Review of the subgenus Polyphylla (Granida) from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae) Author Sehnal, Richard Author Bezdek, Ales text ZooKeys 2011 102 65 76 http://dx.doi.org/10.3897/zookeys.102.1148 journal article http://dx.doi.org/10.3897/zookeys.102.1148 1313-2970-102-65 Polyphylla (Granida) phongsali Zidek , 2006 Figs 16-19 Polyphylla phongsali Zidek , 2006: 10, pl. 3. Type locality. "N. Laos, Phongsali, Gnoi-ou". Type material examined. Holotype (male), labeled: "N. Laos, Phongsali, Gnoi-ou, Li Jingke VI-2003 [h] // BMNH (E) 2006-162 [h] // J. ZIDEK det. 2006 [p] Polyphylla phongsali Zidek [h] HOLOTYPE [p, red label]", in BMNH. Additional material examined. LAO-NE, Hua Phan prov., ~20˚12′N, 104˚01′E, PHU PHAN Mt. 1500-1900m, 17.v.-31.vi. 2007, M. Brancucci leg., 1 male in ABCC; LAOS-NE, Houa Phan prov., 20˚13′09-19″N, 103˚59′54″-104˚00′03″E, 1480-1510m, PHOU PHANE Mt., 22.iv.-14.v.2008, Vit Kubaň leg., 1 male and 1 female in NMPC; LAOS-NE, Houa Phan prov., 20˚13′N 103˚59′E, Ban SALUEI village, 16.vi.2009, 1350 m, at light, Vit . Kubaň leg., 1 male in NMPC; LAOS-NE, Houa Phan prov., 20˚12-13.5′N 103˚59.5′-104˚01′E, Ban Saluei → Phou Pane Mt., 1340-1870 m, 1.v.-16.vi. 2009, Lao collectors leg., 1 male in NMPC; Laos, Houaphan prov., 38 km S of Sam Neua, Saluei 9.-22.5.2009, Martinu lgt. 1350-1900 m, 1 male in JZCP and 1 male in RSCV; Laos, Houaphan prov., 38 km S of Sam Neua, Saluei 9.-22.5.2009, Bednarik lgt. 1350-1900 m, 1 male in RSCV. Diagnosis of female (Fig. 17). Similar to male (Fig. 16), with the following exceptions. The length of the only known female specimen is 28.0 mm (except of pygidium), while the length of males varies between 21.5-24.5 mm. Anterior margin of clypeus only very feebly upturned, nearly flat. Antennal club pentamerous. Outer margin of anterior tibia distinctly tridentate. Upper apical spur of metatibia broad, flattened, blunt apically. Tarsal claws of all pairs of legs equal in length. Figures 16-19. Polyphylla (Granida) phongsali 16 Habitus of male (Hua Phan, Laos, length 22.5 mm), dorsal view 17 Habitus of female (Hua Phan, Laos, length 28.0 mm), dorsal view 18 Male genitalia, dorsal view 19 The same, right lateral view, shaded area indicates overlapping part of left paramere. Scale bar: 5 mm for Figs 18-19. Collecting methods. The female specimen was collected at light. Diagnosis. An easily recognizable Polyphylla (Granida) species because of its maculate elytra. Antennomere 3 long and slender, more than three times longer than antennomere 2. Basal margin of pronotum is almost straight against the scutellum, while convex in other Granida species. It is most likely to be confused only with some Polyphylla (Granida) jessopi specimens bearing the same maculate elytral pattern. Polyphylla (Granida) phongsali in average larger than Polyphylla (Granida) jessopi (the length of males varies between 21.5-24.5 mm versus 18.5-20.0 mm in Polyphylla (Granida) jessopi ). Male genitalia as in Figs 18-19. Distribution. Northern Laos. Remarks. The species was originally described from a single male. Here we recorded eight additional specimens from northern Laos (Fig. 20). The specimens with altitude data were collected between 1350-1900 m a.s.l. Figure 20. Distribution of Polyphylla (Granida) species in continental Asia. Because of imprecise locality data, distributional mark of Polyphylla (Granida) nikodymi in Myanmar is omitted.