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<mods:title id="7B78904B049438F1BF6CB784896217AD">Bryozoan faunas at the Tortonian-Messinian transition. A palaeoenvironmental case study from Crete Island, eastern Mediterranean</mods:title>
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<mods:affiliation id="F30B346CA617B826D25EB0DBDEF20D5B">National &amp; Kapodistrian University of Athens, Faculty of Geology and Geoenvironment, Department of Historical Geology and Paleontology 15784, Athens (Greece) and CR 2 P (CNRS, MNHN, UPMC, Sorbonne Université), Département Origines et Évolution, UMR 7207, Muséum national dHistoire naturelle, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:affiliation id="074D9D8DB2B07E655F34B6171F59D5DC">Géosciences Montpellier, Université de Montpellier Université des Antilles, CNRS, Pointe à Pitre, Guadeloupe, FWI (France)</mods:affiliation>
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<mods:date id="3D9B8A06817C81D17E8D7D52C4B6D75C">2021</mods:date>
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<taxonomicName id="4C0B4D122F0C5365FC43FE74FAFDF4F7" authority="(Lamarck, 1816)" authorityName=", Onychocella" baseAuthorityName="Lamarck" baseAuthorityYear="1816" box="[967,1301,438,466]" class="Stenolaemata" family="Crisiidae" genus="Nellia" kingdom="Animalia" order="Cyclostomatida" pageId="17" pageNumber="1380" phylum="Bryozoa" rank="species" species="tenella">
<emphasis id="B97FEA832F0C5365FC43FE74FBB0F4F4" bold="true" box="[967,1112,438,465]" italics="true" pageId="17" pageNumber="1380">Nellia tenella</emphasis>
(
<bibRefCitation id="EF9A4B602F0C5365FBE2FE75FAE5F4F7" author="LAMARCK J. B. P. A. DE" box="[1126,1293,439,466]" pageId="17" pageNumber="1380" refId="ref26290" refString="LAMARCK J. B. P. A. DE 1816. - Histoire naturelle des Animaux sans Vertebres. Verdiere, Paris, 568 p. https: // doi. org / 10.5962 / bhl. title. 12712" type="book" year="1816">Lamarck, 1816</bibRefCitation>
)
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(
<figureCitation id="13302A142F0C5365FBC3FE14FB7FF4D4" box="[1095,1175,470,497]" captionStart="FIG" captionStartId="18.[132,143,1917,1934]" captionTargetBox="[158,1430,215,1886]" captionTargetId="figure-25@18.[702,1250,773,1340]" captionTargetPageId="18" captionText="FIG. 9. — A, B, Cupuladria cf. canariensis (Busk, 1859); A, frontal view of a whole colony, KER39: AMPG(IV) 3078a; B, dorsal view of a whole colony, KER39: AMPG(IV) 3078b; C, D, Discoporella reussiana (Manzoni,1869); C, frontal view of a whole colony,KER30:AMPG(IV) 3102; D, dorsal view of a whole colony,KER30: AMPG(IV) 3101; E, Nellia tenella (Lamarck, 1816), detail of an internode showing two zooids in frontal view, KER17: AMPG(IV) 3150a; F-H, Canda rectangulata Udin, 1964; F, dorsal view of an internode fragment, FAN35: AMPG(IV) 3506a; G, frontal view of an internode fragment, FAN35: AMPG(IV) 3506b; H, Detail of the same fragment, FAN35: AMPG(IV) 3506b. Scale bars: A-D, 1 mm; E, H, 100 µm; F-G, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.5796589" httpUri="https://zenodo.org/record/5796589/files/figure.png" pageId="17" pageNumber="1380">Fig. 9E</figureCitation>
)
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<treatmentCitation id="0AAA10802F0C5365FCA9FDD2FB64F70D" author="LAMARCK J. B. P. A. DE" box="[813,1164,528,552]" page="135" pageId="17" pageNumber="1380" year="1816">
<taxonomicName id="4C0B4D122F0C5365FCA9FDD2FB64F70D" authority="Lamarck, 1816: 135" authorityName="Lamarck" authorityPageNumber="135" authorityYear="1816" box="[813,1164,528,552]" class="Gymnolaemata" family="Cellariidae" genus="Cellaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="1380" phylum="Bryozoa" rank="species" species="tenella">
<emphasis id="B97FEA832F0C5365FCA9FDD2FC54F70D" box="[813,956,528,552]" italics="true" pageId="17" pageNumber="1380">Cellaria tenella</emphasis>
<bibRefCitation id="EF9A4B602F0C5365FC46FDD2FB64F70D" author="LAMARCK J. B. P. A. DE" box="[962,1164,528,552]" pageId="17" pageNumber="1380" refId="ref26290" refString="LAMARCK J. B. P. A. DE 1816. - Histoire naturelle des Animaux sans Vertebres. Verdiere, Paris, 568 p. https: // doi. org / 10.5962 / bhl. title. 12712" type="book" year="1816">Lamarck, 1816: 135</bibRefCitation>
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.
