859 lines
98 KiB
XML
859 lines
98 KiB
XML
<document id="52B7ED6D63AC68B1E628A1370EB4304B" ID-DOI="10.11646/zootaxa.3946.3.1" ID-GBIF-Dataset="21a54f6c-7b8e-4da6-81ac-05703db5bbf1" ID-ISSN="1175-5326" ID-Zenodo-Dep="244607" ID-ZooBank="91C0214A-7A3D-4DEC-9D24-7B10928AC127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461033981581" checkinUser="plazi" docAuthor="Maryan, Brad, Brennan, Ian G., Adams, Mark & Aplin, Ken P." docDate="2015" docId="03DA87B5FFE78E62FF49FBC02A0C54E5" docLanguage="en" docName="zt03946p330.pdf" docOrigin="Zootaxa 3946 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Delma hebesa Maryan, Brennan, Adams & Aplin, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="325" masterDocId="FFE3FFCDFFF38E7BFFDEFFFF2B475412" masterDocTitle="Molecular and morphological assessment of Delma australis Kluge (Squamata: Pygopodidae), with a description of a new species from the biodiversity ‘ hotspot’ of southwestern Western Australia" masterLastPageNumber="330" masterPageNumber="301" pageNumber="321" updateTime="1698574234490" updateUser="plazi">
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<mods:mods id="901E87B44B60CBDC245D86427D0E731C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6D3792F2F7E1A4F1CD9FCD03ED9350F0">
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<mods:title id="369BC3307CB36D0AEC0AA55B9C66394A">Molecular and morphological assessment of Delma australis Kluge (Squamata: Pygopodidae), with a description of a new species from the biodiversity ‘ hotspot’ of southwestern Western Australia</mods:title>
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</mods:titleInfo>
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<mods:name id="7EB514E261362FF411CBBD87930422B5" type="personal">
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<mods:roleTerm id="9A9E1BD805B30F1E629373A45225B685">Author</mods:roleTerm>
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<mods:namePart id="05E565DA70DB5491BE1E4D40B28137E7">Maryan, Brad</mods:namePart>
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<mods:roleTerm id="4AC080AAE8DB307DEF7E095BC1924B31">Author</mods:roleTerm>
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<mods:namePart id="729A3A8B39E86A37707155433B1FB034">Brennan, Ian G.</mods:namePart>
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<mods:roleTerm id="A987668D97F268B76D6DAE318CCBFAD4">Author</mods:roleTerm>
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<mods:namePart id="8E19D080504C472CA7E12CC41A13BFE5">Adams, Mark</mods:namePart>
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<mods:name id="E33F4432C94CEF17C8ECC354CEC453D0" type="personal">
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<mods:roleTerm id="B91AF111DC4E9B99445FAA25A32FFC04">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FDA1F28F1FE086AA5D8BFACCC625F70C">Aplin, Ken P.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3CC4206E6C87515986D9C3F78EE97D94">text</mods:typeOfResource>
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<mods:title id="A6FBE78C7D9D02266C18428F2072E573">Zootaxa</mods:title>
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<mods:part id="3E4F42AEC79BF6A3BE9BA53CB55F9CD6">
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<mods:date id="7CA5992A4AC90057EDBF7B6697DC3A06">2015</mods:date>
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<mods:detail id="1802E827EFD395E9195A0D953093A25A" type="volume">
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<mods:number id="FA288BAC82560E848A04F0EE00BEB8E8">3946</mods:number>
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<mods:detail id="DF456E1256C68E10ED77F20A442C4854" type="issue">
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<mods:number id="3013EE53028DD867BC1F5758E37452EA">3</mods:number>
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<mods:classification id="EE77430AFF4946F0B008E9D0A82BDA86">journal article</mods:classification>
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<mods:identifier id="105F6CD3F43D27A9A42BB12DA9B562BE" type="DOI">10.11646/zootaxa.3946.3.1</mods:identifier>
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<mods:identifier id="E64A50B4E7C4F0DF21EE75B8ACAF69A0" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03DA87B5FFE78E62FF49FBC02A0C54E5" ID-DOI="http://doi.org/10.5281/zenodo.6109331" ID-GBIF-Taxon="119608713" ID-Zenodo-Dep="6109331" LSID="urn:lsid:plazi:treatment:03DA87B5FFE78E62FF49FBC02A0C54E5" httpUri="http://treatment.plazi.org/id/03DA87B5FFE78E62FF49FBC02A0C54E5" lastPageId="25" lastPageNumber="325" pageId="20" pageNumber="321">
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<subSubSection id="C3696528FFE78E6FFF49FBC02A0E508D" pageId="20" pageNumber="321" type="nomenclature">
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<paragraph id="8BCC36A3FFE78E6FFF49FBC02AE25048" blockId="20.[151,421,1087,1183]" box="[151,421,1087,1114]" pageId="20" pageNumber="321">
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<heading id="D08481CFFFE78E6FFF49FBC02AE25048" bold="true" box="[151,421,1087,1114]" fontSize="11" level="1" pageId="20" pageNumber="321" reason="1">
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<emphasis id="B907EAB1FFE78E6FFF49FBC02AE25048" bold="true" box="[151,421,1087,1114]" pageId="20" pageNumber="321">
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<taxonomicName id="4C734D20FFE78E6FFF49FBC02A06504B" box="[151,321,1087,1113]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
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<emphasis id="B907EAB1FFE78E6FFF49FBC02A06504B" bold="true" box="[151,321,1087,1113]" italics="true" pageId="20" pageNumber="321">Delma hebesa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23457CAFFE78E6FFE96FBBF2AE25048" box="[328,421,1088,1114]" pageId="20" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BCC36A3FFE78E6FFF49FB9D2A6B5069" blockId="20.[151,421,1087,1183]" box="[151,300,1122,1147]" pageId="20" pageNumber="321">
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||
Heath
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<taxonomicName id="4C734D20FFE78E6FFF3FFB9D2A6B5069" box="[225,300,1122,1147]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="genus">Delma</taxonomicName>
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</paragraph>
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||
<paragraph id="8BCC36A3FFE78E6FFF49FB792A0E508D" blockId="20.[151,421,1087,1183]" box="[151,329,1158,1183]" pageId="20" pageNumber="321">
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(
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<figureCitation id="13482A26FFE78E6FFF41FB792BA9508D" box="[159,238,1158,1183]" captionStart="FIGURE 9" captionStartId="21.[151,250,1057,1079]" captionTargetBox="[152,1434,193,1027]" captionTargetId="figure@21.[152,1435,193,1036]" captionTargetPageId="21" captionText="FIGURE 9. Preserved holotype WAM R 144237 of D. hebesa sp. nov. and head scalation in (A) dorsal and (B) lateral views." httpUri="https://zenodo.org/record/244616/files/figure.png" pageId="20" pageNumber="321">Figs. 9</figureCitation>
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,
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<figureCitation id="13482A26FFE78E6FFF24FB782A5F508D" box="[250,280,1159,1183]" captionStart="FIGURE 10" captionStartId="22.[151,250,1688,1710]" captionTargetBox="[398,1187,665,1661]" captionTargetId="figure@22.[393,1193,661,1667]" captionTargetPageId="22" captionText="FIGURE 10. Adult D. hebesa sp. nov. photographed in life from (A) Scaddan and (B) Mettler Lake, Western Australia (images ‒ B. Maryan)." httpUri="https://zenodo.org/record/244617/files/figure.png" pageId="20" pageNumber="321">10</figureCitation>
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, 11)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3696528FFE78E6FFF49FB312FA85214" pageId="20" pageNumber="321" type="materials_examined">
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<paragraph id="8BCC36A3FFE78E6FFF49FB31286B513D" blockId="20.[151,1436,1230,2011]" pageId="20" pageNumber="321">
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<emphasis id="B907EAB1FFE78E6FFF49FB312A4A50F5" bold="true" box="[151,269,1230,1255]" pageId="20" pageNumber="321">
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<typeStatus id="54C88801FFE78E6FFF49FB312A4F50F5" box="[151,264,1230,1255]" pageId="20" pageNumber="321" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectionCode id="ED62AE66FFE78E6FFEC3FB302A2550F5" LSID="urn:lsid:biocol.org:col:34981" box="[285,354,1231,1255]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="321">WAM</collectionCode>
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R144237, male, Bandalup Hill, Ravensthorpe Range (
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<geoCoordinate id="EE475064FFE78E6FFBD8FB302FC250F5" box="[1030,1157,1231,1255]" direction="south" orientation="latitude" pageId="20" pageNumber="321" precision="15" value="-33.674725">33º40'29"S</geoCoordinate>
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<geoCoordinate id="EE475064FFE78E6FFB4AFB312E6350F5" box="[1172,1316,1230,1255]" direction="east" orientation="longitude" pageId="20" pageNumber="321" precision="15" value="120.39833">120º23'54"E</geoCoordinate>
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), Western
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<collectingCountry id="F3647633FFE78E6FFF49FB0D2A465119" box="[151,257,1266,1291]" name="Australia" pageId="20" pageNumber="321">Australia</collectingCountry>
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,
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<collectingCountry id="F3647633FFE78E6FFED3FB0D2A305119" box="[269,375,1266,1291]" name="Australia" pageId="20" pageNumber="321">Australia</collectingCountry>
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, collected by R. Teale & G. Harold,
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<date id="FFCD1063FFE78E6FFCC4FB0C28995119" box="[794,990,1266,1291]" pageId="20" pageNumber="321" value="2000-10-14">14 October 2000</date>
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. Fixed in 10% formalin, stored in 70% ethanol, liver sample stored in ‒80ºC ultrafreezer at
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<collectionCode id="ED62AE66FFE78E6FFD3EFAE8286F513D" LSID="urn:lsid:biocol.org:col:34981" box="[736,808,1303,1327]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="321">WAM</collectionCode>
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.
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</paragraph>
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||
<paragraph id="8BCC36A3FFE78E6FFF19FAC52A8751F1" blockId="20.[151,1436,1230,2011]" pageId="20" pageNumber="321">
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||
<emphasis id="B907EAB1FFE78E6FFF19FAC52A0F5141" bold="true" box="[199,328,1338,1363]" pageId="20" pageNumber="321">
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||
<typeStatus id="54C88801FFE78E6FFF19FAC52A045141" box="[199,323,1338,1363]" pageId="20" pageNumber="321" type="paratype">Paratypes</typeStatus>
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||
:
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</emphasis>
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||
All from Western
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<collectingCountry id="F3647633FFE78E6FFDF7FAC529D35141" box="[553,660,1338,1363]" name="Australia" pageId="20" pageNumber="321">Australia</collectingCountry>
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||
.
