95EF7BF8D2FA566C33" box="[1289,1436,1818,1844]" municipality="Central" name="Castle Forest" pageId="5" pageNumber="253" stateProvince="Baringo">Castle Forest</location>
<collectorName id="26E32CD53010FF95EAE5F8F6FE5F6C5F" box="[151,405,1854,1880]" pageId="5" pageNumber="253">Lodge, D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E3010FF95EBAAF8F6FBBF6C5F" box="[472,1141,1854,1880]" collectingDate="2016-10-27" collectorName="Lodge, D. J. L. &amp; Agassiz" location="Castle Forest" pageId="5" pageNumber="253" specimenCount="1" specimenCount-male="1" stateProvince="Baringo">
<specimenCount id="9D10828A3010FF95EBAAF8F6FDCD6C5F" box="[472,519,1854,1880]" count="1" pageId="5" pageNumber="253" type="male">1 ♂</specimenCount>
,
<date id="FFA86FC33010FF95E867F8F6FD5C6C5F" box="[533,662,1854,1880]" pageId="5" pageNumber="253" value="2016-10-27">
<collectingDate id="EFEC962B3010FF95E867F8F6FD5C6C5F" box="[533,662,1854,1880]" pageId="5" pageNumber="253" value="2016-10-27">27.10.2016</collectingDate>
</date>
,
<location id="8EC91FD83010FF95E8D1F8F6FCF36C5F" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83010FF95E8D1F8F6FCF36C5F" box="[675,825,1854,1880]" name="Castle Forest" pageId="5" pageNumber="253" stateProvince="Baringo">Castle Forest</location>
<collectorName id="26E32CD53010FF95E930F8F6FB8A6C5F" box="[834,1088,1854,1880]" pageId="5" pageNumber="253">Lodge, D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E3010FF95EEF1F8F6FE3D6C7B" collectingDate="2011-11-30" collectorName="D. J. L. Agassiz" location="Naro Moru" municipality="Central" pageId="5" pageNumber="253" specimenCount="1" specimenCount-male="1" stateProvince="Baringo">
<specimenCount id="9D10828A3010FF95EEF1F8F6FB626C5F" box="[1155,1192,1854,1880]" count="1" pageId="5" pageNumber="253" type="male">1♂</specimenCount>
,
<date id="FFA86FC33010FF95EEC4F8F6FAFF6C5F" box="[1206,1333,1854,1880]" pageId="5" pageNumber="253" value="2011-11-30">
<collectingDate id="EFEC962B3010FF95EEC4F8F6FAFF6C5F" box="[1206,1333,1854,1880]" pageId="5" pageNumber="253" value="2011-11-30">30.11.2011</collectingDate>
</date>
,
<collectingMunicipality id="6BCDD3793010FF95EF31F8F6FA526C5F" box="[1347,1432,1854,1880]" pageId="5" pageNumber="253">Central</collectingMunicipality>
,
<location id="8EC91FD83010FF95EAE5F8AAFEDD6C7B" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83010FF95EAE5F8AAFEDD6C7B" box="[151,279,1890,1916]" municipality="Central" name="Naro Moru" pageId="5" pageNumber="253" stateProvince="Baringo">Naro Moru</location>
,
<collectorName id="26E32CD53010FF95EB53F8AAFE0E6C7B" box="[289,452,1890,1916]" pageId="5" pageNumber="253">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
;
<specimenCount id="9D10828A3010FF95E870F8AAFDEF6C7B" box="[514,549,1890,1916]" count="1" pageId="5" pageNumber="253" type="female">1♀</specimenCount>
, female,
<date id="FFA86FC33010FF95E8FEF8AAFCC76C7B" box="[652,781,1890,1916]" pageId="5" pageNumber="253" value="1999-10-10">10.10.1999</date>
,
<materialsCitation id="3B7E435E3010FF95E965F8AAFF096CA7" collectingDate="2000-10-05" collectorName="D. J. L. Agassiz" elevation="2134" location="Western" pageId="5" pageNumber="253" specimenCount="1" specimenCount-female="1" stateProvince="Kericho">
<specimenCount id="9D10828A3010FF95E965F8AAFCF06C7B" box="[791,826,1890,1916]" count="1" pageId="5" pageNumber="253" type="female">1♀</specimenCount>
,
<date id="FFA86FC33010FF95E93AF8AAFC036C7A" box="[840,969,1890,1917]" pageId="5" pageNumber="253" value="2000-10-05">
<collectingDate id="EFEC962B3010FF95E93AF8AAFC036C7A" box="[840,969,1890,1917]" pageId="5" pageNumber="253" value="2000-10-05">10.05.2000</collectingDate>
</date>
,
<location id="8EC91FD83010FF95E9A1F8AAFBFB6C7B" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83010FF95E9A1F8AAFBFB6C7B" box="[979,1073,1890,1916]" name="Western" pageId="5" pageNumber="253" stateProvince="Kericho">Western</location>
,
<collectingRegion id="49D287E13010FF95EE49F8AAFB5F6C7B" box="[1083,1173,1890,1916]" country="Kenya" name="Kericho" pageId="5" pageNumber="253">Kericho</collectingRegion>
<quantity id="4CEEE4E63010FF95EEEEF8AAFB256C7B" box="[1180,1263,1890,1916]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="5" pageNumber="253" unit="ft" value="7000.0">
<elevation id="003BAE303010FF95EEEEF8AAFB256C7B" box="[1180,1263,1890,1916]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="5" pageNumber="253" unit="ft" value="7000.0">7000ft.</elevation>
</quantity>
,
<collectorName id="26E32CD53010FF95EE8BF8AAFA576C7B" box="[1273,1437,1890,1916]" pageId="5" pageNumber="253">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E3010FF95EABDF84EFCC56CA7" box="[207,783,1926,1952]" collectingDate="2013-01-11" collectorName="D. J. L. Agassiz" location="Lewa" municipality="Eastern" pageId="5" pageNumber="253" specimenCount="1" specimenCount-female="1" stateProvince="Kericho">
<specimenCount id="9D10828A3010FF95EABDF84EFF386CA7" box="[207,242,1926,1952]" count="1" pageId="5" pageNumber="253" type="female">1♀</specimenCount>
,
<date id="FFA86FC33010FF95EB72F84EFE4A6CA7" box="[256,384,1926,1952]" pageId="5" pageNumber="253" value="2013-01-11">
<collectingDate id="EFEC962B3010FF95EB72F84EFE4A6CA7" box="[256,384,1926,1952]" pageId="5" pageNumber="253" value="2013-01-11">01.11.2013</collectingDate>
</date>
,
<collectingMunicipality id="6BCDD3793010FF95EBF9F84EFE286CA7" box="[395,482,1926,1952]" pageId="5" pageNumber="253">Eastern</collectingMunicipality>
,
<location id="8EC91FD83010FF95EB9FF84EFDE46CA7" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD83010FF95EB9FF84EFDE46CA7" box="[493,558,1926,1952]" municipality="Eastern" name="Lewa" pageId="5" pageNumber="253" stateProvince="Kericho">Lewa</location>
,
<collectorName id="26E32CD53010FF95E84BF84EFD176CA7" box="[569,733,1926,1952]" pageId="5" pageNumber="253">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
,
</paragraph>
<caption id="DF69198B3013FF96EAEAFC31FEDE6F32" ID-DOI="http://doi.org/10.5281/zenodo.14033550" ID-Zenodo-Dep="14033550" httpUri="https://zenodo.org/record/14033550/files/figure.png" pageId="6" pageNumber="254" startId="6.[152,255,1017,1041]" targetBox="[193,1394,181,993]" targetPageId="6" targetType="figure">
<paragraph id="8BA949033013FF96EAEAFC31FEDE6F32" blockId="6.[151,1437,1017,1077]" pageId="6" pageNumber="254">
<emphasis id="B96295113013FF96EAEAFC31FED06F16" bold="true" box="[152,282,1017,1041]" pageId="6" pageNumber="254">FIGURE 3.</emphasis>
<taxonomicName id="4C1632803013FF96EB50FC31FD936F16" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[290,601,1017,1041]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="254" phylum="Arthropoda" rank="species" species="stichocentra">
<emphasis id="B96295113013FF96EB50FC31FD936F16" box="[290,601,1017,1041]" italics="true" pageId="6" pageNumber="254">Pararhigognostis stichocentra</emphasis>
</taxonomicName>
adult male: labial palpus, Kenya, Rift Valley, Turi, 06.VII.1999, 8000 ft., D.J.L. Agassiz leg.
</paragraph>
</caption>
<caption id="DF69198B3013FF96EAEAF806FBD86CE1" ID-DOI="http://doi.org/10.5281/zenodo.14033552" ID-Zenodo-Dep="14033552" box="[152,1042,1998,2022]" httpUri="https://zenodo.org/record/14033552/files/figure.png" pageId="6" pageNumber="254" startId="6.[152,255,1998,2022]" targetBox="[195,1392,1142,1971]" targetPageId="6" targetType="figure">
<paragraph id="8BA949033013FF96EAEAF806FBD86CE1" blockId="6.[152,1042,1998,2022]" box="[152,1042,1998,2022]" pageId="6" pageNumber="254">
<emphasis id="B96295113013FF96EAEAF806FED26CE1" bold="true" box="[152,280,1998,2022]" pageId="6" pageNumber="254">FIGURE 4.</emphasis>
<taxonomicName id="4C1632803013FF96EB6DF806FD996CE1" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[287,595,1998,2022]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="254" phylum="Arthropoda" rank="species" species="stichocentra">
<emphasis id="B96295113013FF96EB6DF806FD996CE1" box="[287,595,1998,2022]" italics="true" pageId="6" pageNumber="254">Pararhigognostis stichocentra</emphasis>
</taxonomicName>
, wing venations: A, forewing, B, hindwing.
</paragraph>
</caption>
<caption id="DF69198B3012FF97EAEAF85FFAB16CA8" ID-DOI="http://doi.org/10.5281/zenodo.14033554" ID-Zenodo-Dep="14033554" box="[152,1403,1943,1968]" httpUri="https://zenodo.org/record/14033554/files/figure.png" pageId="7" pageNumber="255" startId="7.[152,255,1943,1967]" targetBox="[232,1355,187,1913]" targetPageId="7" targetType="figure">
<paragraph id="8BA949033012FF97EAEAF85FFAB16CA8" blockId="7.[152,1403,1943,1968]" box="[152,1403,1943,1968]" pageId="7" pageNumber="255">
<emphasis id="B96295113012FF97EAEAF85FFED26CB7" bold="true" box="[152,280,1943,1968]" pageId="7" pageNumber="255">FIGURE 5.</emphasis>
<taxonomicName id="4C1632803012FF97EB6DF85FFD996CA8" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[287,595,1943,1967]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="255" phylum="Arthropoda" rank="species" species="stichocentra">
<emphasis id="B96295113012FF97EB6DF85FFD996CA8" box="[287,595,1943,1967]" italics="true" pageId="7" pageNumber="255">Pararhigognostis stichocentra</emphasis>
</taxonomicName>
male genitalia, gen. slide YPO 19/2016: A, general view, B, aedeagus, C, socii.
</paragraph>
</caption>
<caption id="DF69198B301DFF98EAEAF85FFDF86CD4" ID-DOI="http://doi.org/10.5281/zenodo.14033556" ID-Zenodo-Dep="14033556" httpUri="https://zenodo.org/record/14033556/files/figure.png" pageId="8" pageNumber="256" startId="8.[152,255,1943,1967]" targetBox="[252,1335,187,1914]" targetPageId="8" targetType="figure">
<paragraph id="8BA94903301DFF98EAEAF85FFDF86CD4" blockId="8.[151,1436,1943,2003]" pageId="8" pageNumber="256">
<emphasis id="B9629511301DFF98EAEAF85FFED36CA8" bold="true" box="[152,281,1943,1967]" pageId="8" pageNumber="256">FIGURE 6.</emphasis>
<taxonomicName id="4C163280301DFF98EB6DF85FFD9E6CA8" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[287,596,1943,1967]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="256" phylum="Arthropoda" rank="species" species="stichocentra">
<emphasis id="B9629511301DFF98EB6DF85FFD9E6CA8" box="[287,596,1943,1967]" italics="true" pageId="8" pageNumber="256">Pararhigognostis stichocentra</emphasis>
</taxonomicName>
male genitalia, gen. slide YPO 19/2016: A, separated valva, B, tip of aedeagus, C, aedeagus base with ductus ejaculatorius.
</paragraph>
</caption>
<caption id="DF69198B301CFF99EAEAF85FFAED6CA8" ID-DOI="http://doi.org/10.5281/zenodo.14033558" ID-Zenodo-Dep="14033558" box="[152,1319,1943,1968]" httpUri="https://zenodo.org/record/14033558/files/figure.png" pageId="9" pageNumber="257" startId="9.[152,255,1943,1967]" targetBox="[235,1353,187,1913]" targetPageId="9" targetType="figure">
<paragraph id="8BA94903301CFF99EAEAF85FFAED6CA8" blockId="9.[152,1319,1943,1968]" box="[152,1319,1943,1968]" pageId="9" pageNumber="257">
<emphasis id="B9629511301CFF99EAEAF85FFED26CA8" bold="true" box="[152,280,1943,1967]" pageId="9" pageNumber="257">FIGURE 7.</emphasis>
<taxonomicName id="4C163280301CFF99EB6DF85FFD996CA8" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[287,595,1943,1967]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="257" phylum="Arthropoda" rank="species" species="stichocentra">
<emphasis id="B9629511301CFF99EB6DF85FFD996CA8" box="[287,595,1943,1967]" italics="true" pageId="9" pageNumber="257">Pararhigognostis stichocentra</emphasis>
</taxonomicName>
female genitalia, gen. slide: YPO 20/2016: A, general view, B, antrum.
</paragraph>
</caption>
<paragraph id="8BA94903301FFF9AEAB5FF5FFE036BFE" blockId="10.[151,1437,151,825]" pageId="10" pageNumber="258">
<materialsCitation id="3B7E435E301FFF9AEAB5FF5FFB836BB6" box="[199,1097,151,177]" collectingDate="1998-04-12" collectionCode="EB" collectorName="D. J. L. Agassiz" country="Kenya" elevation="2438" location="Prov. Turi" municipality="Rift Valley" pageId="10" pageNumber="258" specimenCount="1" specimenCount-female="1">
<collectionCode id="ED07D1C6301FFF9AEAB5FF5FFF246BB6" box="[199,238,151,177]" pageId="10" pageNumber="258">EB</collectionCode>
;
<collectingCountry id="F3010993301FFF9AEA89FF5FFE826BB6" box="[251,328,151,177]" name="Kenya" pageId="10" pageNumber="258">Kenya</collectingCountry>
:
<specimenCount id="9D10828A301FFF9AEB21FF5FFEBC6BB6" box="[339,374,151,177]" count="1" pageId="10" pageNumber="258" type="female">1♀</specimenCount>
,
<date id="FFA86FC3301FFF9AEBF7FF5FFDCC6BB6" box="[389,518,151,177]" pageId="10" pageNumber="258" value="1998-04-12">
<collectingDate id="EFEC962B301FFF9AEBF7FF5FFDCC6BB6" box="[389,518,151,177]" pageId="10" pageNumber="258" value="1998-04-12">04.12.1998</collectingDate>
</date>
,
<collectingMunicipality id="6BCDD379301FFF9AE860FF5FFD446BB6" box="[530,654,151,177]" pageId="10" pageNumber="258">Rift Valley</collectingMunicipality>
,
<location id="8EC91FD8301FFF9AE8E8FF5FFCC76BB6" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD8301FFF9AE8E8FF5FFCC76BB6" box="[666,781,151,177]" country="Kenya" municipality="Rift Valley" name="Prov. Turi" pageId="10" pageNumber="258">Prov. Turi</location>
,
<quantity id="4CEEE4E6301FFF9AE96BFF5FFCAF6BB6" box="[793,869,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="10" pageNumber="258" unit="ft" value="8000.0">
<elevation id="003BAE30301FFF9AE96BFF5FFCAF6BB6" box="[793,869,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="10" pageNumber="258" unit="ft" value="8000.0">8000ft</elevation>
</quantity>
,
<collectorName id="26E32CD5301FFF9AE903FF5FFBDC6BB6" box="[881,1046,151,177]" pageId="10" pageNumber="258">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E301FFF9AEE24FF5FFEE86BD2" collectingDate="1998-10-16" collectionCode="EB" collectorName="D. J. L. Agassiz" country="Kenya" elevation="2438" location="Prov. Turi" municipality="Rift Valley" pageId="10" pageNumber="258" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D10828A301FFF9AEE24FF5FFBB36BB6" box="[1110,1145,151,177]" count="1" pageId="10" pageNumber="258" type="female">1♀</specimenCount>
,
<date id="FFA86FC3301FFF9AEEFAFF5FFAC36BB6" box="[1160,1289,151,177]" pageId="10" pageNumber="258" value="1998-10-16">
<collectingDate id="EFEC962B301FFF9AEEFAFF5FFAC36BB6" box="[1160,1289,151,177]" pageId="10" pageNumber="258" value="1998-10-16">16.10.1998</collectingDate>
</date>
, ditto,
<collectorName id="26E32CD5301FFF9AEF25FF5FFF3A6BD2" pageId="10" pageNumber="258">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
,
<materialsCitation id="3B7E435E301FFF9AEB5FFF73FC2C6BD2" box="[301,998,187,213]" collectingDate="1999-06-07" collectionCode="EB" collectorName="D. J. L. Agassiz" country="Kenya" elevation="2438" location="Turi" municipality="Rift Valley" pageId="10" pageNumber="258" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D10828A301FFF9AEB5FFF73FE986BD2" box="[301,338,187,213]" count="1" pageId="10" pageNumber="258" type="male">1♂</specimenCount>
,
<date id="FFA86FC3301FFF9AEB2CFF73FE156BD2" box="[350,479,187,213]" pageId="10" pageNumber="258" value="1999-06-07">
<collectingDate id="EFEC962B301FFF9AEB2CFF73FE156BD2" box="[350,479,187,213]" pageId="10" pageNumber="258" value="1999-06-07">06.07.1999</collectingDate>
</date>
,
<collectingMunicipality id="6BCDD379301FFF9AEB98FF73FDAF6BD2" box="[490,613,187,213]" pageId="10" pageNumber="258">Rift Valley</collectingMunicipality>
,
<location id="8EC91FD8301FFF9AE81CFF73FD6A6BD2" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD8301FFF9AE81CFF73FD6A6BD2" box="[622,672,187,213]" country="Kenya" municipality="Rift Valley" name="Turi" pageId="10" pageNumber="258">Turi</location>
,
<quantity id="4CEEE4E6301FFF9AE8D9FF73FCCE6BD2" box="[683,772,187,213]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="10" pageNumber="258" unit="ft" value="8000.0">
<elevation id="003BAE30301FFF9AE8D9FF73FCCE6BD2" box="[683,772,187,213]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="10" pageNumber="258" unit="ft" value="8000.0">8000 ft.</elevation>
</quantity>
,
<collectorName id="26E32CD5301FFF9AE962FF73FC796BD2" box="[784,947,187,213]" pageId="10" pageNumber="258">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E301FFF9AE983FF73FE036BFE" collectingDate="1998-10-31" collectionCode="EB" collectorName="D. J. L. Agassiz" country="Kenya" elevation="2134" location="Western Pr." pageId="10" pageNumber="258" specimenCount="1" specimenCount-male="1" stateProvince="Kericho">
<specimenCount id="9D10828A301FFF9AE983FF73FBDC6BD2" box="[1009,1046,187,213]" count="1" pageId="10" pageNumber="258" type="male">1♂</specimenCount>
,
<date id="FFA86FC3301FFF9AEE50FF73FB696BD2" box="[1058,1187,187,213]" pageId="10" pageNumber="258" value="1998-10-31">
<collectingDate id="EFEC962B301FFF9AEE50FF73FB696BD2" box="[1058,1187,187,213]" pageId="10" pageNumber="258" value="1998-10-31">31.10.1998</collectingDate>
</date>
,
<location id="8EC91FD8301FFF9AEEDFFF73FAFA6BD2" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD8301FFF9AEEDFFF73FAFA6BD2" box="[1197,1328,187,213]" country="Kenya" name="Western Pr." pageId="10" pageNumber="258" stateProvince="Kericho">Western Pr.</location>
,
<collectingRegion id="49D287E1301FFF9AEF49FF73FA526BD2" box="[1339,1432,187,213]" country="Kenya" name="Kericho" pageId="10" pageNumber="258">Kericho</collectingRegion>
,
<quantity id="4CEEE4E6301FFF9AEAE5FF17FF216BFE" box="[151,235,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="10" pageNumber="258" unit="ft" value="7000.0">
<elevation id="003BAE30301FFF9AEAE5FF17FF216BFE" box="[151,235,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="10" pageNumber="258" unit="ft" value="7000.0">7000ft.</elevation>
</quantity>
,
<collectorName id="26E32CD5301FFF9AEA87FF17FE536BFE" box="[245,409,223,249]" pageId="10" pageNumber="258">D.J.L. Agassiz</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BA94903301FFF9AEAB5FECBFB816A62" blockId="10.[151,1437,151,825]" pageId="10" pageNumber="258">
<materialsCitation id="3B7E435E301FFF9AEAB5FECBFF096A46" collectingDate="2008-11-24" collectionCode="NHMO" collectorName="L. Aarvik &amp; D. Agassiz &amp; A. Kingston" country="Kenya" elevation="2110" location="Rift Valley Province" municipality="Gilgil" pageId="10" pageNumber="258" specimenCount="1" specimenCount-female="1">
<collectionCode id="ED07D1C6301FFF9AEAB5FECBFEEA6A1A" box="[199,288,259,285]" country="Norway" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="10" pageNumber="258" type="Museum">NHMO</collectionCode>
;
<collectingCountry id="F3010993301FFF9AEB58FECBFEBD6A1A" box="[298,375,259,285]" name="Kenya" pageId="10" pageNumber="258">Kenya</collectingCountry>
:
<specimenCount id="9D10828A301FFF9AEBF3FECBFE6E6A1A" box="[385,420,259,285]" count="1" pageId="10" pageNumber="258" type="female">1♀</specimenCount>
, 22
<date id="FFA86FC3301FFF9AEBA8FECBFD906A1A" box="[474,602,259,285]" pageId="10" pageNumber="258" value="2008-11-24">
<collectingDate id="EFEC962B301FFF9AEBA8FECBFD906A1A" box="[474,602,259,285]" pageId="10" pageNumber="258" value="2008-11-24">24.11.2008</collectingDate>
</date>
,
<location id="8EC91FD8301FFF9AE816FECBFC806A1A" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD8301FFF9AE816FECBFC806A1A" box="[612,842,259,285]" country="Kenya" municipality="Gilgil" name="Rift Valley Province" pageId="10" pageNumber="258">Rift Valley Province</location>
:
<collectingMunicipality id="6BCDD379301FFF9AE921FECBFC5D6A1A" box="[851,919,259,285]" pageId="10" pageNumber="258">Gilgil</collectingMunicipality>
,
<quantity id="4CEEE4E6301FFF9AE9D3FECBFC326A1A" box="[929,1016,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.11" pageId="10" pageNumber="258" unit="m" value="2110.0">
<elevation id="003BAE30301FFF9AE9D3FECBFC326A1A" box="[929,1016,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.11" pageId="10" pageNumber="258" unit="m" value="2110.0">2110 m</elevation>
</quantity>
.,
<collectorName id="26E32CD5301FFF9AEE75FECBFBBE6A1A" box="[1031,1140,259,285]" pageId="10" pageNumber="258">L. Aarvik</collectorName>
,
<collectorName id="26E32CD5301FFF9AEE0FFECBFB3C6A1A" box="[1149,1270,259,285]" pageId="10" pageNumber="258">D. Agassiz</collectorName>
&amp;
<collectorName id="26E32CD5301FFF9AEF67FECBFA576A1A" box="[1301,1437,259,285]" pageId="10" pageNumber="258">A. Kingston</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E301FFF9AEAA3FEEFFCB86A46" box="[209,882,295,321]" collectingDate="2010-11-27" collectionCode="NHMO" collectorName="L. Aarvik &amp; T. Gilligan" country="Kenya" elevation="2110" location="Rift Valley Province" municipality="Gilgil" pageId="10" pageNumber="258" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D10828A301FFF9AEAA3FEEFFF3E6A46" box="[209,244,295,321]" count="1" pageId="10" pageNumber="258" type="male">1♂</specimenCount>
and
<specimenCount id="9D10828A301FFF9AEB5FFEEFFE9A6A46" box="[301,336,295,321]" count="1" pageId="10" pageNumber="258" type="female">1♀</specimenCount>
,
<date id="FFA86FC3301FFF9AEB12FEEFFE2A6A46" box="[352,480,295,321]" pageId="10" pageNumber="258" value="2010-11-27">
<collectingDate id="EFEC962B301FFF9AEB12FEEFFE2A6A46" box="[352,480,295,321]" pageId="10" pageNumber="258" value="2010-11-27">27.11.2010</collectingDate>
</date>
, ditto,
<collectorName id="26E32CD5301FFF9AE85DFEEFFD566A46" box="[559,668,295,321]" pageId="10" pageNumber="258">L. Aarvik</collectorName>
&amp;
<collectorName id="26E32CD5301FFF9AE8B0FEEFFCF76A46" box="[706,829,295,321]" pageId="10" pageNumber="258">T. Gilligan</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E301FFF9AE90DFEEFFB816A62" collectingDate="2006-11-21" collectionCode="NHMO" collectorName="L. O. Hansen &amp; K. Sund" country="Kenya" county="Rongai" elevation="2206" location="Rift Valley Province" municipality="Chorlim Gate" pageId="10" pageNumber="258" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D10828A301FFF9AE90DFEEFFC726A46" box="[895,952,295,321]" count="2" pageId="10" pageNumber="258" type="female">2♀♀</specimenCount>
, 17
<date id="FFA86FC3301FFF9AE99CFEEFFBBB6A46" box="[1006,1137,295,321]" pageId="10" pageNumber="258" value="2006-11-21">
<collectingDate id="EFEC962B301FFF9AE99CFEEFFBBB6A46" box="[1006,1137,295,321]" pageId="10" pageNumber="258" value="2006-11-21">21.11.2006</collectingDate>
</date>
,
<location id="8EC91FD8301FFF9AEE0FFEEFFAA36A46" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD8301FFF9AEE0FFEEFFAA36A46" box="[1149,1385,295,321]" country="Kenya" county="Rongai" municipality="Chorlim Gate" name="Rift Valley Province" pageId="10" pageNumber="258">Rift Valley Province</location>
:
<location id="8EC91FD8301FFF9AEF06FEEFFE846A62" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD8301FFF9AEF06FEEFFE846A62" country="Kenya" county="Rongai" municipality="Chorlim Gate" name="Mt. Elgon Nat. Park" pageId="10" pageNumber="258">Mt. Elgon Nat. Park</location>
,
<collectingMunicipality id="6BCDD379301FFF9AEB2BFE83FE3F6A62" box="[345,501,331,357]" pageId="10" pageNumber="258">Chorlim Gate</collectingMunicipality>
,
<collectingCounty id="62C8318F301FFF9AE872FE83FD9B6A62" box="[512,593,331,357]" pageId="10" pageNumber="258">Rongai</collectingCounty>
camp,
<quantity id="4CEEE4E6301FFF9AE8D1FE83FD376A62" box="[675,765,331,357]" metricMagnitude="3" metricUnit="m" metricValue="2.206" pageId="10" pageNumber="258" unit="m" value="2206.0">
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</quantity>
.,
<collectorName id="26E32CD5301FFF9AE97EFE83FC516A62" box="[780,923,331,357]" pageId="10" pageNumber="258">L.O. Hansen</collectorName>
&amp;
<collectorName id="26E32CD5301FFF9AE9CCFE83FBD16A62" box="[958,1051,331,357]" pageId="10" pageNumber="258">K. Sund</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BA94903301FFF9AEAB5FEA7FBFD6AAA" blockId="10.[151,1437,151,825]" pageId="10" pageNumber="258">
<materialsCitation id="3B7E435E301FFF9AEAB5FEA7FF096AAA" collectingDate="2007-11-11" collectorName="L. Aarvik &amp; M. Fibiger" country="Uganda" elevation="1700" location="Nyakalengija" municipality="Ruwenzori Mts." pageId="10" pageNumber="258" specimenCount="1" specimenCount-female="1" stateProvince="Kabarole">
<collectingCountry id="F3010993301FFF9AEAB5FEA7FEE96A8E" box="[199,291,367,393]" name="Uganda" pageId="10" pageNumber="258">Uganda</collectingCountry>
:
<specimenCount id="9D10828A301FFF9AEB5FFEA7FE9A6A8E" box="[301,336,367,393]" count="1" pageId="10" pageNumber="258" type="female">1♀</specimenCount>
, 10
<date id="FFA86FC3301FFF9AEBF4FEA7FDCC6A8E" box="[390,518,367,393]" pageId="10" pageNumber="258" value="2007-11-11">
<collectingDate id="EFEC962B301FFF9AEBF4FEA7FDCC6A8E" box="[390,518,367,393]" pageId="10" pageNumber="258" value="2007-11-11">11.11.2007</collectingDate>
</date>
,
<collectingRegion id="49D287E1301FFF9AE863FEA7FD0A6A8E" box="[529,704,367,393]" country="Uganda" name="Kabarole" pageId="10" pageNumber="258">Kabarole Distr.</collectingRegion>
:
<collectingMunicipality id="6BCDD379301FFF9AE8B8FEA7FC4A6A8E" box="[714,896,367,393]" pageId="10" pageNumber="258">Ruwenzori Mts.</collectingMunicipality>
,
<location id="8EC91FD8301FFF9AE9F9FEA7FBE96A8E" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD8301FFF9AE9F9FEA7FBE96A8E" box="[907,1059,367,393]" country="Uganda" municipality="Ruwenzori Mts." name="Nyakalengija" pageId="10" pageNumber="258" stateProvince="Kabarole">Nyakalengija</location>
,
<quantity id="4CEEE4E6301FFF9AEE5CFEA7FB426A8E" box="[1070,1160,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="258" unit="m" value="1700.0">
<elevation id="003BAE30301FFF9AEE5CFEA7FB426A8E" box="[1070,1160,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="258" unit="m" value="1700.0">1700 m</elevation>
</quantity>
.,
<collectorName id="26E32CD5301FFF9AEEE5FEA7FAC86A8E" box="[1175,1282,367,393]" pageId="10" pageNumber="258">L. Aarvik</collectorName>
&amp;
<collectorName id="26E32CD5301FFF9AEF57FEA7FA566A8E" box="[1317,1436,367,393]" pageId="10" pageNumber="258">M. Fibiger</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7E435E301FFF9AEABDFE5BFBFD6AAA" box="[207,1079,403,429]" collectingDate="2007-07-11" collectorName="L. Aarvik &amp; M. Fibiger" country="Uganda" elevation="2330" location="Ruhija" pageId="10" pageNumber="258" specimenCount="1" specimenCount-female="1" stateProvince="Kabale">
<specimenCount id="9D10828A301FFF9AEABDFE5BFF386AAA" box="[207,242,403,429]" count="1" pageId="10" pageNumber="258" type="female">1♀</specimenCount>
, 4
<date id="FFA86FC3301FFF9AEB6EFE5BFE456AAA" box="[284,399,403,429]" pageId="10" pageNumber="258" value="2007-07-11">
<collectingDate id="EFEC962B301FFF9AEB6EFE5BFE456AAA" box="[284,399,403,429]" pageId="10" pageNumber="258" value="2007-07-11">7.11.2007</collectingDate>
</date>
,
<collectingRegion id="49D287E1301FFF9AEBE9FE5BFDF96AAA" box="[411,563,403,429]" country="Uganda" name="Kabale" pageId="10" pageNumber="258">Kabale Distr.</collectingRegion>
:
<location id="8EC91FD8301FFF9AE84FFE5BFD416AAA" LSID="urn:lsid:plazi:treatment:03BFF8153011FF9AEAE5FC6EFC5A683E:8EC91FD8301FFF9AE84FFE5BFD416AAA" box="[573,651,403,429]" country="Uganda" name="Ruhija" pageId="10" pageNumber="258" stateProvince="Kabale">Ruhija</location>
,
<quantity id="4CEEE4E6301FFF9AE8E4FE5BFD3A6AAA" box="[662,752,403,429]" metricMagnitude="3" metricUnit="m" metricValue="2.33" pageId="10" pageNumber="258" unit="m" value="2330.0">
<elevation id="003BAE30301FFF9AE8E4FE5BFD3A6AAA" box="[662,752,403,429]" metricMagnitude="3" metricUnit="m" metricValue="2.33" pageId="10" pageNumber="258" unit="m" value="2330.0">2330 m</elevation>
</quantity>
.,
<collectorName id="26E32CD5301FFF9AE972FE5BFCA16AAA" box="[768,875,403,429]" pageId="10" pageNumber="258">L. Aarvik</collectorName>
&amp;
<collectorName id="26E32CD5301FFF9AE9FDFE5BFBCD6AAA" box="[911,1031,403,429]" pageId="10" pageNumber="258">M. Fibiger</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BA94903301FFF9AEAB5FE7FFC456AF2" blockId="10.[151,1437,151,825]" pageId="10" pageNumber="258">
MHB;
<materialsCitation id="3B7E435E301FFF9AEB6EFE7FFC456AF2" collectingDate="2012-06-11" collectingDateMax="2013-11-26" collectingDateMin="2012-06-11" collectorName="Graaff-Reinet &amp; Asante Sana Game Farm, W &amp; W. Mey" country="South Africa" location="Eastern Cape" pageId="10" pageNumber="258" specimenCount="7" specimenCount-male="7" stateProvince="Eastern Cape">
<collectingCountry id="F3010993301FFF9AEB6EFE7FFE7F6AD6" box="[284,437,439,465]" name="South Africa" pageId="10" pageNumber="258">South Africa</collectingCountry>
:
<specimenCount id="9D10828A301FFF9AEBB4FE7FFDCB6AD6" box="[454,513,439,465]" count="2" pageId="10" pageNumber="258" type="male">2♂♂</specimenCount>
, 05
<date id="FFA86FC3301FFF9AE84EFE7EFD746AD6" box="[572,702,438,465]" pageId="10" pageNumber="258" value="2012-06-11">
<collectingDate id="EFEC962B301FFF9AE84EFE7EFD746AD6" box="[572,702,438,465]" pageId="10" pageNumber="258" value="2012-06-11">06.11.2012</collectingDate>
</date>
,
<specimenCount id="9D10828A301FFF9AE8A2FE7FFCC06AD6" box="[720,778,439,465]" count="2" pageId="10" pageNumber="258" type="male">2♂♂</specimenCount>
, 08
<date id="FFA86FC3301FFF9AE934FE7FFC0D6AD6" box="[838,967,439,465]" pageId="10" pageNumber="258" value="2012-11-13">
<collectingDate id="EFEC962B301FFF9AE934FE7FFC0D6AD6" box="[838,967,439,465]" pageId="10" pageNumber="258" value="2012-11-13">13.11.2012</collectingDate>
</date>
,
<specimenCount id="9D10828A301FFF9AE9ABFE7FFBDE6AD6" box="[985,1044,439,465]" count="2" pageId="10" pageNumber="258" type="male">2♂♂</specimenCount>
,
<date id="FFA86FC3301FFF9AEE57FE7FFB6F6AD6" box="[1061,1189,439,465]" pageId="10" pageNumber="258" value="2012-11-14">
<collectingDate id="EFEC962B301FFF9AEE57FE7FFB6F6AD6" box="[1061,1189,439,465]" pageId="10" pageNumber="258" value="2012-11-14">14.11.2012</collectingDate>
</date>
,
<specimenCount id="9D10828A301FFF9AEEC5FE7FFB116AD6" box="[1207,1243,439,465]" count="1" pageId="10" pageNumber="258" type="male">1♂</specimenCount>
, 22
<date id="FFA86FC3301FFF9AEF6AFE7FFA526AD6" box="[1304,1432,439,465]" pageId="10" pageNumber="258" value="2013-11-26">
<collectingDate id="EFEC962B301FFF9AEF6AFE7FFA526AD6" box="[1304,1432,439,465]" pageId="10" pageNumber="258" value="2013-11-26">26.11.2013</collectingDate>
</date>
,
<collectingRegion id="49D287E1301FFF9AEAE5FE13FEE46AF2" box="[151,302,475,501]" country="South Africa" name="Eastern Cape" pageId="10" pageNumber="258">Eastern Cape</collectingRegion>
,
<collectorName id="26E32CD5301FFF9AEB48FE13FE126AF2" box="[314,472,475,501]" pageId="10" pageNumber="258">Graaff-Reinet</collectorName>
,
<collectorName id="26E32CD5301FFF9AEB93FE13FCE86AF2" box="[481,802,475,501]" pageId="10" pageNumber="258">Asante Sana Game Farm, W</collectorName>
. Mey leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30C1A88301FFF9AEAB5FE36FC5A683E" pageId="10" pageNumber="258" type="biology_ecology">
<paragraph id="8BA94903301FFF9AEAB5FE36FA8A691E" blockId="10.[151,1437,151,825]" box="[199,1344,510,537]" pageId="10" pageNumber="258">
<emphasis id="B9629511301FFF9AEAB5FE36FEED691F" bold="true" box="[199,295,510,536]" pageId="10" pageNumber="258">Biology.</emphasis>
Larvae in November, on
<taxonomicName id="4C163280301FFF9AE837FE37FC26691E" authority="Delile, 1843" authorityName="Delile" authorityYear="1843" box="[581,1004,511,537]" class="Magnoliopsida" family="Asteraceae" genus="Helichrysum" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="258" phylum="Tracheophyta" rank="species" species="citrispinum">
<emphasis id="B9629511301FFF9AE837FE37FC91691F" box="[581,859,511,537]" italics="true" pageId="10" pageNumber="258">Helichrysum citrispinum</emphasis>
Delile, 1843
</taxonomicName>
(
<taxonomicName id="4C163280301FFF9AE989FE37FBB6691E" box="[1019,1148,511,537]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="258" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
), (
<bibRefCitation id="EF8734F2301FFF9AEEE4FE37FAFD691E" author="Meyrick, E." box="[1174,1335,511,537]" pageId="10" pageNumber="258" pagination="107 - 120" refId="ref5877" refString="Meyrick, E. (1932) Entomological expedition to Abyssinia, 1926 - 27 Microlepidoptera. The Transaction of the Entomological Society of London, 80 (1), 107 - 120. https: // doi. org / 10.1111 / j. 1365 - 2311.1932. tb 00085. x" type="journal article" year="1932">Meyrick 1932</bibRefCitation>
)
</paragraph>
<paragraph id="8BA94903301FFF9AEAB5FDEAFE7769F6" blockId="10.[151,1437,151,825]" pageId="10" pageNumber="258">
<emphasis id="B9629511301FFF9AEAB5FDEAFE79693A" bold="true" box="[199,435,546,573]" pageId="10" pageNumber="258">Distributional area.</emphasis>
The species is known so far from
<collectingCountry id="F3010993301FFF9AE94CFDEBFC6A693A" box="[830,928,547,573]" name="Ethiopia" pageId="10" pageNumber="258">Ethiopia</collectingCountry>
,
<collectingCountry id="F3010993301FFF9AE9DEFDEBFC32693A" box="[940,1016,547,573]" name="Kenya" pageId="10" pageNumber="258">Kenya</collectingCountry>
,
<collectingCountry id="F3010993301FFF9AEE76FDEBFB94693A" box="[1028,1118,547,573]" name="Uganda" pageId="10" pageNumber="258">Uganda</collectingCountry>
, and
<collectingCountry id="F3010993301FFF9AEEE9FDEBFAE5693A" box="[1179,1327,547,573]" name="South Africa" pageId="10" pageNumber="258">South Africa</collectingCountry>
. The host plant of caterpillars,
<taxonomicName id="4C163280301FFF9AEBF3FD8FFD526967" authorityName="Delile" authorityYear="1843" box="[385,664,583,609]" class="Magnoliopsida" family="Asteraceae" genus="Helichrysum" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="258" phylum="Tracheophyta" rank="species" species="citrispinum">
<emphasis id="B9629511301FFF9AEBF3FD8FFD526967" box="[385,664,583,609]" italics="true" pageId="10" pageNumber="258">Helichrysum citrispinum</emphasis>
</taxonomicName>
, occurs at high altitudes in mountains (
<bibRefCitation id="EF8734F2301FFF9AEE27FD8FFA9A6966" author="Schweiger, A. H. &amp; Otieno, D. O. &amp; Kulunge, D. O. &amp; Reineking, B. &amp; Tenhunen, J." box="[1109,1360,582,609]" pageId="10" pageNumber="258" pagination="293 - 303" refId="ref6366" refString="Schweiger, A. H., Otieno, D. O., Kulunge, D. O., Reineking, B. &amp; Tenhunen, J. (2015) The Afro-alpine dwarf shrub Helichrysum citrispinum favours undersorey plants through microclimate amelioration. Plant Ecology and Diversity, 8 (3), 293 - 303. https: // doi. org / 10.1080 / 17550874.2015.1014207" type="journal article" year="2015">
Schweiger
<emphasis id="B9629511301FFF9AEEA6FD8FFAC76966" box="[1236,1293,583,609]" italics="true" pageId="10" pageNumber="258">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="EF8734F2301FFF9AEF2EFD8FFEF66982" author="Uhlig, S. K. &amp; Uhlig, K." pageId="10" pageNumber="258" pagination="153 - 156" refId="ref6648" refString="Uhlig, S. K. &amp; Uhlig, K. (1991) Studies on the Altitudinal Zonation of Forests and Alpine Plants in the Central Bale Mountains, Ethiopia. Mountain Research and Development, 11 (2), 153 - 156. https: // doi. org / 10.2307 / 3673574" type="journal article" year="1991">Uhlig &amp; Uhlig 1991</bibRefCitation>
;
<bibRefCitation id="EF8734F2301FFF9AEB38FDA3FE706982" author="Coe, M. J." box="[330,442,619,645]" pageId="10" pageNumber="258" refId="ref5431" refString="Coe, M. J. (1967) The Ecology of the Alpine Zone of Mount Kenya. Uitgeverij Dr. W. Junk, Den Haag, 114 pp. https: // doi. org / 10.1007 / 978 - 94 - 011 - 7831 - 0" type="book" year="1967">Coe 1967</bibRefCitation>
) with a range which includes also
<collectingCountry id="F3010993301FFF9AE921FDA3FC716982" box="[851,955,619,645]" name="Tanzania" pageId="10" pageNumber="258">Tanzania</collectingCountry>
,
<collectingCountry id="F3010993301FFF9AE9BAFDA3FBED6982" box="[968,1063,619,645]" name="Somalia" pageId="10" pageNumber="258">Somalia</collectingCountry>
, and
<collectingCountry id="F3010993301FFF9AEE15FDA3FB0D6982" box="[1127,1223,619,645]" name="Namibia" pageId="10" pageNumber="258">Namibia</collectingCountry>
(
<bibRefCitation id="EF8734F2301FFF9AEEABFDA3FF1B69AE" author="GBIF Secretariat" pageId="10" pageNumber="258" refId="ref5716" refString="GBIF Secretariat (2019) Helichrysum citrispinum. In: GBIF Backbone Taxonomy. Checklist dataset. Available from: https: // www. gbif. org / (accessed 4 March 2020) https: // doi. org / 10.15468 / 39 omei" type="book" year="2019">GBIF Secretariat 2019</bibRefCitation>
) and is more extensive than the currently confirmed distribution range of this lepidopteran. The moth was also recorded in
<collectingCountry id="F3010993301FFF9AEB6FFD7BFE6769CA" box="[285,429,691,717]" name="South Africa" pageId="10" pageNumber="258">South Africa</collectingCountry>
, where
<taxonomicName id="4C163280301FFF9AE876FD7BFD6C69CB" authorityName="Delile" authorityYear="1843" box="[516,678,691,716]" class="Magnoliopsida" family="Asteraceae" genus="Helichrysum" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="258" phylum="Tracheophyta" rank="species" species="citrispinum">
<emphasis id="B9629511301FFF9AE876FD7BFD6C69CB" box="[516,678,691,716]" italics="true" pageId="10" pageNumber="258">H. citrispinum</emphasis>
</taxonomicName>
is absent. Probably caterpillars develop there on other plant species of the genus
<taxonomicName id="4C163280301FFF9AEB55FD1FFE7C69F6" box="[295,438,727,753]" class="Magnoliopsida" family="Asteraceae" genus="Helichrysum" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="258" phylum="Tracheophyta" rank="genus">
<emphasis id="B9629511301FFF9AEB55FD1FFE7C69F6" box="[295,438,727,753]" italics="true" pageId="10" pageNumber="258">Helichrysum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA94903301FFF9AEAB5FD33FC5A683E" blockId="10.[151,1437,151,825]" pageId="10" pageNumber="258">
The localities where the species has been recorded are mostly at altitude
<quantity id="4CEEE4E6301FFF9AEE63FD32FB7A6812" box="[1041,1200,762,789]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="10" pageNumber="258" unit="m" value="2250.0" valueMax="2500.0" valueMin="2000.0">20002500 m</quantity>
except at the higher latitude of
<collectingCountry id="F3010993301FFF9AEB63FCD7FE6B683E" box="[273,417,799,825]" name="South Africa" pageId="10" pageNumber="258">South Africa</collectingCountry>
, reflecting the distribution of its host plants.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,645 @@
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<mods:title id="0D3B1DC8BE7A8B54B794EFC315195CF9">Zootaxa</mods:title>
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<mods:part id="6FB3EDEBB6D6DC9770C37E1460F5BFDD">
<mods:date id="57615430560F42963B0D245418537745">2024</mods:date>
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<mods:number id="E11F1AAC503228AB5669F945D26462D2">2024-11-04</mods:number>
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<mods:detail id="553EBCB0C539F8AA67451C76C71E1C04" type="volume">
<mods:number id="6D903A34EDE0F6D40FA0BFA8E0AD9323">5536</mods:number>
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<mods:url id="107F35516B315CA752E0B54022A23385">http://dx.doi.org/10.11646/zootaxa.5536.2.2</mods:url>
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<mods:classification id="2C54C6581D773D0F2EA723EEE4280419">journal article</mods:classification>
<mods:identifier id="DC7301096D2648361A03FD0D2960CDE0" type="DOI">10.11646/zootaxa.5536.2.2</mods:identifier>
<mods:identifier id="93550648BCDF9F1AC736B8BA7966D5FC" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BFF8153014FF94EAE5FDA2FEC8685A" ID-DOI="http://doi.org/10.5281/zenodo.14033548" ID-Zenodo-Dep="14033548" LSID="urn:lsid:plazi:treatment:03BFF8153014FF94EAE5FDA2FEC8685A" httpUri="http://treatment.plazi.org/id/03BFF8153014FF94EAE5FDA2FEC8685A" lastPageId="4" lastPageNumber="252" pageId="1" pageNumber="249">
<subSubSection id="C30C1A883014FF91EAE5FDA2FE176982" box="[151,477,618,645]" pageId="1" pageNumber="249" type="nomenclature">
<paragraph id="8BA949033014FF91EAE5FDA2FE176982" blockId="1.[151,477,618,645]" box="[151,477,618,645]" pageId="1" pageNumber="249">
<heading id="D0E1FE6F3014FF91EAE5FDA2FE176982" bold="true" box="[151,477,618,645]" fontSize="11" level="1" pageId="1" pageNumber="249" reason="1">
<emphasis id="B96295113014FF91EAE5FDA2FE176982" bold="true" box="[151,477,618,645]" pageId="1" pageNumber="249">
<taxonomicName id="4C1632803014FF91EAE5FDA2FEA26982" authorityName="Baraniak &amp; Agassiz" authorityYear="2024" box="[151,360,618,645]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96295113014FF91EAE5FDA2FEA26982" bold="true" box="[151,360,618,645]" italics="true" pageId="1" pageNumber="249">Pararhigognostis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251286A3014FF91EB1DFDA3FE176982" box="[367,477,619,645]" pageId="1" pageNumber="249" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30C1A883014FF91EAE5FD78FD1369E1" pageId="1" pageNumber="249" type="materials_examined">
<paragraph id="8BA949033014FF91EAE5FD78FC8269C0" blockId="1.[151,840,687,742]" box="[151,840,687,712]" pageId="1" pageNumber="249">
<typeStatus id="54ADF7A13014FF91EAE5FD78FF0369CF" box="[151,201,688,712]" pageId="1" pageNumber="249">Type</typeStatus>
species:
<taxonomicName id="4C1632803014FF91EB54FD67FD5269C0" authority="Meyrick, 1932" authorityName="Meyrick" authorityYear="1932" box="[294,664,687,711]" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="species" species="stichocentra">
<emphasis id="B96295113014FF91EB54FD67FE3069C0" box="[294,506,687,711]" italics="true" pageId="1" pageNumber="249">Plutella stichocentra</emphasis>
<bibRefCitation id="EF8734F23014FF91E872FD67FD5269C0" author="Meyrick, E." box="[512,664,687,711]" pageId="1" pageNumber="249" pagination="107 - 120" refId="ref5877" refString="Meyrick, E. (1932) Entomological expedition to Abyssinia, 1926 - 27 Microlepidoptera. The Transaction of the Entomological Society of London, 80 (1), 107 - 120. https: // doi. org / 10.1111 / j. 1365 - 2311.1932. tb 00085. x" type="journal article" year="1932">Meyrick, 1932</bibRefCitation>
</taxonomicName>
, here designated.
</paragraph>
<paragraph id="8BA949033014FF91EAE5FD06FD1369E1" blockId="1.[151,840,687,742]" box="[151,729,718,742]" pageId="1" pageNumber="249">
The genus
<taxonomicName id="4C1632803014FF91EB74FD06FE7D69E1" authorityName="Baraniak &amp; Agassiz" authorityYear="2024" box="[262,439,718,742]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113014FF91EB74FD06FE7D69E1" box="[262,439,718,742]" italics="true" pageId="1" pageNumber="249">Pararhigognostis</emphasis>
</taxonomicName>
comprises only one species.
</paragraph>
</subSubSection>
<subSubSection id="C30C1A883014FF91EAE5FCDCFD066E05" pageId="1" pageNumber="249" type="diagnosis">
<paragraph id="8BA949033014FF91EAE5FCDCFD066E05" blockId="1.[151,1437,788,2039]" pageId="1" pageNumber="249">
<emphasis id="B96295113014FF91EAE5FCDCFED86829" bold="true" box="[151,274,788,814]" pageId="1" pageNumber="249">Diagnosis.</emphasis>
Superficially, the forewing pattern of
<emphasis id="B96295113014FF91E8B1FCDCFCB36829" box="[707,889,788,814]" italics="true" pageId="1" pageNumber="249">Pararhigonostis</emphasis>
is composed of bands extending along the wing margin, similar to that of
<taxonomicName id="4C1632803014FF91EBCEFCF0FD816854" box="[444,587,824,851]" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96295113014FF91EBCEFCF0FDDF6855" box="[444,533,824,850]" italics="true" pageId="1" pageNumber="249">Plutella</emphasis>
spp.
</taxonomicName>
and
<taxonomicName id="4C1632803014FF91E8F6FCF0FC876854" box="[644,845,824,851]" class="Insecta" family="Plutellidae" genus="Rhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96295113014FF91E8F6FCF0FCDC6855" box="[644,790,824,850]" italics="true" pageId="1" pageNumber="249">Rhigognostis</emphasis>
spp.
</taxonomicName>
However, the male genitalia of
<emphasis id="B96295113014FF91EECEFCF0FAB86855" box="[1212,1394,824,850]" italics="true" pageId="1" pageNumber="249">Pararhigonostis</emphasis>
are remarkably different from those of
<taxonomicName id="4C1632803014FF91E869FC94FDBE6871" authorityName="Schrank" authorityYear="1802" box="[539,628,860,886]" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113014FF91E869FC94FDBE6871" box="[539,628,860,886]" italics="true" pageId="1" pageNumber="249">Plutella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1632803014FF91E8F3FC94FC186871" authority="(Baraniak 2007)" baseAuthorityName="Baraniak" baseAuthorityYear="2007" box="[641,978,860,886]" class="Insecta" family="Plutellidae" genus="Plutelloptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113014FF91E8F3FC94FCDF6871" box="[641,789,860,886]" italics="true" pageId="1" pageNumber="249">Plutelloptera</emphasis>
(
<bibRefCitation id="EF8734F23014FF91E953FC94FC006871" author="Baraniak, E." box="[801,970,860,886]" pageId="1" pageNumber="249" pagination="1 - 122" refId="ref4651" refString="Baraniak, E. (2007) Taxonomic revision of the genus Plutella Schranck, 1802 (Lepidoptera, Plutellidae) from the Palaearctic region with notes on its phylogeny. Polish Journal of Entomology, 76 (Supplement 1), 1 - 122." type="journal article" year="2007">Baraniak 2007</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C1632803014FF91E9AEFC94FB966871" box="[988,1116,860,886]" class="Insecta" family="Plutellidae" genus="Eidophasia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113014FF91E9AEFC94FB966871" box="[988,1116,860,886]" italics="true" pageId="1" pageNumber="249">Eidophasia</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1632803014FF91EEE4FC94FAE26871" baseAuthorityName="Baraniak" baseAuthorityYear="2016" box="[1174,1320,860,886]" class="Insecta" family="Plutellidae" genus="Rhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113014FF91EEE4FC94FAE26871" box="[1174,1320,860,886]" italics="true" pageId="1" pageNumber="249">Rhigognostis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8734F23014FF91EF47FC94FEFA689C" author="Baraniak, E. &amp; Sohn, J. - C." pageId="1" pageNumber="249" pagination="585 - 589" refId="ref4697" refString="Baraniak, E. &amp; Sohn, J. - C. (2015) Redescription of Eidophasia syenitella (Herrich-Schaffer, [1854]) (Lepidoptera, Plutellidae). Zootaxa, 4057 (4), 585 - 589. https: // doi. org / 10.11646 / zootaxa. 4057.4.9" type="journal article" year="2015">Baraniak &amp; Sohn 2015</bibRefCitation>
;
<bibRefCitation id="EF8734F23014FF91EB4EFC48FDAD689C" author="Baraniak, E. &amp; Walczak, U." box="[316,615,896,923]" pageId="1" pageNumber="249" pagination="445 - 448" refId="ref4752" refString="Baraniak, E. &amp; Walczak, U. (2015) Redescription of Eidophasia tauricella Staudinger, 1880 (Lepidoptera, Plutellidae) with first description of female genitalia. Zootaxa, 3956 (3), 445 - 448. https: // doi. org / 10.11646 / zootaxa. 3956.3.10" type="journal article" year="2015">Baraniak &amp; Walczak 2015</bibRefCitation>
;
<bibRefCitation id="EF8734F23014FF91E801FC48FCB3689D" author="Baraniak, E. &amp; Sohn, J. - C." box="[627,889,896,922]" pageId="1" pageNumber="249" pagination="164 - 172" refId="ref4806" refString="Baraniak, E. &amp; Sohn, J. - C. (2016) Revised taxonomic status of Eidophasia zukowskyi Amsel, 1938 (Lepidoptera, Plutellidae) with first description of its male and female genitalia. Zootaxa, 4162 (1), 164 - 172. https: // doi. org / 10.11646 / zootaxa. 4162.1.8" type="journal article" year="2016">Baraniak &amp; Sohn 2016</bibRefCitation>
;
<bibRefCitation id="EF8734F23014FF91E9F6FC48FB41689D" author="Baraniak, E. &amp; Sohn, J. - C." box="[900,1163,896,922]" pageId="1" pageNumber="249" pagination="164 - 172" refId="ref4806" refString="Baraniak, E. &amp; Sohn, J. - C. (2016) Revised taxonomic status of Eidophasia zukowskyi Amsel, 1938 (Lepidoptera, Plutellidae) with first description of its male and female genitalia. Zootaxa, 4162 (1), 164 - 172. https: // doi. org / 10.11646 / zootaxa. 4162.1.8" type="journal article" year="2016">Sohn &amp; Baraniak 2016</bibRefCitation>
). They are characterized by a thick, straight, sclerotized aedeagus which is twice as long as the valva and a vesica that lacks cornuti. The valva is elongate and trapezoidal with a straight costa bearing a very small, hooked apex; and the ventral margin is straight with a distinct, sclerotized, sharp, tooth-like process. The saccus is long and triangular. In the female genitalia of species of
<taxonomicName id="4C1632803014FF91EB79FBD8FE576F2D" baseAuthorityName="Baraniak" baseAuthorityYear="2016" box="[267,413,1040,1066]" class="Insecta" family="Plutellidae" genus="Rhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113014FF91EB79FBD8FE576F2D" box="[267,413,1040,1066]" italics="true" pageId="1" pageNumber="249">Rhigognostis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8734F23014FF91EBD9FBD8FDE56F2D" author="Kyrki, J." box="[427,559,1040,1066]" pageId="1" pageNumber="249" pagination="437 - 453" refId="ref5762" refString="Kyrki, J. (1989) Reassessment of the genus Rhigognostis Zeller, with descriptions of two new and notes on further seven Palaearctic species (Lepidoptera: Plutellidae). Entomologica Scandinavica, 19, 437 - 453. https: // doi. org / 10.1163 / 187631284 X 00064" type="journal article" year="1989">Kyrki 1989</bibRefCitation>
;
<bibRefCitation id="EF8734F23014FF91E84BFBD8FD216F2D" author="Baraniak, E." box="[569,747,1040,1066]" pageId="1" pageNumber="249" pagination="379 - 384" refId="ref4868" refString="Baraniak, E. (2016 a) Description of a new Nepalese Rhigognostis Staudinger, 1857 species (Lepidoptera: Plutellidae). SHILAP Revista de lepidopterologia, 44 (175), 379 - 384. [eISSN: 2340 - 4078] https: // doi. org / 10.57065 / shilap. 701" type="journal article" year="2016">Baraniak 2016a</bibRefCitation>
, b; Baraniak &amp; Larsen 2017; Baraniak
<emphasis id="B96295113014FF91EEE8FBD9FB1B6F2D" box="[1178,1233,1040,1066]" italics="true" pageId="1" pageNumber="249">et al.</emphasis>
2017), the antrum is elongate, terminating in a membranous part, whereas the remaining part is strongly sclerotized, with more or less rounded margins. Only
<taxonomicName id="4C1632803014FF91EBD7FB91FD966F75" box="[421,604,1112,1138]" class="Insecta" family="Plutellidae" genus="Rhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="species" species="husmanniella">
<emphasis id="B96295113014FF91EBD7FB91FD966F75" box="[421,604,1112,1138]" italics="true" pageId="1" pageNumber="249">R. husmanniella</emphasis>
</taxonomicName>
has an elongate antrum, but it is broadest in the middle; other details are characteristic of the genus
<taxonomicName id="4C1632803014FF91EBB4FBB4FCED6F91" authority="(Baraniak 2016 a)" baseAuthorityName="Baraniak" baseAuthorityYear="2016" box="[454,807,1148,1174]" class="Insecta" family="Plutellidae" genus="Rhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113014FF91EBB4FBB4FD926F91" box="[454,600,1148,1174]" italics="true" pageId="1" pageNumber="249">Rhigognostis</emphasis>
(
<bibRefCitation id="EF8734F23014FF91E815FBB4FCEA6F91" author="Baraniak, E." box="[615,800,1148,1174]" pageId="1" pageNumber="249" pagination="379 - 384" refId="ref4868" refString="Baraniak, E. (2016 a) Description of a new Nepalese Rhigognostis Staudinger, 1857 species (Lepidoptera: Plutellidae). SHILAP Revista de lepidopterologia, 44 (175), 379 - 384. [eISSN: 2340 - 4078] https: // doi. org / 10.57065 / shilap. 701" type="journal article" year="2016">Baraniak 2016a</bibRefCitation>
)
</taxonomicName>
. The bursa copulatrix of the genus
<taxonomicName id="4C1632803014FF91EEC7FBB4FA836F91" baseAuthorityName="Baraniak" baseAuthorityYear="2016" box="[1205,1353,1148,1174]" class="Insecta" family="Plutellidae" genus="Rhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="genus">Rhigognostis</taxonomicName>
is with or without signum, and the ductus seminalis opens in its upper part. In
<taxonomicName id="4C1632803014FF91E9D1FB68FBAD6FBD" authorityName="Baraniak &amp; Agassiz" authorityYear="2024" box="[931,1127,1184,1210]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113014FF91E9D1FB68FBAD6FBD" box="[931,1127,1184,1210]" italics="true" pageId="1" pageNumber="249">Pararhigognostis</emphasis>
</taxonomicName>
the antrum is elongate (
<figureCitation id="132D55863014FF91EF1DFB68FF706FD9" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[235,1353,187,1913]" captionTargetId="figure-26@9.[229,1359,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Pararhigognostis stichocentra female genitalia, gen. slide: YPO 20/2016: A, general view, B, antrum." figureDoi="http://doi.org/10.5281/zenodo.14033558" httpUri="https://zenodo.org/record/14033558/files/figure.png" pageId="1" pageNumber="249">Fig. 7B</figureCitation>
), terminating in a membranous part, but its remaining part is broadest near the inception of ductus bursae. Other distinguishing features are summarized in
<tableCitation id="C6947CB83014FF91E800FB20FD0C6E05" box="[626,710,1256,1282]" captionStart="TABLE 1" captionStartId="2.[152,245,468,494]" captionText="TABLE 1. Summary of differences in characters between the genera Plutella, Plutelloptera, Eidopasia messingiella group, E. syenitella group, Rhigognostis and the new genus Pararhigognostis." pageId="1" pageNumber="249">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30C1A883014FF94EAB5FAC4FE4169CA" lastPageId="4" lastPageNumber="252" pageId="1" pageNumber="249" type="description">
<paragraph id="8BA949033014FF91EAB5FAC4FCFE6C18" blockId="1.[151,1437,788,2039]" pageId="1" pageNumber="249">
<emphasis id="B96295113014FF91EAB5FAC4FE9B6E21" bold="true" box="[199,337,1292,1318]" pageId="1" pageNumber="249">Description</emphasis>
(
<figureCitation id="132D55863014FF91EB13FAC4FE6C6E21" box="[353,422,1292,1318]" captionStart="FIGURE 3" captionStartId="6.[152,255,1017,1041]" captionTargetBox="[193,1394,181,993]" captionTargetId="figure-19@6.[193,1394,181,993]" captionTargetPageId="6" captionText="FIGURE 3. Pararhigognostis stichocentra adult male: labial palpus, Kenya, Rift Valley, Turi, 06.VII.1999, 8000 ft., D.J.L. Agassiz leg." figureDoi="http://doi.org/10.5281/zenodo.14033550" httpUri="https://zenodo.org/record/14033550/files/figure.png" pageId="1" pageNumber="249">Fig. 3</figureCitation>
). Head: frons white appressed scales, vertex with white-grey erect scales; ocellus present; chaetosema present; antenna 2/3 of length of forewing; sensory setae of antenna short, sexually dimorphic absent; scape and pedicel white; flagellomeres white, with tiny greyish-yellow bands. Labial palp 3-segmented; outer surface of first segment white, with a narrow of dark brown scales in its upper part; outer surface of second and third segments covered with 3-coloured scales, at the base mostly white ones whereas upper part grey, with scattered brownishyellow scales. Inner side of first and second segments covered mostly with white scales and scattered 3-coloured scales. Tuft on second segment rather small, formed by similarly coloured scales. Third segment longest, curved, pointed apically. Maxillary palp small, inconspicuous, dark grey. Thorax: in part coloured like wing background and white-grey scales erected. Forewing lanceolate, long, wing tip gently rounded (
<figureCitation id="132D55863014FF91EE7EF9E4FBAD6D41" box="[1036,1127,1580,1606]" captionStart="FIGURE 4" captionStartId="6.[152,255,1998,2022]" captionTargetBox="[195,1392,1142,1971]" captionTargetId="figure-44@6.[189,1398,1137,1974]" captionTargetPageId="6" captionText="FIGURE 4. Pararhigognostis stichocentra, wing venations: A, forewing, B, hindwing." figureDoi="http://doi.org/10.5281/zenodo.14033552" httpUri="https://zenodo.org/record/14033552/files/figure.png" pageId="1" pageNumber="249">Fig. 4A</figureCitation>
); pterostigma small, clearly visible but narrow; subcostal vein (Sc) reaching costal margin halfway along its length; all free radial veins, R
<subScript id="17924B463014FF91EF02F9AAFAB26D77" attach="left" box="[1392,1400,1634,1648]" fontSize="6" pageId="1" pageNumber="249">1</subScript>
-R
<subScript id="17924B463014FF91EFE6F9AAFA566D77" attach="left" box="[1428,1436,1634,1648]" fontSize="6" pageId="1" pageNumber="249">4</subScript>
extend to costal margin, only R
<subScript id="17924B463014FF91EB85F94EFE356D93" attach="left" box="[503,511,1670,1684]" fontSize="6" pageId="1" pageNumber="249">5</subScript>
to termen margin (directly below the apex); medial (M
<subScript id="17924B463014FF91EE1FF94EFBBF6D93" attach="left" box="[1133,1141,1670,1684]" fontSize="6" pageId="1" pageNumber="249">1</subScript>
-M
<subScript id="17924B463014FF91EEE5F94EFB556D93" attach="left" box="[1175,1183,1670,1684]" fontSize="6" pageId="1" pageNumber="249">3</subScript>
) and cubital (Cu
<subScript id="17924B463014FF91EF2FF94EFAA66D93" attach="left" box="[1373,1388,1670,1684]" fontSize="6" pageId="1" pageNumber="249">1a</subScript>
and Cu
<subScript id="17924B463014FF91EACAF962FF026DBF" attach="left" box="[184,200,1706,1720]" fontSize="6" pageId="1" pageNumber="249">1b</subScript>
) veins free; postcubital vein poorly visible in middle part of its course; anal veins (A
<subScript id="17924B463014FF91EE0EF962FB4E6DBF" attach="left" box="[1148,1156,1706,1720]" fontSize="6" pageId="1" pageNumber="249">1</subScript>
and A
<subScript id="17924B463014FF91EEBFF962FB1F6DBF" attach="left" box="[1229,1237,1706,1720]" fontSize="6" pageId="1" pageNumber="249">2</subScript>
) free only at base of wings, then fused and conspicuous. Hindwing veins (
<figureCitation id="132D55863014FF91E97FF974FCAE6DD1" box="[781,868,1724,1750]" captionStart="FIGURE 4" captionStartId="6.[152,255,1998,2022]" captionTargetBox="[195,1392,1142,1971]" captionTargetId="figure-44@6.[189,1398,1137,1974]" captionTargetPageId="6" captionText="FIGURE 4. Pararhigognostis stichocentra, wing venations: A, forewing, B, hindwing." figureDoi="http://doi.org/10.5281/zenodo.14033552" httpUri="https://zenodo.org/record/14033552/files/figure.png" pageId="1" pageNumber="249">Fig. 4B</figureCitation>
) Sc+R
<subScript id="17924B463014FF91E9C0F906FC706DDB" attach="left" box="[946,954,1742,1756]" fontSize="6" pageId="1" pageNumber="249">1</subScript>
reaching costal margin at 2/3 of its length; Rs immediately before apex; middle M
<subScript id="17924B463014FF91E822F93AFD926C07" attach="left" box="[592,600,1778,1792]" fontSize="6" pageId="1" pageNumber="249">1</subScript>
to M
<subScript id="17924B463014FF91E8E7F93AFD576C07" attach="left" box="[661,669,1778,1792]" fontSize="6" pageId="1" pageNumber="249">3</subScript>
, cubital Cu
<subScript id="17924B463014FF91E952F93AFCE56C07" attach="left" box="[800,815,1778,1792]" fontSize="6" pageId="1" pageNumber="249">1a</subScript>
and Cu
<subScript id="17924B463014FF91E9F5F93AFC5D6C07" attach="left" box="[903,919,1778,1792]" fontSize="6" pageId="1" pageNumber="249">1b</subScript>
veins free, postcubital vein conspicuous; anal veins free, A
<subScript id="17924B463014FF91EB57F8DEFEE76C23" attach="left" box="[293,301,1814,1828]" fontSize="6" pageId="1" pageNumber="249">1</subScript>
vein, visible only in a small part of its course.
</paragraph>
<paragraph id="8BA949033014FF94EAB5F8E0FC316AD6" blockId="1.[151,1437,788,2039]" lastBlockId="4.[151,1437,151,861]" lastPageId="4" lastPageNumber="252" pageId="1" pageNumber="249">
Abdomen: Male genitalia (prep. gen. no. YPO 19/2016) (
<figureCitation id="132D55863014FF91E947F8E0FC5C6C44" box="[821,918,1832,1859]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="7.[152,255,1943,1967]" captionStartId-1="8.[152,255,1943,1967]" captionTargetBox-0="[232,1355,187,1913]" captionTargetBox-1="[252,1335,187,1914]" captionTargetId-0="figure-26@7.[226,1361,181,1919]" captionTargetId-1="figure-19@8.[246,1341,181,1920]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 5. Pararhigognostis stichocentra male genitalia, gen. slide YPO 19/2016: A, general view, B, aedeagus, C, socii." captionText-1="FIGURE 6. Pararhigognostis stichocentra male genitalia, gen. slide YPO 19/2016: A, separated valva, B, tip of aedeagus, C, aedeagus base with ductus ejaculatorius." figureDoi-0="http://doi.org/10.5281/zenodo.14033554" figureDoi-1="http://doi.org/10.5281/zenodo.14033556" httpUri-0="https://zenodo.org/record/14033554/files/figure.png" httpUri-1="https://zenodo.org/record/14033556/files/figure.png" pageId="1" pageNumber="249">Figs 56</figureCitation>
) with anal tube wide, socius long, membranous (
<figureCitation id="132D55863014FF91EAEDF884FF336C60" box="[159,249,1868,1895]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[232,1355,187,1913]" captionTargetId="figure-26@7.[226,1361,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Pararhigognostis stichocentra male genitalia, gen. slide YPO 19/2016: A, general view, B, aedeagus, C, socii." figureDoi="http://doi.org/10.5281/zenodo.14033554" httpUri="https://zenodo.org/record/14033554/files/figure.png" pageId="1" pageNumber="249">Fig. 5C</figureCitation>
) conspicuously wider in apical part, terminating in 4 distinct, irregularly spaced, pointed outgrowths, covered with fine, scanty spinules (
<figureCitation id="132D55863014FF91E86DF8B8FDB26C8C" box="[543,632,1904,1931]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[232,1355,187,1913]" captionTargetId="figure-26@7.[226,1361,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Pararhigognostis stichocentra male genitalia, gen. slide YPO 19/2016: A, general view, B, aedeagus, C, socii." figureDoi="http://doi.org/10.5281/zenodo.14033554" httpUri="https://zenodo.org/record/14033554/files/figure.png" pageId="1" pageNumber="249">Fig. 5A</figureCitation>
); valva irregular, rhomboid in outline, broadest at 2/3 of its length, distal part constricted, forming a gently rounded apex near costal margin (
<figureCitation id="132D55863014FF91E9FAF85CFC286CA9" box="[904,994,1940,1966]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[252,1335,187,1914]" captionTargetId="figure-19@8.[246,1341,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Pararhigognostis stichocentra male genitalia, gen. slide YPO 19/2016: A, separated valva, B, tip of aedeagus, C, aedeagus base with ductus ejaculatorius." figureDoi="http://doi.org/10.5281/zenodo.14033556" httpUri="https://zenodo.org/record/14033556/files/figure.png" pageId="1" pageNumber="249">Fig. 6A</figureCitation>
); valva longer than wide; in middle part with a conspicuous group of long, pointed, thin thorns, apically adhering to ventral margin, with two hairy areas; costal margin of valva from its basal margin to half its length straight, broadening into a small rounded outgrowth; from outgrowth to apex, costal margin gently rounded; ventral margin straight in proximal part, distal part slightly broader, forming a strongly sclerotized appendage, clearly separated from it, with sparse, minute spinules; appendage pointed apically; very short distal part of ventral margin at half its length with a poorly distinguished, sclerotized appendage; posterior margin oblique, slightly concave at 1/3 of its length, gently rounded to its apex; basal margin of valva short, oblique; processus basalis narrow, short, straight, apex with a conspicuous pointed appendage; apodema absent; vinculum narrow, saccus as long as valva, slightly constricted, with rounded apex. Aedeagus straight, thin, long, twice as long as valva, membranous, of even width, slightly widened basally (
<figureCitation id="132D55863011FF94EE27FEA7FB656A8E" box="[1109,1199,366,393]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[232,1355,187,1913]" captionTargetId="figure-26@7.[226,1361,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Pararhigognostis stichocentra male genitalia, gen. slide YPO 19/2016: A, general view, B, aedeagus, C, socii." figureDoi="http://doi.org/10.5281/zenodo.14033554" httpUri="https://zenodo.org/record/14033554/files/figure.png" pageId="4" pageNumber="252">Fig. 5B</figureCitation>
); vesica with minute spinules in apical part, from base to 2/3 its length markedly sclerotized, with 3 short linear groups of gently rounded spines (
<figureCitation id="132D55863011FF94EA9EFE7FFE8E6AD6" box="[236,324,439,465]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[252,1335,187,1914]" captionTargetId="figure-19@8.[246,1341,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Pararhigognostis stichocentra male genitalia, gen. slide YPO 19/2016: A, separated valva, B, tip of aedeagus, C, aedeagus base with ductus ejaculatorius." figureDoi="http://doi.org/10.5281/zenodo.14033556" httpUri="https://zenodo.org/record/14033556/files/figure.png" pageId="4" pageNumber="252">Fig. 6B</figureCitation>
); bulbus ejaculatorius membranous, large, irregular (
<figureCitation id="132D55863011FF94E9E7FE7FFC246AD6" box="[917,1006,439,465]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[252,1335,187,1914]" captionTargetId="figure-19@8.[246,1341,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Pararhigognostis stichocentra male genitalia, gen. slide YPO 19/2016: A, separated valva, B, tip of aedeagus, C, aedeagus base with ductus ejaculatorius." figureDoi="http://doi.org/10.5281/zenodo.14033556" httpUri="https://zenodo.org/record/14033556/files/figure.png" pageId="4" pageNumber="252">Fig. 6C</figureCitation>
).
