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<document id="256BC6E20565FBA719478CE4C989C82B" ID-DOI="10.5281/zenodo.202791" ID-GBIF-Dataset="fbccfa1d-1095-4a8f-8af3-ac63f906fdd1" ID-ISSN="1175-5326" ID-Zenodo-Dep="202791" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460227336941" checkinUser="plazi" docAuthor="Grossi, Paschoal Coelho" docDate="2011" docId="704BA378B252FF8CFF00561BDEBCFA1C" docLanguage="en" docName="zt02750p014.pdf" docOrigin="Zootaxa 2750" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Incadorcus" docType="key" docVersion="9" lastPageNumber="2" masterDocId="8C72DB00B253FF8DFF975406DB20FFAE" masterDocTitle="Review of Incadorcus Arnaud &amp; Bomans with the description of three new species from the Yungas of Peru and Bolivia (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="2" updateTime="1698238665424" updateUser="plazi">
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<mods:title id="69E329FE7D1707103EEEF92EA6D7EA9F">Review of Incadorcus Arnaud &amp; Bomans with the description of three new species from the Yungas of Peru and Bolivia (Coleoptera: Lucanidae: Lucaninae)</mods:title>
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<mods:date id="5D99B6E1E361F6E1BA376F9CB0E04B6E">2011</mods:date>
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<treatment id="704BA378B252FF8CFF00561BDEBCFA1C" ID-DOI="http://doi.org/10.5281/zenodo.6183953" ID-GBIF-Taxon="119386727" ID-Zenodo-Dep="6183953" LSID="urn:lsid:plazi:treatment:704BA378B252FF8CFF00561BDEBCFA1C" httpUri="http://treatment.plazi.org/id/704BA378B252FF8CFF00561BDEBCFA1C" lastPageNumber="2" pageId="1" pageNumber="2">
<subSubSection id="B0F841E5B252FF8CFF00561BD928FD49" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="F85D126EB252FF8CFF00561BD96AFD99" blockId="1.[151,586,541,567]" box="[151,586,541,567]" pageId="1" pageNumber="2">
<heading id="A315A502B252FF8CFF00561BD96AFD99" bold="true" box="[151,586,541,567]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="CA96CE7CB252FF8CFF00561BD96AFD99" bold="true" box="[151,586,541,567]" pageId="1" pageNumber="2">
Key to adults of
<taxonomicName id="3FE269EDB252FF8CFEF3561BDACDFD99" ID-CoL="543V" box="[356,493,541,567]" class="Insecta" family="Lucanidae" genus="Incadorcus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="CA96CE7CB252FF8CFEF3561BDACDFD99" bold="true" box="[356,493,541,567]" italics="true" pageId="1" pageNumber="2">Incadorcus</emphasis>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="F85D126EB252FF8CFF005662D928FD49" blockId="1.[151,1436,612,743]" pageId="1" pageNumber="2">Identification of males versus females: Male mandibles are strongly asymmetric and are as long as or longer than head length; female mandibles are almost symmetric, shorter than head length. The head of the males is as wide as both elytra; the female head is narrower than the width of both elytra. The external apex of the metatibiae is acute in males and truncate in females.</paragraph>
</subSubSection>
<subSubSection id="B0F841E5B252FF8CFF005711DEBCFA1C" pageId="1" pageNumber="2" type="key">
<keyStep id="43160ACBB252FF8CFF005711DEBCFCE4" pageId="1" pageNumber="2">
<paragraph id="F85D126EB252FF8CFF005711DEBCFC82" blockId="1.[151,1436,791,1458]" box="[151,1436,791,812]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF005711DEBCFC82" box="[151,1436,791,812]" pageId="1" pageNumber="2">1 Surface color usually uniformly dark with no dark red areas at sides............................................. 2</keyLead>
</paragraph>
<paragraph id="F85D126EB252FF8CFF005733DEBCFCE4" blockId="1.[151,1436,791,1458]" box="[151,1436,821,842]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF005733DEBCFCE4" box="[151,1436,821,842]" pageId="1" pageNumber="2">- Surface color usually dark red, especially at sides of pronotum and elytra......................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="43160ACBB252FF8CFF005754DEBBFC11" pageId="1" pageNumber="2">
<paragraph id="F85D126EB252FF8CFF005754DEBCFC2A" blockId="1.[151,1436,791,1458]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF005754DEBCFC2A" pageId="1" pageNumber="2">
2 Third interstriae appearing smooth, impunctate in basal half of disc. Female head and pronotum densely or moderately punc- tate..................................................................................
