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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.102.1148" ID-GBIF-Dataset="7fa288c5-61ce-4f13-b08f-56c96f19d9c0" ID-PMC="PMC3131081" ID-Pensoft-Pub="1313-2970-102-65" ID-PubMed="21747675" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-102-65" ModsDocOrigin="ZooKeys 102" ModsDocTitle="Review of the subgenus Polyphylla (Granida) from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae)" checkinTime="1451250337947" checkinUser="pensoft" docAuthor="Sehnal, Richard &amp; Bezdek, Ales" docDate="2011" docId="A67DCC3BD9A91914D710A03CD26D30D5" docLanguage="en" docName="ZooKeys 102: 65-76" docOrigin="ZooKeys 102" docSource="http://dx.doi.org/10.3897/zookeys.102.1148" docTitle="Polyphylla ((Granida)) phongsali Zidek 2006" docType="treatment" docVersion="3" lastPageNumber="73" masterDocId="FF87FFE2FFB7FFC1BD11FFBF6C6D4F56" masterDocTitle="Review of the subgenus Polyphylla (Granida) from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae)" masterLastPageNumber="76" masterPageNumber="65" pageNumber="72" updateTime="1668149989352" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the subgenus Polyphylla (Granida) from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sehnal, Richard</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezdek, Ales</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>102</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>65</mods:start>
<mods:end>76</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.102.1148</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.102.1148</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-102-65</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152029561" LSID="urn:lsid:plazi:treatment:A67DCC3BD9A91914D710A03CD26D30D5" httpUri="http://treatment.plazi.org/id/A67DCC3BD9A91914D710A03CD26D30D5" lastPageId="8" lastPageNumber="73" pageId="7" pageNumber="72">
<subSubSection pageId="7" pageNumber="72" type="nomenclature">
<paragraph pageId="7" pageNumber="72">
<taxonomicName LSID="http://species-id.net/wiki/Polyphylla_(Granida)_phongsali" authority="Zidek, 2006" authorityName="Zidek" authorityYear="2006" class="Insecta" family="Melolonthidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla ((Granida)) phongsali" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="phongsali" subGenus="(Granida)">
<pageBreakToken pageId="7" pageNumber="72" start="start">Polyphylla</pageBreakToken>
(Granida) phongsali
<normalizedToken originalValue="Zídek">Zidek</normalizedToken>
, 2006
</taxonomicName>
Figs 16-19
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="72" type="reference_group">
<paragraph pageId="7" pageNumber="72">
<taxonomicName class="Insecta" family="Melolonthidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla phongsali" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="phongsali">Polyphylla phongsali</taxonomicName>
<normalizedToken originalValue="Zídek">Zidek</normalizedToken>
, 2006: 10, pl. 3.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="72" type="type locality">
<paragraph pageId="7" pageNumber="72">Type locality.</paragraph>
<paragraph pageId="7" pageNumber="72">&quot;N. Laos, Phongsali, Gnoi-ou&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="72" type="type material examined">
<paragraph pageId="7" pageNumber="72">Type material examined.</paragraph>
<paragraph pageId="7" pageNumber="72">
Holotype (male), labeled: &quot;N. Laos, Phongsali, Gnoi-ou, Li Jingke VI-2003 [h] // BMNH (E) 2006-162 [h] // J. ZIDEK det. 2006 [p] Polyphylla phongsali
<normalizedToken originalValue="Zídek">Zidek</normalizedToken>
[h] HOLOTYPE [p, red label]&quot;, in BMNH.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="72" type="additional material examined">
<paragraph pageId="7" pageNumber="72">Additional material examined.</paragraph>
<paragraph pageId="7" pageNumber="72">
LAO-NE, Hua Phan prov., ~20˚12N, 104˚01E, PHU PHAN Mt. 1500-1900m, 17.v.-31.vi. 2007, M. Brancucci leg., 1 male in ABCC; LAOS-NE, Houa Phan prov., 20˚1309-19″N, 103˚5954″-104˚0003″E, 1480-1510m, PHOU PHANE Mt., 22.iv.-14.v.2008,
<normalizedToken originalValue="Vít">Vit</normalizedToken>
<normalizedToken originalValue="Kubáň">Kubaň</normalizedToken>
leg., 1 male and 1 female in NMPC; LAOS-NE, Houa Phan prov., 20˚13N 103˚59E, Ban SALUEI village, 16.vi.2009, 1350 m, at light,
<normalizedToken originalValue="Vít">Vit</normalizedToken>
.