</paragraph>
<caption id="DF69198B3017FF92EAEAFE1CFE926915" pageId="2" pageNumber="250" startId="2.[152,245,468,494]" targetType="table">
<paragraph id="8BA949033017FF92EAEAFE1CFE926915" blockId="2.[151,2037,468,530]" pageId="2" pageNumber="250">
<emphasis id="B96295113017FF92EAEAFE1CFEC56AE9" bold="true" box="[152,271,468,494]" pageId="2" pageNumber="250">TABLE 1.</emphasis>
Summary of differences in characters between the genera
<taxonomicName id="4C1632803017FF92E9F3FE1CFC1F6AE9" authorityName="Schrank" authorityYear="1802" box="[897,981,468,494]" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113017FF92E9F3FE1CFC1F6AE9" box="[897,981,468,494]" italics="true" pageId="2" pageNumber="250">Plutella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1632803017FF92E990FE1CFBA56AE9" baseAuthorityName="Baraniak" baseAuthorityYear="2007" box="[994,1135,468,494]" class="Insecta" family="Plutellidae" genus="Plutelloptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113017FF92E990FE1CFBA56AE9" box="[994,1135,468,494]" italics="true" pageId="2" pageNumber="250">Plutelloptera</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1632803017FF92EE0EFE1CFABE6AE9" authorityName="Fischer von Röslerstamm" authorityYear="1839" box="[1148,1396,468,494]" class="Insecta" family="Plutellidae" genus="Eidopasia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="species" species="messingiella">
<emphasis id="B96295113017FF92EE0EFE1CFABE6AE9" box="[1148,1396,468,494]" italics="true" pageId="2" pageNumber="250">Eidopasia messingiella</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C1632803017FF92EFB5FE1CF9816AE9" authorityName="Herrich-Schäffer" authorityYear="1855" box="[1479,1611,468,494]" class="Insecta" family="Plutellidae" genus="Eidopasia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="species" species="syenitella">
<emphasis id="B96295113017FF92EFB5FE1CF9816AE9" box="[1479,1611,468,494]" italics="true" pageId="2" pageNumber="250">E. syenitella</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C1632803017FF92ECEFFE1CF8E26AE9" baseAuthorityName="Baraniak" baseAuthorityYear="2016" box="[1693,1832,468,494]" class="Insecta" family="Plutellidae" genus="Rhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113017FF92ECEFFE1CF8E26AE9" box="[1693,1832,468,494]" italics="true" pageId="2" pageNumber="250">Rhigognostis</emphasis>
</taxonomicName>
and the new genus
<taxonomicName id="4C1632803017FF92EAE5FE30FE9B6915" authorityName="Baraniak &amp; Agassiz" authorityYear="2024" box="[151,337,504,530]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113017FF92EAE5FE30FE9B6915" box="[151,337,504,530]" italics="true" pageId="2" pageNumber="250">Pararhigognostis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA949033017FF92EAD0FDD7FD276D92" pageId="2" pageNumber="250">
<table id="F916BBA33017006FEAD0FDD7F8286D92" box="[162,2018,543,1685]" colsContinueIn="3.[151,2013,537,1688]" gridcols="7" gridrows="31" pageId="2" pageNumber="250">
<tr id="35264B413017006FEAD0FDD7F8286931" box="[162,2018,543,566]" gridrow="0" pageId="2" pageNumber="250">
<th id="76F7223D3017006FEAD0FDD7FEB56931" box="[162,383,543,566]" gridcol="0" gridrow="0" pageId="2" pageNumber="250">characters</th>
<th id="76F7223D3017006FEBEAFDD7FDA56931" box="[408,623,543,566]" gridcol="1" gridrow="0" pageId="2" pageNumber="250">
<taxonomicName id="4C1632803017FF92EBEAFDD7FE2E6931" authorityName="Schrank" authorityYear="1802" box="[408,484,543,566]" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113017FF92EBEAFDD7FE2E6931" box="[408,484,543,566]" italics="true" pageId="2" pageNumber="250">Plutella</emphasis>
</taxonomicName>
</th>
<th id="76F7223D3017006FE8F7FDD7FCDC6931" box="[645,790,543,566]" gridcol="2" gridrow="0" pageId="2" pageNumber="250">
<taxonomicName id="4C1632803017FF92E8F7FDD7FCCE6931" baseAuthorityName="Baraniak" baseAuthorityYear="2007" box="[645,772,543,566]" class="Insecta" family="Plutellidae" genus="Plutelloptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113017FF92E8F7FDD7FCCE6931" box="[645,772,543,566]" italics="true" pageId="2" pageNumber="250">Plutelloptera</emphasis>
</taxonomicName>
</th>
<th id="76F7223D3017006FE94FFDD7FBA06931" box="[829,1130,543,566]" gridcol="3" gridrow="0" pageId="2" pageNumber="250">
<taxonomicName id="4C1632803017FF92E94FFDD7FC1A6931" authorityName="Fischer von Röslerstamm" authorityYear="1839" box="[829,976,543,566]" class="Insecta" family="Plutellidae" genus="Eidopasia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="species" species="messingiella">
<emphasis id="B96295113017FF92E94FFDD7FC1A6931" box="[829,976,543,566]" italics="true" pageId="2" pageNumber="250">E. messingiella</emphasis>
</taxonomicName>
-group
</th>
<th id="76F7223D3017006FEEF5FDD7FAA36931" box="[1159,1385,543,566]" gridcol="4" gridrow="0" pageId="2" pageNumber="250">
<emphasis id="B96295113017FF92EEF5FDD7FAD96931" box="[1159,1299,543,566]" italics="true" pageId="2" pageNumber="250">
<taxonomicName id="4C1632803017FF92EEF5FDD7FB306931" authorityName="Herrich-Schäffer" authorityYear="1855" box="[1159,1274,543,566]" class="Insecta" family="Plutellidae" genus="Eidopasia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="species" species="syenitella">E. syenitella</taxonomicName>
</emphasis>
group
</th>
<th id="76F7223D3017006FEFFAFDD7F9B06931" box="[1416,1658,543,566]" gridcol="5" gridrow="0" pageId="2" pageNumber="250">
<taxonomicName id="4C1632803017FF92EFFAFDD7F9CF6931" baseAuthorityName="Baraniak" baseAuthorityYear="2016" box="[1416,1541,543,566]" class="Insecta" family="Plutellidae" genus="Rhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113017FF92EFFAFDD7F9CF6931" box="[1416,1541,543,566]" italics="true" pageId="2" pageNumber="250">Rhigognostis</emphasis>
</taxonomicName>
</th>
<th id="76F7223D3017006FECE0FDD7F8286931" box="[1682,2018,543,566]" gridcol="6" gridrow="0" pageId="2" pageNumber="250">
<taxonomicName id="4C1632803017FF92ECE0FDD7F8F06931" authorityName="Baraniak &amp; Agassiz" authorityYear="2024" box="[1682,1850,543,566]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113017FF92ECE0FDD7F8F06931" box="[1682,1850,543,566]" italics="true" pageId="2" pageNumber="250">Pararhigognostis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35264B413017006FEAD0FD89F8286959" box="[162,2018,577,606]" gridrow="1" pageId="2" pageNumber="250" rowspan-5="1">
<th id="76F7223D3017006FEAD0FD89FEB56959" box="[162,383,577,606]" gridcol="0" gridrow="1" pageId="2" pageNumber="250">course of R1 vein in</th>
<td id="76F7223D3017006FEBEAFD89FCDC6959" box="[408,790,577,606]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="2" pageNumber="250">reaches the costal margin within</td>
<td id="76F7223D3017006FE94FFD89FAA36959" box="[829,1385,577,606]" colspan="2" colspanRight="1" gridcol="3" gridrow="1" pageId="2" pageNumber="250">reaches the costal margin beyond pterostigma</td>
<td id="76F7223D3017006FECE0FD89F8286959" box="[1682,2018,577,606]" gridcol="6" gridrow="1" pageId="2" pageNumber="250">reaches the costal margin within</td>
</tr>
<tr id="35264B413017006FEAD0FDADF828697B" box="[162,2018,613,636]" gridrow="2" pageId="2" pageNumber="250" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76F7223D3017006FEAD0FDADFEB5697B" box="[162,383,613,636]" gridcol="0" gridrow="2" pageId="2" pageNumber="250">foreving</th>
<td id="76F7223D3017006FEBEAFDADFDA5697B" box="[408,623,613,636]" gridcol="1" gridrow="2" pageId="2" pageNumber="250">pterostigma</td>
<td id="76F7223D3017006FECE0FDADF828697B" box="[1682,2018,613,636]" gridcol="6" gridrow="2" pageId="2" pageNumber="250">the pterostigma (Fig. 4A)</td>
</tr>
<tr id="35264B413017006FEAD0FD40F82869CE" box="[162,2018,648,713]" gridrow="3" pageId="2" pageNumber="250" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76F7223D3017006FEAD0FD40FEB569CE" box="[162,383,648,713]" gridcol="0" gridrow="3" pageId="2" pageNumber="250">course veins of M1 and M3 in hindving</th>
<td id="76F7223D3017006FEBEAFD40FDA569CE" box="[408,623,648,713]" gridcol="1" gridrow="3" pageId="2" pageNumber="250">stalked M1 and M2</td>
<td id="76F7223D3017006FE94FFD40FBA069CE" box="[829,1130,648,713]" gridcol="3" gridrow="3" pageId="2" pageNumber="250">free (Fig. 4B)</td>
</tr>
<tr id="35264B413017006FEAD0FD2CF82869FC" box="[162,2018,740,763]" gridrow="4" pageId="2" pageNumber="250">
<th id="76F7223D3017006FEAD0FD2CF82869FC" box="[162,2018,740,763]" colspan="7" colspanRight="6" gridcol="0" gridrow="4" pageId="2" pageNumber="250">male</th>
</tr>
<tr id="35264B413017006FEAD0FCCFF8286819" box="[162,2018,775,798]" gridrow="5" pageId="2" pageNumber="250">
<th id="76F7223D3017006FEAD0FCCFFEB56819" box="[162,383,775,798]" gridcol="0" gridrow="5" pageId="2" pageNumber="250">valva shape</th>
<td id="76F7223D3017006FEBEAFCCFFDA56819" box="[408,623,775,798]" gridcol="1" gridrow="5" pageId="2" pageNumber="250">elongate with rounded</td>
<td id="76F7223D3017006FE8F7FCCFFCDC6819" box="[645,790,775,798]" gridcol="2" gridrow="5" pageId="2" pageNumber="250">broad, square</td>
<td id="76F7223D3017006FE94FFCCFFBA06819" box="[829,1130,775,798]" gridcol="3" gridrow="5" pageId="2" pageNumber="250">broad, obovate</td>
<td id="76F7223D3017006FEEF5FCCFFAA36819" box="[1159,1385,775,798]" gridcol="4" gridrow="5" pageId="2" pageNumber="250">elongate, rectangular in</td>
<td id="76F7223D3017006FEFFAFCCFF9B06819" box="[1416,1658,775,798]" gridcol="5" gridrow="5" pageId="2" pageNumber="250">elongate, gently rounded</td>
<td id="76F7223D3017006FECE0FCCFF8286819" box="[1682,2018,775,798]" gridcol="6" gridrow="5" pageId="2" pageNumber="250">elongate, trapezoid (Fig. 6A)</td>
</tr>
<tr id="35264B413017006FEAD0FCE3F8286845" box="[162,2018,811,834]" gridrow="6" pageId="2" pageNumber="250" rowspan-0="1" rowspan-3="1" rowspan-6="1">
<td id="76F7223D3017006FEBEAFCE3FDA56845" box="[408,623,811,834]" gridcol="1" gridrow="6" pageId="2" pageNumber="250">apex</td>
<td id="76F7223D3017006FE8F7FCE3FCDC6845" box="[645,790,811,834]" gridcol="2" gridrow="6" pageId="2" pageNumber="250">or trapezoid</td>
<td id="76F7223D3017006FEEF5FCE3FAA36845" box="[1159,1385,811,834]" gridcol="4" gridrow="6" pageId="2" pageNumber="250">outline</td>
<td id="76F7223D3017006FEFFAFCE3F9B06845" box="[1416,1658,811,834]" gridcol="5" gridrow="6" pageId="2" pageNumber="250">apex constricted at 2/3 of</td>
</tr>
<tr id="35264B413017006FEAD0FC87F8286861" box="[162,2018,847,870]" gridrow="7" pageId="2" pageNumber="250" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<td id="76F7223D3017006FE8F7FC87FCDC6861" box="[645,790,847,870]" gridcol="2" gridrow="7" pageId="2" pageNumber="250">outline</td>
<td id="76F7223D3017006FEFFAFC87F9B06861" box="[1416,1658,847,870]" gridcol="5" gridrow="7" pageId="2" pageNumber="250">its lenght</td>
</tr>
<tr id="35264B413017006FEAD0FCBAF828688E" box="[162,2018,882,905]" gridrow="8" pageId="2" pageNumber="250">
<th id="76F7223D3017006FEAD0FCBAFEB5688E" box="[162,383,882,905]" gridcol="0" gridrow="8" pageId="2" pageNumber="250">costal margin of valva</th>
<td id="76F7223D3017006FEBEAFCBAFDA5688E" box="[408,623,882,905]" gridcol="1" gridrow="8" pageId="2" pageNumber="250">gently curved</td>
<td id="76F7223D3017006FE8F7FCBAFCDC688E" box="[645,790,882,905]" gridcol="2" gridrow="8" pageId="2" pageNumber="250">straight</td>
<td id="76F7223D3017006FE94FFCBAFBA0688E" box="[829,1130,882,905]" gridcol="3" gridrow="8" pageId="2" pageNumber="250">slightly bent</td>
<td id="76F7223D3017006FEEF5FCBAFAA3688E" box="[1159,1385,882,905]" gridcol="4" gridrow="8" pageId="2" pageNumber="250">straight</td>
<td id="76F7223D3017006FEFFAFCBAF9B0688E" box="[1416,1658,882,905]" gridcol="5" gridrow="8" pageId="2" pageNumber="250">with a gently rounded,</td>
<td id="76F7223D3017006FECE0FCBAF828688E" box="[1682,2018,882,905]" gridcol="6" gridrow="8" pageId="2" pageNumber="250">straight with a very small, hooked</td>
</tr>
<tr id="35264B413017006FEAD0FC5EF82868AA" box="[162,2018,918,941]" gridrow="9" pageId="2" pageNumber="250" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="76F7223D3017006FEFFAFC5EF9B068AA" box="[1416,1658,918,941]" gridcol="5" gridrow="9" pageId="2" pageNumber="250">broad apex, constricted</td>
<td id="76F7223D3017006FECE0FC5EF82868AA" box="[1682,2018,918,941]" gridcol="6" gridrow="9" pageId="2" pageNumber="250">top halfway along (Fig. 6A)</td>
</tr>
<tr id="35264B413017006FEAD0FC72F82868D6" box="[162,2018,954,977]" gridrow="10" pageId="2" pageNumber="250" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<td id="76F7223D3017006FEFFAFC72F9B068D6" box="[1416,1658,954,977]" gridcol="5" gridrow="10" pageId="2" pageNumber="250">at 2/3 of its length</td>
</tr>
<tr id="35264B413017006FEAD0FC15F82868F3" box="[162,2018,989,1012]" gridrow="11" pageId="2" pageNumber="250">
<th id="76F7223D3017006FEAD0FC15FEB568F3" box="[162,383,989,1012]" gridcol="0" gridrow="11" pageId="2" pageNumber="250">posterior margin of</th>
<td id="76F7223D3017006FEBEAFC15FDA568F3" box="[408,623,989,1012]" gridcol="1" gridrow="11" pageId="2" pageNumber="250">distinctly elongated</td>
<td id="76F7223D3017006FE8F7FC15FCDC68F3" box="[645,790,989,1012]" gridcol="2" gridrow="11" pageId="2" pageNumber="250">gently rounded</td>
<td id="76F7223D3017006FE94FFC15FBA068F3" box="[829,1130,989,1012]" gridcol="3" gridrow="11" pageId="2" pageNumber="250">rounded, at the ventral margin</td>
<td id="76F7223D3017006FEEF5FC15FAA368F3" box="[1159,1385,989,1012]" gridcol="4" gridrow="11" pageId="2" pageNumber="250">gently rounded</td>
<td id="76F7223D3017006FEFFAFC15F9B068F3" box="[1416,1658,989,1012]" gridcol="5" gridrow="11" pageId="2" pageNumber="250">distinctly elongated</td>
<td id="76F7223D3017006FECE0FC15F82868F3" box="[1682,2018,989,1012]" gridcol="6" gridrow="11" pageId="2" pageNumber="250">straight with a small recess one</td>
</tr>
<tr id="35264B413017006FEAD0FBC9F8286F1F" box="[162,2018,1025,1048]" gridrow="12" pageId="2" pageNumber="250" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<th id="76F7223D3017006FEAD0FBC9FEB56F1F" box="[162,383,1025,1048]" gridcol="0" gridrow="12" pageId="2" pageNumber="250">valva</th>
<td id="76F7223D3017006FE94FFBC9FBA06F1F" box="[829,1130,1025,1048]" gridcol="3" gridrow="12" pageId="2" pageNumber="250">with a clear concentration of</td>
<td id="76F7223D3017006FECE0FBC9F8286F1F" box="[1682,2018,1025,1048]" gridcol="6" gridrow="12" pageId="2" pageNumber="250">third of the way through (Fig. 6A)</td>
</tr>
<tr id="35264B413017006FEAD0FBEDF8286F3B" box="[162,2018,1061,1084]" gridrow="13" pageId="2" pageNumber="250" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76F7223D3017006FE94FFBEDFBA06F3B" box="[829,1130,1061,1084]" gridcol="3" gridrow="13" pageId="2" pageNumber="250">raised thorns</td>
</tr>
<tr id="35264B413017006FEAD0FB8FF8286F59" box="[162,2018,1095,1118]" gridrow="14" pageId="2" pageNumber="250" rowspan-2="1">
<th id="76F7223D3017006FEAD0FB8FFEB56F59" box="[162,383,1095,1118]" gridcol="0" gridrow="14" pageId="2" pageNumber="250">ventral margin of</th>
<td id="76F7223D3017006FEBEAFB8FFDA56F59" box="[408,623,1095,1118]" gridcol="1" gridrow="14" pageId="2" pageNumber="250">straight</td>
<td id="76F7223D3017006FE94FFB8FFBA06F59" box="[829,1130,1095,1118]" gridcol="3" gridrow="14" pageId="2" pageNumber="250">slightly convex at 2/3 of its</td>
<td id="76F7223D3017006FEEF5FB8FFAA36F59" box="[1159,1385,1095,1118]" gridcol="4" gridrow="14" pageId="2" pageNumber="250">straight</td>
<td id="76F7223D3017006FEFFAFB8FF9B06F59" box="[1416,1658,1095,1118]" gridcol="5" gridrow="14" pageId="2" pageNumber="250">gently rounded</td>
<td id="76F7223D3017006FECE0FB8FF8286F59" box="[1682,2018,1095,1118]" gridcol="6" gridrow="14" pageId="2" pageNumber="250">straight, with a distinct, sclerotized</td>
</tr>
<tr id="35264B413017006FEAD0FBA3F8286F85" box="[162,2018,1131,1154]" gridrow="15" pageId="2" pageNumber="250" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<th id="76F7223D3017006FEAD0FBA3FEB56F85" box="[162,383,1131,1154]" gridcol="0" gridrow="15" pageId="2" pageNumber="250">valva</th>
<td id="76F7223D3017006FE94FFBA3FBA06F85" box="[829,1130,1131,1154]" gridcol="3" gridrow="15" pageId="2" pageNumber="250">lenght</td>
<td id="76F7223D3017006FECE0FBA3F8286F85" box="[1682,2018,1131,1154]" gridcol="6" gridrow="15" pageId="2" pageNumber="250">tooth-shaped appendage</td>
</tr>
<tr id="35264B413017006FEAD0FB47F8286FA1" box="[162,2018,1167,1190]" gridrow="16" pageId="2" pageNumber="250" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="76F7223D3017006FECE0FB47F8286FA1" box="[1682,2018,1167,1190]" gridcol="6" gridrow="16" pageId="2" pageNumber="250">immediately (Fig. 6A)</td>
</tr>
<tr id="35264B413017006FEAD0FB7AF8286FCE" box="[162,2018,1202,1225]" gridrow="17" pageId="2" pageNumber="250" rowspan-3="1" rowspan-5="1">
<th id="76F7223D3017006FEAD0FB7AFEB56FCE" box="[162,383,1202,1225]" gridcol="0" gridrow="17" pageId="2" pageNumber="250">posterior/ costal apex</th>
<td id="76F7223D3017006FEBEAFB7AFDA56FCE" box="[408,623,1202,1225]" gridcol="1" gridrow="17" pageId="2" pageNumber="250">oval</td>
<td id="76F7223D3017006FE8F7FB7AFCDC6FCE" box="[645,790,1202,1225]" gridcol="2" gridrow="17" pageId="2" pageNumber="250">rounded</td>
<td id="76F7223D3017006FEEF5FB7AFAA36FCE" box="[1159,1385,1202,1225]" gridcol="4" gridrow="17" pageId="2" pageNumber="250">oval</td>
<td id="76F7223D3017006FECE0FB7AF8286FCE" box="[1682,2018,1202,1225]" gridcol="6" gridrow="17" pageId="2" pageNumber="250">rounded (Fig. 6A)</td>
</tr>
<tr id="35264B413017006FEAD0FB1DF8286FEB" box="[162,2018,1237,1260]" gridrow="18" pageId="2" pageNumber="250" rowspan-2="1">
<th id="76F7223D3017006FEAD0FB1DFEB56FEB" box="[162,383,1237,1260]" gridcol="0" gridrow="18" pageId="2" pageNumber="250">posterior/ventral apex</th>
<td id="76F7223D3017006FEBEAFB1DFDA56FEB" box="[408,623,1237,1260]" gridcol="1" gridrow="18" pageId="2" pageNumber="250">rounded</td>
<td id="76F7223D3017006FE94FFB1DFBA06FEB" box="[829,1130,1237,1260]" gridcol="3" gridrow="18" pageId="2" pageNumber="250">raised, rounded with a group of</td>
<td id="76F7223D3017006FEEF5FB1DFAA36FEB" box="[1159,1385,1237,1260]" gridcol="4" gridrow="18" pageId="2" pageNumber="250">rounded</td>
<td id="76F7223D3017006FEFFAFB1DF9B06FEB" box="[1416,1658,1237,1260]" gridcol="5" gridrow="18" pageId="2" pageNumber="250">oval</td>
<td id="76F7223D3017006FECE0FB1DF8286FEB" box="[1682,2018,1237,1260]" gridcol="6" gridrow="18" pageId="2" pageNumber="250">triangular (Fig. 6A)</td>
</tr>
<tr id="35264B413017006FEAD0FB31F8286E17" box="[162,2018,1273,1296]" gridrow="19" pageId="2" pageNumber="250" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76F7223D3017006FE94FFB31FBA06E17" box="[829,1130,1273,1296]" gridcol="3" gridrow="19" pageId="2" pageNumber="250">small thorns</td>
</tr>
<tr id="35264B413017006FEAD0FAD4F8286E34" box="[162,2018,1308,1331]" gridrow="20" pageId="2" pageNumber="250" rowspan-6="1">
<th id="76F7223D3017006FEAD0FAD4FEB56E34" box="[162,383,1308,1331]" gridcol="0" gridrow="20" pageId="2" pageNumber="250">sacculus</th>
<td id="76F7223D3017006FEBEAFAD4FDA56E34" box="[408,623,1308,1331]" gridcol="1" gridrow="20" pageId="2" pageNumber="250">fololded with distinct</td>
<td id="76F7223D3017006FE8F7FAD4FCDC6E34" box="[645,790,1308,1331]" gridcol="2" gridrow="20" pageId="2" pageNumber="250">absent</td>
<td id="76F7223D3017006FE94FFAD4FBA06E34" box="[829,1130,1308,1331]" gridcol="3" gridrow="20" pageId="2" pageNumber="250">deeply folded basally, which</td>
<td id="76F7223D3017006FEEF5FAD4FAA36E34" box="[1159,1385,1308,1331]" gridcol="4" gridrow="20" pageId="2" pageNumber="250">long, club-shaped,</td>
<td id="76F7223D3017006FEFFAFAD4F9B06E34" box="[1416,1658,1308,1331]" gridcol="5" gridrow="20" pageId="2" pageNumber="250">absent</td>
</tr>
<tr id="35264B413017006FEAD0FA88F8286E50" box="[162,2018,1344,1367]" gridrow="21" pageId="2" pageNumber="250" rowspan-0="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<td id="76F7223D3017006FEBEAFA88FDA56E50" box="[408,623,1344,1367]" gridcol="1" gridrow="21" pageId="2" pageNumber="250">spines</td>
<td id="76F7223D3017006FE94FFA88FBA06E50" box="[829,1130,1344,1367]" gridcol="3" gridrow="21" pageId="2" pageNumber="250">two complicated arms which</td>
<td id="76F7223D3017006FEEF5FA88FAA36E50" box="[1159,1385,1344,1367]" gridcol="4" gridrow="21" pageId="2" pageNumber="250">margins conspicuously</td>
</tr>
<tr id="35264B413017006FEAD0FAACF8286E7C" box="[162,2018,1380,1403]" gridrow="22" pageId="2" pageNumber="250" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<td id="76F7223D3017006FE94FFAACFBA06E7C" box="[829,1130,1380,1403]" gridcol="3" gridrow="22" pageId="2" pageNumber="250">bear stout spines</td>
<td id="76F7223D3017006FEEF5FAACFAA36E7C" box="[1159,1385,1380,1403]" gridcol="4" gridrow="22" pageId="2" pageNumber="250">denticulate</td>
</tr>
<tr id="35264B413017006FEAD0FA4FF8286E99" box="[162,2018,1415,1438]" gridrow="23" pageId="2" pageNumber="250" rowspan-2="1" rowspan-5="1">
<th id="76F7223D3017006FEAD0FA4FFEB56E99" box="[162,383,1415,1438]" gridcol="0" gridrow="23" pageId="2" pageNumber="250">length of aedeagus as</th>
<td id="76F7223D3017006FEBEAFA4FFDA56E99" box="[408,623,1415,1438]" gridcol="1" gridrow="23" pageId="2" pageNumber="250">shorter</td>
<td id="76F7223D3017006FE94FFA4FFBA06E99" box="[829,1130,1415,1438]" gridcol="3" gridrow="23" pageId="2" pageNumber="250">slightly longer</td>
<td id="76F7223D3017006FEEF5FA4FFAA36E99" box="[1159,1385,1415,1438]" gridcol="4" gridrow="23" pageId="2" pageNumber="250">shorter</td>
<td id="76F7223D3017006FECE0FA4FF8286E99" box="[1682,2018,1415,1438]" gridcol="6" gridrow="23" pageId="2" pageNumber="250">twice as long</td>
</tr>
<tr id="35264B413017006FEAD0FA63F8286EC5" box="[162,2018,1451,1474]" gridrow="24" pageId="2" pageNumber="250" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76F7223D3017006FEAD0FA63FEB56EC5" box="[162,383,1451,1474]" gridcol="0" gridrow="24" pageId="2" pageNumber="250">compare to valve</th>
</tr>
<tr id="35264B413017006FEAD0FA05F8286EE3" box="[162,2018,1485,1508]" gridrow="25" pageId="2" pageNumber="250" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<th id="76F7223D3017006FEAD0FA05FEB56EE3" box="[162,383,1485,1508]" gridcol="0" gridrow="25" pageId="2" pageNumber="250">shape of aedeagus</th>
<td id="76F7223D3017006FEBEAFA05FDA56EE3" box="[408,623,1485,1508]" gridcol="1" gridrow="25" pageId="2" pageNumber="250">straight</td>
<td id="76F7223D3017006FE94FFA05FBA06EE3" box="[829,1130,1485,1508]" gridcol="3" gridrow="25" pageId="2" pageNumber="250">curved</td>
<td id="76F7223D3017006FECE0FA05F8286EE3" box="[1682,2018,1485,1508]" gridcol="6" gridrow="25" pageId="2" pageNumber="250">straight (Fig. 5B)</td>
</tr>
<tr id="35264B413017006FEAD0FA38F8286D00" box="[162,2018,1520,1543]" gridrow="26" pageId="2" pageNumber="250">
<th id="76F7223D3017006FEAD0FA38FEB56D00" box="[162,383,1520,1543]" gridcol="0" gridrow="26" pageId="2" pageNumber="250">base of aedeagus</th>
<td id="76F7223D3017006FEBEAFA38FDA56D00" box="[408,623,1520,1543]" gridcol="1" gridrow="26" pageId="2" pageNumber="250">broadened with a pair</td>
<td id="76F7223D3017006FE8F7FA38FCDC6D00" box="[645,790,1520,1543]" gridcol="2" gridrow="26" pageId="2" pageNumber="250">broadened</td>
<td id="76F7223D3017006FE94FFA38FBA06D00" box="[829,1130,1520,1543]" gridcol="3" gridrow="26" pageId="2" pageNumber="250">slightly dilated</td>
<td id="76F7223D3017006FEEF5FA38FAA36D00" box="[1159,1385,1520,1543]" gridcol="4" gridrow="26" pageId="2" pageNumber="250">expanded on one side</td>
<td id="76F7223D3017006FEFFAFA38F9B06D00" box="[1416,1658,1520,1543]" gridcol="5" gridrow="26" pageId="2" pageNumber="250">unilaterally extended</td>
<td id="76F7223D3017006FECE0FA38F8286D00" box="[1682,2018,1520,1543]" gridcol="6" gridrow="26" pageId="2" pageNumber="250">slightly, dilated (Fig. 6C)</td>
</tr>
<tr id="35264B413017006FEAD0F9DCF8286D2C" box="[162,2018,1556,1579]" gridrow="27" pageId="2" pageNumber="250">
<th id="76F7223D3017006FEAD0F9DCF8286D2C" box="[162,2018,1556,1579]" colspan="7" colspanRight="6" gridcol="0" gridrow="27" pageId="2" pageNumber="250">of lateral hooks</th>
</tr>
<tr id="35264B413017006FEAD0F9FFF8286D49" box="[162,2018,1591,1614]" gridrow="28" pageId="2" pageNumber="250">
<th id="76F7223D3017006FEAD0F9FFFEB56D49" box="[162,383,1591,1614]" gridcol="0" gridrow="28" pageId="2" pageNumber="250">apex of aedeagus</th>
<td id="76F7223D3017006FEBEAF9FFFDA56D49" box="[408,623,1591,1614]" gridcol="1" gridrow="28" pageId="2" pageNumber="250">needle shaped</td>
<td id="76F7223D3017006FE8F7F9FFFCDC6D49" box="[645,790,1591,1614]" gridcol="2" gridrow="28" pageId="2" pageNumber="250">narrowed</td>
<td id="76F7223D3017006FE94FF9FFFBA06D49" box="[829,1130,1591,1614]" gridcol="3" gridrow="28" pageId="2" pageNumber="250">slightly wider</td>
<td id="76F7223D3017006FEEF5F9FFFAA36D49" box="[1159,1385,1591,1614]" gridcol="4" gridrow="28" pageId="2" pageNumber="250">gently tapering</td>
<td id="76F7223D3017006FEFFAF9FFF9B06D49" box="[1416,1658,1591,1614]" gridcol="5" gridrow="28" pageId="2" pageNumber="250">sharpened</td>
<td id="76F7223D3017006FECE0F9FFF8286D49" box="[1682,2018,1591,1614]" gridcol="6" gridrow="28" pageId="2" pageNumber="250">one-sided sharpened (Fig. 6B)</td>
</tr>
<tr id="35264B413017006FEAD0F992F8286D76" box="[162,2018,1626,1649]" gridrow="29" pageId="2" pageNumber="250">
<th id="76F7223D3017006FEAD0F992FEB56D76" box="[162,383,1626,1649]" gridcol="0" gridrow="29" pageId="2" pageNumber="250">aedeagus</th>
<td id="76F7223D3017006FEBEAF992FDA56D76" box="[408,623,1626,1649]" gridcol="1" gridrow="29" pageId="2" pageNumber="250">sclerotized</td>
<td id="76F7223D3017006FE8F7F992FCDC6D76" box="[645,790,1626,1649]" gridcol="2" gridrow="29" pageId="2" pageNumber="250">weakly</td>
<td id="76F7223D3017006FE94FF992FBA06D76" box="[829,1130,1626,1649]" gridcol="3" gridrow="29" pageId="2" pageNumber="250">membranous</td>
<td id="76F7223D3017006FEEF5F992FAA36D76" box="[1159,1385,1626,1649]" gridcol="4" gridrow="29" pageId="2" pageNumber="250">sclerotized</td>
<td id="76F7223D3017006FEFFAF992F9B06D76" box="[1416,1658,1626,1649]" gridcol="5" gridrow="29" pageId="2" pageNumber="250">weakly sclerotized</td>
<td id="76F7223D3017006FECE0F992F8286D76" box="[1682,2018,1626,1649]" gridcol="6" gridrow="29" pageId="2" pageNumber="250">sclerotized (Fig. 5B)</td>
</tr>
<tr id="35264B413017006FEAD0F9B6F8286D92" box="[162,2018,1662,1685]" gridrow="30" pageId="2" pageNumber="250" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76F7223D3017006FE8F7F9B6FCDC6D92" box="[645,790,1662,1685]" gridcol="2" gridrow="30" pageId="2" pageNumber="250">sclerotized</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA949033017FF92ECB1F969F8206DBF" blockId="2.[1731,2026,1697,1720]" box="[1731,2026,1697,1720]" pageId="2" pageNumber="250">
<tableNote id="76F0488D3017FF92ECB1F969F8206DBF" box="[1731,2026,1697,1720]" pageId="2" pageNumber="250" targetBox="[162,2018,543,1685]" targetPageId="2">......continued on the next page</tableNote>
</paragraph>
<caption id="DF69198B3016FF93EAE5FE3FFEB66909" box="[151,380,503,527]" isContinuationCaption="true" pageId="3" pageNumber="251" startId="3.[151,234,503,527]" targetType="table">
<paragraph id="8BA949033016FF93EAE5FE3FFEB66909" blockId="3.[151,380,503,527]" box="[151,380,503,527]" pageId="3" pageNumber="251">
<emphasis id="B96295113016FF93EAE5FE3FFEC96908" bold="true" box="[151,259,503,527]" pageId="3" pageNumber="251">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8BA949033016FF93EAE5FDD1F90E6D9F" pageId="3" pageNumber="251">
<table id="F916BBA33016006FEAE5FDD1F8176D9F" box="[151,2013,537,1688]" colsContinueFrom="2.[162,2018,543,1685]" gridcols="7" gridrows="15" pageId="3" pageNumber="251">
<tr id="35264B413016006FEAE5FDD1F8176937" box="[151,2013,537,560]" gridrow="0" pageId="3" pageNumber="251">
<th id="76F7223D3016006FEAE5FDD1FEB96937" box="[151,371,537,560]" gridcol="0" gridrow="0" pageId="3" pageNumber="251">characters</th>
<th id="76F7223D3016006FEBFEFDD1FD9C6937" box="[396,598,537,560]" gridcol="1" gridrow="0" pageId="3" pageNumber="251">
<taxonomicName id="4C1632803016FF93EBFEFDD1FE126937" authorityName="Schrank" authorityYear="1802" box="[396,472,537,560]" class="Insecta" family="Plutellidae" genus="Plutella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113016FF93EBFEFDD1FE126937" box="[396,472,537,560]" italics="true" pageId="3" pageNumber="251">Plutella</emphasis>
</taxonomicName>
</th>
<th id="76F7223D3016006FE808FDD1FCDE6937" box="[634,788,537,560]" gridcol="2" gridrow="0" pageId="3" pageNumber="251">
<taxonomicName id="4C1632803016FF93E808FDD1FD336937" baseAuthorityName="Baraniak" baseAuthorityYear="2007" box="[634,761,537,560]" class="Insecta" family="Plutellidae" genus="Plutelloptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113016FF93E808FDD1FD336937" box="[634,761,537,560]" italics="true" pageId="3" pageNumber="251">Plutelloptera</emphasis>
</taxonomicName>
</th>
<th id="76F7223D3016006FE940FDD1FB906937" box="[818,1114,537,560]" gridcol="3" gridrow="0" pageId="3" pageNumber="251">
<taxonomicName id="4C1632803016FF93E940FDD1FC0E6937" authorityName="Fischer von Röslerstamm" authorityYear="1839" box="[818,964,537,560]" class="Insecta" family="Plutellidae" genus="Eidopasia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="species" species="messingiella">
<emphasis id="B96295113016FF93E940FDD1FC0E6937" box="[818,964,537,560]" italics="true" pageId="3" pageNumber="251">E. messingiella</emphasis>
</taxonomicName>
-group
</th>
<th id="76F7223D3016006FEE0EFDD1FA876937" box="[1148,1357,537,560]" gridcol="4" gridrow="0" pageId="3" pageNumber="251">
<emphasis id="B96295113016FF93EE0EFDD1FAC26937" box="[1148,1288,537,560]" italics="true" pageId="3" pageNumber="251">
<taxonomicName id="4C1632803016FF93EE0EFDD1FB246937" authorityName="Herrich-Schäffer" authorityYear="1855" box="[1148,1262,537,560]" class="Insecta" family="Plutellidae" genus="Eidopasia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="species" species="syenitella">E. syenitella</taxonomicName>
</emphasis>
group
</th>
<th id="76F7223D3016006FEF0EFDD1F9966937" box="[1404,1628,537,560]" gridcol="5" gridrow="0" pageId="3" pageNumber="251">
<taxonomicName id="4C1632803016FF93EF0EFDD1FA336937" baseAuthorityName="Baraniak" baseAuthorityYear="2016" box="[1404,1529,537,560]" class="Insecta" family="Plutellidae" genus="Rhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113016FF93EF0EFDD1FA336937" box="[1404,1529,537,560]" italics="true" pageId="3" pageNumber="251">Rhigognostis</emphasis>
</taxonomicName>
</th>
<th id="76F7223D3016006FECF5FDD1F8176937" box="[1671,2013,537,560]" gridcol="6" gridrow="0" pageId="3" pageNumber="251">
<taxonomicName id="4C1632803016FF93ECF5FDD1F8E56937" authorityName="Baraniak &amp; Agassiz" authorityYear="2024" box="[1671,1839,537,560]" class="Insecta" family="Plutellidae" genus="Pararhigognostis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="B96295113016FF93ECF5FDD1F8E56937" box="[1671,1839,537,560]" italics="true" pageId="3" pageNumber="251">Pararhigognostis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35264B413016006FEAE5FDF5F817697F" box="[151,2013,573,632]" gridrow="1" pageId="3" pageNumber="251" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76F7223D3016006FEAE5FDF5FEB9697F" box="[151,371,573,632]" gridcol="0" gridrow="1" pageId="3" pageNumber="251">cornuti with aedeagus</th>
<td id="76F7223D3016006FEBFEFDF5FD9C697F" box="[396,598,573,632]" gridcol="1" gridrow="1" pageId="3" pageNumber="251">absent</td>
<td id="76F7223D3016006FEF0EFDF5F996697F" box="[1404,1628,573,632]" gridcol="5" gridrow="1" pageId="3" pageNumber="251">single long dentate, pointed spine</td>
<td id="76F7223D3016006FECF5FDF5F817697F" box="[1671,2013,573,632]" gridcol="6" gridrow="1" pageId="3" pageNumber="251">absent</td>
</tr>
<tr id="35264B413016006FEAE5FD4CF81769E4" box="[151,2013,644,739]" gridrow="2" pageId="3" pageNumber="251">
<th id="76F7223D3016006FEAE5FD4CFEB969E4" box="[151,371,644,739]" gridcol="0" gridrow="2" pageId="3" pageNumber="251">tegumen gnathos complex</th>
<td id="76F7223D3016006FEBFEFD4CFD9C69E4" box="[396,598,644,739]" gridcol="1" gridrow="2" pageId="3" pageNumber="251">gnathal process surrounding of the anal tube</td>
<td id="76F7223D3016006FE808FD4CFCDE69E4" box="[634,788,644,739]" gridcol="2" gridrow="2" pageId="3" pageNumber="251">not isolated</td>
<td id="76F7223D3016006FE940FD4CFB9069E4" box="[818,1114,644,739]" gridcol="3" gridrow="2" pageId="3" pageNumber="251">semi-elliptical</td>
<td id="76F7223D3016006FEE0EFD4CFA8769E4" box="[1148,1357,644,739]" gridcol="4" gridrow="2" pageId="3" pageNumber="251">not isolated</td>
<td id="76F7223D3016006FEF0EFD4CF99669E4" box="[1404,1628,644,739]" gridcol="5" gridrow="2" pageId="3" pageNumber="251">gnathal process surrounding of the anal tube</td>
<td id="76F7223D3016006FECF5FD4CF81769E4" box="[1671,2013,644,739]" gridcol="6" gridrow="2" pageId="3" pageNumber="251">not isolated</td>
</tr>
<tr id="35264B413016006FEAE5FD26F817684A" box="[151,2013,750,845]" gridrow="3" pageId="3" pageNumber="251">
<th id="76F7223D3016006FEAE5FD26FEB9684A" box="[151,371,750,845]" gridcol="0" gridrow="3" pageId="3" pageNumber="251">saccus</th>
<td id="76F7223D3016006FEBFEFD26FD9C684A" box="[396,598,750,845]" gridcol="1" gridrow="3" pageId="3" pageNumber="251">large, long strongly sclerotized</td>
<td id="76F7223D3016006FE808FD26FCDE684A" box="[634,788,750,845]" gridcol="2" gridrow="3" pageId="3" pageNumber="251">large, shorter, strongly slerotized</td>
<td id="76F7223D3016006FE940FD26FB90684A" box="[818,1114,750,845]" gridcol="3" gridrow="3" pageId="3" pageNumber="251">wide, short and slightly curved</td>
<td id="76F7223D3016006FEE0EFD26FA87684A" box="[1148,1357,750,845]" gridcol="4" gridrow="3" pageId="3" pageNumber="251">long and narrow</td>
<td id="76F7223D3016006FEF0EFD26F996684A" box="[1404,1628,750,845]" gridcol="5" gridrow="3" pageId="3" pageNumber="251">very long, narrow</td>
<td id="76F7223D3016006FECF5FD26F817684A" box="[1671,2013,750,845]" gridcol="6" gridrow="3" pageId="3" pageNumber="251">long (Fig. 5A)</td>
</tr>
<tr id="35264B413016006FEAE5FC91F8176877" box="[151,2013,857,880]" gridrow="4" pageId="3" pageNumber="251" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76F7223D3016006FEAE5FC91FEB96877" box="[151,371,857,880]" gridcol="0" gridrow="4" pageId="3" pageNumber="251">apodema</th>
<td id="76F7223D3016006FEBFEFC91FD9C6877" box="[396,598,857,880]" gridcol="1" gridrow="4" pageId="3" pageNumber="251">absent</td>
<td id="76F7223D3016006FEF0EFC91F9966877" box="[1404,1628,857,880]" gridcol="5" gridrow="4" pageId="3" pageNumber="251">present</td>
<td id="76F7223D3016006FECF5FC91F8176877" box="[1671,2013,857,880]" gridcol="6" gridrow="4" pageId="3" pageNumber="251">absent</td>
</tr>
<tr id="35264B413016006FEAE5FC7DF81768CB" box="[151,2013,949,972]" gridrow="5" pageId="3" pageNumber="251" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76F7223D3016006FEAE5FC7DFEB968CB" box="[151,371,949,972]" gridcol="0" gridrow="5" pageId="3" pageNumber="251">female</th>
</tr>
<tr id="35264B413016006FEAE5FC10F81768E8" box="[151,2013,984,1007]" gridrow="6" pageId="3" pageNumber="251" rowspan-4="1">
<th id="76F7223D3016006FEAE5FC10FEB968E8" box="[151,371,984,1007]" gridcol="0" gridrow="6" pageId="3" pageNumber="251">apophyses posteriores</th>
<td id="76F7223D3016006FEBFEFC10FD9C68E8" box="[396,598,984,1007]" gridcol="1" gridrow="6" pageId="3" pageNumber="251">thiny, short</td>
<td id="76F7223D3016006FE808FC10FCDE68E8" box="[634,788,984,1007]" gridcol="2" gridrow="6" pageId="3" pageNumber="251">thiny, longer</td>
<td id="76F7223D3016006FE940FC10FB9068E8" box="[818,1114,984,1007]" gridcol="3" gridrow="6" pageId="3" pageNumber="251">thiny, short</td>
<td id="76F7223D3016006FEF0EFC10F99668E8" box="[1404,1628,984,1007]" gridcol="5" gridrow="6" pageId="3" pageNumber="251">thiny, very short</td>
<td id="76F7223D3016006FECF5FC10F81768E8" box="[1671,2013,984,1007]" gridcol="6" gridrow="6" pageId="3" pageNumber="251">thiny, long (Fig. 7A)</td>
</tr>
<tr id="35264B413016006FEAE5FBFCF8176F68" box="[151,2013,1076,1135]" gridrow="7" pageId="3" pageNumber="251">
<th id="76F7223D3016006FEAE5FBFCFEB96F68" box="[151,371,1076,1135]" gridcol="0" gridrow="7" pageId="3" pageNumber="251">apophyses anteriores</th>
<td id="76F7223D3016006FEBFEFBFCFD9C6F68" box="[396,598,1076,1135]" gridcol="1" gridrow="7" pageId="3" pageNumber="251">thiny, long</td>
<td id="76F7223D3016006FE808FBFCFCDE6F68" box="[634,788,1076,1135]" gridcol="2" gridrow="7" pageId="3" pageNumber="251">thiny, short</td>
<td id="76F7223D3016006FE940FBFCFB906F68" box="[818,1114,1076,1135]" gridcol="3" gridrow="7" pageId="3" pageNumber="251">thiny and curved in half its length</td>
<td id="76F7223D3016006FEE0EFBFCFA876F68" box="[1148,1357,1076,1135]" gridcol="4" gridrow="7" pageId="3" pageNumber="251">thiny</td>
<td id="76F7223D3016006FEF0EFBFCF9966F68" box="[1404,1628,1076,1135]" gridcol="5" gridrow="7" pageId="3" pageNumber="251">thiny, very short</td>
<td id="76F7223D3016006FECF5FBFCF8176F68" box="[1671,2013,1076,1135]" gridcol="6" gridrow="7" pageId="3" pageNumber="251">thiny, long (Fig. 7A)</td>
</tr>
<tr id="35264B413016006FEAE5FBB3F8176FDD" box="[151,2013,1147,1242]" gridrow="8" pageId="3" pageNumber="251">
<th id="76F7223D3016006FEAE5FBB3FEB96FDD" box="[151,371,1147,1242]" gridcol="0" gridrow="8" pageId="3" pageNumber="251">length of a. posteriores as compare to a. anteriores</th>
<td id="76F7223D3016006FEBFEFBB3FD9C6FDD" box="[396,598,1147,1242]" gridcol="1" gridrow="8" pageId="3" pageNumber="251">shorter</td>
<td id="76F7223D3016006FE808FBB3FCDE6FDD" box="[634,788,1147,1242]" gridcol="2" gridrow="8" pageId="3" pageNumber="251">longer</td>
<td id="76F7223D3016006FE940FBB3FB906FDD" box="[818,1114,1147,1242]" gridcol="3" gridrow="8" pageId="3" pageNumber="251">slightly longer</td>
<td id="76F7223D3016006FEE0EFBB3FA876FDD" box="[1148,1357,1147,1242]" gridcol="4" gridrow="8" pageId="3" pageNumber="251">this same lenght</td>
<td id="76F7223D3016006FEF0EFBB3F9966FDD" box="[1404,1628,1147,1242]" gridcol="5" gridrow="8" pageId="3" pageNumber="251">twice as long</td>
<td id="76F7223D3016006FECF5FBB3F8176FDD" box="[1671,2013,1147,1242]" gridcol="6" gridrow="8" pageId="3" pageNumber="251">longer</td>
</tr>
<tr id="35264B413016006FEAE5FB2EF8176E42" box="[151,2013,1254,1349]" gridrow="9" pageId="3" pageNumber="251">
<th id="76F7223D3016006FEAE5FB2EFEB96E42" box="[151,371,1254,1349]" gridcol="0" gridrow="9" pageId="3" pageNumber="251">antrum</th>
<td id="76F7223D3016006FEBFEFB2EFD9C6E42" box="[396,598,1254,1349]" gridcol="1" gridrow="9" pageId="3" pageNumber="251">cup like, thin, only slighty wider than ductus bursa</td>
<td id="76F7223D3016006FE808FB2EFCDE6E42" box="[634,788,1254,1349]" gridcol="2" gridrow="9" pageId="3" pageNumber="251">cup like, clearly wider than ductus bursae</td>
<td id="76F7223D3016006FE940FB2EFB906E42" box="[818,1114,1254,1349]" gridcol="3" gridrow="9" pageId="3" pageNumber="251">elongated, sclerotized</td>
<td id="76F7223D3016006FEE0EFB2EFA876E42" box="[1148,1357,1254,1349]" gridcol="4" gridrow="9" pageId="3" pageNumber="251">cup-shaped, membranous</td>
<td id="76F7223D3016006FEF0EFB2EF9966E42" box="[1404,1628,1254,1349]" gridcol="5" gridrow="9" pageId="3" pageNumber="251">initially narrow, then strongly distended and round</td>
<td id="76F7223D3016006FECF5FB2EF8176E42" box="[1671,2013,1254,1349]" gridcol="6" gridrow="9" pageId="3" pageNumber="251">narrow and elongated, cup-shaped (Fig. 7B)</td>
</tr>
<tr id="35264B413016006FEAE5FA98F8176E8C" box="[151,2013,1360,1419]" gridrow="10" pageId="3" pageNumber="251" rowspan-2="1">
<th id="76F7223D3016006FEAE5FA98FEB96E8C" box="[151,371,1360,1419]" gridcol="0" gridrow="10" pageId="3" pageNumber="251">ductus bursae below antrum</th>
<td id="76F7223D3016006FEBFEFA98FD9C6E8C" box="[396,598,1360,1419]" gridcol="1" gridrow="10" pageId="3" pageNumber="251">sclerotized</td>
<td id="76F7223D3016006FE940FA98FB906E8C" box="[818,1114,1360,1419]" gridcol="3" gridrow="10" pageId="3" pageNumber="251">membranous</td>
<td id="76F7223D3016006FEE0EFA98FA876E8C" box="[1148,1357,1360,1419]" gridcol="4" gridrow="10" pageId="3" pageNumber="251">sclerotized</td>
<td id="76F7223D3016006FEF0EFA98F9966E8C" box="[1404,1628,1360,1419]" gridcol="5" gridrow="10" pageId="3" pageNumber="251">only short section sclerotized</td>
<td id="76F7223D3016006FECF5FA98F8176E8C" box="[1671,2013,1360,1419]" gridcol="6" gridrow="10" pageId="3" pageNumber="251">dilated, markedly sclerotized (Fig. 7A)</td>
</tr>
<tr id="35264B413016006FEAE5FA65F8176EEF" box="[151,2013,1453,1512]" gridrow="11" pageId="3" pageNumber="251">
<th id="76F7223D3016006FEAE5FA65FEB96EEF" box="[151,371,1453,1512]" gridcol="0" gridrow="11" pageId="3" pageNumber="251">ductus bursae</th>
<td id="76F7223D3016006FEBFEFA65FD9C6EEF" box="[396,598,1453,1512]" gridcol="1" gridrow="11" pageId="3" pageNumber="251">short, strongly sclerotized, very thin</td>
<td id="76F7223D3016006FE808FA65FCDE6EEF" box="[634,788,1453,1512]" gridcol="2" gridrow="11" pageId="3" pageNumber="251">very short, sclerotized</td>
<td id="76F7223D3016006FE940FA65FB906EEF" box="[818,1114,1453,1512]" gridcol="3" gridrow="11" pageId="3" pageNumber="251">very short, strengthened at the antrum</td>
<td id="76F7223D3016006FEE0EFA65FA876EEF" box="[1148,1357,1453,1512]" gridcol="4" gridrow="11" pageId="3" pageNumber="251">not very long, membranous</td>
<td id="76F7223D3016006FEF0EFA65F9966EEF" box="[1404,1628,1453,1512]" gridcol="5" gridrow="11" pageId="3" pageNumber="251">long, membranous</td>
<td id="76F7223D3016006FECF5FA65F8176EEF" box="[1671,2013,1453,1512]" gridcol="6" gridrow="11" pageId="3" pageNumber="251">membranous (Fig. 7A)</td>
</tr>
<tr id="35264B413016006FEAE5FA3BF8176D0D" box="[151,2013,1523,1546]" gridrow="12" pageId="3" pageNumber="251">
<th id="76F7223D3016006FEAE5FA3BFEB96D0D" box="[151,371,1523,1546]" gridcol="0" gridrow="12" pageId="3" pageNumber="251">bursa copulatrix</th>
<td id="76F7223D3016006FEBFEFA3BFD9C6D0D" box="[396,598,1523,1546]" gridcol="1" gridrow="12" pageId="3" pageNumber="251">elongate, small,</td>
<td id="76F7223D3016006FE808FA3BFCDE6D0D" box="[634,788,1523,1546]" gridcol="2" gridrow="12" pageId="3" pageNumber="251">elongate, long,</td>
<td id="76F7223D3016006FE940FA3BFB906D0D" box="[818,1114,1523,1546]" gridcol="3" gridrow="12" pageId="3" pageNumber="251">elongated, irregular</td>
<td id="76F7223D3016006FEE0EFA3BFA876D0D" box="[1148,1357,1523,1546]" gridcol="4" gridrow="12" pageId="3" pageNumber="251">small, oval,</td>
<td id="76F7223D3016006FEF0EFA3BF9966D0D" box="[1404,1628,1523,1546]" gridcol="5" gridrow="12" pageId="3" pageNumber="251">large, oval</td>
<td id="76F7223D3016006FECF5FA3BF8176D0D" box="[1671,2013,1523,1546]" gridcol="6" gridrow="12" pageId="3" pageNumber="251">small, round</td>
</tr>
<tr id="35264B413016006FEAE5F9DEF8176D72" box="[151,2013,1558,1653]" gridrow="13" pageId="3" pageNumber="251" rowspan-2="1" rowspan-3="1">
<th id="76F7223D3016006FEAE5F9DEFEB96D72" box="[151,371,1558,1653]" gridcol="0" gridrow="13" pageId="3" pageNumber="251">strengthenedd walls of bursa copulatrix</th>
<td id="76F7223D3016006FEBFEF9DEFD9C6D72" box="[396,598,1558,1653]" gridcol="1" gridrow="13" pageId="3" pageNumber="251">membranous</td>
<td id="76F7223D3016006FEE0EF9DEFA876D72" box="[1148,1357,1558,1653]" gridcol="4" gridrow="13" pageId="3" pageNumber="251">sclerotised, very numerous spines, tiny and short</td>
<td id="76F7223D3016006FEF0EF9DEF9966D72" box="[1404,1628,1558,1653]" gridcol="5" gridrow="13" pageId="3" pageNumber="251">membranous, with small teeth or strongly sclerotized</td>
<td id="76F7223D3016006FECF5F9DEF8176D72" box="[1671,2013,1558,1653]" gridcol="6" gridrow="13" pageId="3" pageNumber="251">membranous, with small teeth (Fig. 7A)</td>
</tr>
<tr id="35264B413016006FEAE5F949F8176D9F" box="[151,2013,1665,1688]" gridrow="14" pageId="3" pageNumber="251" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76F7223D3016006FEAE5F949FEB96D9F" box="[151,371,1665,1688]" gridcol="0" gridrow="14" pageId="3" pageNumber="251">signum</th>
<td id="76F7223D3016006FEBFEF949FD9C6D9F" box="[396,598,1665,1688]" gridcol="1" gridrow="14" pageId="3" pageNumber="251">absent</td>
<td id="76F7223D3016006FEF0EF949F9966D9F" box="[1404,1628,1665,1688]" gridcol="5" gridrow="14" pageId="3" pageNumber="251">present or absent</td>
<td id="76F7223D3016006FECF5F949F8176D9F" box="[1671,2013,1665,1688]" gridcol="6" gridrow="14" pageId="3" pageNumber="251">absent</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA949033011FF94EABCFE13FE4169CA" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="252">
Female genitalia (prep. gen. no. YPO 20/2016) (
<figureCitation id="132D55863011FF94E89FFE13FC8C6AF2" box="[749,838,475,501]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[235,1353,187,1913]" captionTargetId="figure-26@9.[229,1359,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Pararhigognostis stichocentra female genitalia, gen. slide: YPO 20/2016: A, general view, B, antrum." figureDoi="http://doi.org/10.5281/zenodo.14033558" httpUri="https://zenodo.org/record/14033558/files/figure.png" pageId="4" pageNumber="252">Fig. 7A</figureCitation>
), with labii long, triangular, covered with very small, scanty, hair-like bristles; apophyses posteriores very thin, longer than anteriores; apophyses anteriores markedly wider; antrum narrow and elongate, with gently rounded margins, broadening terminally (
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); ductus bursae short, membranous, narrow anterior of antrum, slightly broader and terminally constricted, to inception of ductus seminalis short section of ductus (
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), to inception of bursa copulatrix, slightly dilated; bursa copulatrix small, oval, membranous, with very small sclerites with small teeth on walls of bursa; signum absent; ductus seminalis narrow, membranous.
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The name of the new genus is derived from
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,
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proposed by
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, with the addition of the prefix “para.”
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The newly described genus is represented by one species presently known from several localities in Africa.
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5.
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depository: KRAM. References: present paper (description).
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†Genus
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<emphasis id="B929B23CFFC06663FF3A69B6BA3C5216" bold="true" box="[253,375,1015,1041]" italics="true" pageId="32" pageNumber="233">Damzenia</emphasis>
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6.
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References:
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Comment: Placement of the genus in the tribe
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Genus
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14.
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depository: KRAM. References:
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Comment: Generic placement of the species is somewhat uncertain because ventral side of the
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15.
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depository: CVIA. References:
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<mods:affiliation id="8E149314959AA3437C1EDD35DE6C05C8">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia. &amp; Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. &amp; Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; &amp; Department of Ecology &amp; Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
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Genus
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Motschulsky, 1845
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species:
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<emphasis id="B929B23CFFE26641FEE1698EBA8155E0" box="[294,458,975,999]" italics="true" pageId="2" pageNumber="203">Usechus lacerta</emphasis>
Motschulsky, 1845
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<emphasis id="B929B23CFFE26641FF506E54BAE55228" bold="true" box="[151,430,1045,1071]" pageId="2" pageNumber="203">Taxonomic assignment.</emphasis>
The studied inclusion is placed in the subfamily
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within the family
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based on the combination of the following characters:(1) antennae 11-segmented with 3-segmented club; (2) antennal insertions concealed under lateral expansion of frons; (3) 5-5-4 tarsal formula; (4) abdominal ventrite 1 slightly longer than ventrite 2; and (5) intercoxal process of abdominal ventrite 1 arcuate apically. The fossil specimen is assigned to the tribe
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based on the unique character of the group, namely possessing deep anterolateral antennal cavities on the pronotum. Externally closed procoxal cavities and pronotal disc without transverse basal groove suggest placement in the genus
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<emphasis id="B929B23CFFE26641FD956EACB9FA5300" box="[594,689,1261,1287]" italics="true" pageId="2" pageNumber="203">Usechus</emphasis>
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. Despite several characters of ventral side not being discernible in the beetle under study, we place the new species in
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based on the presence of the above-listed set of external characters.
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<emphasis id="B929B23CFFE26641FF506FE0BA8153BB" bold="true" box="[151,458,1441,1468]" italics="true" pageId="2" pageNumber="203">Usechus andrushchenkoi</emphasis>
Alekseev et Bukejs
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(
<figureCitation id="136672ABFFE26641FF586F84BA4853D8" box="[159,259,1477,1503]" captionStart-0="FIGURES 12" captionStart-1="FIGURES 34" captionStartId-0="4.[151,264,1870,1894]" captionStartId-1="5.[151,264,1852,1876]" captionTargetBox-0="[151,1436,486,1844]" captionTargetBox-1="[151,1435,199,1826]" captionTargetId-0="figure-165@4.[151,1436,486,1844]" captionTargetId-1="figure-17@5.[151,1435,179,1826]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 12. Usechus andrushchenkoi sp. nov., holotype, No. KAM 8820 [KRAM]: 1—habitus, dorsal view; 2—habitus, ventral view. Scale bars = 1.0 mm." captionText-1="FIGURES 34. Usechus andrushchenkoi sp. nov., holotype, No. KAM 8820 [KRAM]: 3—schematic drawing of antenna; 4—anterior part of body, dorsal view. Scale bar = 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14033620" figureDoi-1="http://doi.org/10.5281/zenodo.14033622" httpUri-0="https://zenodo.org/record/14033620/files/figure.png" httpUri-1="https://zenodo.org/record/14033622/files/figure.png" pageId="2" pageNumber="203">Figs 14</figureCitation>
)
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<emphasis id="B929B23CFFE26641FF506C4CBA0D5020" bold="true" box="[151,326,1549,1575]" pageId="2" pageNumber="203">Type material.</emphasis>
<typeStatus id="54E6D08CFFE26641FE896C4CBAF05020" box="[334,443,1549,1575]" pageId="2" pageNumber="203" type="holotype">Holotype</typeStatus>
: No. KAM 8820 [KRAM]; “*
<typeStatus id="54E6D08CFFE26641FCC96C4CB8375020" box="[782,892,1549,1575]" pageId="2" pageNumber="203" type="holotype">Holotype</typeStatus>
/
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</taxonomicName>
/
<taxonomicName id="4C5D15ADFFE26641FBCC6C4CBFF55020" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[1035,1214,1549,1575]" class="Insecta" family="Zopheridae" genus="Usechus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="andrushchenkoi" status="sp. nov.">
<emphasis id="B929B23CFFE26641FBCC6C4CBFF55020" box="[1035,1214,1549,1575]" italics="true" pageId="2" pageNumber="203">andrushchenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A0F47FFE26641FB016C4FBE55502F" box="[1222,1310,1550,1576]" pageId="2" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
/ Alekseev et Bukejs des. 2024” [red handwritten label]; adult, sex unknown; submentum and last abdominal ventrite not observable in the specimen. A complete beetle is included in a parallelepipedic, transparent, yellow amber piece with dimensions of 23×
<quantity id="4CA5C3CBFFE26641FE656C38BABB5093" box="[418,496,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="203" unit="mm" value="15.0">15 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFE26641FC866C38B8CF5093" box="[833,900,1657,1684]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="203" unit="mm" value="6.0">6 mm</quantity>
, preserved without supplementary fixation. The amber piece with the studied specimen was accidentally cracked during mechanical processing and then glued together. As a result, the state of preservation and visibility of the inclusion lying at the junction of the amber layers deteriorated: the cuticle was destructed in some places (antennomeres), and air bubbles appeared in cracks and on the surface of the inclusion in some places (head). Ventral side of specimen is obscured by “milky” opacity. Syninclusions: a few small stellate
<taxonomicName id="4C5D15ADFFE26641FDE56D6FB9C1514F" authorityName="Dumortier" authorityYear="1829" box="[546,650,1838,1864]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="203" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
trichomes.
</paragraph>
<paragraph id="8BE26E2EFFE26641FF006D10BAEF5188" blockId="2.[151,1437,1549,2007]" pageId="2" pageNumber="203">
<emphasis id="B929B23CFFE26641FF006D10BA38516C" bold="true" box="[199,371,1873,1899]" pageId="2" pageNumber="203">
<typeStatus id="54E6D08CFFE26641FF006D10BA49516C" box="[199,258,1873,1899]" pageId="2" pageNumber="203">Type</typeStatus>
stratum.
</emphasis>
Baltic amber from Eocene amber-bearing Blaue Erde deposits; estimated age: middlelate Eocene (Standke 1998).
</paragraph>
<paragraph id="8BE26E2EFFE26641FF006DD8BBA251D0" blockId="2.[151,1437,1549,2007]" pageId="2" pageNumber="203">
<materialsCitation id="3B356473FFE26641FF006DD8BBAE51D0" country="Russia" county="Peninsula" location="Yantarny" municipality="Sambian" pageId="2" pageNumber="203" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B929B23CFFE26641FF006DD8BA2E51B4" bold="true" box="[199,357,1945,1971]" pageId="2" pageNumber="203">
<typeStatus id="54E6D08CFFE26641FF006DD8BA4951B4" box="[199,258,1945,1971]" pageId="2" pageNumber="203">Type</typeStatus>
locality.
</emphasis>
<location id="8E8238F5FFE26641FEAD6DDBBA8451B3" LSID="urn:lsid:plazi:treatment:03F4DF38FFE26647FF506FE0B8A057D6:8E8238F5FFE26641FEAD6DDBBA8451B3" box="[362,463,1946,1972]" country="Russia" county="Peninsula" municipality="Sambian" name="Yantarny" pageId="2" pageNumber="203" stateProvince="Kaliningrad">Yantarny</location>
settlement (formerly Palmnicken),
<collectingMunicipality id="6B86F454FFE26641FC9C6DD8B8F651B4" box="[859,957,1945,1971]" pageId="2" pageNumber="203">Sambian</collectingMunicipality>
(Samland)
<collectingCounty id="628316A2FFE26641FBF96DD8BFE551B4" box="[1086,1198,1945,1971]" pageId="2" pageNumber="203">Peninsula</collectingCounty>
,
<collectingRegion id="4999A0CCFFE26641FB7F6DD8BED351B4" box="[1208,1432,1945,1971]" country="Russia" name="Kaliningrad" pageId="2" pageNumber="203">Kaliningrad Region</collectingRegion>
,
<collectingCountry id="F34A2EBEFFE26641FF506DFCBBAE51D0" box="[151,229,1981,2007]" name="Russia" pageId="2" pageNumber="203">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE36640FF006AD7B9FA539A" pageId="3" pageNumber="204" type="description">
<paragraph id="8BE26E2EFFE36640FF006AD7B80756FE" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">
<emphasis id="B929B23CFFE36640FF006AD7BA1A56B7" bold="true" box="[199,337,150,176]" pageId="3" pageNumber="204">Description</emphasis>
. Measurements: total body length
<quantity id="4CA5C3CBFFE36640FD126AD6B86656B6" box="[725,813,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="204" unit="mm" value="4.1">4.1 mm</quantity>
, maximum body width (across elytra)
<quantity id="4CA5C3CBFFE36640FB1C6AD6BE7F56B6" box="[1243,1332,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="204" unit="mm" value="1.4">1.4 mm</quantity>
; pronotal length
<quantity id="4CA5C3CBFFE36640FF216AFABA0B56D2" box="[230,320,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="204" unit="mm" value="0.9">0.9 mm</quantity>
, maximum pronotal width
<quantity id="4CA5C3CBFFE36640FDBF6AFAB99956D2" box="[632,722,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="204" unit="mm" value="1.3">1.3 mm</quantity>
, basal pronotal width 1.0 mm; elytral length (along the elytral suture, including scutellum)
<quantity id="4CA5C3CBFFE36640FE1F6A9EB97A56FE" box="[472,561,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="204" unit="mm" value="2.9">2.9 mm</quantity>
; antennal length
<quantity id="4CA5C3CBFFE36640FD286A9EB80356FE" box="[751,840,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="204" unit="mm" value="0.6">0.6 mm</quantity>
.
</paragraph>
<paragraph id="8BE26E2EFFE36640FF006B42BE1B5762" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">Body subparallel-sided, elongate, 2.9× longer than wide; weakly convex dorsally and ventrally; unicolorous black (as preserved). Pubescence: pronotum covered with decumbent, narrow, elongate setae directed anteriad and laterad; elytra homogenously covered with shorter, decumbent, narrow, elongate setae, not forming patches.</paragraph>
<paragraph id="8BE26E2EFFE36640FF006B2EB9595482" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">Head partially retracted in pronotum, transverse, widest across eyes, not constricted behind eyes, and without temples. Compound eyes vertical, reniform (emarginate anteriorly at antennal base), flat, not visible in dorsal view, apparently without interfacetal setation. Antennal insertions concealed dorsally by lateral expansion of frons; antennal grooves ventral to eye absent. Antennae short, stout, extending to about middle of pronotum; 11-segmented with 3-segmented weakly flattened club; scape short, subquadrate; pedicel subcylindrical, longest; antennomeres 36 as long as wide, rounded; antennomeres 78 weakly transverse; antennomeres 910 widest, subtrapezoidal, transverse; antennomere 10 shorter than antennomere 9, about 2× as wide as long; antennomere 11 round and weakly flattened, as long as wide.</paragraph>
<paragraph id="8BE26E2EFFE36640FF0068CEBF2F55A2" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">Pronotum nearly hexagonal, transverse, 1.4× as wide as long; deep anterolateral antennal cavities extend to middle of pronotal length and end in round fossa; widest point at middle, coinciding with endpoint of antennal cavities; distinctly narrowed anteriad and posteriad, with maximum pronotal width / basal pronotal width = 1.3; basally bordered. Pronotal disc convex medially, without transverse basal groove; lateral sides explanate, bordered. Pronotal surface coarsely granulate. Anterior and posterior pronotal angles obtuse. Prothoracic notosternal suture complete. Hypomeron densely granulose. Intercoxal prosternal process elonagte, distinctly widened apically and forming posterolateral projections extending to posterior margin of procoxae. Procoxal cavity closed posteriorly by projection of hypomeron and posterolateral projection of intercoxal prosternal process.</paragraph>
<paragraph id="8BE26E2EFFE36640FF0069EEB89255CE" blockId="3.[151,1437,150,2013]" box="[199,985,943,969]" pageId="3" pageNumber="204">Scutellar shield small, cordate, transverse, about 2.0× as wide as long.</paragraph>
<paragraph id="8BE26E2EFFE36640FF006992BF53525E" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">Elytra almost parallel-sided within anterior two-thirds of their length, tapered at apex, convex, 2.1× as long as wide combined, and 3.2× longer than pronotum. Humeral angles rounded. Elytra striate-punctate. Scutellary striole not apparent. Interstriae convex, without tubercles. Epipleura well-developed, widest in basal half, slightly narrowing posteriorly, reaching elytral apices. Metathoracic wings not apparent.</paragraph>
<paragraph id="8BE26E2EFFE36640FF006E22B8E5530A" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">Legs short, rather robust. Procoxa nearly rounded, narrowly separated; metacoxae narrowly oval, transverse, separated by about one longitudinal diameter of metacoxa. Pro- and mesofemora nearly spindle-shaped, slightly widened; metafemora weakly curved; femora and tibiae subequal in length. Tibiae with fringe of setae apically. Tarsal formula 5-5-4; tarsomeres with fine setation ventrally; metatarsomere 4 longest, as long as metatarsomeres 23 combined. Claws simple, large, about 0.3× as long as tarsomere 4.</paragraph>
<paragraph id="8BE26E2EFFE36640FF006F56B9FA539A" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">Abdomen with five visible ventrites, abdominal sutures distinct throughout length. Relative length ratios of ventrites 15 equal to 7:6:4:3:3. Setation and punctation of ventrites not apparent (possibly absent). Abdominal ventrite 1 with apically rounded intercoxal process in form of equilateral triangle, about as wide as metacoxal length. Ventrite 5 with arcuate posterior margin.</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE36640FF006FE6BE47500E" pageId="3" pageNumber="204" type="discussion">
<paragraph id="8BE26E2EFFE36640FF006FE6BE47500E" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">
<emphasis id="B929B23CFFE36640FF006FE6BA6253C6" bold="true" box="[199,297,1447,1473]" pageId="3" pageNumber="204">Remark</emphasis>
. The ventral side of the specimen (mesoventrite, metaventite, abdominal ventrites) is obscured by milky opacity making these areas indistinct or insufficiently visible. The closure of mesocoxal cavities (possibly closed) and the presence of impression on abdominal ventrite 5 (=preapical groove) remain unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE36640FF006C53B81150E6" pageId="3" pageNumber="204" type="diagnosis">
<paragraph id="8BE26E2EFFE36640FF006C53B824509E" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">
<emphasis id="B929B23CFFE36640FF006C53BA85502B" bold="true" box="[199,462,1554,1580]" pageId="3" pageNumber="204">Differential diagnosis.</emphasis>
<taxonomicName id="4C5D15ADFFE36640FE126C52B9A5502A" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[469,750,1555,1581]" class="Insecta" family="Zopheridae" genus="Usechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="andrushchenkoi" status="sp. nov.">
<emphasis id="B929B23CFFE36640FE126C52B9A5502A" box="[469,750,1555,1581]" italics="true" pageId="3" pageNumber="204">Usechus andrushchenkoi</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFE36640FD326C52B805502A" bold="true" box="[757,846,1555,1581]" pageId="3" pageNumber="204">
<taxonomicNameLabel id="A21A0F47FFE36640FD326C52B805502A" box="[757,846,1555,1581]" pageId="3" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all extant congeners in the combination of the following characters: relatively long elytra, about 2.1× as long as wide; weakly developed elytral relief (alternate intervals not costate, tubercles absent); transverse pronotum, 1.4× as wide as long; distinct scutellar shield; and long pedicel, as long as antennomeres 35 combined.
</paragraph>
<paragraph id="8BE26E2EFFE36640FF006CE2B81150E6" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">
The new species can be distinguished from all other described Baltic amber
<taxonomicName id="4C5D15ADFFE36640FBC96CE2BFC550BA" authorityName="Solier" authorityYear="1834" box="[1038,1166,1699,1725]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="family">Zopheridae</taxonomicName>
by the nearly hexagonal pronotum and the deep anterolateral pronotal antennal cavities.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE36640FF006CABB8B7514A" pageId="3" pageNumber="204" type="etymology">
<paragraph id="8BE26E2EFFE36640FF006CABB8B7514A" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="204">
<emphasis id="B929B23CFFE36640FF006CABBAE85103" bold="true" box="[199,419,1770,1796]" pageId="3" pageNumber="204">Derivatio nominis</emphasis>
. Patronymic, the name of new species is dedicated to Konstantin V. Andrushchenko (
<collectingRegion id="4999A0CCFFE36640FF586D4EBA60512E" box="[159,299,1807,1833]" country="Russia" name="Kaliningrad" pageId="3" pageNumber="204">Kaliningrad</collectingRegion>
), a great enthusiast in collecting Baltic amber insect inclusions, who provided the
<typeStatus id="54E6D08CFFE36640FB166D4EBE4A512E" box="[1233,1281,1807,1833]" pageId="3" pageNumber="204">type</typeStatus>
specimen for description and donated the amber from his private collection to the museum.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE36647FF006D17BF9C5746" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204" type="distribution">
<paragraph id="8BE26E2EFFE36647FF006D17BF9C5746" blockId="3.[151,1437,150,2013]" lastBlockId="4.[151,1437,151,465]" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204">
<emphasis id="B929B23CFFE36640FF006D17BABF5176" bold="true" box="[199,500,1878,1905]" pageId="3" pageNumber="204">Distribution of congeners</emphasis>
. The genus
<taxonomicName id="4C5D15ADFFE36640FDBE6D16B9935176" authorityName="Motschulsky" authorityYear="1845" box="[633,728,1879,1905]" class="Insecta" family="Zopheridae" genus="Usechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFE36640FDBE6D16B9935176" box="[633,728,1879,1905]" italics="true" pageId="3" pageNumber="204">Usechus</emphasis>
</taxonomicName>
has a disjunct Recent distribution (
<bibRefCitation id="EFCC13DFFFE36640FB9D6D16BE4B5176" author="Blaisdell, F. E." box="[1114,1280,1879,1905]" pageId="3" pageNumber="204" pagination="1 - 14" refId="ref24659" refString="Blaisdell, F. E. (1929) A revision of the beetles of the tenebrionid tribe Usechini, with descriptions of a new genus and new species. Proceedings of the United States National Museum, 75, 1 - 14. https: // doi. org / 10.5479 / si. 00963801.75 - 2790.1" type="journal article" year="1929">Blaisdell 1929</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFE36640FACC6D16BEDC5176" author="Kulzer, H." box="[1291,1431,1879,1905]" pageId="3" pageNumber="204" pagination="304 - 317" refId="ref26358" refString="Kulzer, H. (1960) Einige neue Tenebrioniden (Col.) (20. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum G. Frey, 11, 304 - 317." type="journal article" year="1960">Kulzer 1960</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFE36640FF506D3ABA5F5192" author="Saito, M." box="[151,276,1915,1941]" pageId="3" pageNumber="204" pagination="103 - 111" refId="ref27732" refString="Saito, M. (1999) Notes on the Japanese species of the genus Usechus (Coleoptera, Zopheridae). Elytra, 27, 103 - 111." type="journal article" year="1999">Saitô 1999</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFE36640FEE66D3DB9175192" author="Slipinski, S. A. &amp; Lawrence, J. F." box="[289,604,1915,1943]" pageId="3" pageNumber="204" pagination="1 - 53" refId="ref27981" refString="Slipinski, S. A. &amp; Lawrence, J. F. (1999) Phylogeny and classification of Zopheridae sensu novo (Coleoptera: Tenebrionoidea) with a review of the genera of Zopherinae (excluding Monommatini). Annales Zoologici, 49, 1 - 53." type="journal article" year="1999">Ślipiński &amp; Lawrence 1999</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFE36640FDAE6D3AB8785192" author="Kim, Y-H. &amp; Ahn, K-J." box="[617,819,1915,1941]" pageId="3" pageNumber="204" pagination="201 - 203" refId="ref26190" refString="Kim, Y-H. &amp; Ahn, K-J. (2009) Usechus chujoi New to Korea (Coleoptera: Zopheridae: Zopherinae). The Korean Society of Systematic Zoology, 25 (2), 201 - 203. https: // doi. org / 10.5635 / KJSZ. 2009.25.2.201" type="journal article" year="2009">Kim &amp; Ahn 2009</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFE36640FC876D3AB8B75192" author="Jung, B. H. &amp; Seung, J. &amp; Park, Y. - G." box="[832,1020,1915,1941]" pageId="3" pageNumber="204" pagination="446 - 449" refId="ref26069" refString="Jung, B. H., Seung, J. &amp; Park, Y. - G. (2023) Three species of the family Zopheridae (Coleoptera) new to Korea. Journal of Asia- Pacific Biodiversity, 16 (3), 446 - 449. https: // doi. org / 10.1016 / j. japb. 2022.11.005" type="journal article" year="2023">
Jung
<emphasis id="B929B23CFFE36640FCBA6D3AB8FB5192" box="[893,944,1915,1941]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2023
</bibRefCitation>
). The group occurs in western North America (2 species:
<taxonomicName id="4C5D15ADFFE36640FE416DDEB99051BE" authority="Motschulsky, 1845" authorityName="Motschulsky" authorityYear="1845" box="[390,731,1950,1977]" class="Insecta" family="Zopheridae" genus="Usechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="lacerta">
<emphasis id="B929B23CFFE36640FE416DDEBAB151BE" box="[390,506,1951,1977]" italics="true" pageId="3" pageNumber="204">U. lacerta</emphasis>
Motschulsky, 1845
</taxonomicName>
—California;
<taxonomicName id="4C5D15ADFFE36640FCBB6DDEBFEA51BE" authority="Casey, 1889" authorityName="Casey" authorityYear="1889" box="[892,1185,1951,1977]" class="Insecta" family="Zopheridae" genus="Usechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="nucleatus">
<emphasis id="B929B23CFFE36640FCBB6DDEBF4751BE" box="[892,1036,1951,1977]" italics="true" pageId="3" pageNumber="204">U. nucleatus</emphasis>
Casey, 1889
</taxonomicName>
—California, Oregon, Washington) and in
<collectingCountry id="F34A2EBEFFE36640FEBD6D82BAA851DA" box="[378,483,1987,2013]" name="China" pageId="3" pageNumber="204">East Asia</collectingCountry>
(4 species:
<taxonomicName id="4C5D15ADFFE36640FDAD6D82B82051DA" authority="Kulzer, 1960" authorityName="Kulzer" authorityYear="1960" box="[618,875,1987,2013]" class="Insecta" family="Zopheridae" genus="Usechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="chujoi">
<emphasis id="B929B23CFFE36640FDAD6D82B99851DA" box="[618,723,1987,2013]" italics="true" pageId="3" pageNumber="204">U. chujoi</emphasis>
<bibRefCitation id="EFCC13DFFFE36640FD1D6D82B82051DA" author="Kulzer, H." box="[730,875,1987,2013]" pageId="3" pageNumber="204" pagination="304 - 317" refId="ref26358" refString="Kulzer, H. (1960) Einige neue Tenebrioniden (Col.) (20. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum G. Frey, 11, 304 - 317." type="journal article" year="1960">Kulzer, 1960</bibRefCitation>
</taxonomicName>
<collectingCountry id="F34A2EBEFFE36640FC4F6D82B88151DA" box="[904,970,1987,2013]" name="Japan" pageId="3" pageNumber="204">Japan</collectingCountry>
,
<collectingCountry id="F34A2EBEFFE36640FC116D82BF5551DA" box="[982,1054,1987,2013]" name="South Korea" pageId="3" pageNumber="204">Korea</collectingCountry>
;
<taxonomicName id="4C5D15ADFFE36640FBEE6D82BE3551DA" authority="Kamiya, 1963" authorityName="Kamiya" authorityYear="1963" box="[1065,1406,1987,2013]" class="Insecta" family="Zopheridae" genus="Usechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="tsushimensis">
<emphasis id="B929B23CFFE36640FBEE6D82BF9151DA" box="[1065,1242,1987,2013]" italics="true" pageId="3" pageNumber="204">U. tsushimensis</emphasis>
Kamiya, 1963
</taxonomicName>
<collectingCountry id="F34A2EBEFFE46647FF506AD6BB9156B6" box="[151,218,151,177]" name="Japan" pageId="4" pageNumber="205">Japan</collectingCountry>
,
<collectingCountry id="F34A2EBEFFE46647FF216AD6BA6556B6" box="[230,302,151,177]" name="South Korea" pageId="4" pageNumber="205">Korea</collectingCountry>
;
<taxonomicName id="4C5D15ADFFE46647FEFC6AD6B92256B6" authority="Sasaji, 1987" authorityName="Sasaji" authorityYear="1987" box="[315,617,151,177]" class="Insecta" family="Zopheridae" genus="Usechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ohdaiensis">
<emphasis id="B929B23CFFE46647FEFC6AD6BA9C56B6" box="[315,471,151,177]" italics="true" pageId="4" pageNumber="205">U. ohdaiensis</emphasis>
Sasaji, 1987
</taxonomicName>
<collectingCountry id="F34A2EBEFFE46647FD416AD6B98256B6" box="[646,713,151,177]" name="Japan" pageId="4" pageNumber="205">Japan</collectingCountry>
;
<taxonomicName id="4C5D15ADFFE46647FD116AD6B88756B6" authority="Saito, 1999" authorityName="Saito" authorityYear="1999" box="[726,972,151,177]" class="Insecta" family="Zopheridae" genus="Usechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="sasajii">
<emphasis id="B929B23CFFE46647FD116AD6B80956B7" box="[726,834,151,176]" italics="true" pageId="4" pageNumber="205">U. sasajii</emphasis>
<bibRefCitation id="EFCC13DFFFE46647FC8C6AD6B88756B6" author="Saito, M." box="[843,972,151,177]" pageId="4" pageNumber="205" pagination="103 - 111" refId="ref27732" refString="Saito, M. (1999) Notes on the Japanese species of the genus Usechus (Coleoptera, Zopheridae). Elytra, 27, 103 - 111." type="journal article" year="1999">Saito, 1999</bibRefCitation>
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<collectingCountry id="F34A2EBEFFE46647FC2D6AD6BF6656B6" box="[1002,1069,151,177]" name="Japan" pageId="4" pageNumber="205">Japan</collectingCountry>
). The fossil species described in the current paper establishes that the tribe
<taxonomicName id="4C5D15ADFFE46647FDB66AFAB99C56D2" authorityName="Horn" authorityYear="1867" box="[625,727,187,213]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="tribe" tribe="Usechini">Usechini</taxonomicName>
, which restricted to the present-day Holarctic realm, dates to at least the Eocene. It also confirms the presence of the tribe in the Western Palearctic in the Paleogene. The Eastern Palearctic, north-eastern Oriental, Western and Eastern Nearctic Regions are generally regarded as typical refuge areas for Eocene Laurasian relicts (e.g.
<bibRefCitation id="EFCC13DFFFE46647FD966B66B9AE5746" author="Andree, K." box="[593,741,294,321]" pageId="4" pageNumber="205" refId="ref24368" refString="Andree, K. (1951) Der Bernstein. Das Bernsteinland und sein Leben. Kosmos, Francksche Verhandlung, Stuttgart, 95 pp." type="book" year="1951">Andrée 1951</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFE46647FD366B66B8C75746" author="Larsson, S. G." box="[753,908,295,321]" pageId="4" pageNumber="205" refId="ref26397" refString="Larsson, S. G. (1978) Baltic Amber - a palaeobiological study. Vol. 1. Scandinavian Science Press Ltd., Klampenborg, 192 pp. https: // doi. org / 10.1163 / 9789004631243" type="book" year="1978">Larsson 1978</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFE46647FC506B66BF815746" author="Weitschat, W. &amp; Wichard, W." box="[919,1226,295,321]" pageId="4" pageNumber="205" refId="ref28436" refString="Weitschat, W. &amp; Wichard, W. (1998) Atlas der Pflanzen und Tiere im baltischen Bernstein. Pfeil, Munchen, 256 pp." type="book" year="1998">Weitschat &amp; Wichard 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
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<emphasis id="B929B23CFFE46647FF006B0ABA8E5762" bold="true" box="[199,453,330,357]" pageId="4" pageNumber="205">Ecology of congeners.</emphasis>
Recent representatives of
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<emphasis id="B929B23CFFE46647FD236B0AB8085762" box="[740,835,331,357]" italics="true" pageId="4" pageNumber="205">Usechus</emphasis>
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occur under the bark and in dead, rotting wood infested by fungi. They have been reported from deciduous (
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<emphasis id="B929B23CFFE46647FD276B2EB85E578E" box="[736,789,367,393]" italics="true" pageId="4" pageNumber="205">Acer</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5D15ADFFE46647FCE46B2EB8CA578E" box="[803,897,367,393]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="205" phylum="Tracheophyta" rank="genus">
<emphasis id="B929B23CFFE46647FCE46B2EB8CA578E" box="[803,897,367,393]" italics="true" pageId="4" pageNumber="205">Quercus</emphasis>
</taxonomicName>
) and coniferous trees (
<bibRefCitation id="EFCC13DFFFE46647FB466B2EBE62578E" author="Blaisdell, F. E." box="[1153,1321,367,393]" pageId="4" pageNumber="205" pagination="1 - 14" refId="ref24659" refString="Blaisdell, F. E. (1929) A revision of the beetles of the tenebrionid tribe Usechini, with descriptions of a new genus and new species. Proceedings of the United States National Museum, 75, 1 - 14. https: // doi. org / 10.5479 / si. 00963801.75 - 2790.1" type="journal article" year="1929">Blaisdell 1929</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFE46647FAF26B2EBA0257AA" author="Doyen, J. T. &amp; Lawrence, J. F." pageId="4" pageNumber="205" pagination="333 - 377" refId="ref25548" refString="Doyen, J. T. &amp; Lawrence, J. F. (1979) Relationships and higher classification of some Tenebrionidae and Zopheridae (Coleoptera). Systematic Entomology, 4, 333 - 377. https: // doi. org / 10.1111 / j. 1365 - 3113.1979. tb 00619. x" type="journal article" year="1979">Doyen &amp; Lawrence 1979</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFE46647FE916BD2BA9857AA" author="Saito, M." box="[342,467,403,429]" pageId="4" pageNumber="205" pagination="103 - 111" refId="ref27732" refString="Saito, M. (1999) Notes on the Japanese species of the genus Usechus (Coleoptera, Zopheridae). Elytra, 27, 103 - 111." type="journal article" year="1999">Saitô 1999</bibRefCitation>
). The Eocene
<taxonomicName id="4C5D15ADFFE46647FDB26BD2B99F57AA" authorityName="Motschulsky" authorityYear="1845" box="[629,724,403,429]" class="Insecta" family="Zopheridae" genus="Usechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFE46647FDB26BD2B99F57AA" box="[629,724,403,429]" italics="true" pageId="4" pageNumber="205">Usechus</emphasis>
</taxonomicName>
representative, described herein, could also be associated with an analogous forest community of coniferous and fagacean wood and fungi.
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Genus
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<emphasis id="B929B23CFFE66645FF296B66BA1B5745" bold="true" box="[238,336,295,322]" italics="true" pageId="6" pageNumber="207">Coxelus</emphasis>
Dejean, 1821
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species:
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Sturm, 1807
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<emphasis id="B929B23CFFE66645FF506BF0BAFA57CC" bold="true" box="[151,433,433,459]" pageId="6" pageNumber="207">Taxonomic assignment.</emphasis>
The inclusion under study possesses a combination of characters corresponding to the diverse tribe
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within the subfamily
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: (1) antennal insertions concealed dorsally; (2) antennae distinctly clubbed, glabrous, lacking scale-like setae; (3) procoxal cavities open posteriorly; (4) all tarsi tetramerous, not dilated; (5) protibiae without apical spurs; and (6) metacoxae separated by triangular process of abdominal ventrite 1. The new extinct species is assigned to the genus
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based on the combination of the following characters: antennae 11-segmented with 2-segmented club; eyes hemispherical; pronotal disc without special sculpture, pronotum narrowed at anterior angles and evenly rounded laterally; short antennal groove present ventrad eye; elytra non carinate; intercoxal process of abdominal ventrite 1 about as wide as metacoxal lenght, and abdominal ventrite 5 without marginal groove; body elongate, and dorsal surface covered with narrow erect setation.
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<mods:location id="029C9E4A0D881AA09CDE406965858D8C">
<mods:url id="D9064CB5A602B60D5145333AA7CA6637">http://dx.doi.org/10.11646/zootaxa.5536.2.1</mods:url>
</mods:location>
<mods:classification id="F6C688A196575F231DC87BBB03989E3C">journal article</mods:classification>
<mods:identifier id="97D26E2714FDB64E6DF31F6752564ECD" type="DOI">10.11646/zootaxa.5536.2.1</mods:identifier>
<mods:identifier id="3D1F103DDA8D25D857FD2600B4134D73" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="02FD308928232A49752D7C4AFE5F968F" type="Zenodo-Dep">14033618</mods:identifier>
<mods:identifier id="C872A2A67A629628EA520F7CFCA526C9" type="ZooBank">5F1FBB59-9C69-4E66-9B0D-69A015F30EAA</mods:identifier>
</mods:mods>
<treatment id="03F4DF38FFE6664BFF50697CBA3257AA" LSID="urn:lsid:plazi:treatment:03F4DF38FFE6664BFF50697CBA3257AA" httpUri="http://treatment.plazi.org/id/03F4DF38FFE6664BFF50697CBA3257AA" lastPageId="8" lastPageNumber="209" pageId="6" pageNumber="207">
<subSubSection id="C3473DA5FFE66645FF50697CB845555F" box="[151,782,829,856]" pageId="6" pageNumber="207" type="nomenclature">
<paragraph id="8BE26E2EFFE66645FF50697CB845555F" blockId="6.[151,782,829,893]" box="[151,782,829,856]" pageId="6" pageNumber="207">
<heading id="D0AAD942FFE66645FF50697CB845555F" bold="true" box="[151,782,829,856]" fontSize="11" level="1" pageId="6" pageNumber="207" reason="1">
<emphasis id="B929B23CFFE66645FF50697CB845555F" bold="true" box="[151,782,829,856]" pageId="6" pageNumber="207">
<taxonomicName id="4C5D15ADFFE66645FF50697CB9E35550" authority="Alekseev et Bukejs" authorityName="Alekseev et Bukejs" authorityYear="2024" box="[151,680,829,856]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="carstengroehni" status="sp. nov.">
<emphasis id="B929B23CFFE66645FF50697CBAFC555F" bold="true" box="[151,439,829,856]" italics="true" pageId="6" pageNumber="207">Coxelus carstengroehni</emphasis>
Alekseev et Bukejs
</taxonomicName>
<taxonomicNameLabel id="A21A0F47FFE66645FD77697FB845555F" box="[688,782,830,856]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE66645FF506923BA40557A" box="[151,267,866,893]" pageId="6" pageNumber="207" type="description">
<paragraph id="8BE26E2EFFE66645FF506923BA40557A" blockId="6.[151,782,829,893]" box="[151,267,866,893]" pageId="6" pageNumber="207">
(
<figureCitation id="136672ABFFE66645FF586923BA48557A" box="[159,259,866,893]" captionStart="FIGURES 56" captionStartId="8.[151,264,1919,1943]" captionTargetBox="[151,1435,486,1893]" captionTargetId="figure-145@8.[151,1435,486,1893]" captionTargetPageId="8" captionText="FIGURES 56. Coxelus carstengroehni sp. nov., holotype, No. 5208 [GPIH]: 5—habitus, dorso-lateral view; 6—habitus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14033624" httpUri="https://zenodo.org/record/14033624/files/figure.png" pageId="6" pageNumber="207">Figs 56</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE66645FF5069EBBBA252E3" pageId="6" pageNumber="207" type="materials_examined">
<paragraph id="8BE26E2EFFE66645FF5069EBBEC35253" blockId="6.[151,1437,937,1972]" pageId="6" pageNumber="207">
<emphasis id="B929B23CFFE66645FF5069EBBA0255C4" bold="true" box="[151,329,937,964]" pageId="6" pageNumber="207">Type material.</emphasis>
<typeStatus id="54E6D08CFFE66645FE9369EBBA8A55C3" box="[340,449,938,964]" pageId="6" pageNumber="207" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED4CF6EBFFE66645FE0969EBB94455C3" box="[462,527,938,964]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="6" pageNumber="207">GPIH</collectionCode>
no. 5208,
<collectionCode id="ED4CF6EBFFE66645FD5569EBB9AB55C3" box="[658,736,938,964]" pageId="6" pageNumber="207">CCGG</collectionCode>
no. 8705 (ex coll. Jonas Damzen JDC-12539); “
<typeStatus id="54E6D08CFFE66645FAD869EBBEC155C3" box="[1311,1418,938,964]" pageId="6" pageNumber="207" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C5D15ADFFE66645FF50698FBAEB55EF" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[151,416,974,1000]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="carstengroehni" status="sp. nov.">
<emphasis id="B929B23CFFE66645FF50698FBAEB55EF" box="[151,416,974,1000]" italics="true" pageId="6" pageNumber="207">Coxelus carstengroehni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A0F47FFE66645FE61698FBAB055EF" box="[422,507,974,1000]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
/ Alekseev et Bukejs des. 2024” [red printed label]; adult, sex unknown.A complete beetle with partially exposed metathoracic wings is included in a transparent, yellow amber piece with 11×
<quantity id="4CA5C3CBFFE66645FAF769B3BE24520B" box="[1328,1391,1010,1036]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="207" unit="mm" value="8.0">8 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFE66645FE726E57BAB25237" box="[437,505,1046,1072]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="207" unit="mm" value="1.0">1 mm</quantity>
, preserved without supplementary fixation. Syninclusions:
<specimenCount id="9D5BA5A7FFE66645FB5F6E57BE745237" box="[1176,1343,1046,1072]" count="2" pageId="6" pageNumber="207" type="generic">two specimens</specimenCount>
of male
<taxonomicName id="4C5D15ADFFE66645FF506E7BBA7C5253" box="[151,311,1082,1108]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
(
<taxonomicName id="4C5D15ADFFE66645FE816E7BBAEB5253" box="[326,416,1082,1108]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
: Nematocera), one of them is partially damaged, and several stellate
<taxonomicName id="4C5D15ADFFE66645FB646E7BBE405253" authorityName="Dumortier" authorityYear="1829" box="[1187,1291,1082,1108]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="207" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
trichomes.
</paragraph>
<paragraph id="8BE26E2EFFE66645FF006E1FBAEF529B" blockId="6.[151,1437,937,1972]" pageId="6" pageNumber="207">
<emphasis id="B929B23CFFE66645FF006E1FBA38527F" bold="true" box="[199,371,1118,1144]" pageId="6" pageNumber="207">
<typeStatus id="54E6D08CFFE66645FF006E1FBA49527F" box="[199,258,1118,1144]" pageId="6" pageNumber="207">Type</typeStatus>
stratum.
</emphasis>
Baltic amber from Eocene amber-bearing Blaue Erde deposits; estimated age: middlelate Eocene (Standke 1998).
</paragraph>
<paragraph id="8BE26E2EFFE66645FF006EE7BBA252E3" blockId="6.[151,1437,937,1972]" pageId="6" pageNumber="207">
<materialsCitation id="3B356473FFE66645FF006EE7BBAE52E3" country="Russia" county="Peninsula" location="Yantarny" municipality="Sambian" pageId="6" pageNumber="207" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B929B23CFFE66645FF006EE7BA2E52B8" bold="true" box="[199,357,1189,1216]" pageId="6" pageNumber="207">
<typeStatus id="54E6D08CFFE66645FF006EE7BA4952C7" box="[199,258,1190,1216]" pageId="6" pageNumber="207">Type</typeStatus>
locality.