<taxonomicName id="3FE269EDB252FF8CFB7E5776DE40FC2B" ID-CoL="3PKZV" box="[1257,1376,879,901]" class="Insecta" family="Lucanidae" genus="Incadorcus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ashaninka" status="sp. nov.">
<emphasis id="CA96CE7CB252FF8CFB7E5776DFD1FC2B" box="[1257,1265,880,901]" italics="true" pageId="1" pageNumber="2">I</emphasis>
.
<emphasis id="CA96CE7CB252FF8CFB6A5776DE40FC2B" box="[1277,1376,880,901]" italics="true" pageId="1" pageNumber="2">ashaninka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1A57307B252FF8CFAF15769DEBCFC2A" box="[1382,1436,879,900]" pageId="1" pageNumber="2" rank="species">
<emphasis id="CA96CE7CB252FF8CFAF15769DEBCFC2A" bold="true" box="[1382,1436,879,900]" pageId="1" pageNumber="2">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="F85D126EB252FF8CFF00578BDEBBFC11" blockId="1.[151,1436,791,1458]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF00578BDEBBFC11" pageId="1" pageNumber="2">
- Third interstriae distinctly punctate in basal half. Females with head and pronotum finely punctate.....................................................................................................
<taxonomicName id="3FE269EDB252FF8CFB0657ACDEBBFC11" ID-CoL="3PL25" authority="Arnaud &amp; Bomans" authorityName="Arnaud &amp; Bomans" box="[1169,1435,938,959]" class="Insecta" family="Lucanidae" genus="Incadorcus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="zugeri">
<emphasis id="CA96CE7CB252FF8CFB0657ACDFB9FC11" box="[1169,1177,938,959]" italics="true" pageId="1" pageNumber="2">I</emphasis>
.
<emphasis id="CA96CE7CB252FF8CFB3357ACDFC0FC11" box="[1188,1248,938,959]" italics="true" pageId="1" pageNumber="2">zugeri</emphasis>
Arnaud &amp; Bomans
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="43160ACBB252FF8CFF0057C1DEBCFBB9" pageId="1" pageNumber="2">
<paragraph id="F85D126EB252FF8CFF0057C1DEBBFC54" blockId="1.[151,1436,791,1458]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF0057C1DEBBFC54" pageId="1" pageNumber="2">
3 Second and fourth interstriae shiny and costate; rest of elytra completely punctate; punctures setose with longer setae at elytral declivity......................................................................
<taxonomicName id="3FE269EDB252FF8CFBEC57E3DEBBFC54" ID-CoL="3PKZY" authority="Arnaud &amp; Bomans" authorityName="Arnaud &amp; Bomans" box="[1147,1435,997,1018]" class="Insecta" family="Lucanidae" genus="Incadorcus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="damasoi">
<emphasis id="CA96CE7CB252FF8CFBEC57E3DFA3FC54" box="[1147,1155,997,1018]" italics="true" pageId="1" pageNumber="2">I</emphasis>
.
