treatments-xml/data/B7/BB/6E/B7BB6E7433D4597DBD1F95EC874B5B10.xml

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<mods:title id="15A12736211077DA8721CF10E316E463">Phylogeny of the old and fragmented genus Austrocoenosia Malloch reveals new evidences on the morphology and evolution of the genera Coenosia Meigen and Neodexiopsis Malloch (Diptera: Muscidae)</mods:title>
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<mods:namePart id="2C7382E534E4132B0A70C604EE517F88">Patitucci, Luciano Damian</mods:namePart>
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<mods:affiliation id="B86613A61AD9FADE18DE3D30E8514129">Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; (MACN), Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="AE54B185AF5C9D6EF234899E33F1052E">Mulieri, Pablo Ricardo</mods:namePart>
<mods:affiliation id="4CAF935473017FA8B13128AF08FC2D60">Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; (MACN), Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="52D4212233DF2FC13901FB8B6A83B107">Couri, Marcia Souto</mods:namePart>
<mods:affiliation id="EA5435BD7EBEFC1C52E07C20136D73D6">Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, RJ, Brazil &amp; Conselho Nacional de Desenvolvimento Cientifico e Tecnologico (CNPq), Brazil</mods:affiliation>
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<mods:namePart id="719621CFDD890EC667FA3C1BB0664780">Dominguez, Martha Cecilia</mods:namePart>
<mods:affiliation id="6285713354BCC64620B8D9D6E124A5F7">Laboratorio de Entomologia, Instituto Argentino de Investigaciones de Zonas Aridas (IADIZA-CONICET) Mendoza, Mendoza, Argentina</mods:affiliation>
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<paragraph id="8E22A8299F2394F75AF5AAC8A49D5543" pageId="0" pageNumber="611">
<taxonomicName id="56DE8DA6180B16AA2D309E8091F3E0FB" LSID="https://zoobank.org/CC840C06-5DAF-4F70-8D3E-65B16D060624" authority="Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez, 2023" authorityName="Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez" authorityYear="2023" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia patagonica" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="patagonica" status="sp. nov.">Coenosia patagonica</taxonomicName>
<taxonomicNameLabel id="EEA01E0FFD228CDECE36341758613676" pageId="0" pageNumber="611">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="3A6A7F79DCB0C737C0C088CFB32AF984" pageId="0" pageNumber="611" type="male (fig. 24a)">
<paragraph id="DB6F778563404B0DAA93AD23CB2AD09A" pageId="0" pageNumber="611">Male (Fig. 24A).</paragraph>
<paragraph id="200159D7919D3C6941236FDA0266503B" pageId="0" pageNumber="611">
Length. Body: 3.75-4.2 mm, wing: 3.70-4.0 mm. 
<emphasis id="A80ABA725CEDD51A0CE6EAB6C5960141" bold="true" italics="true" pageId="0" pageNumber="611">Head</emphasis>
: (Fig. 
<figureCitation id="90E808DA96B5B9E39D659DF5859F2AD5" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coenosia patagonica sp. nov. male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus and surstylus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure24" httpUri="https://binary.pensoft.net/fig/882092" pageId="0" pageNumber="611">24B</figureCitation>
). Frons at vertex about one third of the head width. Frons, fronto-orbital plate, parafacial and gena black with grey pollinosity; 3-4 pairs of frontal setae. Frons longer than wide, with frontal triangle long, light brown, reaching lunula. Gena height similar to the width of postpedicel. Fronto-orbital plate with 3-5 little setulae, close to parafacialia. Antenna black, apical angle of postpedicel acute; in lateral view inserted at the mid-level of the eye; arista with its longest microtrichia shorter than its basal diameter. 
<emphasis id="AD7F6C3B35625324B0E4BB823CACB529" bold="true" italics="true" pageId="0" pageNumber="611">Thorax</emphasis>
: (Fig. 
<figureCitation id="E66015F11303E76D2C24178E2BB8EDA8" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coenosia patagonica sp. nov. male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus and surstylus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure24" httpUri="https://binary.pensoft.net/fig/882092" pageId="0" pageNumber="611">24C</figureCitation>
). Black with grey pollinosity, with two thin dark-brown vitta at dorsocentral rows of setae; anterior and posterior spiracles grey. Chaetotaxy: acr s short and irregular, the anterior presutural acr s with similar length to the extra anterior presutural dorsocentral seta (eaDC); dorsocentrals 1+3, eaDC is less than one third of the aDC. Katepisternum with 1-2 setulae. 
<emphasis id="B7309F5AE4F55A07959EC31A8D63B420" bold="true" italics="true" pageId="0" pageNumber="611">Wing</emphasis>
: Tegula yellow. Both calypters whitish hyaline with white margins; halter yellow. 