</emphasis>
<location id="8E8238F5FFE66645FEAD6EE7BA8452C7" LSID="urn:lsid:plazi:treatment:03F4DF38FFE6664BFF50697CBA3257AA:8E8238F5FFE66645FEAD6EE7BA8452C7" box="[362,463,1190,1216]" country="Russia" county="Peninsula" municipality="Sambian" name="Yantarny" pageId="6" pageNumber="207" stateProvince="Kaliningrad">Yantarny</location>
settlement (formerly Palmnicken),
<collectingMunicipality id="6B86F454FFE66645FC9C6EE7B8F652C7" box="[859,957,1190,1216]" pageId="6" pageNumber="207">Sambian</collectingMunicipality>
(Samland)
<collectingCounty id="628316A2FFE66645FBF96EE7BFE552C7" box="[1086,1198,1190,1216]" pageId="6" pageNumber="207">Peninsula</collectingCounty>
,
<collectingRegion id="4999A0CCFFE66645FB7F6EE7BED352C7" box="[1208,1432,1190,1216]" country="Russia" name="Kaliningrad" pageId="6" pageNumber="207">Kaliningrad Region</collectingRegion>
,
<collectingCountry id="F34A2EBEFFE66645FF506E8BBBAE52E3" box="[151,229,1226,1252]" name="Russia" pageId="6" pageNumber="207">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE66644FF006EACBBA6527A" lastPageId="7" lastPageNumber="208" pageId="6" pageNumber="207" type="description">
<paragraph id="8BE26E2EFFE66645FF006EACB8265357" blockId="6.[151,1437,937,1972]" pageId="6" pageNumber="207">
<emphasis id="B929B23CFFE66645FF006EACBA1A5300" bold="true" box="[199,337,1261,1287]" pageId="6" pageNumber="207">Description</emphasis>
. Measurements: total body length (including visible part of head)
<quantity id="4CA5C3CBFFE66645FBF86EAFBFDC530F" box="[1087,1175,1262,1288]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="207" unit="mm" value="2.3">2.3 mm</quantity>
, maximum body width (across elytra)
<quantity id="4CA5C3CBFFE66645FE876F53BAD1532B" box="[320,410,1298,1324]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="207" unit="mm" value="0.8">0.8 mm</quantity>
; pronotal length
<quantity id="4CA5C3CBFFE66645FD9C6F53B9FF532B" box="[603,692,1298,1324]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="207" unit="mm" value="0.6">0.6 mm</quantity>
, maximum pronotal width
<quantity id="4CA5C3CBFFE66645FC2F6F53BF0A532B" box="[1000,1089,1298,1324]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="207" unit="mm" value="0.7">0.7 mm</quantity>
, basal pronotal width
<quantity id="4CA5C3CBFFE66645FAF96F53BED3532B" box="[1342,1432,1298,1324]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="207" unit="mm" value="0.6">0.6 mm</quantity>
; elytral length (along elytral suture, including scutellum)
<quantity id="4CA5C3CBFFE66645FCD76F77B8225357" box="[784,873,1334,1361]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="207" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
<paragraph id="8BE26E2EFFE66645FF006F1BBA1F53E7" blockId="6.[151,1437,937,1972]" pageId="6" pageNumber="207">Body subparallel-sided, elongate, about 2.9× as long as wide, weakly convex dorsally and ventrally, unicolorous black (as preserved). Pubescence: head with short, erect setae; pronotum sparsely covered with stout, erect setae; elytra sparsely and uniformly covered with longer, erect, narrow, slightly curved setae arranged in rows, and not forming patches.</paragraph>
<paragraph id="8BE26E2EFFE66645FF006FABBA9351B3" blockId="6.[151,1437,937,1972]" pageId="6" pageNumber="207">Head prognathous, transverse; head densely granulose dorsally, granules rather large, about 2× as wide as diameter of one eye facet, and distance between granules distinctly smaller than diameter of one granule, with each granule bearing short, narrow seta; dense, coarse punctation present ventrally. Anterior margin of clypeus rounded. Compound eyes small, hemispherical, convex, without interfacetal setation. Antennal insertions concealed dorsally; antennal grooves ventrad eye present, short. Antennae short, extending to about middle of pronotal length; 11-segmented with distinct, compact, 2-segmented club; covered with sparse, short, semierect setae; scape wide, subcylindrical; pedicel cylindrical, 2× as long as wide, narrower and shorter than scape; antennomere 3 about 2.2× as long as wide, subcylindrical, slightly dilated apically, about 1.4× longer than pedicel; antennomeres 45 slightly elongate, subcylindrical; antennomeres 68 subquadrate, nearly as long as wide; antennomere 9 weakly transverse, 1.3× as wide as long, slightly dilated apically; antennomere 10 subtrapezoidal, dilated apically, widest, transverse, 2.2× wider than antennomere 9; antennomere 11 oval, transverse, 1.5× as wide as long, distinctly narrower than antennomere 10, finely setose. Terminal maxillary palpomere obliquely truncate, pointed apically, slightly longer than penultimate palpomere.</paragraph>
<paragraph id="8BE26E2EFFE76644FF006AD6B869578E" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">Pronotum slightly transverse, 1.2× as wide as long, widest near midlength and slightly narrowed anteriad and posteriad, maximum pronotal width / basal pronotal width = 1.2; pronotal disc weakly convex, with large, oval, longitudinal impression medially; pronotal surface densely covered with granules, bearing erect, stout setae. Lateral margins weakly rounded, with fine crenulation; anterior margin arcuate; posterior margin rounded. Anterior angles prominent, acute, slightly rounded; posterior angles obtuse. Prothoracic notosternal suture complete. Hypomera and prosternum densely and coarsely punctate. Intercoxal prosternal process elongate, narrow, distinctly narrower than one procoxal diameter. Procoxal cavities open posteriorly.</paragraph>
<paragraph id="8BE26E2EFFE76644FF006BD2B85457AA" blockId="7.[151,1437,151,2013]" box="[199,799,403,429]" pageId="7" pageNumber="208">Scutellar shield minute, semioval, slightly transverse.</paragraph>
<paragraph id="8BE26E2EFFE76644FF006BF6BFB454CA" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">Elytra almost parallel-sided in anterior two-thirds of length, tapered at apex, about 1.9× as longe as wide combined, non-carinate, convex, 2.5× longer than pronotum. Elytral base distinctly wider than pronotal posterior margin.Humeral angles rounded. Elytra striate-punctate. Scutellary striole not apparent.Interstriae flat, with elongate granules. Epipleura well-developed, widest in basal half, slightly narrowed posteriad, reaching elytral apices. Meso- and metaventrites densely and coarsely punctate. Mesocoxal cavities closed. Metanepisternum narrow, about 7.9× as long as wide, covered with dense and coarse punctures. Metaventrite with disc slightly convex, slightly shorter than abdominal ventrite 1; with discrimen distinct in posterior two-thirds of length. Relative length ratio of pro- to meso- to metaventrite to abdomen approximately equal to 10:7:8:22. Metathoracic wings developed.</paragraph>
<paragraph id="8BE26E2EFFE76644FF006896BFAD55A2" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">Legs rather short and robust, finely punctate. Procoxa nearly rounded, narrowly separated; mesocoxae relatively round, narrowly separated by about 0.5× diameter of mesocoxa; metacoxae oval, transverse, separated by about one longitudinal diameter of metacoxa. Femora nearly spindle-shaped, flattened, and widened medially; femora and tibiae subequal in length. Tibiae straight, slightly dilated apically, with fringe of short, fine setae apically. Tarsal formula 4-4-4; tarsomeres with sparse, fine setation ventrally; tarsomere 4 longest, longer than metatarsomeres 13 combined, and slightly curved. Claws simple, large, about 0.5× as long as tarsomere 4, and robust.</paragraph>
<paragraph id="8BE26E2EFFE76644FF0069EEBBA6527A" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">Abdomen with five visible ventrites, abdominal sutures distinct throughout length; densely covered with punctures that grade from large to small (ventrites 1 and 2 with distinctly larger punctation than ventrires 35), distance between punctures 0.51.5× diameter of one puncture. Relative length ratios of ventrites 15 equal to 5:5:4:3:3. Abdominal ventrite 1 with small, apically acute intercoxal process in form of equilateral triangle that is about as wide as metacoxal lenght. Abdominal ventrite 5 subtrapezoidal, truncate apically, without marginal groove.</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE76644FF006EC7BA3E509E" pageId="7" pageNumber="208" type="diagnosis">
<paragraph id="8BE26E2EFFE76644FF006EC7BBBD539A" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">
<emphasis id="B929B23CFFE76644FF006EC7BA9F52A7" bold="true" box="[199,468,1158,1184]" pageId="7" pageNumber="208">Differential diagnosis.</emphasis>
<taxonomicName id="4C5D15ADFFE76644FE256EC6B9B852A6" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[482,755,1159,1185]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="carstengroehni" status="sp. nov.">
<emphasis id="B929B23CFFE76644FE256EC6B9B852A6" box="[482,755,1159,1185]" italics="true" pageId="7" pageNumber="208">Coxelus carstengroehni</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFE76644FCC76EC6B82B52A6" bold="true" box="[768,864,1159,1185]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21A0F47FFE76644FCC76EC6B82B52A6" box="[768,864,1159,1185]" pageId="7" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all extant congeners based on the combination of the following characters: eyes without interfacetal setae; metathoracic wings present and metathorax not shortened (in contrast to apterous
<taxonomicName id="4C5D15ADFFE76644FD866E8EB9D052EE" authorityName="Dejean" authorityYear="1821" box="[577,667,1231,1257]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFE76644FD866E8EB9D052EE" box="[577,667,1231,1257]" italics="true" pageId="7" pageNumber="208">Coxelus</emphasis>
</taxonomicName>
of Recent fauna); relatively long elytra, about 1.9× as long as wide; weakly transverse pronotum, 1.2× as wide as long; and elytral setation uniform and not forming patches. The new fossil species resembles the North American
<taxonomicName id="4C5D15ADFFE76644FD406F56B8E35336" authority="(Stephan, 1989)" baseAuthorityName="Stephan" baseAuthorityYear="1989" box="[647,936,1303,1329]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B929B23CFFE76644FD406F56B9BA5336" box="[647,753,1303,1329]" italics="true" pageId="7" pageNumber="208">C. longus</emphasis>
(
<bibRefCitation id="EFCC13DFFFE76644FD396F56B8EB5336" author="Stephan, K. H." box="[766,928,1303,1329]" pageId="7" pageNumber="208" refId="ref28343" refString="Stephan, K. H. (1989) The Bothrideridae and Colydiidae of America north of Mexico (Coleoptera: Clavicornia and Heteromera. Occasional Papers of the Florida State Collection of Arthropods. Vol. 6. Florida Department of Agriculture and Consumer Services, Gainesville, Flordia, xii + 65 pp." type="book" year="1989">Stephan, 1989</bibRefCitation>
)
</taxonomicName>
in its elongate body, but can be distinguished from it based on the smaller body length of
<quantity id="4CA5C3CBFFE76644FD436F7AB9915352" box="[644,730,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="208" unit="mm" value="2.3">2.3 mm</quantity>
(
<quantity id="4CA5C3CBFFE76644FD2E6F7AB80A5352" box="[745,833,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="208" unit="mm" value="2.9">2.9 mm</quantity>
in the
<typeStatus id="54E6D08CFFE76644FC496F7AB8BB5352" box="[910,1008,1339,1365]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5D15ADFFE76644FBD16F7ABFC95352" baseAuthorityName="Stephan" baseAuthorityYear="1989" box="[1046,1154,1339,1365]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B929B23CFFE76644FBD16F7ABFC95352" box="[1046,1154,1339,1365]" italics="true" pageId="7" pageNumber="208">C. longus</emphasis>
</taxonomicName>
according to the original description), larger eyes (eyes reduced, with 2632 facets in
<taxonomicName id="4C5D15ADFFE76644FCF86F1EB8E0537E" baseAuthorityName="Stephan" baseAuthorityYear="1989" box="[831,939,1375,1401]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B929B23CFFE76644FCF86F1EB8E0537E" box="[831,939,1375,1401]" italics="true" pageId="7" pageNumber="208">C. longus</emphasis>
</taxonomicName>
), metathoracic wings present, and elytra not connate.
</paragraph>
<paragraph id="8BE26E2EFFE76644FF006FE6BA3E509E" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">
Among the
<taxonomicName id="4C5D15ADFFE76644FE8C6FE6BAF553C6" authorityName="Erichson" authorityYear="1845" box="[331,446,1447,1473]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="tribe" tribe="Synchitini">Synchitini</taxonomicName>
described from Baltic amber, the new species may be confused only with the apparently similar and probably related
<taxonomicName id="4C5D15ADFFE76644FE2E6F8ABF0353E2" authority="Alekseev et Pankowski, 2020" authorityName="Alekseev et Pankowski" authorityYear="2020" box="[489,1096,1483,1509]" class="Insecta" family="Zopheridae" genus="Semicoxelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="sontagae">
<emphasis id="B929B23CFFE76644FE2E6F8AB9A853E3" box="[489,739,1483,1509]" italics="true" pageId="7" pageNumber="208">Semicoxelus sontagae</emphasis>
<bibRefCitation id="EFCC13DFFFE76644FD2A6F8ABF0353E2" author="Alekseev, V. I. &amp; Pankowski, M." box="[749,1096,1483,1509]" pageId="7" pageNumber="208" pagination="607 - 613" refId="ref24310" refString="Alekseev, V. I. &amp; Pankowski, M. (2020) A new fossil genus of the tribe Synchitini Erichson (Tenebrionoidea: Zopheridae: Colydiinae) from Eocene Baltic amber. Palaeoentomology, 3 (6), 607 - 613. https: // doi. org / 10.11646 / palaeoentomology. 3.6.12" type="journal article" year="2020">Alekseev et Pankowski, 2020</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C5D15ADFFE76644FB916F8ABE1B53E3" authorityName="Alekseev et Pankowski" authorityYear="2020" box="[1110,1360,1483,1509]" class="Insecta" family="Zopheridae" genus="Semicoxelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="sontagae">
<emphasis id="B929B23CFFE76644FB916F8ABE1B53E3" box="[1110,1360,1483,1509]" italics="true" pageId="7" pageNumber="208">Semicoxelus sontagae</emphasis>
</taxonomicName>
(body length of the
<typeStatus id="54E6D08CFFE76644FEF26FAEBADC500E" box="[309,407,1519,1545]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
is
<quantity id="4CA5C3CBFFE76644FE796FAFB96D500E" box="[446,550,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="7" pageNumber="208" unit="mm" value="1.85">1.85 mm</quantity>
) differs from
<taxonomicName id="4C5D15ADFFE76644FD0C6FAEB892500E" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[715,985,1519,1545]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="carstengroehni" status="sp. nov.">
<emphasis id="B929B23CFFE76644FD0C6FAEB892500E" box="[715,985,1519,1545]" italics="true" pageId="7" pageNumber="208">Coxelus carstengroehni</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFE76644FC256FAEBF74500E" bold="true" box="[994,1087,1519,1545]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21A0F47FFE76644FC256FAEBF74500E" box="[994,1087,1519,1545]" pageId="7" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in its vertically elongate eyes (hemispherical in the new species), subantennal grooves absent (present in the new species), antennae stouter and antennal club is more weakly expressed (antennal club is well-defined, and antennomere 10 about 2.2× as wide as antennomere
<quantity id="4CA5C3CBFFE76644FEE86C1ABA105072" box="[303,347,1627,1653]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="7" pageNumber="208" unit="in" value="9.0">9 in</quantity>
<taxonomicName id="4C5D15ADFFE76644FEA56C1AB9675072" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[354,556,1627,1653]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="carstengroehni" status="sp. nov.">
<emphasis id="B929B23CFFE76644FEA56C1AB9675072" box="[354,556,1627,1653]" italics="true" pageId="7" pageNumber="208">C. carstengroehni</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFE76644FDF46C1AB9C65072" bold="true" box="[563,653,1627,1653]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21A0F47FFE76644FDF46C1AB9C65072" box="[563,653,1627,1653]" pageId="7" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), and distinctly curved elytral setation (erect and only slightly curved in the new species).
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE76644FF006CE3BF7D50E6" pageId="7" pageNumber="208" type="etymology">
<paragraph id="8BE26E2EFFE76644FF006CE3BF7D50E6" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">
<emphasis id="B929B23CFFE76644FF006CE3BAD750BB" bold="true" box="[199,412,1698,1724]" pageId="7" pageNumber="208">Derivatio nominis</emphasis>
. The specific epithet is a patronym; the new species is named in honor of Mr. Carsten Gröhn (Glinde,
<collectingCountry id="F34A2EBEFFE76644FF3C6C86BA2350E6" box="[251,360,1735,1761]" name="Germany" pageId="7" pageNumber="208">Germany</collectingCountry>
), who provided the amber piece with the
<typeStatus id="54E6D08CFFE76644FCF16C86B8D350E6" box="[822,920,1735,1761]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
for our study.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE76644FF006CABBAE951DA" pageId="7" pageNumber="208" type="distribution">
<paragraph id="8BE26E2EFFE76644FF006CABBAE951DA" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">
<emphasis id="B929B23CFFE76644FF006CABB94A5102" bold="true" box="[199,513,1770,1797]" pageId="7" pageNumber="208">Distribution of congeners.</emphasis>
The genus
<taxonomicName id="4C5D15ADFFE76644FD406CAAB9AA5102" authorityName="Dejean" authorityYear="1821" box="[647,737,1771,1797]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFE76644FD406CAAB9AA5102" box="[647,737,1771,1797]" italics="true" pageId="7" pageNumber="208">Coxelus</emphasis>
</taxonomicName>
occurs in the western Nearctic and Neotropics (2 species:
<taxonomicName id="4C5D15ADFFE76644FA446CAABADE512E" authority="(Stephan 1989)" baseAuthorityName="Stephan" baseAuthorityYear="1989" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B929B23CFFE76644FA446CAABBA9512E" italics="true" pageId="7" pageNumber="208">C. longus</emphasis>
(
<bibRefCitation id="EFCC13DFFFE76644FF376D4EBAC7512E" author="Stephan, K. H." box="[240,396,1807,1833]" pageId="7" pageNumber="208" refId="ref28343" refString="Stephan, K. H. (1989) The Bothrideridae and Colydiidae of America north of Mexico (Coleoptera: Clavicornia and Heteromera. Occasional Papers of the Florida State Collection of Arthropods. Vol. 6. Florida Department of Agriculture and Consumer Services, Gainesville, Flordia, xii + 65 pp." type="book" year="1989">Stephan 1989</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C5D15ADFFE76644FE0C6D4EB981512E" authority="Horn 1885" authorityName="Horn" authorityYear="1885" box="[459,714,1806,1833]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="B929B23CFFE76644FE0C6D4EB903512F" box="[459,584,1807,1833]" italics="true" pageId="7" pageNumber="208">C. serratus</emphasis>
Horn 1885
</taxonomicName>
), and in temperate and southern areas of the western and eastern parts of Palearctic (6 species:
<taxonomicName id="4C5D15ADFFE76644FE206D72B9AA514A" authority="Dajoz, 1973" authorityName="Dajoz" authorityYear="1973" box="[487,737,1843,1869]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="alinae">
<emphasis id="B929B23CFFE76644FE206D72B905514A" box="[487,590,1843,1869]" italics="true" pageId="7" pageNumber="208">C. alinae</emphasis>
Dajoz, 1973
</taxonomicName>
;
<taxonomicName id="4C5D15ADFFE76644FD2A6D72BFD8514A" authority="(Frivaldszky, 1893)" baseAuthorityName="Frivaldszky" baseAuthorityYear="1893" box="[749,1171,1843,1869]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="bituberculatus">
<emphasis id="B929B23CFFE76644FD2A6D72B8E4514A" box="[749,943,1843,1869]" italics="true" pageId="7" pageNumber="208">C. bituberculatus</emphasis>
(Frivaldszky, 1893)
</taxonomicName>
;
<taxonomicName id="4C5D15ADFFE76644FB676D72BB995176" authority="Reitter, 1885" authorityName="Reitter" authorityYear="1885" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="humeridens">
<emphasis id="B929B23CFFE76644FB676D72BE08514A" box="[1184,1347,1843,1869]" italics="true" pageId="7" pageNumber="208">C. humeridens</emphasis>
Reitter, 1885
</taxonomicName>
;
<taxonomicName id="4C5D15ADFFE76644FF196D16BAB05176" authority="Pie, 1901" authorityName="Pie" authorityYear="1901" box="[222,507,1879,1905]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="luteopilosus">
<emphasis id="B929B23CFFE76644FF196D16BACD5176" box="[222,390,1879,1905]" italics="true" pageId="7" pageNumber="208">C. luteopilosus</emphasis>
Pie, 1901
</taxonomicName>
;
<taxonomicName id="4C5D15ADFFE76644FDC06D16B8595176" authority="(Sturm, 1807)" baseAuthorityName="Sturm" baseAuthorityYear="1807" box="[519,786,1879,1905]" class="Insecta" family="Tenebrionidae" genus="Bolitophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="pictus">
<emphasis id="B929B23CFFE76644FDC06D16B9205177" box="[519,619,1879,1905]" italics="true" pageId="7" pageNumber="208">C. pictus</emphasis>
(Sturm, 1807)
</taxonomicName>
; and
<taxonomicName id="4C5D15ADFFE76644FC896D16BF025176" authority="Slipinski, 1985" authorityName="Slipinski" authorityYear="1985" box="[846,1097,1878,1907]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="yeti">
<emphasis id="B929B23CFFE76644FC896D16B8DC5177" box="[846,919,1879,1905]" italics="true" pageId="7" pageNumber="208">C. yeti</emphasis>
<bibRefCitation id="EFCC13DFFFE76644FC586D19BF025176" author="Slipinski, S. A." box="[927,1097,1878,1907]" pageId="7" pageNumber="208" pagination="181 - 195" refId="ref27901" refString="Slipinski, S. A. (1985) New and little known species of Colydiidae from Asia, Madagascar and Comoro Islands. Annales Zoologici, 39 (10), 181 - 195." type="journal article" year="1985">Ślipinśki, 1985</bibRefCitation>
</taxonomicName>
) (
<bibRefCitation id="EFCC13DFFFE76644FBA66D16BE5A5176" author="Ivie, M. A. &amp; Lord, N. P. &amp; Foley, I. A." box="[1121,1297,1879,1905]" pageId="7" pageNumber="208" pagination="755 - 788" refId="ref25945" refString="Ivie, M. A., Lord, N. P. &amp; Foley, I. A. (2016) Colydiine genera (Coleoptera: Zopheridae: Colydiinae) of the new world: a key and nomenclatural acts 30 years in the making. The Coleopterists Bulletin, 70 (4), 755 - 788. https: // doi. org / 10.1649 / 0010 - 065 X- 70.4.755" type="journal article" year="2016">
Ivie
<emphasis id="B929B23CFFE76644FB526D16BF8C5176" box="[1173,1223,1879,1905]" italics="true" pageId="7" pageNumber="208">et al</emphasis>
. 2016
</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFE76644FADB6D19BA6A5192" author="Slipinski, S. A. &amp; Schuh, R." pageId="7" pageNumber="208" refId="ref28217" refString="Slipinski, S. A. &amp; Schuh, R. (2008) Zopheridae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Apollo Books, Stenstrup, pp. 78 - 87." type="book" year="2008">Ślipiński &amp; Schuh 2008</bibRefCitation>
). The fossil species described in the current paper confirms the presence of the genus in the Paleogene Western Palearctic, which happens to be the area where the genus is represented by the largest number of Recent species (4 spp.) as well.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE8664BFF006AD6BA3257AA" pageId="8" pageNumber="209" type="biology_ecology">
<paragraph id="8BE26E2EFFE8664BFF006AD6BA3257AA" blockId="8.[151,1437,150,429]" pageId="8" pageNumber="209">
<emphasis id="B929B23CFFE8664BFF006AD6BA9A56B6" bold="true" box="[199,465,150,177]" pageId="8" pageNumber="209">Ecology of congeners.</emphasis>
Recent representatives of
<taxonomicName id="4C5D15ADFFE8664BFCC26AD6B82B56B6" authorityName="Dejean" authorityYear="1821" box="[773,864,151,177]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFE8664BFCC26AD6B82B56B6" box="[773,864,151,177]" italics="true" pageId="8" pageNumber="209">Coxelus</emphasis>
</taxonomicName>
are usually found in leaf litter and probably feed on decaying vegetation or fungi (
<bibRefCitation id="EFCC13DFFFE8664BFDE46AFDB82656D2" author="Slipinski, S. A. &amp; Lawrence, J. F." box="[547,877,187,215]" pageId="8" pageNumber="209" refId="ref28031" refString="Slipinski, S. A. &amp; Lawrence, J. F. (2010) 11.9. Zopheridae Solier, 1834. In: Leschen, R. A. B, Beutel, R. G. &amp; Lawrence, J. F. (Eds.), Handbook of zoology. Arthropoda: Insecta. Part 38. Coleoptera, beetles. Vol. 2. Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). Walter de Gruyter, Berlin, pp. 548 - 559. https: // doi. org / 10.1515 / 9783110911213.548" type="book" year="2010">Ślipiński &amp; Lawrence 2010</bibRefCitation>
). They are considered saproxylophagous, being associated with dead wood and woody rotting material (as in
<emphasis id="B929B23CFFE8664BFC9D6A9EB88D56FF" box="[858,966,223,249]" italics="true" pageId="8" pageNumber="209">
<taxonomicName id="4C5D15ADFFE8664BFC9D6A9EB88956FF" baseAuthorityName="Sturm" baseAuthorityYear="1807" box="[858,962,223,249]" class="Insecta" family="Tenebrionidae" genus="Bolitophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="pictus">C. pictus</taxonomicName>
,
</emphasis>
according to
<bibRefCitation id="EFCC13DFFFE8664BFBA26A9EBE6656FE" author="Parisi, F. &amp; Innangi, M. &amp; Tognetti, R. &amp; Lombardi, F. &amp; Chirici, G. &amp; Marchetti, M." box="[1125,1325,223,249]" pageId="8" pageNumber="209" refId="ref27177" refString="Parisi, F., Innangi, M., Tognetti, R., Lombardi, F., Chirici, G. &amp; Marchetti, M. (2021). Forest stand structure and coarse woody debris determine the biodiversity of beetle communities in Mediterranean mountain beech forests. Global Ecology and Conservation, 28, e 01637. https: // doi. org / 10.1016 / j. gecco. 2021. e 01637" type="journal volume" year="2021">
Parisi
<emphasis id="B929B23CFFE8664BFB696A9EBFAB56FE" box="[1198,1248,223,249]" italics="true" pageId="8" pageNumber="209">et al</emphasis>
. 2021
</bibRefCitation>
).
<taxonomicName id="4C5D15ADFFE8664BFA866A9EBB90571B" baseAuthorityName="Sturm" baseAuthorityYear="1807" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="pictus">
<emphasis id="B929B23CFFE8664BFA866A9EBB90571B" italics="true" pageId="8" pageNumber="209">Coxelus pictus</emphasis>
</taxonomicName>
were numerous in samples taken from the decaying small and medium diameter wood debris in two Italian beech forests (
<bibRefCitation id="EFCC13DFFFE8664BFEF86B66B97E5746" author="Macagno, A. L. M. &amp; Hardersen, S. &amp; Nardi, G. &amp; Lo Giudice, G. &amp; Mason, F." box="[319,565,294,321]" pageId="8" pageNumber="209" pagination="510 - 519" refId="ref26737" refString="Macagno, A. L. M., Hardersen, S., Nardi, G., Lo Giudice, G. &amp; Mason, F. (2015) Measuring saproxylic beetle diversity in small and medium diameter dead wood: the &quot; grab-and-go &quot; method. European Journal of Entomology, 112 (3), 510 - 519. https: // doi. org / 10.14411 / eje. 2015.049" type="journal article" year="2015">
Macagno
<emphasis id="B929B23CFFE8664BFE746B66BAAC5746" box="[435,487,295,321]" italics="true" pageId="8" pageNumber="209">et al</emphasis>
. 2015
</bibRefCitation>
),
<taxonomicName id="4C5D15ADFFE8664BFD8D6B66B9805747" authorityName="Horn" authorityYear="1885" box="[586,715,295,321]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="B929B23CFFE8664BFD8D6B66B9805747" box="[586,715,295,321]" italics="true" pageId="8" pageNumber="209">C. serratus</emphasis>
</taxonomicName>
has been reported from redwood duff and under bark of dead Douglas fir, whereas
<taxonomicName id="4C5D15ADFFE8664BFE486B0ABAB65762" baseAuthorityName="Stephan" baseAuthorityYear="1989" box="[399,509,331,357]" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B929B23CFFE8664BFE486B0ABAB65762" box="[399,509,331,357]" italics="true" pageId="8" pageNumber="209">C. longus</emphasis>
</taxonomicName>
has been associated with dead ponderosa pines (
<bibRefCitation id="EFCC13DFFFE8664BFBFF6B0ABF905762" author="Stephan, K. H." box="[1080,1243,331,357]" pageId="8" pageNumber="209" refId="ref28343" refString="Stephan, K. H. (1989) The Bothrideridae and Colydiidae of America north of Mexico (Coleoptera: Clavicornia and Heteromera. Occasional Papers of the Florida State Collection of Arthropods. Vol. 6. Florida Department of Agriculture and Consumer Services, Gainesville, Flordia, xii + 65 pp." type="book" year="1989">Stephan 1989</bibRefCitation>
). The Eocene
<taxonomicName id="4C5D15ADFFE8664BFA456B0ABA0A578E" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" class="Insecta" family="Zopheridae" genus="Coxelus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="carstengroehni" status="sp. nov.">
<emphasis id="B929B23CFFE8664BFA456B0ABA0A578E" italics="true" pageId="8" pageNumber="209">C. carstengroehni</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFE8664BFE8F6B2EBAE9578E" bold="true" box="[328,418,367,393]" pageId="8" pageNumber="209">
<taxonomicNameLabel id="A21A0F47FFE8664BFE8F6B2EBAE9578E" box="[328,418,367,393]" pageId="8" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
could be also associated with an analogous primeval forest community of dead coniferous and fagacean wood.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,88 @@
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<mods:title id="95E147CADD9C42B304FB70585BB3D366">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests</mods:title>
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<emphasis id="B929B23CFFE9664AFF506AD6BA8356B6" bold="true" box="[151,456,151,178]" pageId="9" pageNumber="210">
Genus
<taxonomicName id="4C5D15ADFFE9664AFF296AD6BA8356B6" ID-CoL="63JML" authority="Dajoz, 1984" authorityName="Dajoz" authorityYear="1984" box="[238,456,151,178]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFE9664AFF296AD6BA6556B5" bold="true" box="[238,302,151,178]" italics="true" pageId="9" pageNumber="210">Paha</emphasis>
<bibRefCitation id="EFCC13DFFFE9664AFEF26AD6BA8356B6" author="Dajoz, R." box="[309,456,151,177]" pageId="9" pageNumber="210" pagination="145 - 157" refId="ref25423" refString="Dajoz, R. (1984) Note sur quatre genres de Coleopteres Colydiidae: Nematidium Erichson, Monoedus Horn, Lobogestoria Reitter et Paha n. gen. Bulletin Mensuel de la Societe Linneenne de Lyon, 53 (4), 145 - 157. https: // doi. org / 10.3406 / linly. 1984.10633" type="journal article" year="1984">Dajoz, 1984</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE9664AFF506AA1B9F15466" pageId="9" pageNumber="210" type="materials_examined">
<paragraph id="8BE26E2EFFE9664AFF506AA1B9E256FE" blockId="9.[151,681,223,250]" box="[151,681,223,250]" pageId="9" pageNumber="210">
<typeStatus id="54E6D08CFFE9664AFF506AA1BB8456FD" box="[151,207,224,250]" pageId="9" pageNumber="210">Type</typeStatus>
species:
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<emphasis id="B929B23CFFE9664AFEF26A9EB95156FE" box="[309,538,223,249]" italics="true" pageId="9" pageNumber="210">Paha guadalupensis</emphasis>
<bibRefCitation id="EFCC13DFFFE9664AFDE76A9EB9E256FE" author="Dajoz, R." box="[544,681,223,249]" pageId="9" pageNumber="210" pagination="145 - 157" refId="ref25423" refString="Dajoz, R. (1984) Note sur quatre genres de Coleopteres Colydiidae: Nematidium Erichson, Monoedus Horn, Lobogestoria Reitter et Paha n. gen. Bulletin Mensuel de la Societe Linneenne de Lyon, 53 (4), 145 - 157. https: // doi. org / 10.3406 / linly. 1984.10633" type="journal article" year="1984">Dajoz, 1984</bibRefCitation>
</taxonomicName>
</paragraph>
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<emphasis id="B929B23CFFE9664AFF506B66BAFA5746" bold="true" box="[151,433,295,321]" pageId="9" pageNumber="210">Taxonomic assignment.</emphasis>
The studied inclusion is placed in the tribe
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within the subfamily
<taxonomicName id="4C5D15ADFFE9664AFAE66B66BED75746" authorityName="Erichson" authorityYear="1842" box="[1313,1436,295,321]" class="Insecta" family="Colydiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="family">Colydiinae</taxonomicName>
based on the combination of the following characters: (1) dorsally concealed antennal insertions; (2) antennae clubbed, lacking scale-like setae; (3) procoxal cavities open posteriorly; (4) all tarsi tetramerous, not dilated; (5) protibiae without apical spurs; and (6) metacoxae narrowly separated by triangular process of abdominal ventrite 1. The new extinct species belongs to the genus
<taxonomicName id="4C5D15ADFFE9664AFD166BF6B84757D6" authorityName="Dajoz" authorityYear="1984" box="[721,780,439,465]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFE9664AFD166BF6B84757D6" box="[721,780,439,465]" italics="true" pageId="9" pageNumber="210">Paha</emphasis>
</taxonomicName>
based on its: 10-segmented antennae with 1-segmented, rounded club; antennal groove ventrad eye absent; antennomere 3 not twice as long as antennomere 4; scutellary striole absent; dorsum lacking obvious pubescense (setation short and fine, visible under high magnification only); pronotal disc with elevated central area limited by two parallel longitudinal carinae; odd elytral intervals carinate; and elytral striae with rows of elongate granules.
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View file

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<taxonomicName id="4C5D15ADFFE9664AFF5068F2B93554CA" authority="Alekseev et Bukejs" authorityName="Alekseev et Bukejs" authorityYear="2024" box="[151,638,691,718]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="vanivanitatum" status="sp. nov.">
<emphasis id="B929B23CFFE9664AFF5068F2BAC754CA" bold="true" box="[151,396,691,718]" italics="true" pageId="9" pageNumber="210">Paha vanivanitatum</emphasis>
Alekseev et Bukejs
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<paragraph id="8BE26E2EFFE9664AFF506896BA5254F6" blockId="9.[151,739,691,753]" box="[151,281,727,753]" pageId="9" pageNumber="210">
(
<figureCitation id="136672ABFFE9664AFF586896BA5A54F6" box="[159,273,727,753]" captionStart-0="FIGURES 78" captionStart-1="FIGURES 910" captionStartId-0="10.[151,264,1656,1680]" captionStartId-1="11.[151,264,1563,1587]" captionTargetBox-0="[151,1435,270,1630]" captionTargetBox-1="[151,1435,179,1538]" captionTargetId-0="figure-46@10.[151,1435,270,1630]" captionTargetId-1="figure-17@11.[151,1435,179,1538]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 78. Paha vanivanitatum sp. nov., holotype, No. ACAB 064 [ACAB]: 7—habitus, dorsal view; 8—habitus, ventral view. Scale bar = 0.5 mm." captionText-1="FIGURES 910. Paha vanivanitatum sp. nov., holotype, No. ACAB 064 [ACAB]: 9—details of forebody, dorsal view; 10— prothorax, ventral view. Abbreviations: a2a10—antennomeres 210 respectively; lc—longitudinal carina; pc—procoxa; pf— profemur; pp—prosternal process; pt—protibia. Scale bars = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14033626" figureDoi-1="http://doi.org/10.5281/zenodo.14033628" httpUri-0="https://zenodo.org/record/14033626/files/figure.png" httpUri-1="https://zenodo.org/record/14033628/files/figure.png" pageId="9" pageNumber="210">Figs 710</figureCitation>
)
</paragraph>
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<emphasis id="B929B23CFFE9664AFF50695EBA09553E" bold="true" box="[151,322,799,825]" pageId="9" pageNumber="210">Type material.</emphasis>
<materialsCitation id="3B356473FFE9664AFE80695EBF1E55CE" collectionCode="ACAB" collectorName="No. &amp; Marius Veta &amp; Ventral &amp; Syninclusions" pageId="9" pageNumber="210" specimenCode="ABAC 064" specimenCount="2" typeStatus="holotype">
<typeStatus id="54E6D08CFFE9664AFE80695EBAFF553E" box="[327,436,799,825]" pageId="9" pageNumber="210" type="holotype">Holotype</typeStatus>
:No.
<specimenCode id="DBFBC655FFE9664AFE2F695EB92F553E" box="[488,612,799,825]" collectionCode="ABAC" pageId="9" pageNumber="210">ABAC 064</specimenCode>
[
<collectionCode id="ED4CF6EBFFE9664AFDA8695EB989553E" box="[623,706,799,825]" pageId="9" pageNumber="210">ACAB</collectionCode>
] (
<collectorName id="26A80BF8FFE9664AFD11695EB84B553E" box="[726,768,799,825]" pageId="9" pageNumber="210">No.</collectorName>
3207 ex. coll.
<collectorName id="26A80BF8FFE9664AFC5B695EBF6C553E" box="[924,1063,799,825]" pageId="9" pageNumber="210">Marius Veta</collectorName>
); “
<typeStatus id="54E6D08CFFE9664AFB86695EBFE7553E" box="[1089,1196,799,825]" pageId="9" pageNumber="210" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C5D15ADFFE9664AFB7B695EBED7553E" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[1212,1436,799,825]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="vanivanitatum" status="sp. nov.">
<emphasis id="B929B23CFFE9664AFB7B695EBED7553E" box="[1212,1436,799,825]" italics="true" pageId="9" pageNumber="210">Paha vanivanitatum</emphasis>
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/ Alekseev et Bukejs des. 2024” [red printed label]; adult, sex unknown. A complete beetle with exposed distal part of metathoracic wing included in a transparent, yellow amber piece with 18×
<quantity id="4CA5C3CBFFE9664AFB4E6926BF915585" box="[1161,1242,871,898]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="210" unit="mm" value="12.0">12 mm</quantity>
and a maximum thickness of
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, preserved without supplementary fixation.
<collectorName id="26A80BF8FFE9664AFC8469CAB8DF55A2" box="[835,916,907,933]" pageId="9" pageNumber="210">Ventral</collectorName>
side of head and anterior portion of prosternum completely obscured by milky cover.
<collectorName id="26A80BF8FFE9664AFDFA69EEB99455CE" box="[573,735,943,969]" pageId="9" pageNumber="210">Syninclusions</collectorName>
: few stellate
<taxonomicName id="4C5D15ADFFE9664AFCB769F1B89355CD" authorityName="Dumortier" authorityYear="1829" box="[880,984,944,970]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="210" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
trichomes.
</materialsCitation>
</paragraph>
<paragraph id="8BE26E2EFFE9664AFF006992BAEF5216" blockId="9.[151,1437,799,2014]" pageId="9" pageNumber="210">
<emphasis id="B929B23CFFE9664AFF006992BA3855EA" bold="true" box="[199,371,979,1005]" pageId="9" pageNumber="210">
<typeStatus id="54E6D08CFFE9664AFF006992BA4955EA" box="[199,258,979,1005]" pageId="9" pageNumber="210">Type</typeStatus>
stratum.
</emphasis>
Baltic amber from Eocene amber-bearing Blaue Erde deposits; estimated age: middlelate Eocene (Standke 1998).
</paragraph>
<paragraph id="8BE26E2EFFE9664AFF006E5ABBA2525E" blockId="9.[151,1437,799,2014]" pageId="9" pageNumber="210">
<materialsCitation id="3B356473FFE9664AFF006E5ABBAE525E" country="Russia" county="Peninsula" location="Yantarny" municipality="Sambian" pageId="9" pageNumber="210" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B929B23CFFE9664AFF006E5ABA2E5232" bold="true" box="[199,357,1051,1077]" pageId="9" pageNumber="210">
<typeStatus id="54E6D08CFFE9664AFF006E5ABA495232" box="[199,258,1051,1077]" pageId="9" pageNumber="210">Type</typeStatus>
locality.
</emphasis>
<location id="8E8238F5FFE9664AFEAD6E5DBA845231" LSID="urn:lsid:plazi:treatment:03F4DF38FFE9664FFF5068F2B88F525E:8E8238F5FFE9664AFEAD6E5DBA845231" box="[362,463,1052,1078]" country="Russia" county="Peninsula" municipality="Sambian" name="Yantarny" pageId="9" pageNumber="210" stateProvince="Kaliningrad">Yantarny</location>
settlement (formerly Palmnicken),
<collectingMunicipality id="6B86F454FFE9664AFC9C6E5AB8F65232" box="[859,957,1051,1077]" pageId="9" pageNumber="210">Sambian</collectingMunicipality>
(Samland)
<collectingCounty id="628316A2FFE9664AFBF96E5ABFE55232" box="[1086,1198,1051,1077]" pageId="9" pageNumber="210">Peninsula</collectingCounty>
,
<collectingRegion id="4999A0CCFFE9664AFB7F6E5ABED35232" box="[1208,1432,1051,1077]" country="Russia" name="Kaliningrad" pageId="9" pageNumber="210">Kaliningrad Region</collectingRegion>
,
<collectingCountry id="F34A2EBEFFE9664AFF506E7EBBAE525E" box="[151,229,1087,1113]" name="Russia" pageId="9" pageNumber="210">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFE9664FFF006E22B88A571A" lastPageId="12" lastPageNumber="213" pageId="9" pageNumber="210" type="description">
<paragraph id="8BE26E2EFFE9664AFF006E22BFD852C1" blockId="9.[151,1437,799,2014]" pageId="9" pageNumber="210">
<emphasis id="B929B23CFFE9664AFF006E22BA1A527A" bold="true" box="[199,337,1123,1149]" pageId="9" pageNumber="210">Description</emphasis>
. Measurements: total body length (including visible part of head)
<quantity id="4CA5C3CBFFE9664AFBF86E22BFDC5279" box="[1087,1175,1123,1150]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="210" unit="mm" value="2.4">2.4 mm</quantity>
, maximum body width (across elytra)
<quantity id="4CA5C3CBFFE9664AFE876EC6BAD152A5" box="[320,410,1159,1186]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="210" unit="mm" value="0.9">0.9 mm</quantity>
; pronotal length
<quantity id="4CA5C3CBFFE9664AFD9C6EC6B9FF52A5" box="[603,692,1159,1186]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="210" unit="mm" value="0.6">0.6 mm</quantity>
, maximum pronotal width
<quantity id="4CA5C3CBFFE9664AFC2F6EC6BF0A52A5" box="[1000,1089,1159,1186]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="210" unit="mm" value="0.8">0.8 mm</quantity>
, basal pronotal width
<quantity id="4CA5C3CBFFE9664AFAF96EC6BED352A5" box="[1342,1432,1159,1186]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="210" unit="mm" value="0.7">0.7 mm</quantity>
; elytral length (along elytral suture, including scutellum)
<quantity id="4CA5C3CBFFE9664AFCD76EEAB82252C1" box="[784,873,1195,1222]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="210" unit="mm" value="1.5">1.5 mm</quantity>
; antennal length
<quantity id="4CA5C3CBFFE9664AFBE06EEABFC552C1" box="[1063,1166,1195,1222]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="210" unit="mm" value="0.38">0.38 mm</quantity>
.
</paragraph>
<paragraph id="8BE26E2EFFE9664AFF006E8EBFD05336" blockId="9.[151,1437,799,2014]" pageId="9" pageNumber="210">Body subparallel-sided, elongate, about 2.7× as long as wide, weakly convex dorsally and ventrally, unicolorous black (as preserved). Pubescence: head, pronotum and elytra sparsely covered with short, inconspicuous, curved, semierect setae that are shorter than diameter of puncture or tubercle from which they arise.</paragraph>
<paragraph id="8BE26E2EFFE9664AFF006F7ABA8A509E" blockId="9.[151,1437,799,2014]" pageId="9" pageNumber="210">Head prognathous, transverse; densely granulose dorsally, each round granule bearing short seta. Anterior margin of clypeus rounded. Compound eyes small, slightly convex, apparently hemispherical, without interfacetal setation. Antennal insertions concealed dorsally; antennal grooves ventrad eye apparently absent. Antennae short, extending slightly beyond anterior one-third of pronotal length; 10-segmented with distinct 1-segmented antennal club, antennal club with fine suture between completely fused antennomeres 10 and 11, and sparsely setose; scape almost invisible from above, wide, subcylindrical; pedicel cylindrical, slightly transverse, about as wide as scape; antennomere 3 conical, elongate, about 1.5× as long as wide, about 1.3× longer than antennomere 4; antennomeres 48 subtrapezoidal, slightly dilated apically, as long as wide; antennomere 9 trapezoidal, dilated apically, transverse, 1.3× as wide as long; antennomere 10 widest, widely oval, with widely rounded apex, nearly as long as wide, 2.1× wider than antennomere 9.</paragraph>
<paragraph id="8BE26E2EFFE96649FF006CE5BAFB56D2" blockId="9.[151,1437,799,2014]" lastBlockId="10.[151,1437,151,249]" lastPageId="10" lastPageNumber="211" pageId="9" pageNumber="210">Pronotum slightly transverse, 1.3× as wide as long, widest near midlength, weakly narrowed anteriad and posteriad, with maximum pronotal width / basal pronotal width = 1.1; pronotal surface covered with rounded granules. Pronotal disc with elevated central area delimited by two parallel, longitudinal, slightly curved carinae; elevated area almost flat, with longitudinal shallow impression along midline, and about 0.6× as wide as pronotal maximum width. Anterior pronotal margin sinuate in dorsal view; posterior margin almost straight medially, shallowly concave laterally; lateral margins subparallel, slightly rounded, serrate.Anterior angles slightly prominent, rounded; posterior angles almost rectangular. Prothoracic notosternal suture apparently complete. Prohypomera densely granulate. Prosternum convex, with dense and coarse punctation. Intercoxal prosternal process elongate, extending beyond posterior margin of procoxae, with rounded apical margin, slightly dilated in anterior one-third of length, with lateral margins emarginate, and rather wide, about 1.3× as wide as diameter of procoxa. Procoxal cavities open posteriorly.</paragraph>
<paragraph id="8BE26E2EFFEA6649FF006A9EB90456FE" blockId="10.[151,1437,151,249]" box="[199,591,223,249]" pageId="10" pageNumber="211">Scutellar shield minute, cordiform.</paragraph>
<caption id="DF223EA6FFEA6649FF506C39BAEA50B3" ID-DOI="http://doi.org/10.5281/zenodo.14033626" ID-Zenodo-Dep="14033626" httpUri="https://zenodo.org/record/14033626/files/figure.png" pageId="10" pageNumber="211" startId="10.[151,264,1656,1680]" targetBox="[151,1435,270,1630]" targetPageId="10" targetType="figure">
<paragraph id="8BE26E2EFFEA6649FF506C39BAEA50B3" blockId="10.[151,1436,1656,1716]" pageId="10" pageNumber="211">
<emphasis id="B929B23CFFEA6649FF506C39BA715097" bold="true" box="[151,314,1656,1680]" pageId="10" pageNumber="211">FIGURES 78.</emphasis>
<taxonomicName id="4C5D15ADFFEA6649FE876C39B9465097" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[320,525,1656,1680]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="vanivanitatum" status="sp. nov.">
<emphasis id="B929B23CFFEA6649FE876C39B9465097" box="[320,525,1656,1680]" italics="true" pageId="10" pageNumber="211">Paha vanivanitatum</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFEA6649FDD46C39B9285097" bold="true" box="[531,611,1656,1680]" pageId="10" pageNumber="211">
<taxonomicNameLabel id="A21A0F47FFEA6649FDD46C39B9285097" box="[531,611,1656,1680]" pageId="10" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, No. ACAB 064 [ACAB]: 7—habitus, dorsal view; 8—habitus, ventral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BE26E2EFFEA6649FF006CA3B90551FE" blockId="10.[151,1437,1762,2041]" pageId="10" pageNumber="211">Elytra almost parallel-sided in anterior three-quarters of length and tapered at apex, about 1.8× as long as wide combined, as wide as pronotum basally, convex, carinate, 2.7× longer than pronotum. Humeral angles rounded, weakly serrate. Elytra striate-punctate. Scutellary striole absent. Elytral striae with rows of elongate granules; odd elytral intervals (1, 3, 5, 7) carinate. Epipleura well-developed, widest in basal half, slightly narrowing posteriad, reaching elytral apices, densely covered with round granules. Meso- and metaventrites densely and coarsely punctate. Mesocoxal cavities closed. Metaventrite with disc almost flat, slightly longer than abdominal ventrite 1; discrimen distinct in posterior half. Relative length ratio of pro- to meso- to metaventrite to abdomen approximately equal to 10:6:9:27. Metathoracic wings present.</paragraph>
<caption id="DF223EA6FFEB6648FF506C5AB811507C" ID-DOI="http://doi.org/10.5281/zenodo.14033628" ID-Zenodo-Dep="14033628" httpUri="https://zenodo.org/record/14033628/files/figure.png" pageId="11" pageNumber="212" startId="11.[151,264,1563,1587]" targetBox="[151,1435,179,1538]" targetPageId="11" targetType="figure">
<paragraph id="8BE26E2EFFEB6648FF506C5AB811507C" blockId="11.[151,1436,1563,1660]" pageId="11" pageNumber="212">
<emphasis id="B929B23CFFEB6648FF506C5ABA035034" bold="true" box="[151,328,1563,1587]" pageId="11" pageNumber="212">FIGURES 910.</emphasis>
<taxonomicName id="4C5D15ADFFEB6648FE886C5DB9575034" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[335,540,1564,1588]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="vanivanitatum" status="sp. nov.">
<emphasis id="B929B23CFFEB6648FE886C5DB9575034" box="[335,540,1564,1588]" italics="true" pageId="11" pageNumber="212">Paha vanivanitatum</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFEB6648FDE46C5DB93F5033" bold="true" box="[547,628,1564,1588]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A21A0F47FFEB6648FDE46C5DB93F5033" box="[547,628,1564,1588]" pageId="11" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, No. ACAB 064 [ACAB]: 9—details of forebody, dorsal view; 10— prothorax, ventral view. Abbreviations: a2a10—antennomeres 210 respectively; lc—longitudinal carina; pc—procoxa; pf— profemur; pp—prosternal process; pt—protibia. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BE26E2EFFEB6648FF006CEBB81D519B" blockId="11.[151,1437,1706,1949]" pageId="11" pageNumber="212">Legs rather short and robust, finely punctate and sparsely setose. Procoxa small, nearly rounded, and widely separated by about 1.3× diameter of procoxa; mesocoxae subcircular, separated by 0.7× diameter of mesocoxa; metacoxae widely oval, transverse, narrowly separated by about 0.5× longitudinal diameter of metacoxa. Femora nearly spindle-shaped, slightly flattened, widened medially; femora slightly longer than tibiae. Tibiae slightly curved, with short, fine setae apically; protibiae without apical spurs. Tarsal formula 4-4-4; tarsomeres not dilated, with sparse, fine setation ventrally; tarsomere 4 longest, longer than tarsomeres 13 combined, slightly curved. Claws simple, large, about 0.5× as long as tarsomere 4, robust.</paragraph>
<paragraph id="8BE26E2EFFEC664FFF006AD6B88A571A" blockId="12.[151,1437,151,1113]" pageId="12" pageNumber="213">Abdomen with five visible ventrites, abdominal sutures distinct throughout length; densely covered with large punctation, distance between punctures distinctly smaller than diameter of one puncture. Relative length ratios of ventrites 15 equal to 11:7:5:4:5. Abdominal ventrite 1 with small, narrow, triangular intercoxal process with rounded apex. Abdominal ventrite 5 with widely rounded apical margin.</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFEC664FFF006B67B981543A" pageId="12" pageNumber="213" type="diagnosis">
<paragraph id="8BE26E2EFFEC664FFF006B67BFE257AA" blockId="12.[151,1437,151,1113]" pageId="12" pageNumber="213">
<emphasis id="B929B23CFFEC664FFF006B67BA845747" bold="true" box="[199,463,294,320]" pageId="12" pageNumber="213">Differential diagnosis.</emphasis>
<taxonomicName id="4C5D15ADFFEC664FFE106B66B9F05747" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[471,699,295,321]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="vanivanitatum" status="sp. nov.">
<emphasis id="B929B23CFFEC664FFE106B66B9F05747" box="[471,699,295,321]" italics="true" pageId="12" pageNumber="213">Paha vanivanitatum</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFEC664FFD046B66B8555746" bold="true" box="[707,798,295,321]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="A21A0F47FFEC664FFD046B66B8555746" box="[707,798,295,321]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from extant congeners in the combination of the following characters: larger body size (
<quantity id="4CA5C3CBFFEC664FFD8F6B0AB99B5762" box="[584,720,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="12" pageNumber="213" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.82.1 mm</quantity>
in
<taxonomicName id="4C5D15ADFFEC664FFD366B0AB8E55762" authorityName="Dajoz" authorityYear="1984" box="[753,942,331,357]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="guadalupensis">
<emphasis id="B929B23CFFEC664FFD366B0AB8E55762" box="[753,942,331,357]" italics="true" pageId="12" pageNumber="213">P. guadalupensis</emphasis>
</taxonomicName>
), pronotum slightly transverse, 1.3× as wide as long and widest at midlength (pronotum strongly transverse, twice as wide as long in
<taxonomicName id="4C5D15ADFFEC664FFBA46B2EBE54578E" authorityName="Dajoz" authorityYear="1984" box="[1123,1311,367,393]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="guadalupensis">
<emphasis id="B929B23CFFEC664FFBA46B2EBE54578E" box="[1123,1311,367,393]" italics="true" pageId="12" pageNumber="213">P. guadalupensis</emphasis>
</taxonomicName>
and widest near apical one-quarter of length in
<taxonomicName id="4C5D15ADFFEC664FFDEF6BD2B9ED57AA" baseAuthorityName="LeConte" baseAuthorityYear="1863" box="[552,678,403,429]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B929B23CFFEC664FFDEF6BD2B9ED57AA" box="[552,678,403,429]" italics="true" pageId="12" pageNumber="213">P. laticollis</emphasis>
</taxonomicName>
), and anterior pronotal angles less prominent.
</paragraph>
<paragraph id="8BE26E2EFFEC664FFF006BF6B981543A" blockId="12.[151,1437,151,1113]" pageId="12" pageNumber="213">
The new species can be easily distinguished from other extinct
<taxonomicName id="4C5D15ADFFEC664FFC486BF6BF4157D6" authorityName="Erichson" authorityYear="1842" box="[911,1034,439,465]" class="Insecta" family="Colydiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="family">Colydiinae</taxonomicName>
preserved in Baltic amber based on the combination of generic characters, most obviously in the presence of 10-segmented antennae with 1-segmented club, short and inconspicuous setation of pronotum and elytra, pronotal disc with elevated central area and two longitudinal carinae, and elytral intervals carinate.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFEC664FFF006807BA8354F6" pageId="12" pageNumber="213" type="etymology">
<paragraph id="8BE26E2EFFEC664FFF006807BA8354F6" blockId="12.[151,1437,151,1113]" pageId="12" pageNumber="213">
<emphasis id="B929B23CFFEC664FFF006807BAEB5467" bold="true" box="[199,416,582,608]" pageId="12" pageNumber="213">Derivatio nominis</emphasis>
. The specific epithet
<taxonomicName id="4C5D15ADFFEC664FFD5B6806B8775467" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[668,828,583,608]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="vanivanitatum">
<emphasis id="B929B23CFFEC664FFD5B6806B8775467" box="[668,828,583,608]" italics="true" pageId="12" pageNumber="213">vanivanitatum</emphasis>
</taxonomicName>
is derived from the Latin phrase “
<emphasis id="B929B23CFFEC664FFB176806BA185483" italics="true" pageId="12" pageNumber="213">Vanitas vanitatum et omnia vanitas</emphasis>
” (meaning “vanity of vanities, and all is vanity”), referring (1) to the emptiness of inclusions in amber, (2) to the allegorical genre of painting (Vanitas) which uses symbolism to show the transience of human life and the certainty of death, and (3) to the lack of immediate practical use from amber insects studies. The name is used as noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFEC664FFF0068BBB80755A2" pageId="12" pageNumber="213" type="distribution">
<paragraph id="8BE26E2EFFEC664FFF0068BBB80755A2" blockId="12.[151,1437,151,1113]" pageId="12" pageNumber="213">
<emphasis id="B929B23CFFEC664FFF0068BBB94F5512" bold="true" box="[199,516,762,789]" pageId="12" pageNumber="213">Distribution of congeners.</emphasis>
The genus
<taxonomicName id="4C5D15ADFFEC664FFD4968BAB9825512" authorityName="Dajoz" authorityYear="1984" box="[654,713,763,789]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFEC664FFD4968BAB9825512" box="[654,713,763,789]" italics="true" pageId="12" pageNumber="213">Paha</emphasis>
</taxonomicName>
occurs in Nearctic and Neotropical regions and is represented by four extant species (
<bibRefCitation id="EFCC13DFFFEC664FFE6C695EB97F553E" author="Sharp, D." box="[427,564,799,825]" pageId="12" pageNumber="213" pagination="443 - 488" refId="ref27795" refString="Sharp, D. (1894) Colydiidae. In: Godman, F. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana. Insecta. Coleoptera. Vol. 2. Part 1. Taylor and Francis, London, pp. 443 - 488." type="book chapter" year="1894">Sharp 1894</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFEC664FFD83695EB986553E" author="Dajoz, R." box="[580,717,799,825]" pageId="12" pageNumber="213" pagination="145 - 157" refId="ref25423" refString="Dajoz, R. (1984) Note sur quatre genres de Coleopteres Colydiidae: Nematidium Erichson, Monoedus Horn, Lobogestoria Reitter et Paha n. gen. Bulletin Mensuel de la Societe Linneenne de Lyon, 53 (4), 145 - 157. https: // doi. org / 10.3406 / linly. 1984.10633" type="journal article" year="1984">Dajoz 1984</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFEC664FFD1A695EB8D3553E" author="Ivie, M. A. &amp; Lord, N. P. &amp; Foley, I. A." box="[733,920,799,825]" pageId="12" pageNumber="213" pagination="755 - 788" refId="ref25945" refString="Ivie, M. A., Lord, N. P. &amp; Foley, I. A. (2016) Colydiine genera (Coleoptera: Zopheridae: Colydiinae) of the new world: a key and nomenclatural acts 30 years in the making. The Coleopterists Bulletin, 70 (4), 755 - 788. https: // doi. org / 10.1649 / 0010 - 065 X- 70.4.755" type="journal article" year="2016">
Ivie
<emphasis id="B929B23CFFEC664FFCD3695EB802553E" box="[788,841,799,825]" italics="true" pageId="12" pageNumber="213">et al</emphasis>
. 2016
</bibRefCitation>
):
<taxonomicName id="4C5D15ADFFEC664FFC68695EBE47553E" authority="Dajoz, 1984" authorityName="Dajoz" authorityYear="1984" box="[943,1292,799,825]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="guadalupensis">
<emphasis id="B929B23CFFEC664FFC68695EBF3A553E" box="[943,1137,799,825]" italics="true" pageId="12" pageNumber="213">P. guadalupensis</emphasis>
<bibRefCitation id="EFCC13DFFFEC664FFBBB695EBE47553E" author="Dajoz, R." box="[1148,1292,799,825]" pageId="12" pageNumber="213" pagination="145 - 157" refId="ref25423" refString="Dajoz, R. (1984) Note sur quatre genres de Coleopteres Colydiidae: Nematidium Erichson, Monoedus Horn, Lobogestoria Reitter et Paha n. gen. Bulletin Mensuel de la Societe Linneenne de Lyon, 53 (4), 145 - 157. https: // doi. org / 10.3406 / linly. 1984.10633" type="journal article" year="1984">Dajoz, 1984</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5D15ADFFEC664FFADC695EBA1F555A" authority="(LeConte, 1863)" baseAuthorityName="LeConte" baseAuthorityYear="1863" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B929B23CFFEC664FFADC695EBED7553E" box="[1307,1436,799,825]" italics="true" pageId="12" pageNumber="213">P. laticollis</emphasis>
(LeConte, 1863)
</taxonomicName>
,
<taxonomicName id="4C5D15ADFFEC664FFEA66902B9D7555A" authority="(Hinton, 1935)" baseAuthorityName="Hinton" baseAuthorityYear="1935" box="[353,668,834,861]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="B929B23CFFEC664FFEA66902BAA2555B" box="[353,489,835,860]" italics="true" pageId="12" pageNumber="213">P. mexicana</emphasis>
(Hinton, 1935)
</taxonomicName>
, and
<taxonomicName id="4C5D15ADFFEC664FFD1D6902B8B1555A" authority="(Sharp, 1894)" baseAuthorityName="Sharp" baseAuthorityYear="1894" box="[730,1018,835,861]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B929B23CFFEC664FFD1D6902B818555B" box="[730,851,835,860]" italics="true" pageId="12" pageNumber="213">P. mimetes</emphasis>
(
<bibRefCitation id="EFCC13DFFFEC664FFCA46902B8B9555A" author="Sharp, D." box="[867,1010,835,861]" pageId="12" pageNumber="213" pagination="443 - 488" refId="ref27795" refString="Sharp, D. (1894) Colydiidae. In: Godman, F. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana. Insecta. Coleoptera. Vol. 2. Part 1. Taylor and Francis, London, pp. 443 - 488." type="book chapter" year="1894">Sharp, 1894</bibRefCitation>
)
</taxonomicName>
. The extinct species described in the current paper indicates both the Eocene age of the genus and the presence of representatives of this genus in forest paleoecosystems of the Western Palearctic during the Eocene.
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<emphasis id="B929B23CFFEC664FFF0069EEBA8655CE" bold="true" box="[199,461,942,969]" pageId="12" pageNumber="213">Ecology of congeners.</emphasis>
The natural history of the genus is poorly understood. The single Nearctic species,
<taxonomicName id="4C5D15ADFFEC664FFA4F69EEBBB255EA" baseAuthorityName="LeConte" baseAuthorityYear="1863" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B929B23CFFEC664FFA4F69EEBBB255EA" italics="true" pageId="12" pageNumber="213">P. laticollis</emphasis>
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is associated with oaken logs and old oaks (
<taxonomicName id="4C5D15ADFFEC664FFD2A6992B80055EA" box="[749,843,979,1005]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="genus">
<emphasis id="B929B23CFFEC664FFD2A6992B80055EA" box="[749,843,979,1005]" italics="true" pageId="12" pageNumber="213">Quercus</emphasis>
</taxonomicName>
) and is also reported under bark of the fabacean tree
<taxonomicName id="4C5D15ADFFEC664FFF5069B6BAC15216" box="[151,394,1015,1041]" class="Magnoliopsida" family="Fabaceae" genus="Lysiloma" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="bahamensis">
<emphasis id="B929B23CFFEC664FFF5069B6BAC15216" box="[151,394,1015,1041]" italics="true" pageId="12" pageNumber="213">Lysiloma bahamensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCC13DFFFEC664FFE5C69B6B9075216" author="Blatchley, W. S." box="[411,588,1015,1041]" pageId="12" pageNumber="213" pagination="28 - 35" refId="ref24717" refString="Blatchley, W. S. (1930) Notes on the distribution of Coleoptera in Florida with additions to the known fauna of that state. The Canadian Entomologist, 62, 28 - 35. https: // doi. org / 10.4039 / Ent 6228 - 2" type="journal article" year="1930">Blatchley 1930</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFEC664FFD9D69B6B9B25216" author="Stephan, K. H." box="[602,761,1015,1041]" pageId="12" pageNumber="213" refId="ref28343" refString="Stephan, K. H. (1989) The Bothrideridae and Colydiidae of America north of Mexico (Coleoptera: Clavicornia and Heteromera. Occasional Papers of the Florida State Collection of Arthropods. Vol. 6. Florida Department of Agriculture and Consumer Services, Gainesville, Flordia, xii + 65 pp." type="book" year="1989">Stephan 1989</bibRefCitation>
). The ecology of three other species from Central America (described from
<collectingCountry id="F34A2EBEFFEC664FFE936E5ABAD55232" box="[340,414,1051,1077]" name="Belize" pageId="12" pageNumber="213">Belize</collectingCountry>
, Guadeloupe, and
<collectingCountry id="F34A2EBEFFEC664FFDB46E5AB9815232" box="[627,714,1051,1077]" name="Mexico" pageId="12" pageNumber="213">Mexico</collectingCountry>
) is unknown.
<taxonomicName id="4C5D15ADFFEC664FFCA96E5ABF195233" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[878,1106,1051,1077]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="vanivanitatum" status="sp. nov.">
<emphasis id="B929B23CFFEC664FFCA96E5ABF195233" box="[878,1106,1051,1077]" italics="true" pageId="12" pageNumber="213">Paha vanivanitatum</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFEC664FFB9C6E5ABFFC5232" bold="true" box="[1115,1207,1051,1077]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="A21A0F47FFEC664FFB9C6E5ABFFC5232" box="[1115,1207,1051,1077]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
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could possibly be a member of an Eocene forest community associated with fagacean wood.
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<mods:title id="DA6E589C9CBAD3482F7291ADA6F6606F">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests</mods:title>
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Genus
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<emphasis id="B929B23CFFEC664FFF296EEABA3252C2" bold="true" box="[238,377,1195,1221]" italics="true" pageId="12" pageNumber="213">Lasconotus</emphasis>
Erichson, 1845
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species:
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<taxonomicName id="4C5D15ADFFEC664FFEE16EB1BAD25300" authorityName="Erichson" authorityYear="1845" box="[294,409,1264,1287]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">Lasconotus</taxonomicName>
complex
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LeConte, 1859
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<emphasis id="B929B23CFFEC664FFF506F74BAF95348" bold="true" box="[151,434,1333,1359]" pageId="12" pageNumber="213">Taxonomic assignment.</emphasis>
The studied amber specimen shows a combination of characters corresponding to the tribe
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within the subfamily
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: (1) antennal insertions concealed dorsally; (2) antennae distinctly clubbed, finely and sparsely setose, lacking scale-like setae; (3) all tarsi tetramerous, not dilated; and (4) metacoxae separated by triangular process of abdominal ventrite 1. The new extinct species is placed in the genus
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based on: antennae 11-segmented with 3-segmented club; supraocular carina strong; finely facetted eyes large; subantennal grooves absent; antennomere 3 longer than antennomere 4; apex of protibia spinose; prosternal process not expanded apically beyond midpoint of procoxa; pronotum with paired sublateral carinae; elytra with distinct carinae; and abdominal ventrite 5 with deep preapical groove.
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<subSubSection id="C3473DA5FFF06653FF506CB8B9365114" box="[151,637,1785,1812]" pageId="16" pageNumber="217" type="nomenclature">
<paragraph id="8BE26E2EFFF06653FF506CB8B9365114" blockId="16.[151,637,1785,1812]" box="[151,637,1785,1812]" pageId="16" pageNumber="217">
<heading id="D0AAD942FFF06653FF506CB8B9365114" bold="true" box="[151,637,1785,1812]" fontSize="11" level="1" pageId="16" pageNumber="217" reason="1">
<emphasis id="B929B23CFFF06653FF506CB8B9365114" bold="true" box="[151,637,1785,1812]" pageId="16" pageNumber="217">
Genus
<taxonomicName id="4C5D15ADFFF06653FF296CB8B9365114" authority="Schauffuss, 1882" authorityName="Schauffuss" authorityYear="1882" box="[238,637,1785,1812]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF06653FF296CB8BAEE5113" bold="true" box="[238,421,1785,1812]" italics="true" pageId="16" pageNumber="217">Helioctamenus</emphasis>
Schauffuss, 1882
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFF06652FF506D7FB80C578E" lastPageId="17" lastPageNumber="218" pageId="16" pageNumber="217" type="materials_examined">
<paragraph id="8BE26E2EFFF06653FF506D7FB84A5151" blockId="16.[151,769,1854,1878]" box="[151,769,1854,1878]" pageId="16" pageNumber="217">
<typeStatus id="54E6D08CFFF06653FF506D7FBB825151" box="[151,201,1854,1878]" pageId="16" pageNumber="217">Type</typeStatus>
species:
<taxonomicName id="4C5D15ADFFF06653FEE16D7FB84A5151" authority="Schaufuss, 1882" authorityName="Schaufuss" authorityYear="1882" box="[294,769,1854,1878]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hippopotamus">
<emphasis id="B929B23CFFF06653FEE16D7FB91E5151" box="[294,597,1854,1878]" italics="true" pageId="16" pageNumber="217">Helioctamenus hippopotamus</emphasis>
<bibRefCitation id="EFCC13DFFFF06653FD9C6D7FB84A5151" author="Schaufuss, L. W." box="[603,769,1854,1878]" pageId="16" pageNumber="217" pagination="527 - 560" refId="ref27762" refString="Schaufuss, L. W. (1882) Zoologische Ergebnisse von Excursionen auf den Balearen. III (Addenda und Fortsetzung). Numquam Otiosus, 3, 527 - 560." type="journal article" year="1882">Schaufuss, 1882</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BE26E2EFFF06652FF506DC5B80C578E" blockId="16.[151,1436,1924,2022]" lastBlockId="17.[151,1436,150,393]" lastPageId="17" lastPageNumber="218" pageId="16" pageNumber="217">
<emphasis id="B929B23CFFF06653FF506DC5BAF35199" bold="true" box="[151,440,1924,1950]" pageId="16" pageNumber="217">Taxonomic assignment.</emphasis>
The specimen under consideration belongs to the tribe
<taxonomicName id="4C5D15ADFFF06653FBBD6DC5BFA65199" authorityName="Billberg" authorityYear="1820" box="[1146,1261,1924,1950]" class="Insecta" family="Zopheridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="tribe" tribe="Sarrotriini">Sarrotriini</taxonomicName>
based on the combination of the following characters: (1) dorsally concealed antennal insertions; (2) antennae setose, 10- or 11-segmented, without distinctly separated antennal club; (3) antennal insertions distant from eyes; (4) procoxal cavities open posteriorly; (5) all tarsi tetramerous, not dilated; (6) prosternal process wide, emarginate apically; and (7) metacoxae separated by intercoxal abdominal process with rounded apex. The tribe includes three genera from the Western Palearctic: monotypic
<taxonomicName id="4C5D15ADFFF16652FDE56A9EB86956FE" authority="Reitter, 1882" authorityName="Reitter" authorityYear="1882" box="[546,802,223,249]" class="Insecta" family="Zopheridae" genus="Diplagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF16652FDE56A9EB9CD56FE" box="[546,646,223,249]" italics="true" pageId="17" pageNumber="218">Diplagia</emphasis>
<bibRefCitation id="EFCC13DFFFF16652FD4A6A9EB86956FE" author="Reitter, E." box="[653,802,223,249]" pageId="17" pageNumber="218" pagination="113 - 149" refId="ref27502" refString="Reitter, E. (1882) Bestimmungs-Tabellen der europaischen Coleopteren. VI. Enhaltend die Familien: Colydiidae, Rhysodidae, Trogositidae. Verhandlungen des Naturforschenden Vereines in Brunn, 22, 113 - 149." type="journal article" year="1882">Reitter, 1882</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C5D15ADFFF16652FCE86A9EBF1656FE" authority="Latreille, 1796" authorityName="Latreille" authorityYear="1796" box="[815,1117,223,249]" class="Insecta" family="Zopheridae" genus="Orthocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF16652FCE86A9EB8E456FE" box="[815,943,223,249]" italics="true" pageId="17" pageNumber="218">Orthocerus</emphasis>
Latreille, 1796
</taxonomicName>
with two European species; and comparatively diverse
<taxonomicName id="4C5D15ADFFF16652FE096B42B8E1571A" authority="(Slipinski &amp; Schuh 2008)" baseAuthorityName="Slipinski &amp; Schuh" baseAuthorityYear="2008" box="[462,938,259,287]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF16652FE096B42B93D571A" box="[462,630,259,285]" italics="true" pageId="17" pageNumber="218">Helioctamenus</emphasis>
(
<bibRefCitation id="EFCC13DFFFF16652FD406B45B8E9571A" author="Slipinski, S. A. &amp; Schuh, R." box="[647,930,259,287]" pageId="17" pageNumber="218" refId="ref28217" refString="Slipinski, S. A. &amp; Schuh, R. (2008) Zopheridae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Apollo Books, Stenstrup, pp. 78 - 87." type="book" year="2008">Ślipiński &amp; Schuh 2008</bibRefCitation>
)
</taxonomicName>
. The new extinct species is assigned to the genus
<taxonomicName id="4C5D15ADFFF16652FF246B66BAC05746" baseAuthorityName="Slipinski &amp; Schuh" baseAuthorityYear="2008" box="[227,395,295,321]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF16652FF246B66BAC05746" box="[227,395,295,321]" italics="true" pageId="17" pageNumber="218">Helioctamenus</emphasis>
</taxonomicName>
based on: antennae sparsely and shortly setose (in contrast to
<taxonomicName id="4C5D15ADFFF16652FBAC6B66BF845746" authorityName="Reitter" authorityYear="1882" box="[1131,1231,295,321]" class="Insecta" family="Zopheridae" genus="Diplagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF16652FBAC6B66BF845746" box="[1131,1231,295,321]" italics="true" pageId="17" pageNumber="218">Diplagia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5D15ADFFF16652FACB6B66BEC75746" authorityName="Latreille" authorityYear="1796" box="[1292,1420,295,321]" class="Insecta" family="Zopheridae" genus="Orthocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF16652FACB6B66BEC75746" box="[1292,1420,295,321]" italics="true" pageId="17" pageNumber="218">Orthocerus</emphasis>
</taxonomicName>
), not fusiform (in contrast to
<taxonomicName id="4C5D15ADFFF16652FE1B6B0AB9175762" authorityName="Latreille" authorityYear="1796" box="[476,604,331,357]" class="Insecta" family="Zopheridae" genus="Orthocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF16652FE1B6B0AB9175762" box="[476,604,331,357]" italics="true" pageId="17" pageNumber="218">Orthocerus</emphasis>
</taxonomicName>
); interfacetal setae absent; body sparsely setose dorsally; and pronotum distinctly narrower than elytral base (in contrast to
<taxonomicName id="4C5D15ADFFF16652FD146B2EB87C578E" authorityName="Reitter" authorityYear="1882" box="[723,823,367,393]" class="Insecta" family="Zopheridae" genus="Diplagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF16652FD146B2EB87C578E" box="[723,823,367,393]" italics="true" pageId="17" pageNumber="218">Diplagia</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
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View file

@ -0,0 +1,156 @@
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<mods:title id="11DDE901A47E9490B3049762C8F5DFBE">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests</mods:title>
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<mods:namePart id="C28167E832BBEAF86B693DC2116EED40">Alekseev, Vitalii</mods:namePart>
<mods:affiliation id="559A0E19F60626F88C48034802F02393">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia. &amp; Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. &amp; Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; &amp; Department of Ecology &amp; Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
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<mods:namePart id="DB15A18D09B46B0DBB0C0F822A3267AB">Bukejs, Andris</mods:namePart>
<mods:affiliation id="6FE8C16EE627ACD5D6A8177155E1CE7F">Institute of Life Sciences and Technologies, Daugavpils University, Vienības 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
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<heading id="D0AAD942FFF76654FF506B2FB84D578E" bold="true" box="[151,774,366,393]" fontSize="11" level="1" pageId="23" pageNumber="224" reason="1">
<emphasis id="B929B23CFFF76654FF506B2FB84D578E" bold="true" box="[151,774,366,393]" pageId="23" pageNumber="224">
Genus
<taxonomicName id="4C5D15ADFFF76654FF296B2FB9DB578F" authority="Alekseev et Bukejs" authorityName="Alekseev et Bukejs" authorityYear="2024" box="[238,656,366,393]" class="Magnoliopsida" family="Fagaceae" genus="Thanatoplagia" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="224" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B929B23CFFF76654FF296B2FBAEB578E" bold="true" box="[238,416,366,393]" italics="true" pageId="23" pageNumber="224">Thanatoplagia</emphasis>
Alekseev et Bukejs
</taxonomicName>
<taxonomicNameLabel id="A21A0F47FFF76654FD5F6B2EB84D578E" box="[664,774,367,393]" pageId="23" pageNumber="224" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3473DA5FFF76654FF506BF5BA1354A4" pageId="23" pageNumber="224" type="materials_examined">
<paragraph id="8BE26E2EFFF76654FF506BF5B8BF57CC" blockId="23.[151,1012,435,460]" box="[151,1012,435,460]" pageId="23" pageNumber="224">
<typeStatus id="54E6D08CFFF76654FF506BF5BB8257CB" box="[151,201,436,460]" pageId="23" pageNumber="224">Type</typeStatus>
species:
<taxonomicName id="4C5D15ADFFF76654FEE16BF2B99657CC" authority="Alekseev et Bukejs" authorityName="Alekseev et Bukejs" authorityYear="2024" box="[294,733,435,459]" class="Magnoliopsida" family="Fagaceae" genus="Thanatoplagia" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="224" phylum="Tracheophyta" rank="species" species="tamutisi" status="sp. nov.">
<emphasis id="B929B23CFFF76654FEE16BF2B95F57CC" box="[294,532,435,459]" italics="true" pageId="23" pageNumber="224">Thanatoplagia tamutisi</emphasis>
Alekseev et Bukejs
</taxonomicName>
<taxonomicNameLabel id="A21A0F47FFF76654FD236BF2B87D57CC" box="[740,822,435,459]" pageId="23" pageNumber="224" rank="species">
<emphasis id="B929B23CFFF76654FD236BF2B86457CC" bold="true" box="[740,815,435,459]" pageId="23" pageNumber="224">sp. nov</emphasis>
.
</taxonomicNameLabel>
, designated herein
</paragraph>
<paragraph id="8BE26E2EFFF76654FF506BB8BA1354A4" blockId="23.[151,1437,505,1216]" pageId="23" pageNumber="224">
<emphasis id="B929B23CFFF76654FF506BB8BAE55414" bold="true" box="[151,430,505,531]" pageId="23" pageNumber="224">Taxonomic assignment.</emphasis>
The specimen under study belongs to the tribe
<taxonomicName id="4C5D15ADFFF76654FC796BB8BF7A5414" authorityName="Billberg" authorityYear="1820" box="[958,1073,505,531]" class="Insecta" family="Zopheridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="tribe" tribe="Sarrotriini">Sarrotriini</taxonomicName>
based on the combination of the following characters: (1) dorsally concealed antennal insertions; (2) antennae setose, stout, 10- or 11-segmeneted, without clearly separated antennal club; (3) antennal insertions distant from eyes; (4) procoxal cavities open posteriorly; (5) all tarsi tetramerous, not dilated; and (6) metacoxae separated by intercoxal abdominal process rounded apically.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFF76654FF0068ECBEC9520C" pageId="23" pageNumber="224" type="diagnosis">
<paragraph id="8BE26E2EFFF76654FF0068ECBEC9520C" blockId="23.[151,1437,505,1216]" pageId="23" pageNumber="224">
<emphasis id="B929B23CFFF76654FF0068ECBA8654C0" bold="true" box="[199,461,685,711]" pageId="23" pageNumber="224">Differential diagnosis.</emphasis>
The tribe
<taxonomicName id="4C5D15ADFFF76654FDF868ECB9F954C0" authorityName="Billberg" authorityYear="1820" box="[575,690,685,711]" class="Insecta" family="Zopheridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="tribe" tribe="Sarrotriini">Sarrotriini</taxonomicName>
includes three extant genera from the Western Palearctic:
<taxonomicName id="4C5D15ADFFF76654FAFE68ECBA7A54EC" authority="Reitter, 1882" authorityName="Reitter" authorityYear="1882" class="Insecta" family="Zopheridae" genus="Diplagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF76654FAFE68ECBED654C0" box="[1337,1437,685,711]" italics="true" pageId="23" pageNumber="224">Diplagia</emphasis>
<bibRefCitation id="EFCC13DFFFF76654FF506890BA7A54EC" author="Reitter, E." box="[151,305,721,747]" pageId="23" pageNumber="224" pagination="113 - 149" refId="ref27502" refString="Reitter, E. (1882) Bestimmungs-Tabellen der europaischen Coleopteren. VI. Enhaltend die Familien: Colydiidae, Rhysodidae, Trogositidae. Verhandlungen des Naturforschenden Vereines in Brunn, 22, 113 - 149." type="journal article" year="1882">Reitter, 1882</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5D15ADFFF76654FE846890B93654EC" authority="Latreille, 1796" authorityName="Latreille" authorityYear="1796" box="[323,637,721,747]" class="Insecta" family="Zopheridae" genus="Orthocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF76654FE846890BA8854EC" box="[323,451,721,747]" italics="true" pageId="23" pageNumber="224">Orthocerus</emphasis>
Latreille, 1796
</taxonomicName>
, and
<taxonomicName id="4C5D15ADFFF76654FD036890BE7D54EC" authority="Schauffuss (Slipinski &amp; Schuh 2008)" authorityName="Schauffuss (Slipinski &amp; Schuh" authorityYear="2008" box="[708,1334,721,750]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF76654FD036890B82754EC" box="[708,876,721,747]" italics="true" pageId="23" pageNumber="224">Helioctamenus</emphasis>
Schauffuss (
<bibRefCitation id="EFCC13DFFFF76654FBCF6892BE6554EC" author="Slipinski, S. A. &amp; Schuh, R." box="[1032,1326,721,750]" pageId="23" pageNumber="224" refId="ref28217" refString="Slipinski, S. A. &amp; Schuh, R. (2008) Zopheridae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Apollo Books, Stenstrup, pp. 78 - 87." type="book" year="2008">Ślipiński &amp; Schuh 2008</bibRefCitation>
)
</taxonomicName>
. Despite the apparent similarity, we conclude that the studied fossil cannot be attributed to any of these genera, and thus we propose a new genus for this fossil beetle.
<taxonomicName id="4C5D15ADFFF76654FD756958B81C5534" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[690,855,793,819]" class="Magnoliopsida" family="Fagaceae" genus="Thanatoplagia" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="224" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B929B23CFFF76654FD756958B81C5534" box="[690,855,793,819]" italics="true" pageId="23" pageNumber="224">Thanatoplagia</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFF76654FCA66958B8875534" bold="true" box="[865,972,793,819]" pageId="23" pageNumber="224">
<taxonomicNameLabel id="A21A0F47FFF76654FCA66958B8875534" box="[865,972,793,819]" pageId="23" pageNumber="224" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
clearly differs from
<taxonomicName id="4C5D15ADFFF76654FB7A6958BE6A5534" authorityName="Reitter" authorityYear="1882" box="[1213,1313,793,819]" class="Insecta" family="Zopheridae" genus="Diplagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF76654FB7A6958BE6A5534" box="[1213,1313,793,819]" italics="true" pageId="23" pageNumber="224">Diplagia</emphasis>
</taxonomicName>
in lacking interfacetal setae, but possessing sparsely setose antennae, and pronotum distinctly narrower than elytral base; differs from
<taxonomicName id="4C5D15ADFFF76654FEE16920BAED557C" authorityName="Latreille" authorityYear="1796" box="[294,422,865,891]" class="Insecta" family="Zopheridae" genus="Orthocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF76654FEE16920BAED557C" box="[294,422,865,891]" italics="true" pageId="23" pageNumber="224">Orthocerus</emphasis>
</taxonomicName>
in antennal structure (gradually widened towards apex, not fusiform, sparsely and shortly setose in the new genus); and it differs from
<taxonomicName id="4C5D15ADFFF76654FD5469C4B8705598" authorityName="Schauffuss (Slipinski &amp; Schuh" authorityYear="2008" box="[659,827,901,927]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF76654FD5469C4B8705598" box="[659,827,901,927]" italics="true" pageId="23" pageNumber="224">Helioctamenus</emphasis>
</taxonomicName>
in having stout and strongly transverse antennomeres 49, and pronotum with three distinct impressions (two oval transverse, shallow impressions laterobasally and one large, rounded impression anteromedially). The new fossil genus differs from other similar sarrotriines in a well-developed antennomere 11 (as long as antennomere 10), pronounced humeri and developed metathoracic wings.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFF76654FF006E54B9EF5270" pageId="23" pageNumber="224" type="etymology">
<paragraph id="8BE26E2EFFF76654FF006E54B9EF5270" blockId="23.[151,1437,505,1216]" pageId="23" pageNumber="224">
<emphasis id="B929B23CFFF76654FF006E54BA055228" bold="true" box="[199,334,1045,1071]" pageId="23" pageNumber="224">Etymology.</emphasis>
The name of the new genus is a compound word and combines the old Greek word θάνατος (
<emphasis id="B929B23CFFF76654FF666E78BA4A5254" box="[161,257,1081,1107]" italics="true" pageId="23" pageNumber="224">thánatos</emphasis>
, death) and the old Greek word πλαγία (
<emphasis id="B929B23CFFF76654FD086E78B85C5254" box="[719,791,1081,1107]" italics="true" pageId="23" pageNumber="224">plagía</emphasis>
, slope or hillside) referring to
<taxonomicName id="4C5D15ADFFF76654FBB56E78BF9D5254" authorityName="Reitter" authorityYear="1882" box="[1138,1238,1081,1107]" class="Insecta" family="Zopheridae" genus="Diplagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFF76654FBB56E78BF9D5254" box="[1138,1238,1081,1107]" italics="true" pageId="23" pageNumber="224">Diplagia</emphasis>
</taxonomicName>
, the extant genus of the tribe
<taxonomicName id="4C5D15ADFFF76654FEDE6E1CBADB5270" authorityName="Billberg" authorityYear="1820" box="[281,400,1117,1143]" class="Insecta" family="Zopheridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="tribe" tribe="Sarrotriini">Sarrotriini</taxonomicName>
. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFF76654FF006EC0BA6C52B8" pageId="23" pageNumber="224" type="description">
<paragraph id="8BE26E2EFFF76654FF006EC0BA6C52B8" blockId="23.[151,1437,505,1216]" pageId="23" pageNumber="224">
<emphasis id="B929B23CFFF76654FF006EC0BA12529C" bold="true" box="[199,345,1153,1179]" pageId="23" pageNumber="224">Description.</emphasis>
The new genus is monotypic. Therefore, the generic description considerably overlaps that of the
<typeStatus id="54E6D08CFFF76654FF506EE7BB8C52C7" box="[151,199,1190,1216]" pageId="23" pageNumber="224">type</typeStatus>
species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,191 @@
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<mods:title id="DC45A1A70C98DC2E5FA0AAB1E729B36C">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests</mods:title>
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<mods:namePart id="687AF43756280B9CF00E8FEC10E07EFD">Alekseev, Vitalii</mods:namePart>
<mods:affiliation id="1B6ABC3FEFC00BFD71D65E725401C18D">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia. &amp; Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. &amp; Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; &amp; Department of Ecology &amp; Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
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<mods:namePart id="9E9F630D895A2119B50CE5B9AA96FD11">Bukejs, Andris</mods:namePart>
<mods:affiliation id="C21CA7828231E44B48B5C803D7D06548">Institute of Life Sciences and Technologies, Daugavpils University, Vienības 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
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<subSubSection id="C3473DA5FFF76654FF506F50B842532C" box="[151,777,1297,1324]" pageId="23" pageNumber="224" type="nomenclature">
<paragraph id="8BE26E2EFFF76654FF506F50B842532C" blockId="23.[151,777,1297,1359]" box="[151,777,1297,1324]" pageId="23" pageNumber="224">
<heading id="D0AAD942FFF76654FF506F50B842532C" bold="true" box="[151,777,1297,1324]" fontSize="11" level="1" pageId="23" pageNumber="224" reason="1">
<emphasis id="B929B23CFFF76654FF506F50B842532C" bold="true" box="[151,777,1297,1324]" pageId="23" pageNumber="224">
<taxonomicName id="4C5D15ADFFF76654FF506F50B9EF532C" authority="Alekseev et Bukejs" authorityName="Alekseev et Bukejs" authorityYear="2024" box="[151,676,1297,1324]" class="Magnoliopsida" family="Fagaceae" genus="Thanatoplagia" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="224" phylum="Tracheophyta" rank="species" species="tamutisi" status="sp. nov.">
<emphasis id="B929B23CFFF76654FF506F50BAF9532C" bold="true" box="[151,434,1297,1324]" italics="true" pageId="23" pageNumber="224">Thanatoplagia tamutisi</emphasis>
Alekseev et Bukejs
</taxonomicName>
<taxonomicNameLabel id="A21A0F47FFF76654FD6C6F50B842532C" box="[683,777,1297,1323]" pageId="23" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFF76654FF506F74BA6C5348" box="[151,295,1333,1359]" pageId="23" pageNumber="224" type="description">
<paragraph id="8BE26E2EFFF76654FF506F74BA6C5348" blockId="23.[151,777,1297,1359]" box="[151,295,1333,1359]" pageId="23" pageNumber="224">
(
<figureCitation id="136672ABFFF76654FF586F74BA545348" box="[159,287,1333,1359]" captionStart="FIGURES 2729" captionStartId="24.[151,264,1943,1967]" captionTargetBox="[233,1354,179,1916]" captionTargetId="figure-21@24.[233,1354,179,1916]" captionTargetPageId="24" captionText="FIGURES 2729. Thanatoplagia tamutisi gen. et sp. nov., holotype, No. ACAB 063 [ACAB]: 27—habitus, dorsal view; 28—habitus, ventral view; 29—habitus, right lateral view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14033667" httpUri="https://zenodo.org/record/14033667/files/figure.png" pageId="23" pageNumber="224">Figs 2729</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFF76654FF506F3CBBA250B0" pageId="23" pageNumber="224" type="materials_examined">
<paragraph id="8BE26E2EFFF76654FF506F3CBBAD5020" blockId="23.[151,1436,1405,2008]" pageId="23" pageNumber="224">
<emphasis id="B929B23CFFF76654FF506F3CBA0D5390" bold="true" box="[151,326,1405,1431]" pageId="23" pageNumber="224">Type material.</emphasis>
<materialsCitation id="3B356473FFF76654FE886F3CBBAD5020" collectionCode="ACAB, A" collectorName="No. &amp; Marius Veta &amp; Syninclusions" pageId="23" pageNumber="224" specimenCode="ABAC 063" specimenCount="2" typeStatus="holotype">
<typeStatus id="54E6D08CFFF76654FE886F3CBAF75390" box="[335,444,1405,1431]" pageId="23" pageNumber="224" type="holotype">Holotype</typeStatus>
: No.
