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<document id="8FD4612DFBEBA49569F6B9235C92729B" ID-DOI="10.11646/phytotaxa.381.1.9" ID-ISSN="1179-3163" ID-Zenodo-Dep="13726112" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725675168103" checkinUser="felipe" docAuthor="Sang, Julia, Kiew, Ruth &amp; Yiing, Ling Chea" docDate="2018" docId="03CBE12E707F793395B5A25BCCBDE330" docLanguage="en" docName="phytotaxa.381.1.9.pdf" docOrigin="Phytotaxa 381 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.381.1.9" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Begonia argentii Kiew &amp; S. Julia. A. Habit 2018, spec. nov." docType="treatment" docVersion="2" lastPageNumber="63" masterDocId="FFF29956707C7936953DA56DCD78E70D" masterDocTitle="The Begonia flora of Gunung Mulu and Gunung Buda National Parks, Sarawak, Borneo, including five new species" masterLastPageNumber="79" masterPageNumber="58" pageNumber="61" updateTime="1725688374631" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="2CD5487E1FC833649FA7083282FA3384" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1A7D27CBC1A71791595253F1D6F95D29">
<mods:title id="C05BEBB9C862358D8A4F12B3F0C7497C">The Begonia flora of Gunung Mulu and Gunung Buda National Parks, Sarawak, Borneo, including five new species</mods:title>
</mods:titleInfo>
<mods:name id="ED5AC9F97211DFD79DD662A9822ECC47" type="personal">
<mods:role id="7D3219680982243921EC3A240F621DD2">
<mods:roleTerm id="09711E62D5F081B1201CD45AA572847C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="350D7B3700DDAB586112EE21F563FE4E">Sang, Julia</mods:namePart>
<mods:nameIdentifier id="3FC255BA1611600E9A889B5C670F82CC" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
</mods:name>
<mods:name id="34CF8C6B7211E9BE51151B152B3E7498" type="personal">
<mods:role id="36B786B26BB38A6909D0157E10BD81A6">
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</mods:role>
<mods:namePart id="F03B57BDFC340E9F845D1B4B6F84CCFC">Kiew, Ruth</mods:namePart>
<mods:nameIdentifier id="3E6E49D1D726003EE12AA3BABFAB518D" type="email">ruth@frim.gov.my</mods:nameIdentifier>
</mods:name>
<mods:name id="4174E3A838FFB3C2F99BA35B38299A9D" type="personal">
<mods:role id="C96D5C458DC76316D21C24041A9B5D60">
<mods:roleTerm id="ED2236095CC342D59A2826C399175BB5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F900ABC7468BD73A6488A8C3C3ED99D">Yiing, Ling Chea</mods:namePart>
<mods:nameIdentifier id="51A2FC7CDEE7C283E9E4338C62D6BC8B" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7A0BDA7FCE29C581591655BBE4D1FCAC">text</mods:typeOfResource>
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<mods:titleInfo id="C8316ABFDE0A4CA2AE87550508BE91B7">
<mods:title id="AE12878284555594A066E3C135A8DFD2">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="10F84CC45B6DE85E16659F76C0B6B7C7">
<mods:date id="31CDF456010FB1507063FB9BA5828AB7">2018</mods:date>
<mods:detail id="5B8E8E7A4C5B07D3F4D68DD9B7D2E095" type="pubDate">
<mods:number id="7B6D4A0C41DC7A6E1B5B3D3BCA38391F">2018-12-07</mods:number>
</mods:detail>
<mods:detail id="E02A34443E957A90714CC56BA0B4640D" type="volume">
<mods:number id="BAE36A2F3329C0E0D4CBCF9AC78789D7">381</mods:number>
</mods:detail>
<mods:detail id="41E61F33185AADEF2CE2A02C7AB23D87" type="issue">
<mods:number id="24382D3FDE89BE8237977D4326C5B76A">1</mods:number>