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<collectionCode id="ED62AE66FFE78E6FFD7CFAC429A05141" LSID="urn:lsid:biocol.org:col:34981" box="[674,743,1339,1363]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="321">WAM</collectionCode>
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R129674, male,
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<quantity id="4C8B9B46FFE78E6FFC6CFAC42F435141" box="[946,1028,1338,1363]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="321" unit="km" value="3.8">3.8 km</quantity>
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W of Kundip, (
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||
<geoCoordinate id="EE475064FFE78E6FFB1BFAC42E5C5141" box="[1221,1307,1339,1363]" direction="south" orientation="latitude" pageId="20" pageNumber="321" precision="925" value="-33.683334">33º41'S</geoCoordinate>
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||
<geoCoordinate id="EE475064FFE78E6FFAF8FAC42ED75141" box="[1318,1424,1339,1363]" direction="east" orientation="longitude" pageId="20" pageNumber="321" precision="925" value="120.15">120º09'E</geoCoordinate>
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);
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<collectionCode id="ED62AE66FFE78E6FFF49FAA02B9B5165" LSID="urn:lsid:biocol.org:col:34981" box="[151,220,1375,1399]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="321">WAM</collectionCode>
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R131902, female, Hellfire Bay, Cape Le Grand National Park, (
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<geoCoordinate id="EE475064FFE78E6FFC69FAA12F725165" box="[951,1077,1374,1399]" direction="south" orientation="latitude" pageId="20" pageNumber="321" precision="15" value="-34.004166">34º00'15"S</geoCoordinate>
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<geoCoordinate id="EE475064FFE78E6FFBE2FAA02F8A5165" box="[1084,1229,1375,1399]" direction="east" orientation="longitude" pageId="20" pageNumber="321" precision="15" value="122.17222">122º10'20"E</geoCoordinate>
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);
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<collectionCode id="ED62AE66FFE78E6FFB3CFAA02E605165" LSID="urn:lsid:biocol.org:col:34981" box="[1250,1319,1375,1399]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="321">WAM</collectionCode>
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R132154, female, Duke of Orleans Bay, Wharton Beach, (
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<geoCoordinate id="EE475064FFE78E6FFD3CFA7D287E5189" box="[738,825,1410,1435]" direction="south" orientation="latitude" pageId="20" pageNumber="321" precision="925" value="-33.933334">33º56'S</geoCoordinate>
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||
<geoCoordinate id="EE475064FFE78E6FFC99FA7C28F75189" box="[839,944,1411,1435]" direction="east" orientation="longitude" pageId="20" pageNumber="321" precision="925" value="122.55">122º33'E</geoCoordinate>
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);
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<collectionCode id="ED62AE66FFE78E6FFC15FA7C2F485189" LSID="urn:lsid:biocol.org:col:34981" box="[971,1039,1411,1435]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="321">WAM</collectionCode>
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R144238, male, same details as
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<typeStatus id="54C88801FFE78E6FFF49FA592BBA51AD" box="[151,253,1446,1471]" pageId="20" pageNumber="321" type="holotype">holotype</typeStatus>
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||
;
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||
<collectionCode id="ED62AE66FFE78E6FFECFFA582A1151AD" LSID="urn:lsid:biocol.org:col:34981" box="[273,342,1447,1471]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="321">WAM</collectionCode>
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||
R154234, male, Kundip, (
|
||
<geoCoordinate id="EE475064FFE78E6FFD7AFA59286651AD" box="[676,801,1446,1471]" direction="south" orientation="latitude" pageId="20" pageNumber="321" precision="15" value="-33.67389">33º40'26"S</geoCoordinate>
|
||
<geoCoordinate id="EE475064FFE78E6FFCECFA59288751AD" box="[818,960,1446,1471]" direction="east" orientation="longitude" pageId="20" pageNumber="321" precision="15" value="120.19583">120º11'45"E</geoCoordinate>
|
||
);
|
||
<collectionCode id="ED62AE66FFE78E6FFC00FA582F6451AD" LSID="urn:lsid:biocol.org:col:34981" box="[990,1059,1447,1471]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="321">WAM</collectionCode>
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R156978, male, Canal Rocks, (
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<geoCoordinate id="EE475064FFE78E6FFF41FA352A5A51F1" box="[159,285,1482,1507]" direction="south" orientation="latitude" pageId="20" pageNumber="321" precision="15" value="-33.66278">33º39'46"S</geoCoordinate>
|
||
<geoCoordinate id="EE475064FFE78E6FFEFAFA352AF351F1" box="[292,436,1482,1507]" direction="east" orientation="longitude" pageId="20" pageNumber="321" precision="15" value="115.0125">115º00'45"E</geoCoordinate>
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||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE78E6FFF19FA112FA85214" blockId="20.[151,1436,1230,2011]" box="[199,1263,1518,1543]" pageId="20" pageNumber="321">
|
||
<emphasis id="B907EAB1FFE78E6FFF19FA112AB35215" bold="true" box="[199,500,1518,1543]" pageId="20" pageNumber="321">Other material examined</emphasis>
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||
: A full list of material examined is provided at the end of the paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3696528FFE78E6EFF19F9ED2EC550F1" lastPageId="21" lastPageNumber="322" pageId="20" pageNumber="321" type="diagnosis">
|
||
<paragraph id="8BCC36A3FFE78E6FFF19F9ED29765310" blockId="20.[151,1436,1230,2011]" pageId="20" pageNumber="321">
|
||
<emphasis id="B907EAB1FFE78E6FFF19F9ED2A065239" bold="true" box="[199,321,1554,1579]" pageId="20" pageNumber="321">Diagnosis.</emphasis>
|
||
A small species of
|
||
<taxonomicName id="4C734D20FFE78E6FFDFDF9ED29295239" box="[547,622,1554,1579]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="genus">
|
||
<emphasis id="B907EAB1FFE78E6FFDFDF9ED29295239" box="[547,622,1554,1579]" italics="true" pageId="20" pageNumber="321">Delma</emphasis>
|
||
</taxonomicName>
|
||
(SVL to
|
||
<quantity id="4C8B9B46FFE78E6FFD05F9EC286B5239" box="[731,812,1555,1579]" metricMagnitude="-2" metricUnit="m" metricValue="7.9" pageId="20" pageNumber="321" unit="mm" value="79.0">79 mm</quantity>
|
||
) with: ventral scales not markedly larger than adjacent lateral scales; one pair of supranasals; modally 18 midbody scales; modally 10 hindlimb scales in both sexes; 73‒92 ventral scales (males average 76.8, females 85.5); six upper labials with fourth typically below eye; loreal scale row typically interrupted by a ventral extension of supraloreal scale that contacts upper labials; essentially unpatterned head, sometimes with weak dark variegations on sides of head and indistinct narrow bars or smudges on labial scales, nape and forebody. Diagnostic differences between
|
||
<taxonomicName id="4C734D20FFE78E6FFC42F9382F4A52CD" box="[924,1037,1734,1759]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE78E6FFC42F9382F4A52CD" box="[924,1037,1734,1759]" italics="true" pageId="20" pageNumber="321">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE78E6FFBCBF9392F3752CD" box="[1045,1136,1734,1759]" pageId="20" pageNumber="321" rank="species">
|
||
<emphasis id="B907EAB1FFE78E6FFBCBF9392F3752CD" bold="true" box="[1045,1136,1734,1759]" pageId="20" pageNumber="321">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C734D20FFE78E6FFB76F9382E6A52CD" box="[1192,1325,1734,1759]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFE78E6FFB76F9382E6A52CD" box="[1192,1325,1734,1759]" italics="true" pageId="20" pageNumber="321">D. australis</emphasis>
|
||
</taxonomicName>
|
||
are listed under the foregoing species account.
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE78E6FFF19F8F128DA5381" blockId="20.[151,1436,1230,2011]" pageId="20" pageNumber="321">
|
||
<taxonomicName id="4C734D20FFE78E6FFF19F8F12A2E5335" box="[199,361,1806,1831]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE78E6FFF19F8F12A2E5335" box="[199,361,1806,1831]" italics="true" pageId="20" pageNumber="321">Delma hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE78E6FFEACF8F12A895335" box="[370,462,1806,1831]" pageId="20" pageNumber="321" rank="species">
|
||
<emphasis id="B907EAB1FFE78E6FFEACF8F12A895335" bold="true" box="[370,462,1806,1831]" pageId="20" pageNumber="321">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C734D20FFE78E6FFDB7F8F029AA5334" box="[617,749,1807,1830]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="torquata">
|
||
<emphasis id="B907EAB1FFE78E6FFDB7F8F029AA5334" box="[617,749,1807,1830]" italics="true" pageId="20" pageNumber="321">D. torquata</emphasis>
|
||
</taxonomicName>
|
||
from southeastern Queensland in having a larger adult size (SVL to
|
||
<quantity id="4C8B9B46FFE78E6FFF24F8CC2A0E5358" box="[250,329,1843,1867]" metricMagnitude="-2" metricUnit="m" metricValue="7.9" pageId="20" pageNumber="321" unit="mm" value="79.0">79 mm</quantity>
|
||
<emphasis id="B907EAB1FFE78E6FFE8EF8CB2AD05359" box="[336,407,1844,1867]" italics="true" pageId="20" pageNumber="321">versus</emphasis>
|
||
to
|
||
<quantity id="4C8B9B46FFE78E6FFE62F8CD29575359" box="[444,528,1842,1867]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="20" pageNumber="321" unit="mm" value="63.0">63 mm</quantity>
|
||
), three precloacal scales (
|
||
<emphasis id="B907EAB1FFE78E6FFCEFF8CB283E5359" box="[817,889,1844,1867]" italics="true" pageId="20" pageNumber="321">versus</emphasis>
|
||
two), the fourth upper labial typically below the eye (
|
||
<emphasis id="B907EAB1FFE78E6FFF0BF8A82A5B537C" box="[213,284,1879,1902]" italics="true" pageId="20" pageNumber="321">versus</emphasis>
|
||
typically the third below the eye), modally 18 midbody scale rows (
|
||
<emphasis id="B907EAB1FFE78E6FFBB2F8A82FF3537C" box="[1132,1204,1879,1902]" italics="true" pageId="20" pageNumber="321">versus</emphasis>
|
||
16) and only dark variegations (if present) on head and neck (
|
||
<emphasis id="B907EAB1FFE78E6FFDA2F88329845381" box="[636,707,1916,1939]" italics="true" pageId="20" pageNumber="321">versus</emphasis>
|
||
broad dark bands).
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE78E6FFF19F8612E0B53C9" blockId="20.[151,1436,1230,2011]" pageId="20" pageNumber="321">
|
||
<taxonomicName id="4C734D20FFE78E6FFF19F8612A2C53A5" box="[199,363,1950,1975]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE78E6FFF19F8612A2C53A5" box="[199,363,1950,1975]" italics="true" pageId="20" pageNumber="321">Delma hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE78E6FFEA8F8612A9253A5" box="[374,469,1950,1975]" pageId="20" pageNumber="321" rank="species">
|
||
<emphasis id="B907EAB1FFE78E6FFEA8F8612A9253A5" bold="true" box="[374,469,1950,1975]" pageId="20" pageNumber="321">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from all other Australian species (except
|
||
<taxonomicName id="4C734D20FFE78E6FFBC1F8602FEF53A5" box="[1055,1192,1950,1975]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFE78E6FFBC1F8602FEF53A5" box="[1055,1192,1950,1975]" italics="true" pageId="20" pageNumber="321">D. australis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C734D20FFE78E6FFB65F8602E0653A4" box="[1211,1345,1951,1974]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="torquata">
|
||
<emphasis id="B907EAB1FFE78E6FFB65F8602E0653A4" box="[1211,1345,1951,1974]" italics="true" pageId="20" pageNumber="321">D. torquata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C734D20FFE78E6FFA5FF8602BB953C9" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="concinna">
|
||
<emphasis id="B907EAB1FFE78E6FFA5FF8602BB953C9" italics="true" pageId="20" pageNumber="321">D. concinna</emphasis>
|
||
</taxonomicName>
|
||
) in having ventral scales not markedly larger than adjacent lateral scales (
|
||
<emphasis id="B907EAB1FFE78E6FFBE5F83B2FC453C9" box="[1083,1155,1988,2011]" italics="true" pageId="20" pageNumber="321">versus</emphasis>
|
||
markedly larger).
|
||
</paragraph>
|
||
<caption id="DF0C662BFFE68E6EFF49FBDE2ECC5025" box="[151,1419,1057,1079]" httpUri="https://zenodo.org/record/244616/files/figure.png" pageId="21" pageNumber="322" targetBox="[152,1434,193,1027]" targetPageId="21">
|
||
<paragraph id="8BCC36A3FFE68E6EFF49FBDE2ECC5025" blockId="21.[151,1419,1057,1079]" box="[151,1419,1057,1079]" pageId="21" pageNumber="322">
|
||
<emphasis id="B907EAB1FFE68E6EFF49FBDE2A555025" bold="true" box="[151,274,1057,1079]" pageId="21" pageNumber="322">FIGURE 9.</emphasis>
|
||
Preserved holotype WAM R144237 of
|
||
<taxonomicName id="4C734D20FFE68E6EFD7AFBDD284E5025" box="[676,777,1057,1079]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="322" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE68E6EFD7AFBDD284E5025" box="[676,777,1057,1079]" italics="true" pageId="21" pageNumber="322">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE68E6EFCCEFBDE28275025" box="[784,864,1057,1079]" pageId="21" pageNumber="322" rank="species">
|
||
<emphasis id="B907EAB1FFE68E6EFCCEFBDE28275025" bold="true" box="[784,864,1057,1079]" pageId="21" pageNumber="322">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and head scalation in (A) dorsal and (B) lateral views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC36A3FFE68E6EFF19FBA12EC550F1" blockId="21.[151,1437,1118,2008]" pageId="21" pageNumber="322">
|
||
<taxonomicName id="4C734D20FFE68E6EFF19FBA12A215065" box="[199,358,1118,1143]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="322" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE68E6EFF19FBA12A215065" box="[199,358,1118,1143]" italics="true" pageId="21" pageNumber="322">Delma hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE68E6EFEB2FBA12A815065" box="[364,454,1118,1143]" pageId="21" pageNumber="322" rank="species">
|
||
<emphasis id="B907EAB1FFE68E6EFEB2FBA12A815065" bold="true" box="[364,454,1118,1143]" pageId="21" pageNumber="322">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C734D20FFE68E6EFD86FB9F29825065" box="[600,709,1119,1143]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="322" phylum="Chordata" rank="species" species="fraseri">
|
||
<emphasis id="B907EAB1FFE68E6EFD86FB9F29825065" box="[600,709,1119,1143]" italics="true" pageId="21" pageNumber="322">D. fraseri</emphasis>
|
||
</taxonomicName>
|
||
with which it occurs in sympatry (see below) in having a smaller adult size (SVL to
|
||
<quantity id="4C8B9B46FFE68E6EFEB4FB7C2AFE5088" box="[362,441,1155,1179]" metricMagnitude="-2" metricUnit="m" metricValue="7.9" pageId="21" pageNumber="322" unit="mm" value="79.0">79 mm</quantity>
|
||
<emphasis id="B907EAB1FFE68E6EFE61FB7B29415089" box="[447,518,1156,1179]" italics="true" pageId="21" pageNumber="322">versus</emphasis>
|
||
to
|
||
<quantity id="4C8B9B46FFE68E6EFDF4FB7C29CE5088" box="[554,649,1155,1179]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="21" pageNumber="322" unit="mm" value="140.0">140 mm</quantity>
|
||
,
|
||
<bibRefCitation id="EFE24B52FFE68E6EFD4AFB7D28085089" author="Bush" box="[660,847,1154,1179]" pageId="21" pageNumber="322" refString="Bush, B., Maryan, B., Browne-Cooper & Robinson, D. (2007) Reptiles and Frogs in the bush: Southwestern Australia. University of Western Australia Press, Perth, 302 pp." type="book" year="2007">
|
||
Bush
|
||
<emphasis id="B907EAB1FFE68E6EFD0AFB7B284A5089" box="[724,781,1154,1179]" italics="true" pageId="21" pageNumber="322">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
), one pair of supranasals (
|
||
<emphasis id="B907EAB1FFE68E6EFBADFB7B2FFC5089" box="[1139,1211,1156,1179]" italics="true" pageId="21" pageNumber="322">versus</emphasis>
|
||
two pairs), modally 18 midbody scale rows (
|
||
<emphasis id="B907EAB1FFE68E6EFE77FB572AB750AD" box="[425,496,1192,1215]" italics="true" pageId="21" pageNumber="322">versus</emphasis>
|
||
16), ventral scales not markedly larger than adjacent lateral scales (
|
||
<emphasis id="B907EAB1FFE68E6EFB3AFB572E6B50AD" box="[1252,1324,1192,1215]" italics="true" pageId="21" pageNumber="322">versus</emphasis>
|
||
markedly larger) and only dark variegations (if present) on head and neck (
|
||
<emphasis id="B907EAB1FFE68E6EFCB1FB3328F050F1" box="[879,951,1228,1251]" italics="true" pageId="21" pageNumber="322">versus</emphasis>
|
||
broad dark bands, often faded in adults).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3696528FFE68E6CFF19FB112A7651E1" lastPageId="23" lastPageNumber="324" pageId="21" pageNumber="322" type="description">
|
||
<paragraph id="8BCC36A3FFE68E6EFF19FB112F725139" blockId="21.[151,1437,1118,2008]" pageId="21" pageNumber="322">
|
||
<emphasis id="B907EAB1FFE68E6EFF19FB112A985115" bold="true" box="[199,479,1262,1287]" pageId="21" pageNumber="322">
|
||
Description of
|
||
<typeStatus id="54C88801FFE68E6EFEA6FB112A985115" box="[376,479,1262,1287]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13482A26FFE68E6EFE31FB1129745115" box="[495,563,1262,1287]" captionStart="FIGURE 9" captionStartId="21.[151,250,1057,1079]" captionTargetBox="[152,1434,193,1027]" captionTargetId="figure@21.[152,1435,193,1036]" captionTargetPageId="21" captionText="FIGURE 9. Preserved holotype WAM R 144237 of D. hebesa sp. nov. and head scalation in (A) dorsal and (B) lateral views." httpUri="https://zenodo.org/record/244616/files/figure.png" pageId="21" pageNumber="322">Fig. 9</figureCitation>
|
||
). Measurements (in mm) and meristic values: SVL 57, HD 3.4, HL 6.9, HW 4.5, HLL 3.3, ML 4.9, RL 0.9, RW 1.3, SL 2.6, EW 1.1, HLS 10,
|
||
<collectingCountry id="F3647633FFE68E6EFCA4FAEC28F25139" box="[890,949,1299,1323]" name="Egypt" pageId="21" pageNumber="322">MSR</collectingCountry>
|
||
18, VE 75.