<emphasis id="EEC81909FB4699BD28E2F34F41CDE122" bold="true" italics="true" pageId="0" pageNumber="611">Legs</emphasis>
: Coxae grey. Fore femur grey with apical tip yellow, mid and hind femora yellow with a black apical ring or a dorsal band. Tibiae yellow. Tarsus black. Fore femur with a row of strong pd, a row of strong pv setae, and a row of av setae at basal third. Mid femur with 4-5 setae on anterior surface on basal half, 2-3 pv and 2-3 av setae on basal third, 2 preapical setae on pd to posterior surface; mid tibia with one ad and one pd median setae, with similar length, ad positioned below of the pd seta. Hind femur with a complete row of ad and a row of av setae, with 2-3 pv setae on basal third. Similar size of claws and pulvilli of the three legs. 
<emphasis id="DDB210C4A48056FEBD8FA61F37541304" bold="true" italics="true" pageId="0" pageNumber="611">Abdomen</emphasis>
: (Fig. 
<figureCitation id="EFB9F6406D405787A57FEBAF6D1AAAAE" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coenosia patagonica sp. nov. male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus and surstylus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure24" httpUri="https://binary.pensoft.net/fig/882092" pageId="0" pageNumber="611">24D</figureCitation>
). Black, with grey pruinosity, tergites 1+2-5 with two dark brown spots. Tergite 6 not visible in dorsal. Sternite 5 broader than long, apical margin concave with a 
<normalizedToken id="508B56CC1003674F8979F0F433E2FE52" originalValue="“U”">&quot;U&quot;</normalizedToken>
shape, with a depressed area at the base, and without membrane; setae concentrated on the lobes and some long and strong on apical margin (Fig. 
<figureCitation id="B6E75CB8B0180CC1D7F4862EA2B99705" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coenosia patagonica sp. nov. male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus and surstylus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure24" httpUri="https://binary.pensoft.net/fig/882092" pageId="0" pageNumber="611">24E</figureCitation>
). 
<emphasis id="226514E1A67D715B81550B2E3B91A26A" bold="true" italics="true" pageId="0" pageNumber="611">Terminalia</emphasis>
: Cercus curved in lateral view, keels present at basal half, apical margin little concave. Surstylus longer than broad, narrowing towards the apex, straight, sclerotized (Fig. 
<figureCitation id="47025C1C6B2E6B2CF1A3A88F0F0331C2" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coenosia patagonica sp. nov. male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus and surstylus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure24" httpUri="https://binary.pensoft.net/fig/882092" pageId="0" pageNumber="611">24F-H</figureCitation>
). Hypandrium tubular, wider than long, with several spines on ventral surface. Aedeagus with pregt developed sclerotized; pgt developed; epiphalus long and not sclerotized, and distiph tubular, sclerotized at base, and without spine on ventral surface (Fig. 
<figureCitation id="27AF78E84619D6D471731EAD8889BF1E" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coenosia patagonica sp. nov. male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus and surstylus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure24" httpUri="https://binary.pensoft.net/fig/882092" pageId="0" pageNumber="611">24H, I</figureCitation>
).
</paragraph>
<caption id="FB47A8C4BBE0AE2ED888E29E9623490E" doi="10.3897/asp.81.e104969.figure24" httpUri="https://binary.pensoft.net/fig/882092" pageId="0" pageNumber="611" start="Figure 24" startId="F24">
<paragraph id="01EA3C95955F09240691074214CB801A" pageId="0" pageNumber="611">
<emphasis id="8431530867BD271D3B26B2F5F04956D6" bold="true" pageId="0" pageNumber="611">Figure 24.</emphasis>
<taxonomicName id="3C9FD9F1F094013009201757CC87043C" authorityName="Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez" authorityYear="2023" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia patagonica" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="patagonica">
<emphasis id="A5F83323A4E4667DB92E4B84A7E6DEAF" italics="true" pageId="0" pageNumber="611">Coenosia patagonica</emphasis>
</taxonomicName>
<emphasis id="8B5E7A531C2C82A423BEDB33BB0B0671" bold="true" pageId="0" pageNumber="611">sp. nov.</emphasis>
male. 