<normalizedToken originalValue="Kubáň">Kubaň</normalizedToken>
leg., 1 male in NMPC; LAOS-NE, Houa Phan prov., 20˚12-13.5N 103˚59.5-104˚01E, Ban Saluei → Phou Pane Mt., 1340-1870 m, 1.v.-16.vi. 2009, Lao collectors leg., 1 male in NMPC; Laos, Houaphan prov., 38 km S of Sam Neua, Saluei 9.-22.5.2009,
<normalizedToken originalValue="Martinů">Martinu</normalizedToken>
lgt. 1350-1900 m, 1 male in JZCP and 1 male in RSCV; Laos, Houaphan prov., 38 km S of Sam Neua, Saluei 9.-22.5.2009,
<normalizedToken originalValue="Bednařík">Bednarik</normalizedToken>
lgt. 1350-1900 m, 1 male in RSCV.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="72" type="diagnosis of female">
<paragraph pageId="7" pageNumber="72">Diagnosis of female</paragraph>
<paragraph pageId="7" pageNumber="72">(Fig. 17). Similar to male (Fig. 16), with the following exceptions. The length of the only known female specimen is 28.0 mm (except of pygidium), while the length of males varies between 21.5-24.5 mm. Anterior margin of clypeus only very feebly upturned, nearly flat. Antennal club pentamerous. Outer margin of anterior tibia distinctly tridentate. Upper apical spur of metatibia broad, flattened, blunt apically. Tarsal claws of all pairs of legs equal in length.</paragraph>
<caption pageId="7" pageNumber="72">
<paragraph pageId="7" pageNumber="72">
Figures 16-19.
<taxonomicName class="Insecta" family="Melolonthidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) phongsali" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="phongsali" subGenus="Granida">Polyphylla (Granida) phongsali</taxonomicName>
16 Habitus of male (Hua Phan, Laos, length 22.5 mm), dorsal view 17 Habitus of female (Hua Phan, Laos, length 28.0 mm), dorsal view 18 Male genitalia, dorsal view 19 The same, right lateral view, shaded area indicates overlapping part of left paramere. Scale bar: 5 mm for Figs 18-19.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="7" pageNumber="72" type="collecting methods">
<paragraph pageId="7" pageNumber="72">Collecting methods.</paragraph>
<paragraph pageId="7" pageNumber="72">The female specimen was collected at light.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="72" type="diagnosis">
<paragraph pageId="7" pageNumber="72">Diagnosis.</paragraph>
<paragraph pageId="7" pageNumber="72">
An easily recognizable
<taxonomicName class="Insecta" family="Melolonthidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida)" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="subGenus" subGenus="Granida">Polyphylla (Granida)</taxonomicName>
species because of its maculate elytra. Antennomere 3 long and slender, more than three times longer than antennomere 2. Basal margin of pronotum is almost straight against the scutellum, while convex in other
<taxonomicName class="Insecta" family="Melolonthidae" genus="Granida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Granida" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="genus">Granida</taxonomicName>
species. It is most likely to be confused only with some
<taxonomicName class="Insecta" family="Melolonthidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) jessopi" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="jessopi" subGenus="Granida">Polyphylla (Granida) jessopi</taxonomicName>
specimens bearing the same maculate elytral pattern.
<taxonomicName class="Insecta" family="Melolonthidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) phongsali" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="phongsali" subGenus="Granida">Polyphylla (Granida) phongsali</taxonomicName>
in average larger than
<taxonomicName class="Insecta" family="Melolonthidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) jessopi" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="jessopi" subGenus="Granida">Polyphylla (Granida) jessopi</taxonomicName>
(the length of males varies between 21.5-24.5 mm versus 18.5-20.0 mm in
<taxonomicName class="Insecta" family="Melolonthidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) jessopi" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="jessopi" subGenus="Granida">Polyphylla (Granida) jessopi</taxonomicName>
). Male genitalia as in Figs 18-19.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="72" type="distribution">
<paragraph pageId="7" pageNumber="72">Distribution.</paragraph>
<paragraph pageId="7" pageNumber="72">Northern Laos.</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="73" pageId="7" pageNumber="72" type="remarks">
<paragraph pageId="7" pageNumber="72">Remarks.</paragraph>
<paragraph pageId="7" pageNumber="72">The species was originally described from a single male. Here we recorded eight additional specimens from northern Laos (Fig. 20). The specimens with altitude data were collected between 1350-1900 m a.s.l.</paragraph>
<caption pageId="8" pageNumber="73">
<paragraph pageId="8" pageNumber="73">
<pageBreakToken pageId="8" pageNumber="73" start="start">Figure</pageBreakToken>
20. Distribution of
<taxonomicName class="Insecta" family="Melolonthidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida)" order="Coleoptera" pageId="8" pageNumber="73" phylum="Arthropoda" rank="subGenus" subGenus="Granida">Polyphylla (Granida)</taxonomicName>
species in continental Asia. Because of imprecise locality data, distributional mark of
<taxonomicName class="Insecta" family="Melolonthidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) nikodymi" order="Coleoptera" pageId="8" pageNumber="73" phylum="Arthropoda" rank="species" species="nikodymi" subGenus="Granida">Polyphylla (Granida) nikodymi</taxonomicName>
in Myanmar is omitted.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>