<specimenCode id="DBFBC655FFF76654FE306F3CB93C5390" box="[503,631,1405,1431]" collectionCode="ABAC" pageId="23" pageNumber="224">ABAC 063</specimenCode>
[
<collectionCode id="ED4CF6EBFFF76654FD416F3CB9925390" box="[646,729,1405,1431]" pageId="23" pageNumber="224">ACAB</collectionCode>
] (
<collectorName id="26A80BF8FFF76654FD366F3CB8575390" box="[753,796,1405,1431]" pageId="23" pageNumber="224">No.</collectorName>
3205 ex. coll.
<collectorName id="26A80BF8FFF76654FC006F3CBF1D5390" box="[967,1110,1405,1431]" pageId="23" pageNumber="224">Marius Veta</collectorName>
); “
<typeStatus id="54E6D08CFFF76654FBB36F3CBF945390" box="[1140,1247,1405,1431]" pageId="23" pageNumber="224" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C5D15ADFFF76654FB306F3CBBBA53BC" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" class="Magnoliopsida" family="Fagaceae" genus="Thanatoplagia" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="224" phylum="Tracheophyta" rank="species" species="tamutisi" status="sp. nov.">
<emphasis id="B929B23CFFF76654FB306F3CBBBA53BC" italics="true" pageId="23" pageNumber="224">Thanatoplagia tamutisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A0F47FFF76654FF3D6FE3BA1953BB" box="[250,338,1442,1468]" pageId="23" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
/ Alekseev et Bukejs des. 2024” [red printed label]; adult, sex unknown.
<collectionCode id="ED4CF6EBFFF76654FB526FE3BFE353BB" box="[1173,1192,1442,1468]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="23" pageNumber="224" type="Herbarium">A</collectionCode>
complete beetle with partially exposed distal part of metathoracic wings included in a transparent, minute yellow amber piece with dimensions of 6×
<quantity id="4CA5C3CBFFF76654FE906FA8BADE5003" box="[343,405,1513,1540]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="224" unit="mm" value="3.0">3 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFF76654FD1D6FA8B8505003" box="[730,795,1513,1540]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="224" unit="mm" value="3.0">3 mm</quantity>
, preserved without supplementary fixation.
<collectorName id="26A80BF8FFF76654FB386FA8BED75004" box="[1279,1436,1513,1539]" pageId="23" pageNumber="224">Syninclusions</collectorName>
absent.
</materialsCitation>
</paragraph>
<paragraph id="8BE26E2EFFF76654FF006C70BAEF5068" blockId="23.[151,1436,1405,2008]" pageId="23" pageNumber="224">
<emphasis id="B929B23CFFF76654FF006C70BA38504C" bold="true" box="[199,371,1585,1611]" pageId="23" pageNumber="224">
<typeStatus id="54E6D08CFFF76654FF006C70BA49504C" box="[199,258,1585,1611]" pageId="23" pageNumber="224">Type</typeStatus>
stratum.
</emphasis>
Baltic amber from Eocene amber-bearing Blaue Erde deposits; estimated age: middlelate Eocene (Standke 1998).
</paragraph>
<paragraph id="8BE26E2EFFF76654FF006C38BBA250B0" blockId="23.[151,1436,1405,2008]" pageId="23" pageNumber="224">
<materialsCitation id="3B356473FFF76654FF006C38BBAE50B0" country="Russia" county="Peninsula" location="Yantarny" municipality="Sambian" pageId="23" pageNumber="224" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B929B23CFFF76654FF006C38BA2E5094" bold="true" box="[199,357,1657,1683]" pageId="23" pageNumber="224">
<typeStatus id="54E6D08CFFF76654FF006C38BA495094" box="[199,258,1657,1683]" pageId="23" pageNumber="224">Type</typeStatus>
locality.
</emphasis>
<location id="8E8238F5FFF76654FEAD6C3BBA845093" LSID="urn:lsid:plazi:treatment:03F4DF38FFF7665AFF506F50BE165072:8E8238F5FFF76654FEAD6C3BBA845093" box="[362,463,1658,1684]" country="Russia" county="Peninsula" municipality="Sambian" name="Yantarny" pageId="23" pageNumber="224" stateProvince="Kaliningrad">Yantarny</location>
settlement (formerly Palmnicken),
<collectingMunicipality id="6B86F454FFF76654FC9C6C38B8F65094" box="[859,957,1657,1683]" pageId="23" pageNumber="224">Sambian</collectingMunicipality>
(Samland)
<collectingCounty id="628316A2FFF76654FBF96C38BFE55094" box="[1086,1198,1657,1683]" pageId="23" pageNumber="224">Peninsula</collectingCounty>
,
<collectingRegion id="4999A0CCFFF76654FB7F6C38BED35094" box="[1208,1432,1657,1683]" country="Russia" name="Kaliningrad" pageId="23" pageNumber="224">Kaliningrad Region</collectingRegion>
,
<collectingCountry id="F34A2EBEFFF76654FF506CDCBBAE50B0" box="[151,229,1693,1719]" name="Russia" pageId="23" pageNumber="224">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFF7665AFF006C80BE1A53E2" lastPageId="25" lastPageNumber="226" pageId="23" pageNumber="224" type="description">
<paragraph id="8BE26E2EFFF76654FF006C80BA39514F" blockId="23.[151,1436,1405,2008]" pageId="23" pageNumber="224">
<emphasis id="B929B23CFFF76654FF006C80BA1A50DC" bold="true" box="[199,337,1729,1755]" pageId="23" pageNumber="224">Description</emphasis>
. Measurements: total body length (including visible part of head)
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, maximum body width (across elytra)
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; head length
<quantity id="4CA5C3CBFFF76654FDEA6CA4B9CD5107" box="[557,646,1765,1792]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="224" unit="mm" value="0.5">0.5 mm</quantity>
, head maximum width (across eyes)
<quantity id="4CA5C3CBFFF76654FBEF6CA4BFCA5107" box="[1064,1153,1765,1792]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="224" unit="mm" value="0.6">0.6 mm</quantity>
; pronotal length
<quantity id="4CA5C3CBFFF76654FAF86CA4BED35107" box="[1343,1432,1765,1792]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="224" unit="mm" value="0.7">0.7 mm</quantity>
, maximum pronotal width
<quantity id="4CA5C3CBFFF76654FE016D48B9695123" box="[454,546,1801,1828]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="224" unit="mm" value="0.8">0.8 mm</quantity>
, basal pronotal width
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; elytral length (along elytral suture, including scutellum)
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.
</paragraph>
<paragraph id="8BE26E2EFFF76654FF006D10BA0251DF" blockId="23.[151,1436,1405,2008]" pageId="23" pageNumber="224">Body subparallel-sided, elongate, about 2.2× as long as wide, moderately convex dorsally and weakly convex ventrally, unicolorous black (as preserved). Pubescence: head, pronotum and elytra sparsely covered with uniform, short, strongly curved, semierect setae that are not forming patches; ventral side of specimen with fine, inconspicuous, semierect setae.</paragraph>
<caption id="DF223EA6FFF8665BFF506DD6B88851D4" ID-DOI="http://doi.org/10.5281/zenodo.14033667" ID-Zenodo-Dep="14033667" httpUri="https://zenodo.org/record/14033667/files/figure.png" pageId="24" pageNumber="225" startId="24.[151,264,1943,1967]" targetBox="[233,1354,179,1916]" targetPageId="24" targetType="figure">
<paragraph id="8BE26E2EFFF8665BFF506DD6B88851D4" blockId="24.[151,1436,1943,2003]" pageId="24" pageNumber="225">
<emphasis id="B929B23CFFF8665BFF506DD6BA1351A8" bold="true" box="[151,344,1943,1967]" pageId="24" pageNumber="225">FIGURES 2729.</emphasis>
<taxonomicName id="4C5D15ADFFF8665BFEA66DD6B91951A8" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[353,594,1943,1967]" class="Magnoliopsida" family="Fagaceae" genus="Thanatoplagia" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="225" phylum="Tracheophyta" rank="species" species="tamutisi" status="gen. et sp. nov.">
<emphasis id="B929B23CFFF8665BFEA66DD6B91951A8" box="[353,594,1943,1967]" italics="true" pageId="24" pageNumber="225">Thanatoplagia tamutisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A0F47FFF8665BFD9C6DD6B84A51A8" box="[603,769,1943,1967]" pageId="24" pageNumber="225" rank="species">
<emphasis id="B929B23CFFF8665BFD9C6DD6B9C351A8" bold="true" box="[603,648,1943,1967]" pageId="24" pageNumber="225">gen.</emphasis>
et
<emphasis id="B929B23CFFF8665BFD6A6DD6B84A51A8" bold="true" box="[685,769,1943,1967]" pageId="24" pageNumber="225">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, No. ACAB 063 [ACAB]: 27—habitus, dorsal view; 28—habitus, ventral view; 29—habitus, right lateral view. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BE26E2EFFF9665AFF006AD6BAEB543A" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="226">Head prognathous, transverse; densely granulose dorsally, with granules rather large, about 2× as wide as diameter of one eye facet, and distance between granules distinctly smaller than diameter of one granule. Anterior margin of clypeus rounded.Compound eyes oval, convex, without interfacetal setation.Antennal insertions concealed dorsally; antennal grooves ventrad eye absent. Antennae stout, gradually widened towards apex, short, extending to middle of pronotum; 11-segmented, without clearly separated antennal club; sparsely covered with short, semierect setae, antennomeres 10 and 11 additionally with fine pubescence; densely punctate; scape wide, subcylindrical; pedicel cylindrical, nearly as long as wide, slightly narrower and shorter than scape; antennomere 3 subtrapezoidal, slightly dilated apically, nearly as large as pedicel; antennomeres 49 trapezoidal, strongly transverse, subequal in size and shape; antennomere 10 trapezoidal, dilated apically, strongly transverse, about 2.0× as wide as long, slightly wider than antennomere 9; antennomere 11 widely oval, transverse, 1.3× as wide as long, widely rounded apically, 1.1× as wide as antennomere 10. Terminal maxillary palpomere obliquely truncate, about 2.0× as long as penultimate palpomere.</paragraph>
<paragraph id="8BE26E2EFFF9665AFF006806BAFB5586" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="226">Pronotum slightly transverse, 1.14× as wide as long, subparallel, weakly narrowed posteriad, widest at anterior margin; maximum pronotal width / basal pronotal width = 1.0; pronotal disc convex, with large, deep, oval impression medially and two shallow, oval, transverse impressions laterobasally; lateral sides narrowly explanate; pronotal surface densely covered with granules. Lateral margins almost straight, crenulate; anterior margin arcuate; posterior margin rounded. Anterior pronotal angles nearly rectangular, slightly rounded; posterior angles obtuse. Prothoracic notosternal suture complete. Hypomera and prosternum densely and coarsely granulate, distance between granules smaller than diameter of one granule. Intercoxal prosternal process elongate, extending beyond posterior margin of procoxae, with rounded apical margin, about 1.2× as wide as diameter of procoxa. Procoxal cavities open posteriorly.</paragraph>
<paragraph id="8BE26E2EFFF9665AFF0069CAB9C455A2" blockId="25.[151,1437,151,1653]" box="[199,655,907,933]" pageId="25" pageNumber="226">Scutellar shield minute, oval, transverse.</paragraph>
<paragraph id="8BE26E2EFFF9665AFF0069EEB9E052A6" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="226">Elytra almost parallel-sided within anterior two-thirds of their lengths, tapered at apex, about 1.4× as long as wide combined, convex, 2.6× longer than pronotum, distinctly wider than pronotal posterior margin. Humeral angles rounded. Elytra striate-punctate. Scutellary striole absent. Intervals convex. Epipleura well-developed, widest in basal half of length, slightly narrowed posteriad, reaching elytral apices, densely covered with granules. Mesocoxal cavities closed. Metaventrites with disc slightly convex, densely granulate, distinctly longer than abdominal ventrite 1; discrimen distinct in posterior half. Metanepisternum narrow, 6.3× longer than maximum width, with emarginate lateral margins. Metathoracic wings developed.</paragraph>
<paragraph id="8BE26E2EFFF9665AFF006EEABFCC537E" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="226">Legs rather short and robust, finely pubescent. Procoxa small, nearly rounded, separated by about 1.2× diameter of procoxa; mesocoxae hemispherical, separated by 0.7× diameter of mesocoxa; metacoxae oval, transverse, narrowly separated by about 0.5× longitudinal diameter of metacoxa. Femora elongate oval in shape, slightly dilated medially; femora and tibiae subequal in length. Tibiae straight, slightly dilated apically. Tarsal formula 4-4- 4; tarsomeres not dilated, with sparse, fine setation ventrally; tarsomere 4 longest, longer than metatarsomeres 13 combined, slightly curved. Claws simple, large, about 0.3× as long as tarsomere 4, robust.</paragraph>
<paragraph id="8BE26E2EFFF9665AFF006FC2BE1A53E2" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="226">Abdomen with five visible, similarly articulated ventrites, abdominal sutures distinct throughout length; densely covered with granules. Relative length ratios of ventrites 15 equal to 35:25:20:18:15. Abdominal ventrite 1 with triangular, apically rounded intercoxal process. Abdominal ventrite 5 with widely rounded posterior margin.</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFF9665AFF006FAFB829500E" box="[199,866,1518,1545]" pageId="25" pageNumber="226" type="diagnosis">
<paragraph id="8BE26E2EFFF9665AFF006FAFB829500E" blockId="25.[151,1437,151,1653]" box="[199,866,1518,1545]" pageId="25" pageNumber="226">
<emphasis id="B929B23CFFF9665AFF006FAFBA85500F" bold="true" box="[199,462,1518,1544]" pageId="25" pageNumber="226">Differential diagnosis.</emphasis>
As stated above, for the new genus.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFF9665AFF006C53BE165072" pageId="25" pageNumber="226" type="etymology">
<paragraph id="8BE26E2EFFF9665AFF006C53BE165072" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="226">
<emphasis id="B929B23CFFF9665AFF006C53BAD6502B" bold="true" box="[199,413,1554,1580]" pageId="25" pageNumber="226">Derivatio nominis</emphasis>
. The epithet of this new species is patronymic. The species is named in honor of the Baltic coleopterist, leading researcher of Lithuanian beetles, and our colleague Dr. Vytautas Tamutis (
<collectingRegion id="4999A0CCFFF9665AFB7A6C76BE5E5056" box="[1213,1301,1591,1617]" country="Lithuania" name="Kauno" pageId="25" pageNumber="226">Kaunas</collectingRegion>
,
<collectingCountry id="F34A2EBEFFF9665AFAE66C76BEDA5056" box="[1313,1425,1591,1617]" name="Lithuania" pageId="25" pageNumber="226">Lithuania</collectingCountry>
), for his numerous contributions to the study of entomofauna of
<collectingCountry id="F34A2EBEFFF9665AFC916C1AB88A5072" box="[854,961,1627,1653]" name="Lithuania" pageId="25" pageNumber="226">Lithuania</collectingCountry>
and beetles of the Baltic Sea region.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,135 @@
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<mods:title id="E9E2DECDA3CDC15EEC8847AB296303C3">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests</mods:title>
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<mods:affiliation id="B1B4BB98ABC2AAB56CE7D03750B5B1E5">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia. &amp; Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. &amp; Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; &amp; Department of Ecology &amp; Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
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<mods:namePart id="4EC824FF32FFAC3327E8AEB903288FBC">Bukejs, Andris</mods:namePart>
<mods:affiliation id="5909069DD51D68D7AF45B6106717C312">Institute of Life Sciences and Technologies, Daugavpils University, Vienības 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
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<paragraph id="8BE26E2EFFFA6659FF506AD6B98056B6" blockId="26.[151,715,151,213]" box="[151,715,151,178]" pageId="26" pageNumber="227">
<heading id="D0AAD942FFFA6659FF506AD6B98056B6" bold="true" box="[151,715,151,178]" fontSize="11" level="1" pageId="26" pageNumber="227" reason="1">
<taxonomicName id="4C5D15ADFFFA6659FF506AD6B98056B6" authority="Alekseev et Bukejs, 2016" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[151,715,151,178]" class="Insecta" family="Zopheridae" genus="Endophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="gorskii">
<emphasis id="B929B23CFFFA6659FF506AD6B98056B6" bold="true" box="[151,715,151,178]" pageId="26" pageNumber="227">
<emphasis id="B929B23CFFFA6659FF506AD6BADA56B5" bold="true" box="[151,401,151,178]" italics="true" pageId="26" pageNumber="227">Endophloeus gorskii</emphasis>
<bibRefCitation id="EFCC13DFFFFA6659FE506AD6B98056B6" author="Alekseev, V. I. &amp; Alekseev, P. I." box="[407,715,151,177]" pageId="26" pageNumber="227" pagination="75 - 86" refId="ref23984" refString="Alekseev, V. I. &amp; Alekseev, P. I. (2016) New approaches for reconstruction of the ecosystem of an Eocene amber forest. Biology Bulletin, 43 (1), 75 - 86. https: // doi. org / 10.1134 / S 1062359016010027" type="journal article" year="2016">Alekseev et Bukejs, 2016</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3473DA5FFFA6659FF506AFABA6C56D2" box="[151,295,187,213]" pageId="26" pageNumber="227" type="description">
<paragraph id="8BE26E2EFFFA6659FF506AFABA6C56D2" blockId="26.[151,715,151,213]" box="[151,295,187,213]" pageId="26" pageNumber="227">
(
<figureCitation id="136672ABFFFA6659FF586AFABA5456D2" box="[159,287,187,213]" captionStart="FIGURES 3032" captionStartId="26.[151,264,1539,1563]" captionTargetBox="[151,1435,703,1514]" captionTargetId="figure-200@26.[151,1435,703,1514]" captionTargetPageId="26" captionText="FIGURES 3032. Endophloeus gorskii Alekseev et Bukejs, 2016 in Baltic amber: 30—specimen No. JDC-9315, habitus, dorsal view; 31—idem, habitus, ventral view; 32—specimen No. ABAC 063 [ACAB], habitus, dorsal view. Scale bars = 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.14033669" httpUri="https://zenodo.org/record/14033669/files/figure.png" pageId="26" pageNumber="227">Figs 3032</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFFA6659FF506B42BFC054AD" pageId="26" pageNumber="227" type="materials_examined">
<paragraph id="8BE26E2EFFFA6659FF506B42BBB5578E" blockId="26.[151,1437,259,682]" pageId="26" pageNumber="227">
<emphasis id="B929B23CFFFA6659FF506B42BAC8571A" bold="true" box="[151,387,259,285]" pageId="26" pageNumber="227">Material examined.</emphasis>
<materialsCitation id="3B356473FFFA6659FE4B6B42BBB1578E" collectionCode="JDC" pageId="26" pageNumber="227" specimenCount="1">
<specimenCount id="9D5BA5A7FFFA6659FE4B6B42B965571A" box="[396,558,259,285]" count="1" pageId="26" pageNumber="227" type="generic">One specimen</specimenCount>
with collection number JDC-9315 [
<collectionCode id="ED4CF6EBFFFA6659FC0B6B42BF4E571A" box="[972,1029,259,285]" pageId="26" pageNumber="227">JDC</collectionCode>
], Baltic amber; adult, sex unknown (
<figureCitation id="136672ABFFFA6659FF586B66BA695746" box="[159,290,295,321]" captionStart="FIGURES 3032" captionStartId="26.[151,264,1539,1563]" captionTargetBox="[151,1435,703,1514]" captionTargetId="figure-200@26.[151,1435,703,1514]" captionTargetPageId="26" captionText="FIGURES 3032. Endophloeus gorskii Alekseev et Bukejs, 2016 in Baltic amber: 30—specimen No. JDC-9315, habitus, dorsal view; 31—idem, habitus, ventral view; 32—specimen No. ABAC 063 [ACAB], habitus, dorsal view. Scale bars = 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.14033669" httpUri="https://zenodo.org/record/14033669/files/figure.png" pageId="26" pageNumber="227">Figs 3031</figureCitation>
). A complete beetle is included in a transparent, yellow amber piece with dimensions of 20×
<quantity id="4CA5C3CBFFFA6659FAE66B66BE255745" box="[1313,1390,295,322]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="26" pageNumber="227" unit="mm" value="17.0">17 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFFA6659FE696B0ABABB5761" box="[430,496,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="227" unit="mm" value="5.0">5 mm</quantity>
, preserved without supplementary fixation. Syninclusions: numerous minute detrital particles
</materialsCitation>
.
</paragraph>
<paragraph id="8BE26E2EFFFA6659FF006BD2B9C6543A" blockId="26.[151,1437,259,682]" pageId="26" pageNumber="227">
<materialsCitation id="3B356473FFFA6659FF006BD2B9C3543A" collectionCode="ACAB" collectorName="No. &amp; Jonas Damzen &amp; Baltic &amp; Ventral &amp; Syninclusions" pageId="26" pageNumber="227" specimenCode="ABAC 063" specimenCount="1">
<specimenCount id="9D5BA5A7FFFA6659FF006BD2BA2157AA" box="[199,362,403,429]" count="1" pageId="26" pageNumber="227" type="generic">One specimen</specimenCount>
with collection number
<specimenCode id="DBFBC655FFFA6659FD426BD2B84D57AA" box="[645,774,403,429]" collectionCode="ABAC" pageId="26" pageNumber="227">ABAC 063</specimenCode>
[
<collectionCode id="ED4CF6EBFFFA6659FCD06BD2B82157AA" box="[791,874,403,429]" pageId="26" pageNumber="227">ACAB</collectionCode>
] (
<collectorName id="26A80BF8FFFA6659FC446BD2B8E657AA" box="[899,941,403,429]" pageId="26" pageNumber="227">No.</collectorName>
JDC-6438 ex. coll.
<collectorName id="26A80BF8FFFA6659FB5D6BD5BE0F57AA" box="[1178,1348,403,430]" pageId="26" pageNumber="227">Jonas Damzen</collectorName>
),
<collectorName id="26A80BF8FFFA6659FA9E6BD2BED757AA" box="[1369,1436,403,429]" pageId="26" pageNumber="227">Baltic</collectorName>
amber; adult, male (
<figureCitation id="136672ABFFFA6659FEB96BF6BA9E57D6" box="[382,469,439,465]" captionStart="FIGURES 3032" captionStartId="26.[151,264,1539,1563]" captionTargetBox="[151,1435,703,1514]" captionTargetId="figure-200@26.[151,1435,703,1514]" captionTargetPageId="26" captionText="FIGURES 3032. Endophloeus gorskii Alekseev et Bukejs, 2016 in Baltic amber: 30—specimen No. JDC-9315, habitus, dorsal view; 31—idem, habitus, ventral view; 32—specimen No. ABAC 063 [ACAB], habitus, dorsal view. Scale bars = 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.14033669" httpUri="https://zenodo.org/record/14033669/files/figure.png" pageId="26" pageNumber="227">Fig. 32</figureCitation>
). A complete beetle with partially exposed metathoracic wings and apical portion of aedeagus is included in a transparent, yellow amber piece with dimensions of 14×
<quantity id="4CA5C3CBFFFA6659FBE16B9ABF3F57F1" box="[1062,1140,475,502]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="227" unit="mm" value="10.0">10 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFFA6659FF726BBEBBB3541D" box="[181,248,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="227" unit="mm" value="5.0">5 mm</quantity>
, preserved without supplementary fixation.
<collectorName id="26A80BF8FFFA6659FD256BBEB878541E" box="[738,819,511,537]" pageId="26" pageNumber="227">Ventral</collectorName>
portion of specimen is completely obscured by milky opacity within amber.
<collectorName id="26A80BF8FFFA6659FE556862B97F543A" box="[402,564,547,573]" pageId="26" pageNumber="227">Syninclusions</collectorName>
: absent
</materialsCitation>
.
</paragraph>
<paragraph id="8BE26E2EFFFA6659FF006806BFC054AD" blockId="26.[151,1437,259,682]" pageId="26" pageNumber="227">
<emphasis id="B929B23CFFFA6659FF006806BA4D5466" bold="true" box="[199,262,583,609]" pageId="26" pageNumber="227">Note.</emphasis>
<materialsCitation id="3B356473FFFA6659FECC6806BFCF54AE" collectionCode="ABAC" pageId="26" pageNumber="227" specimenCode="ABAC 063" specimenCount="1" typeStatus="holotype">
Body length of beetle JDC-9315 (including visible part of head)
<quantity id="4CA5C3CBFFFA6659FC086806BF6D5465" box="[975,1062,583,610]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="26" pageNumber="227" unit="mm" value="3.3">3.3 mm</quantity>
. Body length of beetle
<specimenCode id="DBFBC655FFFA6659FAD86806BED75466" box="[1311,1436,583,609]" collectionCode="ABAC" pageId="26" pageNumber="227">ABAC 063</specimenCode>
(including visible part of head)
<quantity id="4CA5C3CBFFFA6659FE3F682AB91A5481" box="[504,593,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="26" pageNumber="227" unit="mm" value="3.5">3.5 mm</quantity>
. Even though the specimens are larger than the
<typeStatus id="54E6D08CFFFA6659FBA1682ABF805482" box="[1126,1227,619,645]" pageId="26" pageNumber="227" type="holotype">holotype</typeStatus>
, no morphological or significant differences were found, and the specimens are assigned to extinct
<taxonomicName id="4C5D15ADFFFA6659FBD068D1BFCF54AE" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[1047,1156,655,681]" class="Insecta" family="Zopheridae" genus="Endophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="gorskii">
<emphasis id="B929B23CFFFA6659FBD068D1BFCF54AE" box="[1047,1156,655,681]" italics="true" pageId="26" pageNumber="227">E. gorskii</emphasis>
</taxonomicName>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,205 @@
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<mods:title id="356140A806FC505F83328106979AABB5">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests</mods:title>
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<mods:affiliation id="CF86C35C91C8E8C30C1DA678F9A1AEB9">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia. &amp; Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. &amp; Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; &amp; Department of Ecology &amp; Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
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<mods:affiliation id="AA9B02E843DA4274FBF432DD08664C2E">Institute of Life Sciences and Technologies, Daugavpils University, Vienības 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
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<subSubSection id="C3473DA5FFFA6659FF506C98B84D50F3" box="[151,774,1753,1780]" pageId="26" pageNumber="227" type="nomenclature">
<paragraph id="8BE26E2EFFFA6659FF506C98B84D50F3" blockId="26.[151,774,1753,1816]" box="[151,774,1753,1780]" pageId="26" pageNumber="227">
<heading id="D0AAD942FFFA6659FF506C98B84D50F3" bold="true" box="[151,774,1753,1780]" fontSize="11" level="1" pageId="26" pageNumber="227" reason="1">
<taxonomicName id="4C5D15ADFFFA6659FF506C98B84D50F3" authority="Alekseev et Bukejs, 2016" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[151,774,1753,1780]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="richardklebsi">
<emphasis id="B929B23CFFFA6659FF506C98B84D50F3" bold="true" box="[151,774,1753,1780]" pageId="26" pageNumber="227">
<emphasis id="B929B23CFFFA6659FF506C98BA8750F3" bold="true" box="[151,460,1753,1780]" italics="true" pageId="26" pageNumber="227">Xylolaemus richardklebsi</emphasis>
<bibRefCitation id="EFCC13DFFFFA6659FE156C9BB84D50F3" author="Alekseev, V. I. &amp; Alekseev, P. I." box="[466,774,1754,1780]" pageId="26" pageNumber="227" pagination="75 - 86" refId="ref23984" refString="Alekseev, V. I. &amp; Alekseev, P. I. (2016) New approaches for reconstruction of the ecosystem of an Eocene amber forest. Biology Bulletin, 43 (1), 75 - 86. https: // doi. org / 10.1134 / S 1062359016010027" type="journal article" year="2016">Alekseev et Bukejs, 2016</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFFA6659FF506CBFBA6C511F" box="[151,295,1790,1816]" pageId="26" pageNumber="227" type="description">
<paragraph id="8BE26E2EFFFA6659FF506CBFBA6C511F" blockId="26.[151,774,1753,1816]" box="[151,295,1790,1816]" pageId="26" pageNumber="227">
(
<figureCitation id="136672ABFFFA6659FF586CBFBA6B511F" box="[159,288,1790,1816]" captionStart="FIGURES 3338" captionStartId="27.[151,264,1905,1929]" captionTargetBox="[151,1435,164,1880]" captionTargetId="figure-17@27.[151,1435,164,1880]" captionTargetPageId="27" captionText="FIGURES 3338. Fossil Xylolaemus in Baltic amber: 33—X. richardklebsi, specimen No. JDC-12545 [JDC], habitus in dorsal view; 34—X. richardklebsi, specimen No. JDC-12546 [JDC], 34—habitus in dorsal view; 35—idem, habitus in ventral view; 36—X. richardklebsi, specimen No. KA-CLP-002 [CKVA], habitus in dorsal view; 37—X. legalovi, specimen No. KA-CLP- 003 [CKVA], 37—habitus in dorsal view, 38—idem, habitus in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14033675" httpUri="https://zenodo.org/record/14033675/files/figure.png" pageId="26" pageNumber="227">Figs 3336</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFFA665FFF506D07BECE55EA" lastPageId="28" lastPageNumber="229" pageId="26" pageNumber="227" type="materials_examined">
<paragraph id="8BE26E2EFFFA6659FF506D07BF7151F7" blockId="26.[151,1436,1862,2032]" pageId="26" pageNumber="227">
<emphasis id="B929B23CFFFA6659FF506D07BAC95167" bold="true" box="[151,386,1862,1888]" pageId="26" pageNumber="227">Material examined.</emphasis>
<specimenCount id="9D5BA5A7FFFA6659FE4D6D07B9605167" box="[394,555,1862,1888]" count="1" pageId="26" pageNumber="227" type="generic">One specimen</specimenCount>
with collection number JDC-12545 [
<collectionCode id="ED4CF6EBFFFA6659FC166D07BF415167" box="[977,1034,1862,1888]" pageId="26" pageNumber="227">JDC</collectionCode>
], Baltic amber; adult, sex unknown (
<figureCitation id="136672ABFFFA6659FF586D2BBBBF5183" box="[159,244,1898,1924]" captionStart="FIGURES 3338" captionStartId="27.[151,264,1905,1929]" captionTargetBox="[151,1435,164,1880]" captionTargetId="figure-17@27.[151,1435,164,1880]" captionTargetPageId="27" captionText="FIGURES 3338. Fossil Xylolaemus in Baltic amber: 33—X. richardklebsi, specimen No. JDC-12545 [JDC], habitus in dorsal view; 34—X. richardklebsi, specimen No. JDC-12546 [JDC], 34—habitus in dorsal view; 35—idem, habitus in ventral view; 36—X. richardklebsi, specimen No. KA-CLP-002 [CKVA], habitus in dorsal view; 37—X. legalovi, specimen No. KA-CLP- 003 [CKVA], 37—habitus in dorsal view, 38—idem, habitus in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14033675" httpUri="https://zenodo.org/record/14033675/files/figure.png" pageId="26" pageNumber="227">Fig. 33</figureCitation>
). A complete beetle is included in a transparent, yellow amber piece with dimensions of 23×
<quantity id="4CA5C3CBFFFA6659FACD6D2BBE135183" box="[1290,1368,1898,1924]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="26" pageNumber="227" unit="mm" value="17.0">17 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFFA6659FE666DCFBAAE51AF" box="[417,485,1934,1960]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="26" pageNumber="227" unit="mm" value="7.0">7 mm</quantity>
, preserved without supplementary fixation. Ventral portion of specimen is obscured by cracks and milky opacity within amber. Syninclusions: one
<taxonomicName id="4C5D15ADFFFA6659FCAD6DF3B8A151CB" box="[874,1002,1970,1996]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="227" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
(
<taxonomicName id="4C5D15ADFFFA6659FC3E6DF3BFDE51CB" box="[1017,1173,1970,1996]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="227" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
) specimen; large leg of?
<taxonomicName id="4C5D15ADFFFA6659FF656D97BA4E51F7" box="[162,261,2006,2032]" class="Pilidiophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="227" phylum="Nemertea" rank="order">Araneae</taxonomicName>
; few stellate
<taxonomicName id="4C5D15ADFFFA6659FE506D97BAB451F7" authorityName="Dumortier" authorityYear="1829" box="[407,511,2006,2032]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="26" pageNumber="227" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
trichomes; and numerous minute detrital particles.
</paragraph>
<caption id="DF223EA6FFFB6658FF506D30BF3C51F2" ID-DOI="http://doi.org/10.5281/zenodo.14033675" ID-Zenodo-Dep="14033675" httpUri="https://zenodo.org/record/14033675/files/figure.png" pageId="27" pageNumber="228" startId="27.[151,264,1905,1929]" targetBox="[151,1435,164,1880]" targetPageId="27" targetType="figure">
<paragraph id="8BE26E2EFFFB6658FF506D30BF3C51F2" blockId="27.[151,1436,1905,2038]" pageId="27" pageNumber="228">
<emphasis id="B929B23CFFFB6658FF506D30BA1F518E" bold="true" box="[151,340,1905,1929]" pageId="27" pageNumber="228">FIGURES 3338.</emphasis>
Fossil
<taxonomicName id="4C5D15ADFFFB6658FE5B6D33B959518D" authorityName="L. Redtenbacher" authorityYear="1857" box="[412,530,1906,1930]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFFB6658FE5B6D33B959518D" box="[412,530,1906,1930]" italics="true" pageId="27" pageNumber="228">Xylolaemus</emphasis>
</taxonomicName>
in Baltic amber: 33—
<taxonomicName id="4C5D15ADFFFB6658FD356D33B8DF518D" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[754,916,1906,1930]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="richardklebsi">
<emphasis id="B929B23CFFFB6658FD356D33B8DF518D" box="[754,916,1906,1930]" italics="true" pageId="27" pageNumber="228">X. richardklebsi</emphasis>
</taxonomicName>
, specimen No. JDC-12545 [JDC], habitus in dorsal view; 34—
<taxonomicName id="4C5D15ADFFFB6658FECE6DD7BAE751A9" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[265,428,1942,1966]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="richardklebsi">
<emphasis id="B929B23CFFFB6658FECE6DD7BAE751A9" box="[265,428,1942,1966]" italics="true" pageId="27" pageNumber="228">X. richardklebsi</emphasis>
</taxonomicName>
, specimen No. JDC-12546 [JDC], 34—habitus in dorsal view; 35—idem, habitus in ventral view; 36—
<taxonomicName id="4C5D15ADFFFB6658FF0D6DFBBA2651D5" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[202,365,1978,2002]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="richardklebsi">
<emphasis id="B929B23CFFFB6658FF0D6DFBBA2651D5" box="[202,365,1978,2002]" italics="true" pageId="27" pageNumber="228">X. richardklebsi</emphasis>
</taxonomicName>
, specimen No. KA-CLP-002 [CKVA], habitus in dorsal view; 37—
<taxonomicName id="4C5D15ADFFFB6658FBEF6DFBBFDC51D5" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[1064,1175,1978,2002]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="legalovi">
<emphasis id="B929B23CFFFB6658FBEF6DFBBFDC51D5" box="[1064,1175,1978,2002]" italics="true" pageId="27" pageNumber="228">X. legalovi</emphasis>
</taxonomicName>
, specimen No. KA-CLP- 003 [CKVA], 37—habitus in dorsal view, 38—idem, habitus in lateral view. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BE26E2EFFFC665FFF006AD6BA465746" blockId="28.[151,1437,150,1005]" pageId="28" pageNumber="229">
<materialsCitation id="3B356473FFFC665FFF006AD6BA425746" collectionCode="JDC" pageId="28" pageNumber="229" specimenCount="1">
<specimenCount id="9D5BA5A7FFFC665FFF006AD6BA2256B6" box="[199,361,151,177]" count="1" pageId="28" pageNumber="229" type="generic">One specimen</specimenCount>
with collection number JDC-12546 [
<collectionCode id="ED4CF6EBFFFC665FFCD46AD6B80756B6" box="[787,844,151,177]" pageId="28" pageNumber="229">JDC</collectionCode>
], Baltic amber; adult, sex unknown (
<figureCitation id="136672ABFFFC665FFB366AD6BE3C56B6" box="[1265,1399,150,177]" captionStart="FIGURES 3338" captionStartId="27.[151,264,1905,1929]" captionTargetBox="[151,1435,164,1880]" captionTargetId="figure-17@27.[151,1435,164,1880]" captionTargetPageId="27" captionText="FIGURES 3338. Fossil Xylolaemus in Baltic amber: 33—X. richardklebsi, specimen No. JDC-12545 [JDC], habitus in dorsal view; 34—X. richardklebsi, specimen No. JDC-12546 [JDC], 34—habitus in dorsal view; 35—idem, habitus in ventral view; 36—X. richardklebsi, specimen No. KA-CLP-002 [CKVA], habitus in dorsal view; 37—X. legalovi, specimen No. KA-CLP- 003 [CKVA], 37—habitus in dorsal view, 38—idem, habitus in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14033675" httpUri="https://zenodo.org/record/14033675/files/figure.png" pageId="28" pageNumber="229">Figs 3435</figureCitation>
). A complete beetle with partially exposed metathoracic wings is included in a transparent, yellow amber piece with dimensions of 25×
<quantity id="4CA5C3CBFFFC665FFEAC6A9FBAF056FE" box="[363,443,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="28" pageNumber="229" unit="mm" value="23.0">23 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFFC665FFCD46A9EB81356FE" box="[787,856,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="229" unit="mm" value="8.0">8 mm</quantity>
, preserved without supplementary fixation. Ventral portion of specimen is visible, partially obscured by gas bubble on right side. Syninclusions: few stellate
<taxonomicName id="4C5D15ADFFFC665FFAF36B42BED7571A" authorityName="Dumortier" authorityYear="1829" box="[1332,1436,259,285]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="28" pageNumber="229" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
trichomes
</materialsCitation>
.
</paragraph>
<paragraph id="8BE26E2EFFFC665FFF006B0ABA0F57D6" blockId="28.[151,1437,150,1005]" pageId="28" pageNumber="229">
<specimenCount id="9D5BA5A7FFFC665FFF006B0ABA2C5762" box="[199,359,331,357]" count="1" pageId="28" pageNumber="229" type="generic">One specimen</specimenCount>
with collection number No. KA-CLP-002 [
<collectionCode id="ED4CF6EBFFFC665FFC896B0AB8E95762" box="[846,930,331,357]" pageId="28" pageNumber="229">CKVA</collectionCode>
], Baltic amber; adult, sex unknown (
<figureCitation id="136672ABFFFC665FFAFB6B0ABEDB5762" box="[1340,1424,331,357]" captionStart="FIGURES 3338" captionStartId="27.[151,264,1905,1929]" captionTargetBox="[151,1435,164,1880]" captionTargetId="figure-17@27.[151,1435,164,1880]" captionTargetPageId="27" captionText="FIGURES 3338. Fossil Xylolaemus in Baltic amber: 33—X. richardklebsi, specimen No. JDC-12545 [JDC], habitus in dorsal view; 34—X. richardklebsi, specimen No. JDC-12546 [JDC], 34—habitus in dorsal view; 35—idem, habitus in ventral view; 36—X. richardklebsi, specimen No. KA-CLP-002 [CKVA], habitus in dorsal view; 37—X. legalovi, specimen No. KA-CLP- 003 [CKVA], 37—habitus in dorsal view, 38—idem, habitus in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14033675" httpUri="https://zenodo.org/record/14033675/files/figure.png" pageId="28" pageNumber="229">Fig. 36</figureCitation>
). A complete beetle included in a transparent, yellow amber piece with dimensions of 20×
<quantity id="4CA5C3CBFFFC665FFB4B6B2EBF97578E" box="[1164,1244,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="28" pageNumber="229" unit="mm" value="27.0">27 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFFC665FFEF76BD2BA3257AA" box="[304,377,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="229" unit="mm" value="4.0">4 mm</quantity>
, preserved without supplementary fixation. Syninclusions:
<specimenCount id="9D5BA5A7FFFC665FFBF36BD2BF9857AA" box="[1076,1235,403,429]" count="1" pageId="28" pageNumber="229" type="generic">one specimen</specimenCount>
of
<taxonomicName id="4C5D15ADFFFC665FFAC36BD2BED757AA" box="[1284,1436,403,429]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
(?
<taxonomicName id="4C5D15ADFFFC665FFF6E6BF6BA7C57D6" box="[169,311,439,465]" class="Insecta" family="Chalcididae" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="family">Chalcididae</taxonomicName>
).
</paragraph>
<paragraph id="8BE26E2EFFFC665FFF006B9AB8475482" blockId="28.[151,1437,150,1005]" pageId="28" pageNumber="229">
<emphasis id="B929B23CFFFC665FFF006B9ABA4D57F2" bold="true" box="[199,262,475,501]" pageId="28" pageNumber="229">Note.</emphasis>
Body length of beetle JDC-12545 (including visible part of head)
<quantity id="4CA5C3CBFFFC665FFBCC6B9ABF2E57F2" box="[1035,1125,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="28" pageNumber="229" unit="mm" value="3.9">3.9 mm</quantity>
, maximum body width
<quantity id="4CA5C3CBFFFC665FFABE6B9ABB8D541E" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="229" unit="mm" value="1.4">1.4 mm</quantity>
. This specimen has a dilated antennomere 10, elytra without fascicules, pronotum with longitudinal impression at middle, and about 1012 small denticles on each side of pronotum laterally. Although the specimen is slightly smaller than the
<typeStatus id="54E6D08CFFFC665FFE886806BAFF5466" box="[335,436,583,609]" pageId="28" pageNumber="229" type="holotype">holotype</typeStatus>
, and it is obscured for study on the ventral side, no morphological differences were found, and the specimen is assigned to extinct
<taxonomicName id="4C5D15ADFFFC665FFD95682AB84E5482" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[594,773,619,645]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="richardklebsi">
<emphasis id="B929B23CFFFC665FFD95682AB84E5482" box="[594,773,619,645]" italics="true" pageId="28" pageNumber="229">X. richardklebsi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE26E2EFFFC665FFF0068CEBFA25512" blockId="28.[151,1437,150,1005]" pageId="28" pageNumber="229">
Body length of beetle JDC-12546 (including visible part of head)
<quantity id="4CA5C3CBFFFC665FFC6068CEB8B454AE" box="[935,1023,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="28" pageNumber="229" unit="mm" value="3.4">3.4 mm</quantity>
, maximum body width
<quantity id="4CA5C3CBFFFC665FFACF68CEBE2A54AE" box="[1288,1377,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="229" unit="mm" value="1.2">1.2 mm</quantity>
. This specimen has dilated antennomere 10, elytra without fascicules, pronotum with distinct longitudinal impression medially, and 12 small denticles on each side of pronotum laterally. While the specimen is smaller than the
<typeStatus id="54E6D08CFFFC665FFAF46896BED354F6" box="[1331,1432,727,753]" pageId="28" pageNumber="229" type="holotype">holotype</typeStatus>
, no morphological differences were found, and the specimen is assigned to extinct
<taxonomicName id="4C5D15ADFFFC665FFBE868BABFA95512" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[1071,1250,763,789]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="richardklebsi">
<emphasis id="B929B23CFFFC665FFBE868BABFA95512" box="[1071,1250,763,789]" italics="true" pageId="28" pageNumber="229">X. richardklebsi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE26E2EFFFC665FFF00695EB9F85586" blockId="28.[151,1437,150,1005]" pageId="28" pageNumber="229">
Body length of beetle KA-CLP-002 (including visible part of head)
<quantity id="4CA5C3CBFFFC665FFC0D695FBF7A553E" box="[970,1073,798,825]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" pageId="28" pageNumber="229" unit="mm" value="4.25">4.25 mm</quantity>
, maximum body width
<quantity id="4CA5C3CBFFFC665FFAF8695FBED2553E" box="[1343,1433,798,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="229" unit="mm" value="1.5">1.5 mm</quantity>
. Although the specimen is slightly larger than the
<typeStatus id="54E6D08CFFFC665FFD236902B802555A" box="[740,841,835,861]" pageId="28" pageNumber="229" type="holotype">holotype</typeStatus>
, no morphological differences were found and the specimen is assigned to extinct
<taxonomicName id="4C5D15ADFFFC665FFE3E6926B9E75586" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[505,684,871,897]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="richardklebsi">
<emphasis id="B929B23CFFFC665FFE3E6926B9E75586" box="[505,684,871,897]" italics="true" pageId="28" pageNumber="229">X. richardklebsi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE26E2EFFFC665FFF0069CABECE55EA" blockId="28.[151,1437,150,1005]" pageId="28" pageNumber="229">
Our study of
<taxonomicName id="4C5D15ADFFFC665FFEAE69CABAA255A2" authorityName="Solier" authorityYear="1834" box="[361,489,907,933]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="family">Zopheridae</taxonomicName>
amber inclusions in Baltic amber (including present material and specimens not considered in the current publication) shows that
<taxonomicName id="4C5D15ADFFFC665FFD7869EEB83A55CE" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[703,881,943,969]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="richardklebsi">
<emphasis id="B929B23CFFFC665FFD7869EEB83A55CE" box="[703,881,943,969]" italics="true" pageId="28" pageNumber="229">X. richardklebsi</emphasis>
</taxonomicName>
was abundant in the Eocene amberiferous forests of Fennosarmatia, and that it is seemingly the most common inclusion among the Baltic amber zopherid species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,100 @@
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<mods:title id="25CF1733691B15B19FED3E91966682C9">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests</mods:title>
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<mods:affiliation id="96FEB4175F68885C77E85E0C199CF085">Institute of Life Sciences and Technologies, Daugavpils University, Vienības 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
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<subSubSection id="C3473DA5FFFC665FFF506C86B88A50E7" box="[151,961,1734,1761]" pageId="28" pageNumber="229" type="nomenclature">
<paragraph id="8BE26E2EFFFC665FFF506C86B88A50E7" blockId="28.[151,961,1734,1797]" box="[151,961,1734,1761]" pageId="28" pageNumber="229">
<heading id="D0AAD942FFFC665FFF506C86B88A50E7" bold="true" box="[151,961,1734,1761]" fontSize="11" level="1" pageId="28" pageNumber="229" reason="1">
<taxonomicName id="4C5D15ADFFFC665FFF506C86B88A50E7" authority="Nabozhenko, Kirejtshuk et Merkl, 2016" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[151,961,1734,1761]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="colydioides">
<emphasis id="B929B23CFFFC665FFF506C86B88A50E7" bold="true" box="[151,961,1734,1761]" pageId="28" pageNumber="229">
<emphasis id="B929B23CFFFC665FFF506C86BA8150E6" bold="true" box="[151,458,1734,1761]" italics="true" pageId="28" pageNumber="229">Yantaroxenos colydioides</emphasis>
<bibRefCitation id="EFCC13DFFFFC665FFE166C86B88A50E7" author="Nabozhenko, M. &amp; Kirejtshuk, A. &amp; Merkl, O." box="[465,961,1734,1761]" pageId="28" pageNumber="229" pagination="563 - 566" refId="ref26965" refString="Nabozhenko, M., Kirejtshuk, A. &amp; Merkl, O. (2016) Yantaroxenos colydioides gen. et sp. nov. (Tenebrionidae: Lagriinae) from Baltic amber. Annales Zoologici, Warszawa, 66 (4), 563 - 566. https: // doi. org / 10.3161 / 00034541 ANZ 2016.66.4.008" type="journal article" year="2016">Nabozhenko, Kirejtshuk et Merkl, 2016</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFFC665FFF506CAABA6C5102" box="[151,295,1771,1797]" pageId="28" pageNumber="229" type="description">
<paragraph id="8BE26E2EFFFC665FFF506CAABA6C5102" blockId="28.[151,961,1734,1797]" box="[151,295,1771,1797]" pageId="28" pageNumber="229">
(
<figureCitation id="136672ABFFFC665FFF586CAABA545102" box="[159,287,1771,1797]" captionStart="FIGURES 3941" captionStartId="29.[151,264,1269,1293]" captionTargetBox="[151,1435,179,1243]" captionTargetId="figure-17@29.[151,1435,179,1243]" captionTargetPageId="29" captionText="FIGURES 3941. Yantaroxenos colydioides in Baltic amber, specimen No. JDC-13221 [JDC]: 39—habitus in dorsal view; 40—idem, habitus in ventral view; 41—idem, habitus in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14033679" httpUri="https://zenodo.org/record/14033679/files/figure.png" pageId="28" pageNumber="229">Figs 3941</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFFC665FFF506D73BFC651DA" pageId="28" pageNumber="229" type="materials_examined">
<paragraph id="8BE26E2EFFFC665FFF506D73BF825192" blockId="28.[151,1437,1842,2013]" pageId="28" pageNumber="229">
<emphasis id="B929B23CFFFC665FFF506D73BACA514B" bold="true" box="[151,385,1842,1868]" pageId="28" pageNumber="229">Material examined.</emphasis>
<materialsCitation id="3B356473FFFC665FFE4E6D72BF8E5192" collectionCode="JDC" pageId="28" pageNumber="229" specimenCount="1">
<specimenCount id="9D5BA5A7FFFC665FFE4E6D72B962514A" box="[393,553,1843,1869]" count="1" pageId="28" pageNumber="229" type="generic">One specimen</specimenCount>
with collection number JDC-13221 [
<collectionCode id="ED4CF6EBFFFC665FFC0A6D72BF4D514A" box="[973,1030,1843,1869]" pageId="28" pageNumber="229">JDC</collectionCode>
], Baltic amber; adult, sex unknown. A rather complete beetle is included in a transparent, yellow amber piece with dimensions of 22×
<quantity id="4CA5C3CBFFFC665FFAC86D16BE185176" box="[1295,1363,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="229" unit="mm" value="6.0">6 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFFC665FFE5A6D3BBAAA5192" box="[413,481,1914,1941]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="28" pageNumber="229" unit="mm" value="5.0">5 mm</quantity>
; preserved without supplementary fixation. Syninclusions: absent
</materialsCitation>
.
</paragraph>
<paragraph id="8BE26E2EFFFC665FFF006DDEBFC651DA" blockId="28.[151,1437,1842,2013]" pageId="28" pageNumber="229">
<emphasis id="B929B23CFFFC665FFF006DDEBA4D51BE" bold="true" box="[199,262,1951,1977]" pageId="28" pageNumber="229">Note.</emphasis>
Body length of beetle JDC-13221 (including visible part of head)
<quantity id="4CA5C3CBFFFC665FFC336DDEBF0651BE" box="[1012,1101,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="28" pageNumber="229" unit="mm" value="6.6">6.6 mm</quantity>
. Specimen JDC-13221 shares with the
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other external characters and is considered conspecific.
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<mods:affiliation id="8CD50F53B5DCF8AC54EBA618D972F127">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia. &amp; Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. &amp; Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; &amp; Department of Ecology &amp; Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
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<emphasis id="B929B23CFFFC665FFF506E7FBAC0525E" bold="true" box="[151,395,1086,1113]" italics="true" pageId="28" pageNumber="229">Xylolaemus legalovi</emphasis>
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(
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)
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<emphasis id="B929B23CFFFC665FFF506EEBBAC852C3" bold="true" box="[151,387,1194,1220]" pageId="28" pageNumber="229">Material examined.</emphasis>
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with collection number No. KA-CLP-003 [
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], Baltic amber; adult, sex unknown. A complete beetle is included in a transparent, yellow amber piece with dimensions of 25×
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and a maximum thickness of
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, preserved without supplementary fixation. Ventral portion of specimen is obscured by cracks and “milky” opacity within amber. Syninclusions: one stellate
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<emphasis id="B929B23CFFFC665FFF006F7ABA4D5352" bold="true" box="[199,262,1339,1365]" pageId="28" pageNumber="229">Note.</emphasis>
Measurements of the specimen (body length
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), dilated antennomere 10, elytra with setal fascicules, pronotum without distinct longitudinal impression at middle, and strongly arcuate anterior margin of pronotum let us assign the specimen to extinct
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Genus
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Erichson, 1842: 214
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1.
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References:
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Bukejs
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. 2019a: 566
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(description).
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Comment: the
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3.
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References:
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Nabozhenko
<emphasis id="B929B23CFFFF665CFE2C6C83B95450DC" box="[491,543,1729,1755]" italics="true" pageId="31" pageNumber="232">et al</emphasis>
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(systematic); present paper (add. rec.).
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Comment: the species and genus were originally described within the tribe
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2.
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†Genus
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<document id="A2C8F17617D8C9A4F826111F833EF163" ID-DOI="10.11646/zootaxa.5536.2.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="14033773" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730713246159" checkinUser="plazi" docAuthor="Barroso, Paulo Cézar Salgado, Somavilla, Alexandre, Broad, Gavin R., Menezes, Rodolpho Santos Telles &amp; Oliveira, Marcio Luiz De" docDate="2024" docId="0E323F5F8459CC00FF62FB47DB3BFDBD" docLanguage="en" docName="zootaxa.5536.2.7.pdf" docOrigin="Zootaxa 5536 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.2.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Angiopolybia obscuriorina Barroso &amp; Somavilla &amp; Broad &amp; Menezes &amp; Oliveira 2024, sp. nov." docType="treatment" docUuid="6AD2C290-0420-4634-9976-274D1BD3E5B7" docUuidSource="ZooBank" docVersion="2" lastPageNumber="319" masterDocId="F20B47278458CC04FFF5FFB4DF62FF80" masterDocTitle="Contributing to the understanding of Angiopolybia Araujo, 1946 (Hymenoptera, Vespidae, Polistinae): a new species, and descriptions of the nest of A. obidensis and male of A. zischkai" masterLastPageNumber="324" masterPageNumber="315" pageNumber="316" updateTime="1730713792271" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B1F3E75B81F600D8D3A8C6251B4DBF17">Contributing to the understanding of Angiopolybia Araujo, 1946 (Hymenoptera, Vespidae, Polistinae): a new species, and descriptions of the nest of A. obidensis and male of A. zischkai</mods:title>
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<mods:namePart id="61332FDB729EBF132CE0DB9A9799D22C">Barroso, Paulo Cézar Salgado</mods:namePart>
<mods:affiliation id="72E224A958F718C010716EC2531EFFD3">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade, Av. André Araújo, 69067 - 375, Manaus, Brazil.</mods:affiliation>
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<mods:namePart id="25FC6E73405419B6DD423B9111FEE687">Somavilla, Alexandre</mods:namePart>
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<mods:affiliation id="3B5D3B21851F2DFB2F72DDFD78EB33FA">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade, Av. André Araújo, 69067 - 375, Manaus, Brazil.</mods:affiliation>
<mods:nameIdentifier id="04DDF90D8552B6C6BE0C8D5C083119D2" type="email">alexandresomavilla@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EEDFA06189728088E47B1B82CBE8C4CA">Broad, Gavin R.</mods:namePart>
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<mods:namePart id="1651838C4D1FB3CD5DDB68ABBFE4B5A0">Menezes, Rodolpho Santos Telles</mods:namePart>
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<mods:affiliation id="3FC015FFA4EC3C2ABA14EA4E640F1220">Universidade Estadual de Santa Cruz, Departamento de Ciências Biológicas, Rodovia Jorge Amado, km 16, 45662 - 900, Ilhéus, Brazil.</mods:affiliation>
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<mods:namePart id="83171CD045A6DE4CC41D48DBC214A2E7">Oliveira, Marcio Luiz De</mods:namePart>
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<mods:affiliation id="39FAD59048C5903926DF4DAD20FA8CA8">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade, Av. André Araújo, 69067 - 375, Manaus, Brazil. &amp; Instituto Nacional de Biodiversidad, División de Entomología, Av. Rumipamba y Av. de los Shyris, RG 77 + GQV, Quito, Ecuador.</mods:affiliation>
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<heading id="DD6C39258459CC05FF62FB47DDD1FA8D" bold="true" box="[151,691,1267,1294]" fontSize="11" level="1" pageId="1" pageNumber="316" reason="1">
<emphasis id="B4EF525B8459CC05FF62FB47DDD1FA8D" bold="true" box="[151,691,1267,1294]" pageId="1" pageNumber="316">
<taxonomicName id="419BF5CA8459CC05FF62FB47DD2AFA8D" authority="Barroso" authorityName="Barroso &amp; Somavilla &amp; Broad &amp; Menezes &amp; Oliveira" authorityYear="2024" box="[151,584,1267,1294]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="obscuriorina" status="sp. nov.">
<emphasis id="B4EF525B8459CC05FF62FB47DEB9FA8E" bold="true" box="[151,475,1267,1294]" italics="true" pageId="1" pageNumber="316">Angiopolybia obscuriorina</emphasis>
Barroso
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,
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<subSubSection id="CE81DDC28459CC05FF62FAA3DE21FAB1" box="[151,323,1303,1329]" pageId="1" pageNumber="316" type="description">
<paragraph id="86248E498459CC05FF62FAA3DE21FAB1" blockId="1.[151,691,1267,1329]" box="[151,323,1303,1329]" pageId="1" pageNumber="316">
(
<figureCitation id="1EA092CC8459CC05FF6AFAA3DE59FAB1" box="[159,315,1303,1329]" captionStart-0="FIGURES 17. 13" captionStart-1="FIGURES 811. 8, 9" captionStartId-0="2.[151,264,1794,1818]" captionStartId-1="3.[151,264,703,727]" captionTargetBox-0="[152,1435,179,1768]" captionTargetBox-1="[191,1395,181,679]" captionTargetId-0="figure-17@2.[152,1435,179,1768]" captionTargetId-1="figure-19@3.[191,1395,181,679]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 17. 13. Angiopolybia obscuriorina sp. nov., holotype, ♀. 46. Stelopolybia paraensisvar.(or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 1, 5. Habitus, lateral view. 2, 6. Habitus, dorsal view. 3, 4. Head, frontal view. 7. Map of distribution (© President and Fellows of Harvard College, MCZ, for images 4, 5, and 6, https://mczbase.mcz. harvard.edu/guid/MCZ:Ent:36021). Scale bar: 1 mm." captionText-1="FIGURES 811. 8, 9. Angiopolybia obscuriorina sp. nov., holotype, ♀. 10, 11. Stelopolybia paraensis var. (or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 8, 10. Metapleural basalar area, lateral view. 9, 11. Tergum II, lateral view. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14033775" figureDoi-1="http://doi.org/10.5281/zenodo.14033777" httpUri-0="https://zenodo.org/record/14033775/files/figure.png" httpUri-1="https://zenodo.org/record/14033777/files/figure.png" pageId="1" pageNumber="316">Figs 13, 79</figureCitation>
)
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<subSubSection id="CE81DDC28459CC05FF62FAEBDC6DF885" pageId="1" pageNumber="316" type="diagnosis">
<paragraph id="86248E498459CC05FF62FAEBDE51F989" blockId="1.[151,1437,1375,1978]" pageId="1" pageNumber="316">
<emphasis id="B4EF525B8459CC05FF62FAEBDE68FAF9" bold="true" box="[151,266,1375,1401]" pageId="1" pageNumber="316">Diagnosis</emphasis>
. Anterior wings
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; tergum II without row of very long bristles on posterior margin; overall color dark brown with only faint posterior bands on terga; eye with very small and sparse bristles; gena angulate with an enlarged lower region; pronotal lamella low on anterior margin, one-fifth of height of the antennal socket; axillary fossa with anterior margin directed anteriorly; posterior submedian translucent mark of the propodeum not inserted in depression.
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<paragraph id="86248E498459CC05FF32F9A7DC6DF885" blockId="1.[151,1437,1375,1978]" pageId="1" pageNumber="316">
<emphasis id="B4EF525B8459CC05FF32F9A7DEB9F9AD" bold="true" box="[199,475,1555,1581]" pageId="1" pageNumber="316">Comparative diagnosis</emphasis>
.
<taxonomicName id="419BF5CA8459CC05FE19F9A7DC7EF9AD" authorityName="Barroso &amp; Somavilla &amp; Broad &amp; Menezes &amp; Oliveira" authorityYear="2024" box="[492,796,1555,1581]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="obscuriorina" status="sp. nov.">
<emphasis id="B4EF525B8459CC05FE19F9A7DC7EF9AD" box="[492,796,1555,1581]" italics="true" pageId="1" pageNumber="316">Angiopolybia obscuriorina</emphasis>
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<taxonomicNameLabel id="AFDCEF208459CC05FCD0F9A7DCE2F9AD" box="[805,896,1555,1581]" pageId="1" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
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resembles
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<emphasis id="B4EF525B8459CC05FBF7F9A7DBF2F9AD" box="[1026,1168,1555,1581]" italics="true" pageId="1" pageNumber="316">A. paraensis</emphasis>
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, particularly the brown variants (
<figureCitation id="1EA092CC8459CC05FF0AF983DE07F9D1" box="[255,357,1591,1617]" captionStart="FIGURES 17. 13" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1435,179,1768]" captionTargetId="figure-17@2.[152,1435,179,1768]" captionTargetPageId="2" captionText="FIGURES 17. 13. Angiopolybia obscuriorina sp. nov., holotype, ♀. 46. Stelopolybia paraensisvar.(or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 1, 5. Habitus, lateral view. 2, 6. Habitus, dorsal view. 3, 4. Head, frontal view. 7. Map of distribution (© President and Fellows of Harvard College, MCZ, for images 4, 5, and 6, https://mczbase.mcz. harvard.edu/guid/MCZ:Ent:36021). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033775" httpUri="https://zenodo.org/record/14033775/files/figure.png" pageId="1" pageNumber="316">Figs 46</figureCitation>
,
<figureCitation id="1EA092CC8459CC05FE85F983DEDAF9D1" box="[368,440,1591,1617]" captionStart="FIGURES 811. 8, 9" captionStartId="3.[151,264,703,727]" captionTargetBox="[191,1395,181,679]" captionTargetId="figure-19@3.[191,1395,181,679]" captionTargetPageId="3" captionText="FIGURES 811. 8, 9. Angiopolybia obscuriorina sp. nov., holotype, ♀. 10, 11. Stelopolybia paraensis var. (or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 8, 10. Metapleural basalar area, lateral view. 9, 11. Tergum II, lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033777" httpUri="https://zenodo.org/record/14033777/files/figure.png" pageId="1" pageNumber="316">10, 11</figureCitation>
), and
<taxonomicName id="419BF5CA8459CC05FE08F983DDE4F9D1" authorityName="Ducke" authorityYear="1904" box="[509,646,1591,1617]" class="Insecta" family="Eumenidae" genus="Polybia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="obidensis">
<emphasis id="B4EF525B8459CC05FE08F983DDE4F9D1" box="[509,646,1591,1617]" italics="true" pageId="1" pageNumber="316">A. obidensis</emphasis>
</taxonomicName>
. However, it differs from
<taxonomicName id="419BF5CA8459CC05FC58F983DB5BF9D1" box="[941,1081,1591,1617]" class="Insecta" family="Eumenidae" genus="Polybia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B8459CC05FC58F983DB5BF9D1" box="[941,1081,1591,1617]" italics="true" pageId="1" pageNumber="316">A. paraensis</emphasis>
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in the absence of a row of very long bristles along the posterior margin of tergum II (
<figureCitation id="1EA092CC8459CC05FD0AF9EFDC27F9F5" box="[767,837,1627,1653]" captionStart="FIGURES 811. 8, 9" captionStartId="3.[151,264,703,727]" captionTargetBox="[191,1395,181,679]" captionTargetId="figure-19@3.[191,1395,181,679]" captionTargetPageId="3" captionText="FIGURES 811. 8, 9. Angiopolybia obscuriorina sp. nov., holotype, ♀. 10, 11. Stelopolybia paraensis var. (or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 8, 10. Metapleural basalar area, lateral view. 9, 11. Tergum II, lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033777" httpUri="https://zenodo.org/record/14033777/files/figure.png" pageId="1" pageNumber="316">Fig. 9</figureCitation>
), which is present in
<taxonomicName id="419BF5CA8459CC05FBCBF9EFDBAFF9F5" box="[1086,1229,1627,1653]" class="Insecta" family="Eumenidae" genus="Polybia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B8459CC05FBCBF9EFDBAFF9F5" box="[1086,1229,1627,1653]" italics="true" pageId="1" pageNumber="316">A. paraensis</emphasis>
</taxonomicName>
(
<figureCitation id="1EA092CC8459CC05FB2BF9EFDA53F9F5" box="[1246,1329,1627,1653]" captionStart="FIGURES 811. 8, 9" captionStartId="3.[151,264,703,727]" captionTargetBox="[191,1395,181,679]" captionTargetId="figure-19@3.[191,1395,181,679]" captionTargetPageId="3" captionText="FIGURES 811. 8, 9. Angiopolybia obscuriorina sp. nov., holotype, ♀. 10, 11. Stelopolybia paraensis var. (or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 8, 10. Metapleural basalar area, lateral view. 9, 11. Tergum II, lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033777" httpUri="https://zenodo.org/record/14033777/files/figure.png" pageId="1" pageNumber="316">Fig. 11</figureCitation>
), and the area of the propodeum anterior to the spiracle projects less in the metapleural basalar area (
<figureCitation id="1EA092CC8459CC05FB7AF9CBDBB0F919" box="[1167,1234,1663,1689]" captionStart="FIGURES 811. 8, 9" captionStartId="3.[151,264,703,727]" captionTargetBox="[191,1395,181,679]" captionTargetId="figure-19@3.[191,1395,181,679]" captionTargetPageId="3" captionText="FIGURES 811. 8, 9. Angiopolybia obscuriorina sp. nov., holotype, ♀. 10, 11. Stelopolybia paraensis var. (or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 8, 10. Metapleural basalar area, lateral view. 9, 11. Tergum II, lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033777" httpUri="https://zenodo.org/record/14033777/files/figure.png" pageId="1" pageNumber="316">Fig. 8</figureCitation>
), while it projects more in
<taxonomicName id="419BF5CA8459CC05FF06F917DE1DF93D" box="[243,383,1699,1725]" class="Insecta" family="Eumenidae" genus="Polybia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B8459CC05FF06F917DE1DF93D" box="[243,383,1699,1725]" italics="true" pageId="1" pageNumber="316">A. paraensis</emphasis>
</taxonomicName>
(
<figureCitation id="1EA092CC8459CC05FE78F917DE82F93D" box="[397,480,1699,1725]" captionStart="FIGURES 811. 8, 9" captionStartId="3.[151,264,703,727]" captionTargetBox="[191,1395,181,679]" captionTargetId="figure-19@3.[191,1395,181,679]" captionTargetPageId="3" captionText="FIGURES 811. 8, 9. Angiopolybia obscuriorina sp. nov., holotype, ♀. 10, 11. Stelopolybia paraensis var. (or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 8, 10. Metapleural basalar area, lateral view. 9, 11. Tergum II, lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033777" httpUri="https://zenodo.org/record/14033777/files/figure.png" pageId="1" pageNumber="316">Fig. 10</figureCitation>
). Additionally, it differs from
<taxonomicName id="419BF5CA8459CC05FCD8F917DCD7F93D" authorityName="Ducke" authorityYear="1904" box="[813,949,1699,1725]" class="Insecta" family="Eumenidae" genus="Polybia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="obidensis">
<emphasis id="B4EF525B8459CC05FCD8F917DCD7F93D" box="[813,949,1699,1725]" italics="true" pageId="1" pageNumber="316">A. obidensis</emphasis>
</taxonomicName>
by the lamella on the anterior margin of the pronotum, which is one-fifth of the height of the antennal socket, the pronotum with a slight prominence in front of the fovea, and the gena not enlarged in the upper region.
</paragraph>
</subSubSection>
<subSubSection id="CE81DDC28459CC07FF32F8BBDD0CFBFB" lastPageId="3" lastPageNumber="318" pageId="1" pageNumber="316" type="materials_examined">
<paragraph id="86248E498459CC07FF32F8BBDE1DFBD7" blockId="1.[151,1437,1375,1978]" lastBlockId="3.[151,1437,871,2041]" lastPageId="3" lastPageNumber="318" pageId="1" pageNumber="316">
<emphasis id="B4EF525B8459CC05FF32F8BBDE11F8A9" bold="true" box="[199,371,1807,1833]" pageId="1" pageNumber="316">Type material</emphasis>
.
<materialsCitation id="36F384148459CC05FE73F8BBDC95F8F1" collectingDate="1982-02-20" collectionCode="AMNH, IV, V" collectorName="J. Carpenter &amp; The" country="Suriname" elevation="500" latitude="4.9666667" location="Brownsberg Nature Park" longLatPrecision="1306" longitude="-55.183334" pageId="1" pageNumber="316" specimenCode="IZC 00178415" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B4EF525B8459CC05FE73F8BBDE91F8A9" bold="true" box="[390,499,1807,1833]" pageId="1" pageNumber="316">
<typeStatus id="592030EB8459CC05FE73F8BBDE91F8A9" box="[390,499,1807,1833]" pageId="1" pageNumber="316" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="909D45C08459CC05FDF0F8A4DD78F8AA" box="[517,538,1808,1834]" count="1" pageId="1" pageNumber="316" type="female"></specimenCount>
,
<collectingCountry id="FE8CCED98459CC05FDD8F8BBDDEDF8A9" box="[557,655,1807,1833]" name="Suriname" pageId="1" pageNumber="316">Surinam</collectingCountry>
,
<location id="8344D8928459CC05FD6AF8BBDCA0F8A9" LSID="urn:lsid:plazi:treatment:0E323F5F8459CC00FF62FB47DB3BFDBD:8344D8928459CC05FD6AF8BBDCA0F8A9" box="[671,962,1807,1834]" country="Suriname" latitude="4.9666667" longLatPrecision="1306" longitude="-55.183334" name="Brownsberg Nature Park" pageId="1" pageNumber="316">Brownsberg Nature Park</location>
,
<quantity id="416323AC8459CC05FC27F8BBDB76F8AA" box="[978,1044,1807,1834]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="316" unit="m" value="500.0">
<elevation id="0DB6697A8459CC05FC27F8BBDB76F8AA" box="[978,1044,1807,1834]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="316" unit="m" value="500.0">500m</elevation>
</quantity>
,
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,
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,
<date id="F225A8898459CC05FAF5F8BBDA17F8A9" box="[1280,1397,1807,1833]" pageId="1" pageNumber="316" value="1982-02-20">
<collectingDate id="E26151618459CC05FAF5F8BBDA17F8A9" box="[1280,1397,1807,1833]" pageId="1" pageNumber="316" value="1982-02-20">20.II.1982</collectingDate>
</date>
(
<collectorName id="2B6EEB9F8459CC05FA7EF8BBDE68F8CD" pageId="1" pageNumber="316">J. Carpenter</collectorName>
),
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_
<specimenCode id="D63D26328459CC05FE71F887DD4EF8CE" box="[388,556,1843,1870]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="316">IZC 00178415</specimenCode>
[
<collectionCode id="E08A168C8459CC05FDCAF887DDFCF8CD" box="[575,670,1843,1869]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="316">AMNH</collectionCode>
].
<collectorName id="2B6EEB9F8459CC05FD46F887DDBDF8CD" box="[691,735,1843,1869]" pageId="1" pageNumber="316">The</collectorName>
<typeStatus id="592030EB8459CC05FD1FF887DC2EF8CD" box="[746,844,1843,1869]" pageId="1" pageNumber="316" type="holotype">holotype</typeStatus>
has the right hind leg broken but associated with the specimen, and tarsomeres
<collectionCode id="E08A168C8459CC05FE05F8ECDD6FF8F2" box="[496,525,1880,1906]" pageId="1" pageNumber="316">IV</collectionCode>
and
<collectionCode id="E08A168C8459CC05FDB2F8ECDD39F8F2" box="[583,603,1880,1906]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="316" type="Museum">V</collectionCode>
of the left hind leg broken and lost
</materialsCitation>
.
<materialsCitation id="36F384148459CC05FBF1F8E3DA01F816" collectionCode="V, FG, NHMUK" collectorName="O. W. Richards Collection &amp; N. Hahn" country="Guyana" elevation="1057" location="Pakaraima Mountains" pageId="1" pageNumber="316" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="B4EF525B8459CC05FBF1F8E3DB1EF8F1" bold="true" box="[1028,1148,1879,1905]" pageId="1" pageNumber="316">
<typeStatus id="592030EB8459CC05FBF1F8E3DB1EF8F1" box="[1028,1148,1879,1905]" pageId="1" pageNumber="316" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="909D45C08459CC05FB79F8E3DBD2F8F1" box="[1164,1200,1879,1905]" count="2" pageId="1" pageNumber="316" type="female">2♀</specimenCount>
,
<collectingCountry id="FE8CCED98459CC05FB4AF8E3DA7BF8F1" box="[1215,1305,1879,1905]" name="Guyana" pageId="1" pageNumber="316">Guyana</collectingCountry>
,
<location id="8344D8928459CC05FAD3F8E3DE70F815" LSID="urn:lsid:plazi:treatment:0E323F5F8459CC00FF62FB47DB3BFDBD:8344D8928459CC05FAD3F8E3DE70F815" country="Guyana" name="Pakaraima Mountains" pageId="1" pageNumber="316">Pakaraima Mountains</location>
,
<quantity id="416323AC8459CC05FED5F8CFDE19F816" box="[288,379,1915,1942]" metricMagnitude="3" metricUnit="m" metricValue="1.057" pageId="1" pageNumber="316" unit="m" value="1057.0">
<elevation id="0DB6697A8459CC05FED5F8CFDE19F816" box="[288,379,1915,1942]" metricMagnitude="3" metricUnit="m" metricValue="1.057" pageId="1" pageNumber="316" unit="m" value="1057.0">1057 m</elevation>
</quantity>
, 2.XI.[19]32 (D.
<collectionCode id="E08A168C8459CC05FDCEF8CFDD33F815" box="[571,593,1915,1941]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="316" type="Museum">V</collectionCode>
.
<collectionCode id="E08A168C8459CC05FDAAF8CFDDE5F815" box="[607,647,1915,1941]" country="Germany" name="Palaontologische Hauptsammlung der Bergakadmie" pageId="1" pageNumber="316">FG</collectionCode>
),
<collectorName id="2B6EEB9F8459CC05FD6BF8CFDCAAF815" box="[670,968,1915,1941]" pageId="1" pageNumber="316">O.W. Richards Collection</collectorName>
, B.M. (
<collectorName id="2B6EEB9F8459CC05FBD6F8CFDB39F815" box="[1059,1115,1915,1941]" name="N. Hahn" pageId="1" pageNumber="316">N.H.</collectorName>
) 1967-510 [
<collectionCode id="E08A168C8459CC05FB1FF8C8DA3FF816" box="[1258,1373,1916,1942]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="1" pageNumber="316" type="Museum">NHMUK</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="36F384148459CC05FA87F8CFDBCBF83A" collectionCode="NHMUK" collectorName="Oronoque &amp; H. Beddington" country="Guyana" location="River Heads" pageId="1" pageNumber="316" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="909D45C08459CC05FA87F8CFDAF7F815" box="[1394,1429,1915,1941]" count="1" pageId="1" pageNumber="316" type="female">1♀</specimenCount>
,
<collectingCountry id="FE8CCED98459CC05FF62F82BDF93F839" box="[151,241,1951,1977]" name="Guyana" pageId="1" pageNumber="316">Guyana</collectingCountry>
,
<collectorName id="2B6EEB9F8459CC05FF08F82BDE0FF839" box="[253,365,1951,1977]" pageId="1" pageNumber="316">Oronoque</collectorName>
&amp; New
<location id="8344D8928459CC05FE24F82BDD02F839" LSID="urn:lsid:plazi:treatment:0E323F5F8459CC00FF62FB47DB3BFDBD:8344D8928459CC05FE24F82BDD02F839" box="[465,608,1951,1977]" country="Guyana" name="River Heads" pageId="1" pageNumber="316">River Heads</location>
, 1938 (
<collectorName id="2B6EEB9F8459CC05FD40F82BDC02F839" box="[693,864,1951,1977]" pageId="1" pageNumber="316">H. Beddington</collectorName>
), B.M. 1938-346 [
<collectionCode id="E08A168C8459CC05FBC5F814DBC1F83A" box="[1072,1187,1952,1978]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="1" pageNumber="316" type="Museum">NHMUK</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="36F384148459CC07FB43F82BDC53FC01" collectionCode="MZUSP" country="Suriname" lastPageId="3" lastPageNumber="318" location="Stelop." pageId="1" pageNumber="316" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="909D45C08459CC05FB43F82BDBBBF839" box="[1206,1241,1951,1977]" count="1" pageId="1" pageNumber="316" type="female">1♀</specimenCount>
,
<collectingCountry id="FE8CCED98459CC05FB1DF82BDA28F839" box="[1256,1354,1951,1977]" name="Suriname" pageId="1" pageNumber="316">Surinam</collectingCountry>
, 1.151,
<emphasis id="B4EF525B845BCC07FF62FCD3DE21FC01" box="[151,323,871,897]" italics="true" pageId="3" pageNumber="318">
<location id="8344D892845BCC07FF62FCD3DF87FC01" LSID="urn:lsid:plazi:treatment:0E323F5F8459CC00FF62FB47DB3BFDBD:8344D892845BCC07FF62FCD3DF87FC01" box="[151,229,871,897]" country="Suriname" name="Stelop." pageId="3" pageNumber="318">Stelop.</location>
paraens
</emphasis>
<taxonomicName id="419BF5CA845BCC07FEBCFCD3DD59FC01" authority=", Ducke" authorityName="Ducke" box="[329,571,871,897]" pageId="3" pageNumber="318" rank="variety" variety="ruficornis">
var.
<emphasis id="B4EF525B845BCC07FE82FCD3DE87FC01" box="[375,485,871,897]" italics="true" pageId="3" pageNumber="318">ruficornis</emphasis>
, Ducke
</taxonomicName>
rev. [19]11 [
<collectionCode id="E08A168C845BCC07FD36FCD3DC4BFC01" box="[707,809,871,897]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="318">MZUSP</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="36F38414845BCC07FCCFFCD3DC86FC25" collectingDate="1982-01-29" collectingDateMax="1982-02-13" collectingDateMin="1982-01-29" collectionCode="AMNH" collectorName="J. Carpenter &amp; D. Trail" country="Suriname" elevation="60" location="Raleighvallen-Voltzberg Nature Reserve" municipality="Voltzberg Camp." pageId="3" pageNumber="318" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="909D45C0845BCC07FCCFFCD3DC3CFC01" box="[826,862,871,897]" count="2" pageId="3" pageNumber="318" type="female">2♀</specimenCount>
,
<collectingCountry id="FE8CCED9845BCC07FC9CFCD3DCA9FC01" box="[873,971,871,897]" name="Suriname" pageId="3" pageNumber="318">Surinam</collectingCountry>
,
<location id="8344D892845BCC07FC21FCD3DAFAFC01" LSID="urn:lsid:plazi:treatment:0E323F5F8459CC00FF62FB47DB3BFDBD:8344D892845BCC07FC21FCD3DAFAFC01" box="[980,1432,871,897]" country="Suriname" municipality="Voltzberg Camp." name="Raleighvallen-Voltzberg Nature Reserve" pageId="3" pageNumber="318">Raleighvallen-Voltzberg Nature Reserve</location>
,
<collectingMunicipality id="66401433845BCC07FF62FC3FDE35FC25" box="[151,343,907,933]" pageId="3" pageNumber="318">Voltzberg Camp.</collectingMunicipality>
,
<quantity id="416323AC845BCC07FE94FC3EDEFEFC25" box="[353,412,906,933]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="318" unit="m" value="60.0">
<elevation id="0DB6697A845BCC07FE94FC3EDEFEFC25" box="[353,412,906,933]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="318" unit="m" value="60.0">60 m</elevation>
</quantity>
,
<date id="F225A889845BCC07FE52FC3FDD35FC25" box="[423,599,907,933]" pageId="3" pageNumber="318" value="1982-01-29" valueMax="1982-02-13" valueMin="1982-01-29">
<collectingDate id="E2615161845BCC07FE52FC3FDD35FC25" box="[423,599,907,933]" pageId="3" pageNumber="318" value="1982-01-29" valueMax="1982-02-13" valueMin="1982-01-29">29.I13.II.1982</collectingDate>
</date>
(
<collectorName id="2B6EEB9F845BCC07FD92FC3FDD8FFC25" box="[615,749,907,933]" pageId="3" pageNumber="318">J. Carpenter</collectorName>
&amp;
<collectorName id="2B6EEB9F845BCC07FCE4FC3FDC0BFC25" box="[785,873,907,933]" pageId="3" pageNumber="318">D. Trail</collectorName>
) [
<collectionCode id="E08A168C845BCC07FC8BFC3FDCBEFC25" box="[894,988,907,933]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="318">AMNH</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="36F38414845BCC07FC1BFC3FDA64FC49" collectingDate="1982-01-26" collectingDateMax="1982-02-15" collectingDateMin="1982-01-26" collectionCode="AMNH" collectorName="J. Carpenter &amp; D. Trail" country="Suriname" latitude="4.7166667" location="Raleighvallen-Voltzberg Nature Reserve" longLatPrecision="1306" longitude="-56.2" municipality="Foengoe" pageId="3" pageNumber="318" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="909D45C0845BCC07FC1BFC3FDB71FC25" box="[1006,1043,907,933]" count="2" pageId="3" pageNumber="318" type="female">2♀</specimenCount>
,
<collectingCountry id="FE8CCED9845BCC07FBEAFC3FDBE3FC25" box="[1055,1153,907,933]" name="Suriname" pageId="3" pageNumber="318">Surinam</collectingCountry>
,
<location id="8344D892845BCC07FB7EFC3FDE2CFC49" LSID="urn:lsid:plazi:treatment:0E323F5F8459CC00FF62FB47DB3BFDBD:8344D892845BCC07FB7EFC3FDE2CFC49" country="Suriname" latitude="4.7166667" longLatPrecision="1306" longitude="-56.2" municipality="Foengoe" name="Raleighvallen-Voltzberg Nature Reserve" pageId="3" pageNumber="318">Raleighvallen-Voltzberg Nature Reserve</location>
,
<collectingMunicipality id="66401433845BCC07FEAAFC1BDEA0FC49" box="[351,450,943,969]" pageId="3" pageNumber="318">Foengoe</collectingMunicipality>
,
<geoCoordinate id="E3AFE88E845BCC07FE21FC1ADD45FC48" box="[468,551,942,968]" degrees="4" direction="north" minutes="43" orientation="latitude" pageId="3" pageNumber="318" precision="925" value="4.7166667">4º43N</geoCoordinate>
,
<geoCoordinate id="E3AFE88E845BCC07FDCCFC1ADDFFFC49" box="[569,669,942,969]" degrees="56" direction="west" minutes="12" orientation="longitude" pageId="3" pageNumber="318" precision="925" value="-56.2">56º12W</geoCoordinate>
,
<date id="F225A889845BCC07FD5AFC1ADC3CFC49" box="[687,862,942,969]" pageId="3" pageNumber="318" value="1982-01-26" valueMax="1982-02-15" valueMin="1982-01-26">
<collectingDate id="E2615161845BCC07FD5AFC1ADC3CFC49" box="[687,862,942,969]" pageId="3" pageNumber="318" value="1982-01-26" valueMax="1982-02-15" valueMin="1982-01-26">26.I15.II.1982</collectingDate>
</date>
(
<collectorName id="2B6EEB9F845BCC07FC80FC1ADB60FC48" box="[885,1026,942,968]" pageId="3" pageNumber="318">J. Carpenter</collectorName>
&amp;
<collectorName id="2B6EEB9F845BCC07FBD3FC1BDBE7FC48" box="[1062,1157,942,969]" pageId="3" pageNumber="318">D. Trail</collectorName>
) [
<collectionCode id="E08A168C845BCC07FB55FC1BDB9CFC49" box="[1184,1278,943,969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="318">AMNH</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="36F38414845BCC07FAE2FC1ADC2CFC6C" country="Suriname" county="Borne" elevation="347" location="French Guiana" municipality="Crique Alama" pageId="3" pageNumber="318" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="909D45C0845BCC07FAE2FC1ADA58FC48" box="[1303,1338,942,968]" count="1" pageId="3" pageNumber="318" type="female">1♀</specimenCount>
,
<location id="8344D892845BCC07FABBFC1ADF89FC6C" LSID="urn:lsid:plazi:treatment:0E323F5F8459CC00FF62FB47DB3BFDBD:8344D892845BCC07FABBFC1ADF89FC6C" country="Suriname" county="Borne" municipality="Crique Alama" name="French Guiana" pageId="3" pageNumber="318">French Guiana</location>
,
<collectingMunicipality id="66401433845BCC07FF0DFC66DEF8FC6C" box="[248,410,978,1004]" pageId="3" pageNumber="318">Crique Alama</collectingMunicipality>
,
<collectingCounty id="6F45F6C5845BCC07FE53FC67DE88FC6D" box="[422,490,979,1005]" pageId="3" pageNumber="318">Borne</collectingCounty>
1, trails A., creek level,
<quantity id="416323AC845BCC07FCFDFC66DC2CFC6C" box="[776,846,978,1004]" metricMagnitude="2" metricUnit="m" metricValue="3.47" pageId="3" pageNumber="318" unit="m" value="347.0">
<specimenCount id="909D45C0845BCC07FCFDFC66DC2CFC6C" box="[776,846,978,1004]" count="347" pageId="3" pageNumber="318" type="generic">
<elevation id="0DB6697A845BCC07FCFDFC66DC2CFC6C" box="[776,846,978,1004]" metricMagnitude="2" metricUnit="m" metricValue="3.47" pageId="3" pageNumber="318" unit="m" value="347.0">347m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation id="36F38414845BCC07FCBBFC66DDC7FB90" collectingDate="2015-03-17" collectionCode="MNHN" collectorName="C. Villemant &amp; Our Planet Reviewed" country="Suriname" latitude="2.24837" longLatPrecision="1" longitude="-54.46895" pageId="3" pageNumber="318" specimenCount="1" typeStatus="paratype">
.,
<geoCoordinate id="E3AFE88E845BCC07FCAAFC66DC89FC6C" box="[863,1003,978,1004]" degrees="02.24837" direction="north" orientation="latitude" pageId="3" pageNumber="318" precision="1" value="2.24837">02.24837°N</geoCoordinate>
,
<geoCoordinate id="E3AFE88E845BCC07FC0DFC66DBE4FC6D" box="[1016,1158,978,1005]" degrees="54.46895" direction="west" orientation="longitude" pageId="3" pageNumber="318" precision="1" value="-54.46895">54.46895°W</geoCoordinate>
,
<date id="F225A889845BCC07FB61FC66DA77FC6D" box="[1172,1301,978,1005]" pageId="3" pageNumber="318" value="2015-03-17">
<collectingDate id="E2615161845BCC07FB61FC66DA77FC6D" box="[1172,1301,978,1005]" pageId="3" pageNumber="318" value="2015-03-17">17.III.2015</collectingDate>
</date>
, hand catch (
<collectorName id="2B6EEB9F845BCC07FF55FC42DE55FB90" box="[160,311,1014,1040]" pageId="3" pageNumber="318">C. Villemant</collectorName>
),
<collectorName id="2B6EEB9F845BCC07FEB8FC42DD21FB90" box="[333,579,1014,1040]" pageId="3" pageNumber="318">Our Planet Reviewed</collectorName>
<collectionCode id="E08A168C845BCC07FDBBFC42DDC7FB90" box="[590,677,1014,1040]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="318">MNHN</collectionCode>
</materialsCitation>
/
<materialsCitation id="36F38414845BCC07FD58FC42DE19FBD7" collectingDate="2015-02-23" collectingDateMax="2015-03-25" collectingDateMin="2015-02-23" collectionCode="AMNH" country="Suriname" determinerName="J. M. Carpenter" location="Mitaraka" pageId="3" pageNumber="318" specimenCode="IZC 00178103" specimenCount="1" typeStatus="paratype">
PNI,
<location id="8344D892845BCC07FD1FFC42DC2DFB90" LSID="urn:lsid:plazi:treatment:0E323F5F8459CC00FF62FB47DB3BFDBD:8344D892845BCC07FD1FFC42DC2DFB90" box="[746,847,1014,1040]" country="Suriname" name="Mitaraka" pageId="3" pageNumber="318">Mitaraka</location>
2015, APA-973-1,
<date id="F225A889845BCC07FBC7FC42DB94FB91" box="[1074,1270,1014,1041]" pageId="3" pageNumber="318" value="2015-02-23" valueMax="2015-03-25" valueMin="2015-02-23">
<collectingDate id="E2615161845BCC07FBC7FC42DB94FB91" box="[1074,1270,1014,1041]" pageId="3" pageNumber="318" value="2015-02-23" valueMax="2015-03-25" valueMin="2015-02-23">23.II25.III.2015</collectingDate>
</date>
,
<taxonomicName id="419BF5CA845BCC07FAF0FC42DE11FBB3" authority="(Spinola)" baseAuthorityName="Spinola" baseAuthorityYear="1851" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845BCC07FAF0FC42DE66FBB3" italics="true" pageId="3" pageNumber="318">Angiopolybia paraensis</emphasis>
(Spinola)
</taxonomicName>
(det.