<emphasis id="50C05947252DFB8D9CDAB89DB47121DD" bold="true" pageId="0" pageNumber="611">A</emphasis>
Lateral view, 
<emphasis id="053C0C5F2C68423F5059AEA46AB4769A" bold="true" pageId="0" pageNumber="611">B</emphasis>
head, frontal view, 
<emphasis id="01EEED2EAAFCE0059E4F08FED115FD6C" bold="true" pageId="0" pageNumber="611">C</emphasis>
thorax, dorsal view, 
<emphasis id="ADB499958FCC5618164A183B1BD6B04A" bold="true" pageId="0" pageNumber="611">D</emphasis>
abdomen, dorsal view, 
<emphasis id="097D8B3743896EDE544A01AA8C9E3070" bold="true" pageId="0" pageNumber="611">E</emphasis>
fifth sternite, 
<emphasis id="643FD63564A14843985ECD9375D9EC0E" bold="true" pageId="0" pageNumber="611">F</emphasis>
cercus and surstylus, dorsal view (SEM), 
<emphasis id="67E4EE74FD13E3DDC5F5F6582B9A7871" bold="true" pageId="0" pageNumber="611">G</emphasis>
cercus and surstylus, dorsal view, 
<emphasis id="C1DC2B0F87373BB25CFDD7111FD3EB8F" bold="true" pageId="0" pageNumber="611">H</emphasis>
cercus, surstylus and phallic complex, lateral view, 
<emphasis id="A3165FCAD1FB14C04C63BCAA96DC9E4C" bold="true" pageId="0" pageNumber="611">I</emphasis>
phallic complex, lateral view. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I). Hypandrium, spicules on ventral surface - red arrows.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="348E4C81B8340ABB32A608A738A5E6ED" pageId="0" pageNumber="611" type="female (fig. 25a, b)">
<paragraph id="E6C2D2FEA9EC9A88BE72C90C420F0983" pageId="0" pageNumber="611">Female (Fig. 25A, B).</paragraph>
<paragraph id="33A5DBC139DC4373668165104141D246" pageId="0" pageNumber="611">
Length. Body: 3.95-4.40 mm, wing: 3.55-4.10 mm. Differs from male as follows: 
<emphasis id="03F3927A3C7C312E9D43829616F80C34" bold="true" italics="true" pageId="0" pageNumber="611">Thorax</emphasis>
: Chaetotaxy: acr s long and irregular, the anterior presutural pair longer than the anterior presutural dorsocentral seta. Katepisternum with 6-8 setula. 
<emphasis id="CCFDB2DC1A3EAEC08E4996F8FA6F58B8" bold="true" italics="true" pageId="0" pageNumber="611">Ovipositor</emphasis>
: Tergites 6 and 7 with two long and thin sclerotized plates, tergite 8 with two long and wide sclerotized plates; epiproct triangular, with hair-like setae, shorter than cercus (Fig. 
<figureCitation id="8ABFC74A614C32DBAFE65760C00A51D7" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coenosia patagonica sp. nov., female. A Lateral view, B head, frontal view, C ovipositor, dorsal view, D ovipositor, ventral view, E spermathecae. Scale bar: 1 mm (A), 0.2 mm (B), 0.5 mm (C, D), 0.1 mm (E)." figureDoi="10.3897/asp.81.e104969.figure25" httpUri="https://binary.pensoft.net/fig/882093" pageId="0" pageNumber="611">25C</figureCitation>
). Sternites 6 and 7 with one long and wide plate; sternite 8 with two small, linear, and sclerotized plates each with 4-5 setae on distal margin; hypoproct triangular, setulose, with several strong setae on distal margin (Fig. 
<figureCitation id="063667922F04DA1BAE38234E2C17BCFC" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coenosia patagonica sp. nov., female. A Lateral view, B head, frontal view, C ovipositor, dorsal view, D ovipositor, ventral view, E spermathecae. Scale bar: 1 mm (A), 0.2 mm (B), 0.5 mm (C, D), 0.1 mm (E)." figureDoi="10.3897/asp.81.e104969.figure25" httpUri="https://binary.pensoft.net/fig/882093" pageId="0" pageNumber="611">25D</figureCitation>
). Spermathecae (Fig. 
<figureCitation id="883C8F8AE23F6F8F959F676FA18126B5" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coenosia patagonica sp. nov., female. A Lateral view, B head, frontal view, C ovipositor, dorsal view, D ovipositor, ventral view, E spermathecae. Scale bar: 1 mm (A), 0.2 mm (B), 0.5 mm (C, D), 0.1 mm (E)." figureDoi="10.3897/asp.81.e104969.figure25" httpUri="https://binary.pensoft.net/fig/882093" pageId="0" pageNumber="611">25E</figureCitation>
).