</mods:detail>
<mods:extent id="52D91C81F9FE25D481187FBEB000C7B3" unit="page">
<mods:start id="0312B1B65606C37F195F755598EB2743">58</mods:start>
<mods:end id="333C1B6FE943F5F127F5D7D455DE8C3B">79</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="E9D2C7EF5DB93DA40FAEA2A797294F09">
<mods:url id="DCC4C66B3F6D406E6E89FD3F17CE7C14">http://dx.doi.org/10.11646/phytotaxa.381.1.9</mods:url>
</mods:location>
<mods:classification id="2D5A57D7ECA77FAAE81145F88AFDE52F">journal article</mods:classification>
<mods:identifier id="6B18FC901328FF1052DB068996CA2589" type="DOI">10.11646/phytotaxa.381.1.9</mods:identifier>
<mods:identifier id="1E79DD79DDE52D749414C03366C991B4" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="B26EE87817239BD02E2F4B234581BCF8" type="Zenodo-Dep">13726112</mods:identifier>
</mods:mods>
<treatment id="03CBE12E707F793395B5A25BCCBDE330" LSID="urn:lsid:plazi:treatment:03CBE12E707F793395B5A25BCCBDE330" httpUri="http://treatment.plazi.org/id/03CBE12E707F793395B5A25BCCBDE330" lastPageId="5" lastPageNumber="63" pageId="3" pageNumber="61">
<subSubSection id="C37803B3707F793595B5A25BCE66E042" box="[136,798,1844,1871]" pageId="3" pageNumber="61" type="nomenclature">
<paragraph id="8BDD5038707F793595B5A25BCE66E042" blockId="3.[136,1452,1844,2085]" box="[136,798,1844,1871]" pageId="3" pageNumber="61">
<heading id="D095E754707F793595B5A25BCE66E042" bold="true" box="[136,798,1844,1871]" fontSize="11" level="1" pageId="3" pageNumber="61" reason="1">
<emphasis id="B9168C2A707F793595B5A25BCE66E042" bold="true" box="[136,798,1844,1871]" pageId="3" pageNumber="61">
2.
<taxonomicName id="4C622BBB707F79359599A259CF5CE042" ID-CoL="7SDR7" authority="Kiew &amp; S.Julia" authorityName="Kiew &amp; S. Julia. A. Habit" authorityYear="2018" box="[164,548,1844,1871]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="61" phylum="Tracheophyta" rank="species" species="argentii" status="sp. nov.">
<emphasis id="B9168C2A707F79359599A259CC1EE043" bold="true" box="[164,358,1844,1870]" italics="true" pageId="3" pageNumber="61">Begonia argentii</emphasis>
Kiew &amp; S.Julia
</taxonomicName>
,
<emphasis id="B9168C2A707F7935970DA259CFE7E043" bold="true" box="[560,671,1844,1870]" italics="true" pageId="3" pageNumber="61">
<taxonomicNameLabel id="A2253151707F7935970DA259CFE7E043" box="[560,671,1844,1870]" pageId="3" pageNumber="61" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13594CBD707F79359793A25BCE6DE042" box="[686,789,1846,1871]" captionStart="FIGURE 3" captionStartId="4.[136,229,1931,1952]" captionTargetBox="[178,1415,232,1895]" captionTargetId="figure-14@4.[151,1436,190,1906]" captionTargetPageId="4" captionText="FIGURE 3. Begonia argentii Kiew &amp; S.Julia. A. Habit; B. Portion of male inflorescence; C. Male flower; D. Stamen; E &amp; F. Female flower; G. Outer tepal; H. Inner tepal; I. Stigma; J. Fruit; K. Transverse section of fruit.All from Argent &amp; Jermy 981." figureDoi="http://doi.org/10.5281/zenodo.13726120" httpUri="https://zenodo.org/record/13726120/files/figure.png" pageId="3" pageNumber="61">Figure 3</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37803B3707F793595B5A231CC05E0D0" pageId="3" pageNumber="61" type="description">
<paragraph id="8BDD5038707F793595B5A231CC41E07C" blockId="3.[136,1452,1844,2085]" box="[136,313,1884,1905]" pageId="3" pageNumber="61">
Sect.