<emphasis id="CA96CE7CB252FF8CFB1857E3DFC0FC54" box="[1167,1248,997,1018]" italics="true" pageId="1" pageNumber="2">damasoi</emphasis>
Arnaud &amp; Bomans
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F85D126EB252FF8CFF005004DEBCFBB9" blockId="1.[151,1436,791,1458]" box="[151,1436,1026,1047]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF005004DEBCFBB9" box="[151,1436,1026,1047]" pageId="1" pageNumber="2">- All interstriae weakly shiny, flat, at most subcostate; punctures appearing glabrous but with short setae.................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="43160ACBB252FF8CFF005019DEBCFBFC" pageId="1" pageNumber="2">
<paragraph id="F85D126EB252FF8CFF005019DEBCFB9A" blockId="1.[151,1436,791,1458]" box="[151,1436,1055,1076]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF005019DEBCFB9A" box="[151,1436,1055,1076]" pageId="1" pageNumber="2">4 Third interstriae almost impunctate; if punctate, punctures fine. Temporal process not distinct in males.................. 5</keyLead>
</paragraph>
<paragraph id="F85D126EB252FF8CFF00503BDEBCFBFC" blockId="1.[151,1436,791,1458]" box="[151,1436,1085,1106]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF00503BDEBCFBFC" box="[151,1436,1085,1106]" pageId="1" pageNumber="2">
- Third interstriae punctate; punctures moderate. Temporal process in males distinct (
<figureCitation id="60D90EEBB252FF8CFB95503BDF68FBFC" box="[1026,1096,1085,1106]" captionStart="FIGURES 19 22" captionStartId="7.[151,264,1459,1482]" captionTargetBox="[300,1288,349,1428]" captionTargetId="figure@7.[283,1304,340,1439]" captionTargetPageId="7" captionText="FIGURES 19 22. Mandibles, head and pronotum of male of Incadorcus species. 19 I. cuzcoensis. 20 I. michellae. 21 I. ayamaranus. 22 I. paulseni." httpUri="https://zenodo.org/record/202795/files/figure.png" pageId="1" pageNumber="2">Fig. 20</figureCitation>
) or not (
<figureCitation id="60D90EEBB252FF8CFB0D503BDFC2FBFC" box="[1178,1250,1085,1106]" captionStart="FIGURES 19 22" captionStartId="7.[151,264,1459,1482]" captionTargetBox="[300,1288,349,1428]" captionTargetId="figure@7.[283,1304,340,1439]" captionTargetPageId="7" captionText="FIGURES 19 22. Mandibles, head and pronotum of male of Incadorcus species. 19 I. cuzcoensis. 20 I. michellae. 21 I. ayamaranus. 22 I. paulseni." httpUri="https://zenodo.org/record/202795/files/figure.png" pageId="1" pageNumber="2">Fig. 21</figureCitation>
)............. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="43160ACBB252FF8CFF00505CDEBBFB69" pageId="1" pageNumber="2">
<paragraph id="F85D126EB252FF8CFF00505CDEBBFB22" blockId="1.[151,1436,791,1458]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF00505CDEBBFB22" pageId="1" pageNumber="2">
5 Femora black. Male head moderately punctate and mandibles thick, head strongly rounded externally at base, external furrow deeply concave.........................................................
<taxonomicName id="3FE269EDB252FF8CFBB5507EDEBBFB22" ID-CoL="3PL24" authority="Arnaud, Noguchi &amp; Bomans" authorityName="Arnaud, Noguchi &amp; Bomans" box="[1058,1435,1143,1165]" class="Insecta" family="Lucanidae" genus="Incadorcus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="shaunai">
<emphasis id="CA96CE7CB252FF8CFBB5507EDF0AFB23" box="[1058,1066,1144,1165]" italics="true" pageId="1" pageNumber="2">I</emphasis>
.
<emphasis id="CA96CE7CB252FF8CFBA1507EDFA2FB23" box="[1078,1154,1144,1165]" italics="true" pageId="1" pageNumber="2">shaunai</emphasis>
Arnaud, Noguchi &amp; Bomans
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F85D126EB252FF8CFF005093DEBBFB69" blockId="1.[151,1436,791,1458]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF005093DEBBFB69" pageId="1" pageNumber="2">
- Femora red. Male head finely punctate and mandibles slender, head weakly rounded externally at base, external furrow shal- low................................................................
<taxonomicName id="3FE269EDB252FF8CFB9F50B4DEBBFB69" ID-CoL="3PKZX" authority="Arnaud, Noguchi &amp; Bomans" authorityName="Arnaud, Noguchi &amp; Bomans" box="[1032,1435,1202,1223]" class="Insecta" family="Lucanidae" genus="Incadorcus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="cuzcoensis">
<emphasis id="CA96CE7CB252FF8CFB9F50B4DF30FB69" box="[1032,1040,1202,1223]" italics="true" pageId="1" pageNumber="2">I</emphasis>
.