</paragraph>
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<paragraph id="B401FAEA540EB119710641140E3DFE24" pageId="0" pageNumber="611">
<emphasis id="85A82C59EA726C868C523552FC547099" bold="true" pageId="0" pageNumber="611">Figure 25.</emphasis>
<taxonomicName id="296E71FEE02F8D0A59263ACA0D33CA1D" authorityName="Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez" authorityYear="2023" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia patagonica" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="patagonica">
<emphasis id="82F10A88A9D2662D6B6B80EDEBA81EAE" italics="true" pageId="0" pageNumber="611">Coenosia patagonica</emphasis>
</taxonomicName>
<emphasis id="D6F0CCEB71AB84E6DFBA77C522AAD165" bold="true" pageId="0" pageNumber="611">sp. nov.</emphasis>
, female. 
<emphasis id="ACE458257486907EAC4A5FEEBD7F38B1" bold="true" pageId="0" pageNumber="611">A</emphasis>
Lateral view, 
<emphasis id="F584390644854F9C5DA5874D1860B05F" bold="true" pageId="0" pageNumber="611">B</emphasis>
head, frontal view, 
<emphasis id="00477C22A09F5AB9DE083E46346959AA" bold="true" pageId="0" pageNumber="611">C</emphasis>
ovipositor, dorsal view, 
<emphasis id="550D1FEEB57E6EF9A2E17DD4007A592F" bold="true" pageId="0" pageNumber="611">D</emphasis>
ovipositor, ventral view, 
<emphasis id="E18BC4E909B36D4D70C31D81BF857338" bold="true" pageId="0" pageNumber="611">E</emphasis>
spermathecae. Scale bar: 1 mm (A), 0.2 mm (B), 0.5 mm (C, D), 0.1 mm (E).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F18DBA828243F58FB325AD90DF26A1E0" pageId="0" pageNumber="611" type="type material">
<paragraph id="0D3B4EACA137AEF4DAA4EA7790CC2A9F" pageId="0" pageNumber="611">Type material examined.</paragraph>
<paragraph id="61B23684EB28597C11BC7334B70226F7" pageId="0" pageNumber="611">
<emphasis id="C9949E9753D2D643DD3C68189F9C6A49" bold="true" pageId="0" pageNumber="611">Holotype.</emphasis>
Male; MACN [pinned]. Original labels: &quot;Arg. Chubut Parque Nacional / Lago Puelo (
<normalizedToken id="FB2DAE276FF1F5F9ADB63A0C938263D7" originalValue="Gendarmería">Gendarmeria</normalizedToken>
Malaise) / 
<geoCoordinate id="2EA4748B439A4DC3C401825718524A7E" degrees="42" direction="south" minutes="05,947" orientation="latitude" precision="1" value="-42.099117">42°05,947'S</geoCoordinate>
<geoCoordinate id="961DA7020CF97D82EA21E6EE7A0F804B" degrees="71" direction="west" minutes="40,937" orientation="longitude" precision="1" value="-71.68228">71°40,937' W</geoCoordinate>
/ I-2012 Mulieri &amp; Patitucci leg.&quot; print. on white paper; &quot;MACN-Ent / 34730&quot; print. on white paper, black frame; 
<normalizedToken id="4C09387B7C20D767F65CACEAC3C6D2D0" originalValue="“Holotype”">&quot;Holotype&quot;</normalizedToken>
print. on red paper, black frame. 
<emphasis id="EC9CAC89580BC5A8660E8E5E76A6EC1A" bold="true" pageId="0" pageNumber="611">Paratypes.</emphasis>
Four females, 5 males; MACN [pinned]. Original labels: &quot;Arg. Chubut Parque Nacional / Lago Puelo (
<normalizedToken id="B11BF7EC93553A6CB4BA9C3E726FAB67" originalValue="Gendarmería">Gendarmeria</normalizedToken>
Malaise) / 
<geoCoordinate id="AC5943EE00395C6FF75F00BEE40732C2" degrees="42" direction="south" minutes="05,947" orientation="latitude" precision="1" value="-42.099117">42°05,947'S</geoCoordinate>
<geoCoordinate id="5A4CE4866470529693740E8963BBE9FB" degrees="71" direction="west" minutes="40,937" orientation="longitude" precision="1" value="-71.68228">71°40,937' W</geoCoordinate>
/ I-2012 Mulieri &amp; Patitucci leg.&quot; print. on White paper; &quot;MACN-Ent / 34731 [to] 34739&quot; print. on white paper, black frame; 
<normalizedToken id="23FBAF58DC0A8C7BC5AB2B711AA8028B" originalValue="“Paratype”">&quot;Paratype&quot;</normalizedToken>
print. on red paper, black frame (MACN). 
<emphasis id="D10D1516B9E11343F738D1C99A1549F6" bold="true" pageId="0" pageNumber="611">Paratypes.</emphasis>
One female and 1 male; MLP [pinned]. Original labels: &quot;Arg. 