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE68E6EFF19FAC9288A51CD" blockId="21.[151,1437,1118,2008]" pageId="21" pageNumber="322">Head short and narrowing gradually forward of eyes, of equal width to body posteriorly; obvious tympanic aperture, indicated by round opening posterior to corner of mouth, opening is more narrow on right side; snout moderately long and rounded in dorsal and lateral profiles; body moderately robust of equal width and round in cross-section; hindlimbs visible as well-developed elongate, rounded flaps adpressed to body at lateral extremes of vent; tail relatively short, tapering very gradually distally to a pointed tip.</paragraph>
|
||
<paragraph id="8BCC36A3FFE68E6EFF19FA15280F539D" blockId="21.[151,1437,1118,2008]" pageId="21" pageNumber="322">
|
||
Head scales smooth, non-imbricate and heterogeneous; rostral rounded anteriorly, wider than long, with obtuse apex projecting between supranasals; one pair of supranasals in broad contact, angled backwards posteromedially behind rostral and in short contact with first upper labial; nostril positioned on posterior junction of supranasal with first upper labial and postnasal; one postnasal, wider than high, angled posteroventrally and in short contact with second upper labial; prefrontals symmetrical and in broad contact; supraloreal much higher than wide, and in broad contact with second upper labial; four loreals, the anterior most slightly larger; five supraciliaries, first and fourth the smallest, second and fifth the largest; two supraoculars, first slightly larger than second; two frontals, the anterior most slightly wider and larger; two parietals, one on left side is slightly larger and longer; occipital present; two upper temporals; six upper labials, fourth the widest and positioned below eye, third the smallest, on left side a small scale intersects suture between second and third upper labial; five lower labials, second the largest and widest, fifth the smallest; mental wider than long with suture starting half-way along first upper labial. General form of head and details of scalation illustrated in
|
||
<figureCitation id="13482A26FFE68E6EFD15F889284B539D" box="[715,780,1910,1935]" captionStart="FIGURE 9" captionStartId="21.[151,250,1057,1079]" captionTargetBox="[152,1434,193,1027]" captionTargetId="figure@21.[152,1435,193,1036]" captionTargetPageId="21" captionText="FIGURE 9. Preserved holotype WAM R 144237 of D. hebesa sp. nov. and head scalation in (A) dorsal and (B) lateral views." httpUri="https://zenodo.org/record/244616/files/figure.png" pageId="21" pageNumber="322">Fig. 9</figureCitation>
|
||
A, B.
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE68E6EFF19F8652F3253C5" blockId="21.[151,1437,1118,2008]" pageId="21" pageNumber="322">Body scales smooth, non-imbricate, homogeneous, and arranged in parallel longitudinal rows; ventral scales only very marginally wider than the adjacent lateral body scales; three precloacal scales.</paragraph>
|
||
<paragraph id="8BCC36A3FFE58E6DFF19FF67299154EA" blockId="22.[151,1436,151,608]" pageId="22" pageNumber="323">
|
||
In preservative, due to the leaching of colours in ethanol after 14 years, the
|
||
<typeStatus id="54C88801FFE58E6DFBC0FF682FC754A2" box="[1054,1152,151,176]" pageId="22" pageNumber="323" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13482A26FFE58E6DFB4EFF682F9454A2" box="[1168,1235,151,176]" captionStart="FIGURE 9" captionStartId="21.[151,250,1057,1079]" captionTargetBox="[152,1434,193,1027]" captionTargetId="figure@21.[152,1435,193,1036]" captionTargetPageId="21" captionText="FIGURE 9. Preserved holotype WAM R 144237 of D. hebesa sp. nov. and head scalation in (A) dorsal and (B) lateral views." httpUri="https://zenodo.org/record/244616/files/figure.png" pageId="22" pageNumber="323">Fig. 9</figureCitation>
|
||
) has lost the grey colouration on the dorsal surface which is now light brown. All other aspects of dark bars or smudges on lower labials and obscure variegations on body remained.
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE58E6DFF19FEFB28765672" blockId="22.[151,1436,151,608]" pageId="22" pageNumber="323">
|
||
<emphasis id="B907EAB1FFE58E6DFF19FEFB2AEB550F" bold="true" box="[199,428,260,285]" pageId="22" pageNumber="323">Colouration in life.</emphasis>
|
||
The following description of colouration in life is based on
|
||
<figureCitation id="13482A26FFE58E6DFB81FEFB2FFD550E" box="[1119,1210,260,285]" captionStart="FIGURE 10" captionStartId="22.[151,250,1688,1710]" captionTargetBox="[398,1187,665,1661]" captionTargetId="figure@22.[393,1193,661,1667]" captionTargetPageId="22" captionText="FIGURE 10. Adult D. hebesa sp. nov. photographed in life from (A) Scaddan and (B) Mettler Lake, Western Australia (images ‒ B. Maryan)." httpUri="https://zenodo.org/record/244617/files/figure.png" pageId="22" pageNumber="323">Figs. 10</figureCitation>
|
||
A, B, 11A and field observations of
|
||
<taxonomicName id="4C734D20FFE58E6DFE95FED62AFB5552" box="[331,444,295,320]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="323" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE58E6DFE95FED62AFB5552" box="[331,444,295,320]" italics="true" pageId="22" pageNumber="323">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE58E6DFE1AFED829595552" box="[452,542,295,320]" pageId="22" pageNumber="323" rank="species">
|
||
<emphasis id="B907EAB1FFE58E6DFE1AFED829595552" bold="true" box="[452,542,295,320]" pageId="22" pageNumber="323">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from the Esperance Plains. Top of head bluish to light grey and unpatterned or with weak dark variegations. Lower labials whitish with blackish bars or smudges, centred on sutures of mental and anterior two lower labial scales, corner of mouth, grading to obscure variegations or weak bars around ear opening and on lateral scales of forebody. There is only very weak ventral extension of black bars or smudges on to chin and throat. Dorsal body surface bluish to light grey anteriorly, gradually merging to light brown body than to light grey tail. Lower flanks are slightly pinkish to reddish brown. Dorsal surface is uniform, except for some indication of obscure dark spots or variegations, particularly on some sutures of body scales. Ventral surface under head and along body whitish, with blackish variegations, becoming less pigmented under tail. The longest tail measured was
|
||
<quantity id="4C8B9B46FFE58E6DFEE2FDB72ADE564D" box="[316,409,584,608]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="323" unit="mm" value="140.0">140 mm</quantity>
|
||
(208% of SVL) on WAM R131902.
|
||
</paragraph>
|
||
<caption id="DF0C662BFFE58E6DFF49F9672A3752DF" httpUri="https://zenodo.org/record/244617/files/figure.png" pageId="22" pageNumber="323" targetBox="[398,1187,665,1661]" targetPageId="22">
|
||
<paragraph id="8BCC36A3FFE58E6DFF49F9672A3752DF" blockId="22.[151,1435,1688,1741]" pageId="22" pageNumber="323">
|
||
<emphasis id="B907EAB1FFE58E6DFF49F9672A6352BC" bold="true" box="[151,292,1688,1710]" pageId="22" pageNumber="323">FIGURE 10.</emphasis>
|
||
Adult
|
||
<taxonomicName id="4C734D20FFE58E6DFEACF9652A9C52BC" box="[370,475,1688,1711]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="323" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE58E6DFEACF9652A9C52BC" box="[370,475,1688,1711]" italics="true" pageId="22" pageNumber="323">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE58E6DFE38F967297D52BC" box="[486,570,1688,1710]" pageId="22" pageNumber="323" rank="species">
|
||
<emphasis id="B907EAB1FFE58E6DFE38F967297D52BC" bold="true" box="[486,570,1688,1710]" pageId="22" pageNumber="323">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
photographed in life from (A) Scaddan and (B) Mettler Lake, Western Australia (images‒B. Maryan).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC36A3FFE58E6DFF19F90A2F785323" blockId="22.[151,1437,1780,2021]" pageId="22" pageNumber="323">
|
||
For comparisons of both
|
||
<taxonomicName id="4C734D20FFE58E6DFE37F909291B531F" box="[489,604,1780,1805]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="323" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE58E6DFE37F909291B531F" box="[489,604,1780,1805]" italics="true" pageId="22" pageNumber="323">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B907EAB1FFE58E6DFDBAF90B2987531F" bold="true" box="[612,704,1780,1805]" pageId="22" pageNumber="323">
|
||
<taxonomicNameLabel id="A23457CAFFE58E6DFDBAF90B2987531F" box="[612,704,1780,1805]" pageId="22" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C734D20FFE58E6DFD25F90928C5531F" box="[763,898,1780,1805]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="323" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFE58E6DFD25F90928C5531F" box="[763,898,1780,1805]" italics="true" pageId="22" pageNumber="323">D. australis</emphasis>
|
||
</taxonomicName>
|
||
in life see
|
||
<bibRefCitation id="EFE24B52FFE58E6DFBDBF90B295F5323" author="Thompson" pageId="22" pageNumber="323" refString="Thompson, S. A. & Thompson, G. G. (2006) Reptiles of the Western Australian Goldfields. Goldfields Environmental Management Group, Kalgoorlie-Boulder, 140 pp." type="book" year="2006">Thompson & Thompson (2006: 44, ‘grey morph’ and ‘brown morph’)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE24B52FFE58E6DFDFDF8E729AF5323" author="Bush" box="[547,744,1816,1841]" pageId="22" pageNumber="323" refString="Bush, B., Maryan, B., Browne-Cooper & Robinson, D. (2007) Reptiles and Frogs in the bush: Southwestern Australia. University of Western Australia Press, Perth, 302 pp." type="book" year="2007">
|
||
Bush
|
||
<emphasis id="B907EAB1FFE58E6DFDBAF8E529DB5323" box="[612,668,1816,1841]" italics="true" pageId="22" pageNumber="323">et al.</emphasis>
|
||
(2007
|
||
</bibRefCitation>
|
||
: 135) and
|
||