<determinerName id="73F8628D845BCC07FE46FBAEDD3DFBB4" box="[435,607,1050,1076]" pageId="3" pageNumber="318">J.M. Carpenter</determinerName>
),
<collectionCode id="E08A168C845BCC07FD9AFBAEDDA7FBB4" box="[623,709,1050,1076]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="318">AMNH</collectionCode>
_
<specimenCode id="D63D2632845BCC07FD26FBAEDC15FBB4" box="[723,887,1050,1076]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="318">IZC 00178103</specimenCode>
[
<collectionCode id="E08A168C845BCC07FC71FBAEDCBDFBB4" box="[900,991,1050,1076]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="318">AMNH</collectionCode>
] [specimen used for DNA extraction by
<bibRefCitation id="E20AF3B8845BCC07FF62FB8ADE16FBD7" author="Barroso, P. C. S. &amp; Menezes, R. S. T. &amp; Oliveira, M. L. &amp; Somavilla, A." box="[151,372,1085,1112]" pageId="3" pageNumber="318" pagination="75 - 97" refId="ref6283" refString="Barroso, P. C. S., Menezes, R. S. T., Oliveira, M. L. &amp; Somavilla, A. (2022) A systematic review of the Neotropical social wasp genus Angiopolybia Araujo, 1946 (Hymenoptera: Vespidae): species delimitation, morphological diagnosis, and geographical distribution. Arthropod Systematics &amp; Phylogeny, 80, 75 - 97. https: // doi. org / 10.3897 / asp. 80. e 71492" type="journal article" year="2022">
Barroso
<emphasis id="B4EF525B845BCC07FF02FB8ADE4DFBD7" box="[247,303,1085,1111]" italics="true" pageId="3" pageNumber="318">et al.</emphasis>
2022
</bibRefCitation>
]
</materialsCitation>
.
</paragraph>
<caption id="D2E4DEC1845ACC06FF62F8B6DDD6F806" ID-DOI="http://doi.org/10.5281/zenodo.14033775" ID-Zenodo-Dep="14033775" httpUri="https://zenodo.org/record/14033775/files/figure.png" pageId="2" pageNumber="317" startId="2.[151,264,1794,1818]" targetBox="[152,1435,179,1768]" targetPageId="2" targetType="figure">
<paragraph id="86248E49845ACC06FF62F8B6DDD6F806" blockId="2.[151,1437,1794,1926]" pageId="2" pageNumber="317">
<emphasis id="B4EF525B845ACC06FF62F8B6DE50F89A" bold="true" box="[151,306,1794,1818]" pageId="2" pageNumber="317">FIGURES 17</emphasis>
.
<emphasis id="B4EF525B845ACC06FEC8F8B6DE01F89A" bold="true" box="[317,355,1794,1818]" pageId="2" pageNumber="317">13</emphasis>
.
<taxonomicName id="419BF5CA845ACC06FE98F8B6DD1FF89A" authorityName="Barroso &amp; Somavilla &amp; Broad &amp; Menezes &amp; Oliveira" authorityYear="2024" box="[365,637,1794,1818]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="317" phylum="Arthropoda" rank="species" species="obscuriorina" status="sp. nov.">
<emphasis id="B4EF525B845ACC06FE98F8B6DD1FF89A" box="[365,637,1794,1818]" italics="true" pageId="2" pageNumber="317">Angiopolybia obscuriorina</emphasis>
</taxonomicName>
<emphasis id="B4EF525B845ACC06FD74F8B6DDADF89A" bold="true" box="[641,719,1794,1818]" pageId="2" pageNumber="317">
<taxonomicNameLabel id="AFDCEF20845ACC06FD74F8B6DDADF89A" box="[641,719,1794,1818]" pageId="2" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ♀.
<emphasis id="B4EF525B845ACC06FCAEF8B6DCE3F89A" bold="true" box="[859,897,1794,1818]" pageId="2" pageNumber="317">46</emphasis>
.
<taxonomicName id="419BF5CA845ACC06FC79F8B6DB10F89A" baseAuthorityName="Spinola" baseAuthorityYear="1851" box="[908,1138,1794,1818]" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="317" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845ACC06FC79F8B6DB10F89A" box="[908,1138,1794,1818]" italics="true" pageId="2" pageNumber="317">Stelopolybia paraensis</emphasis>
</taxonomicName>
var. (or subspecies)
<emphasis id="B4EF525B845ACC06FACFF8B6DAFFF89A" box="[1338,1437,1794,1818]" italics="true" pageId="2" pageNumber="317">obscurior</emphasis>
Bequaert, holotype, ♀ (current
<taxonomicName id="419BF5CA845ACC06FE2CF892DD3BF8BE" baseAuthorityName="Spinola" baseAuthorityYear="1851" box="[473,601,1830,1854]" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="317" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845ACC06FE2CF892DD3BF8BE" box="[473,601,1830,1854]" italics="true" pageId="2" pageNumber="317">A. paraensis</emphasis>
</taxonomicName>
brown).
<emphasis id="B4EF525B845ACC06FD42F892DDA6F8BE" bold="true" box="[695,708,1830,1854]" pageId="2" pageNumber="317">1</emphasis>
,
<emphasis id="B4EF525B845ACC06FD24F892DDBCF8BE" bold="true" box="[721,734,1830,1854]" pageId="2" pageNumber="317">5</emphasis>
. Habitus, lateral view.
<emphasis id="B4EF525B845ACC06FC39F892DCBBF8BE" bold="true" box="[972,985,1830,1854]" pageId="2" pageNumber="317">2</emphasis>
,
<emphasis id="B4EF525B845ACC06FC12F892DC96F8BE" bold="true" box="[999,1012,1830,1854]" pageId="2" pageNumber="317">6</emphasis>
. Habitus, dorsal view.
<emphasis id="B4EF525B845ACC06FB14F892DB8CF8BE" bold="true" box="[1249,1262,1830,1854]" pageId="2" pageNumber="317">3</emphasis>
,
<emphasis id="B4EF525B845ACC06FB0EF892DA6AF8BE" bold="true" box="[1275,1288,1830,1854]" pageId="2" pageNumber="317">4</emphasis>
. Head, frontal view.
<emphasis id="B4EF525B845ACC06FF21F8FEDF83F8E2" bold="true" box="[212,225,1866,1890]" pageId="2" pageNumber="317">7</emphasis>
. Map of distribution (© President and Fellows of Harvard College, MCZ, for images 4, 5, and 6, https://mczbase.mcz. harvard.edu/guid/MCZ:Ent:36021). Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="D2E4DEC1845BCC07FF62FD0BDEA4FC9F" ID-DOI="http://doi.org/10.5281/zenodo.14033777" ID-Zenodo-Dep="14033777" httpUri="https://zenodo.org/record/14033777/files/figure.png" pageId="3" pageNumber="318" startId="3.[151,264,703,727]" targetBox="[191,1395,181,679]" targetPageId="3" targetType="figure">
<paragraph id="86248E49845BCC07FF62FD0BDEA4FC9F" blockId="3.[151,1436,703,800]" pageId="3" pageNumber="318">
<emphasis id="B4EF525B845BCC07FF62FD0BDE20FD57" bold="true" box="[151,322,703,727]" pageId="3" pageNumber="318">FIGURES 811</emphasis>
.
<emphasis id="B4EF525B845BCC07FEA4FD0BDE3CFD57" bold="true" box="[337,350,703,727]" pageId="3" pageNumber="318">8</emphasis>
,
<emphasis id="B4EF525B845BCC07FE98FD0BDE18FD57" bold="true" box="[365,378,703,727]" pageId="3" pageNumber="318">9</emphasis>
.
<taxonomicName id="419BF5CA845BCC07FE7DFD74DDFEFD58" authorityName="Barroso &amp; Somavilla &amp; Broad &amp; Menezes &amp; Oliveira" authorityYear="2024" box="[392,668,704,728]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="obscuriorina" status="sp. nov.">
<emphasis id="B4EF525B845BCC07FE7DFD74DDFEFD58" box="[392,668,704,728]" italics="true" pageId="3" pageNumber="318">Angiopolybia obscuriorina</emphasis>
</taxonomicName>
<emphasis id="B4EF525B845BCC07FD50FD74DD9AFD58" bold="true" box="[677,760,704,728]" pageId="3" pageNumber="318">
<taxonomicNameLabel id="AFDCEF20845BCC07FD50FD74DD9AFD58" box="[677,760,704,728]" pageId="3" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ♀.
<emphasis id="B4EF525B845BCC07FC64FD0BDCC8FD57" bold="true" box="[913,938,703,727]" pageId="3" pageNumber="318">10</emphasis>
,
<emphasis id="B4EF525B845BCC07FC4FFD0BDCB0FD57" bold="true" box="[954,978,703,727]" pageId="3" pageNumber="318">11</emphasis>
.
<taxonomicName id="419BF5CA845BCC07FC14FD74DBA9FD57" baseAuthorityName="Spinola" baseAuthorityYear="1851" box="[993,1227,704,728]" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845BCC07FC14FD74DBA9FD57" box="[993,1227,704,728]" italics="true" pageId="3" pageNumber="318">Stelopolybia paraensis</emphasis>
</taxonomicName>
var. (or subspecies)
<emphasis id="B4EF525B845BCC07FF62FD50DF98FD7C" box="[151,250,740,764]" italics="true" pageId="3" pageNumber="318">obscurior</emphasis>
Bequaert, holotype, ♀ (current
<taxonomicName id="419BF5CA845BCC07FDB6FD50DDA1FD7B" baseAuthorityName="Spinola" baseAuthorityYear="1851" box="[579,707,740,763]" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845BCC07FDB6FD50DDA1FD7B" box="[579,707,740,763]" italics="true" pageId="3" pageNumber="318">A. paraensis</emphasis>
</taxonomicName>
brown).
<emphasis id="B4EF525B845BCC07FCD4FD57DC4CFD7B" bold="true" box="[801,814,739,763]" pageId="3" pageNumber="318">8</emphasis>
,
<emphasis id="B4EF525B845BCC07FCC9FD57DC37FD7B" bold="true" box="[828,853,739,763]" pageId="3" pageNumber="318">10</emphasis>
. Metapleural basalar area, lateral view.
<emphasis id="B4EF525B845BCC07FB05FD57DB9FFD7B" bold="true" box="[1264,1277,739,763]" pageId="3" pageNumber="318">9</emphasis>
,
<emphasis id="B4EF525B845BCC07FAFFFD57DA40FD7B" bold="true" box="[1290,1314,739,763]" pageId="3" pageNumber="318">11</emphasis>
. Tergum II, lateral view. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="86248E49845BCC07FF32FBD5DD0CFBFB" blockId="3.[151,1437,871,2041]" box="[199,622,1121,1148]" pageId="3" pageNumber="318">
<materialsCitation id="36F38414845BCC07FF32FBD5DD09FBFB" box="[199,619,1121,1148]" country="Suriname" location="Brokopondo" pageId="3" pageNumber="318" specimenCount="1" stateProvince="Brokopondo" typeStatus="holotype">
<emphasis id="B4EF525B845BCC07FF32FBD5DE02FBFB" bold="true" box="[199,352,1121,1147]" pageId="3" pageNumber="318">
<typeStatus id="592030EB845BCC07FF32FBD5DE60FBFB" box="[199,258,1121,1147]" pageId="3" pageNumber="318">Type</typeStatus>
locality
</emphasis>
.
<collectingRegion id="445F40AB845BCC07FE9BFBD5DE9FFBFB" box="[366,509,1121,1147]" country="Suriname" name="Brokopondo" pageId="3" pageNumber="318">Brokopondo</collectingRegion>
,
<collectingCountry id="FE8CCED9845BCC07FDFDFBD5DD09FBFB" box="[520,619,1121,1147]" name="Suriname" pageId="3" pageNumber="318">Surinam</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CE81DDC2845BCC07FF32FB30DEE2F831" pageId="3" pageNumber="318" type="description">
<paragraph id="86248E49845BCC07FF32FB30DA53F80D" blockId="3.[151,1437,871,2041]" pageId="3" pageNumber="318">
<emphasis id="B4EF525B845BCC07FF32FB30DE33FB1E" bold="true" box="[199,337,1156,1182]" pageId="3" pageNumber="318">Description</emphasis>
(measurements of the
<typeStatus id="592030EB845BCC07FDA4FB31DDD1FB1F" box="[593,691,1157,1183]" pageId="3" pageNumber="318" type="holotype">holotype</typeStatus>
are given in parentheses). FEMALE.
<emphasis id="B4EF525B845BCC07FBAAFB30DBEDFB1E" bold="true" box="[1119,1167,1156,1182]" pageId="3" pageNumber="318">Size</emphasis>
. Head
<quantity id="416323AC845BCC07FB15FB31DA30FB1F" box="[1248,1362,1157,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="3" pageNumber="318" unit="mm" value="1.95" valueMax="2.0" valueMin="1.9">1.92 mm</quantity>
(1.94) mm long, 3.43.6 (3.57) mm high, and 3.84.2 (3.97) mm wide; mesosoma 6.156.20 (6.20) mm long, fore wing 1516 (15.34) mm long, and hind wing 9.810.5 (10.36) mm long; metasoma 9.59.8 (9.6) mm long.
<emphasis id="B4EF525B845BCC07FAF5FB78DA22FB66" bold="true" box="[1280,1344,1228,1254]" pageId="3" pageNumber="318">Head</emphasis>
. Lateral ocelli 0.280.29 (0.28) mm and median ocellus 0.280.3 (0.28) mm diameter, inserted in a declivity of vertex and lateral ocelli separated from eyes by two times ocellar diameter. Eyes with very small and sparse bristles. Frons with interantennal space 1.7 times height of antennal socket. Anterior tentorial pit closer to antennal socket than to eye. Upper central region of frons with very long bristles. Antennal socket
<quantity id="416323AC845BCC07FC10FAEFDB5EFAF6" box="[997,1084,1371,1398]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="318" unit="mm" value="0.4">0.4 mm</quantity>
high. Clypeus as high as wide, in contact with eyes for approximately height of an antennal socket, with lateral lobe not touching eye. Clypeus with long bristles, but with very long bristles on apical one-third. Gena wider than half width of eye at level of ocular sinus.
<emphasis id="B4EF525B845BCC07FEDFFA72DECAFA60" bold="true" box="[298,424,1478,1504]" pageId="3" pageNumber="318">Mesosoma</emphasis>
. Anterior lamella of pronotum one-fifth height of antennal socket. Pronotal fovea ellipsoid, shallow, and with slight anterior prominence. Mesoscutum convex and 1.1 times longer than wide. Tegula 1.6 times longer than wide. Axillary fossa with anterior margin anteriorly directed. Propodeum with median longitudinal elevation in propodeal furrow, in posterior region. Propodeum with region anterior to spiracle projecting shallowly into metapleural basalar area. Propodeum without translucent posterior submedian mark, anterior to propodeal valve. Propodeal valve complete and expanded, median region half height of antennal socket. Fore wing with prestigma as long as wide. Hind wing with 1214 (12) hamuli.
<emphasis id="B4EF525B845BCC07FC9EF929DC85F937" bold="true" box="[875,999,1693,1719]" pageId="3" pageNumber="318">Metasoma</emphasis>
. Tergum I 1.6 times longer than wide, angled at posterior third in lateral view. Tergum II 0.7 times as long as broad and without row of very long bristles on posterior margin.
<emphasis id="B4EF525B845BCC07FEABF950DEC0F97E" bold="true" box="[350,418,1764,1790]" pageId="3" pageNumber="318">Color</emphasis>
. Dark brown in general. Brown: mandibles, gena, radicle of scape, flagellomere 1 dorsally, mesepisternum (except for region posterior to scrobal furrow), metapleural basalar area, coxae, tibiae, and tarsi. Yellow: lateral region of frons and spot on tegula. Yellowish-brown: flagellomeres 110. Wings with yellowish-brown cells in proximal halves and yellow cells in distal halves, and yellowish-brown venation, except for reddish-brown veins costa, subcosta+radius, radial sector, prestigma, and base of pterostigma. MALE. Unknown.
</paragraph>
<paragraph id="86248E49845BCC07FF32F823DEE2F831" blockId="3.[151,1437,871,2041]" box="[199,384,1943,1969]" pageId="3" pageNumber="318">
<emphasis id="B4EF525B845BCC07FF32F823DF9EF831" bold="true" box="[199,252,1943,1969]" pageId="3" pageNumber="318">Nest</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="CE81DDC2845BCC07FF32F80EDD26F879" pageId="3" pageNumber="318" type="distribution">
<paragraph id="86248E49845BCC07FF32F80EDD26F879" blockId="3.[151,1437,871,2041]" pageId="3" pageNumber="318">
<emphasis id="B4EF525B845BCC07FF32F80EDE3BF854" bold="true" box="[199,345,1978,2004]" pageId="3" pageNumber="318">Distribution</emphasis>
.
<collectingCountry id="FE8CCED9845BCC07FE98F80FDEAAF855" box="[365,456,1979,2005]" name="Guyana" pageId="3" pageNumber="318">Guyana</collectingCountry>
: Pakaraima Mountains, East
<collectingCountry id="FE8CCED9845BCC07FCD7F80FDC19F855" box="[802,891,1979,2005]" name="Guyana" pageId="3" pageNumber="318">Berbice</collectingCountry>
Corentyne;
<collectingCountry id="FE8CCED9845BCC07FBFAF80FDB10F855" box="[1039,1138,1979,2005]" name="Suriname" pageId="3" pageNumber="318">Surinam</collectingCountry>
:
<collectingRegion id="445F40AB845BCC07FB77F80FDA73F855" box="[1154,1297,1979,2005]" country="Suriname" name="Brokopondo" pageId="3" pageNumber="318">Brokopondo</collectingRegion>
,
<collectingRegion id="445F40AB845BCC07FAD4F80FDAFAF855" box="[1313,1432,1979,2005]" country="Suriname" name="Sipaliwini" pageId="3" pageNumber="318">Sipaliwini</collectingRegion>
; French Guiana: Maripassoula (
<figureCitation id="1EA092CC845BCC07FE07F86BDD54F879" box="[498,566,2015,2041]" captionStart="FIGURES 17. 13" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1435,179,1768]" captionTargetId="figure-17@2.[152,1435,179,1768]" captionTargetPageId="2" captionText="FIGURES 17. 13. Angiopolybia obscuriorina sp. nov., holotype, ♀. 46. Stelopolybia paraensisvar.(or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 1, 5. Habitus, lateral view. 2, 6. Habitus, dorsal view. 3, 4. Head, frontal view. 7. Map of distribution (© President and Fellows of Harvard College, MCZ, for images 4, 5, and 6, https://mczbase.mcz. harvard.edu/guid/MCZ:Ent:36021). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033775" httpUri="https://zenodo.org/record/14033775/files/figure.png" pageId="3" pageNumber="318">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CE81DDC2845CCC00FF32FF23DED4FF79" pageId="4" pageNumber="319" type="etymology">
<paragraph id="86248E49845CCC00FF32FF23DED4FF79" blockId="4.[151,1437,151,573]" pageId="4" pageNumber="319">
<emphasis id="B4EF525B845CCC00FF32FF23DE2AFF31" bold="true" box="[199,328,151,177]" pageId="4" pageNumber="319">Etymology</emphasis>
. The name
<taxonomicName id="419BF5CA845CCC00FE3AFF23DD0BFF31" authorityName="Barroso &amp; Somavilla &amp; Broad &amp; Menezes &amp; Oliveira" authorityYear="2024" box="[463,617,151,177]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="obscuriorina">
<emphasis id="B4EF525B845CCC00FE2DFF23DD0BFF31" box="[472,617,151,177]" italics="true" pageId="4" pageNumber="319">obscuriorina</emphasis>
</taxonomicName>
is a fusion of two Latin words:
<emphasis id="B4EF525B845CCC00FC29FF23DB2BFF31" box="[988,1097,151,177]" italics="true" pageId="4" pageNumber="319">obscurior</emphasis>
= subspecific epithet of an
<taxonomicName id="419BF5CA845CCC00FA71FF23DE66FF54" baseAuthorityName="Spinola" baseAuthorityYear="1851" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845CCC00FA71FF23DE66FF54" italics="true" pageId="4" pageNumber="319">A. paraensis</emphasis>
</taxonomicName>
variant, designated a morph by
<bibRefCitation id="E20AF3B8845CCC00FD9BFF0FDC41FF55" author="Richards, O. W." box="[622,803,187,213]" pageId="4" pageNumber="319" refId="ref7469" refString="Richards, O. W. (1978) The social wasps of the Americas, excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards (1978)</bibRefCitation>
+
<emphasis id="B4EF525B845CCC00FCB7FF0FDC04FF54" box="[834,870,187,212]" italics="true" pageId="4" pageNumber="319">ina</emphasis>
= likeness. The name is based on the resemblance to the variant
<emphasis id="B4EF525B845CCC00FEB7FF6BDECDFF79" box="[322,431,223,249]" italics="true" pageId="4" pageNumber="319">obscurior</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="CE81DDC2845CCC00FF32FEB6DB3BFDBD" pageId="4" pageNumber="319" type="discussion">
<paragraph id="86248E49845CCC00FF32FEB6DB3BFDBD" blockId="4.[151,1437,151,573]" pageId="4" pageNumber="319">
<emphasis id="B4EF525B845CCC00FF32FEB6DE2AFE9C" bold="true" box="[199,328,258,284]" pageId="4" pageNumber="319">Comments</emphasis>
. While the specimens designated as
<typeStatus id="592030EB845CCC00FD2BFEB7DC2BFE9D" box="[734,841,259,285]" pageId="4" pageNumber="319" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="419BF5CA845CCC00FC98FEB7DA6DFE9D" box="[877,1295,259,285]" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="variety" species="paraensis" variety="obscurior">
<emphasis id="B4EF525B845CCC00FC98FEB7DB0EFE9C" box="[877,1132,259,285]" italics="true" pageId="4" pageNumber="319">Stelopolybia paraensis</emphasis>
var.
<emphasis id="B4EF525B845CCC00FB57FEB7DA6DFE9D" box="[1186,1295,259,285]" italics="true" pageId="4" pageNumber="319">obscurior</emphasis>
</taxonomicName>
by
<bibRefCitation id="E20AF3B8845CCC00FAC2FEB7DF80FEC1" author="Bequaert, J. C." pageId="4" pageNumber="319" pagination="247 - 304" refId="ref6433" refString="Bequaert, J. C. (1944) The social Vespidae of the Guianas, particularly of British Guiana. Bulletin of the Museum of Comparative Zoology, 94, 247 - 304." type="journal article" year="1944">Bequaert (1944)</bibRefCitation>
from
<collectingCountry id="FE8CCED9845CCC00FED0FE93DE1EFEC1" box="[293,380,295,321]" name="Guyana" pageId="4" pageNumber="319">Guyana</collectingCountry>
(Mountain Roraima and
<collectingCountry id="FE8CCED9845CCC00FD66FE93DC64FEC1" box="[659,774,295,321]" name="Guyana" pageId="4" pageNumber="319">Essequibo</collectingCountry>
River) and
<collectingCountry id="FE8CCED9845CCC00FC7CFE93DC8AFEC1" box="[905,1000,295,321]" name="Suriname" pageId="4" pageNumber="319">Surinam</collectingCountry>
(Brownsberg) have not been examined in detail, because of their geographical distribution we believe that these specimens also correspond to the new species,
<taxonomicName id="419BF5CA845CCC00FF02FEDBDECBFE09" authorityName="Barroso &amp; Somavilla &amp; Broad &amp; Menezes &amp; Oliveira" authorityYear="2024" box="[247,425,367,393]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="obscuriorina" status="sp. nov.">
<emphasis id="B4EF525B845CCC00FF02FEDBDECBFE09" box="[247,425,367,393]" italics="true" pageId="4" pageNumber="319">A. obscuriorina</emphasis>
</taxonomicName>
<emphasis id="B4EF525B845CCC00FE44FEDBDD6EFE09" bold="true" box="[433,524,367,393]" pageId="4" pageNumber="319">
<taxonomicNameLabel id="AFDCEF20845CCC00FE44FEDBDD6EFE09" box="[433,524,367,393]" pageId="4" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Also, specimens from Pakaraima Mountains in
<collectingCountry id="FE8CCED9845CCC00FBC5FEDBDBE5FE09" box="[1072,1159,367,393]" name="Guyana" pageId="4" pageNumber="319">Guyana</collectingCountry>
were excluded from the distribution map due to the challenges in precisely determining the locality. Moreover, the distribution map (
<figureCitation id="1EA092CC845CCC00FA9AFE27DFC4FE51" captionStart="FIGURES 17. 13" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1435,179,1768]" captionTargetId="figure-17@2.[152,1435,179,1768]" captionTargetPageId="2" captionText="FIGURES 17. 13. Angiopolybia obscuriorina sp. nov., holotype, ♀. 46. Stelopolybia paraensisvar.(or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 1, 5. Habitus, lateral view. 2, 6. Habitus, dorsal view. 3, 4. Head, frontal view. 7. Map of distribution (© President and Fellows of Harvard College, MCZ, for images 4, 5, and 6, https://mczbase.mcz. harvard.edu/guid/MCZ:Ent:36021). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033775" httpUri="https://zenodo.org/record/14033775/files/figure.png" pageId="4" pageNumber="319">Fig. 7</figureCitation>
) highlights distinct geographic discontinuity between brown specimens of
<taxonomicName id="419BF5CA845CCC00FC03FE03DBE1FE50" baseAuthorityName="Spinola" baseAuthorityYear="1851" box="[1014,1155,439,465]" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845CCC00FC03FE03DBE1FE50" box="[1014,1155,439,465]" italics="true" pageId="4" pageNumber="319">A. paraensis</emphasis>
</taxonomicName>
and the newly identified
<taxonomicName id="419BF5CA845CCC00FF62FE6FDE28FE75" authorityName="Barroso &amp; Somavilla &amp; Broad &amp; Menezes &amp; Oliveira" authorityYear="2024" box="[151,330,475,501]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="obscuriorina" status="sp. nov.">
<emphasis id="B4EF525B845CCC00FF62FE6FDE28FE75" box="[151,330,475,501]" italics="true" pageId="4" pageNumber="319">A. obscuriorina</emphasis>
</taxonomicName>
<emphasis id="B4EF525B845CCC00FEA1FE6FDED2FE75" bold="true" box="[340,432,475,501]" pageId="4" pageNumber="319">
<taxonomicNameLabel id="AFDCEF20845CCC00FEA1FE6FDED2FE75" box="[340,432,475,501]" pageId="4" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
While
<taxonomicName id="419BF5CA845CCC00FDFDFE6FDDF5FE74" baseAuthorityName="Spinola" baseAuthorityYear="1851" box="[520,663,475,501]" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845CCC00FDFDFE6FDDF5FE74" box="[520,663,475,501]" italics="true" pageId="4" pageNumber="319">A. paraensis</emphasis>
</taxonomicName>
predominantly inhabits regions proximal to the Andes of
<collectingCountry id="FE8CCED9845CCC00FACCFE6FDAFAFE75" box="[1337,1432,475,501]" name="Ecuador" pageId="4" pageNumber="319">Ecuador</collectingCountry>
,
<collectingCountry id="FE8CCED9845CCC00FF62FE4BDE6AFD99" box="[151,264,511,537]" name="Colombia" pageId="4" pageNumber="319">Colombia</collectingCountry>
, and northwestern
<collectingRegion id="445F40AB845CCC00FE12FE4BDD02FD99" box="[487,608,511,537]" country="Colombia" name="Amazonas" pageId="4" pageNumber="319">Amazonas</collectingRegion>
,
<collectingCountry id="FE8CCED9845CCC00FD9AFE4BDDD4FD99" box="[623,694,511,537]" name="Brazil" pageId="4" pageNumber="319">Brazil</collectingCountry>
, the newly discovered species has a confined distribution, with populations exclusively found in the plains of
<collectingCountry id="FE8CCED9845CCC00FD6BFD97DD9AFDBD" box="[670,760,547,573]" name="Guyana" pageId="4" pageNumber="319">Guyana</collectingCountry>
,
<collectingCountry id="FE8CCED9845CCC00FCF6FD97DC13FDBD" box="[771,881,547,573]" name="Suriname" pageId="4" pageNumber="319">Suriname</collectingCountry>
, and French Guiana.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,213 @@
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<mods:title id="0067DF871ECD054083BBD22252FA5466">Contributing to the understanding of Angiopolybia Araujo, 1946 (Hymenoptera, Vespidae, Polistinae): a new species, and descriptions of the nest of A. obidensis and male of A. zischkai</mods:title>
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<mods:namePart id="45D4D49BAAB9FE9C84487E917301DF46">Barroso, Paulo Cézar Salgado</mods:namePart>
<mods:affiliation id="CDFA46D669BCA0E76E193F5036223867">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade, Av. André Araújo, 69067 - 375, Manaus, Brazil.</mods:affiliation>
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<mods:namePart id="C6BAA0D0BB6BB86140E951720C1CB902">Somavilla, Alexandre</mods:namePart>
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<mods:affiliation id="CD29F51B2AD6EAD0F2EB4ACCB60D9112">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade, Av. André Araújo, 69067 - 375, Manaus, Brazil.</mods:affiliation>
<mods:nameIdentifier id="284BCCD91F932892F14B9E0D45C7181B" type="email">alexandresomavilla@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FA8D6FCDD109B2D8D421DF64DDA690F8">Broad, Gavin R.</mods:namePart>
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<mods:affiliation id="4B27D3A1FC710FBD8C8AA1288F9227BC">Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
<mods:nameIdentifier id="451F6A801CBE3CED5BB94C4A08554E64" type="email">g.broad@nhm.ac.uk</mods:nameIdentifier>
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<mods:namePart id="3B34D08DD1010F04A7F906468B17E41C">Menezes, Rodolpho Santos Telles</mods:namePart>
<mods:nameIdentifier id="E3FF1B83B50A5EB4704706DC0755907E" type="ORCID">0000-0002-6612-3543</mods:nameIdentifier>
<mods:affiliation id="96FFC77245221AF433F9E33B65B8CA23">Universidade Estadual de Santa Cruz, Departamento de Ciências Biológicas, Rodovia Jorge Amado, km 16, 45662 - 900, Ilhéus, Brazil.</mods:affiliation>
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<mods:namePart id="1EF6A1118D2883295CC5ED3DA9BEE71F">Oliveira, Marcio Luiz De</mods:namePart>
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<mods:affiliation id="9CFCAD9CA2CDC3B2B4D6A522227BC7D2">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade, Av. André Araújo, 69067 - 375, Manaus, Brazil. &amp; Instituto Nacional de Biodiversidad, División de Entomología, Av. Rumipamba y Av. de los Shyris, RG 77 + GQV, Quito, Ecuador.</mods:affiliation>
<mods:nameIdentifier id="F081097C493A94EF10870D9593938711" type="email">marciooliveirainpa@gmail.com</mods:nameIdentifier>
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<subSubSection id="CE81DDC2845CCC00FF62FD3ADD06FD28" box="[151,612,654,681]" pageId="4" pageNumber="319" type="nomenclature">
<paragraph id="86248E49845CCC00FF62FD3ADD06FD28" blockId="4.[151,612,654,717]" box="[151,612,654,681]" pageId="4" pageNumber="319">
<heading id="DD6C3925845CCC00FF62FD3ADD06FD28" bold="true" box="[151,612,654,681]" fontSize="11" level="1" pageId="4" pageNumber="319" reason="1">
<taxonomicName id="419BF5CA845CCC00FF62FD3ADD06FD28" ID-CoL="DZ9Z" authority="(Ducke, 1904)" baseAuthorityName="Ducke" baseAuthorityYear="1904" box="[151,612,654,681]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="obidensis">
<emphasis id="B4EF525B845CCC00FF62FD3ADD06FD28" bold="true" box="[151,612,654,681]" pageId="4" pageNumber="319">
<emphasis id="B4EF525B845CCC00FF62FD3ADED2FD29" bold="true" box="[151,432,654,681]" italics="true" pageId="4" pageNumber="319">Angiopolybia obidensis</emphasis>
(
<bibRefCitation id="E20AF3B8845CCC00FE35FD3ADD3EFD28" author="Ducke, A." box="[448,604,654,680]" pageId="4" pageNumber="319" pagination="317 - 374" refId="ref7065" refString="Ducke, A. (1904) Sobre as vespidas sociais do Para. Extracto do Boletim do Museu Goeldi, 4, 317 - 374. https: // doi. org / 10.5962 / bhl. title. 100023" type="journal article" year="1904">Ducke, 1904</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="CE81DDC2845CCC00FF62FD07DBC5F8CF" pageId="4" pageNumber="319" type="description">
<paragraph id="86248E49845CCC00FF62FD07DE45FD4D" blockId="4.[151,612,654,717]" box="[151,295,691,717]" pageId="4" pageNumber="319">
(
<figureCitation id="1EA092CC845CCC00FF6AFD07DE7DFD4D" box="[159,287,691,717]" captionStart="FIGURES 1214" captionStartId="4.[151,264,1558,1582]" captionTargetBox="[191,1395,993,1535]" captionTargetId="figure-312@4.[191,1395,993,1535]" captionTargetPageId="4" captionText="FIGURES 1214. Nests of Angiopolybia obidensis (Ducke). 12. Location where the nests were found. 13. Occupied nest. 14. Abandoned nest." figureDoi="http://doi.org/10.5281/zenodo.14033779" httpUri="https://zenodo.org/record/14033779/files/figure.png" pageId="4" pageNumber="319">Figs 1214</figureCitation>
)
</paragraph>
<paragraph id="86248E49845CCC00FF62FD4FDE07FC49" blockId="4.[151,1437,763,969]" pageId="4" pageNumber="319">
<emphasis id="B4EF525B845CCC00FF62FD4FDFAEFC95" bold="true" box="[151,204,763,789]" pageId="4" pageNumber="319">Nest</emphasis>
. Although the nest of
<taxonomicName id="419BF5CA845CCC00FE49FD4FDD21FC95" baseAuthorityName="Ducke" baseAuthorityYear="1904" box="[444,579,763,789]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="obidensis">
<emphasis id="B4EF525B845CCC00FE49FD4FDD21FC95" box="[444,579,763,789]" italics="true" pageId="4" pageNumber="319">A. obidensis</emphasis>
</taxonomicName>
could not be collected due to its location in a cave protected by environmental law (
<figureCitation id="1EA092CC845CCC00FF25FCABDE46FCB9" box="[208,292,799,825]" captionStart="FIGURES 1214" captionStartId="4.[151,264,1558,1582]" captionTargetBox="[191,1395,993,1535]" captionTargetId="figure-312@4.[191,1395,993,1535]" captionTargetPageId="4" captionText="FIGURES 1214. Nests of Angiopolybia obidensis (Ducke). 12. Location where the nests were found. 13. Occupied nest. 14. Abandoned nest." figureDoi="http://doi.org/10.5281/zenodo.14033779" httpUri="https://zenodo.org/record/14033779/files/figure.png" pageId="4" pageNumber="319">Fig. 12</figureCitation>
), certain characteristics can still be described. The nest is approximately
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high and
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wide, and connected to the substrate by a
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diameter base. The envelope is oval-shaped, made of flexible paper with long fibers, and has an entrance about
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wide that does not project downward as a tube. The general color of the nest is amber with some brown fibers (
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). Additionally, the overlapping combs of an abandoned nest can be seen in
<figureCitation id="1EA092CC845CCC00FEFBFC1BDE03FC49" box="[270,353,943,969]" captionStart="FIGURES 1214" captionStartId="4.[151,264,1558,1582]" captionTargetBox="[191,1395,993,1535]" captionTargetId="figure-312@4.[191,1395,993,1535]" captionTargetPageId="4" captionText="FIGURES 1214. Nests of Angiopolybia obidensis (Ducke). 12. Location where the nests were found. 13. Occupied nest. 14. Abandoned nest." figureDoi="http://doi.org/10.5281/zenodo.14033779" httpUri="https://zenodo.org/record/14033779/files/figure.png" pageId="4" pageNumber="319">Fig. 14</figureCitation>
.
</paragraph>
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<paragraph id="86248E49845CCC00FF62F9A2DE20F9D2" blockId="4.[151,1436,1558,1618]" pageId="4" pageNumber="319">
<emphasis id="B4EF525B845CCC00FF62F9A2DE2CF9AE" bold="true" box="[151,334,1558,1582]" pageId="4" pageNumber="319">FIGURES 1214</emphasis>
. Nests of
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<emphasis id="B4EF525B845CCC00FE43F9A2DDC4F9AE" box="[438,678,1558,1582]" italics="true" pageId="4" pageNumber="319">Angiopolybia obidensis</emphasis>
(Ducke)
</taxonomicName>
.
<emphasis id="B4EF525B845CCC00FCF9F9A2DC47F9AE" bold="true" box="[780,805,1558,1582]" pageId="4" pageNumber="319">12</emphasis>
. Location where the nests were found.
<emphasis id="B4EF525B845CCC00FB4DF9A2DBB3F9AE" bold="true" box="[1208,1233,1558,1582]" pageId="4" pageNumber="319">13</emphasis>
. Occupied nest.
<emphasis id="B4EF525B845CCC00FA88F9A2DAF4F9AE" bold="true" box="[1405,1430,1558,1582]" pageId="4" pageNumber="319">14</emphasis>
. Abandoned nest.
</paragraph>
</caption>
<paragraph id="86248E49845CCC00FF32F911DBC5F8CF" blockId="4.[151,1437,1701,2015]" pageId="4" pageNumber="319">
The nest of
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<emphasis id="B4EF525B845CCC00FEA6F911DEBCF93F" box="[339,478,1701,1727]" italics="true" pageId="4" pageNumber="319">A. obidensis</emphasis>
</taxonomicName>
differs from the typical form of
<taxonomicName id="419BF5CA845CCC00FCAEF911DCAFF93F" baseAuthorityName="Lepeletier" baseAuthorityYear="1836" box="[859,973,1701,1727]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B4EF525B845CCC00FCAEF911DCAFF93F" box="[859,973,1701,1727]" italics="true" pageId="4" pageNumber="319">A. pallens</emphasis>
</taxonomicName>
by having an oval-shaped envelope and lacking a tubular projection at the entrance. These features also appear to differ from those of
<taxonomicName id="419BF5CA845CCC00FB24F97DDA2EF963" authorityName="Richards" authorityYear="1978" box="[1233,1356,1737,1763]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis id="B4EF525B845CCC00FB24F97DDA2EF963" box="[1233,1356,1737,1763]" italics="true" pageId="4" pageNumber="319">A. zischkai</emphasis>
</taxonomicName>
, based on the comments by
<bibRefCitation id="E20AF3B8845CCC00FE7FF959DDE4F887" author="Valverde, J. P. &amp; Hanson, P. &amp; Carpenter, J." box="[394,646,1773,1799]" pageId="4" pageNumber="319" pagination="174 - 199" refId="ref8007" refString="Valverde, J. P., Hanson, P. &amp; Carpenter, J. (2019) Keys to the Costa Rican species of paper wasps (Hymenoptera: Vespidae: Polistinae). Revista de Biologia Tropical, 67 (2), 174 - 199. https: // doi. org / 10.15517 / rbt. v 67 i 2 SUPL. 37229" type="journal article" year="2019">
Valverde
<emphasis id="B4EF525B845CCC00FE02F95ADD50F887" box="[503,562,1773,1799]" italics="true" pageId="4" pageNumber="319">et al.</emphasis>
(2019)
</bibRefCitation>
in the identification key to social wasps of
<collectingCountry id="FE8CCED9845CCC00FB76F959DA60F887" box="[1155,1282,1773,1799]" name="Costa Rica" pageId="4" pageNumber="319">Costa Rica</collectingCountry>
, that the nest envelope is like an inverted flask. However, the nest of
<taxonomicName id="419BF5CA845CCC00FD0BF8A5DCE5F8AB" baseAuthorityName="Ducke" baseAuthorityYear="1904" box="[766,903,1809,1835]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="obidensis">
<emphasis id="B4EF525B845CCC00FD0BF8A5DCE5F8AB" box="[766,903,1809,1835]" italics="true" pageId="4" pageNumber="319">A. obidensis</emphasis>
</taxonomicName>
resembles that of
<taxonomicName id="419BF5CA845CCC00FBA4F8A5DBBEF8AB" baseAuthorityName="Spinola" baseAuthorityYear="1851" box="[1105,1244,1809,1835]" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845CCC00FBA4F8A5DBBEF8AB" box="[1105,1244,1809,1835]" italics="true" pageId="4" pageNumber="319">A. paraensis</emphasis>
</taxonomicName>
, despite the brief descriptions provided by
<bibRefCitation id="E20AF3B8845CCC00FE46F881DD33F8CF" author="Schulz, W. A." box="[435,593,1845,1871]" pageId="4" pageNumber="319" pagination="757 - 832" refId="ref7501" refString="Schulz, W. A. (1903) Hymenopteren Amazoniens. Sitzungsberichte der mathematisch-physikalischen Klasse, 33, 757 - 832." type="journal article" year="1903">Schulz (1903)</bibRefCitation>
and
<bibRefCitation id="E20AF3B8845CCC00FD72F881DC46F8CF" author="Ducke, A." box="[647,804,1845,1871]" pageId="4" pageNumber="319" pagination="449 - 544" refId="ref7135" refString="Ducke, A. (1910) Revision des guepes sociales polygames d'Amerique. Annales Musei Nationalis Hungarici, 8, 449 - 544." type="journal article" year="1910">Ducke (1910)</bibRefCitation>
, as noted by
<bibRefCitation id="E20AF3B8845CCC00FC42F881DBC1F8CF" author="Barroso, P. C. S. &amp; Menezes, R. S. T. &amp; Oliveira, M. L. &amp; Somavilla, A." box="[951,1187,1845,1871]" pageId="4" pageNumber="319" pagination="75 - 97" refId="ref6283" refString="Barroso, P. C. S., Menezes, R. S. T., Oliveira, M. L. &amp; Somavilla, A. (2022) A systematic review of the Neotropical social wasp genus Angiopolybia Araujo, 1946 (Hymenoptera: Vespidae): species delimitation, morphological diagnosis, and geographical distribution. Arthropod Systematics &amp; Phylogeny, 80, 75 - 97. https: // doi. org / 10.3897 / asp. 80. e 71492" type="journal article" year="2022">
Barroso
<emphasis id="B4EF525B845CCC00FBE3F882DB2DF8CF" box="[1046,1103,1845,1871]" italics="true" pageId="4" pageNumber="319">et al.</emphasis>
(2022)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CE81DDC2845CCC01FF32F8ECDEDEFE9D" lastPageId="5" lastPageNumber="320" pageId="4" pageNumber="319" type="discussion">
<paragraph id="86248E49845CCC01FF32F8ECDEDEFE9D" blockId="4.[151,1437,1701,2015]" lastBlockId="5.[151,1436,151,285]" lastPageId="5" lastPageNumber="320" pageId="4" pageNumber="319">
<emphasis id="B4EF525B845CCC00FF32F8ECDE2AF8F2" bold="true" box="[199,328,1880,1906]" pageId="4" pageNumber="319">Comments</emphasis>
. The nest of
<taxonomicName id="419BF5CA845CCC00FE2BF8EDDD0BF8F3" baseAuthorityName="Ducke" baseAuthorityYear="1904" box="[478,617,1881,1907]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="obidensis">
<emphasis id="B4EF525B845CCC00FE2BF8EDDD0BF8F3" box="[478,617,1881,1907]" italics="true" pageId="4" pageNumber="319">A. obidensis</emphasis>
</taxonomicName>
was found in a cave in the Presidente Figueiredo Municipality of
<collectingCountry id="FE8CCED9845CCC00FAADF8EDDAFEF8F3" box="[1368,1436,1881,1907]" name="Brazil" pageId="4" pageNumber="319">Brazil</collectingCountry>
(Caverna Refúgio do Maroaga,
<geoCoordinate id="E3AFE88E845CCC00FDF6F8C9DDEDF817" box="[515,655,1917,1943]" degrees="2" direction="south" minutes="03" orientation="latitude" pageId="4" pageNumber="319" precision="1" seconds="03.6" value="-2.0509999">2°0303.6”S</geoCoordinate>
<geoCoordinate id="E3AFE88E845CCC00FD6DF8C9DC22F818" box="[664,832,1917,1944]" degrees="59" direction="west" minutes="58" orientation="longitude" pageId="4" pageNumber="319" precision="1" seconds="14.0" value="-59.970554">59°5814.0”W</geoCoordinate>
). It was attached to the wall at a height of about 2.2 meters above the ground. This location resembles the one described by
<bibRefCitation id="E20AF3B8845CCC00FC40F815DB36F83B" author="Schulz, W. A." box="[949,1108,1953,1979]" pageId="4" pageNumber="319" pagination="757 - 832" refId="ref7501" refString="Schulz, W. A. (1903) Hymenopteren Amazoniens. Sitzungsberichte der mathematisch-physikalischen Klasse, 33, 757 - 832." type="journal article" year="1903">Schulz (1903)</bibRefCitation>
, where a nest of
<taxonomicName id="419BF5CA845CCC00FAE5F815DAFEF83B" baseAuthorityName="Spinola" baseAuthorityYear="1851" box="[1296,1436,1953,1979]" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845CCC00FAE5F815DAFEF83B" box="[1296,1436,1953,1979]" italics="true" pageId="4" pageNumber="319">A. paraensis</emphasis>
</taxonomicName>
was found beneath a large stone park table in what is now Rodrigues Alves Park, Belém,
<collectingCountry id="FE8CCED9845CCC00FB75F871DBAAF85F" box="[1152,1224,1989,2015]" name="Brazil" pageId="4" pageNumber="319">Brazil</collectingCountry>
. The nests of these species might be difficult to find due to their nesting preferences, as they build nests in protected environments and on rocky substrates, which are uncommon in the Amazon Forest. Another possibility is that these species nest in cavities, not necessarily in rocky substrates, making their nests difficult to locate, similar to most species of
<taxonomicName id="419BF5CA845DCC01FAB0FF6BDE6BFE9D" authority="Lepeletier" authorityName="Lepeletier" class="Insecta" family="Vespidae" genus="Agelaia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B4EF525B845DCC01FAB0FF6BDAFEFF79" box="[1349,1436,223,249]" italics="true" pageId="5" pageNumber="320">Agelaia</emphasis>
Lepeletier
</taxonomicName>
(
<bibRefCitation id="E20AF3B8845DCC01FEE2FEB7DECDFE9D" author="Wenzel, J. W." box="[279,431,259,285]" pageId="5" pageNumber="320" pagination="1 - 39" refId="ref8075" refString="Wenzel, J. W. (1998) A generic key to the nests of hornets, yellowjackets, and paper wasps worldwide (Vespidae: Vespinae, Polistinae). American Museum Novitates, 3224, 1 - 39." type="journal article" year="1998">Wenzel 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,272 @@
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<mods:title id="0874B5C5965AD96134ED915DEB5662AA">Contributing to the understanding of Angiopolybia Araujo, 1946 (Hymenoptera, Vespidae, Polistinae): a new species, and descriptions of the nest of A. obidensis and male of A. zischkai</mods:title>
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<mods:location id="7DA7EC828275860B283DBFE3ABF63B84">
<mods:url id="7C3AB81F438CAE0B795A010E8DFB4044">http://dx.doi.org/10.11646/zootaxa.5536.2.7</mods:url>
</mods:location>
<mods:classification id="1F66C6CDA6085827851E3080C2C9914E">journal article</mods:classification>
<mods:identifier id="CB0255AA97ADE680DF74ED39BF0BC401" type="DOI">10.11646/zootaxa.5536.2.7</mods:identifier>
<mods:identifier id="61DBA96C17C9DC7E29B3F26A55E6C828" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C3498107664578848259E5DBACD9B88F" type="Zenodo-Dep">14033773</mods:identifier>
<mods:identifier id="F9355CF122BE9F7EC5B41E2B54E8A9D9" type="ZooBank">AF4A514F-1DC2-4D70-AEAB-8C55DBB9CA2A</mods:identifier>
</mods:mods>
<treatment id="0E323F5F845DCC02FF62FEDADEC9FC01" ID-DOI="http://doi.org/10.5281/zenodo.14033751" ID-Zenodo-Dep="14033751" LSID="urn:lsid:plazi:treatment:0E323F5F845DCC02FF62FEDADEC9FC01" httpUri="http://treatment.plazi.org/id/0E323F5F845DCC02FF62FEDADEC9FC01" lastPageId="6" lastPageNumber="321" pageId="5" pageNumber="320">
<subSubSection id="CE81DDC2845DCC01FF62FEDADD07FE08" box="[151,613,366,393]" pageId="5" pageNumber="320" type="nomenclature">
<paragraph id="86248E49845DCC01FF62FEDADD07FE08" blockId="5.[151,613,366,429]" box="[151,613,366,393]" pageId="5" pageNumber="320">
<heading id="DD6C3925845DCC01FF62FEDADD07FE08" bold="true" box="[151,613,366,393]" fontSize="11" level="1" pageId="5" pageNumber="320" reason="1">
<taxonomicName id="419BF5CA845DCC01FF62FEDADD07FE08" ID-CoL="66R66" authority="Richards, 1978" authorityName="Richards" authorityYear="1978" box="[151,613,366,393]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis id="B4EF525B845DCC01FF62FEDADD07FE08" bold="true" box="[151,613,366,393]" pageId="5" pageNumber="320">
<emphasis id="B4EF525B845DCC01FF62FEDADEC0FE09" bold="true" box="[151,418,366,393]" italics="true" pageId="5" pageNumber="320">Angiopolybia zischkai</emphasis>
<bibRefCitation id="E20AF3B8845DCC01FE5CFEDADD07FE08" author="Richards, O. W." box="[425,613,366,392]" pageId="5" pageNumber="320" refId="ref7469" refString="Richards, O. W. (1978) The social wasps of the Americas, excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards, 1978</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CE81DDC2845DCC01FF62FE27DE45FE2D" box="[151,295,402,429]" pageId="5" pageNumber="320" type="description">
<paragraph id="86248E49845DCC01FF62FE27DE45FE2D" blockId="5.[151,613,366,429]" box="[151,295,402,429]" pageId="5" pageNumber="320">
(
<figureCitation id="1EA092CC845DCC01FF6AFE27DE7DFE2D" box="[159,287,402,429]" captionStart="FIGURES 1522" captionStartId="5.[151,264,1312,1336]" captionTargetBox="[190,1397,669,1289]" captionTargetId="figure-171@5.[190,1397,669,1289]" captionTargetPageId="5" captionText="FIGURES 1522. Angiopolybia zischkai Richards, ♂. 15. Habitus, lateral view. 16. Habitus, dorsal view. 17. Head, frontal view. 18. Paramere. 19. Aedeagus, lateral view. 20. Aedeagus, ventral view. 21. Digitus. 22. Cuspis. Scale bar: 1 mm for 1517 and 0.5 mm for 1822." figureDoi="http://doi.org/10.5281/zenodo.14033781" httpUri="https://zenodo.org/record/14033781/files/figure.png" pageId="5" pageNumber="320">Figs 1522</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CE81DDC2845DCC01FF62FE6EDC9EFD05" pageId="5" pageNumber="320" type="diagnosis">
<paragraph id="86248E49845DCC01FF62FE6EDC9EFD05" blockId="5.[151,1436,474,645]" pageId="5" pageNumber="320">
<emphasis id="B4EF525B845DCC01FF62FE6EDEC9FE74" bold="true" box="[151,427,474,500]" pageId="5" pageNumber="320">Comparative diagnosis</emphasis>
. The male of
<taxonomicName id="419BF5CA845DCC01FDA5FE6FDDA9FE75" authorityName="Richards" authorityYear="1978" box="[592,715,475,501]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis id="B4EF525B845DCC01FDA5FE6FDDA9FE75" box="[592,715,475,501]" italics="true" pageId="5" pageNumber="320">A. zischkai</emphasis>
</taxonomicName>
is particularly similar to that of
<taxonomicName id="419BF5CA845DCC01FBB6FE6FDBD7FE75" baseAuthorityName="Lepeletier" baseAuthorityYear="1836" box="[1091,1205,475,501]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B4EF525B845DCC01FBB6FE6FDBD7FE75" box="[1091,1205,475,501]" italics="true" pageId="5" pageNumber="320">A. pallens</emphasis>
</taxonomicName>
. However, the male of
<taxonomicName id="419BF5CA845DCC01FF42FE4BDE53FD99" authorityName="Richards" authorityYear="1978" box="[183,305,511,537]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis id="B4EF525B845DCC01FF42FE4BDE53FD99" box="[183,305,511,537]" italics="true" pageId="5" pageNumber="320">A. zischkai</emphasis>
</taxonomicName>
differs by the lobe of the paramere, which is not curved downwards; the parameral spine with long bristles; apical portion of aedeagus wider in the anterior half; aedeagus denticulation in the form of large, wide denticles in the posterior half and small, slender denticles in the anterior half; digitus without a projection adjacent to the basal articulation; cuspis with small bristles, dense throughout the area.
</paragraph>
</subSubSection>
<caption id="D2E4DEC1845DCC01FF62FA94DEE2FA00" ID-DOI="http://doi.org/10.5281/zenodo.14033781" ID-Zenodo-Dep="14033781" httpUri="https://zenodo.org/record/14033781/files/figure.png" pageId="5" pageNumber="320" startId="5.[151,264,1312,1336]" targetBox="[190,1397,669,1289]" targetPageId="5" targetType="figure">
<paragraph id="86248E49845DCC01FF62FA94DEE2FA00" blockId="5.[151,1437,1312,1408]" pageId="5" pageNumber="320">
<emphasis id="B4EF525B845DCC01FF62FA94DE32FAB8" bold="true" box="[151,336,1312,1336]" pageId="5" pageNumber="320">FIGURES 1522</emphasis>
.
<taxonomicName id="419BF5CA845DCC01FEAAFA94DDC5FAB8" authority="Richards" authorityName="Richards" authorityYear="1978" box="[351,679,1312,1336]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis id="B4EF525B845DCC01FEAAFA94DD20FAB8" box="[351,578,1312,1336]" italics="true" pageId="5" pageNumber="320">Angiopolybia zischkai</emphasis>
Richards
</taxonomicName>
, ♂.
<emphasis id="B4EF525B845DCC01FD20FA94DD8CFAB8" bold="true" box="[725,750,1312,1336]" pageId="5" pageNumber="320">15</emphasis>
. Habitus, lateral view.
<emphasis id="B4EF525B845DCC01FC14FA94DC98FAB8" bold="true" box="[993,1018,1312,1336]" pageId="5" pageNumber="320">16</emphasis>
. Habitus, dorsal view.
<emphasis id="B4EF525B845DCC01FB19FA94DA67FAB8" bold="true" box="[1260,1285,1312,1336]" pageId="5" pageNumber="320">17</emphasis>
. Head, frontal view.
<emphasis id="B4EF525B845DCC01FF26FAF0DF8EFADC" bold="true" box="[211,236,1348,1372]" pageId="5" pageNumber="320">18</emphasis>
. Paramere.
<emphasis id="B4EF525B845DCC01FE90FAF0DE1CFADC" bold="true" box="[357,382,1348,1372]" pageId="5" pageNumber="320">19</emphasis>
. Aedeagus, lateral view.
<emphasis id="B4EF525B845DCC01FD89FAF0DDF7FADC" bold="true" box="[636,661,1348,1372]" pageId="5" pageNumber="320">20</emphasis>
. Aedeagus, ventral view.
<emphasis id="B4EF525B845DCC01FC6FFAF0DCD1FADC" bold="true" box="[922,947,1348,1372]" pageId="5" pageNumber="320">21</emphasis>
. Digitus.
<emphasis id="B4EF525B845DCC01FBE2FAF0DB52FADC" bold="true" box="[1047,1072,1348,1372]" pageId="5" pageNumber="320">22</emphasis>
. Cuspis. Scale bar: 1 mm for 1517 and 0.5 mm for 1822.
</paragraph>
</caption>
<subSubSection id="CE81DDC2845DCC01FF32FA66DD74F990" pageId="5" pageNumber="320" type="materials_examined">
<paragraph id="86248E49845DCC01FF32FA66DD74F990" blockId="5.[151,1437,1490,2021]" pageId="5" pageNumber="320">
<emphasis id="B4EF525B845DCC01FF32FA66DECCFA6C" bold="true" box="[199,430,1490,1516]" pageId="5" pageNumber="320">Material examined</emphasis>
.
<materialsCitation id="36F38414845DCC01FE35FA66DD70F990" collectingDate="1991-01-06" collectionCode="AMNH" collectorName="F. D. Parker" country="Costa Rica" location="Bijagua" pageId="5" pageNumber="320" specimenCode="IZC 00390604" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela">
<specimenCount id="909D45C0845DCC01FE35FA66DE87FA6C" box="[448,485,1490,1516]" count="1" pageId="5" pageNumber="320" type="male">1♂</specimenCount>
,
<collectingCountry id="FE8CCED9845DCC01FE03FA66DD15FA6C" box="[502,631,1490,1516]" name="Costa Rica" pageId="5" pageNumber="320">Costa Rica</collectingCountry>
,
<collectingRegion id="445F40AB845DCC01FD70FA66DC37FA6C" box="[645,853,1490,1516]" country="Costa Rica" name="Alajuela" pageId="5" pageNumber="320">Alajuela Province</collectingRegion>
,
<location id="8344D892845DCC01FC90FA66DCDDFA6C" LSID="urn:lsid:plazi:treatment:0E323F5F845DCC02FF62FEDADEC9FC01:8344D892845DCC01FC90FA66DCDDFA6C" box="[869,959,1490,1516]" country="Costa Rica" name="Bijagua" pageId="5" pageNumber="320" stateProvince="Alajuela">Bijagua</location>
,
<locationDeviation id="9796976B845DCC01FC3AFA66DBE4FA6C" box="[975,1158,1490,1516]" location="Upala" pageId="5" pageNumber="320">
<quantity id="416323AC845DCC01FC3AFA66DB78FA6C" box="[975,1050,1490,1516]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="320" unit="km" value="20.0">20 km</quantity>
S Upala
</locationDeviation>
,
<date id="F225A889845DCC01FB63FA66DB91FA6C" box="[1174,1267,1490,1516]" pageId="5" pageNumber="320" value="1991-01-06">
<collectingDate id="E2615161845DCC01FB63FA66DB91FA6C" box="[1174,1267,1490,1516]" pageId="5" pageNumber="320" value="1991-01-06">6.I.1991</collectingDate>
</date>
(
<collectorName id="2B6EEB9F845DCC01FAF2FA66DAF3FA6C" box="[1287,1425,1490,1516]" pageId="5" pageNumber="320">F.D. Parker</collectorName>
),
<collectionCode id="E08A168C845DCC01FF62FA42DF8FF990" box="[151,237,1526,1552]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="5" pageNumber="320">AMNH</collectionCode>
_
<specimenCode id="D63D2632845DCC01FF0EFA42DEFDF990" box="[251,415,1526,1552]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="5" pageNumber="320">IZC 00390604</specimenCode>
[
<collectionCode id="E08A168C845DCC01FE59FA42DD69F990" box="[428,523,1526,1552]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="5" pageNumber="320">AMNH</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CE81DDC2845DCC02FF32F9AEDF87FD29" lastPageId="6" lastPageNumber="321" pageId="5" pageNumber="320" type="description">
<paragraph id="86248E49845DCC02FF32F9AEDF87FD29" blockId="5.[151,1437,1490,2021]" lastBlockId="6.[151,1437,151,897]" lastPageId="6" lastPageNumber="321" pageId="5" pageNumber="320">
<emphasis id="B4EF525B845DCC01FF32F9AEDED5F9B4" bold="true" box="[199,439,1562,1588]" pageId="5" pageNumber="320">Description of male</emphasis>
(hithertho unkonown).
<emphasis id="B4EF525B845DCC01FD3EF9AEDD99F9B4" bold="true" box="[715,763,1562,1588]" pageId="5" pageNumber="320">Size</emphasis>
. Head
<quantity id="416323AC845DCC01FCA7F9AEDCDBF9B5" box="[850,953,1562,1589]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="5" pageNumber="320" unit="mm" value="1.12">1.12 mm</quantity>
long,
<quantity id="416323AC845DCC01FBF3F9AEDB0FF9B5" box="[1030,1133,1562,1589]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="5" pageNumber="320" unit="mm" value="2.03">2.03 mm</quantity>
high, and
<quantity id="416323AC845DCC01FB18F9AEDA36F9B5" box="[1261,1364,1562,1589]" metricMagnitude="-3" metricUnit="m" metricValue="2.39" pageId="5" pageNumber="320" unit="mm" value="2.39">2.39 mm</quantity>
wide; mesosoma
<quantity id="416323AC845DCC01FEE0F98ADE18F9D9" box="[277,378,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="4.42" pageId="5" pageNumber="320" unit="mm" value="4.42">4.42 mm</quantity>
long, fore wing
<quantity id="416323AC845DCC01FDCDF98ADDECF9D9" box="[568,654,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="5" pageNumber="320" unit="mm" value="7.1">7.1 mm</quantity>
long, and hind wing
<quantity id="416323AC845DCC01FC74F98ADC84F9D9" box="[897,998,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="5.44" pageId="5" pageNumber="320" unit="mm" value="5.44">5.44 mm</quantity>
long; metasoma
<quantity id="416323AC845DCC01FB5DF98ADA6FF9D9" box="[1192,1293,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="6.08" pageId="5" pageNumber="320" unit="mm" value="6.08">6.08 mm</quantity>
long.
<emphasis id="B4EF525B845DCC01FAA0F98ADAF7F9D8" bold="true" box="[1365,1429,1598,1624]" pageId="5" pageNumber="320">Head</emphasis>
. Lateral ocelli
<quantity id="416323AC845DCC01FEC0F9D6DEF9F9FD" box="[309,411,1634,1661]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="5" pageNumber="320" unit="mm" value="0.18">0.18 mm</quantity>
, and median ocellus
<quantity id="416323AC845DCC01FD7FF9D6DD83F9FD" box="[650,737,1634,1661]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="320" unit="mm" value="0.2">0.2 mm</quantity>
diameter. Frons with interantennal space 1.63 times height of antennal socket. Anterior tentorial pit closer to eye than to antennal socket. Antennal socket
<quantity id="416323AC845DCC01FB6DF932DB99F921" box="[1176,1275,1670,1697]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="320" unit="mm" value="0.24">0.24 mm</quantity>
high. Clypeus 1.1 times higher than wide and apex less acute than in female. Pubescence stronger than in female. Gena less than half of width of eye at level of ocular sinus.
<emphasis id="B4EF525B845DCC01FD69F97ADC78F968" bold="true" box="[668,794,1742,1768]" pageId="5" pageNumber="320">Mesosoma</emphasis>
. Propodeum with translucent posterior submedian mark anterior to propodeal valve, not inserted in a round depression. Propodeal valve complete and expanded, median region one-third of height of antennal socket. Fore wing with prestigma as long as wide. Hind wing with six hamuli.
<emphasis id="B4EF525B845DCC01FF62F88EDE71F8D4" bold="true" box="[151,275,1850,1876]" pageId="5" pageNumber="320">Metasoma</emphasis>
. Tergum I 1.8 times longer than broad. Tergum II 0.75 times longer than wide.
<emphasis id="B4EF525B845DCC01FB75F88EDBA6F8D4" bold="true" box="[1152,1220,1850,1876]" pageId="5" pageNumber="320">Color</emphasis>
. Yellow in general. Black: central large spot on vertex, lateral large band across frons, from ocellar area to upper margin of antennal socket, transverse large band anterior to occipital carina, mesoscutum, except two median bands, spot on upper region of mesepisternum, terga IIIVII, except posterior thin bands, and metasomal sterna IVVI, except posterior thin bands. Brown: large band anterior to pronotal carina and dorso-lateral band on pronotum, small spot above dorsal groove and spot posterior to scrobal furrow, tegula, inner region of subalar metapleural area, scutellum, posterior half of metanotum, longitudinal median band and lateral band on propodeum, and terga I and II, except posterior thin band on tergum II. Yellowish brown: antennal segments, except inner region of pedicel. Fore wing with yellowish cells, hind wing with hyaline cells; yellowish-brown venation, except reddish-brown veins costa, subcosta+radius, media+cubitus, media and radial sector.
<emphasis id="B4EF525B845ECC02FCD4FE92DCEDFEC0" bold="true" box="[801,911,294,320]" pageId="6" pageNumber="321">Genitalia</emphasis>
(
<figureCitation id="1EA092CC845ECC02FC6BFE93DB40FEC1" box="[926,1058,295,321]" captionStart="FIGURES 1522" captionStartId="5.[151,264,1312,1336]" captionTargetBox="[190,1397,669,1289]" captionTargetId="figure-171@5.[190,1397,669,1289]" captionTargetPageId="5" captionText="FIGURES 1522. Angiopolybia zischkai Richards, ♂. 15. Habitus, lateral view. 16. Habitus, dorsal view. 17. Head, frontal view. 18. Paramere. 19. Aedeagus, lateral view. 20. Aedeagus, ventral view. 21. Digitus. 22. Cuspis. Scale bar: 1 mm for 1517 and 0.5 mm for 1822." figureDoi="http://doi.org/10.5281/zenodo.14033781" httpUri="https://zenodo.org/record/14033781/files/figure.png" pageId="6" pageNumber="321">Figs 1822</figureCitation>
).
<emphasis id="B4EF525B845ECC02FBC0FE93DBC9FEC1" bold="true" box="[1077,1195,295,321]" pageId="6" pageNumber="321">Paramere</emphasis>
<quantity id="416323AC845ECC02FB46FE92DA6BFEC1" box="[1203,1289,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="321" unit="mm" value="1.5">1.5 mm</quantity>
long and
<quantity id="416323AC845ECC02FA8CFE93DFA1FEE5" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="321" unit="mm" value="0.7">0.7 mm</quantity>
wide; parameral spine one-seventh length of paramere, curved upwards and with long and sparse bristles; lobe with a rounded apex and not curved downwards.
<emphasis id="B4EF525B845ECC02FD4EFEDADC4CFE08" bold="true" box="[699,814,366,392]" pageId="6" pageNumber="321">Aedeagus</emphasis>
<quantity id="416323AC845ECC02FCC0FEDBDCE8FE09" box="[821,906,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="321" unit="mm" value="1.4">1.4 mm</quantity>
long; enlarged valve with a small emargination at tip; apical portion
<quantity id="416323AC845ECC02FE88FE26DE82FE2D" box="[381,480,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="321" unit="mm" value="0.53">0.53 mm</quantity>
long and straight, ventral margin with denticles directed anteriorly; denticulation with large denticles in posterior half and small denticles in anterior half, more sclerotized than the rest of apical portion; small bristles closer in lower half, sparse in upper half; median expansion without denticles and with acute apex; lateral apodeme not flattened dorsoventrally at apex; basal apodeme arched to venter.
<emphasis id="B4EF525B845ECC02FBA0FE4ADBC9FD98" bold="true" box="[1109,1195,510,536]" pageId="6" pageNumber="321">Digitus</emphasis>
1.8 times longer than wide; apical process not curved in upper half and with medium bristles and sparse; rounded anteroventral lobe with strip of black scale-like bristles crossing it obliquely at base of digitus; bristles absent in basal articulation.
<emphasis id="B4EF525B845ECC02FABEFDF2DAFEFDE0" bold="true" box="[1355,1436,582,608]" pageId="6" pageNumber="321">Cuspis</emphasis>
approximately
<quantity id="416323AC845ECC02FEB1FDDEDEC9FD05" box="[324,427,618,645]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="321" unit="mm" value="0.45">0.45 mm</quantity>
long, with six black scale-like bristles on lateral lobe, small dense bristles throughout cuspis.
</paragraph>
</subSubSection>
<subSubSection id="CE81DDC2845ECC02FF32FD06DEC9FC01" pageId="6" pageNumber="321" type="discussion">
<paragraph id="86248E49845ECC02FF32FD06DEC9FC01" blockId="6.[151,1437,151,897]" pageId="6" pageNumber="321">
<emphasis id="B4EF525B845ECC02FF32FD06DE2AFD4C" bold="true" box="[199,328,690,716]" pageId="6" pageNumber="321">Comments</emphasis>
. The described male of
<taxonomicName id="419BF5CA845ECC02FDA7FD07DC2BFD4D" authorityName="Richards" authorityYear="1978" box="[594,841,691,717]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis id="B4EF525B845ECC02FDA7FD07DC2BFD4D" box="[594,841,691,717]" italics="true" pageId="6" pageNumber="321">Angiopolybia zischkai</emphasis>
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is from
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, where the species was previously recorded by
<bibRefCitation id="E20AF3B8845ECC02FED7FD63DD74FD71" author="Valverde, J. P. &amp; Hanson, P. &amp; Carpenter, J." box="[290,534,727,753]" pageId="6" pageNumber="321" pagination="174 - 199" refId="ref8007" refString="Valverde, J. P., Hanson, P. &amp; Carpenter, J. (2019) Keys to the Costa Rican species of paper wasps (Hymenoptera: Vespidae: Polistinae). Revista de Biologia Tropical, 67 (2), 174 - 199. https: // doi. org / 10.15517 / rbt. v 67 i 2 SUPL. 37229" type="journal article" year="2019">
Valverde
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(2019)
</bibRefCitation>
and
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Barroso
<emphasis id="B4EF525B845ECC02FD5EFD63DD81FD71" box="[683,739,727,753]" italics="true" pageId="6" pageNumber="321">et al.</emphasis>
(2022)
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. Despite some differences between males and females, such as the male having a slenderer clypeus with stronger pubescence, a narrower gena, and slightly shorter terga I and II (female redescribed by
<bibRefCitation id="E20AF3B8845ECC02FE1DFCABDDA1FCB9" author="Barroso, P. C. S. &amp; Menezes, R. S. T. &amp; Oliveira, M. L. &amp; Somavilla, A." box="[488,707,799,825]" pageId="6" pageNumber="321" pagination="75 - 97" refId="ref6283" refString="Barroso, P. C. S., Menezes, R. S. T., Oliveira, M. L. &amp; Somavilla, A. (2022) A systematic review of the Neotropical social wasp genus Angiopolybia Araujo, 1946 (Hymenoptera: Vespidae): species delimitation, morphological diagnosis, and geographical distribution. Arthropod Systematics &amp; Phylogeny, 80, 75 - 97. https: // doi. org / 10.3897 / asp. 80. e 71492" type="journal article" year="2022">
Barroso
<emphasis id="B4EF525B845ECC02FDBDFCABDDE2FCB9" box="[584,640,799,825]" italics="true" pageId="6" pageNumber="321">et al.</emphasis>
2022
</bibRefCitation>
), the male specimen was associated with the female based on the collection of females from the same locality, as well as the identification key, redescription and diagnosis provided by
<bibRefCitation id="E20AF3B8845ECC02FF4FFCD3DEC5FC01" author="Barroso, P. C. S. &amp; Menezes, R. S. T. &amp; Oliveira, M. L. &amp; Somavilla, A." box="[186,423,871,897]" pageId="6" pageNumber="321" pagination="75 - 97" refId="ref6283" refString="Barroso, P. C. S., Menezes, R. S. T., Oliveira, M. L. &amp; Somavilla, A. (2022) A systematic review of the Neotropical social wasp genus Angiopolybia Araujo, 1946 (Hymenoptera: Vespidae): species delimitation, morphological diagnosis, and geographical distribution. Arthropod Systematics &amp; Phylogeny, 80, 75 - 97. https: // doi. org / 10.3897 / asp. 80. e 71492" type="journal article" year="2022">
Barroso
<emphasis id="B4EF525B845ECC02FEEFFCD3DE30FC01" box="[282,338,871,897]" italics="true" pageId="6" pageNumber="321">et al.</emphasis>
(2022)
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.
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View file

@ -0,0 +1,211 @@
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Key to the species of
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<emphasis id="B4EF525B845ECC02FE68FC66DD5CFC6D" bold="true" box="[413,574,978,1005]" italics="true" pageId="6" pageNumber="321">Angiopolybia</emphasis>
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This identification key is modified from
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Barroso
<emphasis id="B4EF525B845ECC02FD4AFBAFDD95FBB5" box="[703,759,1051,1077]" italics="true" pageId="6" pageNumber="321">et al.</emphasis>
(2022)
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.
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. Fore wing
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long; eyes with medium bristles, between 0.05 and
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long; lateral lobe of clypeus touching the eye; anterior margin of the axillary fossa directed to the posterior region.............................................
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Fore wing
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long; eyes with tiny bristles, approximately
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long; lateral lobe of clypeus not touching the eye; anterior margin of the axillary fossa directed to the anterior region..............................................
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. Pronotum with lobe developed below the pronotal fovea; metapleural basalar area with parallel upper and lower margins................................................................................
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Pronotum without lobe below the pronotal fovea; metapleural basalar area with divergent upper and lower margins......................................................................................
<taxonomicName id="419BF5CA845ECC02FB6CFA99DAFEFAC2" ID-CoL="66R66" authority="Richards, 1978" authorityName="Richards" authorityYear="1978" box="[1177,1436,1324,1347]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis id="B4EF525B845ECC02FB6CFA99DA67FAC2" bold="true" box="[1177,1285,1324,1346]" italics="true" pageId="6" pageNumber="321">A. zischkai</emphasis>
<bibRefCitation id="E20AF3B8845ECC02FAFEFA99DAFEFAC2" author="Richards, O. W." box="[1291,1436,1325,1347]" pageId="6" pageNumber="321" refId="ref7469" refString="Richards, O. W. (1978) The social wasps of the Americas, excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards, 1978</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D6F96EC845ECC02FF62FAFEDAFEFA37" pageId="6" pageNumber="321">
<paragraph id="86248E49845ECC02FF62FAFEDAFEFAFD" blockId="6.[151,1437,1119,1582]" pageId="6" pageNumber="321">
<keyLead id="3D6A2D7C845ECC02FF62FAFEDAFEFAFD" pageId="6" pageNumber="321">
<emphasis id="B4EF525B845ECC02FF62FAFEDFC1FADF" bold="true" box="[151,163,1354,1375]" pageId="6" pageNumber="321">3</emphasis>
. Pronotum with high lamella along anterior margin, about one-third of height of antennal socket; gena enlarged in the upper region.........................................................................
<taxonomicName id="419BF5CA845ECC02FB61FADCDAFEFAFD" ID-CoL="DZ9Z" authority="(Ducke, 1904)" baseAuthorityName="Ducke" baseAuthorityYear="1904" box="[1172,1436,1383,1406]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="obidensis">
<emphasis id="B4EF525B845ECC02FB61FADCDA6EFAFD" bold="true" box="[1172,1292,1383,1405]" italics="true" pageId="6" pageNumber="321">A. obidensis</emphasis>
(
<bibRefCitation id="E20AF3B8845ECC02FAEDFADCDAF7FAFD" author="Ducke, A." box="[1304,1429,1384,1406]" pageId="6" pageNumber="321" pagination="317 - 374" refId="ref7065" refString="Ducke, A. (1904) Sobre as vespidas sociais do Para. Extracto do Boletim do Museu Goeldi, 4, 317 - 374. https: // doi. org / 10.5962 / bhl. title. 100023" type="journal article" year="1904">Ducke, 1904</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="86248E49845ECC02FF62FA31DAFEFA37" blockId="6.[151,1437,1119,1582]" pageId="6" pageNumber="321">
<keyLead id="3D6A2D7C845ECC02FF62FA31DAFEFA37" pageId="6" pageNumber="321">
<emphasis id="B4EF525B845ECC02FF62FA31DFC1FA1A" bold="true" box="[151,163,1413,1434]" pageId="6" pageNumber="321"></emphasis>
Pronotum with low lamella along anterior margin, about one-fifth of height of antennal socket; gena not enlarged in the upper region..............................................................................................
<emphasis id="B4EF525B845ECC02FA65FA16DAFEFA37" bold="true" box="[1424,1436,1442,1463]" pageId="6" pageNumber="321">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D6F96EC845ECC02FF62FA0BDAFEF9AD" pageId="6" pageNumber="321">
<paragraph id="86248E49845ECC02FF62FA0BDAFEFA73" blockId="6.[151,1437,1119,1582]" pageId="6" pageNumber="321">
<keyLead id="3D6A2D7C845ECC02FF62FA0BDAFEFA73" pageId="6" pageNumber="321">
<emphasis id="B4EF525B845ECC02FF62FA0BDFC1FA54" bold="true" box="[151,163,1471,1492]" pageId="6" pageNumber="321">4</emphasis>
. Tergum II with a row of very long bristles along posterior margin (
<figureCitation id="1EA092CC845ECC02FCBBFA74DCF1FA55" box="[846,915,1472,1494]" captionStart="FIGURES 811. 8, 9" captionStartId="3.[151,264,703,727]" captionTargetBox="[191,1395,181,679]" captionTargetId="figure-19@3.[191,1395,181,679]" captionTargetPageId="3" captionText="FIGURES 811. 8, 9. Angiopolybia obscuriorina sp. nov., holotype, ♀. 10, 11. Stelopolybia paraensis var. (or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 8, 10. Metapleural basalar area, lateral view. 9, 11. Tergum II, lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033777" httpUri="https://zenodo.org/record/14033777/files/figure.png" pageId="6" pageNumber="321">Fig. 11</figureCitation>
); propodeum with region anterior to spiracle projecting more in metapleural basalar area (
<figureCitation id="1EA092CC845ECC02FDDDFA69DD0CFA72" box="[552,622,1501,1523]" captionStart="FIGURES 811. 8, 9" captionStartId="3.[151,264,703,727]" captionTargetBox="[191,1395,181,679]" captionTargetId="figure-19@3.[191,1395,181,679]" captionTargetPageId="3" captionText="FIGURES 811. 8, 9. Angiopolybia obscuriorina sp. nov., holotype, ♀. 10, 11. Stelopolybia paraensis var. (or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 8, 10. Metapleural basalar area, lateral view. 9, 11. Tergum II, lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033777" httpUri="https://zenodo.org/record/14033777/files/figure.png" pageId="6" pageNumber="321">Fig. 10</figureCitation>
)...........................................
<taxonomicName id="419BF5CA845ECC02FB73FA69DAFEFA73" authority="(Spinola, 1851)" baseAuthorityName="Spinola" baseAuthorityYear="1851" box="[1158,1436,1501,1523]" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="paraensis">
<emphasis id="B4EF525B845ECC02FB73FA69DA63FA72" bold="true" box="[1158,1281,1501,1522]" italics="true" pageId="6" pageNumber="321">A. paraensis</emphasis>
(
<bibRefCitation id="E20AF3B8845ECC02FAFBFA69DAF7FA73" author="Spinola, M." box="[1294,1429,1501,1523]" pageId="6" pageNumber="321" pagination="19 - 94" refId="ref7965" refString="Spinola, M. (1851) Compte rendu des Hymenopteres inedits: provenants du voyage entomologique de M. Ghiliani dans le Para em 1846. Memorie della Reale Accademia delle Scienze di Torino, 13, 19 - 94." type="journal article" year="1851">Spinola, 1851</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="86248E49845ECC02FF62FA4EDAFEF9AD" blockId="6.[151,1437,1119,1582]" pageId="6" pageNumber="321">
<keyLead id="3D6A2D7C845ECC02FF62FA4EDAFEF9AD" pageId="6" pageNumber="321">
<emphasis id="B4EF525B845ECC02FF62FA4EDFC1F98F" bold="true" box="[151,163,1530,1551]" pageId="6" pageNumber="321"></emphasis>
Tergum II without a row of very long bristles along the posterior margin (
<figureCitation id="1EA092CC845ECC02FC41FA4EDC92F98F" box="[948,1008,1530,1552]" captionStart="FIGURES 811. 8, 9" captionStartId="3.[151,264,703,727]" captionTargetBox="[191,1395,181,679]" captionTargetId="figure-19@3.[191,1395,181,679]" captionTargetPageId="3" captionText="FIGURES 811. 8, 9. Angiopolybia obscuriorina sp. nov., holotype, ♀. 10, 11. Stelopolybia paraensis var. (or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 8, 10. Metapleural basalar area, lateral view. 9, 11. Tergum II, lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033777" httpUri="https://zenodo.org/record/14033777/files/figure.png" pageId="6" pageNumber="321">Fig. 9</figureCitation>
); propodeum with the region anterior to the spiracle projecting less in metapleural basalar area (
<figureCitation id="1EA092CC845ECC02FD26F9ACDC6EF9AD" box="[723,780,1560,1582]" captionStart="FIGURES 811. 8, 9" captionStartId="3.[151,264,703,727]" captionTargetBox="[191,1395,181,679]" captionTargetId="figure-19@3.[191,1395,181,679]" captionTargetPageId="3" captionText="FIGURES 811. 8, 9. Angiopolybia obscuriorina sp. nov., holotype, ♀. 10, 11. Stelopolybia paraensis var. (or subspecies) obscurior Bequaert, holotype, ♀ (current A. paraensis brown). 8, 10. Metapleural basalar area, lateral view. 9, 11. Tergum II, lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033777" httpUri="https://zenodo.org/record/14033777/files/figure.png" pageId="6" pageNumber="321">Fig. 8</figureCitation>
)..........................