<normalizedToken id="7DBE18DEADA4DB3FE9F348151AB4EC45" originalValue="Neuquén">Neuquen</normalizedToken>
, Villa Pehuenia / 1187 m a.s.l. Malaise / 
<geoCoordinate id="FF87869712DC112F3AA4E44F389A9166" degrees="38.884503" direction="south" orientation="latitude" precision="1" value="-38.884502">-38.884503</geoCoordinate>
, 
<geoCoordinate id="769C66A063CEEC8E636AF3C12296D016" degrees="71.166194" direction="west" orientation="longitude" precision="1" value="-71.16619">-71.166194</geoCoordinate>
/ 9/11.II.2018 Mulieri leg.&quot; Print. on white paper, 
<normalizedToken id="9FBC50E4BF512899DA98469294F77FB8" originalValue="“Paratype”">&quot;Paratype&quot;</normalizedToken>
print. on red paper, black frame. One female; MLP [pinned]. Original labels: &quot;Arg. 
<normalizedToken id="0B4F78ABA93C17D191E27BC04B94AE54" originalValue="Río">Rio</normalizedToken>
Negro, PNN 
<normalizedToken id="E4543D32E0D05B3244CDB2136EE0CD76" originalValue="Huapí">Huapi</normalizedToken>
/ 
<normalizedToken id="C7B112B71725F4FD2B9F5CB9146AD212" originalValue="Mallín">Mallin</normalizedToken>
Playa negra 797 m / 
<geoCoordinate id="2C37D1ABB8BFBF958C73421837917CAE" degrees="41,357809" direction="south" orientation="latitude" precision="1" value="-41.357807">-41,357809</geoCoordinate>
, 
<geoCoordinate id="C0AE0C118681B4A9904F35828041F72E" degrees="71,571683" direction="west" orientation="longitude" precision="1" value="-71.571686">-71,571683</geoCoordinate>
/ 19/I-05/II/2009 / 
<normalizedToken id="DC9BA478A224287E27375E5B9DDD1D43" originalValue="Garré">Garre</normalizedToken>
&amp; M.de Oca leg. / ARG.DWN-280 / Malaise trap. MLP&quot; print. on white paper, 
<normalizedToken id="8705B3E31371B9D126D8C58CD8CB24A3" originalValue="“Paratype”">&quot;Paratype&quot;</normalizedToken>
print. on red paper, black frama. 
<emphasis id="98D69B416C3151A290F5BEB33777F61A" bold="true" pageId="0" pageNumber="611">Paratypes.</emphasis>
Two females; IFML [pinned]. Original labels: &quot;R. Argentina / 
<normalizedToken id="24F42191CCE07C95BE9DA2C22F9FB955" originalValue="Neuquén">Neuquen</normalizedToken>
/ Truran Kura / 8 km NW S.M. de los / andes- 1.000 m.s.n.m. / 18/X/1986 / col. M. y P. Gentili&quot; print. on white paper, black frame; &quot;Trampa / Malaise&quot; print. on white paper; 
<normalizedToken id="969CD7D7ADE96A9587DE669EBA6C1B6D" originalValue="“Colección">&quot;Coleccion</normalizedToken>
/ Inst. Fund. M. Lillo / (4000) S.M. de 
<normalizedToken id="1B1F5CD0729BEEFF04DF47B38747259E" originalValue="Tucumán">Tucuman</normalizedToken>
/ 
<normalizedToken id="1451800E80468A5A6070AFF416B7377E" originalValue="Tucumán">Tucuman</normalizedToken>
- Argentina&quot; print. on green paper; 
<normalizedToken id="534AC65CC43F7EB3881511B87C07970B" originalValue="“Paratype”">&quot;Paratype&quot;</normalizedToken>
print. on red paper, black frame. 4 females, pinned, labels: &quot;R. Argentina / 
<normalizedToken id="637D880CD245FEEA3F449C9F42301F6F" originalValue="Neuquén">Neuquen</normalizedToken>
/ Truran Kura / 8 km NW S.M. de los / andes- 1.000 m.s.n.m. / 25/X/1986 / col. M. y P. Gentili&quot; print. on white paper, black frame; &quot;Trampa / Malaise&quot; print. on white paper; 
<normalizedToken id="412553CA9FA327A340DFA0EC21E5C2B1" originalValue="“Colección">&quot;Coleccion</normalizedToken>
/ Inst. Fund. M. Lillo / (4000) S.M. de 
<normalizedToken id="9874E4AD0D6D8F17D04005164016C05E" originalValue="Tucumán">Tucuman</normalizedToken>
/ 
<normalizedToken id="AEBAC5AC23612C57424FE9C2D8F230E6" originalValue="Tucumán">Tucuman</normalizedToken>
- Argentina&quot; print. on green paper; 
<normalizedToken id="87DB31D89B73E23E46D848CAE65F80DA" originalValue="“Paratype”">&quot;Paratype&quot;</normalizedToken>
print. on red paper, black frame. 1 female, IFML [pinned]. Original labels: &quot;R. Argentina / Santa Cruz /Lago Argentino / Brazo Rico / 12-19-II-1980 / col. A. Willink&quot; print. on white paper, black frame; &quot;Trampa / Malaise&quot; print. on white paper; 
<normalizedToken id="F4B777B5A4314B0B360C9FE8C740B441" originalValue="“Colección">&quot;Coleccion</normalizedToken>
/ Inst. Fund. M. Lillo / (4000) S.M. de 
<normalizedToken id="1725783231DF2955AC2CBBDBAB86B628" originalValue="Tucumán">Tucuman</normalizedToken>
/ 
<normalizedToken id="0B24F4474F08595FD11A10CBAAB466B3" originalValue="Tucumán">Tucuman</normalizedToken>
- Argentina&quot; print. on green paper; 
<normalizedToken id="9EEA9ADE9654F88D1A89BC12D6F9BA29" originalValue="“Paratype”">&quot;Paratype&quot;</normalizedToken>
print. on red paper, black frame.