<bibRefCitation id="EFE24B52FFE58E6DFC83F8E72F7C5323" author="Henkel" box="[861,1083,1816,1841]" pageId="22" pageNumber="323" refString="Henkel, F. W. (2010) Terralog Vol. 10: Geckos of Australia. Edition Chimaira, Frankfurt am Main, Germany, 160 pp." type="book" year="2010">Henkel (2010: 141)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE58E6CFF19F8C329F7550F" blockId="22.[151,1437,1780,2021]" lastBlockId="23.[151,1437,151,285]" lastPageId="23" lastPageNumber="324" pageId="22" pageNumber="323">
|
||
<emphasis id="B907EAB1FFE58E6DFF19F8C32A7C5347" bold="true" box="[199,315,1852,1877]" pageId="22" pageNumber="323">Variation.</emphasis>
|
||
<tableCitation id="C6F10318FFE58E6DFE9BF8C32ADF5347" box="[325,408,1852,1877]" captionStart="TABLE 5" captionStartId="8.[151,239,1329,1352]" captionText="TABLE 5. Descriptive statistics for D. hebesa sp. nov. and each of three regional populations of D. australis. Values shown are the sample size (N), the mean, ± one standard deviation (S. D.), and the minimum (Min.) and maximum (Max.) values. See Material and Methods for abbreviations of measurements and counts." pageId="22" pageNumber="323">Table 5</tableCitation>
|
||
presents the means, standard deviations and ranges of the characters counted and measured for each sex of
|
||
<taxonomicName id="4C734D20FFE58E6DFE97F89D2AFD536B" box="[329,442,1888,1913]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="323" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE58E6DFE97F89D2AFD536B" box="[329,442,1888,1913]" italics="true" pageId="22" pageNumber="323">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE58E6DFE1DF89F295A536B" box="[451,541,1888,1913]" pageId="22" pageNumber="323" rank="species">
|
||
<emphasis id="B907EAB1FFE58E6DFE1DF89F295A536B" bold="true" box="[451,541,1888,1913]" pageId="22" pageNumber="323">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<tableCitation id="C6F10318FFE58E6DFDFBF89F293D536B" box="[549,634,1888,1913]" captionStart="TABLE 8" captionStartId="23.[151,239,332,355]" captionTargetBox="[159,1410,414,754]" captionTargetPageId="23" captionText="TABLE 8. Individual measurements (mm) and meristic counts for the type series of D. hebesa sp. nov. The holotype is marked with an asterisk." httpUri="http://table.plazi.org/id/DF0C662BFFE48E6CFF49FEB32ADB5591" pageId="22" pageNumber="323" tableUuid="DF0C662BFFE48E6CFF49FEB32ADB5591">Table 8</tableCitation>
|
||
presents the individual measurements and meristic counts for the
|
||
<typeStatus id="54C88801FFE58E6DFAB5F89E2EDB536B" box="[1387,1436,1889,1913]" pageId="22" pageNumber="323">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C734D20FFE58E6DFF23F8792A29538F" box="[253,366,1924,1949]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="323" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE58E6DFF23F8792A29538F" box="[253,366,1924,1949]" italics="true" pageId="22" pageNumber="323">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE58E6DFEA8F87B2A97538F" box="[374,464,1924,1949]" pageId="22" pageNumber="323" rank="species">
|
||
<emphasis id="B907EAB1FFE58E6DFEA8F87B2A97538F" bold="true" box="[374,464,1924,1949]" pageId="22" pageNumber="323">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<collectingCountry id="F3647633FFE58E6DFE06F87A2AB0538F" box="[472,503,1925,1949]" name="American Samoa" pageId="22" pageNumber="323">As</collectingCountry>
|
||
noted in
|
||
<taxonomicName id="4C734D20FFE58E6DFDBCF87929A0538F" box="[610,743,1924,1949]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="323" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFE58E6DFDBCF87929A0538F" box="[610,743,1924,1949]" italics="true" pageId="22" pageNumber="323">D. australis</emphasis>
|
||
</taxonomicName>
|
||
, the variation in the contact of the supraloreal with the upper labials and interrupting the loreal scale row or not, is similarly recorded in
|
||
<taxonomicName id="4C734D20FFE58E6DFBDEF8552F3453D3" box="[1024,1139,1960,1985]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="323" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE58E6DFBDEF8552F3453D3" box="[1024,1139,1960,1985]" italics="true" pageId="22" pageNumber="323">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE58E6DFBA2F8572F9F53D3" box="[1148,1240,1960,1985]" pageId="22" pageNumber="323" rank="species">
|
||
<emphasis id="B907EAB1FFE58E6DFBA2F8572F9F53D3" bold="true" box="[1148,1240,1960,1985]" pageId="22" pageNumber="323">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
as the following conditions: the supraloreal contacts second upper labial on both sides (as in
|
||
<typeStatus id="54C88801FFE58E6DFC3DF8332F0053F7" box="[995,1095,1996,2021]" pageId="22" pageNumber="323" type="holotype">holotype</typeStatus>
|
||
) in WAM R129674 and WAM R154234; it contacts second and third upper labial on both sides in WAM R131902 and WAM R144238, and is separated from upper labials by either one large loreal and an elongate postnasal or two smaller loreals in WAM R156978. One specimen (WAM R154234) has five upper labials unilaterally, with the third under eye. All the
|
||
<typeStatus id="54C88801FFE48E6CFAB2FF1F2EDB54EA" box="[1388,1436,224,248]" pageId="23" pageNumber="324">type</typeStatus>
|
||
material has 18 midbody scales and 2‒4 loreals.
|
||
</paragraph>
|
||
<caption id="DF0C662BFFE48E6CFF49FEB32ADB5591" ID-Table-UUID="DF0C662BFFE48E6CFF49FEB32ADB5591" httpUri="http://table.plazi.org/id/DF0C662BFFE48E6CFF49FEB32ADB5591" pageId="23" pageNumber="324" targetBox="[159,1410,414,754]" targetIsTable="true" targetPageId="23">
|
||
<paragraph id="8BCC36A3FFE48E6CFF49FEB32ADB5591" blockId="23.[151,1436,331,387]" pageId="23" pageNumber="324">
|
||
<emphasis id="B907EAB1FFE48E6CFF49FEB32A4C5571" bold="true" box="[151,267,332,355]" pageId="23" pageNumber="324">TABLE 8.</emphasis>
|
||
Individual measurements (mm) and meristic counts for the type series of
|
||
<taxonomicName id="4C734D20FFE48E6CFBFBFEB32FD75571" box="[1061,1168,331,355]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE48E6CFBFBFEB32FD75571" box="[1061,1168,331,355]" italics="true" pageId="23" pageNumber="324">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE48E6CFB49FEB32FAA5571" box="[1175,1261,332,355]" pageId="23" pageNumber="324" rank="species">
|
||
<emphasis id="B907EAB1FFE48E6CFB49FEB32FAA5571" bold="true" box="[1175,1261,332,355]" pageId="23" pageNumber="324">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
The holotype is marked with an asterisk.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC36A3FFE48E6CFF41FE602E3D56E0" pageId="23" pageNumber="324">
|
||
<table id="F973C403FFE47184FF41FE612EC556E0" box="[159,1410,414,754]" gridcols="13" gridrows="8" pageId="23" pageNumber="324">
|
||
<tr id="354334E1FFE47184FF41FE612EC555A6" box="[159,1410,414,436]" gridrow="0" pageId="23" pageNumber="324">
|
||
<th id="76925D9DFFE47184FF41FE612BB055A6" box="[159,247,414,436]" gridcol="0" gridrow="0" pageId="23" pageNumber="324">WAM #</th>
|
||
<th id="76925D9DFFE47184FEFAFE612A0D55A6" box="[292,330,414,436]" gridcol="1" gridrow="0" pageId="23" pageNumber="324">Sex</th>
|
||
<th id="76925D9DFFE47184FEB2FE612ADB55A6" box="[364,412,414,436]" gridcol="2" gridrow="0" pageId="23" pageNumber="324">SVL</th>
|
||
<th id="76925D9DFFE47184FE1EFE612AA255A6" box="[448,485,414,436]" gridcol="3" gridrow="0" pageId="23" pageNumber="324">HD</th>
|
||
<th id="76925D9DFFE47184FDC8FE61297F55A6" box="[534,568,414,436]" gridcol="4" gridrow="0" pageId="23" pageNumber="324">HL</th>
|
||
<th id="76925D9DFFE47184FDB6FE6129D555A6" box="[616,658,414,436]" gridcol="5" gridrow="0" pageId="23" pageNumber="324">HW</th>
|
||
<th id="76925D9DFFE47184FD67FE6129AD55A6" box="[697,746,414,436]" gridcol="6" gridrow="0" pageId="23" pageNumber="324">HLL</th>
|
||
<th id="76925D9DFFE47184FCCDFE61287E55A6" box="[787,825,414,436]" gridcol="7" gridrow="0" pageId="23" pageNumber="324">ML</th>
|
||
<th id="76925D9DFFE47184FCB5FE6128CC55A6" box="[875,907,414,436]" gridcol="8" gridrow="0" pageId="23" pageNumber="324">RL</th>
|
||
<th id="76925D9DFFE47184FC65FE6128A455A6" box="[955,995,414,436]" gridcol="9" gridrow="0" pageId="23" pageNumber="324">RW</th>
|
||
<th id="76925D9DFFE47184FBC9FE612F7055A6" box="[1047,1079,414,436]" gridcol="10" gridrow="0" pageId="23" pageNumber="324">SL</th>
|
||
<th id="76925D9DFFE47184FBB8FE612FCA55A6" box="[1126,1165,414,436]" gridcol="11" gridrow="0" pageId="23" pageNumber="324">EW</th>
|
||
<th id="76925D9DFFE47184FB1FFE612EC555A6" box="[1217,1410,414,436]" gridcol="12" gridrow="0" pageId="23" pageNumber="324">HLS MSR VE</th>
|
||
</tr>
|
||
<tr id="354334E1FFE47184FF41FE332EC555F0" box="[159,1410,460,482]" gridrow="1" pageId="23" pageNumber="324">
|
||
<th id="76925D9DFFE47184FF41FE332BB055F0" box="[159,247,460,482]" gridcol="0" gridrow="1" pageId="23" pageNumber="324">129674</th>
|
||
<td id="76925D9DFFE47184FEFAFE332A0D55F0" box="[292,330,460,482]" gridcol="1" gridrow="1" pageId="23" pageNumber="324">♂</td>
|
||
<td id="76925D9DFFE47184FEB2FE332ADB55F0" box="[364,412,460,482]" gridcol="2" gridrow="1" pageId="23" pageNumber="324">61</td>
|
||
<td id="76925D9DFFE47184FE1EFE332AA255F0" box="[448,485,460,482]" gridcol="3" gridrow="1" pageId="23" pageNumber="324">3.2</td>
|
||
<td id="76925D9DFFE47184FDC8FE33297F55F0" box="[534,568,460,482]" gridcol="4" gridrow="1" pageId="23" pageNumber="324">6.6</td>
|
||
<td id="76925D9DFFE47184FDB6FE3329D555F0" box="[616,658,460,482]" gridcol="5" gridrow="1" pageId="23" pageNumber="324">4.3</td>
|
||
<td id="76925D9DFFE47184FD67FE3329AD55F0" box="[697,746,460,482]" gridcol="6" gridrow="1" pageId="23" pageNumber="324">3.1</td>
|
||
<td id="76925D9DFFE47184FCCDFE33287E55F0" box="[787,825,460,482]" gridcol="7" gridrow="1" pageId="23" pageNumber="324">5.2</td>
|
||
<td id="76925D9DFFE47184FCB5FE3328CC55F0" box="[875,907,460,482]" gridcol="8" gridrow="1" pageId="23" pageNumber="324">0.9</td>
|
||
<td id="76925D9DFFE47184FC65FE3328A455F0" box="[955,995,460,482]" gridcol="9" gridrow="1" pageId="23" pageNumber="324">1.5</td>