<taxonomicName id="4C622BBB707F79359580A231CC41E07C" box="[189,313,1884,1905]" class="Magnoliopsida" family="Begoniaceae" genus="Petermannia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="61" phylum="Tracheophyta" rank="genus">
<emphasis id="B9168C2A707F79359580A231CC41E07C" box="[189,313,1884,1905]" italics="true" pageId="3" pageNumber="61">Petermannia</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BDD5038707F793595B5A2EDCC05E0D0" blockId="3.[136,1452,1844,2085]" pageId="3" pageNumber="61">
It is most similar to
<taxonomicName id="4C622BBB707F79359474A2EDCE78E098" authority="Kiew &amp; S. Julia (2007: 223)" authorityName="Kiew &amp; S. Julia" authorityPageNumber="223" authorityYear="2007" box="[329,768,1920,1941]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="61" phylum="Tracheophyta" rank="species" species="punchak">
<emphasis id="B9168C2A707F79359474A2EDCC97E098" box="[329,495,1920,1941]" italics="true" pageId="3" pageNumber="61">Begonia punchak</emphasis>
Kiew &amp; S.Julia (2007: 223)
</taxonomicName>
in its habit and leaf characters; but
<taxonomicName id="4C622BBB707F7935916BA2EDC9C3E098" authorityName="Kiew &amp; S. Julia. A. Habit" authorityYear="2018" box="[1110,1211,1920,1941]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="61" phylum="Tracheophyta" rank="species" species="argentii">
<emphasis id="B9168C2A707F7935916BA2EDC9C3E098" box="[1110,1211,1920,1941]" italics="true" pageId="3" pageNumber="61">B. argentii</emphasis>
</taxonomicName>
is different in its smaller stipules 9–16 ×
<quantity id="4C9AFDDD707F79359467A2C9CCD3E0B4" box="[346,427,1956,1977]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="3" pageNumber="61" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
(vs. ca. 22 ×
<quantity id="4C9AFDDD707F79359709A2C9CF08E0B4" box="[564,624,1956,1977]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="61" unit="mm" value="8.0">8 mm</quantity>
), 2-tepaled male flower (vs. 4-tepaled), and larger capsules, 19–20 ×
<quantity id="4C9AFDDD707F79359025A2C9C8F9E0B4" box="[1304,1409,1956,1977]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="3" pageNumber="61" unit="mm" value="20.0" valueMax="22.0" valueMin="18.0">18–22 mm</quantity>
(vs. 10–13
<emphasis id="B9168C2A707F793595C2A2A5CC75E0D0" bold="true" box="[255,269,1992,2013]" pageId="3" pageNumber="61">×</emphasis>
<quantity id="4C9AFDDD707F7935942EA2A5CC0AE0D0" box="[275,370,1992,2013]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="3" pageNumber="61" unit="mm" value="12.0" valueMax="15.0" valueMin="9.0">9–15 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C37803B3707F793395B5A281C873E5BC" lastPageId="5" lastPageNumber="63" pageId="3" pageNumber="61" type="materials_examined">
<paragraph id="8BDD5038707F793595B5A281CCACEF28" blockId="3.[136,1452,1844,2085]" pageId="3" pageNumber="61">
<emphasis id="B9168C2A707F793595B5A281CDBAEF0C" bold="true" box="[136,194,2028,2049]" pageId="3" pageNumber="61">Type:</emphasis>
—
<materialsCitation id="3B0A5A65707F793595E6A281CCA9EF28" collectingDate="1978-04-14" collectionCode="KEP, K" collectorName="Argent &amp; Jermy" country="Malaysia" county="Borneo" location="Gunung Mulu National Park" municipality="Marudi District" pageId="3" pageNumber="61" specimenCount="2" stateProvince="Sarawak" typeStatus="holotype">
<collectingCountry id="F37510A8707F793595E6A281CC22EF0C" box="[219,346,2028,2049]" name="Malaysia" pageId="3" pageNumber="61">MALAYSIA</collectingCountry>
.