</paragraph>
</subSubSection>
<subSubSection id="4DC1EDB34A2F4E6FF68981212C483716" pageId="0" pageNumber="611" type="materials_examined">
<paragraph id="D6C0572AF9BD613A703755B790D9C275" pageId="0" pageNumber="611">Additional material examined.</paragraph>
<paragraph id="E2B5782F9DEDEBE2B2E0732650BA5853" pageId="0" pageNumber="611">
<materialsCitation id="D1CE6768883AFD67232B8AC2A061C16B" collectingDate="2013-02" collectorName="Mulieri, Patitucci, Olea" country="ARGENTINA" elevation="532" latitude="-42.8885" location="Cabana La Cascada" longLatPrecision="1" longitude="-71.59238" specimenCount="15" specimenCount-female="2" specimenCount-male="13" stateProvince="Chubut province">
<collectingCountry id="15335B16EE951643391F83415E5F20E4" name="Argentina">ARGENTINA</collectingCountry>
- 
<emphasis id="D58E240C148BE890EDAF60777929FD40" bold="true" pageId="0" pageNumber="611">
<collectingRegion id="DC28F806C9B762D843B97CDEC0F6D2E7" country="Argentina" name="Chubut">Chubut province</collectingRegion>
</emphasis>
• 
<specimenCount id="91E3AFEB200EB188FBA77BB03E374366" type="female">2 females</specimenCount>
, 
<specimenCount id="5375927D1B3BCD15C16C5F21969D0B9C" type="male">13 males</specimenCount>
, PNLA, 
<location id="0782CA8C86309D27601C7FE4627C3AA2" LSID="urn:lsid:plazi:treatment:B7BB6E7433D4597DBD1F95EC874B5B10:0782CA8C86309D27601C7FE4627C3AA2" country="ARGENTINA" latitude="-42.8885" longLatPrecision="1" longitude="-71.59238" name="Cabana La Cascada" stateProvince="Chubut province">
<normalizedToken id="C1C20BF2D74D844DBA6A3F57886B5361" originalValue="Cabaña">Cabana</normalizedToken>
La Cascada
</location>
; 
<geoCoordinate id="B783924A0F493DCAF42F6A163FC98070" degrees="42.888499" direction="south" orientation="latitude" precision="1" value="-42.8885">-42.888499</geoCoordinate>
, 
<geoCoordinate id="26C5933FCE681A546DC08ADB0AEA22A6" degrees="71.592376" direction="west" orientation="longitude" precision="1" value="-71.59238">-71.592376</geoCoordinate>
; 
<elevation id="A2A01DCB73156DA97BB9855DD785F9AA" metricMagnitude="2" metricUnit="m" metricValue="5.32" unit="m" value="532.0">
<quantity id="1962AA325E6F59D66851DE2A9C04944E" metricMagnitude="2" metricUnit="m" metricValue="5.32" unit="m" value="532.0">532 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate id="4341B5B8FD6F053803E41B37CCF527C5" value="2013-02">Feb 2013</collectingDate>
; 
<collectorName id="8C43F575FB7F09E307039E0A942ABD4D">Mulieri</collectorName>
, 
<collectorName id="3B8D7D78BF02FD7A3172D1502F553F7F">Patitucci</collectorName>
&amp; 
<collectorName id="7AFBDE8D5BDA57F09581CED08BD84B7D">Olea</collectorName>
leg.; MACN
</materialsCitation>
. - 
<materialsCitation id="43E41EC1FE512F38CDE2BAA23C408557" collectingDate="2018-02-13" collectingMethod="Malaise trap" collectorName="Compagnucci, Patitucci" country="Argentina" elevation="1262" latitude="-36.992245" location="Las Ovejas" longLatPrecision="1" longitude="-70.7495" specimenCount="2" specimenCount-male="2" stateProvince="Neuquen province">
<emphasis id="E21BBE24D0AC3D71375EC617F53D804B" bold="true" pageId="0" pageNumber="611">
<collectingRegion id="7D87B85ED0E9A302689919EE80C22A5E" country="Argentina" name="Neuquen">
<normalizedToken id="5C6867B55CD12DDD41165A3C8C51A71A" originalValue="Neuquén">Neuquen</normalizedToken>
province
</collectingRegion>
</emphasis>
• 
<specimenCount id="F5C83A8BC638F0DF051BBB0A737191A9" type="male">2 males</specimenCount>
; 
<location id="DB551097A76161ADE3E1787449E6E723" LSID="urn:lsid:plazi:treatment:B7BB6E7433D4597DBD1F95EC874B5B10:DB551097A76161ADE3E1787449E6E723" country="Argentina" latitude="-36.992245" longLatPrecision="1" longitude="-70.7495" name="Las Ovejas" stateProvince="Neuquen province">Las Ovejas</location>