<emphasis id="CA96CE7CB252FF8CFB8B50B4DFA2FB69" box="[1052,1154,1202,1223]" italics="true" pageId="1" pageNumber="2">cuzcoensis</emphasis>
Arnaud, Noguchi &amp; Bomans
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="43160ACBB252FF8CFF0050C9DEBCFA92" pageId="1" pageNumber="2">
<paragraph id="F85D126EB252FF8CFF0050C9DEBBFAAC" blockId="1.[151,1436,791,1458]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF0050C9DEBBFAAC" pageId="1" pageNumber="2">
6 Temporal process distinctly projected laterally. Third interstriae completely punctate, punctures moderate in size. Male mandi- bles with external furrow exceeding half of mandibular length.........................
<taxonomicName id="3FE269EDB252FF8CFBF850EBDEBBFAAC" authority="Arnaud &amp; Bomans" authorityName="Arnaud &amp; Bomans" box="[1135,1435,1261,1282]" class="Insecta" family="Lucanidae" genus="Incadorcus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="michellae">
<emphasis id="CA96CE7CB252FF8CFBF850EBDFC0FAAC" box="[1135,1248,1261,1282]" italics="true" pageId="1" pageNumber="2">I. michellae</emphasis>
Arnaud &amp; Bomans
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F85D126EB252FF8CFF00510CDEBCFA92" blockId="1.[151,1436,791,1458]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF00510CDEBCFA92" pageId="1" pageNumber="2">- Temporal process not projected laterally, simply rounded or almost absent. Third interstriae completely punctate or not. Male mandibles with external furrow exceeding half of mandibular length or confined to basal third....................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="43160ACBB252FF8CFF005143DEBCFA1C" pageId="1" pageNumber="2">
<paragraph id="F85D126EB252FF8CFF005143DEBCFAD9" blockId="1.[151,1436,791,1458]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF005143DEBCFAD9" pageId="1" pageNumber="2">
<specimenCount id="EEE4D9E7B252FF8CFF005143DA32FAF4" box="[151,274,1349,1370]" pageId="1" pageNumber="2" type="male">7 Male</specimenCount>
mandibles with external furrow confined to basal third. Third interstriae not totally punctate; punctures ending at apex of basal third............................................................................
<taxonomicName id="3FE269EDB252FF8CFB425164DE7FFAD9" ID-CoL="3PKZW" box="[1237,1375,1378,1399]" class="Insecta" family="Lucanidae" genus="Incadorcus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ayamaranus" status="sp. nov.">
<emphasis id="CA96CE7CB252FF8CFB425164DE7FFAD9" box="[1237,1375,1378,1399]" italics="true" pageId="1" pageNumber="2">I. ayamaranus</emphasis>
</taxonomicName>
<emphasis id="CA96CE7CB252FF8CFAF15164DEBCFAD9" bold="true" box="[1382,1436,1378,1399]" pageId="1" pageNumber="2">
<taxonomicNameLabel id="D1A57307B252FF8CFAF15164DEBCFAD9" box="[1382,1436,1378,1399]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F85D126EB252FF8CFF005179DEBCFA1C" blockId="1.[151,1436,791,1458]" pageId="1" pageNumber="2">
<keyLead id="4313B15BB252FF8CFF005179DEBCFA1C" pageId="1" pageNumber="2">
- Male mandibles with external furrow exceeding half of mandibular length. Third interstriae completely and uniformly punc- tate; punctures fine.......................................................................
<taxonomicName id="3FE269EDB252FF8CFB6C519BDE7FFA1C" ID-CoL="3PL23" box="[1275,1375,1437,1458]" class="Insecta" family="Lucanidae" genus="Incadorcus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="paulseni" status="sp. nov.">
<emphasis id="CA96CE7CB252FF8CFB6C519BDE7FFA1C" box="[1275,1375,1437,1458]" italics="true" pageId="1" pageNumber="2">I. paulseni</emphasis>
</taxonomicName>
<emphasis id="CA96CE7CB252FF8CFAF1519BDEBCFA1C" bold="true" box="[1382,1436,1437,1458]" pageId="1" pageNumber="2">
<taxonomicNameLabel id="D1A57307B252FF8CFAF1519BDEBCFA1C" box="[1382,1436,1437,1458]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
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