<collectingCounty id="62BC28B4707F7935945EA281CCD4EF0C" box="[355,428,2028,2049]" pageId="3" pageNumber="61">Borneo</collectingCounty>
,
<collectingRegion id="49A69EDA707F7935948BA281CF73EF0C" box="[438,523,2028,2049]" country="Malaysia" name="Sarawak" pageId="3" pageNumber="61">Sarawak</collectingRegion>
,
<collectingMunicipality id="6BB9CA42707F79359729A281CFD5EF0C" box="[532,685,2028,2049]" pageId="3" pageNumber="61">Marudi District</collectingMunicipality>
:
<location id="8EBD06E3707F7935978BA281CEB3EF0C" LSID="urn:lsid:plazi:treatment:03CBE12E707F793395B5A25BCCBDE330:8EBD06E3707F7935978BA281CEB3EF0C" box="[694,971,2028,2049]" country="Malaysia" county="Borneo" municipality="Marudi District" name="Gunung Mulu National Park" pageId="3" pageNumber="61" stateProvince="Sarawak">Gunung Mulu National Park</location>
,
<location id="8EBD06E3707F793596E9A281C934EF0C" LSID="urn:lsid:plazi:treatment:03CBE12E707F793395B5A25BCCBDE330:8EBD06E3707F793596E9A281C934EF0C" box="[980,1100,2028,2049]" country="Malaysia" county="Borneo" municipality="Marudi District" name="Gunung Api" pageId="3" pageNumber="61" stateProvince="Sarawak">Gunung Api</location>
,
<date id="FFDC76F8707F7935916BA281C9A6EF0C" box="[1110,1246,2028,2049]" pageId="3" pageNumber="61" value="1978-04-14">
<collectingDate id="EF988F10707F7935916BA281C9A6EF0C" box="[1110,1246,2028,2049]" pageId="3" pageNumber="61" value="1978-04-14">14 April 1978</collectingDate>
</date>
,
<emphasis id="B9168C2A707F793591DAA281C8D3EF0C" box="[1255,1451,2028,2049]" italics="true" pageId="3" pageNumber="61">
<collectorName id="269735EE707F793591DAA281C85FEF0C" box="[1255,1319,2028,2049]" pageId="3" pageNumber="61">Argent</collectorName>
&amp;
<collectorName id="269735EE707F7935907AA281C8FAEF0C" box="[1351,1410,2028,2049]" pageId="3" pageNumber="61">Jermy</collectorName>
981
</emphasis>
(
<typeStatus id="54D9EE9A707F793595F9AD7DCC62EF28" box="[196,282,2064,2085]" pageId="3" pageNumber="61" type="holotype">holotype</typeStatus>
<collectionCode id="ED73C8FD707F79359422AD7DCC2CEF28" box="[287,340,2064,2085]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="3" pageNumber="61" type="Herbarium">KEP</collectionCode>
!;
<typeStatus id="54D9EE9A707F7935945CAD7DCCDEEF28" box="[353,422,2064,2085]" pageId="3" pageNumber="61" type="isotype">isotype</typeStatus>
<collectionCode id="ED73C8FD707F79359490AD7DCCBCEF28" box="[429,452,2064,2085]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="61" type="Herbarium">K</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<caption id="DF1D00B07078793295B5A2E6C87CE0C9" ID-DOI="http://doi.org/10.5281/zenodo.13726120" ID-Zenodo-Dep="13726120" httpUri="https://zenodo.org/record/13726120/files/figure.png" pageId="4" pageNumber="62" startId="4.[136,229,1931,1952]" targetBox="[178,1415,232,1895]" targetPageId="4" targetType="figure">
<paragraph id="8BDD50387078793295B5A2E6C87CE0C9" blockId="4.[136,1452,1930,1988]" pageId="4" pageNumber="62">
<emphasis id="B9168C2A7078793295B5A2E6CD87E0AD" bold="true" box="[136,255,1931,1952]" pageId="4" pageNumber="62">FIGURE 3.</emphasis>
<taxonomicName id="4C622BBB70787932943BA2E7CFD4E0AD" authority="Kiew &amp; S. Julia. A. Habit" authorityName="Kiew &amp; S. Julia. A. Habit" authorityYear="2018" box="[262,684,1930,1952]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="62" phylum="Tracheophyta" rank="species" species="argentii">
<emphasis id="B9168C2A70787932943BA2E7CCD5E092" bold="true" box="[262,429,1930,1951]" italics="true" pageId="4" pageNumber="62">Begonia argentii</emphasis>
Kiew &amp; S.Julia.