|
||
<td id="76925D9DFFE47184FBC9FE332F7055F0" box="[1047,1079,460,482]" gridcol="10" gridrow="1" pageId="23" pageNumber="324">2.8</td>
|
||
<td id="76925D9DFFE47184FBB8FE332FCA55F0" box="[1126,1165,460,482]" gridcol="11" gridrow="1" pageId="23" pageNumber="324">1.2</td>
|
||
<td id="76925D9DFFE47184FB1FFE332EC555F0" box="[1217,1410,460,482]" gridcol="12" gridrow="1" pageId="23" pageNumber="324">10 18 76</td>
|
||
</tr>
|
||
<tr id="354334E1FFE47184FF41FE062EC5561D" box="[159,1410,505,527]" gridrow="2" pageId="23" pageNumber="324">
|
||
<th id="76925D9DFFE47184FF41FE062BB0561D" box="[159,247,505,527]" gridcol="0" gridrow="2" pageId="23" pageNumber="324">131902</th>
|
||
<td id="76925D9DFFE47184FEFAFE062A0D561D" box="[292,330,505,527]" gridcol="1" gridrow="2" pageId="23" pageNumber="324">♀</td>
|
||
<td id="76925D9DFFE47184FEB2FE062ADB561D" box="[364,412,505,527]" gridcol="2" gridrow="2" pageId="23" pageNumber="324">67</td>
|
||
<td id="76925D9DFFE47184FE1EFE062AA2561D" box="[448,485,505,527]" gridcol="3" gridrow="2" pageId="23" pageNumber="324">2.6</td>
|
||
<td id="76925D9DFFE47184FDC8FE06297F561D" box="[534,568,505,527]" gridcol="4" gridrow="2" pageId="23" pageNumber="324">6.6</td>
|
||
<td id="76925D9DFFE47184FDB6FE0629D5561D" box="[616,658,505,527]" gridcol="5" gridrow="2" pageId="23" pageNumber="324">3.7</td>
|
||
<td id="76925D9DFFE47184FD67FE0629AD561D" box="[697,746,505,527]" gridcol="6" gridrow="2" pageId="23" pageNumber="324">2.2</td>
|
||
<td id="76925D9DFFE47184FCCDFE06287E561D" box="[787,825,505,527]" gridcol="7" gridrow="2" pageId="23" pageNumber="324">5.4</td>
|
||
<td id="76925D9DFFE47184FCB5FE0628CC561D" box="[875,907,505,527]" gridcol="8" gridrow="2" pageId="23" pageNumber="324">0.7</td>
|
||
<td id="76925D9DFFE47184FC65FE0628A4561D" box="[955,995,505,527]" gridcol="9" gridrow="2" pageId="23" pageNumber="324">1.3</td>
|
||
<td id="76925D9DFFE47184FBC9FE062F70561D" box="[1047,1079,505,527]" gridcol="10" gridrow="2" pageId="23" pageNumber="324">2.3</td>
|
||
<td id="76925D9DFFE47184FBB8FE062FCA561D" box="[1126,1165,505,527]" gridcol="11" gridrow="2" pageId="23" pageNumber="324">1.1</td>
|
||
<td id="76925D9DFFE47184FB1FFE062EC5561D" box="[1217,1410,505,527]" gridcol="12" gridrow="2" pageId="23" pageNumber="324">10 18 84</td>
|
||
</tr>
|
||
<tr id="354334E1FFE47184FF41FDD92EC5562E" box="[159,1410,550,572]" gridrow="3" pageId="23" pageNumber="324">
|
||
<th id="76925D9DFFE47184FF41FDD92BB0562E" box="[159,247,550,572]" gridcol="0" gridrow="3" pageId="23" pageNumber="324">132154</th>
|
||
<td id="76925D9DFFE47184FEFAFDD92A0D562E" box="[292,330,550,572]" gridcol="1" gridrow="3" pageId="23" pageNumber="324">♀</td>
|
||
<td id="76925D9DFFE47184FEB2FDD92ADB562E" box="[364,412,550,572]" gridcol="2" gridrow="3" pageId="23" pageNumber="324">78</td>
|
||
<td id="76925D9DFFE47184FE1EFDD92AA2562E" box="[448,485,550,572]" gridcol="3" gridrow="3" pageId="23" pageNumber="324">3.4</td>
|
||
<td id="76925D9DFFE47184FDC8FDD9297F562E" box="[534,568,550,572]" gridcol="4" gridrow="3" pageId="23" pageNumber="324">7.6</td>
|
||
<td id="76925D9DFFE47184FDB6FDD929D5562E" box="[616,658,550,572]" gridcol="5" gridrow="3" pageId="23" pageNumber="324">5</td>
|
||
<td id="76925D9DFFE47184FD67FDD929AD562E" box="[697,746,550,572]" gridcol="6" gridrow="3" pageId="23" pageNumber="324">2.1</td>
|
||
<td id="76925D9DFFE47184FCCDFDD9287E562E" box="[787,825,550,572]" gridcol="7" gridrow="3" pageId="23" pageNumber="324">5.3</td>
|
||
<td id="76925D9DFFE47184FCB5FDD928CC562E" box="[875,907,550,572]" gridcol="8" gridrow="3" pageId="23" pageNumber="324">1</td>
|
||
<td id="76925D9DFFE47184FC65FDD928A4562E" box="[955,995,550,572]" gridcol="9" gridrow="3" pageId="23" pageNumber="324">1.4</td>
|
||
<td id="76925D9DFFE47184FBC9FDD92F70562E" box="[1047,1079,550,572]" gridcol="10" gridrow="3" pageId="23" pageNumber="324">3.2</td>
|
||
<td id="76925D9DFFE47184FBB8FDD92FCA562E" box="[1126,1165,550,572]" gridcol="11" gridrow="3" pageId="23" pageNumber="324">1.3</td>
|
||
<td id="76925D9DFFE47184FB1FFDD92EC5562E" box="[1217,1410,550,572]" gridcol="12" gridrow="3" pageId="23" pageNumber="324">10 18 85</td>
|
||
</tr>
|
||
<tr id="354334E1FFE47184FF41FDAB2EC55678" box="[159,1410,596,618]" gridrow="4" pageId="23" pageNumber="324">
|
||
<th id="76925D9DFFE47184FF41FDAB2BB05678" box="[159,247,596,618]" gridcol="0" gridrow="4" pageId="23" pageNumber="324">144237*</th>
|
||
<td id="76925D9DFFE47184FEFAFDAB2A0D5678" box="[292,330,596,618]" gridcol="1" gridrow="4" pageId="23" pageNumber="324">♂</td>
|
||
<td id="76925D9DFFE47184FEB2FDAB2ADB5678" box="[364,412,596,618]" gridcol="2" gridrow="4" pageId="23" pageNumber="324">57</td>
|
||
<td id="76925D9DFFE47184FE1EFDAB2AA25678" box="[448,485,596,618]" gridcol="3" gridrow="4" pageId="23" pageNumber="324">3.4</td>
|
||
<td id="76925D9DFFE47184FDC8FDAB297F5678" box="[534,568,596,618]" gridcol="4" gridrow="4" pageId="23" pageNumber="324">6.9</td>
|
||
<td id="76925D9DFFE47184FDB6FDAB29D55678" box="[616,658,596,618]" gridcol="5" gridrow="4" pageId="23" pageNumber="324">4.5</td>
|
||
<td id="76925D9DFFE47184FD67FDAB29AD5678" box="[697,746,596,618]" gridcol="6" gridrow="4" pageId="23" pageNumber="324">3.3</td>
|
||
<td id="76925D9DFFE47184FCCDFDAB287E5678" box="[787,825,596,618]" gridcol="7" gridrow="4" pageId="23" pageNumber="324">4.9</td>
|
||
<td id="76925D9DFFE47184FCB5FDAB28CC5678" box="[875,907,596,618]" gridcol="8" gridrow="4" pageId="23" pageNumber="324">0.9</td>
|
||
<td id="76925D9DFFE47184FC65FDAB28A45678" box="[955,995,596,618]" gridcol="9" gridrow="4" pageId="23" pageNumber="324">1.3</td>
|
||
<td id="76925D9DFFE47184FBC9FDAB2F705678" box="[1047,1079,596,618]" gridcol="10" gridrow="4" pageId="23" pageNumber="324">2.6</td>
|
||
<td id="76925D9DFFE47184FBB8FDAB2FCA5678" box="[1126,1165,596,618]" gridcol="11" gridrow="4" pageId="23" pageNumber="324">1.1</td>
|
||
<td id="76925D9DFFE47184FB1FFDAB2EC55678" box="[1217,1410,596,618]" gridcol="12" gridrow="4" pageId="23" pageNumber="324">10 18 75</td>
|
||
</tr>
|
||
<tr id="354334E1FFE47184FF41FD7E2EC55685" box="[159,1410,641,663]" gridrow="5" pageId="23" pageNumber="324">
|
||
<th id="76925D9DFFE47184FF41FD7E2BB05685" box="[159,247,641,663]" gridcol="0" gridrow="5" pageId="23" pageNumber="324">144238</th>
|
||
<td id="76925D9DFFE47184FEFAFD7E2A0D5685" box="[292,330,641,663]" gridcol="1" gridrow="5" pageId="23" pageNumber="324">♂</td>
|
||
<td id="76925D9DFFE47184FEB2FD7E2ADB5685" box="[364,412,641,663]" gridcol="2" gridrow="5" pageId="23" pageNumber="324">66</td>
|
||
<td id="76925D9DFFE47184FE1EFD7E2AA25685" box="[448,485,641,663]" gridcol="3" gridrow="5" pageId="23" pageNumber="324">3.2</td>
|
||
<td id="76925D9DFFE47184FDC8FD7E297F5685" box="[534,568,641,663]" gridcol="4" gridrow="5" pageId="23" pageNumber="324">7.5</td>
|
||
<td id="76925D9DFFE47184FDB6FD7E29D55685" box="[616,658,641,663]" gridcol="5" gridrow="5" pageId="23" pageNumber="324">4.7</td>
|
||
<td id="76925D9DFFE47184FD67FD7E29AD5685" box="[697,746,641,663]" gridcol="6" gridrow="5" pageId="23" pageNumber="324">3.6</td>
|
||
<td id="76925D9DFFE47184FCCDFD7E287E5685" box="[787,825,641,663]" gridcol="7" gridrow="5" pageId="23" pageNumber="324">5.8</td>
|
||
<td id="76925D9DFFE47184FCB5FD7E28CC5685" box="[875,907,641,663]" gridcol="8" gridrow="5" pageId="23" pageNumber="324">1</td>
|
||
<td id="76925D9DFFE47184FC65FD7E28A45685" box="[955,995,641,663]" gridcol="9" gridrow="5" pageId="23" pageNumber="324">1.7</td>
|
||
<td id="76925D9DFFE47184FBC9FD7E2F705685" box="[1047,1079,641,663]" gridcol="10" gridrow="5" pageId="23" pageNumber="324">2.9</td>
|
||
<td id="76925D9DFFE47184FBB8FD7E2FCA5685" box="[1126,1165,641,663]" gridcol="11" gridrow="5" pageId="23" pageNumber="324">1.2</td>
|
||
<td id="76925D9DFFE47184FB1FFD7E2EC55685" box="[1217,1410,641,663]" gridcol="12" gridrow="5" pageId="23" pageNumber="324">10 18 78</td>
|
||
</tr>
|
||
<tr id="354334E1FFE47184FF41FD512EC556D6" box="[159,1410,686,708]" gridrow="6" pageId="23" pageNumber="324">
|
||
<th id="76925D9DFFE47184FF41FD512BB056D6" box="[159,247,686,708]" gridcol="0" gridrow="6" pageId="23" pageNumber="324">154234</th>
|
||
<td id="76925D9DFFE47184FEFAFD512A0D56D6" box="[292,330,686,708]" gridcol="1" gridrow="6" pageId="23" pageNumber="324">♂</td>
|
||
<td id="76925D9DFFE47184FEB2FD512ADB56D6" box="[364,412,686,708]" gridcol="2" gridrow="6" pageId="23" pageNumber="324">49</td>
|
||
<td id="76925D9DFFE47184FE1EFD512AA256D6" box="[448,485,686,708]" gridcol="3" gridrow="6" pageId="23" pageNumber="324">2.5</td>
|
||
<td id="76925D9DFFE47184FDC8FD51297F56D6" box="[534,568,686,708]" gridcol="4" gridrow="6" pageId="23" pageNumber="324">5.9</td>
|
||
<td id="76925D9DFFE47184FDB6FD5129D556D6" box="[616,658,686,708]" gridcol="5" gridrow="6" pageId="23" pageNumber="324">3.7</td>
|
||
<td id="76925D9DFFE47184FD67FD5129AD56D6" box="[697,746,686,708]" gridcol="6" gridrow="6" pageId="23" pageNumber="324">2.4</td>
|
||
<td id="76925D9DFFE47184FCCDFD51287E56D6" box="[787,825,686,708]" gridcol="7" gridrow="6" pageId="23" pageNumber="324">4.4</td>
|
||
<td id="76925D9DFFE47184FCB5FD5128CC56D6" box="[875,907,686,708]" gridcol="8" gridrow="6" pageId="23" pageNumber="324">0.8</td>
|
||
<td id="76925D9DFFE47184FC65FD5128A456D6" box="[955,995,686,708]" gridcol="9" gridrow="6" pageId="23" pageNumber="324">1.3</td>
|
||
<td id="76925D9DFFE47184FBC9FD512F7056D6" box="[1047,1079,686,708]" gridcol="10" gridrow="6" pageId="23" pageNumber="324">2.6</td>
|
||
<td id="76925D9DFFE47184FBB8FD512FCA56D6" box="[1126,1165,686,708]" gridcol="11" gridrow="6" pageId="23" pageNumber="324">1.2</td>
|
||
<td id="76925D9DFFE47184FB1FFD512EC556D6" box="[1217,1410,686,708]" gridcol="12" gridrow="6" pageId="23" pageNumber="324">10 18 77</td>
|
||
</tr>
|
||
<tr id="354334E1FFE47184FF41FD232EC556E0" box="[159,1410,732,754]" gridrow="7" pageId="23" pageNumber="324">
|
||