; 
<geoCoordinate id="09875A005E727FCDABC251734647F091" degrees="36.992243" direction="south" orientation="latitude" precision="1" value="-36.992245">-36.992243</geoCoordinate>
, 
<geoCoordinate id="AA06A874B0F8BEA7458AF51FEB86607A" degrees="70.749499" direction="west" orientation="longitude" precision="1" value="-70.7495">-70.749499</geoCoordinate>
; 
<elevation id="5328930FF31B9B6099F6D861DF7B885E" metricMagnitude="3" metricUnit="m" metricValue="1.262" unit="m" value="1262.0">
<quantity id="44E58CC4E0A5EAF7F2588FB728779D61" metricMagnitude="3" metricUnit="m" metricValue="1.262" unit="m" value="1262.0">1262 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate id="44AA2F7AAD51B9287582FC67CCFE47C5" value="2018-02-13">13 Feb 2018</collectingDate>
; 
<collectorName id="7A0AC080E538850CF4F87CC0D2D817C8">Compagnucci</collectorName>
&amp; 
<collectorName id="B7AB693EBC99175B7AA3C9B46902A713">Patitucci</collectorName>
leg.; 
<collectingMethod id="677C014D933B3D812736B0A3D3D77D9B">Malaise trap</collectingMethod>
; MACN
</materialsCitation>
• 
<materialsCitation id="3BBC6ED281797796FBC0C55EDA4E51E8" collectingDate="2013-01-09" collectorName="Mulieri, Patitucci, Olea" country="Argentina" elevation="1070" latitude="-39.14693" location="Norquinco" longLatPrecision="1" longitude="-71.23272" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Neuquen">
<specimenCount id="04EA3D093BDBAEF5BA31E20D936C9938" type="female">2 females</specimenCount>
, 
<specimenCount id="B4B56C1697EEE11314EDBC68F71650B9" type="male">1 male</specimenCount>
; PNL, 
<normalizedToken id="BA1E0996844CD45E8E88E4D5AC14AD3A" originalValue="Ñorquinco">
<location id="2CF9C61D89520622D859939AC7D8F748" LSID="urn:lsid:plazi:treatment:B7BB6E7433D4597DBD1F95EC874B5B10:2CF9C61D89520622D859939AC7D8F748" country="Argentina" latitude="-39.14693" longLatPrecision="1" longitude="-71.23272" name="Norquinco" stateProvince="Neuquen">Norquinco</location>
</normalizedToken>
; 
<geoCoordinate id="D143E4CB6A37361D42EDF60DE460662A" degrees="39.146931" direction="south" orientation="latitude" precision="1" value="-39.14693">-39.146931</geoCoordinate>
, 
<geoCoordinate id="8B4CA3CF5670EC89A34D1E2161DF5565" degrees="71.232717" direction="west" orientation="longitude" precision="1" value="-71.23272">-71.232717</geoCoordinate>
; 
<elevation id="279A361680ED9F558CE290CF18CE3297" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" unit="m" value="1070.0">
<quantity id="1262F495AF14BABA56C2108CE7E92D6B" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" unit="m" value="1070.0">1070 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate id="E6369795248E200FA9D573BA1D1E129D" value="2013-01-09">09 Jan 2013</collectingDate>
; 
<collectorName id="1DB5098F8433D912B9A0BB7B4BCF6C38">Mulieri</collectorName>
, 
<collectorName id="759F871629A0E51F970B444A17B12CE7">Patitucci</collectorName>
&amp; 
<collectorName id="3BCA7C3A3E6977D004515B810FEA507C">Olea</collectorName>
leg.; MACN
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C0C4437F0CC0B0B837D921EB53961752" pageId="0" pageNumber="611" type="distribution">
<paragraph id="FCFA86C686035CB33479D25C92354124" pageId="0" pageNumber="611">Distribution (Fig. 27C).</paragraph>
<paragraph id="FB153CD86DB936B0D9C807129F02C2DA" pageId="0" pageNumber="611">
ARGENTINA: Chubut (new record), 
<normalizedToken id="73E0BB2EE22481C2EC91E2A885A55F77" originalValue="Neuquén">Neuquen</normalizedToken>
(new record), 
<normalizedToken id="6CCBBF892BB25343A1C7602E683D41B9" originalValue="Río">Rio</normalizedToken>
Negro (new record), Santa Cruz (new record).