<emphasis id="B9168C2A70787932976BA2E6CF15E0AD" bold="true" box="[598,621,1931,1952]" pageId="4" pageNumber="62">A.</emphasis>
Habit
</taxonomicName>
;
<emphasis id="B9168C2A70787932978AA2E6CFB5E0AD" bold="true" box="[695,717,1931,1952]" pageId="4" pageNumber="62">B.</emphasis>
Portion of male inflorescence;
<emphasis id="B9168C2A70787932913CA2E6C960E0AD" bold="true" box="[1025,1048,1931,1952]" pageId="4" pageNumber="62">C.</emphasis>
Male flower;
<emphasis id="B9168C2A707879329199A2E6C9C4E0AD" bold="true" box="[1188,1212,1931,1952]" pageId="4" pageNumber="62">D.</emphasis>
Stamen;
<emphasis id="B9168C2A707879329024A2E6C825E0AD" bold="true" box="[1305,1373,1931,1952]" pageId="4" pageNumber="62">E &amp; F.</emphasis>
Female flower;
<emphasis id="B9168C2A7078793295EEA2C2CD94E0C9" bold="true" box="[211,236,1967,1988]" pageId="4" pageNumber="62">G.</emphasis>
Oute
<emphasis id="B9168C2A70787932941CA2C2CC54E0C9" bold="true" box="[289,300,1967,1988]" pageId="4" pageNumber="62">r</emphasis>
tepal;
<emphasis id="B9168C2A707879329451A2C2CCFDE0C9" bold="true" box="[364,389,1967,1988]" pageId="4" pageNumber="62">H.</emphasis>
Inner tepal;
<emphasis id="B9168C2A7078793294C2A2C2CF76E0C9" bold="true" box="[511,526,1967,1988]" pageId="4" pageNumber="62">I.</emphasis>
Stigma;
<emphasis id="B9168C2A707879329758A2C2CF0FE0C9" bold="true" box="[613,631,1967,1988]" pageId="4" pageNumber="62">J.</emphasis>
Frui
<emphasis id="B9168C2A707879329798A2C2CFACE0C9" bold="true" box="[677,724,1967,1988]" pageId="4" pageNumber="62">t; K.</emphasis>
Transverse section of fruit. All from Argent &amp; Jermy 981.
</paragraph>
</caption>
<paragraph id="8BDD50387079793395B5A5CDC873E5BC" blockId="5.[136,1452,160,1085]" pageId="5" pageNumber="63">
Erect herb, whole plant glabrous. Stem little branched, quite woody,
<quantity id="4C9AFDDD7079793396A5A5CDCE8EE7B4" box="[920,1014,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="63" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
diameter, internodes
<quantity id="4C9AFDDD7079793391D1A5CDC812E7B4" box="[1260,1386,160,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="5" pageNumber="63" unit="cm" value="1.2" valueMax="1.6" valueMin="0.8">0.8–1.6 cm</quantity>
long. Stipules lanceolate, 9–16 ×
<quantity id="4C9AFDDD7079793394FDA5A9CF58E7D0" box="[448,544,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="63" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
, margin entire, apex acute, caducous.
<emphasis id="B9168C2A7079793396F1A5A9C966E7D0" bold="true" box="[972,1054,196,221]" pageId="5" pageNumber="63">Leaves</emphasis>
alternate, distant. Petiole
<quantity id="4C9AFDDD70797933907EA5A9C8D3E7D0" box="[1347,1451,196,221]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="5" pageNumber="63" unit="cm" value="1.7" valueMax="2.0" valueMin="1.4">1.4–2 cm</quantity>
long on lower leaves,
<quantity id="4C9AFDDD707979339441A585CC80E60C" box="[380,504,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="5" pageNumber="63" unit="cm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 cm</quantity>
long on upper leaves, slightly grooved. Laminas plain green, stiff, thickly coriaceous when dried, oblique, ovate, 4–5.2 ×
<quantity id="4C9AFDDD707979339727A461CFE2E628" box="[538,666,268,293]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="5" pageNumber="63" unit="cm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 cm</quantity>
, asymmetric, broad side
<quantity id="4C9AFDDD707979339692A461C956E628" box="[943,1070,268,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="5" pageNumber="63" unit="cm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7–1.9 cm</quantity>
, base cordate but not overlapping, basal lobes
<quantity id="4C9AFDDD707979339430A45DCCF6E644" box="[269,398,304,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="5" pageNumber="63" unit="cm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 cm</quantity>
, margin slightly dentate, apex sharply acute; veins palmate-pinnate, palmate ones, branching once or twice about halfway to margin, 3 veins on either side of midrib, 2 veins in basal lobes, prominent and slightly raised on both sides. Inflorescences protogynous, paniculate, axillary from the upper leaf axils, to
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long, ca.