<taxonomicName id="419BF5CA845ECC02FBA2F9ACDA24F9AD" authority="Barroso" authorityName="Barroso &amp; Somavilla &amp; Broad &amp; Menezes &amp; Oliveira" authorityYear="2024" box="[1111,1350,1559,1581]" class="Insecta" family="Vespidae" genus="Angiopolybia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="obscuriorina" status="sp. nov.">
<emphasis id="B4EF525B845ECC02FBA2F9ACDB90F9AD" bold="true" box="[1111,1266,1559,1581]" italics="true" pageId="6" pageNumber="321">A. obscuriorina</emphasis>
Barroso
</taxonomicName>
,
<emphasis id="B4EF525B845ECC02FAA5F9ACDAFEF9AD" bold="true" box="[1360,1436,1560,1581]" pageId="6" pageNumber="321">
<taxonomicNameLabel id="AFDCEF20845ECC02FAA5F9ACDAFEF9AD" box="[1360,1436,1560,1581]" pageId="6" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,372 @@
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<mods:title id="DC51BE49CDFD5A06F2E8E1A84102A4B8">Psammobiont halacarid mites (Acari: Halacaridae) from Türkiye: description of two new species of the genus Atelopsalis Trouessart, 1896 and Scaptognathus Trouessart, 1889 and two new records</mods:title>
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<paragraph id="AD5D36F15002FF83FF1C4CA4FDEAFA60" blockId="10.[151,1184,1482,1581]" box="[151,589,1482,1509]" pageId="10" pageNumber="271">
<heading id="F615819D5002FF83FF1C4CA4FDEAFA60" bold="true" box="[151,589,1482,1509]" fontSize="11" level="1" pageId="10" pageNumber="271" reason="1">
<emphasis id="9F96EAE35002FF83FF1C4CA4FDEAFA60" bold="true" box="[151,589,1482,1509]" pageId="10" pageNumber="271">
<taxonomicName id="6AE24D725002FF83FF1C4CA4FE4FFA60" authorityName="Durucan" authorityYear="2024" box="[151,488,1482,1509]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="antalyaensis" status="sp. nov.">
<emphasis id="9F96EAE35002FF83FF1C4CA4FE4FFA60" bold="true" box="[151,488,1482,1509]" italics="true" pageId="10" pageNumber="271">Scaptognathus antalyaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84A557985002FF83FE644CA5FDEAFA60" box="[495,589,1483,1509]" pageId="10" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<emphasis id="9F96EAE35002FF83FF1C4C81FED5F98C" bold="true" box="[151,370,1519,1545]" pageId="10" pageNumber="271">Zoobank number:</emphasis>
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urn:lsid:zoobank.org:act:
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<paragraph id="AD5D36F15002FF83FF1C4F7CFE39F9A9" blockId="10.[151,1184,1482,1581]" box="[151,414,1554,1581]" pageId="10" pageNumber="271">
<emphasis id="9F96EAE35002FF83FF1C4F7CFE39F9A9" bold="true" box="[151,414,1554,1581]" pageId="10" pageNumber="271">
<figureCitation id="35D92A745002FF83FF1C4F7CFEA0F9A9" box="[151,263,1554,1580]" captionStart="FIGURE 6" captionStartId="12.[152,255,1856,1880]" captionTargetBox="[213,1373,184,1826]" captionTargetId="figure-23@12.[209,1378,179,1830]" captionTargetPageId="12" captionText="FIGURE 6. Scaptognathus antalyaensis sp. nov.: Line art illustrations of female (AE) (FDHAL-24/106) and male (FI) (FDHAL-24/107): A Idiosoma, dorsal view B Idiosoma, ventral view C Genitoanal plate, detailed D Gnathosoma, dorsal view E Right palp, dorsal view F Idiosoma, dorsal view G Idiosoma, ventral view H Genitoanal plate, detailed I Spermatopositor (S. antalyaensis sp. nov.) J Spermatopositor (S. hallezi) K Spermatopositor (S. sabularius). Scale bars: A, B, D, F, G=100 µm; C, H=50 µm; E= 50 µm; IK= 25 µm." figureDoi="http://doi.org/10.5281/zenodo.14033584" httpUri="https://zenodo.org/record/14033584/files/figure.png" pageId="10" pageNumber="271">Figures 6</figureCitation>
,
<figureCitation id="35D92A745002FF83FE9F4F7DFE85F9A8" box="[276,290,1555,1581]" captionStart="FIGURE 7" captionStartId="13.[152,255,1429,1453]" captionTargetBox="[151,1435,179,1404]" captionTargetId="figure-16@13.[151,1435,179,1404]" captionTargetPageId="13" captionText="FIGURE 7. Scaptognathus antalyaensis sp. nov.: Line art illustrations of female (FDHAL-24/106) A Leg I, medial view B Leg II, ventral view C Leg III, ventral view D Leg IV, lateral view. Scale bars: AD=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14033588" httpUri="https://zenodo.org/record/14033588/files/figure.png" pageId="10" pageNumber="271">7</figureCitation>
&amp;
<tableCitation id="E060034A5002FF83FECC4F7DFE39F9A9" box="[327,414,1554,1581]" captionStart="TABLE 2" captionStartId="11.[152,244,188,212]" captionText="TABLE 2. Measurements for Scaptognathus antalyaensis sp. nov. (in micrometers)." pageId="10" pageNumber="271">Table 2</tableCitation>
</emphasis>
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<paragraph id="AD5D36F15002FF83FF1C4F34FE01F938" blockId="10.[151,1437,1626,2013]" pageId="10" pageNumber="271">
<emphasis id="9F96EAE35002FF83FF1C4F34FED8F9F1" bold="true" box="[151,383,1626,1652]" pageId="10" pageNumber="271">Material examined.</emphasis>
<materialsCitation id="1D8A3CAC5002FF83FE0E4F35FE05F938" collectingDate="2018-07-03" collectionCode="FDHAL" elevation="3" latitude="36.84936" location="Kundu" longLatPrecision="1" longitude="30.839" municipality="All" pageId="10" pageNumber="271" specimenCount="2" stateProvince="Antalya" typeStatus="holotype">
<collectingMunicipality id="4D39AC8B5002FF83FE0E4F35FE0EF9F0" box="[389,425,1627,1653]" pageId="10" pageNumber="271">All</collectingMunicipality>
individuals obtained at St.3 (
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,
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), well sorted soft sand,
<elevation id="26CFD1C25002FF83FB2F4F35FAB3F9F0" box="[1188,1300,1627,1653]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="271" unit="m" value="3.0">
<quantity id="6A1A9B145002FF83FB2F4F35FB69F9F0" box="[1188,1230,1627,1653]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="271" unit="m" value="3.0">3 m</quantity>
depth
</elevation>
,
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,
<geoCoordinate id="C8D650365002FF83FF1C4F11FEB6F91C" box="[151,273,1663,1689]" degrees="30.839000" direction="east" orientation="longitude" pageId="10" pageNumber="271" precision="1" value="30.839">30.839000</geoCoordinate>
,
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<collectingDate id="C918E9D95002FF83FE944F11FE0DF91C" box="[287,426,1663,1689]" pageId="10" pageNumber="271" value="2018-07-03">3 July 2018</collectingDate>
</date>
, dissected and mounted into one slide:
<emphasis id="9F96EAE35002FF83FCF94F11FC4FF91C" bold="true" box="[882,1000,1663,1689]" pageId="10" pageNumber="271">
<typeStatus id="725988535002FF83FCF94F11FC44F91C" box="[882,995,1663,1689]" pageId="10" pageNumber="271" type="holotype">Holotype</typeStatus>
:
</emphasis>
Female (FDHAL-24/106);
<emphasis id="9F96EAE35002FF83FAAD4F11FA3BF91C" bold="true" box="[1318,1436,1663,1689]" pageId="10" pageNumber="271">
<typeStatus id="725988535002FF83FAAD4F11FA30F91C" box="[1318,1431,1663,1689]" pageId="10" pageNumber="271" type="paratype">Paratype</typeStatus>
:
</emphasis>
Male (
<collectionCode id="CBF3AE345002FF83FF544FCDFE99F938" box="[223,318,1699,1725]" pageId="10" pageNumber="271">FDHAL</collectionCode>
<emphasis id="9F96EAE35002FF83FEB54FCDFEE0F938" bold="true" box="[318,327,1699,1725]" pageId="10" pageNumber="271">-</emphasis>
24/107)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5F8657A5002FF83FF4C4FA9FCDFF964" box="[199,888,1735,1761]" pageId="10" pageNumber="271" type="etymology">
<paragraph id="AD5D36F15002FF83FF4C4FA9FCDFF964" blockId="10.[151,1437,1626,2013]" box="[199,888,1735,1761]" pageId="10" pageNumber="271">
<emphasis id="9F96EAE35002FF83FF4C4FA9FEE9F964" bold="true" box="[199,334,1735,1761]" pageId="10" pageNumber="271">Etymology.</emphasis>
The species is named for its presence in
<collectingRegion id="6F26F8135002FF83FC934FA9FCD3F964" box="[792,884,1735,1761]" country="Turkey" name="Antalya" pageId="10" pageNumber="271">Antalya</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="E5F8657A5002FF82FF4C4F84FE60F98B" lastPageId="11" lastPageNumber="272" pageId="10" pageNumber="271" type="description">
<paragraph id="AD5D36F15002FF83FF4C4F84FD55F880" blockId="10.[151,1437,1626,2013]" box="[199,754,1770,1797]" pageId="10" pageNumber="271">
<emphasis id="9F96EAE35002FF83FF4C4F84FEFEF881" bold="true" box="[199,345,1770,1796]" pageId="10" pageNumber="271">Description.</emphasis>
Measurements presented in
<tableCitation id="E060034A5002FF83FD134F85FD49F880" box="[664,750,1771,1797]" captionStart="TABLE 2" captionStartId="11.[152,244,188,212]" captionText="TABLE 2. Measurements for Scaptognathus antalyaensis sp. nov. (in micrometers)." pageId="10" pageNumber="271">Table 2</tableCitation>
.
</paragraph>
<paragraph id="AD5D36F15002FF83FF4C4E60FAF8F858" blockId="10.[151,1437,1626,2013]" pageId="10" pageNumber="271">
<emphasis id="9F96EAE35002FF83FF4C4E60FE01F8AD" bold="true" box="[199,422,1806,1833]" pageId="10" pageNumber="271">
<emphasis id="9F96EAE35002FF83FF4C4E60FEB9F8AC" bold="true" box="[199,286,1806,1833]" italics="true" pageId="10" pageNumber="271">Female</emphasis>
(
<typeStatus id="725988535002FF83FEA64E60FE3EF8AD" box="[301,409,1806,1832]" pageId="10" pageNumber="271" type="holotype">holotype</typeStatus>
).
</emphasis>
Dorsal plates irregularly shaped pits arranged conspicuously within polygons. AD and PD large. Dorsum with seven pairs of dorsal setae. All dorsal and ventral setae are short and almost the same length. Glp-1 on AD. Pairs of ds-1 and glp-1 at the same level medial to gland pores. Ds-2 on AD. Pairs of ds-3 to ds-
<quantity id="6A1A9B145002FF83FA044E38FF0AF810" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="271" unit="in" value="5.0">5 in</quantity>
membranous integument. Ds-6 and ds-7 on PD. Anterior margin of AD truncate, posteriorly convex. OC small, wider than long with large central pore and few minute pits. Cornea not seen. Anterior margin of PD truncate, lateral margins parallel. No setae in posterior margin of PD or on anal plate. Pair of glp-
<quantity id="6A1A9B145002FF83FBAE4EADFBE9F858" box="[1061,1102,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="271" unit="in" value="3.0">3 in</quantity>
posterior margin of PD.
</paragraph>
<caption id="F99D66795003FF82FF1349D2FB84FF51" box="[152,1059,188,213]" pageId="11" pageNumber="272" startId="11.[152,244,188,212]" targetType="table">
<paragraph id="AD5D36F15003FF82FF1349D2FB84FF51" blockId="11.[152,1059,188,213]" box="[152,1059,188,213]" pageId="11" pageNumber="272">
<emphasis id="9F96EAE35003FF82FF1349D2FEA8FF51" bold="true" box="[152,271,188,212]" pageId="11" pageNumber="272">TABLE 2.</emphasis>
Measurements for
<taxonomicName id="6AE24D725003FF82FE5549D2FCA1FF51" authorityName="Durucan" authorityYear="2024" box="[478,774,188,212]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="antalyaensis" status="sp. nov.">
<emphasis id="9F96EAE35003FF82FE5549D2FCA1FF51" box="[478,774,188,212]" italics="true" pageId="11" pageNumber="272">Scaptognathus antalyaensis</emphasis>
</taxonomicName>
<emphasis id="9F96EAE35003FF82FC8649D3FCC4FF50" bold="true" box="[781,867,189,213]" pageId="11" pageNumber="272">
<taxonomicNameLabel id="84A557985003FF82FC8649D3FCC4FF50" box="[781,867,189,213]" pageId="11" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(in micrometers).
</paragraph>
</caption>
<paragraph id="AD5D36F15003FF82FF364997FB5EFD2E" pageId="11" pageNumber="272">
<table id="DFE2C45150030076FF36498BFABAFD2E" box="[189,1309,229,683]" gridcols="5" gridrows="11" pageId="11" pageNumber="272">
<tr id="13D234B350030076FF36498BFABAFEA7" box="[189,1309,229,290]" gridrow="0" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF36498BFDBBFEA7" box="[189,540,229,290]" gridcol="0" gridrow="0" pageId="11" pageNumber="272">
<taxonomicName id="6AE24D725003FF82FF364997FE6FFE8B" authorityName="Durucan" authorityYear="2024" box="[189,456,249,270]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="antalyaensis" status="sp. nov.">
<emphasis id="9F96EAE35003FF82FF364997FE6FFE8B" box="[189,456,249,270]" italics="true" pageId="11" pageNumber="272">Scaptognathus antalyaensis</emphasis>
</taxonomicName>
<emphasis id="9F96EAE35003FF82FE454997FDBBFE8B" bold="true" box="[462,540,249,270]" pageId="11" pageNumber="272">
<taxonomicNameLabel id="84A557985003FF82FE454997FDBBFE8B" box="[462,540,249,270]" pageId="11" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="50035DCF50030076FDC2498BFD33FEA7" box="[585,660,229,290]" gridcol="1" gridrow="0" pageId="11" pageNumber="272">
<emphasis id="9F96EAE35003FF82FDC2498BFD33FF7F" bold="true" box="[585,660,229,250]" pageId="11" pageNumber="272">Female</emphasis>
Length
</th>
<th id="50035DCF50030076FCAE498BFCC6FEA7" box="[805,865,229,290]" gridcol="2" gridrow="0" pageId="11" pageNumber="272">Width</th>
<th id="50035DCF50030076FB8D498BFBEDFEA7" box="[1030,1098,229,290]" gridcol="3" gridrow="0" pageId="11" pageNumber="272">
<emphasis id="9F96EAE35003FF82FB8D498BFB9DFF7F" bold="true" box="[1030,1082,229,250]" pageId="11" pageNumber="272">Male</emphasis>
Length
</th>
<th id="50035DCF50030076FB6A498BFABAFEA7" box="[1249,1309,229,290]" gridcol="4" gridrow="0" pageId="11" pageNumber="272">Width</th>
</tr>
<tr id="13D234B350030076FF36485AFABAFECF" box="[189,1309,308,330]" gridrow="1" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF36485AFDBBFECF" box="[189,540,308,330]" gridcol="0" gridrow="1" pageId="11" pageNumber="272">Idiosoma</th>
<td id="50035DCF50030076FDC2485AFD33FECF" box="[585,660,308,330]" gridcol="1" gridrow="1" pageId="11" pageNumber="272">325</td>
<td id="50035DCF50030076FCAE485AFCC6FECF" box="[805,865,308,330]" gridcol="2" gridrow="1" pageId="11" pageNumber="272">200</td>
<td id="50035DCF50030076FB8D485AFBEDFECF" box="[1030,1098,308,330]" gridcol="3" gridrow="1" pageId="11" pageNumber="272">300</td>
<td id="50035DCF50030076FB6A485AFABAFECF" box="[1249,1309,308,330]" gridcol="4" gridrow="1" pageId="11" pageNumber="272">200</td>
</tr>
<tr id="13D234B350030076FF364835FABAFEF4" box="[189,1309,347,369]" gridrow="2" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF364835FDBBFEF4" box="[189,540,347,369]" gridcol="0" gridrow="2" pageId="11" pageNumber="272">Gnathosoma</th>
<td id="50035DCF50030076FDC24835FD33FEF4" box="[585,660,347,369]" gridcol="1" gridrow="2" pageId="11" pageNumber="272">236</td>
<td id="50035DCF50030076FCAE4835FCC6FEF4" box="[805,865,347,369]" gridcol="2" gridrow="2" pageId="11" pageNumber="272">136</td>
<td id="50035DCF50030076FB8D4835FBEDFEF4" box="[1030,1098,347,369]" gridcol="3" gridrow="2" pageId="11" pageNumber="272">211</td>
<td id="50035DCF50030076FB6A4835FABAFEF4" box="[1249,1309,347,369]" gridcol="4" gridrow="2" pageId="11" pageNumber="272">120</td>
</tr>
<tr id="13D234B350030076FF3648ECFABAFE1D" box="[189,1309,386,408]" gridrow="3" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF3648ECFDBBFE1D" box="[189,540,386,408]" gridcol="0" gridrow="3" pageId="11" pageNumber="272">Ro</th>
<td id="50035DCF50030076FDC248ECFD33FE1D" box="[585,660,386,408]" gridcol="1" gridrow="3" pageId="11" pageNumber="272">110</td>
<td id="50035DCF50030076FCAE48ECFCC6FE1D" box="[805,865,386,408]" gridcol="2" gridrow="3" pageId="11" pageNumber="272">25</td>
<td id="50035DCF50030076FB8D48ECFBEDFE1D" box="[1030,1098,386,408]" gridcol="3" gridrow="3" pageId="11" pageNumber="272">110</td>
<td id="50035DCF50030076FB6A48ECFABAFE1D" box="[1249,1309,386,408]" gridcol="4" gridrow="3" pageId="11" pageNumber="272">23</td>
</tr>
<tr id="13D234B350030076FF3648C4FABAFE45" box="[189,1309,426,448]" gridrow="4" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF3648C4FDBBFE45" box="[189,540,426,448]" gridcol="0" gridrow="4" pageId="11" pageNumber="272">AD</th>
<td id="50035DCF50030076FDC248C4FD33FE45" box="[585,660,426,448]" gridcol="1" gridrow="4" pageId="11" pageNumber="272">140</td>
<td id="50035DCF50030076FCAE48C4FCC6FE45" box="[805,865,426,448]" gridcol="2" gridrow="4" pageId="11" pageNumber="272">135</td>
<td id="50035DCF50030076FB8D48C4FBEDFE45" box="[1030,1098,426,448]" gridcol="3" gridrow="4" pageId="11" pageNumber="272">125</td>
<td id="50035DCF50030076FB6A48C4FABAFE45" box="[1249,1309,426,448]" gridcol="4" gridrow="4" pageId="11" pageNumber="272">137</td>
</tr>
<tr id="13D234B350030076FF3648BFFABAFE62" box="[189,1309,465,487]" gridrow="5" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF3648BFFDBBFE62" box="[189,540,465,487]" gridcol="0" gridrow="5" pageId="11" pageNumber="272">OC</th>
<td id="50035DCF50030076FDC248BFFD33FE62" box="[585,660,465,487]" gridcol="1" gridrow="5" pageId="11" pageNumber="272">14</td>
<td id="50035DCF50030076FCAE48BFFCC6FE62" box="[805,865,465,487]" gridcol="2" gridrow="5" pageId="11" pageNumber="272">12</td>
<td id="50035DCF50030076FB8D48BFFBEDFE62" box="[1030,1098,465,487]" gridcol="3" gridrow="5" pageId="11" pageNumber="272">17</td>
<td id="50035DCF50030076FB6A48BFFABAFE62" box="[1249,1309,465,487]" gridcol="4" gridrow="5" pageId="11" pageNumber="272">15</td>
</tr>
<tr id="13D234B350030076FF364897FABAFD8B" box="[189,1309,505,526]" gridrow="6" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF364897FDBBFD8B" box="[189,540,505,526]" gridcol="0" gridrow="6" pageId="11" pageNumber="272">PD</th>
<td id="50035DCF50030076FDC24897FD33FD8B" box="[585,660,505,526]" gridcol="1" gridrow="6" pageId="11" pageNumber="272">138</td>
<td id="50035DCF50030076FCAE4897FCC6FD8B" box="[805,865,505,526]" gridcol="2" gridrow="6" pageId="11" pageNumber="272">90</td>
<td id="50035DCF50030076FB8D4897FBEDFD8B" box="[1030,1098,505,526]" gridcol="3" gridrow="6" pageId="11" pageNumber="272">137</td>
<td id="50035DCF50030076FB6A4897FABAFD8B" box="[1249,1309,505,526]" gridcol="4" gridrow="6" pageId="11" pageNumber="272">100</td>
</tr>
<tr id="13D234B350030076FF364B4EFABAFDB3" box="[189,1309,544,566]" gridrow="7" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF364B4EFDBBFDB3" box="[189,540,544,566]" gridcol="0" gridrow="7" pageId="11" pageNumber="272">AE</th>
<td id="50035DCF50030076FDC24B4EFD33FDB3" box="[585,660,544,566]" gridcol="1" gridrow="7" pageId="11" pageNumber="272">135</td>
<td id="50035DCF50030076FCAE4B4EFCC6FDB3" box="[805,865,544,566]" gridcol="2" gridrow="7" pageId="11" pageNumber="272">175</td>
<td id="50035DCF50030076FB8D4B4EFBEDFDB3" box="[1030,1098,544,566]" gridcol="3" gridrow="7" pageId="11" pageNumber="272">112</td>
<td id="50035DCF50030076FB6A4B4EFABAFDB3" box="[1249,1309,544,566]" gridcol="4" gridrow="7" pageId="11" pageNumber="272">182</td>
</tr>
<tr id="13D234B350030076FF364B29FABAFDD8" box="[189,1309,583,605]" gridrow="8" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF364B29FDBBFDD8" box="[189,540,583,605]" gridcol="0" gridrow="8" pageId="11" pageNumber="272">PE</th>
<td id="50035DCF50030076FDC24B29FD33FDD8" box="[585,660,583,605]" gridcol="1" gridrow="8" pageId="11" pageNumber="272">150</td>
<td id="50035DCF50030076FCAE4B29FCC6FDD8" box="[805,865,583,605]" gridcol="2" gridrow="8" pageId="11" pageNumber="272">80</td>
<td id="50035DCF50030076FB8D4B29FBEDFDD8" box="[1030,1098,583,605]" gridcol="3" gridrow="8" pageId="11" pageNumber="272">152</td>
<td id="50035DCF50030076FB6A4B29FABAFDD8" box="[1249,1309,583,605]" gridcol="4" gridrow="8" pageId="11" pageNumber="272">63</td>
</tr>
<tr id="13D234B350030076FF364B00FABAFD01" box="[189,1309,622,644]" gridrow="9" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF364B00FDBBFD01" box="[189,540,622,644]" gridcol="0" gridrow="9" pageId="11" pageNumber="272">GA</th>
<td id="50035DCF50030076FDC24B00FD33FD01" box="[585,660,622,644]" gridcol="1" gridrow="9" pageId="11" pageNumber="272">105</td>
<td id="50035DCF50030076FCAE4B00FCC6FD01" box="[805,865,622,644]" gridcol="2" gridrow="9" pageId="11" pageNumber="272">80</td>
<td id="50035DCF50030076FB8D4B00FBEDFD01" box="[1030,1098,622,644]" gridcol="3" gridrow="9" pageId="11" pageNumber="272">137</td>
<td id="50035DCF50030076FB6A4B00FABAFD01" box="[1249,1309,622,644]" gridcol="4" gridrow="9" pageId="11" pageNumber="272">82</td>
</tr>
<tr id="13D234B350030076FF364BF8FABAFD2E" box="[189,1309,662,683]" gridrow="10" pageId="11" pageNumber="272">
<th id="50035DCF50030076FF364BF8FDBBFD2E" box="[189,540,662,683]" gridcol="0" gridrow="10" pageId="11" pageNumber="272">GO</th>
<td id="50035DCF50030076FDC24BF8FD33FD2E" box="[585,660,662,683]" gridcol="1" gridrow="10" pageId="11" pageNumber="272">38</td>
<td id="50035DCF50030076FCAE4BF8FCC6FD2E" box="[805,865,662,683]" gridcol="2" gridrow="10" pageId="11" pageNumber="272">21</td>
<td id="50035DCF50030076FB8D4BF8FBEDFD2E" box="[1030,1098,662,683]" gridcol="3" gridrow="10" pageId="11" pageNumber="272">40</td>
<td id="50035DCF50030076FB6A4BF8FABAFD2E" box="[1249,1309,662,683]" gridcol="4" gridrow="10" pageId="11" pageNumber="272">24</td>
</tr>
</table>
</paragraph>
<paragraph id="AD5D36F15003FF82FF4C4A6EFB18FC2F" blockId="11.[151,1437,767,1946]" pageId="11" pageNumber="272">
AE with a pair of epimeral pores and three pairs of setae. Posterior margin of AE convex. Ornamentation of ventral plates same as that of the dorsal plate. AE with 8 pairs of circular markings which are muscle strings. PE with one lateral seta and three ventral setae. PE extending beyond insertion of leg IV. Female GA bipartite, both parts almost equal in length.
<emphasis id="9F96EAE35003FF82FE514A02FD21FC00" box="[474,646,875,901]" italics="true" pageId="11" pageNumber="272">Pars sclerosum</emphasis>
with ornamentation while
<emphasis id="9F96EAE35003FF82FC384A02FB37FC00" box="[947,1168,875,901]" italics="true" pageId="11" pageNumber="272">pars membranosum</emphasis>
without ornamentation. Posterior part of GA is reticulated with four pairs of pgs; genital sclerites with two pairs of sgs.
</paragraph>
<paragraph id="AD5D36F15003FF82FF4C4ADDFCE4FBBC" blockId="11.[151,1437,767,1946]" pageId="11" pageNumber="272">Gnathosoma 0.7 of length of idiosoma. Dorsal gnathosomal base with same ornamentation of idiosomal plates. Rostrum shorter than gnathosomal base. Pharyngeal field small. Rostrum spatula-shaped, apically truncated. Both pairs of maxillary setae situated in anterior postion of the rostrum, truncate apex with two pairs of rostral setae. Palps attached dorsally and two segmented. P3 and P4 fused.</paragraph>
<paragraph id="AD5D36F15003FF82FF4C4D2AFA29FADF" blockId="11.[151,1437,767,1946]" pageId="11" pageNumber="272">
Legs (
<figureCitation id="35D92A745003FF82FE9B4D2DFE3FFBDB" box="[272,408,1091,1118]" captionStart="FIGURE 7" captionStartId="13.[152,255,1429,1453]" captionTargetBox="[151,1435,179,1404]" captionTargetId="figure-16@13.[151,1435,179,1404]" captionTargetPageId="13" captionText="FIGURE 7. Scaptognathus antalyaensis sp. nov.: Line art illustrations of female (FDHAL-24/106) A Leg I, medial view B Leg II, ventral view C Leg III, ventral view D Leg IV, lateral view. Scale bars: AD=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14033588" httpUri="https://zenodo.org/record/14033588/files/figure.png" pageId="11" pageNumber="272">Figs. 7AD</figureCitation>
) shorter than idiosoma. Telofemur I longer than tibia I. Telofemora slender, their length: height ratio 3.9, 3.7, 2.4 and 2.3 from leg I to IV, respectively.All telofemora with coarse reticulation. Leg chaetotaxy (
<figureCitation id="35D92A745003FF82FF144DE5FE83FB23" box="[159,292,1163,1190]" captionStart="FIGURE 7" captionStartId="13.[152,255,1429,1453]" captionTargetBox="[151,1435,179,1404]" captionTargetId="figure-16@13.[151,1435,179,1404]" captionTargetPageId="13" captionText="FIGURE 7. Scaptognathus antalyaensis sp. nov.: Line art illustrations of female (FDHAL-24/106) A Leg I, medial view B Leg II, ventral view C Leg III, ventral view D Leg IV, lateral view. Scale bars: AD=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14033588" httpUri="https://zenodo.org/record/14033588/files/figure.png" pageId="11" pageNumber="272">Figs. 7AD</figureCitation>
) (from trochanter to tarsus) (solenidia, famuli and pas excluded), leg I, 1, 2, 5, 4, 9, 4; leg II, 1, 2, 4, 4, 7, 4; leg III, 1, 2, 3, 3, 78, 3; leg IV, 1, 1, 2, 3, 7, 3. Telofemur I with one ventral and one ventrolateral bipectinate seta. Genu to tarsus I with 2, 6, 1 bipectinate ventral setae, genu to tarsus II with 1, 4, 1 and genua to tarsi III and IV with 0, 4, 0 such setae each. Tarsi I to IV with 2, 1, 0, 0 ventral setae; on tarsi I and II one of ventral setae bipectinate. Solenidion of tarsus I is almost 6 long, setiform, and dorsolateral position. Solenidion on tarsus II in dorsomedial position. Tip of tarsi with pas singlets. Paired claws are weakly sclerotized, with accessory process but no pecten.
</paragraph>
<paragraph id="AD5D36F15003FF82FF4C4C0DFE60F98B" blockId="11.[151,1437,767,1946]" pageId="11" pageNumber="272">
<emphasis id="9F96EAE35003FF82FF4C4C0DFEAEFAFB" bold="true" box="[199,265,1379,1406]" pageId="11" pageNumber="272">
<emphasis id="9F96EAE35003FF82FF4C4C0DFEA5FAFB" bold="true" box="[199,258,1379,1406]" italics="true" pageId="11" pageNumber="272">Male</emphasis>
.
</emphasis>
Male is similar to the female except for the genital plate. GA oval, not bipartite, consisting only
<emphasis id="9F96EAE35003FF82FAE14C0AFEAFFA24" italics="true" pageId="11" pageNumber="272">pars sclerosum</emphasis>
and extending to about the level of middle angles of PE. GA furnished panels surrounding genital foramen. Male with 55 setae surrounding of genital foramen. Subgenital setae short, three pairs on genital sclerites. Spermatopositor large, reaching anteriorly to the level of insertions of leg IV. One pair of outlying setae at 0.44 on GA. Adanal setae not seen.
</paragraph>
</subSubSection>
<subSubSection id="E5F8657A5003FF82FF4C4F76FEA2F81F" pageId="11" pageNumber="272" type="discussion">
<paragraph id="AD5D36F15003FF82FF4C4F76FEA2F81F" blockId="11.[151,1437,767,1946]" pageId="11" pageNumber="272">
<emphasis id="9F96EAE35003FF82FF4C4F76FE9CF9B7" bold="true" box="[199,315,1560,1586]" pageId="11" pageNumber="272">Remarks.</emphasis>
<taxonomicName id="6AE24D725003FF82FECD4F79FD25F9B4" authorityName="Durucan" authorityYear="2024" box="[326,642,1559,1585]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="antalyaensis" status="sp. nov.">
<emphasis id="9F96EAE35003FF82FECD4F79FD25F9B4" box="[326,642,1559,1585]" italics="true" pageId="11" pageNumber="272">Scaptognathus antalyaensis</emphasis>
</taxonomicName>
<emphasis id="9F96EAE35003FF82FD064F76FD4DF9B7" bold="true" box="[653,746,1560,1586]" pageId="11" pageNumber="272">
<taxonomicNameLabel id="84A557985003FF82FD064F76FD4DF9B7" box="[653,746,1560,1586]" pageId="11" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is characterized by the combination of: dorsum with short setae, all plates reticulate sculpturing, posterior margin of AD and AE convex, ds-2 on AD, female GA bipartite with four pairs of pgs, male GA uniform, claws with accessory process, number of bipectinate seta on tarsi IIV 1, 1, 0, 0; tibiae 6, 4, 4, 4; genua IIV 1, 1, 0, 0; telofemora IIV 2, 0, 0, 0. The new species has affinities to other previously known Mediterranean species (
<taxonomicName id="6AE24D725003FF82FD304FC6FC4CF947" authority="Trouessart, 1894" authorityName="Trouessart" authorityYear="1894" box="[699,1003,1703,1730]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="hallezi">
<emphasis id="9F96EAE35003FF82FD304FC6FC84F944" box="[699,803,1703,1730]" italics="true" pageId="11" pageNumber="272">S. hallezi</emphasis>
<bibRefCitation id="C9734B005003FF82FCA74FC6FC4CF947" author="Trouessart, E." box="[812,1003,1704,1730]" pageId="11" pageNumber="272" pagination="139 - 175" refId="ref9163" refString="Trouessart, E. (1894) Note sur les acariens marins (Halacaridae) recoltes par M. Henri Gadeau de Kerville sur le littoral du department de la Manche (Juillet-Aout 1893). Bulletin de la Societe Amis des Sciences Naturelles, Rouen, 9, 139 - 175." type="journal article" year="1894">Trouessart, 1894</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="6AE24D725003FF82FC734FC6FA84F944" authority="Andre, 1961" authorityName="Andre" authorityYear="1961" box="[1016,1315,1703,1730]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="sabularius">
<emphasis id="9F96EAE35003FF82FC734FC6FB2CF944" box="[1016,1163,1703,1730]" italics="true" pageId="11" pageNumber="272">S. sabularius</emphasis>
André, 1961
</taxonomicName>
,
<taxonomicName id="6AE24D725003FF82FABA4FC6FEF5F963" authority="Trouessart, 1889" authorityName="Trouessart" authorityYear="1889" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="9F96EAE35003FF82FABA4FC6FA3BF944" box="[1329,1436,1703,1730]" italics="true" pageId="11" pageNumber="272">S. tridens</emphasis>
Trouessart, 1889
</taxonomicName>
and
<taxonomicName id="6AE24D725003FF82FE024FA2FD14F960" authority="Bartsch, 1986" authorityName="Bartsch" authorityYear="1986" box="[393,691,1739,1766]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="tereninus">
<emphasis id="9F96EAE35003FF82FE024FA2FDABF960" box="[393,524,1740,1766]" italics="true" pageId="11" pageNumber="272">S. tereninus</emphasis>
Bartsch, 1986
</taxonomicName>
). However, the new species easily differs from them by having
<emphasis id="9F96EAE35003FF82FAF74FA5FA3BF960" bold="true" box="[1404,1436,1739,1765]" pageId="11" pageNumber="272">(1)</emphasis>
larger spermatopositor (see
<figureCitation id="35D92A745003FF82FE444F81FDEDF88C" box="[463,586,1775,1801]" captionStart="FIGURE 6" captionStartId="12.[152,255,1856,1880]" captionTargetBox="[213,1373,184,1826]" captionTargetId="figure-23@12.[209,1378,179,1830]" captionTargetPageId="12" captionText="FIGURE 6. Scaptognathus antalyaensis sp. nov.: Line art illustrations of female (AE) (FDHAL-24/106) and male (FI) (FDHAL-24/107): A Idiosoma, dorsal view B Idiosoma, ventral view C Genitoanal plate, detailed D Gnathosoma, dorsal view E Right palp, dorsal view F Idiosoma, dorsal view G Idiosoma, ventral view H Genitoanal plate, detailed I Spermatopositor (S. antalyaensis sp. nov.) J Spermatopositor (S. hallezi) K Spermatopositor (S. sabularius). Scale bars: A, B, D, F, G=100 µm; C, H=50 µm; E= 50 µm; IK= 25 µm." figureDoi="http://doi.org/10.5281/zenodo.14033584" httpUri="https://zenodo.org/record/14033584/files/figure.png" pageId="11" pageNumber="272">Figs. 6JL</figureCitation>
),
<emphasis id="9F96EAE35003FF82FDD64F81FDD9F88C" bold="true" box="[605,638,1775,1801]" pageId="11" pageNumber="272">(2)</emphasis>
striking reticulate sculptures on the plates,
<emphasis id="9F96EAE35003FF82FBEC4F81FB2FF88C" bold="true" box="[1127,1160,1775,1801]" pageId="11" pageNumber="272">(3)</emphasis>
shorter ds-1, and
<emphasis id="9F96EAE35003FF82FAD94F81FAD4F88C" bold="true" box="[1362,1395,1775,1801]" pageId="11" pageNumber="272">(4)</emphasis>
the number of male perigenital setae (in
<taxonomicName id="6AE24D725003FF82FDB14E7AFD05F8A8" authorityName="Trouessart" authorityYear="1894" box="[570,674,1811,1838]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="hallezi">
<emphasis id="9F96EAE35003FF82FDB14E7AFD05F8A8" box="[570,674,1811,1838]" italics="true" pageId="11" pageNumber="272">S. hallezi</emphasis>
</taxonomicName>
2+3138; in
<taxonomicName id="6AE24D725003FF82FCB04E7AFC69F8A8" authorityName="Andre" authorityYear="1961" box="[827,974,1811,1838]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="sabularius">
<emphasis id="9F96EAE35003FF82FCB04E7AFC69F8A8" box="[827,974,1811,1838]" italics="true" pageId="11" pageNumber="272">S. sabularius</emphasis>
</taxonomicName>
2+24; in
<taxonomicName id="6AE24D725003FF82FBB54E7AFB0EF8A8" authorityName="Trouessart" authorityYear="1889" box="[1086,1193,1811,1838]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="9F96EAE35003FF82FBB54E7AFB0EF8A8" box="[1086,1193,1811,1838]" italics="true" pageId="11" pageNumber="272">S. tridens</emphasis>
</taxonomicName>
2+16; in
<taxonomicName id="6AE24D725003FF82FA934E7AFA3BF8A8" authorityName="Bartsch" authorityYear="1986" box="[1304,1436,1812,1838]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="tereninus">
<emphasis id="9F96EAE35003FF82FA934E7AFA3BF8A8" box="[1304,1436,1812,1838]" italics="true" pageId="11" pageNumber="272">S. tereninus</emphasis>
</taxonomicName>
2+2230
<emphasis id="9F96EAE35003FF82FE8F4E56FEBCF8D4" box="[260,283,1848,1873]" italics="true" pageId="11" pageNumber="272">vs</emphasis>
in
<taxonomicName id="6AE24D725003FF82FECF4E56FE48F8D4" authorityName="Durucan" authorityYear="2024" box="[324,495,1847,1874]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="antalyaensis" status="sp. nov.">
<emphasis id="9F96EAE35003FF82FECF4E56FE48F8D4" box="[324,495,1847,1874]" italics="true" pageId="11" pageNumber="272">S. antalyaensis</emphasis>
</taxonomicName>
<emphasis id="9F96EAE35003FF82FE734E56FDF3F8D7" bold="true" box="[504,596,1848,1874]" pageId="11" pageNumber="272">
<taxonomicNameLabel id="84A557985003FF82FE734E56FDF3F8D7" box="[504,596,1848,1874]" pageId="11" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
2+55. Moreover, tibia II has four bipectinate setae in
<taxonomicName id="6AE24D725003FF82FB434E56FAD4F8D4" authorityName="Durucan" authorityYear="2024" box="[1224,1395,1847,1874]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="antalyaensis" status="sp. nov.">
<emphasis id="9F96EAE35003FF82FB434E56FAD4F8D4" box="[1224,1395,1847,1874]" italics="true" pageId="11" pageNumber="272">S. antalyaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84A557985003FF82FAF74E56FF60F8F3" pageId="11" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="9F96EAE35003FF82FF464E32FF43F8F0" box="[205,228,1884,1909]" italics="true" pageId="11" pageNumber="272">vs</emphasis>
three in
<taxonomicName id="6AE24D725003FF82FECC4E32FE09F8F0" authorityName="Trouessart" authorityYear="1894" box="[327,430,1883,1910]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="hallezi">
<emphasis id="9F96EAE35003FF82FECC4E32FE09F8F0" box="[327,430,1883,1910]" italics="true" pageId="11" pageNumber="272">S. hallezi</emphasis>
</taxonomicName>
,
<taxonomicName id="6AE24D725003FF82FE374E32FDEAF8F0" authorityName="Andre" authorityYear="1961" box="[444,589,1883,1910]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="sabularius">
<emphasis id="9F96EAE35003FF82FE374E32FDEAF8F0" box="[444,589,1883,1910]" italics="true" pageId="11" pageNumber="272">S. sabularius</emphasis>
</taxonomicName>
and
<taxonomicName id="6AE24D725003FF82FD0F4E32FCA0F8F0" authorityName="Bartsch" authorityYear="1986" box="[644,775,1884,1910]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="tereninus">
<emphasis id="9F96EAE35003FF82FD0F4E32FCA0F8F0" box="[644,775,1884,1910]" italics="true" pageId="11" pageNumber="272">S. tereninus</emphasis>
</taxonomicName>
and gnathosoma larger than
<taxonomicName id="6AE24D725003FF82FBC14E32FB16F8F0" authorityName="Trouessart" authorityYear="1894" box="[1098,1201,1883,1910]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="hallezi">
<emphasis id="9F96EAE35003FF82FBC14E32FB16F8F0" box="[1098,1201,1883,1910]" italics="true" pageId="11" pageNumber="272">S. hallezi</emphasis>
</taxonomicName>
,
<taxonomicName id="6AE24D725003FF82FB344E32FAF6F8F0" authorityName="Andre" authorityYear="1961" box="[1215,1361,1883,1910]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="sabularius">
<emphasis id="9F96EAE35003FF82FB344E32FAF6F8F0" box="[1215,1361,1883,1910]" italics="true" pageId="11" pageNumber="272">S. sabularius</emphasis>
</taxonomicName>
and
<taxonomicName id="6AE24D725003FF82FA0C4E32FF59F81C" authorityName="Bartsch" authorityYear="1986" class="Arachnida" family="Halacaridae" genus="Scaptognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="272" phylum="Arthropoda" rank="species" species="tereninus">
<emphasis id="9F96EAE35003FF82FA0C4E32FF59F81C" italics="true" pageId="11" pageNumber="272">S. tereninus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,183 @@
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<subSubSection id="A79665027942FFE7CED9F921FC09F90C" box="[151,909,1698,1725]" pageId="20" pageNumber="421" type="nomenclature">
<paragraph id="EF3336897942FFE7CED9F921FC09F90C" blockId="20.[151,909,1698,1725]" box="[151,909,1698,1725]" pageId="20" pageNumber="421">
<heading id="B47B81E57942FFE7CED9F921FC09F90C" bold="true" box="[151,909,1698,1725]" fontSize="11" level="1" pageId="20" pageNumber="421" reason="1">
<emphasis id="DDF8EA9B7942FFE7CED9F921FC09F90C" bold="true" box="[151,909,1698,1725]" pageId="20" pageNumber="421">
<taxonomicName id="288C4D0A7942FFE7CED9F921FD36F90C" ID-CoL="C2D2" authority="(Brinkhurst 1964)" baseAuthorityName="Brinkhurst" baseAuthorityYear="1964" box="[151,690,1698,1725]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="20" pageNumber="421" phylum="Annelida" rank="species" species="americanus">
<emphasis id="DDF8EA9B7942FFE7CED9F921FE4DF90D" bold="true" box="[151,457,1698,1725]" italics="true" pageId="20" pageNumber="421">Brinkhurstia americanus</emphasis>
(
<bibRefCitation id="8B1D4B787942FFE7CF96F921FD2DF90C" author="Brinkhurst, R. O." box="[472,681,1698,1724]" pageId="20" pageNumber="421" pagination="527 - 536" refId="ref22581" refString="Brinkhurst, R. O. (1964) A taxonomic revision of the Alluroididae (Oligochaeta). Proceedings of the Zoological Society of London, 142, 527 - 536. https: // doi. org / 10.1111 / j. 1469 - 7998.1964. tb 04512. x" type="journal article" year="1964">Brinkhurst 1964</bibRefCitation>
)
</taxonomicName>
(See
<figureCitation id="77B72A0C7942FFE7CCBBF921FCBBF90C" box="[757,831,1698,1724]" captionStart="FIGURE 3" captionStartId="7.[151,250,1800,1824]" captionTargetBox="[357,1231,1173,1769]" captionTargetId="figure-517@7.[357,1231,1173,1777]" captionTargetPageId="7" captionText="FIGURE 3. Brinkhurstia americanus (Brinkhurst 1964). Longitudinal section through the male terminalia. at.ch, terminal chamber of atrium; at.d, atrial duct; gl.sh, glandular sheath; m.sh, muscular sheath; o.v, ovary; pro, protractor muscle; p.s, penial seta; re, retractor muscle; sep, septum; v.d, vas deferens. From Jamieson (1968)." figureDoi="http://doi.org/10.5281/zenodo.14022633" httpUri="https://zenodo.org/record/14022633/files/figure.png" pageId="20" pageNumber="421">Figs 3</figureCitation>
&amp;
<figureCitation id="77B72A0C7942FFE7CD28F921FC00F90C" box="[870,900,1698,1724]" captionStart="FIGURE 10" captionStartId="21.[151,250,861,885]" captionTargetBox="[536,1038,320,822]" captionTargetId="figure-65@21.[508,1080,273,838]" captionTargetPageId="21" captionText="FIGURE 10. Brinkhurstia americanus. (Brinkhurst 1964). Spermatheca opening middorsally in the anterior of IX. Abbreviations: a, ampulla; cv, commissural vessel. From Omodeo &amp; Coates (2001), reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.14022653" httpUri="https://zenodo.org/record/14022653/files/figure.png" pageId="20" pageNumber="421">10</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A79665027942FFE7CED9F964FF7AF8C9" pageId="20" pageNumber="421" type="reference_group">
<paragraph id="EF3336897942FFE7CED9F964FD55F94F" blockId="20.[151,1437,1767,1914]" box="[151,721,1767,1791]" pageId="20" pageNumber="421">
<treatmentCitationGroup id="CF9C11A77942FFE7CED9F964FD55F94F" box="[151,721,1767,1791]" pageId="20" pageNumber="421">
<taxonomicName id="288C4D0A7942FFE7CED9F964FEF8F94F" ID-CoL="C2D2" authorityName="Brinkhurst" authorityYear="1964" box="[151,380,1767,1791]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="20" pageNumber="421" phylum="Annelida" rank="species" species="americanus">
<emphasis id="DDF8EA9B7942FFE7CED9F964FEF8F94F" box="[151,380,1767,1791]" italics="true" pageId="20" pageNumber="421">Alluroides americanus</emphasis>
</taxonomicName>
nom. nud.
<treatmentCitation id="6E2D10987942FFE7CFA1F964FD49F94F" author="Cernosvitov, L." box="[495,717,1767,1791]" page="19" pageId="20" pageNumber="421" year="1936">
<bibRefCitation id="8B1D4B787942FFE7CFA1F964FD49F94F" author="Cernosvitov, L." box="[495,717,1767,1791]" pageId="20" pageNumber="421" pagination="16 - 19" refId="ref22730" refString="Cernosvitov, L. (1936) Notes sur la distribution mondiale de quelques oligochetes. Meimoires de la Socieitei Zoologique Tcheicoslovaque de Prague, 3, 16 - 19." type="journal article" year="1936">Cernosvitov 1936: 19</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="EF3336897942FFE7CED9F885FD7EF8AE" blockId="20.[151,1437,1767,1914]" box="[151,762,1798,1822]" pageId="20" pageNumber="421">
<treatmentCitationGroup id="CF9C11A77942FFE7CED9F885FD7EF8AE" box="[151,762,1798,1822]" pageId="20" pageNumber="421">
<treatmentCitation id="6E2D10987942FFE7CED9F885FD16F8AE" author="Brinkhurst, R. O." box="[151,658,1798,1822]" page="534" pageId="20" pageNumber="421" year="1964">
<taxonomicName id="288C4D0A7942FFE7CED9F885FD16F8AE" ID-CoL="C2D2" authority="Brinkhurst 1964: 533 - 534" authorityName="Brinkhurst" authorityPageNumber="533 - 534" authorityYear="1964" box="[151,658,1798,1822]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="20" pageNumber="421" phylum="Annelida" rank="species" species="americanus">
<emphasis id="DDF8EA9B7942FFE7CED9F885FEF8F8AD" box="[151,380,1798,1822]" italics="true" pageId="20" pageNumber="421">Alluroides americanus</emphasis>
<bibRefCitation id="8B1D4B787942FFE7CFCDF885FD16F8AE" author="Brinkhurst, R. O." box="[387,658,1798,1822]" pageId="20" pageNumber="421" pagination="527 - 536" refId="ref22581" refString="Brinkhurst, R. O. (1964) A taxonomic revision of the Alluroididae (Oligochaeta). Proceedings of the Zoological Society of London, 142, 527 - 536. https: // doi. org / 10.1111 / j. 1469 - 7998.1964. tb 04512. x" type="journal article" year="1964">Brinkhurst 1964: 533534</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<figureCitation id="77B72A0C7942FFE7CCD5F885FD72F8AE" box="[667,758,1798,1822]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="13.[151,250,1798,1822]" captionStartId-1="14.[151,250,1657,1681]" captionStartId-2="16.[151,250,1164,1188]" captionTargetBox-0="[205,1383,202,1774]" captionTargetBox-1="[176,1411,1183,1620]" captionTargetBox-2="[473,1052,399,1141]" captionTargetId-0="figure-19@13.[205,1383,201,1775]" captionTargetId-1="figure-399@14.[158,1430,1174,1636]" captionTargetId-2="figure-134@16.[473,1115,381,1141]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="16" captionText-0="FIGURE 5. Alluroides brinkhursti brinkhursti Jamieson (1968). A. Transverse section (TS) of clitellum, showing single cell layer; the cells with conspicuous secretory granules and each with a basal nucleus. B. TS through the wall of the atrium, showing a group of atrial gland cells with ductule penetrating the muscular sheath of the atrium. C. Longitudinal section through the male pore, showing the ectal end of the atrium, which forms a penis with muscular sheath, ciliated epithelium and rope of spermatozoa in the lumen, forming in the ectal chamber a sperm mass. D. Alluroides pordagei. Oblique section through the atrial bulb, containing a large sperm mass, and the associated atrium. From Jamieson (2006)." captionText-1="FIGURE 6. Alluroides brinkhursti abyssinicus Jamieson (1968). Longitudinal section of holotype. Abbreviations: epi, epithelium of atrium; gl sh, glandular sheath of atrium; m, mouth; msh, muscular sheath of atrium; oe, oesophagus; oo, oocyte: ph m, pharyngeal musculature; s.ves, seminal vesicle; sp, spermatheca; vbv, ventral blood vessel; vd, vas deferens; v n, ventral nerve cord. Slightly modified from Jamieson (1968)." captionText-2="FIGURE 7. Alluroides lauzannei Ljungström, 1971. Detail of an atrium. Note the vas deferens entering the apex of the atrium via canaliculi. Relabelled from Lauzanne (1968)." figureDoi-0="http://doi.org/10.5281/zenodo.14022639" figureDoi-1="http://doi.org/10.5281/zenodo.14022641" figureDoi-2="http://doi.org/10.5281/zenodo.14022643" httpUri-0="https://zenodo.org/record/14022639/files/figure.png" httpUri-1="https://zenodo.org/record/14022641/files/figure.png" httpUri-2="https://zenodo.org/record/14022643/files/figure.png" pageId="20" pageNumber="421">Figs.57</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="EF3336897942FFE7CED9F8A7FF7AF8C9" blockId="20.[151,1437,1767,1914]" pageId="20" pageNumber="421">
<treatmentCitationGroup id="CF9C11A77942FFE7CED9F8A7FA18F88D" box="[151,1436,1828,1853]" pageId="20" pageNumber="421">
<taxonomicName id="288C4D0A7942FFE7CED9F8A7FD9FF88C" ID-CoL="C2D2" authority="(Brinkhurst)" baseAuthorityName="Brinkhurst" baseAuthorityYear="1964" box="[151,539,1828,1852]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="20" pageNumber="421" phylum="Annelida" rank="species" species="americanus">
<emphasis id="DDF8EA9B7942FFE7CED9F8A7FE10F88C" box="[151,404,1828,1852]" italics="true" pageId="20" pageNumber="421">Brinkhurstia americanus</emphasis>
(Brinkhurst)
</taxonomicName>
,
<treatmentCitation id="6E2D10987942FFE7CC68F8A7FC89F88D" author="Jamieson, B. G. M." box="[550,781,1828,1853]" page="80" pageId="20" pageNumber="421" year="1968">
<bibRefCitation id="8B1D4B787942FFE7CC68F8A7FC89F88D" author="Jamieson, B. G. M." box="[550,781,1828,1853]" pageId="20" pageNumber="421" pagination="55 - 86" refId="ref23501" refString="Jamieson, B. G. M. (1968) A taxonometric investigation of the Alluroididae (Oligochaeta). Journal of Zoology, 155, 55 - 86. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03029. x" type="journal article" year="1968">Jamieson 1968: 8081</bibRefCitation>
</treatmentCitation>
,
<figureCitation id="77B72A0C7942FFE7CD56F8A7FCDBF88D" box="[792,863,1828,1853]" captionStart="FIGURE 3" captionStartId="7.[151,250,1800,1824]" captionTargetBox="[357,1231,1173,1769]" captionTargetId="figure-517@7.[357,1231,1173,1777]" captionTargetPageId="7" captionText="FIGURE 3. Brinkhurstia americanus (Brinkhurst 1964). Longitudinal section through the male terminalia. at.ch, terminal chamber of atrium; at.d, atrial duct; gl.sh, glandular sheath; m.sh, muscular sheath; o.v, ovary; pro, protractor muscle; p.s, penial seta; re, retractor muscle; sep, septum; v.d, vas deferens. From Jamieson (1968)." figureDoi="http://doi.org/10.5281/zenodo.14022633" httpUri="https://zenodo.org/record/14022633/files/figure.png" pageId="20" pageNumber="421">Figs. 3</figureCitation>
,
<figureCitation id="77B72A0C7942FFE7CD25F8A6FC03F88D" box="[875,903,1829,1853]" captionStart="FIGURE 12" captionStartId="23.[151,250,1189,1213]" captionTargetBox="[366,1270,294,1138]" captionTargetId="figure-61@23.[287,1301,273,1155]" captionTargetPageId="23" captionText="FIGURE 12. Brinkhurstia donaldi Omodeo &amp; Coates 2001. Spermatheca, sagittal section. Abbreviations: a, ampulla; d, duct; di, diverticulum; se, septum with muscle fibres; spz, spermatozoa; ss, sponge spicules. From Omodeo and Coates (2001), reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.14022659" httpUri="https://zenodo.org/record/14022659/files/figure.png" pageId="20" pageNumber="421">12</figureCitation>
;
<treatmentCitation id="6E2D10987942FFE7CDDCF8A7FAA5F88C" author="Brinkhurst, R. O. &amp; Jamieson, B. G. M." box="[914,1313,1828,1853]" page="716" pageId="20" pageNumber="421" year="1971">
<bibRefCitation id="8B1D4B787942FFE7CDDCF8A7FAA5F88C" author="Brinkhurst, R. O. &amp; Jamieson, B. G. M." box="[914,1313,1828,1853]" pageId="20" pageNumber="421" pagination="708 - 722" refId="ref22671" refString="Brinkhurst, R. O. &amp; Jamieson, B. G. M. (1971) 14. Family Alluroididae. In: Brinkhurst, R. O. &amp; Jamieson, B. G. M., Aquatic Oligochaeta of the World. Oliver and Boyd, Edinburgh, pp. 708 - 722." type="book chapter" year="1971">Brinkhurst &amp; Jamieson 1971: 716717</bibRefCitation>
</treatmentCitation>
,
<figureCitation id="77B72A0C7942FFE7CB65F8A7FA18F88D" box="[1323,1436,1828,1853]" captionStart="FIGURE 14" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[206,1382,421,1809]" captionTargetId="figure-158@26.[201,1386,417,1813]" captionTargetPageId="26" captionText="FIGURE 14. Lacandodrilus paludosus Fragoso &amp; Rojas 2023. A. External view of the anterior ventral region of a clitellate adult. B. Internal view of the anterior region of an adult. C. Spermatheca. Abbreviations: a, ampulla: clit, clitellum; d, duct; dv, dorsal vessel; gg, genital glands (prostates?); img, internal male gonoducts; mf, male funnels; n, nephridia; oe, oesophagus; sg, septal glands. Modified from Fragoso &amp; Rojas (2023)." figureDoi="http://doi.org/10.5281/zenodo.14022665" httpUri="https://zenodo.org/record/14022665/files/figure.png" pageId="20" pageNumber="421">
Fig.
<geoCoordinate id="8AB8504E7942FFE7CB14F8A6FA1CF88D" box="[1370,1432,1829,1853]" degrees="14.1" direction="east" orientation="longitude" pageId="20" pageNumber="421" precision="5555" value="14.1">14.1E</geoCoordinate>
,
</figureCitation>
</treatmentCitationGroup>
F;
<bibRefCitation id="8B1D4B787942FFE7CEADF8C0FE59F8EB" author="Righi, G. &amp; Ayres, I. &amp; Bittencourt, E. C. R." box="[227,477,1859,1883]" pageId="20" pageNumber="421" pagination="1 - 49" refId="ref24391" refString="Righi, G., Ayres, I. &amp; Bittencourt, E. C. R. (1978) Oligochaeta (Annelida) do Instituto Nacional de Pesquisas da Amazonia. Acta Amazonica, 8, 1 - 49. https: // doi. org / 10.1590 / 1809 - 439219780831 s 005" type="journal article" year="1978">
Righi
<emphasis id="DDF8EA9B7942FFE7CF6CF8C7FECBF8EB" box="[290,335,1859,1883]" italics="true" pageId="20" pageNumber="421">et al</emphasis>
. 1978: 1112
</bibRefCitation>
,
<figureCitation id="77B72A0C7942FFE7CFA6F8C0FDBDF8EB" box="[488,569,1859,1883]" captionStart="FIGURE 11" captionStartId="22.[151,250,1028,1052]" captionTargetBox="[297,1290,345,997]" captionTargetId="figure-124@22.[285,1302,342,1007]" captionTargetPageId="22" captionText="FIGURE 11. Brinkhurstia donaldi Omodeo and Coates (2001).A. Scheme showing arrangement of genital organs and vascular commissures. B. Cross section through spermathecal duct. C. Cross section of the muscular sac ensheathing penial seta. D. Section through ental part of prostate. E. Cross section of prostatic duct. F. Dorsal view of the arrangement of the penial setae within IXXIII. Abbreviations: a, spermathecal ampulla, cv, commissural vessels, d, spermathecal duct, di, spermathecal diverticulum, e, egg, if female funnel, md, male duct, pc, penial seta, pr, prostate. From Omodeo &amp; Coates (2001), reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.14022655" httpUri="https://zenodo.org/record/14022655/files/figure.png" pageId="20" pageNumber="421">Figs. 11</figureCitation>
21;
<bibRefCitation id="8B1D4B787942FFE7CC23F8C0FCD7F8EB" author="Stacey, D. F. &amp; Coates, K. A." box="[621,851,1859,1883]" pageId="20" pageNumber="421" pagination="17 - 29" refId="ref24568" refString="Stacey, D. F. &amp; Coates, K. A. (1996) Oligochaetes (Naididae, Tubificidae, Opistocystidae, Enchytraeidae, Sparganophilidae and Alluroididae) of Guyana. Hydrobiologia, 334, 17 - 29. https: // doi. org / 10.1007 / BF 00017350" type="journal article" year="1996">Stacey &amp; Coates 1996</bibRefCitation>
:
<tableCitation id="A20E03327942FFE7CD13F8C0FC37F8EB" box="[861,947,1859,1883]" captionStart="TABLE 1" captionStartId="11.[151,243,849,875]" captionTargetPageId="11" captionText="TABLE 1. The most significant characters distinguishing the genera of the Alluroidinae (modified and expanded from Omodeo and Coates, 2001)." pageId="20" pageNumber="421">Tables 1</tableCitation>
and 2;
<bibRefCitation id="8B1D4B787942FFE7CDB1F8C0FAD2F8EB" author="Omodeo, P. &amp; Coates, K. A." box="[1023,1366,1859,1883]" pageId="20" pageNumber="421" pagination="39 - 47" refId="ref24287" refString="Omodeo, P. &amp; Coates, K. A. (2001) New alluroidids (Annelida, Clitellata) from Guyana. Hydrobiologia, 463, 39 - 47. https: // doi. org / 10.1023 / A: 1013178902798" type="journal article" year="2001">Omodeo and Coates 2001: 3940</bibRefCitation>
,
<figureCitation id="77B72A0C7942FFE7CB2FF8C0FA19F8EB" box="[1377,1437,1859,1883]" captionStart="FIGURE 1" captionStartId="1.[151,250,1089,1113]" captionTargetBox="[277,1306,185,1056]" captionTargetId="figure-19@1.[271,1316,179,1063]" captionTargetPageId="1" captionText="FIGURE 1A. Hennigram (morphological phylogram) for all families of the opisthoporous oligochaetes constructed by nearestneighbour clustering. For details see Jamieson (1978). Redrawn." figureDoi="http://doi.org/10.5281/zenodo.14022625" httpUri="https://zenodo.org/record/14022625/files/figure.png" pageId="20" pageNumber="421">Fig. 1</figureCitation>
A, B.
</paragraph>
</subSubSection>
<subSubSection id="A79665027942FFE6CED9F82BFACEF9F6" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421" type="description">
<paragraph id="EF3336897942FFE6CED9F82BFCB4FF49" blockId="20.[151,1436,1960,2023]" lastBlockId="21.[151,1436,150,249]" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421">
Length
<quantity id="28749B6C7942FFE7CEA0F82BFEEDF873" box="[238,361,1960,1987]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.0" metricValueMin="2.4" pageId="20" pageNumber="421" unit="mm" value="27.0" valueMax="30.0" valueMin="24.0">2430 mm</quantity>
, width
<quantity id="28749B6C7942FFE7CFF5F82BFDE5F873" box="[443,609,1960,1987]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="7.5" metricValueMin="5.6" pageId="20" pageNumber="421" unit="mm" value="0.655" valueMax="0.75" valueMin="0.56">0.560.75 mm</quantity>
, segments 100130. Prostomium short or large and rounded. Ventral setal couples of XIII replaced by a single penial seta on each side; in the ovarian segments
<emphasis id="DDF8EA9B7942FFE7CA7CF84FFB65F856" box="[1074,1249,1996,2022]" italics="true" pageId="20" pageNumber="421">aa:ab:bc:cd:dd</emphasis>
=4.7:1:5:1.1:6.2;
<emphasis id="DDF8EA9B7943FFE6CED9FF14FF37FF01" box="[151,179,151,177]" italics="true" pageId="21" pageNumber="422">dd</emphasis>
=025 u; length of a lateral seta from XI, base to node=87 µm. Clitellum annular, slightly less developed ventrally; segments 1/2XIIXV. Lateral lines present. Male pores ventrolateral in XIII. Female pores anterior in XIV. Spermathecal pore single, middorsal, anteriorly in IX.
</paragraph>
<caption id="BBF366017943FFE6CED9FCDEFA97FC2A" ID-DOI="http://doi.org/10.5281/zenodo.14022653" ID-Zenodo-Dep="14022653" httpUri="https://zenodo.org/record/14022653/files/figure.png" pageId="21" pageNumber="422" startId="21.[151,250,861,885]" targetBox="[536,1038,320,822]" targetPageId="21" targetType="figure">
<paragraph id="EF3336897943FFE6CED9FCDEFA97FC2A" blockId="21.[151,1437,861,922]" pageId="21" pageNumber="422">
<emphasis id="DDF8EA9B7943FFE6CED9FCDEFDC0FCC5" bold="true" box="[151,580,861,885]" pageId="21" pageNumber="422">
FIGURE 10.
<taxonomicName id="288C4D0A7943FFE6CF7BFCDEFDC0FCC5" baseAuthorityName="Brinkhurst" baseAuthorityYear="1964" box="[309,580,861,885]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="21" pageNumber="422" phylum="Annelida" rank="species" species="americanus">
<emphasis id="DDF8EA9B7943FFE6CF7BFCDEFDC0FCC5" bold="true" box="[309,580,861,885]" italics="true" pageId="21" pageNumber="422">Brinkhurstia americanus</emphasis>
</taxonomicName>
</emphasis>
. (
<bibRefCitation id="8B1D4B787943FFE6CC11FCDDFC96FCC5" author="Brinkhurst, R. O." box="[607,786,861,886]" pageId="21" pageNumber="422" pagination="527 - 536" refId="ref22581" refString="Brinkhurst, R. O. (1964) A taxonomic revision of the Alluroididae (Oligochaeta). Proceedings of the Zoological Society of London, 142, 527 - 536. https: // doi. org / 10.1111 / j. 1469 - 7998.1964. tb 04512. x" type="journal article" year="1964">Brinkhurst 1964</bibRefCitation>
). Spermatheca opening middorsally in the anterior of IX. Abbreviations: a, ampulla; cv, commissural vessel. From
<bibRefCitation id="8B1D4B787943FFE6CC90FC01FC63FC2A" author="Omodeo, P. &amp; Coates, K. A." box="[734,999,898,922]" pageId="21" pageNumber="422" pagination="39 - 47" refId="ref24287" refString="Omodeo, P. &amp; Coates, K. A. (2001) New alluroidids (Annelida, Clitellata) from Guyana. Hydrobiologia, 463, 39 - 47. https: // doi. org / 10.1023 / A: 1013178902798" type="journal article" year="2001">Omodeo &amp; Coates (2001)</bibRefCitation>
, reproduced with permission.
</paragraph>
</caption>
<paragraph id="EF3336897943FFE6CE89FC6FFACEF9F6" blockId="21.[151,1437,1004,1642]" pageId="21" pageNumber="422">Septum 3/4 excessively delicate and doubtfully complete; 4/5 and 14/15 hardly appreciably, 6/78/9 fairly strongly thickened (two tiers of muscle fibres) the remainder intermediate; 10/11 has one tier of muscle fibres and is not backwardly deflected. Septal glands in VVIII. Alimentary canal expanding greatly in XVI but no definite transition from oesophagus to intestine. Gizzard, oesophageal diverticula and typhlosole absent. Perienteric blood sinus in IX posteriad, not appreciable in front of this. Testes discrete elongated tongues of tissue dependent from the anterior wall of X. Gonadial tissue absent from XI. Seminal vesicles absent or present in XI. Vas deferens running on body wall in XI and XII. Atria very thin much coiled tubes extending from midXIII into XIV; discharging in the close proximity of the corresponding vas deferens through the posterior wall of a globular non-muscular ectal chamber; a simple, straight, unornamented penial seta in a follicle approximately twice the length of the chamber, penetrates the anterior wall of the chamber. Each prostate has a cuboidal lining epithelium surrounded in turn by a very thin muscular sheath and an external layer of prostatic cells only one to a few cells thick, though considerable masses of prostatic cells are formed between adjacent coils. Width muscular tube of atrium (external) 28 µm; width ectal chamber 37 µm; length penial seta 115 µm; greatest width 18 µm. Ovaries dependent from the anterior wall of XIII and extending posteriad into ovisacs formed by the backward evagination of septum 13/14 through as many as 8 segments: diameter of the largest oocytenuclei 33 µm. First nephridia (from peritoneal masses) in IX or X. Spermatheca in IX a simple elongated pouch, 87 µm wide, with no distinct ampulla, lined internally by a short columnar epithelium invested in a single layer of circular muscle and an outer glandular? peritoneal sheath.</paragraph>
</subSubSection>
<subSubSection id="A79665027943FFE6CE89F9CCFC35F9DA" box="[199,945,1615,1642]" pageId="21" pageNumber="422" type="distribution">
<paragraph id="EF3336897943FFE6CE89F9CCFC35F9DA" blockId="21.[151,1437,1004,1642]" box="[199,945,1615,1642]" pageId="21" pageNumber="422">
<emphasis id="DDF8EA9B7943FFE6CE89F9CCFEDDF9D9" bold="true" box="[199,345,1615,1641]" pageId="21" pageNumber="422">Distribution</emphasis>
.
<collectingCountry id="979B76197943FFE6CF29F9D3FE2BF9DA" box="[359,431,1616,1642]" name="Brazil" pageId="21" pageNumber="422">Brazil</collectingCountry>
; Saint Lucia (Caribbean);
<collectingCountry id="979B76197943FFE6CC9BF9D3FCCCF9DA" box="[725,840,1616,1642]" name="Argentina" pageId="21" pageNumber="422">Argentina</collectingCountry>
;
<collectingCountry id="979B76197943FFE6CD1DF9D3FC2EF9DA" box="[851,938,1616,1642]" name="Guyana" pageId="21" pageNumber="422">Guyana</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="ED6146E98F6825081109D592FB449D2A">A monograph of the Oligochaete family Alluroididae</mods:title>
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<paragraph id="EF3336897943FFE6CED9F938FC1CF965" blockId="21.[151,920,1723,1750]" box="[151,920,1723,1750]" pageId="21" pageNumber="422">
<heading id="B47B81E57943FFE6CED9F938FC1CF965" bold="true" box="[151,920,1723,1750]" fontSize="11" level="1" pageId="21" pageNumber="422" reason="1">
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<taxonomicName id="288C4D0A7943FFE6CED9F938FD49F966" ID-CoL="NBD9" authority="Omodeo &amp; Coates, 2001" authorityName="Omodeo &amp; Coates" authorityYear="2001" box="[151,717,1723,1750]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="21" pageNumber="422" phylum="Annelida" rank="species" species="donaldi">
<emphasis id="DDF8EA9B7943FFE6CED9F938FE12F966" bold="true" box="[151,406,1723,1750]" italics="true" pageId="21" pageNumber="422">Brinkhurstia donaldi</emphasis>
<bibRefCitation id="8B1D4B787943FFE6CFD0F938FD49F966" author="Omodeo, P. &amp; Coates, K. A." box="[414,717,1723,1750]" pageId="21" pageNumber="422" pagination="39 - 47" refId="ref24287" refString="Omodeo, P. &amp; Coates, K. A. (2001) New alluroidids (Annelida, Clitellata) from Guyana. Hydrobiologia, 463, 39 - 47. https: // doi. org / 10.1023 / A: 1013178902798" type="journal article" year="2001">Omodeo &amp; Coates, 2001</bibRefCitation>
</taxonomicName>
(see
<figureCitation id="77B72A0C7943FFE6CD45F938FCE7F966" box="[779,867,1723,1750]" captionStart="FIGURE 11" captionStartId="22.[151,250,1028,1052]" captionTargetBox="[297,1290,345,997]" captionTargetId="figure-124@22.[285,1302,342,1007]" captionTargetPageId="22" captionText="FIGURE 11. Brinkhurstia donaldi Omodeo and Coates (2001).A. Scheme showing arrangement of genital organs and vascular commissures. B. Cross section through spermathecal duct. C. Cross section of the muscular sac ensheathing penial seta. D. Section through ental part of prostate. E. Cross section of prostatic duct. F. Dorsal view of the arrangement of the penial setae within IXXIII. Abbreviations: a, spermathecal ampulla, cv, commissural vessels, d, spermathecal duct, di, spermathecal diverticulum, e, egg, if female funnel, md, male duct, pc, penial seta, pr, prostate. From Omodeo &amp; Coates (2001), reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.14022655" httpUri="https://zenodo.org/record/14022655/files/figure.png" pageId="21" pageNumber="422">Figs 11</figureCitation>
,
<figureCitation id="77B72A0C7943FFE6CD3EF938FC0AF965" box="[880,910,1723,1749]" captionStart="FIGURE 12" captionStartId="23.[151,250,1189,1213]" captionTargetBox="[366,1270,294,1138]" captionTargetId="figure-61@23.[287,1301,273,1155]" captionTargetPageId="23" captionText="FIGURE 12. Brinkhurstia donaldi Omodeo &amp; Coates 2001. Spermatheca, sagittal section. Abbreviations: a, ampulla; d, duct; di, diverticulum; se, septum with muscle fibres; spz, spermatozoa; ss, sponge spicules. From Omodeo and Coates (2001), reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.14022659" httpUri="https://zenodo.org/record/14022659/files/figure.png" pageId="21" pageNumber="422">12</figureCitation>
)
</emphasis>
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<subSubSection id="A79665027943FFE6CED9F883FAB7F856" pageId="21" pageNumber="422" type="reference_group">
<paragraph id="EF3336897943FFE6CED9F883FD6BF8A8" blockId="21.[151,818,1792,1878]" box="[151,751,1792,1816]" pageId="21" pageNumber="422">
<treatmentCitationGroup id="CF9C11A77943FFE6CED9F883FD6BF8A8" box="[151,751,1792,1816]" pageId="21" pageNumber="422">
<taxonomicName id="288C4D0A7943FFE6CED9F883FEBFF8A8" box="[151,315,1792,1816]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="21" pageNumber="422" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="DDF8EA9B7943FFE6CED9F883FE93F8A8" box="[151,279,1792,1816]" italics="true" pageId="21" pageNumber="422">Brinkhurstia</emphasis>
sp.
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<treatmentCitation id="6E2D10987943FFE6CF0FF883FDCFF8A8" author="Coates, K. A. &amp; Stacey, D. F." box="[321,587,1792,1816]" page="81" pageId="21" pageNumber="422" year="1994">
<bibRefCitation id="8B1D4B787943FFE6CF0FF883FDCFF8A8" author="Coates, K. A. &amp; Stacey, D. F." box="[321,587,1792,1816]" pageId="21" pageNumber="422" pagination="79 - 84" refId="ref22798" refString="Coates, K. A. &amp; Stacey, D. F. (1994) Oligochaetes (Naididae, Tubificidae, Enchytraeidae and Alluroididae) of Guyana, Peru and Ecuador. Hydrobiologia, 278, 79 - 84. https: // doi. org / 10.1007 / BF 00142313" type="journal article" year="1994">Coates &amp; Stacey 1994: 81</bibRefCitation>
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,
<tableCitation id="A20E03327943FFE6CC18F883FD25F8A8" box="[598,673,1792,1816]" captionStart="TABLE 1" captionStartId="11.[151,243,849,875]" captionTargetPageId="11" captionText="TABLE 1. The most significant characters distinguishing the genera of the Alluroidinae (modified and expanded from Omodeo and Coates, 2001)." pageId="21" pageNumber="422">Table 1</tableCitation>
,
<figureCitation id="77B72A0C7943FFE6CCE3F883FD6FF8A8" box="[685,747,1792,1816]" captionStart="FIGURE 2" captionStartId="2.[151,250,1933,1957]" captionTargetId="figure-21@2.[353,1234,181,1909]" captionTargetPageId="2" captionText="FIGURE 2. A phylogenetic classification of the Clitellata. (From Jamieson 2006)." pageId="21" pageNumber="422">Fig. 2</figureCitation>
.
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</paragraph>
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<taxonomicName id="288C4D0A7943FFE6CED9F89CFED4F887" box="[151,336,1823,1847]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="21" pageNumber="422" phylum="Annelida" rank="species" species="undefined-2">
<emphasis id="DDF8EA9B7943FFE6CED9F89CFE93F887" box="[151,279,1823,1847]" italics="true" pageId="21" pageNumber="422">Brinkhurstia</emphasis>
sp. 2
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.
<bibRefCitation id="8B1D4B787943FFE6CF14F89CFDBBF887" author="Stacey, D. F. &amp; Coates, K. A." box="[346,575,1823,1847]" pageId="21" pageNumber="422" pagination="17 - 29" refId="ref24568" refString="Stacey, D. F. &amp; Coates, K. A. (1996) Oligochaetes (Naididae, Tubificidae, Opistocystidae, Enchytraeidae, Sparganophilidae and Alluroididae) of Guyana. Hydrobiologia, 334, 17 - 29. https: // doi. org / 10.1007 / BF 00017350" type="journal article" year="1996">Stacey &amp; Coates 1996</bibRefCitation>
:
<tableCitation id="A20E03327943FFE6CC07F89CFD10F887" box="[585,660,1823,1847]" captionStart="TABLE 1" captionStartId="11.[151,243,849,875]" captionTargetPageId="11" captionText="TABLE 1. The most significant characters distinguishing the genera of the Alluroidinae (modified and expanded from Omodeo and Coates, 2001)." pageId="21" pageNumber="422">Table 1</tableCitation>
and
<figureCitation id="77B72A0C7943FFE6CC8BF89CFCABF887" box="[709,815,1823,1847]" captionStart="FIGURE 4" captionStartId="12.[151,250,1906,1930]" captionTargetBox="[242,1363,1542,1866]" captionTargetId="figure-568@12.[215,1371,1530,1885]" captionTargetPageId="12" captionText="FIGURE 4. Alluroides brinkhursti brinkhursti Jamieson (1968). Longitudinal section of the holotype. Abbreviations: br, brain; bv, blood vessel; m, mouth; msh, muscular sheath of atrium; oe, oesophagus; oo, oocyte: os, ovisac; ph m, pharyngeal musculature; pr c, prostatic cells; sep, septum; sep gl, septal gland; sp p, spermathecal pore; s. ves, seminal vesicle; sp, spermatheca; sp f, sperm funnel; tes, testis; vd, vas deferens; v n, ventral nerve cord. Adapted from Jamieson (1968)." figureDoi="http://doi.org/10.5281/zenodo.14022635" httpUri="https://zenodo.org/record/14022635/files/figure.png" pageId="21" pageNumber="422">Fig. 4DF</figureCitation>
.
</paragraph>
<paragraph id="EF3336897943FFE6CED9F8BDFCA2F8E6" blockId="21.[151,818,1792,1878]" box="[151,806,1854,1878]" pageId="21" pageNumber="422">
<treatmentCitationGroup id="CF9C11A77943FFE6CED9F8BDFCA2F8E6" box="[151,806,1854,1878]" pageId="21" pageNumber="422">
<treatmentCitation id="6E2D10987943FFE6CED9F8BDFD32F8E6" author="Omodeo, P. &amp; Coates, K. A." box="[151,694,1854,1878]" page="43" pageId="21" pageNumber="422" year="2001">
<taxonomicName id="288C4D0A7943FFE6CED9F8BDFD32F8E6" ID-CoL="NBD9" authority="Omodeo &amp; Coates 2001: 40 - 43" authorityName="Omodeo &amp; Coates" authorityPageNumber="40 - 43" authorityYear="2001" box="[151,694,1854,1878]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="21" pageNumber="422" phylum="Annelida" rank="species" species="donaldi">
<emphasis id="DDF8EA9B7943FFE6CED9F8BDFEEFF8E6" box="[151,363,1854,1878]" italics="true" pageId="21" pageNumber="422">Brinkhurstia donaldi</emphasis>
<bibRefCitation id="8B1D4B787943FFE6CF3FF8BDFD32F8E6" author="Omodeo, P. &amp; Coates, K. A." box="[369,694,1854,1878]" pageId="21" pageNumber="422" pagination="39 - 47" refId="ref24287" refString="Omodeo, P. &amp; Coates, K. A. (2001) New alluroidids (Annelida, Clitellata) from Guyana. Hydrobiologia, 463, 39 - 47. https: // doi. org / 10.1023 / A: 1013178902798" type="journal article" year="2001">Omodeo &amp; Coates 2001: 4043</bibRefCitation>
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, Figs, 23.
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<paragraph id="EF3336897943FFE6CED9F807FAB7F856" blockId="21.[151,1436,1924,2022]" pageId="21" pageNumber="422">
Length
<quantity id="28749B6C7943FFE6CEA4F807FEE6F82E" box="[234,354,1924,1950]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="21" pageNumber="422" unit="mm" value="31.0" valueMax="32.0" valueMin="30.0">3032mm</quantity>
, width
<quantity id="28749B6C7943FFE6CFE0F807FDD5F82E" box="[430,593,1924,1950]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" metricValueMax="9.0" metricValueMin="8.2" pageId="21" pageNumber="422" unit="mm" value="0.86" valueMax="0.9" valueMin="0.82">0.820.90mm</quantity>
; segments 105, 109, 131, 135,150,multiannulated.Lateral lines recognizable only on microscope slides: Body slender, cylindrical, tapering posteriorly; a series of buttons formed by 68 hyaline cells situated amid the longitudinal muscle fibres in a line included between setae
<emphasis id="DDF8EA9B7943FFE6CA61F84FFBB9F856" box="[1071,1085,1996,2022]" italics="true" pageId="21" pageNumber="422">b</emphasis>
and
<emphasis id="DDF8EA9B7943FFE6CA3DF84EFBFBF856" box="[1139,1151,1997,2022]" italics="true" pageId="21" pageNumber="422">c</emphasis>
. Anus terminal.
</paragraph>
</subSubSection>
<subSubSection id="A79665027940FFE5CE89FF14FAC9F866" pageId="22" pageNumber="423" type="description">
<paragraph id="EF3336897940FFE5CE89FF14FBD0FEF0" blockId="22.[151,1436,151,321]" pageId="22" pageNumber="423">
Clitellum annular, 1/3XII1/2XV, male pores in the middle of XIII aligned with setae
<emphasis id="DDF8EA9B7940FFE5CAE3FF14FB3FFF00" box="[1197,1211,151,176]" italics="true" pageId="22" pageNumber="423">a</emphasis>
; in a few cases the terminal chambers and the penial setae protrude through them. Female pores in furrow 13/14, slightly ventral to setal line
<emphasis id="DDF8EA9B7940FFE5CF4AFF5CFE96FF48" box="[260,274,223,248]" italics="true" pageId="22" pageNumber="423">a</emphasis>
. Spermathecal pore a middorsal cleft in furrow 7/8. Setae sigmoid, slender, unsculptured; the longest ones in XI and measure respectively:
<emphasis id="DDF8EA9B7940FFE5CC75FE80FDCDFEAC" box="[571,585,259,284]" italics="true" pageId="22" pageNumber="423">a</emphasis>
328 µm,
<emphasis id="DDF8EA9B7940FFE5CCFAFE80FD46FEAD" box="[692,706,259,285]" italics="true" pageId="22" pageNumber="423">b</emphasis>
284 µm,
<emphasis id="DDF8EA9B7940FFE5CD63FE80FCBDFEAC" box="[813,825,259,284]" italics="true" pageId="22" pageNumber="423">c</emphasis>
164 µm,
<emphasis id="DDF8EA9B7940FFE5CDEAFE80FC36FEAD" box="[932,946,259,285]" italics="true" pageId="22" pageNumber="423">d</emphasis>
280 µm; their lengths decreasing anteriorly and posteriorly; in XX,
<emphasis id="DDF8EA9B7940FFE5CFECFEA4FE3AFEF0" box="[418,446,295,320]" italics="true" pageId="22" pageNumber="423">aa</emphasis>
:
<emphasis id="DDF8EA9B7940FFE5CF88FEA4FE66FEF1" box="[454,482,295,321]" italics="true" pageId="22" pageNumber="423">ab</emphasis>
:
<emphasis id="DDF8EA9B7940FFE5CFA7FEA4FD87FEF1" box="[489,515,295,321]" italics="true" pageId="22" pageNumber="423">bc</emphasis>
:
<emphasis id="DDF8EA9B7940FFE5CC42FEA4FDA2FEF1" box="[524,550,295,321]" italics="true" pageId="22" pageNumber="423">cd</emphasis>
:
<emphasis id="DDF8EA9B7940FFE5CC60FEA4FDCEFEF1" box="[558,586,295,321]" italics="true" pageId="22" pageNumber="423">dd</emphasis>
=6.1:1.0:9.0:1.0:8.2. Cuticle about 2 µm thick.
</paragraph>
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<paragraph id="EF3336897940FFE5CED9FB87FEC4FB7F" blockId="22.[151,1437,1027,1232]" pageId="22" pageNumber="423">
<emphasis id="DDF8EA9B7940FFE5CED9FB87FE99FBAC" bold="true" box="[151,285,1028,1052]" pageId="22" pageNumber="423">FIGURE 11.</emphasis>
<taxonomicName id="288C4D0A7940FFE5CF6CFB80FC9DFBAC" authority="Omodeo and Coates (2001)" authorityName="Omodeo and Coates" authorityYear="2001" box="[290,793,1027,1052]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="22" pageNumber="423" phylum="Annelida" rank="species" species="donaldi">
<emphasis id="DDF8EA9B7940FFE5CF6CFB80FE79FBAB" bold="true" box="[290,509,1027,1051]" italics="true" pageId="22" pageNumber="423">Brinkhurstia donaldi</emphasis>
<bibRefCitation id="8B1D4B787940FFE5CC4CFB87FC9DFBAC" author="Omodeo, P. &amp; Coates, K. A." box="[514,793,1028,1052]" pageId="22" pageNumber="423" pagination="39 - 47" refId="ref24287" refString="Omodeo, P. &amp; Coates, K. A. (2001) New alluroidids (Annelida, Clitellata) from Guyana. Hydrobiologia, 463, 39 - 47. https: // doi. org / 10.1023 / A: 1013178902798" type="journal article" year="2001">Omodeo and Coates (2001)</bibRefCitation>
</taxonomicName>
.
<emphasis id="DDF8EA9B7940FFE5CD6EFB87FCB6FBAC" bold="true" box="[800,818,1028,1052]" pageId="22" pageNumber="423">A</emphasis>
. Scheme showing arrangement of genital organs and vascular commissures.
<emphasis id="DDF8EA9B7940FFE5CF64FBABFEBFFBF0" bold="true" box="[298,315,1064,1088]" pageId="22" pageNumber="423">B</emphasis>
. Cross section through spermathecal duct.
<emphasis id="DDF8EA9B7940FFE5CCB3FBABFC8BFBF0" bold="true" box="[765,783,1064,1088]" pageId="22" pageNumber="423">C</emphasis>
. Cross section of the muscular sac ensheathing penial seta.
<emphasis id="DDF8EA9B7940FFE5CBCAFBABFA12FBF0" bold="true" box="[1412,1430,1064,1088]" pageId="22" pageNumber="423">D</emphasis>
. Section through ental part of prostate.
<emphasis id="DDF8EA9B7940FFE5CC7EFBCFFDC5FBD4" bold="true" box="[560,577,1100,1124]" pageId="22" pageNumber="423">E</emphasis>
. Cross section of prostatic duct.
<emphasis id="DDF8EA9B7940FFE5CDEEFBCFFC2BFBD4" bold="true" box="[928,943,1100,1124]" pageId="22" pageNumber="423">F</emphasis>
. Dorsal view of the arrangement of the penial setae within IXXIII. Abbreviations: a, spermathecal ampulla, cv, commissural vessels, d, spermathecal duct, di, spermathecal diverticulum, e, egg, if female funnel, md, male duct, pc, penial seta, pr, prostate. From
<bibRefCitation id="8B1D4B787940FFE5CA56FB17FAA4FB1C" author="Omodeo, P. &amp; Coates, K. A." box="[1048,1312,1172,1196]" pageId="22" pageNumber="423" pagination="39 - 47" refId="ref24287" refString="Omodeo, P. &amp; Coates, K. A. (2001) New alluroidids (Annelida, Clitellata) from Guyana. Hydrobiologia, 463, 39 - 47. https: // doi. org / 10.1023 / A: 1013178902798" type="journal article" year="2001">Omodeo &amp; Coates (2001)</bibRefCitation>
, reproduced with permission.