</paragraph>
</subSubSection>
<subSubSection id="5EEF64BA87BA4852089F15C75AA2EB47" pageId="0" pageNumber="611" type="remarks">
<paragraph id="D7F74EB90DE46B6C2A55434377D9D52E" pageId="0" pageNumber="611">Remarks.</paragraph>
<paragraph id="0C3D453C137C4A29B03D970D1AA8B5CD" pageId="0" pageNumber="611">
<taxonomicName id="88A0D0FCEEE8C24BC96F5A2B9D48D3A3" authorityName="Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez" authorityYear="2023" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia patagonica" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="patagonica">
<emphasis id="2AF074FBD98190BF021960216FF841FC" italics="true" pageId="0" pageNumber="611">Coenosia patagonica</emphasis>
</taxonomicName>
<emphasis id="09FDB5A7FE5C1E54261F4FB7EDC92C93" bold="true" pageId="0" pageNumber="611">sp. nov.</emphasis>
is distinguished from its congeners by the coloration of its legs and its chaetotaxy. In addition, the male cercus has keels present at the basal half and the apical margin little concave, and surstylus longer than broad, whereas the female ovipositor is in tergite 8 with two long and wide sclerotized plates. In 
<normalizedToken id="C57CC46C2A9A01512FA9F5B4DCCAD2C6" originalValue="Stein’s">Stein's</normalizedToken>
key (1911), 
<taxonomicName id="26B4806A1FACB1EA5698FCF462C2B8C5" lsidName="C. patagonica" pageId="0" pageNumber="611" rank="species" species="patagonica">
<emphasis id="40F24B90E11DEFDFFB9389AAAD6A7746" italics="true" pageId="0" pageNumber="611">C. patagonica</emphasis>
</taxonomicName>
comes close to 
<taxonomicName id="A67F9189CB35E6412D99F87733C268BC" authorityName="Stein" authorityYear="1911" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia rotundiventris" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="rotundiventris">
<emphasis id="66B7A5B81F5EEC9B7038C74CAD160804" italics="true" pageId="0" pageNumber="611">Coenosia rotundiventris</emphasis>
</taxonomicName>
<bibRefCitation id="1EC6CD8DB653B2D8D41DE799E56AF858" author="Stein, P" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="0" pageNumber="611" pagination="61 - 189" refId="B67" refString="Stein, P, 1911. Die von Schnuse in Suedamerika gefangenen Anthomyiden. Archiv fuer Naturgeschichte 77: 61 - 189" title="Die von Schnuse in Suedamerika gefangenen Anthomyiden." volume="77" year="1911">Stein 1911</bibRefCitation>
, but differs in the coloration of legs. In 
<normalizedToken id="0E3E2D78B5F1BC31E7818EA6F3188A88" originalValue="Malloch’s">Malloch's</normalizedToken>
key (1934), 
<taxonomicName id="421A4AE4F7E5415650C9F93F48C93801" lsidName="C. patagonica" pageId="0" pageNumber="611" rank="species" species="patagonica">
<emphasis id="71FE14510E810F332F6E1DB2E7D258BB" italics="true" pageId="0" pageNumber="611">C. patagonica</emphasis>
</taxonomicName>
comes close to 
<taxonomicName id="3F47502569F5BCBA175725603DB74A41" lsidName="C. ignobilis" pageId="0" pageNumber="611" rank="species" species="ignobilis">
<emphasis id="FF771AE2293CF97C7B8E68E381E348E4" italics="true" pageId="0" pageNumber="611">C. ignobilis</emphasis>
</taxonomicName>
Stein, 1911, but the coloration of the hind femur and the male terminalia can separate it from the latter.
</paragraph>
</subSubSection>
</treatment>
</document>