<specimenCount id="9D649BB17079793390A3A415CDACE6B8" count="2" pageId="5" pageNumber="63" type="female">2 female</specimenCount>
flowers below, much branched with many male flowers above. Bracts lanceolate, ca. 12 ×
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, margin entire.
<emphasis id="B9168C2A7079793395B5A4ADCC6FE6D4" bold="true" box="[136,279,448,473]" pageId="5" pageNumber="63">Male flower</emphasis>
: pedicel ca.
<quantity id="4C9AFDDD70797933949FA4ADCC9AE6D4" box="[418,482,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="63" unit="mm" value="5.0">5 mm</quantity>
long; tepals 2, pink, broadly oval, 3.5–4 ×
<quantity id="4C9AFDDD707979339683A4ADC978E6D4" box="[958,1024,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="63" unit="mm" value="4.0">4 mm</quantity>
, margin entire, apex rounded; stamens ca. 25, cluster globose, sessile; filament ca.
<quantity id="4C9AFDDD70797933974EA489CFB1E6F0" box="[627,713,484,509]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="63" unit="mm" value="0.5">0.5 mm</quantity>
long; anthers oblong, ca. 0.8
<emphasis id="B9168C2A707979339129A489C95CE6F0" bold="true" box="[1044,1060,484,509]" pageId="5" pageNumber="63">×</emphasis>
<quantity id="4C9AFDDD707979339116A489C9FBE6F0" box="[1067,1155,484,509]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="63" unit="mm" value="0.5">0.5 mm</quantity>
, apex emarginate.
<emphasis id="B9168C2A707979339069A489CDACE52C" bold="true" pageId="5" pageNumber="63">Female flower</emphasis>
: pedicel
<quantity id="4C9AFDDD707979339404A765CCEDE52C" box="[313,405,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="63" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long, pinkish; ovary broadly ellipsoid, ca. 5 ×
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, wings 3, equal, locules 3, placentas 2 per locule; tepals 5, pink, broadly elliptic; outer 4 tepals 6–7 ×
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; inner tepal elliptic, ca. 6 ×
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, margin entire, apex broadly acute; styles 3, to
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long, bifurcating; stigma anchor-shaped, papillose forming a continuous twisted band.
<emphasis id="B9168C2A7079793395F1A719CC6DE580" bold="true" box="[204,277,628,653]" pageId="5" pageNumber="63">Fruits</emphasis>
obovate, 19–20 ×
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, glabrous, locules 3, wings equal, truncate or rounded distally, broadly acute proximally,
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wide, dehiscing between the wings and locules; pendent, pedicel
<quantity id="4C9AFDDD707979339170A7F5C9B2E5BC" box="[1101,1226,664,689]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="5" pageNumber="63" unit="cm" value="1.85" valueMax="2.2" valueMin="1.5">1.5–2.2 cm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="C37803B3707979339580A7D1CC61E44C" pageId="5" pageNumber="63" type="distribution">
<paragraph id="8BDD5038707979339580A7D1C9B3E5D8" blockId="5.[136,1452,160,1085]" box="[189,1227,700,725]" pageId="5" pageNumber="63">
<emphasis id="B9168C2A707979339580A7D1CC21E5D8" bold="true" box="[189,345,700,725]" pageId="5" pageNumber="63">Distribution:</emphasis>
—
<collectingCountry id="F37510A870797933944BA7D1CF72E5D8" box="[374,522,700,725]" name="Malaysia" pageId="5" pageNumber="63">MALAYSIA</collectingCountry>
. Borneo. Endemic in
<collectingRegion id="49A69EDA7079793397C1A7D1CE18E5D8" box="[764,864,700,725]" country="Malaysia" name="Sarawak" pageId="5" pageNumber="63">Sarawak</collectingRegion>
. Known only from Gunung Api.