<th id="76925D9DFFE47184FF41FD232BB056E0" box="[159,247,732,754]" gridcol="0" gridrow="7" pageId="23" pageNumber="324">156978</th>
|
||
<td id="76925D9DFFE47184FEFAFD232A0D56E0" box="[292,330,732,754]" gridcol="1" gridrow="7" pageId="23" pageNumber="324">♂</td>
|
||
<td id="76925D9DFFE47184FEB2FD232ADB56E0" box="[364,412,732,754]" gridcol="2" gridrow="7" pageId="23" pageNumber="324">66</td>
|
||
<td id="76925D9DFFE47184FE1EFD232AA256E0" box="[448,485,732,754]" gridcol="3" gridrow="7" pageId="23" pageNumber="324">3.2</td>
|
||
<td id="76925D9DFFE47184FDC8FD23297F56E0" box="[534,568,732,754]" gridcol="4" gridrow="7" pageId="23" pageNumber="324">7.6</td>
|
||
<td id="76925D9DFFE47184FDB6FD2329D556E0" box="[616,658,732,754]" gridcol="5" gridrow="7" pageId="23" pageNumber="324">4.4</td>
|
||
<td id="76925D9DFFE47184FD67FD2329AD56E0" box="[697,746,732,754]" gridcol="6" gridrow="7" pageId="23" pageNumber="324">3.6</td>
|
||
<td id="76925D9DFFE47184FCCDFD23287E56E0" box="[787,825,732,754]" gridcol="7" gridrow="7" pageId="23" pageNumber="324">5</td>
|
||
<td id="76925D9DFFE47184FCB5FD2328CC56E0" box="[875,907,732,754]" gridcol="8" gridrow="7" pageId="23" pageNumber="324">1</td>
|
||
<td id="76925D9DFFE47184FC65FD2328A456E0" box="[955,995,732,754]" gridcol="9" gridrow="7" pageId="23" pageNumber="324">1.3</td>
|
||
<td id="76925D9DFFE47184FBC9FD232F7056E0" box="[1047,1079,732,754]" gridcol="10" gridrow="7" pageId="23" pageNumber="324">3.1</td>
|
||
<td id="76925D9DFFE47184FBB8FD232FCA56E0" box="[1126,1165,732,754]" gridcol="11" gridrow="7" pageId="23" pageNumber="324">1.1</td>
|
||
<td id="76925D9DFFE47184FB1FFD232EC556E0" box="[1217,1410,732,754]" gridcol="12" gridrow="7" pageId="23" pageNumber="324">11 18 78</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE48E6CFF49FB352BB1510F" pageId="23" pageNumber="324">
|
||
<table id="F973C403FFE47184FF49FB352EDB510F" box="[151,1436,1226,1309]" gridcols="4" gridrows="3" pageId="23" pageNumber="324">
|
||
<tr id="354334E1FFE47184FF49FB352EDB50F2" box="[151,1436,1226,1248]" gridrow="0" pageId="23" pageNumber="324">
|
||
<th id="76925D9DFFE47184FF49FB352AA450F2" box="[151,483,1226,1248]" gridcol="0" gridrow="0" pageId="23" pageNumber="324">
|
||
<emphasis id="B907EAB1FFE48E6CFF49FB352A5950F2" bold="true" box="[151,286,1226,1248]" pageId="23" pageNumber="324">FIGURE 11.</emphasis>
|
||
Head patterning of
|
||
</th>
|
||
<th id="76925D9DFFE47184FE34FB35299650F2" box="[490,721,1226,1248]" gridcol="1" gridrow="0" pageId="23" pageNumber="324">
|
||
(A)
|
||
<taxonomicName id="4C734D20FFE48E6CFDCAFB34293D50F2" box="[532,634,1226,1248]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE48E6CFDCAFB34293D50F2" box="[532,634,1226,1248]" italics="true" pageId="23" pageNumber="324">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE48E6CFD5EFB35299650F2" box="[640,721,1226,1248]" pageId="23" pageNumber="324" rank="species">
|
||
<emphasis id="B907EAB1FFE48E6CFD5EFB35299650F2" bold="true" box="[640,721,1226,1248]" pageId="23" pageNumber="324">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</th>
|
||
<th id="76925D9DFFE47184FD06FB35280450F2" box="[728,835,1226,1248]" gridcol="2" gridrow="0" pageId="23" pageNumber="324">adult from</th>
|
||
<th id="76925D9DFFE47184FC94FB352EDB50F2" box="[842,1436,1226,1248]" gridcol="3" gridrow="0" pageId="23" pageNumber="324">
|
||
Kundip, Western Australia and (B)
|
||
<taxonomicName id="4C734D20FFE48E6CFB6DFB342E6C50F2" box="[1203,1323,1226,1248]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFE48E6CFB6DFB342E6C50F2" box="[1203,1323,1226,1248]" italics="true" pageId="23" pageNumber="324">D. australis</emphasis>
|
||
</taxonomicName>
|
||
adult from
|
||
</th>
|
||
</tr>
|
||
<tr id="354334E1FFE47184FF49FB162EDB50ED" box="[151,1436,1257,1279]" gridrow="1" pageId="23" pageNumber="324">
|
||
<th id="76925D9DFFE47184FF49FB162AA450ED" box="[151,483,1257,1279]" gridcol="0" gridrow="1" pageId="23" pageNumber="324">Cowell, Eyre Peninsula, South</th>
|
||
<td id="76925D9DFFE47184FE34FB16299650ED" box="[490,721,1257,1279]" gridcol="1" gridrow="1" pageId="23" pageNumber="324">Australia, and arrows</td>
|
||
<td id="76925D9DFFE47184FD06FB16280450ED" box="[728,835,1257,1279]" gridcol="2" gridrow="1" pageId="23" pageNumber="324">indicating</td>
|
||
<td id="76925D9DFFE47184FC94FB162EDB50ED" box="[842,1436,1257,1279]" gridcol="3" gridrow="1" pageId="23" pageNumber="324">pigmented or clear suture on lower labials (images‒B.</td>
|
||
</tr>
|
||
<tr id="354334E1FFE47184FF49FAF72EDB510F" box="[151,1436,1288,1309]" gridrow="2" pageId="23" pageNumber="324" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76925D9DFFE47184FF49FAF72AA4510F" box="[151,483,1288,1309]" gridcol="0" gridrow="2" pageId="23" pageNumber="324">Maryan).</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE48E6CFF19FAB52A7651E1" blockId="23.[151,1436,1354,2027]" pageId="23" pageNumber="324">
|
||
<emphasis id="B907EAB1FFE48E6CFF19FAB529CF5171" bold="true" box="[199,648,1354,1379]" pageId="23" pageNumber="324">Variation in colouration and pattern.</emphasis>
|
||
Most
|
||
<emphasis id="B907EAB1FFE48E6CFD0DFAB529995171" bold="true" box="[723,734,1354,1379]" pageId="23" pageNumber="324">s</emphasis>
|
||
pecimens are similarly coloured to the
|
||
<typeStatus id="54C88801FFE48E6CFB43FAB52FB85171" box="[1181,1279,1354,1379]" pageId="23" pageNumber="324" type="holotype">holotype</typeStatus>
|
||
in life and in preservative. Occasional preserved individuals (e.g. WAM R131902) have more pronounced dark bars on the lower labials and lateral scales of forebody, while others (e.g. WAM R154234) are a very uniform dark grey on head, body and tail. In life, hatchlings of
|
||
<taxonomicName id="4C734D20FFE48E6CFDF2FA4829DA51DD" box="[556,669,1462,1487]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE48E6CFDF2FA4829DA51DD" box="[556,669,1462,1487]" italics="true" pageId="23" pageNumber="324">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE48E6CFD7BFA49284651DD" box="[677,769,1462,1487]" pageId="23" pageNumber="324" rank="species">
|
||
<emphasis id="B907EAB1FFE48E6CFD7BFA49284651DD" bold="true" box="[677,769,1462,1487]" pageId="23" pageNumber="324">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have a similar colour and head pattern to adults (B. Bush, pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3696528FFE48E6CFF19FA0129BB5229" pageId="23" pageNumber="324" type="etymology">
|
||
<paragraph id="8BCC36A3FFE48E6CFF19FA0129BB5229" blockId="23.[151,1436,1354,2027]" pageId="23" pageNumber="324">
|
||
<emphasis id="B907EAB1FFE48E6CFF19FA012A095205" bold="true" box="[199,334,1534,1559]" pageId="23" pageNumber="324">Etymology.</emphasis>
|
||
The specific name
|
||
<taxonomicName id="4C734D20FFE48E6CFDF4FA01293F5205" box="[554,632,1534,1559]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="hebesa">
|
||
<emphasis id="B907EAB1FFE48E6CFDF4FA01293F5205" box="[554,632,1534,1559]" italics="true" pageId="23" pageNumber="324">hebesa</emphasis>
|
||
</taxonomicName>
|
||
is derived from the Latin adjective
|
||
<emphasis id="B907EAB1FFE48E6CFBD4FA012F0D5205" box="[1034,1098,1534,1559]" italics="true" pageId="23" pageNumber="324">hebes</emphasis>
|
||
, meaning dull, alluding to the matt body texture, without much shine, of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3696528FFE48E63FF19F9B929A455C2" lastPageId="24" lastPageNumber="325" pageId="23" pageNumber="324" type="distribution">
|
||
<paragraph id="8BCC36A3FFE48E6CFF19F9B92A9E52FD" blockId="23.[151,1436,1354,2027]" pageId="23" pageNumber="324">
|
||
<emphasis id="B907EAB1FFE48E6CFF19F9B92954524D" bold="true" box="[199,531,1606,1631]" pageId="23" pageNumber="324">Distribution and sympatry.</emphasis>
|
||
<taxonomicName id="4C734D20FFE48E6CFDC0F9B92986524D" box="[542,705,1606,1631]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE48E6CFDC0F9B92986524D" box="[542,705,1606,1631]" italics="true" pageId="23" pageNumber="324">Delma hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE48E6CFD12F9B9286E524D" box="[716,809,1606,1631]" pageId="23" pageNumber="324" rank="species">
|
||
<emphasis id="B907EAB1FFE48E6CFD12F9B9286E524D" bold="true" box="[716,809,1606,1631]" pageId="23" pageNumber="324">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is widespread on the Esperance Plains bioregion and patchily distributed on the Warren and southern Jarrah Forest bioregions (
|
||
<bibRefCitation id="EFE24B52FFE48E6CFBCDF9952E375291" author="Thackway" box="[1043,1392,1642,1667]" pageId="23" pageNumber="324" refString="Thackway, R. & Cresswell, I. D. (1995) An Interim Biogeographic Regionalisation for Australia: A Framework for Setting Priorities in the National Reserves System Cooperative Program Version 4.0. Australian Nature Conservation Agency, Reserves Systems Unit, Canberra." type="book" year="1995">Thackway & Cresswell 1995</bibRefCitation>
|
||
) in southwestern Western
|
||
<collectingCountry id="F3647633FFE48E6CFE4AF9712ABC52B5" box="[404,507,1678,1703]" name="Australia" pageId="23" pageNumber="324">Australia</collectingCountry>
|
||
(
|
||
<figureCitation id="13482A26FFE48E6CFDD5F971290952B5" box="[523,590,1678,1703]" captionStart="FIGURE 8" captionStartId="19.[151,250,1403,1425]" captionTargetBox="[153,1433,484,1378]" captionTargetId="figure@19.[151,1437,481,1382]" captionTargetPageId="19" captionText="FIGURE 8. Map of Australia showing distribution of the D. australis species-group and specimens examined in the DNA and allozyme analyses. Specimens examined in the morphological analysis are indicated by a symbol with a central dot." httpUri="https://zenodo.org/record/244615/files/figure.png" pageId="23" pageNumber="324">Fig. 8</figureCitation>
|
||
). Records extend east to the vicinity of Thomas River and Cape Arid, west to Canal Rocks and near Busselton, and inland to Stirling Range National Park, Ongerup, Ravensthorpe Range, Scaddan and Mount Burdett.