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<emphasis id="B97FEA832F0C5365FCA9FD82FC46F77D" box="[813,942,576,600]" italics="true" pageId="17" pageNumber="1380">Nellia oculata</emphasis>
<bibRefCitation id="EF9A4B602F0C5365FC43FD82FBBEF77D" author="BUSK G." box="[967,1110,575,600]" pageId="17" pageNumber="1380" pagination="1 - 54" refId="ref20868" refString="BUSK G. 1852. - Catalogue of the Marine Polyzoa in the British Museum. Pt. I. Cheilostomata (part). London: 1 - 54. https: // www. biodiversitylibrary. org / page / 18304571" type="book chapter" year="1852">Busk 1852: 18</bibRefCitation>
, pl. 64, fig. 6; pl. 65 (bis), fig. 4. —
<bibRefCitation id="EF9A4B602F0C5365FCA8FD98FBE3F757" author="MOISSETTE P. &amp; DELRIEU B. &amp; TSAGARIS S." box="[812,1035,602,626]" pageId="17" pageNumber="1380" pagination="75 - 123" refId="ref27989" refString="MOISSETTE P., DELRIEU B. &amp; TSAGARIS S. 1993. - Bryozoaires du bassin neogene d'Heraklion (Crete centrale, Grece). Le Miocene superieur: premiers resultats. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 190: 75 - 123." type="journal article" year="1993">
Moissette
<emphasis id="B97FEA832F0C5365FC0DFD99FC5CF757" box="[905,948,602,626]" italics="true" pageId="17" pageNumber="1380">et al.</emphasis>
1993: 98
</bibRefCitation>
, fig.5i. —
<bibRefCitation id="EF9A4B602F0C5365FBE8FD98FAC4F757" author="MOISSETTE P." box="[1132,1324,602,626]" pageId="17" pageNumber="1380" pagination="181 - 203" refId="ref27753" refString="MOISSETTE P. 1997. - Bryozoaires recoltes dans les unites messiniennes de sondages offshore dans le golfe de Gabes (Tunisie). Revue de Micropaleontologie 40: 181 - 203. https: // doi. org / 10.1016 / S 0035 - 1598 (97) 90554 - 7" type="journal article" year="1997">Moissette 1997: 192</bibRefCitation>
, pl. 2, figs 5-6.
</paragraph>
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<taxonomicName id="4C0B4D122F0C5365FCA9FD48FC41F787" authorityName=", Onychocella" baseAuthorityName="Lamarck" baseAuthorityYear="1816" box="[813,937,650,674]" class="Stenolaemata" family="Crisiidae" genus="Nellia" kingdom="Animalia" order="Cyclostomatida" pageId="17" pageNumber="1380" phylum="Bryozoa" rank="species" species="tenella">
<emphasis id="B97FEA832F0C5365FCA9FD48FC41F787" box="[813,937,650,674]" italics="true" pageId="17" pageNumber="1380">Nellia tenella</emphasis>
</taxonomicName>
Ziko
<emphasis id="B97FEA832F0C5365FC7CFD49FBCFF787" box="[1016,1063,650,674]" italics="true" pageId="17" pageNumber="1380">et al.</emphasis>
2016: 18, pl. 2, fig. 6.