</paragraph>
<paragraph id="8BDD5038707979339580A78DCC61E44C" blockId="5.[136,1452,160,1085]" pageId="5" pageNumber="63">
<emphasis id="B9168C2A707979339580A78DCC5BE5F4" bold="true" box="[189,291,736,761]" pageId="5" pageNumber="63">Habitat:</emphasis>
—Limestone forest at
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elevation, growing on a ridge that has been burned within living memory and is now covered by
<taxonomicName id="4C622BBB7079793394A9A669CEFEE410" authority="St. John ex B. C. Stone" authorityName="B. C. Stone" box="[404,902,772,797]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="5" pageNumber="63" phylum="Tracheophyta" rank="species" species="calcinactus">
<emphasis id="B9168C2A7079793394A9A669CFF5E410" box="[404,653,772,797]" italics="true" pageId="5" pageNumber="63">Pandanus calcinactus</emphasis>
St.John
<emphasis id="B9168C2A7079793397CCA669CE72E410" box="[753,778,772,797]" italics="true" pageId="5" pageNumber="63">ex</emphasis>
B.C.Stone
</taxonomicName>
and
<taxonomicName id="4C622BBB7079793396FDA669C99EE410" authority="B. C. Stone" authorityName="B. C. Stone" box="[960,1254,772,797]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="5" pageNumber="63" phylum="Tracheophyta" rank="species" species="lepatophilus">
<emphasis id="B9168C2A7079793396FDA669C91FE410" box="[960,1127,772,797]" italics="true" pageId="5" pageNumber="63">P. lepatophilus</emphasis>
B.C.Stone
</taxonomicName>
, both endemic to Gunung Api.
</paragraph>
</subSubSection>
<subSubSection id="C37803B3707979339580A621CCBDE330" pageId="5" pageNumber="63" type="etymology">
<paragraph id="8BDD5038707979339580A621CFB7E484" blockId="5.[136,1452,160,1085]" pageId="5" pageNumber="63">
<emphasis id="B9168C2A707979339580A621CC30E468" bold="true" box="[189,328,844,869]" pageId="5" pageNumber="63">Etymology:</emphasis>
—Named for Dr G. C. George Argent, b. 1941, who collected widely in
<collectingCountry id="F37510A8707979339146A621C9ACE468" box="[1147,1236,844,869]" name="Malaysia" pageId="5" pageNumber="63">Malesia</collectingCountry>
including this new species and is an expert in
<taxonomicName id="4C622BBB707979339489A61DCF59E484" box="[436,545,880,905]" class="Magnoliopsida" family="Ericaceae" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="63" phylum="Tracheophyta" rank="family">Ericaceae</taxonomicName>
and
<taxonomicName id="4C622BBB707979339765A61DCFB4E484" box="[600,716,880,905]" class="Liliopsida" family="Musaceae" kingdom="Plantae" order="Zingiberales" pageId="5" pageNumber="63" phylum="Tracheophyta" rank="family">Musaceae</taxonomicName>
.
</paragraph>
<paragraph id="8BDD5038707979339580A6F9CC99E4F8" blockId="5.[136,1452,160,1085]" pageId="5" pageNumber="63">
<emphasis id="B9168C2A707979339580A6F9CC71E4A0" bold="true" box="[189,265,916,941]" pageId="5" pageNumber="63">Notes:</emphasis>
—Measurements in the above description are based on dried herbarium specimens. Among the Borneo begonias, it is similar to
<taxonomicName id="4C622BBB707979339499A6D5CE40E4DC" authority="K. G. Pearce (2003: 82)" authorityName="K. G. Pearce" authorityPageNumber="82" authorityYear="2003" box="[420,824,952,977]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="63" phylum="Tracheophyta" rank="species" species="subisensis">
<emphasis id="B9168C2A707979339499A6D5CF4FE4DC" box="[420,567,952,977]" italics="true" pageId="5" pageNumber="63">B. subisensis</emphasis>
K.G.Pearce (2003:82)
</taxonomicName>
in its habit but is generally less robust and has shorter internodes and smaller lamina.
</paragraph>
<paragraph id="8BDD5038707979339580A16DCCBDE330" blockId="5.[136,1452,160,1085]" pageId="5" pageNumber="63">Although it is known from a single collection, it may not be rare but is under-collected because it grows on a ridge that is dangerous to explore.</paragraph>
</subSubSection>
</treatment>
</document>
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