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE48E6CFF19F9052FD853D5" blockId="23.[151,1436,1354,2027]" pageId="23" pageNumber="324">
|
||
The geographic distributions of
|
||
<taxonomicName id="4C734D20FFE48E6CFDEEF90429E75301" box="[560,672,1786,1811]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE48E6CFDEEF90429E75301" box="[560,672,1786,1811]" italics="true" pageId="23" pageNumber="324">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE48E6CFD76F90528455301" box="[680,770,1786,1811]" pageId="23" pageNumber="324" rank="species">
|
||
<emphasis id="B907EAB1FFE48E6CFD76F90528455301" bold="true" box="[680,770,1786,1811]" pageId="23" pageNumber="324">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C734D20FFE48E6CFCE7F90428F95301" box="[825,958,1786,1811]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFE48E6CFCE7F90428F95301" box="[825,958,1786,1811]" italics="true" pageId="23" pageNumber="324">D. australis</emphasis>
|
||
</taxonomicName>
|
||
appear to be parapatric (
|
||
<figureCitation id="13482A26FFE48E6CFB08F9052E5D5301" box="[1238,1306,1786,1811]" captionStart="FIGURE 8" captionStartId="19.[151,250,1403,1425]" captionTargetBox="[153,1433,484,1378]" captionTargetId="figure@19.[151,1437,481,1382]" captionTargetPageId="19" captionText="FIGURE 8. Map of Australia showing distribution of the D. australis species-group and specimens examined in the DNA and allozyme analyses. Specimens examined in the morphological analysis are indicated by a symbol with a central dot." httpUri="https://zenodo.org/record/244615/files/figure.png" pageId="23" pageNumber="324">Fig. 8</figureCitation>
|
||
). Currently the two species are known to occur within
|
||
<quantity id="4C8B9B46FFE48E6CFD5DF8E0298B5325" box="[643,716,1822,1847]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="324" unit="km" value="80.0">80 km</quantity>
|
||
of each other in the east: WAM R91740 of
|
||
<taxonomicName id="4C734D20FFE48E6CFB18F8E02E7F5325" box="[1222,1336,1822,1847]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE48E6CFB18F8E02E7F5325" box="[1222,1336,1822,1847]" italics="true" pageId="23" pageNumber="324">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE48E6CFA9FF8E12EDB5325" box="[1345,1436,1822,1847]" pageId="23" pageNumber="324" rank="species">
|
||
<emphasis id="B907EAB1FFE48E6CFA9FF8E12EDB5325" bold="true" box="[1345,1436,1822,1847]" pageId="23" pageNumber="324">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from Cape Arid
|
||
<emphasis id="B907EAB1FFE48E6CFE90F8BC2AD25348" box="[334,405,1859,1882]" italics="true" pageId="23" pageNumber="324">versus</emphasis>
|
||
WAM R36229 of
|
||
<taxonomicName id="4C734D20FFE48E6CFDBBF8BC29AE5349" box="[613,745,1858,1883]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFE48E6CFDBBF8BC29AE5349" box="[613,745,1858,1883]" italics="true" pageId="23" pageNumber="324">D. australis</emphasis>
|
||
</taxonomicName>
|
||
from Pine Hill and within
|
||
<quantity id="4C8B9B46FFE48E6CFBC8F8BC2F2D5349" box="[1046,1130,1858,1883]" metricMagnitude="5" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="324" unit="km" value="130.0">130 km</quantity>
|
||
in the west: WAM R42637 of
|
||
<taxonomicName id="4C734D20FFE48E6CFF67F8982A6B536D" box="[185,300,1894,1919]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE48E6CFF67F8982A6B536D" box="[185,300,1894,1919]" italics="true" pageId="23" pageNumber="324">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE48E6CFEE8F8992AD3536D" box="[310,404,1894,1919]" pageId="23" pageNumber="324" rank="species">
|
||
<emphasis id="B907EAB1FFE48E6CFEE8F8992AD3536D" bold="true" box="[310,404,1894,1919]" pageId="23" pageNumber="324">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from Ongerup
|
||
<emphasis id="B907EAB1FFE48E6CFD95F89829D5536C" box="[587,658,1895,1918]" italics="true" pageId="23" pageNumber="324">versus</emphasis>
|
||
WAM R12604 of
|
||
<taxonomicName id="4C734D20FFE48E6CFCAFF89828BF536D" box="[881,1016,1894,1919]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFE48E6CFCAFF89828BF536D" box="[881,1016,1894,1919]" italics="true" pageId="23" pageNumber="324">D. australis</emphasis>
|
||
</taxonomicName>
|
||
from Wagin. Specimens from these parapatric localities do not show any indication of morphological intermediacy. The only recorded instance of sympatry involving
|
||
<taxonomicName id="4C734D20FFE48E6CFEA6F8502AAE53D5" box="[376,489,1966,1991]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE48E6CFEA6F8502AAE53D5" box="[376,489,1966,1991]" italics="true" pageId="23" pageNumber="324">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE48E6CFE31F851290E53D5" box="[495,585,1966,1991]" pageId="23" pageNumber="324" rank="species">
|
||
<emphasis id="B907EAB1FFE48E6CFE31F851290E53D5" bold="true" box="[495,585,1966,1991]" pageId="23" pageNumber="324">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is with
|
||
<taxonomicName id="4C734D20FFE48E6CFD7CF850284853D5" box="[674,783,1967,1991]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="fraseri">
|
||
<emphasis id="B907EAB1FFE48E6CFD7CF850284853D5" box="[674,783,1967,1991]" italics="true" pageId="23" pageNumber="324">D. fraseri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFE24B52FFE48E6CFCC0F85128DD53D5" author="Bush" box="[798,922,1966,1991]" pageId="23" pageNumber="324" refString="Bush, B. (1981) Reptiles of the Kalgoorlie-Esperance Region. B. Bush, Perth, 46 pp." type="book" year="1981">Bush 1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE24B52FFE48E6CFC78F85028A753D5" author="Bush" box="[934,992,1967,1991]" pageId="23" pageNumber="324" refString="Bush, B. (1984) Seasonal aggregation behaviour in a mixed population of legless lizards, Delma australis and D. fraseri. Herpetofauna, 16, 1 - 6." type="journal article" year="1984">1984</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="4C734D20FFE48E6CFBD4F8502FC853D5" box="[1034,1167,1966,1991]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFE48E6CFBD4F8502FC853D5" box="[1034,1167,1966,1991]" italics="true" pageId="23" pageNumber="324">D. australis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFE48E63FF11F82D29A455C2" blockId="23.[151,1436,1354,2027]" lastBlockId="24.[151,1436,151,464]" lastPageId="24" lastPageNumber="325" pageId="23" pageNumber="324">
|
||
<emphasis id="B907EAB1FFE48E6CFF11F82D2A7553F9" bold="true" box="[207,306,2002,2027]" pageId="23" pageNumber="324">Habitat.</emphasis>
|
||
<taxonomicName id="4C734D20FFE48E6CFEE5F82D2A9B53F9" box="[315,476,2002,2027]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFE48E6CFEE5F82D2A9B53F9" box="[315,476,2002,2027]" italics="true" pageId="23" pageNumber="324">Delma hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFE48E6CFE3AF82D297853F9" box="[484,575,2002,2027]" pageId="23" pageNumber="324" rank="species">
|
||
<emphasis id="B907EAB1FFE48E6CFE3AF82D297853F9" bold="true" box="[484,575,2002,2027]" pageId="23" pageNumber="324">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
occupies the proteaceous scrub and mallee heath on south coast sandplains (
|
||
<bibRefCitation id="EFE24B52FFEB8E63FF40FF682A6354A2" author="Beard" box="[158,292,151,176]" pageId="24" pageNumber="325" refString="Beard, J. S. (1990) Plant Life of Western Australia. Kangaroo Press, Sydney, 317 pp." type="book" year="1990">Beard 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE24B52FFEB8E63FEF1FF672ABD54A2" author="Comer" box="[303,506,151,176]" pageId="24" pageNumber="325" refString="Comer, S., Gilfillan, S., Barrett, S., Grant, M., Tiedemann, K. & Anderson, L. (2001) Esperance 2 (ESP 2 - Recherche subregion). A Biodiversity Audit of Western Australia's 53 Biogeographical Subregions in 2002. Department of Conservation and Land Management, Perth. [unkown pagination]" type="journal article" year="2001">
|
||
Comer
|
||
<emphasis id="B907EAB1FFEB8E63FE5CFF662AFD54A2" box="[386,442,151,176]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
;
|
||
<figureCitation id="13482A26FFEB8E63FDDBFF68291054A2" box="[517,599,151,176]" captionStart="FIGURE 12" captionStartId="24.[151,250,1395,1417]" captionTargetBox="[151,1436,568,1373]" captionTargetId="figure@24.[151,1436,517,1373]" captionTargetPageId="24" captionText="FIGURE 12. The habitat of D. hebesa sp. nov., dense scrub heath on sandplain at Stirling Range National Park, Western Australia (image ‒ B. Maryan)." httpUri="https://zenodo.org/record/244618/files/figure.png" pageId="24" pageNumber="325">Fig. 12</figureCitation>
|
||
). This habitat preference is exemplified of 65 records of
|
||
<taxonomicName id="4C734D20FFEB8E63FB13FF662E7A54A2" box="[1229,1341,151,176]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFEB8E63FB13FF662E7A54A2" box="[1229,1341,151,176]" italics="true" pageId="24" pageNumber="325">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFEB8E63FA9DFF682EDB54A2" box="[1347,1436,151,176]" pageId="24" pageNumber="325" rank="species">
|
||
<emphasis id="B907EAB1FFEB8E63FA9DFF682EDB54A2" bold="true" box="[1347,1436,151,176]" pageId="24" pageNumber="325">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from 18 sites, in which 79% were recorded from mallee heath (
|
||
<bibRefCitation id="EFE24B52FFEB8E63FC80FF432F7D54C6" author="Sanders" box="[862,1082,188,213]" pageId="24" pageNumber="325" refString="Sanders, A., Chapman, A., Teale, R. J. & Harold, G. (2012) Vertebrate fauna of the Fitzgerald Biosphere Reserve, Western Australia. The Western Australian Naturalist, 28, 141 - 253." type="journal article" year="2012">
|
||
Sanders
|
||
<emphasis id="B907EAB1FFEB8E63FC1EFF4228BF54C7" box="[960,1016,188,213]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="4C734D20FFEB8E63FBBAFF422FAE54C7" box="[1124,1257,188,213]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFEB8E63FBBAFF422FAE54C7" box="[1124,1257,188,213]" italics="true" pageId="24" pageNumber="325">D. australis</emphasis>
|
||
</taxonomicName>
|
||
). These diverse vegetation communities provide ample cover for
|
||
<taxonomicName id="4C734D20FFEB8E63FD11FF1E280554EA" box="[719,834,223,248]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFEB8E63FD11FF1E280554EA" box="[719,834,223,248]" italics="true" pageId="24" pageNumber="325">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFEB8E63FC92FF2028EF54EA" box="[844,936,223,248]" pageId="24" pageNumber="325" rank="species">
|
||
<emphasis id="B907EAB1FFEB8E63FC92FF2028EF54EA" bold="true" box="[844,936,223,248]" pageId="24" pageNumber="325">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, where most specimens, including the
|
||
<typeStatus id="54C88801FFEB8E63FAB2FF1F2EDB54EA" box="[1388,1436,224,248]" pageId="24" pageNumber="325">type</typeStatus>
|
||
series, have been pit-trapped and raked from leaf litter, spoil-heaps, and mats of dead vegetation and inside abandoned stick-ant (
|
||
<taxonomicName id="4C734D20FFEB8E63FE57FED829F25552" authority="Forel" authorityName="Forel" box="[393,693,295,320]" class="Insecta" family="Formicidae" genus="Iridomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="conifer">
|
||
<emphasis id="B907EAB1FFEB8E63FE57FED829375552" box="[393,624,295,320]" italics="true" pageId="24" pageNumber="325">Iridomyrmex conifer</emphasis>
|
||
Forel
|
||
</taxonomicName>
|
||
) nests. It has also been found under logs, mallee roots, rocks and rubbish, especially pieces of corrugated iron in disturbed areas adjacent to uncleared heath (
|
||
<bibRefCitation id="EFE24B52FFEB8E63FB63FEB32E795576" author="Bush" box="[1213,1342,332,357]" pageId="24" pageNumber="325" refString="Bush, B. (1981) Reptiles of the Kalgoorlie-Esperance Region. B. Bush, Perth, 46 pp." type="book" year="1981">Bush 1981</bibRefCitation>
|
||
). It also occupies the granitic heath where it is occasionally found under exfoliated granite slabs (B. Maryan, pers. obs.). There are no habitat data associated with specimens from the lower southwestern corner of Western
|
||
<collectingCountry id="F3647633FFEB8E63FAEBFE6B2EDB55BF" box="[1333,1436,404,429]" name="Australia" pageId="24" pageNumber="325">Australia</collectingCountry>
|
||
(WAM R129003, WAM R156978, WAM R172507).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0C662BFFEB8E63FF49FA8C2A8E51B5" httpUri="https://zenodo.org/record/244618/files/figure.png" pageId="24" pageNumber="325" targetBox="[151,1436,568,1373]" targetPageId="24">
|
||
<paragraph id="8BCC36A3FFEB8E63FF49FA8C2A8E51B5" blockId="24.[151,1435,1395,1447]" pageId="24" pageNumber="325">
|
||
<emphasis id="B907EAB1FFEB8E63FF49FA8C2A65519B" bold="true" box="[151,290,1395,1417]" pageId="24" pageNumber="325">FIGURE 12.</emphasis>
|
||
The habitat of
|
||
<taxonomicName id="4C734D20FFEB8E63FE17FA8B2976519B" box="[457,561,1395,1417]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFEB8E63FE17FA8B2976519B" box="[457,561,1395,1417]" italics="true" pageId="24" pageNumber="325">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFEB8E63FDE5FA8C29C8519B" box="[571,655,1395,1417]" pageId="24" pageNumber="325" rank="species">
|
||
<emphasis id="B907EAB1FFEB8E63FDE5FA8C29C8519B" bold="true" box="[571,655,1395,1417]" pageId="24" pageNumber="325">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, dense scrub heath on sandplain at Stirling Range National Park, Western Australia (image‒B. Maryan).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3696528FFEB8E62FF19FA2B2A0C54E5" lastPageId="25" lastPageNumber="326" pageId="24" pageNumber="325" type="discussion">
|
||
<paragraph id="8BCC36A3FFEB8E63FF19FA2B29D652FB" blockId="24.[151,1437,1492,2021]" pageId="24" pageNumber="325">