</paragraph>
</caption>
<paragraph id="EF3336897940FFE5CE89FA99FA99FA55" blockId="22.[151,1437,1305,2043]" pageId="22" pageNumber="423">
Septum 3/4 incomplete, septa 4/512/13 fairly thick. Septal glands in V through VIII. Perienteric blood sinus commencing in IX. Nephridia avesiculate beginning in XI. Dorsal and ventral vessels with large diameters; commissural vessels in VXI tortuous with thin walls, diameter
<emphasis id="DDF8EA9B7940FFE5CD12FAE2FCF2FACA" box="[860,886,1377,1402]" italics="true" pageId="22" pageNumber="423">ca</emphasis>
17 µm; their dorsal loops in VVIII embedded in the septal glands; commissures in X and XI compressed and hidden by large gregarine gamonts; numerous capillaries beneath and among the epidermal cells, especially in the clitellar region, and among the glandular cells covering the prostates; diameter
<emphasis id="DDF8EA9B7940FFE5CF3EFA4FFE0EFA55" box="[368,394,1484,1509]" italics="true" pageId="22" pageNumber="423">ca</emphasis>
3.5 µm. Muscular fibres of the body wall ribbonshaped, 3 µm thick, 21 µm high.
</paragraph>
<paragraph id="EF3336897940FFE5CE89FA6CFC23F8DB" blockId="22.[151,1437,1305,2043]" pageId="22" pageNumber="423">
<materialsCitation id="5FE43CD47940FFE5CE89FA6CFC27F8DB" collectionCode="IX" pageId="22" pageNumber="423" specimenCount="2" specimenCount-egg="2">
Testes and seminal funnels in X; male ducts thin in XI and XII; in XIII becoming sinuous and filled with sperm, forming a highly convoluted epididymis, and finally connecting with the prostate at an intermediate point. Prostates long, convoluted, confined to XIII; with an inner layer of epithelial (glandular?) cells surrounded by a thick layer of muscle cells and covered by large, clubshaped gland cells which diminishes and then disappears near the outlet into the globular terminal chamber, which opens through as a small penial papilla. In each terminal chamber emerges a ribbonlike penial seta, whose proximal end lies in the anterior dorsal part of
<collectionCode id="899DAE4C7940FFE5CAC8F923FB2CF90A" box="[1158,1192,1696,1722]" pageId="22" pageNumber="423">IX</collectionCode>
: the two penial setae, each ensheathed in a muscular sac, run parallel near the middorsal line in
<collectionCode id="899DAE4C7940FFE5CDA6F947FB8CF96E" box="[1000,1032,1732,1758]" pageId="22" pageNumber="423">IX</collectionCode>
, X and XI; then in XII they diverge sidewards and ventralwards, arcing around the terminal chamber in XIII and exiting through it. Ovaries in XIII;
<specimenCount id="F98AFD007940FFE5CB3CF964FF3BF895" count="2" pageId="22" pageNumber="423" type="egg">two egg</specimenCount>
sacs arise near the female funnels and extending backwards to XVI; the histology of the egg sac resembling that of
<taxonomicName id="288C4D0A7940FFE5CEF8F8AEFE27F8F7" authorityName="Omodeo" authorityYear="1996" box="[182,419,1837,1863]" class="Clitellata" family="Alluroididae" genus="Kathrynella" kingdom="Animalia" order="Alluroidida" pageId="22" pageNumber="423" phylum="Annelida" rank="species" species="guyanae">
<emphasis id="DDF8EA9B7940FFE5CEF8F8AEFE27F8F7" box="[182,419,1837,1863]" italics="true" pageId="22" pageNumber="423">Kathrynella guyanae</emphasis>
</taxonomicName>
. Ovaries and egg sacs disappear in fully mature specimens leaving two or three free large ellipsoid eggs (420 by 420 by 300 µm) in the coelom of XV and XVI
</materialsCitation>
.
</paragraph>
<paragraph id="EF3336897940FFE5CE89F8F7FAC9F866" blockId="22.[151,1437,1305,2043]" pageId="22" pageNumber="423">Spermatheca with a thick, spindleshaped muscular duct that runs forward to the middle of VIII abutting on a very large, free, globular ampulla that almost fills the cavity of this segment. Near the insertion of the duct is attached a pedunculated diverticulum. Sperm contained in the diverticulum and in the ental part of the duct.</paragraph>
</subSubSection>
<subSubSection id="A79665027940FFE4CE89F862FBD8FF65" lastPageId="23" lastPageNumber="424" pageId="22" pageNumber="423" type="discussion">
<paragraph id="EF3336897940FFE4CE89F862FBD8FF65" blockId="22.[151,1437,1305,2043]" lastBlockId="23.[151,1436,151,249]" lastPageId="23" lastPageNumber="424" pageId="22" pageNumber="423">
<emphasis id="DDF8EA9B7940FFE5CE89F862FEB0F84B" bold="true" box="[199,308,2017,2043]" pageId="22" pageNumber="423">Remarks</emphasis>
.
<taxonomicName id="288C4D0A7940FFE5CF0DF862FE3EF84A" authorityName="Omodeo and Coates" authorityYear="2001" box="[323,442,2016,2042]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="22" pageNumber="423" phylum="Annelida" rank="species" species="donaldi">
<emphasis id="DDF8EA9B7940FFE5CF0DF862FE3EF84A" box="[323,442,2016,2042]" italics="true" pageId="22" pageNumber="423">B. donaldi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="288C4D0A7940FFE5CC1FF862FD70F84A" baseAuthorityName="Brinkhurst" baseAuthorityYear="1964" box="[593,756,2017,2042]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="22" pageNumber="423" phylum="Annelida" rank="species" species="americanus">
<emphasis id="DDF8EA9B7940FFE5CC1FF862FD70F84A" box="[593,756,2017,2042]" italics="true" pageId="22" pageNumber="423">B. americanus</emphasis>
</taxonomicName>
, the only described congener, in the form, size and peculiar insertion of the penial setae. Its spermatheca has a diverticulum and a much longer duct (correlated with the length of the penial setae) and opens one segment more anteriorly (
<bibRefCitation id="8B1D4B787941FFE4CD74FF38FBCBFF65" author="Omodeo, P. &amp; Coates, K. A." box="[826,1103,187,213]" pageId="23" pageNumber="424" pagination="39 - 47" refId="ref24287" refString="Omodeo, P. &amp; Coates, K. A. (2001) New alluroidids (Annelida, Clitellata) from Guyana. Hydrobiologia, 463, 39 - 47. https: // doi. org / 10.1023 / A: 1013178902798" type="journal article" year="2001">Omodeo &amp; Coates 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A79665027941FFE4CE89FF5DFC6AFF49" box="[199,1006,222,249]" pageId="23" pageNumber="424" type="distribution">
<paragraph id="EF3336897941FFE4CE89FF5DFC6AFF49" blockId="23.[151,1436,151,249]" box="[199,1006,222,249]" pageId="23" pageNumber="424">
<emphasis id="DDF8EA9B7941FFE4CE89FF5DFEDDFF48" bold="true" box="[199,345,222,248]" pageId="23" pageNumber="424">Distribution</emphasis>
.
<collectingCountry id="979B76197941FFE4CF29FF5CFE45FF49" box="[359,449,223,249]" name="Guyana" pageId="23" pageNumber="424">Guyana</collectingCountry>
, Kurupukari creek, flowing into
<collectingCountry id="979B76197941FFE4CD7EFF5CFC27FF49" box="[816,931,223,249]" name="Guyana" pageId="23" pageNumber="424">Essequibo</collectingCountry>
River.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,114 @@
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<mods:title id="F02D908299CCC88BA67292168F919280">A monograph of the Oligochaete family Alluroididae</mods:title>
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<subSubSection id="A79665027945FFE0CED9FDE9FD24FD34" box="[151,672,618,645]" pageId="19" pageNumber="420" type="nomenclature">
<paragraph id="EF3336897945FFE0CED9FDE9FD24FD34" blockId="19.[151,672,618,645]" box="[151,672,618,645]" pageId="19" pageNumber="420">
<heading id="B47B81E57945FFE0CED9FDE9FD24FD34" bold="true" box="[151,672,618,645]" fontSize="11" level="1" pageId="19" pageNumber="420" reason="1">
<emphasis id="DDF8EA9B7945FFE0CED9FDE9FD24FD34" bold="true" box="[151,672,618,645]" pageId="19" pageNumber="420">
Genus
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<emphasis id="DDF8EA9B7945FFE0CEA0FDE8FE44FD35" bold="true" box="[238,448,619,645]" italics="true" pageId="19" pageNumber="420">Barriejamiesonia</emphasis>
<bibRefCitation id="8B1D4B787945FFE0CF86FDE9FD24FD34" author="Ljungstrom, P. O." box="[456,672,618,644]" pageId="19" pageNumber="420" pagination="313 - 325" refId="ref23844" refString="Ljungstrom, P. O. (1971) New species of aquatic earthworms from Malawi and South Africa. Revue d'Ecologie et de Biologie du Sol, 8 (2), 313 - 325." type="journal article" year="1971">Ljungström 1971</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A79665027945FFE0CED9FD33FBACFD77" box="[151,1064,687,711]" pageId="19" pageNumber="420" type="reference_group">
<paragraph id="EF3336897945FFE0CED9FD33FBACFD77" blockId="19.[151,1064,687,711]" box="[151,1064,687,711]" pageId="19" pageNumber="420">
<treatmentCitationGroup id="CF9C11A77945FFE0CED9FD33FBACFD77" box="[151,1064,687,711]" pageId="19" pageNumber="420">
<treatmentCitation id="6E2D10987945FFE0CED9FD33FDB0FD77" author="Ljungstrom, P. O." box="[151,564,687,711]" page="321" pageId="19" pageNumber="420" year="1971">
<taxonomicName id="288C4D0A7945FFE0CED9FD33FDB0FD77" authority="Ljungstrom 1971: 321" authorityName="Ljungstrom" authorityPageNumber="321" authorityYear="1971" box="[151,564,687,711]" class="Clitellata" family="Alluroididae" genus="Barriejamiesonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alluroidida" pageId="19" pageNumber="420" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B7945FFE0CED9FD33FECCFD77" box="[151,328,688,711]" italics="true" pageId="19" pageNumber="420">Barriejamiesonia</emphasis>
<bibRefCitation id="8B1D4B787945FFE0CF01FD2CFDB0FD77" author="Ljungstrom, P. O." box="[335,564,687,711]" pageId="19" pageNumber="420" pagination="313 - 325" refId="ref23844" refString="Ljungstrom, P. O. (1971) New species of aquatic earthworms from Malawi and South Africa. Revue d'Ecologie et de Biologie du Sol, 8 (2), 313 - 325." type="journal article" year="1971">Ljungström 1971: 321</bibRefCitation>
</taxonomicName>
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. (Correction for
<taxonomicName id="288C4D0A7945FFE0CC93FD33FC03FD77" ID-CoL="84LRC" box="[733,903,688,711]" class="Clitellata" family="Alluroididae" genus="Barryjamiesonia" kingdom="Animalia" order="Alluroidida" pageId="19" pageNumber="420" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B7945FFE0CC93FD33FC03FD77" box="[733,903,688,711]" italics="true" pageId="19" pageNumber="420">Barryjamiesonia</emphasis>
</taxonomicName>
of Ljungström)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A79665027945FFE0CED9FD76FDA3FAC4" pageId="19" pageNumber="420" type="discussion">
<paragraph id="EF3336897945FFE0CED9FD76FB5DFC10" blockId="19.[151,1437,757,1396]" pageId="19" pageNumber="420">
<taxonomicName id="288C4D0A7945FFE0CED9FD76FEA1FCBF" box="[151,293,757,783]" class="Clitellata" family="Alluroididae" kingdom="Animalia" order="Alluroidida" pageId="19" pageNumber="411" phylum="Annelida" rank="subFamily" subFamily="Alluroidinae">Alluroidinae</taxonomicName>
in which the atria are divided into three regions: an ectal muscular spindleshaped chamber, a short narrow duct and an ental gland; vas deferens opening independently from the corresponding atrial duct and not thickened. Male pores ventral just in front of or slightly lateral to
<emphasis id="DDF8EA9B7945FFE0CDD8FCBDFC2EFCE7" box="[918,938,830,855]" italics="true" pageId="19" pageNumber="420">a.</emphasis>
Setae
<emphasis id="DDF8EA9B7945FFE0CDB4FCBEFB92FCE7" box="[1018,1046,829,855]" italics="true" pageId="19" pageNumber="420">ab</emphasis>
of XIII present but not modified. Genital setae absent. Spermathecal pores lateral on extreme posterior margin of VIII. First septum 5/6; 10/11 not attenuated. First visible nephridia in VII. Intestine commencing in XIV. Seminal vesicles present.
</paragraph>
<paragraph id="EF3336897945FFE0CE89FC2AFE67FAB8" blockId="19.[151,1437,757,1396]" pageId="19" pageNumber="420">
<emphasis id="DDF8EA9B7945FFE0CE89FC2AFEB0FC73" bold="true" box="[199,308,937,963]" pageId="19" pageNumber="420">Remarks</emphasis>
. Ljungström distinguished this genus from
<taxonomicName id="288C4D0A7945FFE0CD66FC2AFC18FC73" authorityName="Beddard" authorityYear="1894" box="[808,924,937,963]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="19" pageNumber="420" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B7945FFE0CD66FC2AFC18FC73" box="[808,924,937,963]" italics="true" pageId="19" pageNumber="420">Alluroides</emphasis>
</taxonomicName>
by the atria not being wide coiled tubes, the vasa deferentia opening ectally rather than entally; the atria being divided into a muscular duct and a glandular prostate; first septum 5/6 rather than 3/4; first nephridia in VII rather than in XIIIXIV. From
<taxonomicName id="288C4D0A7945FFE0CAEFFC72FAABFBBB" authorityName="Jamieson" authorityYear="1968" box="[1185,1327,1009,1035]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="19" pageNumber="420" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B7945FFE0CAEFFC72FAABFBBB" box="[1185,1327,1009,1035]" italics="true" pageId="19" pageNumber="420">Brinkhurstia</emphasis>
</taxonomicName>
in having a muscular rather than nonmuscular chamber; the prostate cells not surrounding the muscular duct; penial setae absent; male pores ventral rather than ventrolateral; setae
<emphasis id="DDF8EA9B7945FFE0CD65FBBAFCC3FBE3" box="[811,839,1081,1107]" italics="true" pageId="19" pageNumber="420">ab</emphasis>
present in XIII; spermathecal pores paired ventral on VII rather than?IX or further back. From
<taxonomicName id="288C4D0A7945FFE0CC3CFBDEFD5BFBC7" authorityName="Jamieson" authorityYear="1968" box="[626,735,1117,1143]" class="Clitellata" family="Alluroididae" genus="Standeria" kingdom="Animalia" order="Alluroidida" pageId="19" pageNumber="420" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B7945FFE0CC3CFBDEFD5BFBC7" box="[626,735,1117,1143]" italics="true" pageId="19" pageNumber="420">Standeria</emphasis>
</taxonomicName>
in having the atria divided into a glandular and muscular duct rather than a muscular duct lined internally with prostatic cells; vas deferens opening ectally rather than entally into the chamber; setae
<emphasis id="DDF8EA9B7945FFE0CF3FFB26FE09FB0F" box="[369,397,1189,1215]" italics="true" pageId="19" pageNumber="420">ab</emphasis>
present in XIII; spermathecal pores on VIII rather than IX; first septum 5/6 rather than 4/5; 10/11 not attenuated; genital setae absent; first nephridia in VII rather than XI; intestinal origin in XIV rather than XV; seminal vesicles present.
</paragraph>
<paragraph id="EF3336897945FFE0CE89FA92FDA3FAC4" blockId="19.[151,1437,757,1396]" pageId="19" pageNumber="420">Further differences from other alluroidine genera were that the atria are more like the prostates of the Ocnerodrilinae; the wide separation of the dorsal setae at 0.44 rather than 0.2 of the circumference; and the absence of a visible latero-muscular groove.</paragraph>
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<mods:title id="625817C293AA4F1D90DA3ADE3842FAE6">A monograph of the Oligochaete family Alluroididae</mods:title>
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SUBFAMILY
<taxonomicName id="288C4D0A795CFFF9CF09FC75FDAAFBA0" authority="Timm &amp; Martin, 2015" authorityName="Timm &amp; Martin" authorityYear="2015" box="[327,558,1014,1040]" class="Clitellata" family="Alluroididae" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="subFamily" subFamily="Alluroidinae">ALLUROIDINAE</taxonomicName>
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<typeStatus id="3037882B795CFFF9CED9FBBCFF50FBE9" box="[151,212,1087,1113]" pageId="10" pageNumber="411">Type</typeStatus>
genus:
<taxonomicName id="288C4D0A795CFFF9CF7BFBBDFDEDFBE8" ID-CoL="84L9X" authority="Beddard 1894" authorityName="Beddard" authorityYear="1894" box="[309,617,1086,1113]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CF7BFBBDFE37FBE9" bold="true" box="[309,435,1086,1113]" italics="true" pageId="10" pageNumber="411">Alluroides</emphasis>
<bibRefCitation id="8B1D4B78795CFFF9CFF4FBBCFDEDFBE8" author="Beddard, F. E." box="[442,617,1086,1113]" pageId="10" pageNumber="411" pagination="201 - 269" refId="ref22462" refString="Beddard, F. E. (1894) A contribution to our knowledge of the Oligochaeta of tropical eastern Africa. Quarterly Journal of Microscopical Science, 36, 201 - 269. https: // doi. org / 10.1242 / jcs. s 2 - 36.142.201" type="journal article" year="1894">Beddard 1894</bibRefCitation>
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Key to the genera of the
<taxonomicName id="288C4D0A795CFFF9CF86FB05FDE2FB10" box="[456,614,1158,1184]" class="Clitellata" family="Alluroididae" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="subFamily" subFamily="Alluroidinae">Alluroidinae</taxonomicName>
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<keyLead id="547D95BC795CFFF9CEEDFB57FC71FB5C" box="[163,1013,1236,1260]" pageId="10" pageNumber="411">1 Spermathecal pore(s) middorsal 2</keyLead>
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<paragraph id="EF333689795CFFF9CEBFFB7FFC71FAA4" blockId="10.[163,1013,1236,1300]" box="[241,1013,1276,1300]" pageId="10" pageNumber="411">
<keyLead id="547D95BC795CFFF9CEBFFB7FFC71FAA4" box="[241,1013,1276,1300]" pageId="10" pageNumber="411">Spermathecal pores paired lateral 4</keyLead>
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<keyStep id="54782E2C795CFFF9CEEDFACEFBC1FA3D" pageId="10" pageNumber="411">
<paragraph id="EF333689795CFFF9CEEDFACEFC71FAD5" blockId="10.[163,1093,1357,1421]" box="[163,1013,1357,1381]" pageId="10" pageNumber="411">
<keyLead id="547D95BC795CFFF9CEEDFACEFC71FAD5" box="[163,1013,1357,1381]" pageId="10" pageNumber="411">2 Testes in X 3</keyLead>
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<keyLead id="547D95BC795CFFF9CEBFFAF6FBC1FA3D" box="[241,1093,1397,1421]" pageId="10" pageNumber="411">
Testes in XI
<taxonomicName id="288C4D0A795CFFF9CDA6FAF6FBC1FA3D" box="[1000,1093,1397,1421]" class="Clitellata" family="Alluroididae" genus="Righiella" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CDA6FAF6FBC1FA3D" box="[1000,1093,1397,1421]" italics="true" pageId="10" pageNumber="411">Righiella</emphasis>
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3 Penial setae present
<taxonomicName id="288C4D0A795CFFF9CDA6FA46FBECFA6D" authority="Penial" authorityName="Penial" box="[1000,1128,1477,1501]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CDA6FA46FBECFA6D" box="[1000,1128,1477,1501]" italics="true" pageId="10" pageNumber="411">Brinkhurstia</emphasis>
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Penial setae absent
<taxonomicName id="288C4D0A795CFFF9CDA6FA6DFBD5F9B6" authorityName="Beddard" authorityYear="1894" box="[1000,1105,1518,1542]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CDA6FA6DFBD5F9B6" box="[1000,1105,1518,1542]" italics="true" pageId="10" pageNumber="411">Alluroides</emphasis>
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<keyLead id="547D95BC795CFFF9CEEDF9BDFBF8F9E6" box="[163,1148,1598,1622]" pageId="10" pageNumber="411">
4 Testes in X and XI
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<emphasis id="DDF8EA9B795CFFF9CDA6F9BDFBF8F9E6" box="[1000,1148,1598,1622]" italics="true" pageId="10" pageNumber="411">Lacandodrilus</emphasis>
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<paragraph id="EF333689795CFFF9CEBFF9E4FC71F9CE" blockId="10.[163,1148,1598,1663]" box="[241,1013,1638,1663]" pageId="10" pageNumber="411">
<keyLead id="547D95BC795CFFF9CEBFF9E4FC71F9CE" box="[241,1013,1638,1663]" pageId="10" pageNumber="411">Testes in X or XI 5</keyLead>
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<paragraph id="EF333689795CFFF9CEEDF934FC71F97F" blockId="10.[163,1121,1719,1783]" box="[163,1013,1719,1743]" pageId="10" pageNumber="411">
<keyLead id="547D95BC795CFFF9CEEDF934FC71F97F" box="[163,1013,1719,1743]" pageId="10" pageNumber="411">5 Testes in X 6</keyLead>
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<paragraph id="EF333689795CFFF9CEBFF963FBE5F947" blockId="10.[163,1121,1719,1783]" box="[241,1121,1759,1783]" pageId="10" pageNumber="411">
<keyLead id="547D95BC795CFFF9CEBFF963FBE5F947" box="[241,1121,1759,1783]" pageId="10" pageNumber="411">
Testes in XI
<taxonomicName id="288C4D0A795CFFF9CDA6F95CFBE5F947" authorityName="Omodeo" authorityYear="1996" box="[1000,1121,1759,1783]" class="Clitellata" family="Alluroididae" genus="Kathrynella" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CDA6F95CFBE5F947" box="[1000,1121,1759,1783]" italics="true" pageId="10" pageNumber="411">Kathrynella</emphasis>
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<keyStep id="54782E2C795CFFF9CEEDF8B3FBCFF871" pageId="10" pageNumber="411">
<keyLead id="547D95BC795CFFF9CEEDF8B3FB1DF8C0" pageId="10" pageNumber="411">
<paragraph id="EF333689795CFFF9CEEDF8B3FE38F8F8" blockId="10.[163,1177,1840,1985]" box="[163,444,1840,1864]" pageId="10" pageNumber="411">6 Genital setae absent</paragraph>
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Seminal vesicles present
<taxonomicName id="288C4D0A795CFFF9CDA6F8DAFB1DF8C0" authorityName="Ljungstrom" authorityYear="1971" box="[1000,1177,1881,1904]" class="Clitellata" family="Alluroididae" genus="Barriejamiesonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CDA6F8DAFB1DF8C0" box="[1000,1177,1881,1904]" italics="true" pageId="10" pageNumber="411">Barriejamiesonia</emphasis>
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<paragraph id="EF333689795CFFF9CEBFF803FE40F828" blockId="10.[163,1177,1840,1985]" box="[241,452,1920,1944]" pageId="10" pageNumber="411">Genital setae present</paragraph>
<paragraph id="EF333689795CFFF9CEBFF82BFBCFF871" blockId="10.[163,1177,1840,1985]" box="[241,1099,1960,1985]" pageId="10" pageNumber="411">
Seminal vesicles absent
<taxonomicName id="288C4D0A795CFFF9CDA6F82AFBCFF871" authorityName="Jamieson" authorityYear="1968" box="[1000,1099,1961,1985]" class="Clitellata" family="Alluroididae" genus="Standeria" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CDA6F82AFBCFF871" box="[1000,1099,1961,1985]" italics="true" pageId="10" pageNumber="411">Standeria</emphasis>
</taxonomicName>
</paragraph>
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<emphasis id="DDF8EA9B795DFFF8CED9FCD2FE8AFCDB" bold="true" box="[151,270,849,875]" pageId="11" pageNumber="412">TABLE 1.</emphasis>
The most significant characters distinguishing the genera of the
<taxonomicName id="288C4D0A795DFFF8CDF0FCD2FBC1FCDB" box="[958,1093,849,875]" class="Clitellata" family="Alluroididae" kingdom="Animalia" order="Alluroidida" pageId="11" pageNumber="411" phylum="Annelida" rank="subFamily" subFamily="Alluroidinae">Alluroidinae</taxonomicName>
(modified and expanded from
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).
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<taxonomicName id="288C4D0A795DFFF8CFD9FCF9FDACFC1F" authority="Beddard, 1894" authorityName="Beddard" authorityYear="1894" box="[407,552,890,943]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="11" pageNumber="412" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795DFFF8CFD9FCF9FE79FC23" box="[407,509,890,915]" italics="true" pageId="11" pageNumber="412">Alluroides</emphasis>
<bibRefCitation id="8B1D4B78795DFFF8CFD9FC15FDACFC1F" author="Beddard, F. E." box="[407,552,918,943]" pageId="11" pageNumber="412" pagination="201 - 269" refId="ref22462" refString="Beddard, F. E. (1894) A contribution to our knowledge of the Oligochaeta of tropical eastern Africa. Quarterly Journal of Microscopical Science, 36, 201 - 269. https: // doi. org / 10.1242 / jcs. s 2 - 36.142.201" type="journal article" year="1894">Beddard, 1894</bibRefCitation>
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</th>
<th id="126D5DB7795D000CCC3CFCF9FC90FC07" box="[626,788,890,951]" gridcol="2" gridrow="0" pageId="11" pageNumber="412">
<taxonomicName id="288C4D0A795DFFF8CC3CFCF9FC8FFC1F" authority="Jamieson, 1968" authorityName="Jamieson" authorityYear="1968" box="[626,779,890,943]" class="Clitellata" family="Alluroididae" genus="Standeria" kingdom="Animalia" order="Alluroidida" pageId="11" pageNumber="412" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795DFFF8CC3CFCF9FD56FC23" box="[626,722,890,915]" italics="true" pageId="11" pageNumber="412">Standeria</emphasis>
<bibRefCitation id="8B1D4B78795DFFF8CC3CFC15FC8FFC1F" author="Jamieson, B. G. M." box="[626,779,918,943]" pageId="11" pageNumber="412" pagination="55 - 86" refId="ref23501" refString="Jamieson, B. G. M. (1968) A taxonometric investigation of the Alluroididae (Oligochaeta). Journal of Zoology, 155, 55 - 86. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03029. x" type="journal article" year="1968">Jamieson, 1968</bibRefCitation>
</taxonomicName>
</th>
<th id="126D5DB7795D000CCD70FCF9FC7CFC07" box="[830,1016,890,951]" gridcol="3" gridrow="0" pageId="11" pageNumber="412">
<taxonomicName id="288C4D0A795DFFF8CD70FCF9FC53FC1F" authority="Jamieson, 1968" authorityName="Jamieson" authorityYear="1968" box="[830,983,890,943]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="11" pageNumber="412" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795DFFF8CD70FCF9FC3FFC23" box="[830,955,890,915]" italics="true" pageId="11" pageNumber="412">Brinkhurstia</emphasis>
<bibRefCitation id="8B1D4B78795DFFF8CD70FC15FC53FC1F" author="Jamieson, B. G. M." box="[830,983,918,943]" pageId="11" pageNumber="412" pagination="55 - 86" refId="ref23501" refString="Jamieson, B. G. M. (1968) A taxonometric investigation of the Alluroididae (Oligochaeta). Journal of Zoology, 155, 55 - 86. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03029. x" type="journal article" year="1968">Jamieson, 1968</bibRefCitation>
</taxonomicName>
</th>
<th id="126D5DB7795D000CCA64FCF9FAA5FC07" box="[1066,1313,890,951]" gridcol="4" gridrow="0" pageId="11" pageNumber="412">
<taxonomicName id="288C4D0A795DFFF8CA64FCF9FAA5FC1F" authority="Omdeo &amp; Coates, 2001" authorityName="Omodeo &amp; Coates" authorityYear="2001" box="[1066,1313,890,943]" class="Clitellata" family="Alluroididae" genus="Righiella" kingdom="Animalia" order="Alluroidida" pageId="11" pageNumber="412" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795DFFF8CA64FCF9FB01FC23" box="[1066,1157,890,915]" italics="true" pageId="11" pageNumber="412">Righiella</emphasis>
Omdeo &amp; Coates, 2001
</taxonomicName>
</th>
<th id="126D5DB7795D000CCB0EFCF9FA69FC07" box="[1344,1517,890,951]" gridcol="5" gridrow="0" pageId="11" pageNumber="412">
<taxonomicName id="288C4D0A795DFFF8CB0EFCF9FA57FC1F" authority="Omodeo, 1996" authorityName="Omodeo" authorityYear="1996" box="[1344,1491,890,943]" class="Clitellata" family="Alluroididae" genus="Kathrynella" kingdom="Animalia" order="Alluroidida" pageId="11" pageNumber="412" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795DFFF8CB0EFCF9FA32FC23" box="[1344,1462,890,915]" italics="true" pageId="11" pageNumber="412">Kathrynella</emphasis>
<bibRefCitation id="8B1D4B78795DFFF8CB0EFC15FA57FC1F" author="Omodeo, P." box="[1344,1491,918,943]" pageId="11" pageNumber="412" pagination="11 - 15" refId="ref24252" refString="Omodeo, P. (1996) Kathrynella, a new oligochaete genus from Guyana. Hydrobiologia, 334. 11 - 15. https: // doi. org / 10.1007 / BF 00017349" type="journal article" year="1996">Omodeo, 1996</bibRefCitation>
</taxonomicName>
</th>
<th id="126D5DB7795D000CC849FCF9F93EFC07" box="[1543,1722,890,951]" gridcol="6" gridrow="0" pageId="11" pageNumber="412">
<taxonomicName id="288C4D0A795DFFF8C849FCF9F93EFC1F" authority="Ljungstrom, 1971" authorityName="Ljungstrom" authorityYear="1971" box="[1543,1722,890,943]" class="Clitellata" family="Alluroididae" genus="Barriejamiesonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alluroidida" pageId="11" pageNumber="412" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795DFFF8C849FCF9F930FC23" box="[1543,1716,890,915]" italics="true" pageId="11" pageNumber="412">Barriejamiesonia</emphasis>
<bibRefCitation id="8B1D4B78795DFFF8C849FC15F93EFC1F" author="Ljungstrom, P. O." box="[1543,1722,918,943]" pageId="11" pageNumber="412" pagination="313 - 325" refId="ref23844" refString="Ljungstrom, P. O. (1971) New species of aquatic earthworms from Malawi and South Africa. Revue d'Ecologie et de Biologie du Sol, 8 (2), 313 - 325." type="journal article" year="1971">Ljungström, 1971</bibRefCitation>
</taxonomicName>
</th>
<th id="126D5DB7795D000CC8B6FCF9F867FC07" box="[1784,2019,890,951]" gridcol="7" gridrow="0" pageId="11" pageNumber="412">
<taxonomicName id="288C4D0A795DFFF8C8B6FCF9F867FC07" authority="Fragoso &amp; Rojas, 2023" authorityName="Fragoso &amp; Rojas" authorityYear="2023" box="[1784,2019,890,951]" class="Clitellata" family="Alluroididae" genus="Lacandodrilus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Alluroidida" pageId="11" pageNumber="412" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795DFFF8C8B6FCF9F808FC23" box="[1784,1932,890,915]" italics="true" pageId="11" pageNumber="412">Lacandodrilus</emphasis>
<bibRefCitation id="8B1D4B78795DFFF8C8B6FC1DF867FC07" author="Fragoso, C. &amp; Rojas, P." box="[1784,2019,926,951]" pageId="11" pageNumber="412" pagination="136 - 156" refId="ref23321" refString="Fragoso, C., &amp; Rojas, P. (2023) A new semiaquatic worm (Annelida, Oligochaeta) from southeastern Mexican tropical rain forests. Zootaxa, 5255 (1), 136 - 156. https: // doi. org / 10.11646 / zootaxa. 5255.1.16" type="journal article" year="2023">Fragoso &amp; Rojas, 2023</bibRefCitation>
</taxonomicName>
</th>
</tr>
<tr id="51BC34CB795D000CCEECFC44F867FC50" box="[162,2019,967,992]" gridrow="1" pageId="11" pageNumber="412">
<th id="126D5DB7795D000CCEECFC44FEFCFC50" box="[162,376,967,992]" gridcol="0" gridrow="1" pageId="11" pageNumber="412">Male pores</th>
<td id="126D5DB7795D000CCFD9FC44FDC2FC50" box="[407,582,967,992]" gridcol="1" gridrow="1" pageId="11" pageNumber="412">XIII</td>
<td id="126D5DB7795D000CCC3CFC44FC90FC50" box="[626,788,967,992]" gridcol="2" gridrow="1" pageId="11" pageNumber="412">XIII</td>
<td id="126D5DB7795D000CCD70FC44FC7CFC50" box="[830,1016,967,992]" gridcol="3" gridrow="1" pageId="11" pageNumber="412">XIII</td>
<td id="126D5DB7795D000CCA64FC44FAA5FC50" box="[1066,1313,967,992]" gridcol="4" gridrow="1" pageId="11" pageNumber="412">XIII</td>
<td id="126D5DB7795D000CCB0EFC44FA69FC50" box="[1344,1517,967,992]" gridcol="5" gridrow="1" pageId="11" pageNumber="412">XIV</td>
<td id="126D5DB7795D000CC849FC44F93EFC50" box="[1543,1722,967,992]" gridcol="6" gridrow="1" pageId="11" pageNumber="412">XIII</td>
<td id="126D5DB7795D000CC8B6FC44F867FC50" box="[1784,2019,967,992]" gridcol="7" gridrow="1" pageId="11" pageNumber="412">XIII</td>
</tr>
<tr id="51BC34CB795D000CCEECFC6CF867FB94" box="[162,2019,1007,1060]" gridrow="2" pageId="11" pageNumber="412">
<th id="126D5DB7795D000CCEECFC6CFEFCFB94" box="[162,376,1007,1060]" gridcol="0" gridrow="2" pageId="11" pageNumber="412">Spermathecal pores</th>
<td id="126D5DB7795D000CCFD9FC6CFDC2FB94" box="[407,582,1007,1060]" gridcol="1" gridrow="2" pageId="11" pageNumber="412">Mid-dorsal, some or all of VI-VIII</td>
<td id="126D5DB7795D000CCC3CFC6CFC90FB94" box="[626,788,1007,1060]" gridcol="2" gridrow="2" pageId="11" pageNumber="412">Lateral 8/9</td>
<td id="126D5DB7795D000CCD70FC6CFC7CFB94" box="[830,1016,1007,1060]" gridcol="3" gridrow="2" pageId="11" pageNumber="412">Mid-dorsal 7/8, IX</td>
<td id="126D5DB7795D000CCA64FC6CFAA5FB94" box="[1066,1313,1007,1060]" gridcol="4" gridrow="2" pageId="11" pageNumber="412">Mid-dorsal 7/8</td>
<td id="126D5DB7795D000CCB0EFC6CFA69FB94" box="[1344,1517,1007,1060]" gridcol="5" gridrow="2" pageId="11" pageNumber="412">Lateral 6/7</td>
<td id="126D5DB7795D000CC849FC6CF93EFB94" box="[1543,1722,1007,1060]" gridcol="6" gridrow="2" pageId="11" pageNumber="412">Lateral 8/9</td>
<td id="126D5DB7795D000CC8B6FC6CF867FB94" box="[1784,2019,1007,1060]" gridcol="7" gridrow="2" pageId="11" pageNumber="412">Lateral 9/10</td>
</tr>
<tr id="51BC34CB795D000CCEECFBB0F867FBFC" box="[162,2019,1075,1100]" gridrow="3" pageId="11" pageNumber="412">
<th id="126D5DB7795D000CCEECFBB0FEFCFBFC" box="[162,376,1075,1100]" gridcol="0" gridrow="3" pageId="11" pageNumber="412">Testes in</th>
<td id="126D5DB7795D000CCFD9FBB0FDC2FBFC" box="[407,582,1075,1100]" gridcol="1" gridrow="3" pageId="11" pageNumber="412">X</td>
<td id="126D5DB7795D000CCC3CFBB0FC90FBFC" box="[626,788,1075,1100]" gridcol="2" gridrow="3" pageId="11" pageNumber="412">X</td>
<td id="126D5DB7795D000CCD70FBB0FC7CFBFC" box="[830,1016,1075,1100]" gridcol="3" gridrow="3" pageId="11" pageNumber="412">X</td>
<td id="126D5DB7795D000CCA64FBB0FAA5FBFC" box="[1066,1313,1075,1100]" gridcol="4" gridrow="3" pageId="11" pageNumber="412">XI</td>
<td id="126D5DB7795D000CCB0EFBB0FA69FBFC" box="[1344,1517,1075,1100]" gridcol="5" gridrow="3" pageId="11" pageNumber="412">XI</td>
<td id="126D5DB7795D000CC849FBB0F93EFBFC" box="[1543,1722,1075,1100]" gridcol="6" gridrow="3" pageId="11" pageNumber="412">X</td>
<td id="126D5DB7795D000CC8B6FBB0F867FBFC" box="[1784,2019,1075,1100]" gridcol="7" gridrow="3" pageId="11" pageNumber="412">X and XI</td>
</tr>
<tr id="51BC34CB795D000CCEECFBD8F867FB20" box="[162,2019,1115,1168]" gridrow="4" pageId="11" pageNumber="412">
<th id="126D5DB7795D000CCEECFBD8FEFCFB20" box="[162,376,1115,1168]" gridcol="0" gridrow="4" pageId="11" pageNumber="412">Male ducts insertion</th>
<td id="126D5DB7795D000CCFD9FBD8FDC2FB20" box="[407,582,1115,1168]" gridcol="1" gridrow="4" pageId="11" pageNumber="412">In ectal terminal chamber</td>
<td id="126D5DB7795D000CCC3CFBD8FC90FB20" box="[626,788,1115,1168]" gridcol="2" gridrow="4" pageId="11" pageNumber="412">In ectal terminal chamber</td>
<td id="126D5DB7795D000CCD70FBD8FC7CFB20" box="[830,1016,1115,1168]" gridcol="3" gridrow="4" pageId="11" pageNumber="412">In ectal terminal chamber</td>
<td id="126D5DB7795D000CCA64FBD8FAA5FB20" box="[1066,1313,1115,1168]" gridcol="4" gridrow="4" pageId="11" pageNumber="412">On the ental tip of prostates</td>
<td id="126D5DB7795D000CCB0EFBD8FA69FB20" box="[1344,1517,1115,1168]" gridcol="5" gridrow="4" pageId="11" pageNumber="412">On the ental tip of prostates</td>
<td id="126D5DB7795D000CC849FBD8F93EFB20" box="[1543,1722,1115,1168]" gridcol="6" gridrow="4" pageId="11" pageNumber="412">In ectal terminal chamber</td>
<td id="126D5DB7795D000CC8B6FBD8F867FB20" box="[1784,2019,1115,1168]" gridcol="7" gridrow="4" pageId="11" pageNumber="412">Not seen</td>
</tr>
<tr id="51BC34CB795D000CCEECFB1CF867FB08" box="[162,2019,1183,1208]" gridrow="5" pageId="11" pageNumber="412">
<th id="126D5DB7795D000CCEECFB1CFEFCFB08" box="[162,376,1183,1208]" gridcol="0" gridrow="5" pageId="11" pageNumber="412">Penial setae</th>
<td id="126D5DB7795D000CCFD9FB1CFDC2FB08" box="[407,582,1183,1208]" gridcol="1" gridrow="5" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CCC3CFB1CFC90FB08" box="[626,788,1183,1208]" gridcol="2" gridrow="5" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CCD70FB1CFC7CFB08" box="[830,1016,1183,1208]" gridcol="3" gridrow="5" pageId="11" pageNumber="412">yes</td>
<td id="126D5DB7795D000CCA64FB1CFAA5FB08" box="[1066,1313,1183,1208]" gridcol="4" gridrow="5" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CCB0EFB1CFA69FB08" box="[1344,1517,1183,1208]" gridcol="5" gridrow="5" pageId="11" pageNumber="412">yes</td>
<td id="126D5DB7795D000CC849FB1CF93EFB08" box="[1543,1722,1183,1208]" gridcol="6" gridrow="5" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CC8B6FB1CF867FB08" box="[1784,2019,1183,1208]" gridcol="7" gridrow="5" pageId="11" pageNumber="412">no</td>
</tr>
<tr id="51BC34CB795D000CCEECFB44F867FB50" box="[162,2019,1223,1248]" gridrow="6" pageId="11" pageNumber="412">
<th id="126D5DB7795D000CCEECFB44FEFCFB50" box="[162,376,1223,1248]" gridcol="0" gridrow="6" pageId="11" pageNumber="412">Secondary annulation</th>
<td id="126D5DB7795D000CCFD9FB44FDC2FB50" box="[407,582,1223,1248]" gridcol="1" gridrow="6" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CCC3CFB44FC90FB50" box="[626,788,1223,1248]" gridcol="2" gridrow="6" pageId="11" pageNumber="412">yes</td>
<td id="126D5DB7795D000CCD70FB44FC7CFB50" box="[830,1016,1223,1248]" gridcol="3" gridrow="6" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CCA64FB44FAA5FB50" box="[1066,1313,1223,1248]" gridcol="4" gridrow="6" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CCB0EFB44FA69FB50" box="[1344,1517,1223,1248]" gridcol="5" gridrow="6" pageId="11" pageNumber="412">yes</td>
<td id="126D5DB7795D000CC849FB44F93EFB50" box="[1543,1722,1223,1248]" gridcol="6" gridrow="6" pageId="11" pageNumber="412">VIII</td>
<td id="126D5DB7795D000CC8B6FB44F867FB50" box="[1784,2019,1223,1248]" gridcol="7" gridrow="6" pageId="11" pageNumber="412">yes</td>
</tr>
<tr id="51BC34CB795D000CCEECFB6CF867FAB8" box="[162,2019,1263,1288]" gridrow="7" pageId="11" pageNumber="412">
<th id="126D5DB7795D000CCEECFB6CFEFCFAB8" box="[162,376,1263,1288]" gridcol="0" gridrow="7" pageId="11" pageNumber="412">Genital papillae</th>
<td id="126D5DB7795D000CCFD9FB6CFDC2FAB8" box="[407,582,1263,1288]" gridcol="1" gridrow="7" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CCC3CFB6CFC90FAB8" box="[626,788,1263,1288]" gridcol="2" gridrow="7" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CCD70FB6CFC7CFAB8" box="[830,1016,1263,1288]" gridcol="3" gridrow="7" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CCA64FB6CFAA5FAB8" box="[1066,1313,1263,1288]" gridcol="4" gridrow="7" pageId="11" pageNumber="412">no</td>
<td id="126D5DB7795D000CCB0EFB6CFA69FAB8" box="[1344,1517,1263,1288]" gridcol="5" gridrow="7" pageId="11" pageNumber="412">yes</td>
<td id="126D5DB7795D000CC849FB6CF93EFAB8" box="[1543,1722,1263,1288]" gridcol="6" gridrow="7" pageId="11" pageNumber="412">yes</td>
<td id="126D5DB7795D000CC8B6FB6CF867FAB8" box="[1784,2019,1263,1288]" gridcol="7" gridrow="7" pageId="11" pageNumber="412">no</td>
</tr>
<tr id="51BC34CB795D000CCEECFA9BF867FAFC" box="[162,2019,1304,1356]" gridrow="8" pageId="11" pageNumber="412">
<th id="126D5DB7795D000CCEECFA9BFEFCFAFC" box="[162,376,1304,1356]" gridcol="0" gridrow="8" pageId="11" pageNumber="412">Geographical Distribution</th>
<td id="126D5DB7795D000CCFD9FA9BFDC2FAFC" box="[407,582,1304,1356]" gridcol="1" gridrow="8" pageId="11" pageNumber="412">Central Africa</td>
<td id="126D5DB7795D000CCC3CFA9BFC90FAFC" box="[626,788,1304,1356]" gridcol="2" gridrow="8" pageId="11" pageNumber="412">Central Africa</td>
<td id="126D5DB7795D000CCD70FA9BFC7CFAFC" box="[830,1016,1304,1356]" gridcol="3" gridrow="8" pageId="11" pageNumber="412">South America</td>
<td id="126D5DB7795D000CCA64FA9BFAA5FAFC" box="[1066,1313,1304,1356]" gridcol="4" gridrow="8" pageId="11" pageNumber="412">Guyana</td>
<td id="126D5DB7795D000CCB0EFA9BFA69FAFC" box="[1344,1517,1304,1356]" gridcol="5" gridrow="8" pageId="11" pageNumber="412">Guyana</td>
<td id="126D5DB7795D000CC849FA9BF93EFAFC" box="[1543,1722,1304,1356]" gridcol="6" gridrow="8" pageId="11" pageNumber="412">Central Africa</td>
<td id="126D5DB7795D000CC8B6FA9BF867FAFC" box="[1784,2019,1304,1356]" gridcol="7" gridrow="8" pageId="11" pageNumber="412">Mexico</td>
</tr>
</table>
</paragraph>
<paragraph id="EF333689795AFFFFCED9FF14FE78FD8D" blockId="12.[151,1437,151,645]" pageId="12" pageNumber="413">
A longitudinal lateral line (lateromuscular groove) present on each side between the ventral and dorsal setal couples, or absent. Dorsal median intersetal distance (
<emphasis id="DDF8EA9B795AFFFFCCDDFF38FD2BFF65" box="[659,687,187,213]" italics="true" pageId="12" pageNumber="413">dd</emphasis>
:u),
<emphasis id="DDF8EA9B795AFFFFCC93FF38FD73FF64" box="[733,759,187,212]" italics="true" pageId="12" pageNumber="413">ca</emphasis>
0.2 to 0.5 of the circumference of the body; genital setae or penial setae present or absent. Clitellum commencing in XII or XIII, maximally extending for five segments. Male pores ventral to lateral in XIII or exceptionally XIV. Female pores at or near the anterior border of XIV in line with the male pores or nearly so or exceptionally near the ventral midline. Spermathecal pores lateral to dorsal but never in line with the male pores, paired or dorsal median and single in VIIX, maximally in three of these segments. Dorsal pores absent. Subneural blood vessel absent. Nephridia avesiculate. Prostates (atria) tubular, lobulate or bulbous, receiving the male ducts, or discharging with the latter but separately from them into a terminal chamber; internal epithelium surrounded by a muscular sheath outside which prostatic cells are usually present. Gizzards absent. Proandric, testes in X only; or less commonly holandric, testes in X and XI, or metandric, testes in X, Seminal vesicles projecting into XI, or more elongate, or absent. Ovisacs extending posteriorly through several segments; eggs large and yolky.
</paragraph>
</subSubSection>
</keyLead>
</keyStep>
</key>
</subSubSection>
<subSubSection id="A7966502795AFFFFCE89FDC5FB73FD35" pageId="12" pageNumber="413" type="distribution">
<paragraph id="EF333689795AFFFFCE89FDC5FB73FD35" blockId="12.[151,1437,151,645]" pageId="12" pageNumber="413">
<emphasis id="DDF8EA9B795AFFFFCE89FDC5FEDDFDD0" bold="true" box="[199,345,582,608]" pageId="12" pageNumber="413">Distribution</emphasis>
.
<collectingCountry id="979B7619795AFFFFCF28FDC4FE5CFDD1" box="[358,472,583,609]" name="Colombia" pageId="12" pageNumber="413">Colombia</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCFACFDC4FDAEFDD1" box="[482,554,583,609]" name="Brazil" pageId="12" pageNumber="413">Brazil</collectingCountry>
; Saint Lucia (Caribbean);
<collectingCountry id="979B7619795AFFFFCD04FDC4FC27FDD1" box="[842,931,583,609]" name="Mexico" pageId="12" pageNumber="413">Mexico</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCDE3FDC4FB91FDD1" box="[941,1045,583,609]" name="Tanzania" pageId="12" pageNumber="413">Tanzania</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCA50FDC4FBFDFDD1" box="[1054,1145,583,609]" name="Uganda" pageId="12" pageNumber="413">Uganda</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCACDFDC4FB7AFDD1" box="[1155,1278,583,609]" name="Zimbabwe" pageId="12" pageNumber="413">Zimbabwe</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCB46FDC4FA1CFDD1" box="[1288,1432,583,609]" name="South Africa" pageId="12" pageNumber="413">South Africa</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCED9FDE8FF7EFD35" box="[151,250,619,645]" name="Ethiopia" pageId="12" pageNumber="413">Ethiopia</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCF4DFDE8FEF2FD35" box="[259,374,619,645]" name="Argentina" pageId="12" pageNumber="413">Argentina</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCFCCFDE8FE59FD35" box="[386,477,619,645]" name="Guyana" pageId="12" pageNumber="413">Guyana</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCFA9FDE8FDA2FD35" box="[487,550,619,645]" name="Chad" pageId="12" pageNumber="413">Chad</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCC7FFDE8FD19FD35" box="[561,669,619,645]" name="Paraguay" pageId="12" pageNumber="413">Paraguay</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCCE6FDE8FD71FD35" box="[680,757,619,645]" name="Kenya" pageId="12" pageNumber="413">Kenya</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCD4EFDE8FB0FFD35" box="[768,1163,619,645]" name="Democratic Republic of the Congo" pageId="12" pageNumber="413">Democratic Republic of the Congo</collectingCountry>
;
<collectingCountry id="979B7619795AFFFFCAD9FDE8FB77FD35" box="[1175,1267,619,645]" name="Panama" pageId="12" pageNumber="413">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,336 @@
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<subSubSection id="A7966502795FFFFACED9FB4DFC71FB58" box="[151,1013,1230,1257]" pageId="9" pageNumber="410" type="nomenclature">
<paragraph id="EF333689795FFFFACED9FB4DFC71FB58" blockId="9.[151,1013,1230,1257]" box="[151,1013,1230,1257]" pageId="9" pageNumber="410">
<heading id="B47B81E5795FFFFACED9FB4DFC71FB58" bold="true" box="[151,1013,1230,1257]" fontSize="11" level="1" pageId="9" pageNumber="410" reason="1">
<emphasis id="DDF8EA9B795FFFFACED9FB4DFC71FB58" bold="true" box="[151,1013,1230,1257]" pageId="9" pageNumber="410">
FAMILY
<taxonomicName id="288C4D0A795FFFFACF40FB4DFD4AFB58" ID-CoL="84JV9" authority="Michaelsen 1900" authorityName="Michaelsen" authorityYear="1900" box="[270,718,1230,1256]" class="Clitellata" family="Alluroididae" kingdom="Animalia" order="Alluroidida" pageId="9" pageNumber="410" phylum="Annelida" rank="family">
ALLUROIDIDAE
<bibRefCitation id="8B1D4B78795FFFFACFB3FB4DFD4AFB58" author="Michaelsen, W." box="[509,718,1230,1256]" pageId="9" pageNumber="410" refId="ref23961" refString="Michaelsen, W. (1900) Oligochaeta. Das Tierreich. Vol. 10. Lief. Friedlander und Sohn, Berlin, 575 pp. https: // doi. org / 10.5962 / bhl. title. 11605" type="book" year="1900">Michaelsen 1900</bibRefCitation>
</taxonomicName>
, emend.
<bibRefCitation id="8B1D4B78795FFFFACD73FB4DFC71FB58" author="Jamieson, B. G. M." box="[829,1013,1230,1256]" pageId="9" pageNumber="410" pagination="55 - 86" refId="ref23501" refString="Jamieson, B. G. M. (1968) A taxonometric investigation of the Alluroididae (Oligochaeta). Journal of Zoology, 155, 55 - 86. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03029. x" type="journal article" year="1968">Jamieson 1968</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A7966502795FFFFACED9FA94FE40F8B5" pageId="9" pageNumber="410" type="type_taxon">
<paragraph id="EF333689795FFFFACED9FA94FDEDFA80" blockId="9.[151,617,1302,1329]" box="[151,617,1302,1329]" pageId="9" pageNumber="410">
<emphasis id="DDF8EA9B795FFFFACED9FA94FDEDFA80" bold="true" box="[151,617,1302,1329]" pageId="9" pageNumber="410">
<typeStatus id="3037882B795FFFFACED9FA94FF50FA81" box="[151,212,1303,1329]" pageId="9" pageNumber="410">Type</typeStatus>
genus:
<taxonomicName id="288C4D0A795FFFFACF7BFA95FDEDFA80" ID-CoL="84L9X" authority="Beddard 1894" authorityName="Beddard" authorityYear="1894" box="[309,617,1302,1329]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="9" pageNumber="410" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795FFFFACF7BFA95FE37FA81" bold="true" box="[309,435,1302,1329]" italics="true" pageId="9" pageNumber="410">Alluroides</emphasis>
<bibRefCitation id="8B1D4B78795FFFFACFF4FA94FDEDFA80" author="Beddard, F. E." box="[442,617,1302,1329]" pageId="9" pageNumber="410" pagination="201 - 269" refId="ref22462" refString="Beddard, F. E. (1894) A contribution to our knowledge of the Oligochaeta of tropical eastern Africa. Quarterly Journal of Microscopical Science, 36, 201 - 269. https: // doi. org / 10.1242 / jcs. s 2 - 36.142.201" type="journal article" year="1894">Beddard 1894</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="EF333689795FFFFACED9FADCFE40F8B5" blockId="9.[151,1437,1375,2013]" pageId="9" pageNumber="410">
Setae singlepointed, sigmoid, 4 pairs per segment commencing on II; genital or penial setae sometimes present. Lateral line (lateromuscular groove) present or absent. Clitellum one cell thick, commencing on XI, XII or XIII and occupying 26 segments. Male pores one pair, intraclitellar, either at the anterior border of XIII, or in its setal arc, or in XIV. Female pores one pair, at the anterior border, or more posterior in XIV. Spermathecal pores on one to three of segments VIIX, at or near their anterior margins, lateral to dorsal, paired or single. Dorsal pores absent. Alimentary canal with or without oesophageal gizzards; lacking diverticula or other appendages. Nephridia holonephridia. Testes in X or in X and XI or in XI only (proandric, holandric or metandric). Seminal vesicles, when present, projecting into XI only or extending posteriorly through several segments as in microdriles. Prostate glands (euprostates or atria) either discharging separately from the male pores or through the latter, in some cases receiving the vasa deferentia entally. Ovaries anterior in XIII. Mature oocytes large and yolky, in ovisacs which extend posteriorly through several segments. Spermathecae without diverticula, sometimes (
<taxonomicName id="288C4D0A795FFFFACAE6F944FA17F951" ID-CoL="NBD9" authorityName="Omodeo &amp; Coates" authorityYear="2001" box="[1192,1427,1735,1761]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="9" pageNumber="410" phylum="Annelida" rank="species" species="donaldi">
<emphasis id="DDF8EA9B795FFFFACAE6F944FA17F951" box="[1192,1427,1735,1761]" italics="true" pageId="9" pageNumber="410">Brinkhurstia donaldi</emphasis>
</taxonomicName>
) with a single diverticulum.
</paragraph>
</subSubSection>
<subSubSection id="A7966502795FFFF9CE89F88DFE25FC15" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410" type="distribution">
<paragraph id="EF333689795FFFFACE89F88DFD3DF8C1" blockId="9.[151,1437,1375,2013]" pageId="9" pageNumber="410">
<emphasis id="DDF8EA9B795FFFFACE89F88DFEDDF898" bold="true" box="[199,345,1806,1832]" pageId="9" pageNumber="410">Distribution</emphasis>
.
<taxonomicName id="288C4D0A795FFFFACF25F88CFE7FF899" box="[363,507,1807,1833]" class="Clitellata" family="Alluroididae" kingdom="Animalia" order="Alluroidida" pageId="9" pageNumber="411" phylum="Annelida" rank="subFamily" subFamily="Alluroidinae">Alluroidinae</taxonomicName>
,
<collectingCountry id="979B7619795FFFFACC42F88CFDFAF899" box="[524,638,1807,1833]" name="Colombia" pageId="9" pageNumber="410">Colombia</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACCC1F88CFD53F899" box="[655,727,1807,1833]" name="Brazil" pageId="9" pageNumber="410">Brazil</collectingCountry>
; Saint Lucia;
<collectingCountry id="979B7619795FFFFACDCFF88CFC5EF899" box="[897,986,1807,1833]" name="Mexico" pageId="9" pageNumber="410">Mexico</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACDA5F88CFBD7F899" box="[1003,1107,1807,1833]" name="Tanzania" pageId="9" pageNumber="410">Tanzania</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACA2AF88CFB3BF899" box="[1124,1215,1807,1833]" name="Uganda" pageId="9" pageNumber="410">Uganda</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACA81F88CFACEF899" box="[1231,1354,1807,1833]" name="Zimbabwe" pageId="9" pageNumber="410">Zimbabwe</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACB15F88CFF66F8FD" name="South Africa" pageId="9" pageNumber="410">South Africa</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACEBAF8B0FED3F8FD" box="[244,343,1843,1869]" name="Ethiopia" pageId="9" pageNumber="410">Ethiopia</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACF29F8B0FE5EF8FD" box="[359,474,1843,1869]" name="Argentina" pageId="9" pageNumber="410">Argentina</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACFA2F8B0FDC3F8FD" box="[492,583,1843,1869]" name="Guyana" pageId="9" pageNumber="410">Guyana</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACC16F8B0FD13F8FD" box="[600,663,1843,1869]" name="Chad" pageId="9" pageNumber="410">Chad</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACCE7F8B0FC91F8FD" box="[681,789,1843,1869]" name="Paraguay" pageId="9" pageNumber="410">Paraguay</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACD68F8B0FCF7F8FD" box="[806,883,1843,1869]" name="Kenya" pageId="9" pageNumber="410">Kenya</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACDCBF8B0FAAEF8FD" box="[901,1322,1843,1869]" name="Democratic Republic of the Congo" pageId="9" pageNumber="410">Democratic Republic of the Congo</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACB73F8B0FA1DF8FD" box="[1341,1433,1843,1869]" name="Panama" pageId="9" pageNumber="410">Panama</collectingCountry>
.
<taxonomicName id="288C4D0A795FFFFACED9F8D4FED7F8C1" box="[151,339,1879,1905]" class="Clitellata" family="Syngenodrilidae" kingdom="Animalia" order="Alluroidida" pageId="9" pageNumber="433" phylum="Annelida" rank="subFamily" subFamily="Syngenodrilinae">Syngenodrilinae</taxonomicName>
:
<collectingCountry id="979B7619795FFFFACF10F8D4FE2FF8C1" box="[350,427,1879,1905]" name="Kenya" pageId="9" pageNumber="410">Kenya</collectingCountry>
;
<collectingCountry id="979B7619795FFFFACFFBF8D4FD99F8C1" box="[437,541,1879,1905]" name="Tanzania" pageId="9" pageNumber="410">Tanzania</collectingCountry>
; Martinique?.
</paragraph>
<paragraph id="EF333689795FFFF9CE89F8F8FED8FED5" blockId="9.[151,1437,1375,2013]" lastBlockId="10.[151,1437,151,933]" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410">
The family
<taxonomicName id="288C4D0A795FFFFACF1AF8F8FE66F825" authorityName="Michaelsen" authorityYear="1900" box="[340,482,1915,1941]" class="Clitellata" family="Alluroididae" kingdom="Animalia" order="Alluroidida" pageId="9" pageNumber="410" phylum="Annelida" rank="family">Alluroididae</taxonomicName>
was established by
<bibRefCitation id="8B1D4B78795FFFFACC94F8F8FC36F825" author="Michaelsen, W." box="[730,946,1915,1941]" pageId="9" pageNumber="410" refId="ref23961" refString="Michaelsen, W. (1900) Oligochaeta. Das Tierreich. Vol. 10. Lief. Friedlander und Sohn, Berlin, 575 pp. https: // doi. org / 10.5962 / bhl. title. 11605" type="book" year="1900">Michaelsen (1900)</bibRefCitation>
for
<taxonomicName id="288C4D0A795FFFFACDA3F8F8FA1CF825" authority="Beddard (1894)" authorityName="Beddard" authorityYear="1894" box="[1005,1432,1915,1941]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="9" pageNumber="410" phylum="Annelida" rank="species" species="pordagei">
<emphasis id="DDF8EA9B795FFFFACDA3F8F8FB56F825" box="[1005,1234,1915,1941]" italics="true" pageId="9" pageNumber="410">Alluroides pordagei</emphasis>
<bibRefCitation id="8B1D4B78795FFFFACA91F8F8FA1CF825" author="Beddard, F. E." box="[1247,1432,1915,1941]" pageId="9" pageNumber="410" pagination="201 - 269" refId="ref22462" refString="Beddard, F. E. (1894) A contribution to our knowledge of the Oligochaeta of tropical eastern Africa. Quarterly Journal of Microscopical Science, 36, 201 - 269. https: // doi. org / 10.1242 / jcs. s 2 - 36.142.201" type="journal article" year="1894">Beddard (1894)</bibRefCitation>
</taxonomicName>
. Subsequent additions were
<taxonomicName id="288C4D0A795FFFFACF9BF81CFCC6F809" authority="Beddard (1906)" authorityName=", Beddard" authorityYear="1906" box="[469,834,1951,1977]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="9" pageNumber="410" phylum="Annelida" rank="species" species="tanganyikae">
<emphasis id="DDF8EA9B795FFFFACF9BF81CFD05F809" box="[469,641,1951,1977]" italics="true" pageId="9" pageNumber="410">A. tanganyikae</emphasis>
<bibRefCitation id="8B1D4B78795FFFFACCC2F81CFCC6F809" author="Beddard, F. E. &amp; Cunnington, W. A. &amp; Report on the Oligochaeta &amp; Proceedings of the Zoological Society of London" box="[652,834,1951,1977]" pageId="9" pageNumber="410" pagination="206 - 218" refId="ref22511" refString="Beddard, F. E. (190 a 6) Zoological results of the Third Tanganyika Expedition, conducted by Cunnington, W. A. (1904 - 1905) Report on the Oligochaeta. Proceedings of the Zoological Society of London, 1906, 206 - 218. https: // doi. org / 10.1111 / j. 1469 - 7998.1906. tb 08432. x" type="book chapter" year="1906">Beddard (1906)</bibRefCitation>
</taxonomicName>
, and two species erected by
<bibRefCitation id="8B1D4B78795FFFFACAD9F81CFAE1F809" author="Brinkhurst, R. O." box="[1175,1381,1951,1977]" pageId="9" pageNumber="410" pagination="527 - 536" refId="ref22581" refString="Brinkhurst, R. O. (1964) A taxonomic revision of the Alluroididae (Oligochaeta). Proceedings of the Zoological Society of London, 142, 527 - 536. https: // doi. org / 10.1111 / j. 1469 - 7998.1964. tb 04512. x" type="journal article" year="1964">Brinkhurst (1964)</bibRefCitation>
in a revision of the family, namely
<taxonomicName id="288C4D0A795FFFFACFB4F840FD41F86C" authorityName="Brinkhurst" authorityYear="1964" box="[506,709,1987,2013]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="9" pageNumber="410" phylum="Annelida" rank="species" species="ruwenzoriensis">
<emphasis id="DDF8EA9B795FFFFACFB4F840FD41F86C" box="[506,709,1987,2013]" italics="true" pageId="9" pageNumber="410">A. ruwenzoriensis</emphasis>
</taxonomicName>
and
<taxonomicName id="288C4D0A795FFFFACD4EF840FC20F86C" baseAuthorityName="Brinkhurst" baseAuthorityYear="1964" box="[768,932,1987,2013]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="9" pageNumber="410" phylum="Annelida" rank="species" species="americanus">
<emphasis id="DDF8EA9B795FFFFACD4EF840FC20F86C" box="[768,932,1987,2013]" italics="true" pageId="9" pageNumber="410">A. americanus</emphasis>
</taxonomicName>
. The latter species, referred to a new genus
<taxonomicName id="288C4D0A795CFFF9CED9FF14FEA1FF01" box="[151,293,151,177]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CED9FF14FEA1FF01" box="[151,293,151,177]" italics="true" pageId="10" pageNumber="411">Brinkhurstia</emphasis>
</taxonomicName>
by
<bibRefCitation id="8B1D4B78795CFFF9CF1EFF14FD89FF01" author="Jamieson, B. G. M." box="[336,525,151,177]" pageId="10" pageNumber="411" pagination="55 - 86" refId="ref23501" refString="Jamieson, B. G. M. (1968) A taxonometric investigation of the Alluroididae (Oligochaeta). Journal of Zoology, 155, 55 - 86. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03029. x" type="journal article" year="1968">Jamieson (1968)</bibRefCitation>
, was the only American record for an otherwise African group until descriptions from
<collectingCountry id="979B7619795CFFF9CE99FF38FEAAFF65" box="[215,302,187,213]" name="Guyana" pageId="10" pageNumber="411">Guyana</collectingCountry>
of
<taxonomicName id="288C4D0A795CFFF9CF17FF38FDC3FF64" authorityName="Omodeo" authorityYear="1996" box="[345,583,187,213]" class="Clitellata" family="Alluroididae" genus="Kathrynella" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="guyanae">
<emphasis id="DDF8EA9B795CFFF9CF17FF38FDC3FF64" box="[345,583,187,213]" italics="true" pageId="10" pageNumber="411">Kathrynella guyanae</emphasis>
</taxonomicName>
by
<bibRefCitation id="8B1D4B78795CFFF9CC38FF38FCAFFF65" author="Omodeo, P." box="[630,811,187,213]" pageId="10" pageNumber="411" pagination="11 - 15" refId="ref24252" refString="Omodeo, P. (1996) Kathrynella, a new oligochaete genus from Guyana. Hydrobiologia, 334. 11 - 15. https: // doi. org / 10.1007 / BF 00017349" type="journal article" year="1996">Omodeo (1996)</bibRefCitation>
and
<taxonomicName id="288C4D0A795CFFF9CD28FF38FBC7FF64" authorityName="Omodeo &amp; Coates" authorityYear="2001" box="[870,1091,187,213]" class="Clitellata" family="Alluroididae" genus="Righiella" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="jamiesoni">
<emphasis id="DDF8EA9B795CFFF9CD28FF38FBC7FF64" box="[870,1091,187,213]" italics="true" pageId="10" pageNumber="411">Righiella jamiesoni</emphasis>
</taxonomicName>
by
<bibRefCitation id="8B1D4B78795CFFF9CA3CFF38FA18FF65" author="Omodeo, P. &amp; Coates, K. A." box="[1138,1436,187,213]" pageId="10" pageNumber="411" pagination="39 - 47" refId="ref24287" refString="Omodeo, P. &amp; Coates, K. A. (2001) New alluroidids (Annelida, Clitellata) from Guyana. Hydrobiologia, 463, 39 - 47. https: // doi. org / 10.1023 / A: 1013178902798" type="journal article" year="2001">Omodeo &amp; Coates (2001)</bibRefCitation>
with redescription of
<taxonomicName id="288C4D0A795CFFF9CFC7FF5CFD25FF48" baseAuthorityName="Brinkhurst" baseAuthorityYear="1964" box="[393,673,223,249]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="americanus">
<emphasis id="DDF8EA9B795CFFF9CFC7FF5CFD25FF48" box="[393,673,223,249]" italics="true" pageId="10" pageNumber="411">Brinkhurstia americanus</emphasis>
</taxonomicName>
and addition of
<emphasis id="DDF8EA9B795CFFF9CD14FF5CFA92FF49" box="[858,1302,223,249]" italics="true" pageId="10" pageNumber="411">
<taxonomicName id="288C4D0A795CFFF9CD14FF5CFC56FF49" authorityName="Omodeo &amp; Coates" authorityYear="2001" box="[858,978,223,249]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="donaldi">B. donaldi</taxonomicName>
.
<taxonomicName id="288C4D0A795CFFF9CD93FF5CFA92FF49" authorityName="Ljungstrom" authorityYear="1971" box="[989,1302,223,249]" class="Clitellata" family="Alluroididae" genus="Barriejamiesonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="aberratus">Barriejamiesonia aberratus</taxonomicName>
</emphasis>
from
<collectingCountry id="979B7619795CFFF9CB15FF5CFF5BFEAD" name="South Africa" pageId="10" pageNumber="411">South Africa</collectingCountry>
and
<taxonomicName id="288C4D0A795CFFF9CF5CFE80FE7DFEAD" authorityName="Ljungstrom" authorityYear="1971" box="[274,505,259,285]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="lauzannei">
<emphasis id="DDF8EA9B795CFFF9CF5CFE80FE7DFEAD" box="[274,505,259,285]" italics="true" pageId="10" pageNumber="411">Alluroides lauzannei</emphasis>
</taxonomicName>
(described as
<taxonomicName id="288C4D0A795CFFF9CCDBFE80FCB8FEAD" authorityName=", Beddard" authorityYear="1906" box="[661,828,259,285]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="tanganyikae">
<emphasis id="DDF8EA9B795CFFF9CCDBFE80FCB8FEAD" box="[661,828,259,285]" italics="true" pageId="10" pageNumber="411">A. tanganyikae</emphasis>
</taxonomicName>
from Lakes
<collectingCountry id="979B7619795CFFF9CD88FE80FB80FEAD" box="[966,1028,259,285]" name="Chad" pageId="10" pageNumber="411">Chad</collectingCountry>
, Léré and Fianga by
<bibRefCitation id="8B1D4B78795CFFF9CAA6FE80FA10FEAD" author="Lauzanne, L." box="[1256,1428,259,285]" pageId="10" pageNumber="411" pagination="83 - 110" refId="ref23807" refString="Lauzanne, L. (1968) Inventaire preliminaire des Oligochetes du Lac Tchad, Cahiers d'Office de la Recherche Scientifique et Technique Outre-Mer, Serie Hydrobiologie, 2 (1), 83 - 110." type="journal article" year="1968">Lauzanne 1968</bibRefCitation>
) were added by
<bibRefCitation id="8B1D4B78795CFFF9CF0FFEA4FD9DFEF1" author="Ljungstrom, P. O." box="[321,537,295,321]" pageId="10" pageNumber="411" pagination="313 - 325" refId="ref23844" refString="Ljungstrom, P. O. (1971) New species of aquatic earthworms from Malawi and South Africa. Revue d'Ecologie et de Biologie du Sol, 8 (2), 313 - 325." type="journal article" year="1971">Ljungström (1971)</bibRefCitation>
. Recently
<taxonomicName id="288C4D0A795CFFF9CCC0FEA4FC2EFEF1" authorityName="Fragoso &amp; Rojas" authorityYear="2023" box="[654,938,295,321]" class="Clitellata" family="Alluroididae" genus="Lacandodrilus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="paludosus">
<emphasis id="DDF8EA9B795CFFF9CCC0FEA4FC2EFEF1" box="[654,938,295,321]" italics="true" pageId="10" pageNumber="411">Lacandodrilus paludosus</emphasis>
</taxonomicName>
was added by
<bibRefCitation id="8B1D4B78795CFFF9CA1EFEA4FADBFEF1" author="Fragoso, C. &amp; Rojas, P." box="[1104,1375,295,321]" pageId="10" pageNumber="411" pagination="136 - 156" refId="ref23321" refString="Fragoso, C., &amp; Rojas, P. (2023) A new semiaquatic worm (Annelida, Oligochaeta) from southeastern Mexican tropical rain forests. Zootaxa, 5255 (1), 136 - 156. https: // doi. org / 10.11646 / zootaxa. 5255.1.16" type="journal article" year="2023">Fragoso &amp; Rojas (2023)</bibRefCitation>
from southern
<collectingCountry id="979B7619795CFFF9CEB1FEC8FEDCFED5" box="[255,344,331,357]" name="Mexico" pageId="10" pageNumber="411">Mexico</collectingCountry>
.
</paragraph>
<paragraph id="EF333689795CFFF9CE89FEECFE2AFCA5" blockId="10.[151,1437,151,933]" pageId="10" pageNumber="411">
A number of forms have been described (
<bibRefCitation id="8B1D4B78795CFFF9CCEDFEECFCEEFE39" author="Michaelsen, W." box="[675,874,367,393]" pageId="10" pageNumber="411" pagination="1 - 32" refId="ref24002" refString="Michaelsen, W. (1913) Oligochaten vom tropischen und sudlichsubtropischen Afrika. Zoologica, Stuttgart 68, 1 - 32." type="journal article" year="1913">Michaelsen 1913</bibRefCitation>
,
<bibRefCitation id="8B1D4B78795CFFF9CD38FEECFC41FE39" author="Michaelsen, W." box="[886,965,367,393]" pageId="10" pageNumber="411" pagination="81 - 127" refId="ref24025" refString="Michaelsen, W. (1914 a) Oligochaten vom tropischen Afrika. Mitteilungen aus dem Zoologischen Museum in Hamburg, 21, 81 - 127." type="journal article" year="1914">1914 a</bibRefCitation>
,
<bibRefCitation id="8B1D4B78795CFFF9CD9AFEECFBA1FE39" author="Michaelsen, W." box="[980,1061,367,393]" pageId="10" pageNumber="411" refId="ref24052" refString="Michaelsen, W. (1914 b) Oligochaeta, Beitrage zur Kenntnis der Land und Siisswasser Fauna Deutsch-Sudwestafrika, Hamburg Deutsch Sudwestafrika Studienreise, 1, 137. https: // doi. org / 10.5962 / bhl. title. 7352" type="journal volume" year="1914">1914 b</bibRefCitation>
,
<bibRefCitation id="8B1D4B78795CFFF9CA7CFEEDFBE9FE39" author="Michaelsen, W." box="[1074,1133,366,393]" pageId="10" pageNumber="411" pagination="23 - 42" refId="ref24095" refString="Michaelsen, W. (1915) Oligochaeta. In: Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale. 1911 - 1912. Resultats scientifiques. Vermes, 2, pp. 23 - 42." type="journal article" year="1915">1915</bibRefCitation>
,
<bibRefCitation id="8B1D4B78795CFFF9CA34FEEDFB31FE39" author="Michaelsen, W." box="[1146,1205,366,393]" pageId="10" pageNumber="411" pagination="33 - 95" refId="ref24136" refString="Michaelsen, W. (1935) Oligochaten von Belgisch-Kongo. Revue de Zoologie et de Botanie Africaines, 27, 33 - 95." type="journal article" year="1935">1935</bibRefCitation>
,
<bibRefCitation id="8B1D4B78795CFFF9CA8CFEECFB7FFE39" author="Michaelsen, W." box="[1218,1275,367,393]" pageId="10" pageNumber="411" pagination="37 - 71" refId="ref24161" refString="Michaelsen, W. (1936) Oligochaten von Belgisch-Kongo III. Revue de Zoologie et de Botanie Africaines, 29, 37 - 71." type="journal article" year="1936">1936</bibRefCitation>
) which are of uncertain status but are referred to
<taxonomicName id="288C4D0A795CFFF9CC54FE10FD19FE1D" authorityName="Beddard" authorityYear="1894" box="[538,669,403,429]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="pordagei">
<emphasis id="DDF8EA9B795CFFF9CC54FE10FD19FE1D" box="[538,669,403,429]" italics="true" pageId="10" pageNumber="411">A. pordagei</emphasis>
</taxonomicName>
(as did
<bibRefCitation id="8B1D4B78795CFFF9CCBBFE10FC3CFE1D" author="Michaelsen, W." box="[757,952,403,429]" pageId="10" pageNumber="411" pagination="37 - 71" refId="ref24161" refString="Michaelsen, W. (1936) Oligochaten von Belgisch-Kongo III. Revue de Zoologie et de Botanie Africaines, 29, 37 - 71." type="journal article" year="1936">Michaelsen 1936</bibRefCitation>
). In a taxonometric investigation,
<bibRefCitation id="8B1D4B78795CFFF9CB7AFE10FF66FE61" author="Jamieson, B. G. M." pageId="10" pageNumber="411" pagination="55 - 86" refId="ref23501" refString="Jamieson, B. G. M. (1968) A taxonometric investigation of the Alluroididae (Oligochaeta). Journal of Zoology, 155, 55 - 86. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03029. x" type="journal article" year="1968">Jamieson (1968)</bibRefCitation>
tentatively included, in the
<taxonomicName id="288C4D0A795CFFF9CC5FFE34FD25FE61" authorityName="Michaelsen" authorityYear="1900" box="[529,673,439,465]" class="Clitellata" family="Alluroididae" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="family">Alluroididae</taxonomicName>
, the subfamily
<taxonomicName id="288C4D0A795CFFF9CD1BFE34FB8BFE61" box="[853,1039,439,465]" class="Clitellata" family="Syngenodrilinae" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="family">Syngenodrilinae</taxonomicName>
, a monotypic group containing only
<taxonomicName id="288C4D0A795CFFF9CED9FE58FD27FE45" authority="Smith &amp; Green 1919" authorityName="Smith &amp; Green" authorityYear="1919" box="[151,675,475,501]" class="Clitellata" family="Syngenodrilidae" genus="Syngenodrilus" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="lamuensis">
<emphasis id="DDF8EA9B795CFFF9CED9FE58FE2BFE45" box="[151,431,475,501]" italics="true" pageId="10" pageNumber="411">Syngenodrilus lamuensis</emphasis>
<bibRefCitation id="8B1D4B78795CFFF9CFF8FE58FD27FE45" author="Smith, F. &amp; Green, B. R." box="[438,675,475,501]" pageId="10" pageNumber="411" pagination="145 - 166" refId="ref24512" refString="Smith, F. &amp; Green, B. R. (1919) Descriptions of new African earthworms including a new genus of Moniligastridae. Proceedings of the U. S. National Museum, 55, 145 - 166. https: // doi. org / 10.5479 / si. 00963801.55 - 2263.145" type="journal article" year="1919">Smith &amp; Green 1919</bibRefCitation>
</taxonomicName>
, which those authors had placed in the
<taxonomicName id="288C4D0A795CFFF9CA15FE58FA94FE45" box="[1115,1296,475,501]" class="Clitellata" family="Moniligastridae" kingdom="Animalia" order="Moniligastrida" pageId="10" pageNumber="411" phylum="Annelida" rank="family">Moniligastridae</taxonomicName>
. In the same work (
<bibRefCitation id="8B1D4B78795CFFF9CE90FE7CFE03FDA9" author="Jamieson, B. G. M." box="[222,391,511,537]" pageId="10" pageNumber="411" pagination="55 - 86" refId="ref23501" refString="Jamieson, B. G. M. (1968) A taxonometric investigation of the Alluroididae (Oligochaeta). Journal of Zoology, 155, 55 - 86. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03029. x" type="journal article" year="1968">Jamieson 1968</bibRefCitation>
) the
<typeStatus id="3037882B795CFFF9CFF3FE7CFE69FDA9" box="[445,493,511,537]" pageId="10" pageNumber="411">type</typeStatus>
of a new monotypic alluroidine genus,
<taxonomicName id="288C4D0A795CFFF9CDEDFE7CFB33FDA9" authorityName="Jamieson" authorityYear="1968" box="[931,1207,511,537]" class="Clitellata" family="Alluroididae" genus="Standeria" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="transvaalensis">
<emphasis id="DDF8EA9B795CFFF9CDEDFE7CFB33FDA9" box="[931,1207,511,537]" italics="true" pageId="10" pageNumber="411">Standeria transvaalensis</emphasis>
</taxonomicName>
(from Lake Chrissie in the former Transvaal, now the Province of Mpumalanga) was described,
<taxonomicName id="288C4D0A795CFFF9CDB6FDA0FB7CFD8C" authorityName="Brinkhurst" authorityYear="1964" box="[1016,1272,547,573]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="americanus">
<emphasis id="DDF8EA9B795CFFF9CDB6FDA0FB7CFD8C" box="[1016,1272,547,573]" italics="true" pageId="10" pageNumber="411">Alluroides americanus</emphasis>
</taxonomicName>
was made the
<typeStatus id="3037882B795CFFF9CED9FDC4FF43FDD1" box="[151,199,583,609]" pageId="10" pageNumber="411">type</typeStatus>
of the new, monotypic genus,
<taxonomicName id="288C4D0A795CFFF9CC7AFDC4FD46FDD1" box="[564,706,583,609]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CC7AFDC4FD46FDD1" box="[564,706,583,609]" italics="true" pageId="10" pageNumber="411">Brinkhurstia</emphasis>
</taxonomicName>
;
<emphasis id="DDF8EA9B795CFFF9CC9AFDC4FC11FDD0" box="[724,917,583,609]" italics="true" pageId="10" pageNumber="411">
<taxonomicName id="288C4D0A795CFFF9CC9AFDC4FCFBFDD1" authorityName=", Beddard" authorityYear="1906" box="[724,895,583,609]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="tanganyikae">A. tanganyikae</taxonomicName>
s
</emphasis>
.
<bibRefCitation id="8B1D4B78795CFFF9CDE8FDC4FBE7FDD1" author="Brinkhurst, R. O." box="[934,1123,583,609]" pageId="10" pageNumber="411" pagination="527 - 536" refId="ref22581" refString="Brinkhurst, R. O. (1964) A taxonomic revision of the Alluroididae (Oligochaeta). Proceedings of the Zoological Society of London, 142, 527 - 536. https: // doi. org / 10.1111 / j. 1469 - 7998.1964. tb 04512. x" type="journal article" year="1964">Brinkhurst 1964</bibRefCitation>
, (Mt. Elgon) was removed from
<taxonomicName id="288C4D0A795CFFF9CE96FDE8FE68FD35" authority="Beddard" authorityName=", Beddard" authorityYear="1906" box="[216,492,619,645]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="tanganyikae">
<emphasis id="DDF8EA9B795CFFF9CE96FDE8FE07FD35" box="[216,387,619,645]" italics="true" pageId="10" pageNumber="411">A. tanganyikae</emphasis>
Beddard
</taxonomicName>
and made the
<typeStatus id="3037882B795CFFF9CCD5FDE8FD4FFD35" box="[667,715,619,645]" pageId="10" pageNumber="411">type</typeStatus>
of a new species,
<taxonomicName id="288C4D0A795CFFF9CDE6FDE8FBC2FD35" authorityName="Jamieson" authorityYear="1968" box="[936,1094,619,645]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="species" species="brinkhursti">
<emphasis id="DDF8EA9B795CFFF9CDE6FDE8FBC2FD35" box="[936,1094,619,645]" italics="true" pageId="10" pageNumber="411">A. brinkhursti</emphasis>
</taxonomicName>
of which a new subspecies,
<taxonomicName id="288C4D0A795CFFF9CED9FD0CFE3AFD19" authorityName="Jamieson" authorityYear="1968" box="[151,446,655,681]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="subSpecies" species="brinkhursti" subSpecies="abyssinicus">
<emphasis id="DDF8EA9B795CFFF9CED9FD0CFE3AFD19" box="[151,446,655,681]" italics="true" pageId="10" pageNumber="411">A. brinkhursti abyssinicus</emphasis>
</taxonomicName>
was described from
<collectingCountry id="979B7619795CFFF9CCE3FD0CFC94FD19" box="[685,784,655,681]" name="Ethiopia" pageId="10" pageNumber="411">Ethiopia</collectingCountry>
.
<taxonomicName id="288C4D0A795CFFF9CD52FD0CFC64FD18" authorityName="Ljungstrom" authorityYear="1971" box="[796,992,655,680]" class="Clitellata" family="Alluroididae" genus="Barriejamiesonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CD52FD0CFC64FD18" box="[796,992,655,680]" italics="true" pageId="10" pageNumber="411">Barriejamiesonia</emphasis>
</taxonomicName>
,
<taxonomicName id="288C4D0A795CFFF9CDA1FD0CFBE7FD19" authorityName="Beddard" authorityYear="1894" box="[1007,1123,655,681]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CDA1FD0CFBE7FD19" box="[1007,1123,655,681]" italics="true" pageId="10" pageNumber="411">Alluroides</emphasis>
</taxonomicName>
,
<emphasis id="DDF8EA9B795CFFF9CA3CFD0CFA18FD19" box="[1138,1436,655,681]" italics="true" pageId="10" pageNumber="411">
<taxonomicName id="288C4D0A795CFFF9CA3CFD0CFA86FD19" box="[1138,1282,655,681]" class="Clitellata" family="Alluroididae" genus="Brinkhurstia" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">Brinkhurstia</taxonomicName>
,
<taxonomicName id="288C4D0A795CFFF9CB41FD0CFA1CFD19" authorityName="Omodeo" authorityYear="1996" box="[1295,1432,655,681]" class="Clitellata" family="Alluroididae" genus="Kathrynella" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">Kathrynella</taxonomicName>
,
</emphasis>
<taxonomicName id="288C4D0A795CFFF9CED9FD30FEADFD7C" authority=", La" authorityName="La" box="[151,297,691,717]" class="Clitellata" family="Alluroididae" genus="Righiella" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CED9FD30FF7AFD7D" box="[151,254,691,717]" italics="true" pageId="10" pageNumber="411">Righiella</emphasis>
,
<emphasis id="DDF8EA9B795CFFF9CF45FD30FEADFD7C" box="[267,297,691,716]" italics="true" pageId="10" pageNumber="411">La</emphasis>
</taxonomicName>
c
<emphasis id="DDF8EA9B795CFFF9CF7BFD30FE2AFD7D" box="[309,430,691,717]" italics="true" pageId="10" pageNumber="411">andodrilus</emphasis>
and
<taxonomicName id="288C4D0A795CFFF9CFABFD30FDD6FD7D" authorityName="Jamieson" authorityYear="1968" box="[485,594,691,717]" class="Clitellata" family="Alluroididae" genus="Standeria" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="genus">
<emphasis id="DDF8EA9B795CFFF9CFABFD30FDD6FD7D" box="[485,594,691,717]" italics="true" pageId="10" pageNumber="411">Standeria</emphasis>
</taxonomicName>
constitute the subfamily
<taxonomicName id="288C4D0A795CFFF9CD39FD30FB81FD7D" box="[887,1029,691,717]" class="Clitellata" family="Alluroidinae" kingdom="Animalia" order="Alluroidida" pageId="10" pageNumber="411" phylum="Annelida" rank="family">Alluroidinae</taxonomicName>
while the
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, as
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had suggested it should, forms the other subfamily of the
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. The two subfamilies may be distinguished as follows:
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1. Outer setal couples dorsal or lateral. Gizzards absent. Prostates (atria) 1 pair; pores combined with the male pores, or (
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<emphasis id="DDF8EA9B795CFFF9CF47FCC0FE28FCED" box="[265,428,835,861]" italics="true" pageId="10" pageNumber="411">Lacandodrilus</emphasis>
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) absent.
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.
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2. Outer setal couples lateral, not dorsal. Gizzards present. Prostates (atria) 3 pairs, pores separate from the male pores.
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.
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gen sp.
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gen sp.
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, 134, 2132.
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<paragraph id="EF3336897976FFD3CE89FDC2FE2AFD5B" blockId="32.[151,1437,469,1000]" pageId="32" pageNumber="433">Pharynx long, welldeveloped in IV and V. Septal glands 4 pairs, in V, VI, VII and VIII. Septa from 4/5 strongly developed and bulging posteriad (excepting 10/11), Testes in X and XI, with corresponding sperm funnels. Male genital tract not determinable. Ovaries one pair, in XIII; female funnel welldeveloped on septum 13/14. One pair of lateral spermatheca in IX. A second individual with the genitalia shifted 2 segments anteriorly (testes in VIII and IX, female pores in XII).</paragraph>
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<emphasis id="DDF8EA9B7976FFD3CE89FD76FEB0FCBF" bold="true" box="[199,308,757,783]" pageId="32" pageNumber="433">Remarks</emphasis>
. The lateral position of the dorsal setae and the presence of two pairs of testes (in X and XI) led Chagné &amp; Giani to place the specimens in the subfamily
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only two were mature but that their state of conservation did not permit identification. As they did not report on prostate structures the identification as
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<mods:identifier id="FBF9D7F9253AE2FC2194325B67D8E1B3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="77A35351A94E50B65D0390ECC1C09853" type="Zenodo-Dep">14033526</mods:identifier>
<mods:identifier id="04CF3832F21187548B97B4CA2C1CF0BD" type="ZooBank">8EC4885C-D44D-4C08-9B2D-9EF7D1D5AF0B</mods:identifier>
</mods:mods>
<treatment id="9E63878BFFB7FF9EFF5AF9F38E41FEC5" ID-DOI="http://doi.org/10.5281/zenodo.14033538" ID-Zenodo-Dep="14033538" LSID="urn:lsid:plazi:treatment:9E63878BFFB7FF9EFF5AF9F38E41FEC5" httpUri="http://treatment.plazi.org/id/9E63878BFFB7FF9EFF5AF9F38E41FEC5" lastPageId="4" lastPageNumber="349" pageId="1" pageNumber="346">
<subSubSection id="5ED06516FFB7FF9BFF5AF9F38E47FD27" box="[151,844,741,768]" pageId="1" pageNumber="346" type="nomenclature">
<paragraph id="1675369DFFB7FF9BFF5AF9F38E47FD27" blockId="1.[151,844,741,803]" box="[151,844,741,768]" pageId="1" pageNumber="346">
<heading id="4D3D81F1FFB7FF9BFF5AF9F38E47FD27" bold="true" box="[151,844,741,768]" fontSize="11" level="1" pageId="1" pageNumber="346" reason="1">
<emphasis id="24BEEA8FFFB7FF9BFF5AF9F38E47FD27" bold="true" box="[151,844,741,768]" pageId="1" pageNumber="346">
<taxonomicName id="D1CA4D1EFFB7FF9BFF5AF9F38FEAFD27" authority="Amal &amp; Girish Kumar" authorityName="Amal &amp; Girish Kumar" authorityYear="2024" box="[151,737,741,768]" class="Insecta" family="Crabronidae" genus="Arpactophilus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="346" phylum="Arthropoda" rank="species" species="pulawskii" status="sp. nov.">
<emphasis id="24BEEA8FFFB7FF9BFF5AF9F38CB5FCD8" bold="true" box="[151,446,741,768]" italics="true" pageId="1" pageNumber="346">Arpactophilus pulawskii</emphasis>
Amal &amp; Girish Kumar
</taxonomicName>
,
<taxonomicNameLabel id="3F8D57F4FFB7FF9BFD22F9F38E47FD27" box="[751,844,741,767]" pageId="1" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5ED06516FFB7FF9BFF5AF81F8C13FCFB" box="[151,280,777,803]" pageId="1" pageNumber="346" type="description">
<paragraph id="1675369DFFB7FF9BFF5AF81F8C13FCFB" blockId="1.[151,844,741,803]" box="[151,280,777,803]" pageId="1" pageNumber="346">
(
<figureCitation id="8EF12A18FFB7FF9BFF52F81F8C04FCFB" box="[159,271,777,803]" captionStart-0="FIGURES 15" captionStart-1="FIGURES 611" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1943,1967]" captionTargetBox-0="[186,1401,191,1909]" captionTargetBox-1="[185,1401,191,1909]" captionTargetId-0="figure-17@2.[176,1411,181,1919]" captionTargetId-1="figure-17@3.[175,1411,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 15. Arpactophilus pulawskii Amal &amp; Girish Kumar sp. nov., holotype male. 1, Habitus, lateral view; 2, Head, ventral view; 3, Head, dorsal view; 4, Head, lateral view; 5, Antenna." captionText-1="FIGURES 611. Arpactophilus pulawskii Amal &amp; Girish Kumar sp. nov., holotype male. 6, Habitus, ventral view; 7, Habitus, dorsal view; 8, Head, frontal view; 9, Propodeum, dorsal view; 10, Mesosoma, lateral view; 11, Forewing." figureDoi-0="http://doi.org/10.5281/zenodo.14033532" figureDoi-1="http://doi.org/10.5281/zenodo.14033536" httpUri-0="https://zenodo.org/record/14033532/files/figure.png" httpUri-1="https://zenodo.org/record/14033536/files/figure.png" pageId="1" pageNumber="346">Figs 111</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5ED06516FFB7FF9BFF5AF8478962FC57" pageId="1" pageNumber="346" type="materials_examined">
<paragraph id="1675369DFFB7FF9BFF5AF8478962FC57" blockId="1.[151,1437,849,2028]" pageId="1" pageNumber="346">
<emphasis id="24BEEA8FFFB7FF9BFF5AF8478C5AFCB3" bold="true" box="[151,337,849,875]" pageId="1" pageNumber="346">Type materials.</emphasis>
<materialsCitation id="A6A23CC0FFB7FF9BFE94F8478962FC57" collectingDate="2021-11-06" collectionCode="ZSIK, WGRC" collectorName="A. Rameshkumar &amp; Regd. No." country="India" elevation="1842" latitude="25.529722" location="Eden Garden" longLatPrecision="21" longitude="94.129166" municipality="Khuzama district" pageId="1" pageNumber="346" specimenCount="1" specimenCount-male="1" stateProvince="Nagaland" typeStatus="holotype">
<typeStatus id="C971883FFFB7FF9BFE94F8478CCAFCB3" box="[345,449,849,875]" pageId="1" pageNumber="346" type="holotype">Holotype</typeStatus>
<specimenCount id="00CCFD14FFB7FF9BFE07F8448CEAFCB4" box="[458,481,850,876]" count="1" pageId="1" pageNumber="346" type="male"></specimenCount>
,
<emphasis id="24BEEA8FFFB7FF9BFE23F8478F25FCB3" bold="true" box="[494,558,849,875]" pageId="1" pageNumber="346">
<collectingCountry id="6EDD760DFFB7FF9BFE23F8478F25FCB3" box="[494,558,849,875]" name="India" pageId="1" pageNumber="346">India</collectingCountry>
</emphasis>
:
<collectingRegion id="D40EF87FFFB7FF9BFDF3F8478FA7FCB3" box="[574,684,849,875]" country="India" name="Nagaland" pageId="1" pageNumber="346">Nagaland</collectingRegion>
,
<collectingMunicipality id="F611ACE7FFB7FF9BFD74F8478E76FCB3" box="[697,893,849,875]" pageId="1" pageNumber="346">Khuzama district</collectingMunicipality>
,
<location id="13156046FFB7FF9BFC44F8478916FCB3" LSID="urn:lsid:plazi:treatment:9E63878BFFB7FF9EFF5AF9F38E41FEC5:13156046FFB7FF9BFC44F8478916FCB3" box="[905,1053,849,875]" country="India" latitude="25.529722" longLatPrecision="21" longitude="94.129166" municipality="Khuzama district" name="Eden Garden" pageId="1" pageNumber="346" stateProvince="Nagaland">Eden Garden</location>
(
<geoCoordinate id="73FE505AFFB7FF9BFBE2F84789B1FCB4" box="[1071,1210,849,876]" degrees="25" direction="north" minutes="31" orientation="latitude" pageId="1" pageNumber="346" precision="15" seconds="47" value="25.529722">25°3147”N</geoCoordinate>
,
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,
<quantity id="D1329B78FFB7FF9BFA90F8478DBBFC57" metricMagnitude="3" metricUnit="m" metricValue="1.8419999999999999" pageId="1" pageNumber="346" unit="m" value="1842.0">
<elevation id="9DE7D1AEFFB7FF9BFA90F8478DBBFC57" metricMagnitude="3" metricUnit="m" metricValue="1.8419999999999999" pageId="1" pageNumber="346" unit="m" value="1842.0">1,842 m</elevation>
</quantity>
),
<date id="6274105DFFB7FF9BFF09F8638C3BFC57" box="[196,304,885,911]" pageId="1" pageNumber="346" value="2021-11-06">
<collectingDate id="7230E9B5FFB7FF9BFF09F8638C3BFC57" box="[196,304,885,911]" pageId="1" pageNumber="346" value="2021-11-06">6.xi.2021</collectingDate>
</date>
,
<collectorName id="BB3F534BFFB7FF9BFEF6F8608CF4FC57" box="[315,511,885,912]" pageId="1" pageNumber="346">A. Rameshkumar</collectorName>
collector,
<collectionCode id="70DBAE58FFB7FF9BFDB9F8638FB9FC57" box="[628,690,885,911]" pageId="1" pageNumber="346">ZSIK</collectionCode>
<collectorName id="BB3F534BFFB7FF9BFD74F8638E27FC48" box="[697,812,885,912]" pageId="1" pageNumber="346">Regd. No.</collectorName>
ZSI/
<collectionCode id="70DBAE58FFB7FF9BFCA8F8638EB0FC57" box="[869,955,885,911]" pageId="1" pageNumber="346">WGRC</collectionCode>
/IR-INV.27149.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5ED06516FFB7FF9BFF0AF88F8E20FB9C" pageId="1" pageNumber="346" type="diagnosis">
<paragraph id="1675369DFFB7FF9BFF0AF88F8E20FB9C" blockId="1.[151,1437,849,2028]" pageId="1" pageNumber="346">
<emphasis id="24BEEA8FFFB7FF9BFF0AF88F8C49FC6B" bold="true" box="[199,322,921,947]" pageId="1" pageNumber="346">Diagnosis.</emphasis>
This new species differs from all other species in having the following combination of characters: head square-shaped in dorsal view, somewhat prognathic; clypeus with median area of apical margin deeply emarginate (inverted V-shaped), uplifted; eyes distinctly small, confined to lower half of head, not reaching level of anterior ocellus; crescent-shaped depression present on either side of lateral ocellus; top of vertex deeply impressed medially; and dorsal side of propodeum irregularly rugose.
</paragraph>
</subSubSection>
<subSubSection id="5ED06516FFB7FF9EFF0AFF5B8CA9FF69" lastPageId="4" lastPageNumber="349" pageId="1" pageNumber="346" type="description">
<paragraph id="1675369DFFB7FF9BFF0AFF5B89C9FBB0" blockId="1.[151,1437,849,2028]" box="[199,1218,1101,1128]" pageId="1" pageNumber="346">
<emphasis id="24BEEA8FFFB7FF9BFF0AFF5B8C52FBBF" bold="true" box="[199,345,1101,1127]" pageId="1" pageNumber="346">Description.</emphasis>
<typeStatus id="C971883FFFB7FF9BFEADFF5B8CC3FBBF" box="[352,456,1101,1127]" pageId="1" pageNumber="346" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="8EF12A18FFB7FF9BFDD9FF5B8F8CFBBF" box="[532,647,1101,1127]" captionStart-0="FIGURES 15" captionStart-1="FIGURES 611" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1943,1967]" captionTargetBox-0="[186,1401,191,1909]" captionTargetBox-1="[185,1401,191,1909]" captionTargetId-0="figure-17@2.[176,1411,181,1919]" captionTargetId-1="figure-17@3.[175,1411,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 15. Arpactophilus pulawskii Amal &amp; Girish Kumar sp. nov., holotype male. 1, Habitus, lateral view; 2, Head, ventral view; 3, Head, dorsal view; 4, Head, lateral view; 5, Antenna." captionText-1="FIGURES 611. Arpactophilus pulawskii Amal &amp; Girish Kumar sp. nov., holotype male. 6, Habitus, ventral view; 7, Habitus, dorsal view; 8, Head, frontal view; 9, Propodeum, dorsal view; 10, Mesosoma, lateral view; 11, Forewing." figureDoi-0="http://doi.org/10.5281/zenodo.14033532" figureDoi-1="http://doi.org/10.5281/zenodo.14033536" httpUri-0="https://zenodo.org/record/14033532/files/figure.png" httpUri-1="https://zenodo.org/record/14033536/files/figure.png" pageId="1" pageNumber="346">Figs 111</figureCitation>
). Body length
<quantity id="D1329B78FFB7FF9BFCE1FF5B8E98FBB0" box="[812,915,1101,1128]" metricMagnitude="-3" metricUnit="m" metricValue="6.98" pageId="1" pageNumber="346" unit="mm" value="6.98">6.98 mm</quantity>
; forewing length
<quantity id="D1329B78FFB7FF9BFB9AFF5B89B5FBB0" box="[1111,1214,1101,1128]" metricMagnitude="-3" metricUnit="m" metricValue="4.83" pageId="1" pageNumber="346" unit="mm" value="4.83">4.83 mm</quantity>
.
</paragraph>
<paragraph id="1675369DFFB7FF9BFF0AFF678C7AFAE7" blockId="1.[151,1437,849,2028]" pageId="1" pageNumber="346">
<emphasis id="24BEEA8FFFB7FF9BFF0AFF678C13FB54" bold="true" box="[199,280,1137,1164]" italics="true" pageId="1" pageNumber="346">Colour</emphasis>
. Head ferruginous except lower vertex, upper frons, most of occiput (extending to middle region of hypostoma) brownish black. Lower frons, clypeus (margin brownish black), mandible (apical margins brownish black) pale brownish yellow.
<taxonomicName id="D1CA4D1EFFB7FF9BFE22FFAC8F63FB0C" box="[495,616,1210,1236]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="346" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
black except pronotal tubercle and tegula pale yellow. Legs yellowish brown. Metasoma black except apex of T3T7 with ferruginous band, S2 with ferruginous markings and S3S7 almost entirely ferruginous. Wings uniformly hyaline except costal cell of forewing flavo-hyaline, veins brown, stigma dark brown.
</paragraph>
<paragraph id="1675369DFFB7FF9BFF0AFE5F8F5CF913" blockId="1.[151,1437,849,2028]" pageId="1" pageNumber="346">
<emphasis id="24BEEA8FFFB7FF9BFF0AFE5F8C0EFABC" bold="true" box="[199,261,1353,1380]" italics="true" pageId="1" pageNumber="346">Head</emphasis>
. Square-shaped in dorsal view, somewhat prognathic; clypeus with median area of apical margin deeply emarginate (inverted V-shaped) with two teeth-like projections at apex, uplifted; median carina complete, slightly extending towards lower front; mandible bidentate, teeth of almost equal length; eyes distinctly small, confined to lower half of head, not reaching level of anterior ocellus; lower frons sloping towards clypeus (invaginated medially); front imbricate, with minute sparse punctures; POL 2.8 × OOL; POL 4.4 × hind ocellar diameter; OOL 1.57 × hind ocellar diameter; lower vertex punctured as frons; upper vertex almost shiny; crescent-shaped depression present on either side of lateral ocellus on vertex; top of vertex deeply impressed medially; gena shiny as upper vertex; occipital carina and hypostomal carina absent. Antenna with pedicel distinctly longer than broad; basal flagellomeres and apical flagellomere longer than broad and remaining flagellomeres as long as broad to broader than long; length: width of scape: pedicel: F1: F2: F3: F4: F5: F6: F7: F8: F9: F10: F11 = 2.96: 1.81: 1.63: 1.38: 1.09: 1.02: 1: 1: 1: 0.98: 0.98: 0.98: 1.8.
</paragraph>
<paragraph id="1675369DFFB7FF9BFF0AFDC0897FF81F" blockId="1.[151,1437,849,2028]" pageId="1" pageNumber="346">
<taxonomicName id="D1CA4D1EFFB7FF9BFF0AFDC08C49F928" box="[199,322,1750,1776]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="24BEEA8FFFB7FF9BFF0AFDC08C49F928" bold="true" box="[199,322,1750,1776]" italics="true" pageId="1" pageNumber="346">Mesosoma</emphasis>
</taxonomicName>
. Median length of mesosoma 1.31 × as long as broad (including tegula) in dorsal view; pronotum anteromedially with few inconspicuous ridges, complete transverse carina present near mesoscutum; mesoscutum imbricate with minute punctures, parapsidal line present; scutellum, metanotum, and mesopleuron punctate as mesoscutum; scutellar groove distinct; episternal sulcus prominent, omaulus present, acetabular carina absent; dorsal surface of propodeum irregularly rugose; posterior surface of propodeum with narrow transverse ridges forming reticulation towards dorsal side; lateral surface of propodeum irregularly reticulate, striate towards metapleuron. Hind femur 2.95 × as long as its maximum width. Forewing with two submarginal cells.
</paragraph>
<paragraph id="1675369DFFB7FF9BFF0AFCC48849F834" blockId="1.[151,1437,849,2028]" box="[199,1346,2001,2028]" pageId="1" pageNumber="346">
<emphasis id="24BEEA8FFFB7FF9BFF0AFCC48C34F834" bold="true" box="[199,319,2002,2028]" italics="true" pageId="1" pageNumber="346">Metasoma</emphasis>
. Micro-imbricate with sparse punctures. S2 without bulging. S8 triangular, pointed at apex.
</paragraph>
<caption id="42B56615FFB4FF98FF5AFC818E5DF80C" ID-DOI="http://doi.org/10.5281/zenodo.14033532" ID-Zenodo-Dep="14033532" httpUri="https://zenodo.org/record/14033532/files/figure.png" pageId="2" pageNumber="347" startId="2.[151,264,1943,1967]" targetBox="[186,1401,191,1909]" targetPageId="2" targetType="figure">
<paragraph id="1675369DFFB4FF98FF5AFC818E5DF80C" blockId="2.[151,1436,1943,2004]" pageId="2" pageNumber="347">
<emphasis id="24BEEA8FFFB4FF98FF5AFC818C36F877" bold="true" box="[151,317,1943,1967]" pageId="2" pageNumber="347">FIGURES 15.</emphasis>
<taxonomicName id="D1CA4D1EFFB4FF98FE8BFC818E38F877" authority="Amal &amp; Girish Kumar" authorityName="Amal &amp; Girish Kumar" authorityYear="2024" box="[326,819,1943,1968]" class="Insecta" family="Crabronidae" genus="Arpactophilus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="347" phylum="Arthropoda" rank="species" species="pulawskii" status="sp. nov.">
<emphasis id="24BEEA8FFFB4FF98FE8BFC818F34F877" box="[326,575,1943,1967]" italics="true" pageId="2" pageNumber="347">Arpactophilus pulawskii</emphasis>
Amal &amp; Girish Kumar
</taxonomicName>
<emphasis id="24BEEA8FFFB4FF98FCF1FC8E8E84F868" bold="true" box="[828,911,1944,1968]" pageId="2" pageNumber="347">
<taxonomicNameLabel id="3F8D57F4FFB4FF98FCF1FC8E8E84F868" box="[828,911,1944,1968]" pageId="2" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. 1, Habitus, lateral view; 2, Head, ventral view; 3, Head, dorsal view; 4, Head, lateral view; 5, Antenna.
</paragraph>
</caption>
<caption id="42B56615FFB5FF99FF5AFC8189C5F80B" ID-DOI="http://doi.org/10.5281/zenodo.14033536" ID-Zenodo-Dep="14033536" httpUri="https://zenodo.org/record/14033536/files/figure.png" pageId="3" pageNumber="348" startId="3.[151,264,1943,1967]" targetBox="[185,1401,191,1909]" targetPageId="3" targetType="figure">
<paragraph id="1675369DFFB5FF99FF5AFC8189C5F80B" blockId="3.[151,1436,1943,2004]" pageId="3" pageNumber="348">
<emphasis id="24BEEA8FFFB5FF99FF5AFC818C4DF877" bold="true" box="[151,326,1943,1967]" pageId="3" pageNumber="348">FIGURES 611.</emphasis>
<taxonomicName id="D1CA4D1EFFB5FF99FE81FC818E27F877" authority="Amal &amp; Girish Kumar" authorityName="Amal &amp; Girish Kumar" authorityYear="2024" box="[332,812,1943,1968]" class="Insecta" family="Crabronidae" genus="Arpactophilus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="pulawskii" status="sp. nov.">
<emphasis id="24BEEA8FFFB5FF99FE81FC818F48F877" box="[332,579,1943,1967]" italics="true" pageId="3" pageNumber="348">Arpactophilus pulawskii</emphasis>
Amal &amp; Girish Kumar
</taxonomicName>
<emphasis id="24BEEA8FFFB5FF99FCFEFC8E8E8FF868" bold="true" box="[819,900,1944,1968]" pageId="3" pageNumber="348">
<taxonomicNameLabel id="3F8D57F4FFB5FF99FCFEFC8E8E8FF868" box="[819,900,1944,1968]" pageId="3" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. 6, Habitus, ventral view; 7, Habitus, dorsal view; 8, Head, frontal view; 9, Propodeum, dorsal view; 10,
<taxonomicName id="D1CA4D1EFFB5FF99FC8FFCAA8EB9F80B" box="[834,946,1980,2003]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="346" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
, lateral view; 11, Forewing.
</paragraph>
</caption>
<paragraph id="1675369DFFB2FF9EFF0AFB818CA9FF69" blockId="4.[151,1436,151,285]" box="[199,418,151,177]" pageId="4" pageNumber="349">
<emphasis id="24BEEA8FFFB2FF9EFF0AFB818C15FF69" bold="true" box="[199,286,151,177]" pageId="4" pageNumber="349">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="5ED06516FFB2FF9EFF0AFBAC8F28FF0D" box="[199,547,186,213]" pageId="4" pageNumber="349" type="distribution">
<paragraph id="1675369DFFB2FF9EFF0AFBAC8F28FF0D" blockId="4.[151,1436,151,285]" box="[199,547,186,213]" pageId="4" pageNumber="349">
<emphasis id="24BEEA8FFFB2FF9EFF0AFBAC8C52FF0C" bold="true" box="[199,345,186,212]" pageId="4" pageNumber="349">Distribution</emphasis>
.
<collectingCountry id="6EDD760DFFB2FF9EFEAAFBAD8CAEFF0D" box="[359,421,187,213]" name="India" pageId="4" pageNumber="349">India</collectingCountry>
:
<collectingRegion id="D40EF87FFFB2FF9EFE7DFBAD8F14FF0D" box="[432,543,187,213]" country="India" name="Nagaland" pageId="4" pageNumber="349">Nagaland</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="5ED06516FFB2FF9EFF0AFBC98E41FEC5" pageId="4" pageNumber="349" type="etymology">
<paragraph id="1675369DFFB2FF9EFF0AFBC98E41FEC5" blockId="4.[151,1436,151,285]" pageId="4" pageNumber="349">
<emphasis id="24BEEA8FFFB2FF9EFF0AFBC98C43FF21" bold="true" box="[199,328,223,249]" pageId="4" pageNumber="349">Etymology</emphasis>
. The species is named in honor of Dr. Wojciech J. Pulawski,
<collectingRegion id="D40EF87FFFB2FF9EFC3DFBC98969FF21" box="[1008,1122,223,249]" country="United States of America" name="California" pageId="4" pageNumber="349">California</collectingRegion>
Academy of Sciences,
<collectingCountry id="6EDD760DFFB2FF9EFAABFBC98897FF21" box="[1382,1436,223,249]" name="United States of America" pageId="4" pageNumber="349">USA</collectingCountry>
for his valuable contribution to the taxonomy of
<taxonomicName id="D1CA4D1EFFB2FF9EFD75FA158E4DFEC5" authorityName="Latreille" authorityYear="1802" box="[696,838,259,285]" class="Insecta" family="Crabronidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="349" phylum="Arthropoda" rank="family">Crabronidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C1D1B33E92AF1B23863EAAB1E862131C">A fly in the ointment: the re-discovered holotype of Trypeta cylindrica Walker, 1853 belongs to the genus Coelopacidia Enderlein (Diptera: Tephritidae), not Chyliza Fallén (Psilidae)</mods:title>
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<taxonomicName id="8CF73B94FFDBC425FF50FAC4FDFAFA9A" ID-CoL="V4BS" authority="Lamb, 1917" authorityName="Lamb" authorityYear="1917" box="[151,522,1346,1373]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="pallidipes" status="stat. nov.">
<emphasis id="79839C05FFDBC425FF50FAC4FE9EFA9B" bold="true" box="[151,366,1346,1373]" italics="true" pageId="4" pageNumber="597">Chyliza pallidipes</emphasis>
<bibRefCitation id="2F663DE6FFDBC425FEB2FAC4FDFAFA9A" author="Lamb" box="[373,522,1346,1372]" firstAuthor="Lamb" pageId="4" pageNumber="597" pagination="266 - 273" refId="ref3900" refString="Lamb, C. G. (1917) Notes on exotic Helomyzidae, Sciomyzidae, and Psilidae. Annals and Magazine of Natural History, Series 8, 19, 266 - 273. https: // doi. org / 10.1080 / 00222931709486933" type="journal article" year="1917">Lamb, 1917</bibRefCitation>
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,
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<figureCitation id="D3CC5C92FFDBC425FF50FAE0FF2DFA46" box="[151,221,1382,1408]" captionStart="FIGURE 3" captionStartId="4.[151,250,772,796]" captionTargetBox="[340,1247,181,748]" captionTargetId="figure-21@4.[340,1247,181,748]" captionTargetPageId="4" captionText="FIGURE 3. The misidentified type of Chyliza cylindrica (Walker, 1853), NHMUK 013444153, dorsal view. Scale bar 5mm." figureDoi="http://doi.org/10.5281/zenodo.14022617" httpUri="https://zenodo.org/record/14022617/files/figure.png" pageId="4" pageNumber="597">Figs 3</figureCitation>
,
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<emphasis id="79839C05FFDBC425FF50FA2DFF14FA05" box="[151,228,1451,1475]" italics="true" pageId="4" pageNumber="597">Chyliza</emphasis>
sp.
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:
<treatmentCitation id="CA566606FFDBC425FED2FA2DFE22FA05" author="Walker, F." box="[277,466,1451,1475]" page="231" pageId="4" pageNumber="597" year="1858">
<bibRefCitation id="2F663DE6FFDBC425FED2FA2DFE22FA05" author="Walker" box="[277,466,1451,1475]" firstAuthor="Walker" page="231" pageId="4" pageNumber="597" pagination="190 - 235" refId="ref4345" refString="Walker, F. (1858) Characters of undescribed Diptera in the collection of W. W. Saunders, Esq., F. R. S. Transactions of the Entomological Society of London, Series 2, 4, 190 - 235. https: // doi. org / 10.1111 / j. 1365 - 2311.1858. tb 01822. x" type="journal article" year="1858">Walker, 1858: 231</bibRefCitation>
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.
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<emphasis id="79839C05FFDBC425FF50FA4FFEBFFA27" box="[151,335,1481,1505]" italics="true" pageId="4" pageNumber="597">Chyliza pallidipes</emphasis>
<bibRefCitation id="2F663DE6FFDBC425FE92FA4FFDF6FA24" author="Lamb" box="[341,518,1481,1506]" firstAuthor="Lamb" page="271" pageId="4" pageNumber="597" pagination="266 - 273" refId="ref3900" refString="Lamb, C. G. (1917) Notes on exotic Helomyzidae, Sciomyzidae, and Psilidae. Annals and Magazine of Natural History, Series 8, 19, 266 - 273. https: // doi. org / 10.1080 / 00222931709486933" type="journal article" year="1917">Lamb, 1917: 271</bibRefCitation>
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; Peradeniya,
<collectingCountry id="33E00087FFDBC425FD49FA4FFD04FA27" box="[654,756,1481,1505]" name="Sri Lanka" pageId="4" pageNumber="597">Sri Lanka</collectingCountry>
.
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<treatmentCitationGroup id="6BE76739FFDBC425FF50FA6FFC90F9C6" box="[151,864,1512,1536]" pageId="4" pageNumber="597">
<taxonomicName id="8CF73B94FFDBC425FF50FA6FFE55F9C6" authority="Walker" authorityName="Walker" authorityYear="1853" box="[151,421,1512,1536]" class="Insecta" family="Tephritidae" genus="Trypeta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDBC425FF50FA6FFEA2F9C6" box="[151,338,1512,1536]" italics="true" pageId="4" pageNumber="597">Trypeta cylindrica</emphasis>
Walker
</taxonomicName>
:
<treatmentCitation id="CA566606FFDBC425FE6AFA6EFD91F9C6" author="Hardy, D. E." box="[429,609,1512,1536]" page="213" pageId="4" pageNumber="597" year="1959">
<bibRefCitation id="2F663DE6FFDBC425FE6AFA6EFD91F9C6" author="Hardy" box="[429,609,1512,1536]" firstAuthor="Hardy" page="213" pageId="4" pageNumber="597" pagination="159 - 242" refId="ref3726" refString="Hardy, D. E. (1959) The Walker types of fruit flies (Tephritidae-Diptera) in the British Museum collection. Bulletin of the British Museum (Natural History), Entomology, 8 (5), 159 - 242." type="journal article" year="1959">Hardy, 1959: 213</bibRefCitation>
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;
<collectingCountry id="33E00087FFDBC425FDACFA6EFD52F9C6" box="[619,674,1512,1536]" name="India" pageId="4" pageNumber="597">India</collectingCountry>
; misidentification.
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<paragraph id="4B484017FFDBC425FF50F981FCA9F9D9" blockId="4.[151,1300,1451,1720]" box="[151,857,1543,1567]" pageId="4" pageNumber="597">
Unplaced
<taxonomicName id="8CF73B94FFDBC425FEC9F981FE37F9D9" authority="Walker" authorityName="Walker" authorityYear="1853" box="[270,455,1543,1567]" class="Insecta" family="Tephritidae" genus="Trypeta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDBC425FEC9F981FE85F9D9" box="[270,373,1543,1567]" italics="true" pageId="4" pageNumber="597">cylindrica</emphasis>
Walker
</taxonomicName>
:
<bibRefCitation id="2F663DE6FFDBC425FE17F981FD6CF9D9" author="Steyskal" box="[464,668,1543,1567]" firstAuthor="Steyskal" page="164" pageId="4" pageNumber="597" refId="ref4103" refString="Steyskal, G. C. (1977) Family Platystomatidae. In: Delfinado, M. D. &amp; Hardy, D. E. (Eds.), A catalog of the Diptera of the Oriental Region. Vol. 3. University Press of Hawaii, Honolulu, pp. 135 - 164." type="book" year="1977">Steyskal, 1977: 164</bibRefCitation>
; misidentification.
</paragraph>
<paragraph id="4B484017FFDBC425FF50F9A3FBB1F9FB" blockId="4.[151,1300,1451,1720]" box="[151,1089,1573,1598]" pageId="4" pageNumber="597">
<treatmentCitationGroup id="6BE76739FFDBC425FF50F9A3FBB1F9FB" box="[151,1089,1573,1598]" pageId="4" pageNumber="597">
<taxonomicName id="8CF73B94FFDBC425FF50F9A3FE01F9FB" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[151,497,1573,1597]" class="Insecta" family="Tephritidae" genus="Coelopacidia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDBC425FF50F9A3FE7DF9FB" box="[151,397,1573,1597]" italics="true" pageId="4" pageNumber="597">Coelopacidia cylindrica</emphasis>
(Walker)
</taxonomicName>
:
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<bibRefCitation id="2F663DE6FFDBC425FE3DF9A0FD5DF9F8" author="Kapoor" box="[506,685,1574,1598]" firstAuthor="Kapoor" page="29" pageId="4" pageNumber="597" refId="ref3868" refString="Kapoor, V. C. (1993) Indian fruit flies (lnsecta: Diptera: Tephritidae). Oxford and IBH Publishing Company, New Delhi, 228 pp." type="book" year="1993">Kapoor, 1993: 29</bibRefCitation>
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, 90; East
<collectingCountry id="33E00087FFDBC425FCD7F9A3FCB7F9FB" box="[784,839,1573,1597]" name="India" pageId="4" pageNumber="597">India</collectingCountry>
;
<emphasis id="79839C05FFDBC425FC95F9A0FC72F9FB" box="[850,898,1574,1597]" italics="true" pageId="4" pageNumber="597">Pars</emphasis>
, misidentification.
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<treatmentCitationGroup id="6BE76739FFDBC425FF50F9C2FAE4F99A" box="[151,1300,1604,1628]" pageId="4" pageNumber="597">
<taxonomicName id="8CF73B94FFDBC425FF50F9C2FE45F99A" ID-CoL="V49Z" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[151,437,1604,1628]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDBC425FF50F9C2FEA2F99A" box="[151,338,1604,1628]" italics="true" pageId="4" pageNumber="597">Chyliza cylindrica</emphasis>
(Walker)
</taxonomicName>
:
<treatmentCitation id="CA566606FFDBC425FE78F9C2FD36F99A" author="Shatalkin &amp; A. I" box="[447,710,1604,1628]" page="101" pageId="4" pageNumber="597" year="1998">
<bibRefCitation id="2F663DE6FFDBC425FE78F9C2FD36F99A" author="Shatalkin" box="[447,710,1604,1628]" firstAuthor="Shatalkin" page="101 - 102" pageId="4" pageNumber="597" pagination="89 - 111" refId="ref4001" refString="Shatalkin A. I. (1998 [for 1997]) Review of the Asian species of Chyliza Fallen (Diptera, Psilidae). Russian Entomological Journal, 6 (1 - 2), 89 - 111." type="book chapter" year="1998">Shatalkin, 1998: 101102</bibRefCitation>
</treatmentCitation>
, figs 15, 31;
<collectingCountry id="33E00087FFDBC425FC8DF9C2FC71F99A" box="[842,897,1604,1628]" name="India" pageId="4" pageNumber="597">India</collectingCountry>
,
<collectingCountry id="33E00087FFDBC425FC4DF9C2FC00F99A" box="[906,1008,1604,1628]" name="Sri Lanka" pageId="4" pageNumber="597">Sri Lanka</collectingCountry>
,
<collectingCountry id="33E00087FFDBC425FC3DF9C2FBA7F99A" box="[1018,1111,1604,1628]" name="Thailand" pageId="4" pageNumber="597">Thailand</collectingCountry>
; misidentification.
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</paragraph>
<paragraph id="4B484017FFDBC425FF50F9E5FBABF9BD" blockId="4.[151,1300,1451,1720]" box="[151,1115,1635,1659]" pageId="4" pageNumber="597">
<treatmentCitationGroup id="6BE76739FFDBC425FF50F9E5FBABF9BD" box="[151,1115,1635,1659]" pageId="4" pageNumber="597">
<taxonomicName id="8CF73B94FFDBC425FF50F9E5FE42F9BD" ID-CoL="V49Z" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[151,434,1635,1659]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDBC425FF50F9E5FEA2F9BD" box="[151,338,1635,1659]" italics="true" pageId="4" pageNumber="597">Chyliza cylindrica</emphasis>
(Walker)
</taxonomicName>
(
<taxonomicName id="8CF73B94FFDBC425FE78F9E5FD8FF9BD" ID-CoL="V4BS" authority="Lamb" authorityName="Lamb" authorityYear="1917" box="[447,639,1635,1659]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="pallidipes">
=
<emphasis id="79839C05FFDBC425FE12F9E5FDC9F9BD" box="[469,569,1635,1659]" italics="true" pageId="4" pageNumber="597">pallidipes</emphasis>
Lamb
</taxonomicName>
):
<bibRefCitation id="2F663DE6FFDBC425FD57F9E5FC6DF9BD" author="Hancock, D. L. &amp; Drew, R. A. I." box="[656,925,1635,1659]" pageId="4" pageNumber="597" refId="ref3516" refString="Hancock, D. L. &amp; Drew, R. A. I. (2005) New genera, species and records of Adramini (Diptera: Tephritidae: Trypetinae) from the" title="New genera, species and records of Adramini (Diptera: Tephritidae: Trypetinae) from the" type="book" year="2005">Hancock &amp; Drew, 2005: 5</bibRefCitation>
; misidentification.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4B484017FFDBC425FF50F907FC99F95F" blockId="4.[151,1300,1451,1720]" box="[151,873,1665,1690]" pageId="4" pageNumber="597">
<treatmentCitationGroup id="6BE76739FFDBC425FF50F907FC99F95F" box="[151,873,1665,1690]" pageId="4" pageNumber="597">
<taxonomicName id="8CF73B94FFDBC425FF50F907FE45F95F" ID-CoL="V49Z" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[151,437,1665,1689]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDBC425FF50F907FEA2F95F" box="[151,338,1665,1689]" italics="true" pageId="4" pageNumber="597">Chyliza cylindrica</emphasis>
(Walker)
</taxonomicName>
:
<treatmentCitation id="CA566606FFDBC425FE78F907FD5BF95F" author="David, K. J. &amp; Hancock, D. L. &amp; Freidberg, A. &amp; Goodger, K. F. M." box="[447,683,1665,1690]" page="456" pageId="4" pageNumber="597" year="2013">
<bibRefCitation id="2F663DE6FFDBC425FE78F907FD5BF95F" author="David" box="[447,683,1665,1690]" etAl="et al." firstAuthor="David" page="456" pageId="4" pageNumber="597" refId="ref3516" refString="3, 53 - 168. David, K. J., Hancock, D. L., Freidberg, A. &amp; Goodger, K. F. M. (2013) New species and records of Euphranta Loew and other" type="book" year="2013">
David
<emphasis id="79839C05FFDBC425FDC4F904FDC0F95F" box="[515,560,1665,1689]" italics="true" pageId="4" pageNumber="597">et al</emphasis>
., 2013: 456
</bibRefCitation>
</treatmentCitation>
; misidentification.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4B484017FFDBC425FF50F926FC9AF97E" blockId="4.[151,1300,1451,1720]" box="[151,874,1696,1720]" pageId="4" pageNumber="597">
<treatmentCitationGroup id="6BE76739FFDBC425FF50F926FC9AF97E" box="[151,874,1696,1720]" pageId="4" pageNumber="597">
<taxonomicName id="8CF73B94FFDBC425FF50F926FE45F97E" ID-CoL="V49Z" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[151,437,1696,1720]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDBC425FF50F926FEA2F97E" box="[151,338,1696,1720]" italics="true" pageId="4" pageNumber="597">Chyliza cylindrica</emphasis>
(Walker)
</taxonomicName>
:
<treatmentCitation id="CA566606FFDBC425FE78F926FD5DF97E" author="Shatalkin, A. I." box="[447,685,1696,1720]" page="152" pageId="4" pageNumber="597" year="2014">
<bibRefCitation id="2F663DE6FFDBC425FE78F926FD90F97E" author="Shatalkin" box="[447,608,1696,1720]" firstAuthor="Shatalkin" pageId="4" pageNumber="597" pagination="151 - 154" refId="ref4042" refString="Shatalkin, A. I. (2014) Chyliza gavryushini sp. n. (Psilidae) and Cypselosoma tibialis sp. n. (Cypselosomatidae) - two new dipterous species from Vietnam. Russian Entomological Journal, 23 (2), 151 - 154. https: // doi. org / 10.15298 / rusentj. 23.2.08" type="journal article" year="2014">Shatalkin, 2014</bibRefCitation>
, p. 152
</treatmentCitation>
; misidentification.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="03ED139CFFDBC425FF50F966FE18F8A0" pageId="4" pageNumber="597" type="materials_examined">
<paragraph id="4B484017FFDBC425FF50F966FD93F93C" blockId="4.[151,1436,1759,1894]" box="[151,611,1759,1786]" pageId="4" pageNumber="597">
<materialsCitation id="FB9F4A4AFFDBC425FF50F966FD93F93C" box="[151,611,1759,1786]" collectionCode="NHMUK" pageId="4" pageNumber="597" specimenCode="NHMUK 013444154" specimenCount="1" typeStatus="holotype">
<emphasis id="79839C05FFDBC425FF50F966FEFBF93C" bold="true" box="[151,267,1760,1786]" pageId="4" pageNumber="597">
<typeStatus id="944CFEB5FFDBC425FF50F966FEF6F93C" box="[151,262,1760,1786]" pageId="4" pageNumber="597" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="1B51E86CFFDBC425FED5F966FDF4F93C" box="[274,516,1759,1786]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="597" type="Museum">NHMUK 013444154</specimenCode>
, female.
</materialsCitation>
</paragraph>
<paragraph id="4B484017FFDBC425FF00F882FD9CF8D8" blockId="4.[151,1436,1759,1894]" box="[199,620,1795,1822]" pageId="4" pageNumber="597">
<emphasis id="79839C05FFDBC425FF00F882FE96F8DB" bold="true" box="[199,358,1795,1822]" pageId="4" pageNumber="597">Type locality.</emphasis>
Peradeniya,
<collectingCountry id="33E00087FFDBC425FE30F882FD98F8D8" box="[503,616,1796,1822]" name="Sri Lanka" pageId="4" pageNumber="597">Sri Lanka</collectingCountry>
.
</paragraph>
<paragraph id="4B484017FFDBC425FF00F8A1FE18F8A0" blockId="4.[151,1436,1759,1894]" pageId="4" pageNumber="597">
<emphasis id="79839C05FFDBC425FF00F8A1FE72F887" bold="true" box="[199,386,1831,1857]" pageId="4" pageNumber="597">Other material.</emphasis>
Specimens listed by
<bibRefCitation id="2F663DE6FFDBC425FDB5F8AEFCDEF884" author="Shatalkin" box="[626,814,1832,1858]" firstAuthor="Shatalkin" pageId="4" pageNumber="597" pagination="89 - 111" refId="ref4001" refString="Shatalkin A. I. (1998 [for 1997]) Review of the Asian species of Chyliza Fallen (Diptera, Psilidae). Russian Entomological Journal, 6 (1 - 2), 89 - 111." type="book chapter" year="1998">Shatalkin (1998)</bibRefCitation>
under
<taxonomicName id="8CF73B94FFDBC425FC40F8AEFBA6F884" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[903,1110,1832,1858]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDBC425FC40F8AEFBA6F884" box="[903,1110,1832,1858]" italics="true" pageId="4" pageNumber="597">Chyliza cylindrica</emphasis>
</taxonomicName>
, including the misidentified type (
<collectionCode id="2DE6D8D2FFDBC425FF21F8CAFEA2F8A0" box="[230,338,1868,1894]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="597" type="Museum">NHMUK</collectionCode>
013444153).
</paragraph>
</subSubSection>
<caption id="1F88109FFFDAC424FF50FAABFEB0FAAF" ID-DOI="http://doi.org/10.5281/zenodo.14022619" ID-Zenodo-Dep="14022619" httpUri="https://zenodo.org/record/14022619/files/figure.png" pageId="5" pageNumber="598" startId="5.[151,250,1325,1349]" targetBox="[340,1247,181,1301]" targetPageId="5" targetType="figure">
<paragraph id="4B484017FFDAC424FF50FAABFEB0FAAF" blockId="5.[151,1437,1325,1385]" pageId="5" pageNumber="598">
<emphasis id="79839C05FFDAC424FF50FAABFEE4FA83" bold="true" box="[151,276,1325,1349]" pageId="5" pageNumber="598">FIGURE 4.</emphasis>
The holotype of
<taxonomicName id="8CF73B94FFDAC424FE02FAABFCF2FA83" authority="Lamb, 1917" authorityName="Lamb" authorityYear="1917" box="[453,770,1325,1349]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="pallidipes">
<emphasis id="79839C05FFDAC424FE02FAABFD8EFA83" box="[453,638,1325,1349]" italics="true" pageId="5" pageNumber="598">Chyliza pallidipes</emphasis>
<bibRefCitation id="2F663DE6FFDAC424FD42FAABFCF2FA83" author="Lamb" box="[645,770,1325,1349]" firstAuthor="Lamb" pageId="5" pageNumber="598" pagination="266 - 273" refId="ref3900" refString="Lamb, C. G. (1917) Notes on exotic Helomyzidae, Sciomyzidae, and Psilidae. Annals and Magazine of Natural History, Series 8, 19, 266 - 273. https: // doi. org / 10.1080 / 00222931709486933" type="journal article" year="1917">Lamb, 1917</bibRefCitation>
</taxonomicName>
, NHMUK 013444154.
<emphasis id="79839C05FFDAC424FC33FAABFBF6FA83" bold="true" box="[1012,1030,1325,1349]" pageId="5" pageNumber="598">A</emphasis>
, dorsal view.
<emphasis id="79839C05FFDAC424FB51FAABFB57FA83" bold="true" box="[1174,1191,1325,1349]" pageId="5" pageNumber="598">B</emphasis>
, lateral view (left side). Scale bars 5mm.
</paragraph>
</caption>
<subSubSection id="03ED139CFFDAC424FF50FA3DFDB6F9FD" pageId="5" pageNumber="598" type="description">
<paragraph id="4B484017FFDAC424FF50FA3DFED8FA13" blockId="5.[151,296,1467,1493]" box="[151,296,1467,1493]" pageId="5" pageNumber="598">
<emphasis id="79839C05FFDAC424FF50FA3DFED8FA13" bold="true" box="[151,296,1467,1493]" pageId="5" pageNumber="598">Description</emphasis>
</paragraph>
<paragraph id="4B484017FFDAC424FF50FA7BFDB6F9FD" blockId="5.[151,1436,1532,1595]" pageId="5" pageNumber="598">
See
<bibRefCitation id="2F663DE6FFDAC424FF0FFA7AFE74F9D1" author="Shatalkin" box="[200,388,1532,1559]" firstAuthor="Shatalkin" pageId="5" pageNumber="598" pagination="89 - 111" refId="ref4001" refString="Shatalkin A. I. (1998 [for 1997]) Review of the Asian species of Chyliza Fallen (Diptera, Psilidae). Russian Entomological Journal, 6 (1 - 2), 89 - 111." type="book chapter" year="1998">Shatalkin (1998)</bibRefCitation>
under
<emphasis id="79839C05FFDAC424FE19FA7AFD47F9D0" box="[478,695,1532,1558]" italics="true" pageId="5" pageNumber="598">
<taxonomicName id="8CF73B94FFDAC424FE19FA7AFDC4F9D0" authorityName="Fallen" authorityYear="1820" box="[478,564,1532,1558]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="genus">Chyliza</taxonomicName>
cylindrica
</emphasis>
.
<typeStatus id="944CFEB5FFDAC424FD01FA7AFCC1F9D0" box="[710,817,1532,1558]" pageId="5" pageNumber="598" type="holotype">Holotype</typeStatus>
, length from tip of head (excluding antennae) to tip of abdomen
<quantity id="8C0FEDF2FFDAC424FEC2F9A6FEA6F9FD" box="[261,342,1568,1595]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="598" unit="mm" value="5.2">5.2mm</quantity>
, wing-length
<quantity id="8C0FEDF2FFDAC424FE37F9A7FDB2F9FD" box="[496,578,1569,1595]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="598" unit="mm" value="3.8">3.8mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="03ED139CFFDAC427FF50F90BFE03FE17" lastPageId="6" lastPageNumber="599" pageId="5" pageNumber="598" type="discussion">
<paragraph id="4B484017FFDAC424FF50F90BFEF9F961" blockId="5.[151,265,1677,1703]" box="[151,265,1677,1703]" pageId="5" pageNumber="598">
<emphasis id="79839C05FFDAC424FF50F90BFEF9F961" bold="true" box="[151,265,1677,1703]" pageId="5" pageNumber="598">Remarks</emphasis>
</paragraph>
<paragraph id="4B484017FFDAC424FF50F952FC74F82D" blockId="5.[151,1437,1748,2027]" pageId="5" pageNumber="598">
Specimen 013444153 matches
<bibRefCitation id="2F663DE6FFDAC424FE30F952FD52F929" author="Hardy's" box="[503,674,1748,1775]" firstAuthor="Hardy's" pageId="5" pageNumber="598" pagination="159 - 242" refId="ref3726" refString="Hardy, D. E. (1959) The Walker types of fruit flies (Tephritidae-Diptera) in the British Museum collection. Bulletin of the British Museum (Natural History), Entomology, 8 (5), 159 - 242." type="journal article" year="1959">Hardys (1959)</bibRefCitation>
description of the “type” of
<taxonomicName id="8CF73B94FFDAC424FC20F953FAFFF928" authority="Walker" authorityName="Walker" authorityYear="1853" box="[999,1295,1748,1774]" class="Insecta" family="Tephritidae" genus="Trypeta" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDAC424FC20F953FB47F928" box="[999,1207,1748,1774]" italics="true" pageId="5" pageNumber="598">Trypeta cylindrica</emphasis>
Walker
</taxonomicName>
, particularly in that it is in very poor condition with completely hyaline wings and the thorax is densely covered with recumbent white pile, but does not match the original description by
<bibRefCitation id="2F663DE6FFDAC424FC6CF89AFBBEF8F1" author="Walker" box="[939,1102,1820,1847]" firstAuthor="Walker" pageId="5" pageNumber="598" refId="ref4193" refString="Walker, F. (1853) Diptera (Part IV). In: Insecta Saundersiana or characters of undescribed insects in the collection of William Wilson Saunders, Esq., F. R. S., F. L. S. Vol. 1. John van Voorst, London, pp. 253 - 414. https: // doi. org / 10.5962 / bhl. title. 9150" type="book" year="1853">Walker (1853)</bibRefCitation>
and figures by Westwood, so was misidentified. When
<bibRefCitation id="2F663DE6FFDAC424FE75F8C6FD92F89D" author="Steyskal" box="[434,610,1856,1883]" firstAuthor="Steyskal" pageId="5" pageNumber="598" refId="ref4103" refString="Steyskal, G. C. (1977) Family Platystomatidae. In: Delfinado, M. D. &amp; Hardy, D. E. (Eds.), A catalog of the Diptera of the Oriental Region. Vol. 3. University Press of Hawaii, Honolulu, pp. 135 - 164." type="book" year="1977">Steyskal (1977)</bibRefCitation>
listed
<taxonomicName id="8CF73B94FFDAC424FD6CF8C7FCD7F89C" authorityName="Walker" authorityYear="1853" box="[683,807,1856,1883]" class="Insecta" family="Tephritidae" genus="Trypeta" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDAC424FD72F8C6FCD7F89C" box="[693,807,1856,1882]" italics="true" pageId="5" pageNumber="598">cylindrica</emphasis>
</taxonomicName>
under unplaced species of Platystomatidae he referred to
<bibRefCitation id="2F663DE6FFDAC424FF71F8E2FEA2F8B9" author="Hardy" box="[182,338,1892,1919]" firstAuthor="Hardy" pageId="5" pageNumber="598" pagination="159 - 242" refId="ref3726" refString="Hardy, D. E. (1959) The Walker types of fruit flies (Tephritidae-Diptera) in the British Museum collection. Bulletin of the British Museum (Natural History), Entomology, 8 (5), 159 - 242." type="journal article" year="1959">Hardy (1959)</bibRefCitation>
, so was therefore referring to this specimen. This specimen was subsequently referred to as the type of
<taxonomicName id="8CF73B94FFDAC424FF38F80EFDC8F864" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[255,568,1928,1954]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDAC424FF38F80EFE3EF864" box="[255,462,1928,1954]" italics="true" pageId="5" pageNumber="598">Chyliza cylindrica</emphasis>
(Walker)
</taxonomicName>
and the species was re-described by
<bibRefCitation id="2F663DE6FFDAC424FC1BF80EFB6AF865" author="Shatalkin" box="[988,1178,1928,1955]" firstAuthor="Shatalkin" pageId="5" pageNumber="598" pagination="89 - 111" refId="ref4001" refString="Shatalkin A. I. (1998 [for 1997]) Review of the Asian species of Chyliza Fallen (Diptera, Psilidae). Russian Entomological Journal, 6 (1 - 2), 89 - 111." type="book chapter" year="1998">Shatalkin (1998)</bibRefCitation>
, who included
<taxonomicName id="8CF73B94FFDAC424FA80F80EFE65F801" authority="Lamb, 1917" authorityName="Lamb" authorityYear="1917" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="pallidipes">
<emphasis id="79839C05FFDAC424FA80F80EFEF5F800" italics="true" pageId="5" pageNumber="598">Chyliza pallidipes</emphasis>
<bibRefCitation id="2F663DE6FFDAC424FECAF82AFE65F801" author="Lamb" box="[269,405,1964,1991]" firstAuthor="Lamb" pageId="5" pageNumber="598" pagination="266 - 273" refId="ref3900" refString="Lamb, C. G. (1917) Notes on exotic Helomyzidae, Sciomyzidae, and Psilidae. Annals and Magazine of Natural History, Series 8, 19, 266 - 273. https: // doi. org / 10.1080 / 00222931709486933" type="journal article" year="1917">Lamb, 1917</bibRefCitation>
</taxonomicName>
as a junior synonym. Shatalkins description was based on a total of
<specimenCount id="5DF18B9EFFDAC424FB59F82BFAC7F800" box="[1182,1335,1964,1991]" count="12" pageId="5" pageNumber="598" type="generic">12 specimens</specimenCount>
from
<collectingCountry id="33E00087FFDAC424FABBF82AFF10F82C" name="Sri Lanka" pageId="5" pageNumber="598">Sri Lanka</collectingCountry>
,
<collectingCountry id="33E00087FFDAC424FF2CF856FED5F82C" box="[235,293,2000,2026]" name="India" pageId="5" pageNumber="598">India</collectingCountry>
and
<collectingCountry id="33E00087FFDAC424FE9CF856FE31F82C" box="[347,449,2000,2026]" name="Thailand" pageId="5" pageNumber="598">Thailand</collectingCountry>
, including the
<typeStatus id="944CFEB5FFDAC424FDA1F856FD38F82C" box="[614,712,2000,2026]" pageId="5" pageNumber="598" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8CF73B94FFDAC424FD2AF857FC8CF82C" authorityName="Lamb" authorityYear="1917" box="[749,892,2000,2027]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="pallidipes">
<emphasis id="79839C05FFDAC424FD2AF857FC8CF82C" box="[749,892,2000,2027]" italics="true" pageId="5" pageNumber="598">C. pallidipes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4B484017FFD9C427FF00FF11FB55FE87" blockId="6.[151,1437,151,465]" pageId="6" pageNumber="599">
Given that the type had been misidentified and the name combination
<taxonomicName id="8CF73B94FFD9C427FC3BFF11FACAFF77" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[1020,1338,151,177]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFD9C427FC3BFF11FB3CFF77" box="[1020,1228,151,177]" italics="true" pageId="6" pageNumber="599">Chyliza cylindrica</emphasis>
(Walker)
</taxonomicName>
was not formally described as a valid species then the Type and Holotype labels were therefore incorrectly attributed. The green-circled Type label was added in 1940 when the collection was moved to safety at the beginning of World War II and the red-bordered Holotype label very recently (A. Pont, pers. comm.). However, the green-circled Type labels were added arbitrarily and were often placed on specimens that were not
<typeStatus id="944CFEB5FFD9C427FBA5FEA1FB51FE87" box="[1122,1185,295,321]" pageId="6" pageNumber="599">types</typeStatus>
.
</paragraph>
<paragraph id="4B484017FFD9C427FF00FECDFE03FE17" blockId="6.[151,1437,151,465]" pageId="6" pageNumber="599">
The
<typeStatus id="944CFEB5FFD9C427FF39FECDFE90FEA3" box="[254,352,331,357]" pageId="6" pageNumber="599" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8CF73B94FFD9C427FE49FECDFDD1FEA3" authorityName="Lamb" authorityYear="1917" box="[398,545,331,357]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="pallidipes">
<emphasis id="79839C05FFD9C427FE49FECDFDD1FEA3" box="[398,545,331,357]" italics="true" pageId="6" pageNumber="599">C. pallidipes</emphasis>
</taxonomicName>
(NHMUK 013444154,
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), and the misidentified type are clearly conspecific and are very similar in length, from tip of head (excluding antennae) to tip of abdomen, being
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Lamb
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is thus taken out of synonymy and is the valid name for this species.
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View file

@ -0,0 +1,371 @@
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<mods:title id="9F6A22B184B14B396B943B71592C07EE">A fly in the ointment: the re-discovered holotype of Trypeta cylindrica Walker, 1853 belongs to the genus Coelopacidia Enderlein (Diptera: Tephritidae), not Chyliza Fallén (Psilidae)</mods:title>
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<emphasis id="79839C05FFDDC423FF50FD34FE46FD0B" bold="true" box="[151,438,690,717]" italics="true" pageId="2" pageNumber="595">Coelopacidia cylindrica</emphasis>
(
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,
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<emphasis id="79839C05FFDDC423FF50FC9AFEA2FCF5" box="[151,338,795,819]" italics="true" pageId="2" pageNumber="595">Trypeta cylindrica</emphasis>
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, pl. 8, figs 6, 6a; East Indies.
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<emphasis id="79839C05FFDDC423FF50FCBDFEA2FC94" box="[151,338,826,850]" italics="true" pageId="2" pageNumber="595">Trypeta cylindrica</emphasis>
Walker
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.
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<emphasis id="79839C05FFDDC423FF50FCDFFEA2FCB6" box="[151,338,856,880]" italics="true" pageId="2" pageNumber="595">Trypeta cylindrica</emphasis>
Walker
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:
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; East
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.
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<emphasis id="79839C05FFDDC423FF50FCF1FE7DFC49" box="[151,397,887,911]" italics="true" pageId="2" pageNumber="595">Coelopacidia cylindrica</emphasis>
(Walker)
</taxonomicName>
:
<treatmentCitation id="CA566606FFDDC423FE3DFCF1FD50FC49" author="Hardy, D. E." box="[506,672,887,911]" page="79" pageId="2" pageNumber="595" year="1977">
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, N. comb.;
<collectingCountry id="33E00087FFDDC423FCE1FCF1FCADFC49" box="[806,861,887,911]" name="India" pageId="2" pageNumber="595">India</collectingCountry>
.
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<emphasis id="79839C05FFDDC423FF50FC10FE7DFC68" box="[151,397,918,942]" italics="true" pageId="2" pageNumber="595">Coelopacidia cylindrica</emphasis>
(Walker)
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:
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, 90; East
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;
<emphasis id="79839C05FFDDC423FC95FC11FC79FC68" box="[850,905,919,942]" italics="true" pageId="2" pageNumber="595">Pars.</emphasis>
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<paragraph id="4B484017FFDDC423FF50FC32FD53FC0A" blockId="2.[151,905,795,1034]" box="[151,675,948,973]" pageId="2" pageNumber="595">
<taxonomicName id="8CF73B94FFDDC423FF50FC32FEB6FC0A" box="[151,326,948,972]" class="Insecta" family="Tephritidae" genus="Coelopacidia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="595" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="79839C05FFDDC423FF50FC32FED0FC0A" box="[151,288,948,972]" italics="true" pageId="2" pageNumber="595">Coelopacidia</emphasis>
sp.
</taxonomicName>
:
<bibRefCitation id="2F663DE6FFDDC423FE97FC32FDADFC0A" author="Hancock, D. L. &amp; Drew, R. A. I." box="[336,605,948,973]" pageId="2" pageNumber="595" refId="ref3516" refString="Hancock, D. L. &amp; Drew, R. A. I. (2005) New genera, species and records of Adramini (Diptera: Tephritidae: Trypetinae) from the" title="New genera, species and records of Adramini (Diptera: Tephritidae: Trypetinae) from the" type="book" year="2005">Hancock &amp; Drew, 2005: 5</bibRefCitation>
;
<collectingCountry id="33E00087FFDDC423FDAFFC32FD6FFC0A" box="[616,671,948,972]" name="India" pageId="2" pageNumber="595">India</collectingCountry>
.
</paragraph>
<paragraph id="4B484017FFDDC423FF50FC55FD0BFC2D" blockId="2.[151,905,795,1034]" box="[151,763,979,1003]" pageId="2" pageNumber="595">
<treatmentCitationGroup id="6BE76739FFDDC423FF50FC55FD0BFC2D" box="[151,763,979,1003]" pageId="2" pageNumber="595">
<taxonomicName id="8CF73B94FFDDC423FF50FC55FEB6FC2D" box="[151,326,979,1003]" class="Insecta" family="Tephritidae" genus="Coelopacidia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="595" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="79839C05FFDDC423FF50FC55FED0FC2D" box="[151,288,979,1003]" italics="true" pageId="2" pageNumber="595">Coelopacidia</emphasis>
sp.
</taxonomicName>
:
<treatmentCitation id="CA566606FFDDC423FE97FC55FDCCFC2D" author="David, K. J. &amp; Hancock, D. L. &amp; Freidberg, A. &amp; Goodger, K. F. M." box="[336,572,979,1003]" page="456" pageId="2" pageNumber="595" year="2013">
<bibRefCitation id="2F663DE6FFDDC423FE97FC55FDCCFC2D" author="David" box="[336,572,979,1003]" etAl="et al." firstAuthor="David" page="456" pageId="2" pageNumber="595" refId="ref3516" refString="3, 53 - 168. David, K. J., Hancock, D. L., Freidberg, A. &amp; Goodger, K. F. M. (2013) New species and records of Euphranta Loew and other" type="book" year="2013">
David
<emphasis id="79839C05FFDDC423FE53FC52FE31FC2D" box="[404,449,979,1003]" italics="true" pageId="2" pageNumber="595">et al</emphasis>
., 2013: 456
</bibRefCitation>
</treatmentCitation>
, figs 5759;
<collectingCountry id="33E00087FFDDC423FD07FC55FD07FC2D" box="[704,759,979,1003]" name="India" pageId="2" pageNumber="595">India</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4B484017FFDDC423FF50FC74FCD4FBCC" blockId="2.[151,905,795,1034]" box="[151,804,1010,1034]" pageId="2" pageNumber="595">
<taxonomicName id="8CF73B94FFDDC423FF50FC74FDF4FBCC" ID-CoL="3RWC" box="[151,516,1010,1034]" class="Insecta" family="Tephritidae" genus="Coelopacidia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="595" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="79839C05FFDDC423FF50FC74FED0FBCC" box="[151,288,1010,1034]" italics="true" pageId="2" pageNumber="595">Coelopacidia</emphasis>
undescribed species
</taxonomicName>
:
<bibRefCitation id="2F663DE6FFDDC423FDC9FC74FD2EFBCC" author="Hancock, D. L." box="[526,734,1010,1034]" journalOrPublisher="Israel Journal of Entomology" pageId="2" pageNumber="595" pagination="215 - 222" part="49" refId="ref3662" refString="Hancock, D. L. (2019) Comments on the genus Coelopacidia Enderlein (Diptera: Tephritidae: Trypetinae: Adramini), with a key to known species. Israel Journal of Entomology, 49 (2), 215 - 222." title="Comments on the genus Coelopacidia Enderlein (Diptera: Tephritidae: Trypetinae: Adramini), with a key to known species" type="journal article" year="2019">Hancock, 2019: 221</bibRefCitation>
;
<collectingCountry id="33E00087FFDDC423FD2FFC74FCEFFBCC" box="[744,799,1010,1034]" name="India" pageId="2" pageNumber="595">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="03ED139CFFDDC423FF50FBBEFD1AFBB0" pageId="2" pageNumber="595" type="materials_examined">
<paragraph id="4B484017FFDDC423FF50FBBEFD1AFBB0" blockId="2.[151,926,1080,1143]" pageId="2" pageNumber="595">
<materialsCitation id="FB9F4A4AFFDDC423FF50FBBEFD1AFBB0" collectionCode="NHMUK" country="India" location="India" pageId="2" pageNumber="595" specimenCode="NHMUK 013448450" specimenCount="1" typeStatus="holotype">
<emphasis id="79839C05FFDDC423FF50FBBEFEFBFB94" bold="true" box="[151,267,1080,1106]" pageId="2" pageNumber="595">
<typeStatus id="944CFEB5FFDDC423FF50FBBEFEF6FB94" box="[151,262,1080,1106]" pageId="2" pageNumber="595" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="1B51E86CFFDDC423FED5FBBFFDF4FB95" box="[274,516,1080,1107]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="595" type="Museum">NHMUK 013448450</specimenCode>
, male, William Saunders Collection.
<emphasis id="79839C05FFDDC423FF00FBDAFE96FBB0" bold="true" box="[199,358,1116,1142]" pageId="2" pageNumber="595">Type locality.</emphasis>
<collectingCountry id="33E00087FFDDC423FEABFBDAFE58FBB0" box="[364,424,1116,1142]" name="India" pageId="2" pageNumber="595">India</collectingCountry>
, precise location not known.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="03ED139CFFDDC423FF50FB4EFDD7F970" pageId="2" pageNumber="595" type="description">
<paragraph id="4B484017FFDDC423FF50FB4EFEB5FB24" blockId="2.[151,325,1224,1250]" box="[151,325,1224,1250]" pageId="2" pageNumber="595">
<heading id="1000F77BFFDDC423FF50FB4EFEB5FB24" bold="true" box="[151,325,1224,1250]" fontSize="11" level="1" pageId="2" pageNumber="595" reason="1">
<emphasis id="79839C05FFDDC423FF50FB4EFEB5FB24" bold="true" box="[151,325,1224,1250]" pageId="2" pageNumber="595">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="4B484017FFDDC423FF50FA96FDD7F970" blockId="2.[151,1436,1296,1718]" pageId="2" pageNumber="595">
Male. Length from tip of head (excluding antennae) to tip of abdomen
<quantity id="8C0FEDF2FFDDC423FC07FA96FBD0FAEC" box="[960,1056,1296,1322]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="2" pageNumber="595" unit="mm" value="10.2">10.2mm</quantity>
, wing length
<quantity id="8C0FEDF2FFDDC423FB7CFA96FAFCFAEC" box="[1211,1292,1296,1322]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="2" pageNumber="595" unit="mm" value="9.4">9.4mm</quantity>
; head length
<quantity id="8C0FEDF2FFDDC423FF50FAB2FF16FA88" box="[151,230,1332,1358]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="595" unit="mm" value="1.6">1.6mm</quantity>
(excluding antennae). Head with 2 pairs of frontal and 1 pair of orbital setae, all weak; ocellar setae very weak and hair-like; antennae about as long as face, first flagellomere fulvous, third flagellomere pale yellow, elongate and apically rounded; arista long, black and bare. Thorax largely reddish brown with postpronotal lobes and narrow postsutural lateral vittae yellow; anepisternal stripe broadly yellow and extending as a narrow dorsal stripe to postpronotal lobe; chaetotaxy with well developed postpronotal, anterior and posterior notopleural, supra-alar and postalar setae and moderately developed dorsocentral setae placed behind line of supra-alar setae; I anepisternal seta; scutellum red-brown with 2 pairs of setae, basal and apical. Wing largely hyaline with a very faint yellowish tinge; pterostigma yellow; r-m crossvein at apical third of cell dm; dm-cu crossvein with a narrow infuscation, broadest posteriorly; apical brown spot across vein R4+5 as broad as long in apical portion of cell r2+3. Legs yellow; fore femora with 2 short, black anterior spines in distal half; mid tibiae with 2 distinct apical spines. Abdomen elongate, reddish brown with darker staining.
</paragraph>
</subSubSection>
<subSubSection id="03ED139CFFDDC425FF50F88EFE9FFBDF" lastPageId="4" lastPageNumber="597" pageId="2" pageNumber="595" type="discussion">
<paragraph id="4B484017FFDDC423FF50F88EFEF9F8E4" blockId="2.[151,265,1800,1826]" box="[151,265,1800,1826]" pageId="2" pageNumber="595">
<emphasis id="79839C05FFDDC423FF50F88EFEF9F8E4" bold="true" box="[151,265,1800,1826]" pageId="2" pageNumber="595">Remarks</emphasis>
</paragraph>
<paragraph id="4B484017FFDDC423FF50F8D6FE68F83C" blockId="2.[151,1437,1872,2043]" pageId="2" pageNumber="595">
The
<taxonomicName id="8CF73B94FFDDC423FF09F8D6FE95F8AC" authorityName="Enderlein" authorityYear="1911" box="[206,357,1872,1898]" class="Insecta" family="Tephritidae" genus="Coelopacidia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="79839C05FFDDC423FF09F8D6FE95F8AC" box="[206,357,1872,1898]" italics="true" pageId="2" pageNumber="595">Coelopacidia</emphasis>
</taxonomicName>
specimen,
<collectionCode id="2DE6D8D2FFDDC423FE2BF8D7FDA6F8AD" box="[492,598,1873,1899]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="595" type="Museum">NHMUK</collectionCode>
013448450, is very similar to
<bibRefCitation id="2F663DE6FFDDC423FC02F8D6FB8FF8AD" author="Walker's" box="[965,1151,1872,1899]" firstAuthor="Walker's" pageId="2" pageNumber="595" refId="ref4193" refString="Walker, F. (1853) Diptera (Part IV). In: Insecta Saundersiana or characters of undescribed insects in the collection of William Wilson Saunders, Esq., F. R. S., F. L. S. Vol. 1. John van Voorst, London, pp. 253 - 414. https: // doi. org / 10.5962 / bhl. title. 9150" type="book" year="1853">Walkers (1853)</bibRefCitation>
original description and Westwoods figures of
<taxonomicName id="8CF73B94FFDDC423FE64F8F3FD89F848" authorityName="Walker" authorityYear="1853" box="[419,633,1908,1934]" class="Insecta" family="Tephritidae" genus="Trypeta" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="595" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDDC423FE64F8F3FE09F848" box="[419,505,1909,1934]" italics="true" pageId="2" pageNumber="595">Trypeta</emphasis>
<emphasis id="79839C05FFDDC423FDC0F8F2FD89F848" box="[519,633,1908,1934]" italics="true" pageId="2" pageNumber="595">cylindrica</emphasis>
</taxonomicName>
in general habitus and particularly in that it possesses brown wing-tips. The specimen also has distinct pterostigmata on the wings, long, black setae on the thorax and head, and a long, curved arista on the antenna, which are clearly seen in Westwoods figures. From this it is clear that this specimen is the correct
<typeStatus id="944CFEB5FFDDC423FEE8F866FE65F83C" box="[303,405,2016,2042]" pageId="2" pageNumber="595" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="1F88109FFFDCC422FF50FA4BFB78F9CF" ID-DOI="http://doi.org/10.5281/zenodo.14022613" ID-Zenodo-Dep="14022613" httpUri="https://zenodo.org/record/14022613/files/figure.png" pageId="3" pageNumber="596" startId="3.[151,250,1485,1509]" targetBox="[340,1247,181,1461]" targetPageId="3" targetType="figure">
<paragraph id="4B484017FFDCC422FF50FA4BFB78F9CF" blockId="3.[151,1437,1485,1545]" pageId="3" pageNumber="596">
<emphasis id="79839C05FFDCC422FF50FA4BFEE2FA23" bold="true" box="[151,274,1485,1509]" pageId="3" pageNumber="596">FIGURE 2.</emphasis>
The re-discovered holotype of
<taxonomicName id="8CF73B94FFDCC422FD89FA4BFC13FA23" authority="(Walker, 1853)" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[590,995,1485,1509]" class="Insecta" family="Tephritidae" genus="Coelopacidia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDCC422FD89FA4BFCB4FA23" box="[590,836,1485,1509]" italics="true" pageId="3" pageNumber="596">Coelopacidia cylindrica</emphasis>
(
<bibRefCitation id="2F663DE6FFDCC422FC97FA4BFC2BFA23" author="Walker" box="[848,987,1485,1509]" firstAuthor="Walker" pageId="3" pageNumber="596" refId="ref4193" refString="Walker, F. (1853) Diptera (Part IV). In: Insecta Saundersiana or characters of undescribed insects in the collection of William Wilson Saunders, Esq., F. R. S., F. L. S. Vol. 1. John van Voorst, London, pp. 253 - 414. https: // doi. org / 10.5962 / bhl. title. 9150" type="book" year="1853">Walker, 1853</bibRefCitation>
)
</taxonomicName>
, NHMUK 013448450.
<emphasis id="79839C05FFDCC422FB17FA4BFB12FA23" bold="true" box="[1232,1250,1485,1509]" pageId="3" pageNumber="596">A</emphasis>
, dorsal view; note the wings are not flat in this image.
<emphasis id="79839C05FFDCC422FDC4FA77FDE4F9CF" bold="true" box="[515,532,1521,1545]" pageId="3" pageNumber="596">B</emphasis>
, right side of the head. Scale bars 5mm for
<emphasis id="79839C05FFDCC422FC08FA77FC11F9CF" bold="true" box="[975,993,1521,1545]" pageId="3" pageNumber="596">A</emphasis>
and 1mm for
<emphasis id="79839C05FFDCC422FBB6FA77FB72F9CF" bold="true" box="[1137,1154,1521,1545]" pageId="3" pageNumber="596">B</emphasis>
.
</paragraph>
</caption>
<paragraph id="4B484017FFDCC422FF00F9D2FA69F91D" blockId="3.[151,1437,1620,2043]" pageId="3" pageNumber="596">
Although the number of specimens was not expressly stated, the precise nature of Walkers original description and subsequent illustrations implied only a single specimen was available, here accepted as the
<typeStatus id="944CFEB5FFDCC422FACFF9FEFA85F954" box="[1288,1397,1656,1682]" pageId="3" pageNumber="596" type="holotype">Holotype</typeStatus>
by monotypy under Article 73.1.2 of the International Code of Zoological Nomenclature (
<bibRefCitation id="2F663DE6FFDCC422FB97F91BFB23F971" author="ICZN" box="[1104,1235,1693,1719]" firstAuthor="ICZN" pageId="3" pageNumber="596" refId="ref3835" refString="ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. Fourth Edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
). This specimen, a male, also bears a handwritten [original?] label that is largely indecipherable but appears to indicate
<collectingRegion id="89338EF5FFDCC422FB38F947FACDF91D" box="[1279,1341,1729,1755]" country="South Africa" name="Western Cape" pageId="3" pageNumber="596">Cape</collectingRegion>
Camp.
</paragraph>
<paragraph id="4B484017FFDCC422FF00F962FE00F83C" blockId="3.[151,1437,1620,2043]" pageId="3" pageNumber="596">
<bibRefCitation id="2F663DE6FFDCC422FF00F962FEABF939" author="Bezzi" box="[199,347,1764,1791]" firstAuthor="Bezzi" pageId="3" pageNumber="596" refId="ref3488" refString="Bezzi, M. (1913) Indian trypaneids (fruit-flies) in the collection of the Indian Museum, Calcutta. Memoirs of the Indian Museum," type="book" year="1913">Bezzi (1913)</bibRefCitation>
suggested a relationship with
<taxonomicName id="8CF73B94FFDCC422FD7DF962FC9DF938" authority="Walker" authorityName="Walker" authorityYear="1859" box="[698,877,1764,1790]" class="Insecta" family="Tephritidae" genus="Adrama" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="79839C05FFDCC422FD7DF962FCE4F938" box="[698,788,1764,1790]" italics="true" pageId="3" pageNumber="596">Adrama</emphasis>
Walker
</taxonomicName>
and his statement of Type in London therefore indicates a reference to this specimen.
<bibRefCitation id="2F663DE6FFDCC422FD90F88EFCF4F8E5" author="Hardy's" box="[599,772,1800,1827]" firstAuthor="Hardy's" pageId="3" pageNumber="596" pagination="159 - 242" refId="ref3726" refString="Hardy, D. E. (1959) The Walker types of fruit flies (Tephritidae-Diptera) in the British Museum collection. Bulletin of the British Museum (Natural History), Entomology, 8 (5), 159 - 242." type="journal article" year="1959">Hardys (1959)</bibRefCitation>
redescription does not match the original description and illustrations, so he had misidentified the “type” (see below). Later,
<bibRefCitation id="2F663DE6FFDCC422FC67F8AAFBCAF881" author="Hardy" box="[928,1082,1836,1863]" firstAuthor="Hardy" pageId="3" pageNumber="596" refId="ref3771" refString="Hardy, D. E. (1977) Family Tephritidae (Trypetidae, Trupaneidae). In: Delfinado, M. D. &amp; Hardy, D. E. (Eds.), A catalog of the Diptera of the Oriental Region. Vol. 3. University Press of Hawaii, Honolulu, pp. 44 - 134." type="book" year="1977">Hardy (1977)</bibRefCitation>
listed
<taxonomicName id="8CF73B94FFDCC422FB4DF8AAFF0AF8AC" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1853" class="Insecta" family="Tephritidae" genus="Coelopacidia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDCC422FB4DF8AAFA6DF880" box="[1162,1437,1836,1862]" italics="true" pageId="3" pageNumber="596">Coelopacidia cylindrica</emphasis>
(Walker)
</taxonomicName>
as a new combination in the
<taxonomicName id="8CF73B94FFDCC422FD83F8D6FD36F8AC" authorityName="Macquart" authorityYear="1835" box="[580,710,1872,1898]" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="family">Tephritidae</taxonomicName>
, so he had subsequently correctly identified the
<typeStatus id="944CFEB5FFDCC422FB23F8D6FAB6F8AC" box="[1252,1350,1872,1898]" pageId="3" pageNumber="596" type="holotype">holotype</typeStatus>
but did not refer to it as such.
<bibRefCitation id="2F663DE6FFDCC422FE56F8F3FDC6F849" author="Kapoor" box="[401,566,1909,1935]" firstAuthor="Kapoor" pageId="3" pageNumber="596" refId="ref3868" refString="Kapoor, V. C. (1993) Indian fruit flies (lnsecta: Diptera: Tephritidae). Oxford and IBH Publishing Company, New Delhi, 228 pp." type="book" year="1993">Kapoor (1993)</bibRefCitation>
also listed
<taxonomicName id="8CF73B94FFDCC422FD72F8F2FC35F848" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[693,965,1908,1934]" class="Insecta" family="Tephritidae" genus="Coelopacidia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDCC422FD72F8F2FC35F848" box="[693,965,1908,1934]" italics="true" pageId="3" pageNumber="596">Coelopacidia cylindrica</emphasis>
</taxonomicName>
in the
<taxonomicName id="8CF73B94FFDCC422FBD6F8F2FB63F848" authorityName="Macquart" authorityYear="1835" box="[1041,1171,1908,1934]" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="family">Tephritidae</taxonomicName>
, however he referred to both
<bibRefCitation id="2F663DE6FFDCC422FF16F81EFE9AF875" author="Hardy" box="[209,362,1944,1971]" firstAuthor="Hardy" pageId="3" pageNumber="596" pagination="159 - 242" refId="ref3726" refString="Hardy, D. E. (1959) The Walker types of fruit flies (Tephritidae-Diptera) in the British Museum collection. Bulletin of the British Museum (Natural History), Entomology, 8 (5), 159 - 242." type="journal article" year="1959">Hardy (1959)</bibRefCitation>
and
<bibRefCitation id="2F663DE6FFDCC422FE64F81EFDCEF875" author="Hardy" box="[419,574,1944,1971]" firstAuthor="Hardy" pageId="3" pageNumber="596" refId="ref3771" refString="Hardy, D. E. (1977) Family Tephritidae (Trypetidae, Trupaneidae). In: Delfinado, M. D. &amp; Hardy, D. E. (Eds.), A catalog of the Diptera of the Oriental Region. Vol. 3. University Press of Hawaii, Honolulu, pp. 44 - 134." type="book" year="1977">Hardy (1977)</bibRefCitation>
, effectively referring to both specimens.
<bibRefCitation id="2F663DE6FFDCC422FBD6F81EFADEF875" author="Hancock, D. L. &amp; Drew, R. A. I." box="[1041,1326,1944,1971]" pageId="3" pageNumber="596" refId="ref3516" refString="Hancock, D. L. &amp; Drew, R. A. I. (2005) New genera, species and records of Adramini (Diptera: Tephritidae: Trypetinae) from the" title="New genera, species and records of Adramini (Diptera: Tephritidae: Trypetinae) from the" type="book" year="2005">Hancock &amp; Drew (2005)</bibRefCitation>
,
<bibRefCitation id="2F663DE6FFDCC422FAFCF81EFF0FF811" author="David" etAl="et al." firstAuthor="David" pageId="3" pageNumber="596" refId="ref3516" refString="3, 53 - 168. David, K. J., Hancock, D. L., Freidberg, A. &amp; Goodger, K. F. M. (2013) New species and records of Euphranta Loew and other" type="book" year="2013">
David
<emphasis id="79839C05FFDCC422FA4FF81FFF44F810" italics="true" pageId="3" pageNumber="596">et al.</emphasis>
(2013
</bibRefCitation>
, with figures) and
<bibRefCitation id="2F663DE6FFDCC422FE16F83AFD76F811" author="Hancock, D. L." box="[465,646,1980,2007]" journalOrPublisher="Israel Journal of Entomology" pageId="3" pageNumber="596" pagination="215 - 222" part="49" refId="ref3662" refString="Hancock, D. L. (2019) Comments on the genus Coelopacidia Enderlein (Diptera: Tephritidae: Trypetinae: Adramini), with a key to known species. Israel Journal of Entomology, 49 (2), 215 - 222." title="Comments on the genus Coelopacidia Enderlein (Diptera: Tephritidae: Trypetinae: Adramini), with a key to known species" type="journal article" year="2019">Hancock (2019)</bibRefCitation>
referred to this specimen but they did not consider it was the
<typeStatus id="944CFEB5FFDCC422FAFDF83AFA6CF810" box="[1338,1436,1980,2006]" pageId="3" pageNumber="596" type="holotype">holotype</typeStatus>
of a previously named species.
</paragraph>
<caption id="1F88109FFFDBC425FF50FC82FF22FC86" ID-DOI="http://doi.org/10.5281/zenodo.14022617" ID-Zenodo-Dep="14022617" httpUri="https://zenodo.org/record/14022617/files/figure.png" pageId="4" pageNumber="597" startId="4.[151,250,772,796]" targetBox="[340,1247,181,748]" targetPageId="4" targetType="figure">
<paragraph id="4B484017FFDBC425FF50FC82FF22FC86" blockId="4.[151,1436,772,832]" pageId="4" pageNumber="597">
<emphasis id="79839C05FFDBC425FF50FC82FEE7FCDA" bold="true" box="[151,279,772,796]" pageId="4" pageNumber="597">FIGURE 3.</emphasis>
The misidentified type of
<taxonomicName id="8CF73B94FFDBC425FD8AFC82FC4AFCDA" authority="' (Walker, 1853)" authorityName="' (Walker" authorityYear="1853" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[589,954,772,796]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="cylindrica">
<emphasis id="79839C05FFDBC425FD8AFC82FCFCFCDA" box="[589,780,772,796]" italics="true" pageId="4" pageNumber="597">Chyliza cylindrica</emphasis>
(
<bibRefCitation id="2F663DE6FFDBC425FCE5FC82FC42FCDA" author="Walker" box="[802,946,772,796]" firstAuthor="Walker" pageId="4" pageNumber="597" refId="ref4193" refString="Walker, F. (1853) Diptera (Part IV). In: Insecta Saundersiana or characters of undescribed insects in the collection of William Wilson Saunders, Esq., F. R. S., F. L. S. Vol. 1. John van Voorst, London, pp. 253 - 414. https: // doi. org / 10.5962 / bhl. title. 9150" type="book" year="1853">Walker, 1853</bibRefCitation>
)
</taxonomicName>
, NHMUK 013444153, dorsal view. Scale bar 5mm.
</paragraph>
</caption>
<paragraph id="4B484017FFDBC425FF00FC14FE9FFBDF" blockId="4.[151,1436,914,1049]" pageId="4" pageNumber="597">
This species is very similar to
<taxonomicName id="8CF73B94FFDBC425FDE5FC15FCDCFC6A" authority="(Munro, 1932)" baseAuthorityName="Munro" baseAuthorityYear="1932" box="[546,812,914,941]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="vivax">
<emphasis id="79839C05FFDBC425FDE5FC15FD8EFC6A" box="[546,638,915,941]" italics="true" pageId="4" pageNumber="597">C. vivax</emphasis>
(Munro, 1932)
</taxonomicName>
from
<collectingCountry id="33E00087FFDBC425FCB4FC14FBF4FC6A" box="[883,1028,914,940]" name="South Africa" pageId="4" pageNumber="597">South Africa</collectingCountry>
, differing only in the broader apical spot on the wing. The locality
<collectingRegion id="89338EF5FFDBC425FE36FC30FDDFFC16" box="[497,559,950,976]" country="South Africa" name="Western Cape" pageId="4" pageNumber="597">Cape</collectingRegion>
Camp is ambiguous and the possibility remains that the
<typeStatus id="944CFEB5FFDBC425FB75FC31FB12FC17" box="[1202,1250,951,977]" pageId="4" pageNumber="597">type</typeStatus>
locality of East Indies [
<collectingCountry id="33E00087FFDBC425FF2CFC5CFEDCFC32" box="[235,300,986,1012]" name="India" pageId="4" pageNumber="597">India</collectingCountry>
] is erroneous and the specimen is actually from the
<collectingRegion id="89338EF5FFDBC425FC43FC5CFBDAFC32" box="[900,1066,986,1012]" country="South Africa" name="Western Cape" pageId="4" pageNumber="597">Cape Province</collectingRegion>
of
<collectingCountry id="33E00087FFDBC425FB96FC5CFB13FC32" box="[1105,1251,986,1012]" name="South Africa" pageId="4" pageNumber="597">South Africa</collectingCountry>
, with
<taxonomicName id="8CF73B94FFDBC425FAEDFC5DFA77FC32" baseAuthorityName="Munro" baseAuthorityYear="1932" box="[1322,1415,987,1013]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="vivax">
<emphasis id="79839C05FFDBC425FAEDFC5DFA77FC32" box="[1322,1415,987,1013]" italics="true" pageId="4" pageNumber="597">C. vivax</emphasis>
</taxonomicName>
a potential synonym.
</paragraph>
</subSubSection>
</treatment>
</document>