</paragraph>
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OCCURRENCE. — Eocene:
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,
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,
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,
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,
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(
<bibRefCitation id="EF9A4B602F0C5365FCB1FD17FC4DF7C9" author="BRAGA G." box="[821,933,724,749]" pageId="17" pageNumber="1380" pagination="71 - 92" refId="ref20646" refString="BRAGA G. 2008. - Atlas of Cenozoic Bryozoa of north-eastern Italy (Venetia Region). Lavori della Societa Veneziana di Scienze Naturale 33: 71 - 92." type="journal article" year="2008">Braga 2008</bibRefCitation>
). Oligocene:
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,
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,
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(
<bibRefCitation id="EF9A4B602F0C5365FB74FD16FC88F622" author="EL SAFORI Y. A. &amp; MUFTAH A. M." pageId="17" pageNumber="1380" pagination="315 - 334" refId="ref23245" refString="EL SAFORI Y. A. &amp; MUFTAH A. M. 2007. - Oligocene bryozoans from Al Jabal al Akhdar, Libya. Egyptian Journal of Paleontology 7: 315 - 334." type="journal article" year="2007">El Safori &amp; Muftah 2007</bibRefCitation>
),
<collectingCountry id="F31C76012F0C5365FCF0FD2DFC4DF622" box="[884,933,751,775]" name="United Arab Emirates" pageId="17" pageNumber="1380">UAE</collectingCountry>
(
<bibRefCitation id="EF9A4B602F0C5365FC37FD2DFB9AF622" author="BRAGA G. &amp; BAHR S. A." box="[947,1138,751,775]" pageId="17" pageNumber="1380" pagination="241 - 265" refId="ref20677" refString="BRAGA G. &amp; BAHR S. A. 2003. - Bryozoa from the Oligocene succession of Jabal Hafit, Al Ain, United Arab Emirates and Arabian Peninsula. Bollettino della Societa Paleontologica Italiana 42: 241 - 265." type="journal article" year="2003">Braga &amp; Bahr 2003</bibRefCitation>
). Early Miocene:
<collectingCountry id="F31C76012F0C5365FAA5FD2DFAB8F622" box="[1313,1360,751,775]" name="United States of America" pageId="17" pageNumber="1380">USA</collectingCountry>
(
<bibRefCitation id="EF9A4B602F0C5365FADAFD2DFC21F604" author="DI MARTINO E. &amp; TAYLOR P. D. &amp; PORTELL R. W." pageId="17" pageNumber="1380" pagination="97 - 200" refId="ref22659" refString="DI MARTINO E., TAYLOR P. D. &amp; PORTELL R. W. 2017. - Bryozoans from the lower Miocene Chipola Formation, Calhoun County, Florida, USA. Bulletin of the Florida Museum of Natural History 53 (4): 97 - 200." type="journal article" year="2017">
Di Martino
<emphasis id="B97FEA832F0C5365FCD9FCC8FC67F604" box="[861,911,777,801]" italics="true" pageId="17" pageNumber="1380">et al.</emphasis>
2017
</bibRefCitation>
),
<collectingCountry id="F31C76012F0C5365FC65FCCBFB5EF604" box="[993,1206,777,801]" name="Dominican Republic" pageId="17" pageNumber="1380">Dominican Republic</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FB46FCC8FAEFF604" box="[1218,1287,778,801]" name="France" pageId="17" pageNumber="1380">France</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FA96FCCBFA80F604" box="[1298,1384,777,801]" name="Portugal" pageId="17" pageNumber="1380">Portugal</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FAF0FCC8FA46F607" box="[1396,1454,778,802]" name="Egypt" pageId="17" pageNumber="1380">Egypt</collectingCountry>
(El Safori &amp; El-Sorogy 1999). Middle Miocene:
<collectingCountry id="F31C76012F0C5365FA85FCE1FAA7F61E" box="[1281,1359,803,827]" name="Jamaica" pageId="17" pageNumber="1380">Jamaica</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FADCFCE6FA47F619" box="[1368,1455,804,828]" name="Hungary" pageId="17" pageNumber="1380">Hungary</collectingCountry>
(
<bibRefCitation id="EF9A4B602F0C5365FCB1FCFCFC13F673" author="MOISSETTE P. &amp; DULAI A. &amp; MULLER P." box="[821,1019,830,854]" pageId="17" pageNumber="1380" pagination="300 - 314" refId="ref28080" refString="MOISSETTE P., DULAI A. &amp; MULLER P. 2006. - Bryozoan faunas in the Middle Miocene of Hungary: biodiversity and biogeography. Palaeogeography, Palaeoclimatology, Palaeoecology 233: 300 - 314. https: // doi. org / 10.1016 / j. palaeo. 2005.10.001" type="journal article" year="2006">
Moissette
<emphasis id="B97FEA832F0C5365FC13FCFDFC2CF673" box="[919,964,830,854]" italics="true" pageId="17" pageNumber="1380">et al.</emphasis>
2006
</bibRefCitation>
),
<collectingCountry id="F31C76012F0C5365FB8AFCFCFBBEF673" box="[1038,1110,830,854]" name="Austria" pageId="17" pageNumber="1380">Austria</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FBDBFCFCFB4BF670" box="[1119,1187,830,853]" name="France" pageId="17" pageNumber="1380">France</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FB2FFCFCFAE8F673" box="[1195,1280,830,854]" name="Portugal" pageId="17" pageNumber="1380">Portugal</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FA8CFCFCFAAAF673" box="[1288,1346,830,854]" name="Egypt" pageId="17" pageNumber="1380">Egypt</collectingCountry>
(Ziko
<emphasis id="B97FEA832F0C5365FA06FCFDFA58F673" box="[1410,1456,830,854]" italics="true" pageId="17" pageNumber="1380">et al.</emphasis>
2016). Late Miocene:
<collectingCountry id="F31C76012F0C5365FC7AFC9BFBBDF655" box="[1022,1109,857,880]" name="Morocco" pageId="17" pageNumber="1380">Morocco</collectingCountry>
(
<bibRefCitation id="EF9A4B602F0C5365FBE4FC9AFB19F654" author="EL HAJJAJI K." box="[1120,1265,856,881]" pageId="17" pageNumber="1380" pagination="1 - 355" refId="ref23176" refString="EL HAJJAJI K. 1992. - Les Bryozoaires du Miocene superieur du Maroc nord-Oriental. Documents des Laboratoires de Geologie de Lyon 123: 1 - 355." type="journal article" year="1992">El Hajjaji 1992</bibRefCitation>
),
<collectingCountry id="F31C76012F0C5365FA80FC9AFAAFF655" box="[1284,1351,856,880]" name="Algeria" pageId="17" pageNumber="1380">Algeria</collectingCountry>
(
<bibRefCitation id="EF9A4B602F0C5365FAD7FC9AFC88F6AE" author="MOISSETTE P." pageId="17" pageNumber="1380" pagination="1 - 351" refId="ref27655" refString="MOISSETTE P. 1988. - Faunes de bryozoaires du Messinien d'Algerie occidentale. Documents des Laboratoires de Geologie de Lyon 102: 1 - 351." type="journal article" year="1988">Moissette 1988</bibRefCitation>
), Crete,
<collectingCountry id="F31C76012F0C5365FC3EFCB1FBE8F6AE" box="[954,1024,883,907]" name="Turkey" pageId="17" pageNumber="1380">Turkey</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FB8FFCB1FBBBF6AE" box="[1035,1107,883,907]" name="Tunisia" pageId="17" pageNumber="1380">Tunisia</collectingCountry>
(
<bibRefCitation id="EF9A4B602F0C5365FBE7FCB1FB15F6AE" author="MOISSETTE P." box="[1123,1277,883,907]" pageId="17" pageNumber="1380" pagination="181 - 203" refId="ref27753" refString="MOISSETTE P. 1997. - Bryozoaires recoltes dans les unites messiniennes de sondages offshore dans le golfe de Gabes (Tunisie). Revue de Micropaleontologie 40: 181 - 203. https: // doi. org / 10.1016 / S 0035 - 1598 (97) 90554 - 7" type="journal article" year="1997">Moissette 1997</bibRefCitation>
). Pliocene:
<collectingCountry id="F31C76012F0C5365FAF2FCB1FCB6F680" name="Costa Rica" pageId="17" pageNumber="1380">Costa Rica</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FCEEFC4CFC54F680" box="[874,956,910,933]" name="Panama" pageId="17" pageNumber="1380">Panama</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FC4CFC4FFB49F680" box="[968,1185,909,933]" name="Dominican Republic" pageId="17" pageNumber="1380">Dominican Republic</collectingCountry>
,
<collectingCountry id="F31C76012F0C5365FB2AFC4FFB14F680" box="[1198,1276,909,933]" name="Jamaica" pageId="17" pageNumber="1380">Jamaica</collectingCountry>
(
<bibRefCitation id="EF9A4B602F0C5365FA8FFC4FFCB7F6E5" author="TAYLOR P. D. &amp; FOSTER T. S." pageId="17" pageNumber="1380" pagination="63 - 83" refId="ref31149" refString="TAYLOR P. D. &amp; FOSTER T. S. 1998. - Bryozoans from the Pliocene Bowden shell bed of Jamaica. Contributions to Tertiary and Quaternary Geology 35: 63 - 83." type="journal article" year="1998">Taylor &amp; Foster 1998</bibRefCitation>
;
<bibRefCitation id="EF9A4B602F0C5365FCEEFC6AFBD0F6E5" author="CHEETHAM A. H. &amp; JACKSON J. B. C. &amp; SANNER J. &amp; VENTOCILLA Y." box="[874,1080,936,960]" pageId="17" pageNumber="1380" pagination="159 - 192" refId="ref21747" refString="CHEETHAM A. H., JACKSON J. B. C., SANNER J., VENTOCILLA Y. 1999. - Neogene cheilostome Bryozoa of tropical America: Comparison and contrast between the Central American isthmus (Panama, Costa Rica) and the North-Central Caribbean (Dominican Republic), in COLLINS L. S. &amp; COATES A. G. (eds), A paleobiotic survey of caribbean faunas from the Neogene of the isthmus of Panama. Bulletins of American Paleontology 357: 159 - 192." type="journal article" year="1999">
Cheetham
<emphasis id="B97FEA832F0C5365FC50FC6BFBEAF6E5" box="[980,1026,936,960]" italics="true" pageId="17" pageNumber="1380">et al.</emphasis>
1999
</bibRefCitation>
). Pleistocene: Rhodes (PM, pers. obs.). Recent: eastern and western Atlantic, Indian Ocean, Pacific. This species is widely distributed in tropical to subtropical waters around the world. It lives at depths between 0 and
<quantity id="4CF39B742F0C5365FB48FC35FAE4F12A" box="[1228,1292,1015,1039]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="1380" unit="m" value="250.0">250 m</quantity>
(
<bibRefCitation id="EF9A4B602F0C5365FA98FC3AFA49F12A" author="FRANSEN C. H. J. M." box="[1308,1441,1015,1039]" pageId="17" pageNumber="1380" pagination="1 - 119" refId="ref23326" refString="FRANSEN C. H. J. M. 1986. - Caribbean Bryozoa: Anasca and Ascophora imperfecta of the inner bays of Curacao and Bonaire. Studies on the fauna of Curacao and other Caribbean Islands 210: 1 - 119. https: // repository. naturalis. nl / pub / 506057" type="journal article" year="1986">Fransen 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C311651A2F0C5365FCA9FBF4FB98F02A" pageId="17" pageNumber="1380" type="description">
<paragraph id="8BB436912F0C5365FCA9FBF4FC55F16B" blockId="17.[811,1458,1078,1296]" box="[813,957,1078,1104]" pageId="17" pageNumber="1380">DESCRIPTION</paragraph>
<paragraph id="8BB436912F0C5365FCA9FB94FB98F02A" blockId="17.[811,1458,1078,1296]" pageId="17" pageNumber="1380">Colony cellariiform. Internodes straight with quadrangular section and elongate zooids separated by shallow grooves. Opesia oval, occupying the greatest part of the zooidal frontal wall. Gymnocyst and cryptocyst well-developed proximally. Two small avicularia occur on the proximal part of the gymnocyst. Ovicells not observed.</paragraph>
</subSubSection>
<subSubSection id="C311651A2F0C5365FCA9FAF4FB5BF0CA" pageId="17" pageNumber="1380" type="discussion">
<paragraph id="8BB436912F0C5365FCA9FAF4FC79F06B" blockId="17.[811,1456,1334,1519]" box="[813,913,1334,1360]" pageId="17" pageNumber="1380">REMARKS</paragraph>
<paragraph id="8BB436912F0C5365FCA9FA97FB5BF0CA" blockId="17.[811,1456,1334,1519]" pageId="17" pageNumber="1380">
<emphasis id="B97FEA832F0C5365FCA9FA97FC56F04A" box="[813,958,1365,1391]" italics="true" pageId="17" pageNumber="1380">Nellia oculata</emphasis>
Busk is now regarded as a junior synonym of
<emphasis id="B97FEA832F0C5365FCA9FAB4FCA1F0AA" box="[813,841,1398,1423]" italics="true" pageId="17" pageNumber="1380">N.</emphasis>
<taxonomicName id="4C0B4D122F0C5365FCD5FAB7FBE7F0B5" authority="(Lamarck)" authorityName=", Onychocella" baseAuthorityName="Lamarck" baseAuthorityYear="1816" box="[849,1039,1397,1424]" class="Stenolaemata" family="Crisiidae" genus="Nellia" kingdom="Animalia" order="Cyclostomatida" pageId="17" pageNumber="1380" phylum="Bryozoa" rank="species" species="tenella">
<emphasis id="B97FEA832F0C5365FCD5FAB7FC7DF0AA" box="[849,917,1397,1423]" italics="true" pageId="17" pageNumber="1380">tenella</emphasis>
(Lamarck)
</taxonomicName>
. However, their very broad ecological, geographical and stratigraphical records may indicate either a living fossil species or a “cryptic species complex” (
<bibRefCitation id="EF9A4B602F0C5365FAD5FA77FC77F0CA" author="WINSTON J. E. &amp; VIEIRA L. M. &amp; WOOLLACOTT R. M." pageId="17" pageNumber="1380" pagination="139 - 239" refId="ref31899" refString="WINSTON J. E., VIEIRA L. M. &amp; WOOLLACOTT R. M. 2014. - Scientific Results of the Hassler Expedition. Bryozoa. No. 2. Brazil. Bulletin of the Museum of Comparative Zoology 161: 139 - 239. https: // doi. org / 10.3099 / MCZ 14.1" type="journal article" year="2014">
Winston
<emphasis id="B97FEA832F0C5365FCA9FA14FC88F0CA" box="[813,864,1493,1519]" italics="true" pageId="17" pageNumber="1380">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="EF9A4B602F0C5365FC29FA17FB4BF0CA" author="DI MARTINO E. &amp; TAYLOR P. D. &amp; PORTELL R. W." box="[941,1187,1493,1519]" pageId="17" pageNumber="1380" pagination="97 - 200" refId="ref22659" refString="DI MARTINO E., TAYLOR P. D. &amp; PORTELL R. W. 2017. - Bryozoans from the lower Miocene Chipola Formation, Calhoun County, Florida, USA. Bulletin of the Florida Museum of Natural History 53 (4): 97 - 200." type="journal article" year="2017">
Di Martino
<emphasis id="B97FEA832F0C5365FBB5FA14FB8DF0CA" box="[1073,1125,1493,1519]" italics="true" pageId="17" pageNumber="1380">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>