|
||
<emphasis id="B907EAB1FFEB8E63FF19FA2B2A7C51FF" bold="true" box="[199,315,1492,1517]" pageId="24" pageNumber="325">Remarks.</emphasis>
|
||
<bibRefCitation id="EFE24B52FFEB8E63FE9CFA2B2ABD51FF" author="Bush" box="[322,506,1492,1517]" pageId="24" pageNumber="325" refString="Bush, B. (1981) Reptiles of the Kalgoorlie-Esperance Region. B. Bush, Perth, 46 pp." type="book" year="1981">Bush (1981: 21)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE24B52FFEB8E63FDD8FA2B282F51FF" author="Wilson" box="[518,872,1492,1517]" pageId="24" pageNumber="325" refString="Wilson, S. K. & Knowles, D. G. (1988) Australia's Reptiles. A Photographic Reference to the Terrestrial Reptiles of Australia. Collins, Sydney, 447 pp." type="book" year="1988">Wilson & Knowles (1988: 246)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFE24B52FFEB8E63FC40FA2B2F2351FF" author="Bush" box="[926,1124,1492,1517]" pageId="24" pageNumber="325" refString="Bush, B., Maryan, B., Browne-Cooper & Robinson, D. (2007) Reptiles and Frogs in the bush: Southwestern Australia. University of Western Australia Press, Perth, 302 pp." type="book" year="2007">
|
||
Bush
|
||
<emphasis id="B907EAB1FFEB8E63FC01FA2A2F5051FF" box="[991,1047,1492,1517]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
|
||
(2007
|
||
</bibRefCitation>
|
||
: 135a) illustrated
|
||
<taxonomicName id="4C734D20FFEB8E63FAF2FA2A2EDB51FF" box="[1324,1436,1492,1517]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFEB8E63FAF2FA2A2EDB51FF" box="[1324,1436,1492,1517]" italics="true" pageId="24" pageNumber="325">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFEB8E63FF49FA072BB55203" box="[151,242,1528,1553]" pageId="24" pageNumber="325" rank="species">
|
||
<emphasis id="B907EAB1FFEB8E63FF49FA072BB55203" bold="true" box="[151,242,1528,1553]" pageId="24" pageNumber="325">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
as
|
||
<taxonomicName id="4C734D20FFEB8E63FEC4FA062AD85203" box="[282,415,1528,1553]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFEB8E63FEC4FA062AD85203" box="[282,415,1528,1553]" italics="true" pageId="24" pageNumber="325">D. australis</emphasis>
|
||
</taxonomicName>
|
||
. Additionally,
|
||
<bibRefCitation id="EFE24B52FFEB8E63FD94FA0728E45203" author="Wilson" box="[586,931,1528,1553]" pageId="24" pageNumber="325" refString="Wilson, S. K. & Knowles, D. G. (1988) Australia's Reptiles. A Photographic Reference to the Terrestrial Reptiles of Australia. Collins, Sydney, 447 pp." type="book" year="1988">Wilson & Knowles (1988: 96)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFE24B52FFEB8E63FC02FA072E5A5202" author="Wilson" box="[988,1309,1528,1553]" pageId="24" pageNumber="325" refString="Wilson, S. & Swan, G. (2013) A Complete Guide to Reptiles of Australia. Fourth Edition. New Holland Publishers, Sydney, 592 pp." type="book" year="2013">Wilson & Swan (2013: 144)</bibRefCitation>
|
||
refer to
|
||
<taxonomicName id="4C734D20FFEB8E63FA5FFA062BBE5227" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFEB8E63FA5FFA062BBE5227" italics="true" pageId="24" pageNumber="325">D. australis</emphasis>
|
||
</taxonomicName>
|
||
individuals from the far southwest of range as bluish grey, a colouration feature consistent with
|
||
<taxonomicName id="4C734D20FFEB8E63FAF2F9E22EDB5227" box="[1324,1436,1564,1589]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFEB8E63FAF2F9E22EDB5227" box="[1324,1436,1564,1589]" italics="true" pageId="24" pageNumber="325">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFEB8E63FF49F9BF2BB5524B" box="[151,242,1600,1625]" pageId="24" pageNumber="325" rank="species">
|
||
<emphasis id="B907EAB1FFEB8E63FF49F9BF2BB5524B" bold="true" box="[151,242,1600,1625]" pageId="24" pageNumber="325">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Historically,
|
||
<taxonomicName id="4C734D20FFEB8E63FE55F9BE2957524B" box="[395,528,1600,1625]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFEB8E63FE55F9BE2957524B" box="[395,528,1600,1625]" italics="true" pageId="24" pageNumber="325">D. australis</emphasis>
|
||
</taxonomicName>
|
||
has been documented as not occurring in the humid deep southwest of Western
|
||
<collectingCountry id="F3647633FFEB8E63FF49F99B2BB9526F" box="[151,254,1636,1661]" name="Australia" pageId="24" pageNumber="325">Australia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFE24B52FFEB8E63FED2F99B2AD2526F" author="Kluge" box="[268,405,1636,1661]" pageId="24" pageNumber="325" refString="Kluge, A. G. (1974) A taxonomic revision of the lizard family Pygopodidae. Miscellaneous Publications of the Museum of Zoology, University of Michigan No. 152, 1 - 72." type="book chapter" year="1974">Kluge 1974</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE24B52FFEB8E63FE7EF99B29FD526F" author="Wilson" box="[416,698,1636,1661]" pageId="24" pageNumber="325" refString="Wilson, S. K. & Knowles, D. G. (1988) Australia's Reptiles. A Photographic Reference to the Terrestrial Reptiles of Australia. Collins, Sydney, 447 pp." type="book" year="1988">Wilson & Knowles 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE24B52FFEB8E63FD18F99A28C7526F" author="Storr" box="[710,896,1636,1661]" pageId="24" pageNumber="325" refString="Storr, G. M., Smith, L. A. & Johnstone, R. E. (1990) Lizards of Western Australia. III. Geckos & Pygopods. Western Australian Museum, Perth, 141 pp." type="book" year="1990">
|
||
Storr
|
||
<emphasis id="B907EAB1FFEB8E63FCDAF99A287B526F" box="[772,828,1636,1661]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
|
||
1990
|
||
</bibRefCitation>
|
||
). However, there are a few records of
|
||
<taxonomicName id="4C734D20FFEB8E63FAF2F99A2EDB526F" box="[1324,1436,1636,1661]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFEB8E63FAF2F99A2EDB526F" box="[1324,1436,1636,1661]" italics="true" pageId="24" pageNumber="325">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFEB8E63FF49F9772BB752B3" box="[151,240,1672,1697]" pageId="24" pageNumber="325" rank="species">
|
||
<emphasis id="B907EAB1FFEB8E63FF49F9772BB752B3" bold="true" box="[151,240,1672,1697]" pageId="24" pageNumber="325">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in this area where they appear to be scarce when compared to the Esperance Plains. This apparent scarcity is documented by
|
||
<bibRefCitation id="EFE24B52FFEB8E63FEB1F952290352D7" box="[367,580,1708,1733]" pageId="24" pageNumber="325" refString="How, R. A., Dell, J. & Humphreys, W. F. (1987) The ground vertebrate fauna of coastal areas between Busselton and Albany, Western Australia. Records of the Western Australian Museum, 13, 553 - 574." type="journal article">
|
||
How
|
||
<emphasis id="B907EAB1FFEB8E63FE6FF9522AAA52D7" box="[433,493,1708,1733]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
|
||
(1987)
|
||
</bibRefCitation>
|
||
, who recorded ‘
|
||
<taxonomicName id="4C734D20FFEB8E63FCDBF95228C952D7" box="[773,910,1708,1733]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFEB8E63FCDBF95228C952D7" box="[773,910,1708,1733]" italics="true" pageId="24" pageNumber="325">D. australis</emphasis>
|
||
</taxonomicName>
|
||
’ from only Two Peoples Bay in a survey of herpetofauna between Busselton and Albany.
|
||
</paragraph>
|
||
<paragraph id="8BCC36A3FFEB8E62FF19F90B2A0C54E5" blockId="24.[151,1437,1492,2021]" lastBlockId="25.[151,1436,151,248]" lastPageId="25" lastPageNumber="326" pageId="24" pageNumber="325">
|
||
Biological surveys conducted on the Esperance Plains to date indicate
|
||
<taxonomicName id="4C734D20FFEB8E63FC04F90A2F0D531F" box="[986,1098,1780,1805]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFEB8E63FC04F90A2F0D531F" box="[986,1098,1780,1805]" italics="true" pageId="24" pageNumber="325">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFEB8E63FB8EF90B2FEE531F" box="[1104,1193,1780,1805]" pageId="24" pageNumber="325" rank="species">
|
||
<emphasis id="B907EAB1FFEB8E63FB8EF90B2FEE531F" bold="true" box="[1104,1193,1780,1805]" pageId="24" pageNumber="325">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
to be widespread and locally abundant in areas of suitable habitat (
|
||
<bibRefCitation id="EFE24B52FFEB8E63FD75F8E728FB5323" author="Chapman" box="[683,956,1816,1841]" pageId="24" pageNumber="325" refString="Chapman, A. & Dell, J. (1975) Reptiles, Amphibians and Fishes. In: A biological survey of Cape Le Grand National Park. Records of the Western Australian Museum, 1 (Supplement), pp. 34 - 38." type="journal article" year="1975">Chapman & Dell 1975</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE24B52FFEB8E63FC12F8E72F0B5323" author="Bush" box="[972,1100,1816,1841]" pageId="24" pageNumber="325" refString="Bush, B. (1985) Some reptiles and frogs recorded in Stokes National Park. The Western Australian Naturalist, 16, 52." type="journal article" year="1985">Bush 1985</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE24B52FFEB8E63FB82F8E72ED05323" author="Chapman" box="[1116,1431,1816,1841]" pageId="24" pageNumber="325" refString="Chapman, A. & Newbey, K. R. (1995) A biological survey of the Fitzgerald area, Western Australia. CALM Science, 3 (Supplement), 1 - 258." type="journal article" year="1995">Chapman & Newbey 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE24B52FFEB8E63FF49F8C32AC65347" author="Sanders" box="[151,385,1852,1877]" pageId="24" pageNumber="325" refString="Sanders, A., Chapman, A., Teale, R. J. & Harold, G. (2012) Vertebrate fauna of the Fitzgerald Biosphere Reserve, Western Australia. The Western Australian Naturalist, 28, 141 - 253." type="journal article" year="2012">
|
||
Sanders
|
||
<emphasis id="B907EAB1FFEB8E63FF22F8C22A7D5347" box="[252,314,1852,1877]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="4C734D20FFEB8E63FE6BF8C229785347" box="[437,575,1852,1877]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFEB8E63FE6BF8C229785347" box="[437,575,1852,1877]" italics="true" pageId="24" pageNumber="325">D. australis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<bibRefCitation id="EFE24B52FFEB8E63FD82F8C329A25347" author="Bush" box="[604,741,1852,1877]" pageId="24" pageNumber="325" refString="Bush, B. (1983) A record of reproduction in captive Delma australis and D. fraseri (Lacertilia: Pygopodidae). Herpetofauna, 15, 11 - 12." type="journal article" year="1983">Bush (1983</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE24B52FFEB8E63FD29F8C228755347" author="Bush" box="[759,818,1853,1877]" pageId="24" pageNumber="325" refString="Bush, B. (1984) Seasonal aggregation behaviour in a mixed population of legless lizards, Delma australis and D. fraseri. Herpetofauna, 16, 1 - 6." type="journal article" year="1984">1984</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="4C734D20FFEB8E63FCB8F8C228B75347" box="[870,1008,1852,1877]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="B907EAB1FFEB8E63FCB8F8C228B75347" box="[870,1008,1852,1877]" italics="true" pageId="24" pageNumber="325">D. australis</emphasis>
|
||
</taxonomicName>
|
||
) provides additional information on reproduction in captivity and field observations of winter aggregations of
|
||
<taxonomicName id="4C734D20FFEB8E63FC3CF89E2F14536B" box="[994,1107,1888,1913]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFEB8E63FC3CF89E2F14536B" box="[994,1107,1888,1913]" italics="true" pageId="24" pageNumber="325">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFEB8E63FB82F89F2FF0536B" box="[1116,1207,1888,1913]" pageId="24" pageNumber="325" rank="species">
|
||
<emphasis id="B907EAB1FFEB8E63FB82F89F2FF0536B" bold="true" box="[1116,1207,1888,1913]" pageId="24" pageNumber="325">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from the Esperance Plains. The Esperance Plains bioregion is a biogeographically significant area rich in endemic plants, rare ecosystems, and vulnerable and specially protected fauna (
|
||
<bibRefCitation id="EFE24B52FFEB8E63FC84F8572F0753D2" author="Comer" box="[858,1088,1960,1985]" pageId="24" pageNumber="325" refString="Comer, S., Gilfillan, S., Barrett, S., Grant, M., Tiedemann, K. & Anderson, L. (2001) Esperance 2 (ESP 2 - Recherche subregion). A Biodiversity Audit of Western Australia's 53 Biogeographical Subregions in 2002. Department of Conservation and Land Management, Perth. [unkown pagination]" type="journal article" year="2001">
|
||
Comer
|
||
<emphasis id="B907EAB1FFEB8E63FC68F85628B153D3" box="[950,1014,1960,1985]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
). Approximately 87% of the Esperance Plains bioregion has been cleared and developed for intensive agriculture. However, much of the remaining vegetation is afforded statutory protection, including many nature reserves and national parks (Cape Arid, Cape Le Grand, Stokes, Fitzgerald River and Stirling Range National Parks) where
|
||
<taxonomicName id="4C734D20FFEA8E62FB78FF422E5E54C7" box="[1190,1305,188,213]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="326" phylum="Chordata" rank="species" species="hebesa" status="sp. nov.">
|
||
<emphasis id="B907EAB1FFEA8E62FB78FF422E5E54C7" box="[1190,1305,188,213]" italics="true" pageId="25" pageNumber="326">D. hebesa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457CAFFEA8E62FAFDFF432E3854C7" box="[1315,1407,188,213]" pageId="25" pageNumber="326" rank="species">
|
||
<emphasis id="B907EAB1FFEA8E62FAFDFF432E3854C7" bold="true" box="[1315,1407,188,213]" pageId="25" pageNumber="326">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is known to occur.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |