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<mods:title id="E94712DCBDB9070BCC8D9AF952B06F23">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<emphasis id="6E9B9D47FFD8FFF08C92FE19FEBF01D1" bold="true" box="[151,310,439,465]" italics="true" pageId="45" pageNumber="46">Pusia vexans</emphasis>
(
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)
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<figureCitation id="C4D45DD0FFD8FFF08C92FE75FF7901F5" box="[151,240,475,501]" captionStart="FIGURE 4" captionStartId="10.[151,250,1550,1574]" captionTargetBox="[162,1423,181,1527]" captionTargetId="figure-19@10.[162,1425,181,1527]" captionTargetPageId="10" captionText="FIGURE 4. Protoconchs of Paratethyan Costellariidae. A. Antithala filipescui nov. sp., Lăpugiu de Sus (Romania), NHMW 1876/0011/0012). B. Antithala sturi (Hoernes &amp; Auinger, 1880), Lăpugiu de Sus (Romania), NHMW 2020/0058/0001. C. Bellardithala kovaci nov. sp., Steinebrunn (Austria), NHMW 2020/0062/0001. D. Bellardithala partschi (Hörnes, 1852), Steinebrunn (Austria), NHMW 2020/0060/0003. E. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1876/0011/0008. F. Ebenomitra pseudopyramidella (Boettger, 1906), Steinebrunn (Austria), NHMW 1857/0014/0009. G. Fedosovia fuchsi (Hoernes &amp; Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1857/0024/0012. H. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. I. Pusia? confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. J. Pusia pseudorecticosta (Boettger, 1906), Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0005. K. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. L. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. M. Vexillum intermittens (Hoernes &amp; Auinger, 1880), paralectotype, Baden or Bad Vöslau (Austria), NHMW 2020/0087/0003. N. Vexillum michelottii (Hörnes, 1852), lectotype, Baden (Austria), NHMW 1851/0013/0001. O. Tosapusia pseudocupressina Bałuk, 1997, Lăpugiu de Sus (Romania), NHMW 1870/0033/0037." figureDoi="http://doi.org/10.5281/zenodo.4921850" httpUri="https://zenodo.org/record/4921850/files/figure.png" pageId="45" pageNumber="46">Figs 4L</figureCitation>
,
<figureCitation id="C4D45DD0FFD8FFF08CF9FE74FEA801F5" box="[252,289,474,501]" captionStart="FIGURE 15" captionStartId="44.[151,250,1686,1710]" captionTargetBox="[151,1436,236,1663]" captionTargetId="figure-33@44.[151,1436,236,1663]" captionTargetPageId="44" captionText="FIGURE 15 A1A2. Pusia pseudorecticosta (Boettger, 1906), lectotype, Steinebrunn (Austria), NHMW 2020/0076/0002. B1B2. Pusia pseudorecticosta (Boettger, 1906), paralectotype, Steinebrunn (Austria), NHMW 2020/0076/0001. C1C2. Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0004. D. Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0003. E1E2. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. F1F2. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0003. G1G2. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0001. H. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0079/0001. I1I2. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. J1J2. Pusia vexans (Boettger, 1902), lectotype, Coşteiu de Sus (Romania), SMF 360339. K1K2. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2018/0248/0327." figureDoi="http://doi.org/10.5281/zenodo.4921872" httpUri="https://zenodo.org/record/4921872/files/figure.png" pageId="45" pageNumber="46">15I</figureCitation>
<figureCitation id="C4D45DD0FFD8FFF08D24FE43FEC801F5" box="[289,321,475,507]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="45" pageNumber="46">
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I
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, J
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J
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, K
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<subScript id="C06B4310FFD8FFF08DA8FE43FE3C01FB" attach="left" box="[429,437,493,507]" fontSize="6" pageId="45" pageNumber="46">1</subScript>
K
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<taxonomicName id="9BEF3AD6FFD8FFF08CAFFD8EFD0D0237" box="[170,644,543,567]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="46" phylum="Mollusca" rank="variety" species="recticosta" subGenus="Costellaria" variety="vexans">
<emphasis id="6E9B9D47FFD8FFF08CAFFD8EFF6D0237" box="[170,228,544,567]" italics="true" pageId="45" pageNumber="46">Mitra</emphasis>
(
<emphasis id="6E9B9D47FFD8FFF08CF6FDB1FEEC0237" box="[243,357,543,567]" italics="true" pageId="45" pageNumber="46">Costellaria</emphasis>
)
<emphasis id="6E9B9D47FFD8FFF08D76FD8EFE5F0237" box="[371,470,544,567]" italics="true" pageId="45" pageNumber="46">recticosta</emphasis>
Bell. var.
<emphasis id="6E9B9D47FFD8FFF08E3AFD8EFD0D0237" box="[575,644,544,567]" italics="true" pageId="45" pageNumber="46">vexans</emphasis>
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n.—
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.
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<emphasis id="6E9B9D47FFD8FFF08C92FD90FF580255" box="[151,209,574,597]" italics="true" pageId="45" pageNumber="46">Mitra</emphasis>
(
<emphasis id="6E9B9D47FFD8FFF08CE5FD90FEDB0256" box="[224,338,574,598]" italics="true" pageId="45" pageNumber="46">Costellaria</emphasis>
)
<emphasis id="6E9B9D47FFD8FFF08D65FD90FD820256" box="[352,523,574,598]" italics="true" pageId="45" pageNumber="46">pseudorecticosta</emphasis>
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n. nom.
<taxonomicName id="9BEF3AD6FFD8FFF08E60FD90FC9D0256" authority="Bttgr." authorityName="Bttgr." box="[613,788,574,598]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="46" phylum="Mollusca" rank="variety" species="recticosta" variety="vexans">
var.
<emphasis id="6E9B9D47FFD8FFF08E95FD91FD5C0256" box="[656,725,575,598]" italics="true" pageId="45" pageNumber="46">vexans</emphasis>
Bttgr.
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<treatmentCitation id="DD4E6744FFD8FFF08F28FD90FC6E0256" author="Boettger, O." box="[813,999,574,598]" page="10" pageId="45" pageNumber="46" year="1906">
<bibRefCitation id="387E3CA4FFD8FFF08F28FD90FC6E0256" author="Boettger, O." box="[813,999,574,598]" pageId="45" pageNumber="46" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger 1906: 10</bibRefCitation>
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.
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</paragraph>
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<emphasis id="6E9B9D47FFD8FFF08C92FDF2FF670274" box="[151,238,604,628]" italics="true" pageId="45" pageNumber="46">Vexillum</emphasis>
(
<emphasis id="6E9B9D47FFD8FFF08CF8FDF2FEE60274" box="[253,367,604,628]" italics="true" pageId="45" pageNumber="46">Costellaria</emphasis>
)
<emphasis id="6E9B9D47FFD8FFF08D7BFDF2FDFD0274" box="[382,628,604,628]" italics="true" pageId="45" pageNumber="46">pseudorecticosta vexans</emphasis>
(Boettger)
</taxonomicName>
<treatmentCitation id="DD4E6744FFD8FFF08EFEFDF2FC290274" author="Zilch, A." box="[763,928,604,629]" page="259" pageId="45" pageNumber="46" year="1934">
<bibRefCitation id="387E3CA4FFD8FFF08EFEFDF2FC290274" author="Zilch, A." box="[763,928,604,629]" pageId="45" pageNumber="46" pagination="193 - 302" refId="ref63342" refString="Zilch, A. (1934) Zur Fauna des Mittel-Miocans von Kostej (Banat). Typus-Bestimmung und Tafeln zu O. Boettger's Bearbeitungen. Senckenbergiana, 16, 193 - 302." type="journal article" year="1934">Zilch 1934: 259</bibRefCitation>
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, pl. 17, fig. 16.
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</paragraph>
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<taxonomicName id="9BEF3AD6FFD8FFF08C92FDD5FDE50293" authority="(Boettger, 1901)" baseAuthorityName="Boettger" baseAuthorityYear="1901" box="[151,620,635,659]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="vexans" subGenus="Costellaria">
<emphasis id="6E9B9D47FFD8FFF08C92FDD5FF670293" box="[151,238,635,659]" italics="true" pageId="45" pageNumber="46">Vexillum</emphasis>
(
<emphasis id="6E9B9D47FFD8FFF08CF8FDD5FEE60293" box="[253,367,635,659]" italics="true" pageId="45" pageNumber="46">Costellaria</emphasis>
)
<emphasis id="6E9B9D47FFD8FFF08D7BFDD2FE4A0293" box="[382,451,636,659]" italics="true" pageId="45" pageNumber="46">vexans</emphasis>
(Boettger, 1901)
</taxonomicName>
<treatmentCitation id="DD4E6744FFD8FFF08E82FDD5FCAE0293" author="Baluk, W." box="[647,807,635,659]" page="37" pageId="45" pageNumber="46" year="1997">
<bibRefCitation id="387E3CA4FFD8FFF08E82FDD5FCAE0293" author="Baluk, W." box="[647,807,635,659]" pageId="45" pageNumber="46" pagination="1 - 75" refId="ref54188" refString="Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]" type="journal article" year="1997">Bałuk 1997: 37</bibRefCitation>
</treatmentCitation>
, pl. 10, figs 12.
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</paragraph>
<paragraph id="5C504155FFD8FFF08C92FD34FB3E02B2" blockId="45.[151,1207,543,690]" box="[151,1207,666,690]" pageId="45" pageNumber="46">
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<taxonomicName id="9BEF3AD6FFD8FFF08C92FD34FC9702B2" authority="(Boettger, 1906)" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[151,798,666,690]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="46" phylum="Mollusca" rank="subSpecies" species="pseudorecticosta" subGenus="Costellaria" subSpecies="vexans">
<emphasis id="6E9B9D47FFD8FFF08C92FD34FF6702B2" box="[151,238,666,690]" italics="true" pageId="45" pageNumber="46">Vexillum</emphasis>
(
<emphasis id="6E9B9D47FFD8FFF08CF8FD34FEE602B2" box="[253,367,666,690]" italics="true" pageId="45" pageNumber="46">Costellaria</emphasis>
)
<emphasis id="6E9B9D47FFD8FFF08D7BFD34FDFD02B2" box="[382,628,666,690]" italics="true" pageId="45" pageNumber="46">pseudorecticosta vexans</emphasis>
(
<bibRefCitation id="387E3CA4FFD8FFF08E84FD34FC9E02B2" author="Boettger, O." box="[641,791,666,690]" pageId="45" pageNumber="46" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
</taxonomicName>
<treatmentCitation id="DD4E6744FFD8FFF08F3DFD34FBBF02B2" author="Popa, M. &amp; Ianoliu, C." box="[824,1078,666,690]" page="86" pageId="45" pageNumber="46" year="2000">
<bibRefCitation id="387E3CA4FFD8FFF08F3DFD34FBBF02B2" author="Popa, M. &amp; Ianoliu, C." box="[824,1078,666,690]" pageId="45" pageNumber="46" pagination="79 - 92" refId="ref60686" refString="Popa, M. &amp; Ianoliu, C. (2000) Badenian mollusks from Rahchitova (Hațeg Depression, Romania). Studia universitatis Babes- Bolyai, Geologia, 45 (2), 79 - 92. [http: // scholarcommons. usf. edu / cgi / viewcontent. cgi? article = 1123 &amp; context = geologia] https: // doi. org / 10.5038 / 1937 - 8602.45.2.6" type="journal article" year="2000">Popa &amp; Ianoliu 2000: 86</bibRefCitation>
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, pl. 3, fig. 5.
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</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFD8FFF08C92FD4EFE2B055E" pageId="45" pageNumber="46" type="materials_examined">
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<emphasis id="6E9B9D47FFD8FFF08C92FD4EFECD02FA" bold="true" box="[151,324,736,762]" pageId="45" pageNumber="46">Type material.</emphasis>
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<typeStatus id="8354FFF7FFD8FFF08D4CFD4FFE3402FB" box="[329,445,737,763]" pageId="45" pageNumber="46" type="lectotype">Lectotype</typeStatus>
:
<specimenCode id="0C49E92EFFD8FFF08DC2FD4EFDD002FA" box="[455,601,736,762]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="45" pageNumber="46">SMF 360339</specimenCode>
(=
<collectionCode id="3AFED990FFD8FFF08E78FD4EFD3C02FA" box="[637,693,736,762]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="45" pageNumber="46">SMF</collectionCode>
<date id="28516795FFD8FFF08EBEFD4FFCAD02FA" box="[699,804,736,763]" pageId="45" pageNumber="46" value="2213-12">
<collectingDate id="38159E7DFFD8FFF08EBEFD4FFCAD02FA" box="[699,804,736,763]" pageId="45" pageNumber="46" value="2213-12">XII 2213</collectingDate>
</date>
a), Senckenberg Museum, Frankfurt/Main,
<collectingCountry id="24F801C5FFD8FFF08904FD4EFAE302FA" box="[1281,1386,736,762]" name="Germany" pageId="45" pageNumber="46">Germany</collectingCountry>
, SL:
<quantity id="9B17ECB0FFD8FFF08C92FCAAFF52031F" box="[151,219,772,799]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="45" pageNumber="46" unit="mm" value="6.0">6 mm</quantity>
, MD:
<quantity id="9B17ECB0FFD8FFF08D21FCAAFEF4031F" box="[292,381,772,799]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="45" pageNumber="46" unit="mm" value="2.4">2.4 mm</quantity>
,
<location id="5930178EFFD8FFF08D8CFCAAFDB8031E" LSID="urn:lsid:plazi:treatment:D446F043FFD8FFF08C92FE19FEBF06A2:5930178EFFD8FFF08D8CFCAAFDB8031E" box="[393,561,772,798]" country="Germany" name="Costeiu de Sus" pageId="45" pageNumber="46">
Coşteiu
<collectorName id="F11A2483FFD8FFF08DE2FCAAFDB8031E" box="[487,561,772,798]" pageId="45" pageNumber="46">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFD8FFF08E44FCAAFD21031E" box="[577,680,772,798]" name="Romania" pageId="45" pageNumber="46">Romania</collectingCountry>
<emphasis id="6E9B9D47FFD8FFF08EADFCAAFD38031E" bold="true" box="[680,689,772,798]" pageId="45" pageNumber="46">)</emphasis>
, illustrated in
<bibRefCitation id="387E3CA4FFD8FFF08F50FCAAFC55031E" author="Zilch, A." box="[853,988,772,798]" pageId="45" pageNumber="46" pagination="193 - 302" refId="ref63342" refString="Zilch, A. (1934) Zur Fauna des Mittel-Miocans von Kostej (Banat). Typus-Bestimmung und Tafeln zu O. Boettger's Bearbeitungen. Senckenbergiana, 16, 193 - 302." type="journal article" year="1934">Zilch (1934</bibRefCitation>
, pl. 17, fig. 16), figs 15J
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J
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.
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<typeStatus id="8354FFF7FFD8FFF0892EFCAAFF5F0343" pageId="45" pageNumber="46" type="paralectotype">Paralectotypes</typeStatus>
:
<collectionCode id="3AFED990FFD8FFF08CE4FC86FE900342" box="[225,281,808,834]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="45" pageNumber="46">SMF</collectionCode>
<date id="28516795FFD8FFF08D25FC87FE010342" box="[288,392,808,835]" pageId="45" pageNumber="46" value="2213-12">
<collectingDate id="38159E7DFFD8FFF08D25FC87FE010342" box="[288,392,808,835]" pageId="45" pageNumber="46" value="2213-12">XII 2213</collectingDate>
</date>
bc,
<location id="5930178EFFD8FFF08DB9FC86FD300343" LSID="urn:lsid:plazi:treatment:D446F043FFD8FFF08C92FE19FEBF06A2:5930178EFFD8FFF08DB9FC86FD300343" box="[444,697,808,835]" country="Germany" county="Main" municipality="Frankfurt" name="Senckenberg Museum" pageId="45" pageNumber="46">Senckenberg Museum</location>
,
<collectingMunicipality id="BC34DB2FFFD8FFF08EC1FC86FCB80342" box="[708,817,808,834]" pageId="45" pageNumber="46">Frankfurt</collectingMunicipality>
/
<collectingCounty id="B53139D9FFD8FFF08F3CFC86FCFD0342" box="[825,884,808,834]" pageId="45" pageNumber="46">Main</collectingCounty>
,
<collectingCountry id="24F801C5FFD8FFF08F7BFC86FC6E0342" box="[894,999,808,834]" name="Germany" pageId="45" pageNumber="46">Germany</collectingCountry>
</materialsCitation>
,
<materialsCitation id="EC874B08FFD8FFF08FF6FC86FA950343" ID-GBIF-Occurrence="3311814391" box="[1011,1308,808,835]" collectionCode="SMF" collectorName="de Sus" country="Romania" location="Costeiu de Sus" pageId="45" pageNumber="46" specimenCount="1" typeStatus="paralectotype">
<location id="5930178EFFD8FFF08FF6FC86FB100342" LSID="urn:lsid:plazi:treatment:D446F043FFD8FFF08C92FE19FEBF06A2:5930178EFFD8FFF08FF6FC86FB100342" box="[1011,1177,808,834]" country="Romania" name="Costeiu de Sus" pageId="45" pageNumber="46">
Coşteiu
<collectorName id="F11A2483FFD8FFF0884AFC86FB100342" box="[1103,1177,808,834]" pageId="45" pageNumber="46">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFD8FFF088ADFC86FA9D0343" box="[1192,1300,808,835]" name="Romania" pageId="45" pageNumber="46">Romania</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFD8FFF08CC2FCE2FCA603AF" blockId="45.[151,1437,736,1698]" pageId="45" pageNumber="46">
<emphasis id="6E9B9D47FFD8FFF08CC2FCE2FE320366" bold="true" box="[199,443,844,870]" pageId="45" pageNumber="46">Additional material.</emphasis>
<materialsCitation id="EC874B08FFD8FFF08DC6FCE3FADC0367" ID-GBIF-Occurrence="3311814347" box="[451,1365,844,871]" collectionCode="NHMW" country="Romania" location="Costeiu de Sus" pageId="45" pageNumber="46" specimenCode="NHMW 2020" specimenCount="1">
<specimenCode id="0C49E92EFFD8FFF08DC6FCE3FDE80366" box="[451,609,844,871]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="45" pageNumber="46" type="Museum">NHMW 2020</specimenCode>
/0081/0001, SL:
<quantity id="9B17ECB0FFD8FFF08F1AFCE2FCF10367" box="[799,888,844,871]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="45" pageNumber="46" unit="mm" value="5.4">5.4 mm</quantity>
, MD:
<quantity id="9B17ECB0FFD8FFF08FC7FCE2FB920367" box="[962,1051,844,871]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="45" pageNumber="46" unit="mm" value="2.1">2.1 mm</quantity>
,
<location id="5930178EFFD8FFF0882DFCE2FB580366" LSID="urn:lsid:plazi:treatment:D446F043FFD8FFF08C92FE19FEBF06A2:5930178EFFD8FFF0882DFCE2FB580366" box="[1064,1233,844,870]" country="Romania" name="Costeiu de Sus" pageId="45" pageNumber="46">Coşteiu de Sus</location>
(
<collectingCountry id="24F801C5FFD8FFF088E4FCE2FAC50367" box="[1249,1356,844,871]" name="Romania" pageId="45" pageNumber="46">Romania</collectingCountry>
)
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, 15I
<subScript id="C06B4310FFD8FFF08983FCF0FA07036C" attach="left" box="[1414,1422,862,876]" fontSize="6" pageId="45" pageNumber="46">1</subScript>
15I
<subScript id="C06B4310FFD8FFF08CB8FC2CFF4C0390" attach="left" box="[189,197,898,912]" fontSize="6" pageId="45" pageNumber="46">2</subScript>
, 4L;
<materialsCitation id="EC874B08FFD8FFF08D0CFCDFFB35038B" ID-GBIF-Occurrence="3311814366" box="[265,1212,880,907]" collectionCode="NHMW" country="Romania" location="Costeiu de Sus" pageId="45" pageNumber="46" specimenCode="NHMW 2020" specimenCount="1">
<specimenCode id="0C49E92EFFD8FFF08D0CFCDFFE20038A" box="[265,425,880,907]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="45" pageNumber="46" type="Museum">NHMW 2020</specimenCode>
/0081/0002, SL: 6.0 mm, MD:
<quantity id="9B17ECB0FFD8FFF08F1FFCDEFCFF038B" box="[794,886,880,907]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="45" pageNumber="46" unit="mm" value="2.5">2.5 mm</quantity>
,
<location id="5930178EFFD8FFF08F83FCDEFBBC038A" LSID="urn:lsid:plazi:treatment:D446F043FFD8FFF08C92FE19FEBF06A2:5930178EFFD8FFF08F83FCDEFBBC038A" box="[902,1077,880,906]" country="Romania" name="Costeiu de Sus" pageId="45" pageNumber="46">Coşteiu de Sus</location>
(
<collectingCountry id="24F801C5FFD8FFF0884CFCDEFB3D038B" box="[1097,1204,880,907]" name="Romania" pageId="45" pageNumber="46">Romania</collectingCountry>
)
</materialsCitation>
, 15K
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K
<subScript id="C06B4310FFD8FFF08923FC2CFAA70390" attach="left" box="[1318,1326,898,912]" fontSize="6" pageId="45" pageNumber="46">2</subScript>
;
<materialsCitation id="EC874B08FFD8FFF08944FCDFFCA203AF" ID-GBIF-Occurrence="3311814477" collectionCode="NHMW" country="Romania" location="Costeiu de Sus" pageId="45" pageNumber="46" specimenCode="NHMW 2018" specimenCount="47">
<specimenCode id="0C49E92EFFD8FFF08944FCDFFF4603AE" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="45" pageNumber="46" type="Museum">NHMW 2018</specimenCode>
/0248/0327,
<specimenCount id="4AE98ADCFFD8FFF08D58FC3AFE7103AE" box="[349,504,916,942]" pageId="45" pageNumber="46" type="generic">47 specimens</specimenCount>
,
<location id="5930178EFFD8FFF08E07FC3AFD2103AE" LSID="urn:lsid:plazi:treatment:D446F043FFD8FFF08C92FE19FEBF06A2:5930178EFFD8FFF08E07FC3AFD2103AE" box="[514,680,916,942]" country="Romania" name="Costeiu de Sus" pageId="45" pageNumber="46">Coşteiu de Sus</location>
(
<collectingCountry id="24F801C5FFD8FFF08EBDFC3AFCAD03AF" box="[696,804,916,943]" name="Romania" pageId="45" pageNumber="46">Romania</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFD8FFF08CC2FC16FB830516" blockId="45.[151,1437,736,1698]" pageId="45" pageNumber="46">
<emphasis id="6E9B9D47FFD8FFF08CC2FC16FE3203D2" bold="true" box="[199,443,952,978]" pageId="45" pageNumber="46">Revised description.</emphasis>
Shell small, moderately broad fusiform, with high conical spire. Protoconch (fig. 4L) multispiral, high conical, mammillate, of about 3.5 weakly convex whorls (height: 730 μm, diameter: 550 μm). Teleoconch consisting of five convex whorls, with periphery in abapical third, separated by impressed suture. Last whorl convex, somewhat inflated, constricted at base. Axial sculpture of swollen, convex axial ribs separated by concave interspaces of roughly equal width. Axial ribs becoming slightly more prominent towards abapical suture. Axial ribs and interspaces widening on last whorl, subobsolete towards aperture in some specimens. Aperture moderately narrow, ovoid, posteriorly narrowly angulated, with weak parietal swelling. Columellar callus narrow. Columella with four prominent columellar folds, weakening abapically. Outer lip thin, bearing about five prominent lirae within. Five to seven prominent spiral cords on weak fasciole. Siphonal canal moderately short to moderately long, narrow, straight or bent slightly to left, with very shallow siphonal notch.
</paragraph>
<paragraph id="5C504155FFD8FFF08CC2FA8EFE2B055E" blockId="45.[151,1437,736,1698]" pageId="45" pageNumber="46">
<emphasis id="6E9B9D47FFD8FFF08CC2FA8EFDB7053A" bold="true" box="[199,574,1312,1338]" pageId="45" pageNumber="46">Shell measurements and ratios.</emphasis>
SL: 4.96.0 mm, MD:
<quantity id="9B17ECB0FFD8FFF08F4CFA8EFC5A053B" box="[841,979,1312,1339]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="45" pageNumber="46" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.92.2 mm</quantity>
; AA: 3841°, SL/MD: 2.52.6, AL/AW: 3.44.3, AH/S: 2.42.7.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFD8FFF08CC2FAC6FDA80637" pageId="45" pageNumber="46" type="discussion">
<paragraph id="5C504155FFD8FFF08CC2FAC6FC1505EE" blockId="45.[151,1437,736,1698]" pageId="45" pageNumber="46">
<emphasis id="6E9B9D47FFD8FFF08CC2FAC6FEC50582" bold="true" box="[199,332,1384,1410]" pageId="45" pageNumber="46">Discussion.</emphasis>
This species is characterised by its small size, convex whorls and swollen axial ribs. There are no similar species in Paratethyan faunas.
<taxonomicName id="9BEF3AD6FFD8FFF08E4DFA22FA1105A6" authority="Landau, Ceulemans &amp; Van Dingenen, 2019" authorityName="Landau, Ceulemans &amp; Van Dingenen" authorityYear="2019" box="[584,1432,1420,1447]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="renauleauensis">
<emphasis id="6E9B9D47FFD8FFF08E4DFA22FD4505A6" box="[584,716,1420,1446]" italics="true" pageId="45" pageNumber="46">Ebenomitra</emphasis>
cf.
<emphasis id="6E9B9D47FFD8FFF08EFCFA22FC2B05A6" box="[761,930,1420,1446]" italics="true" pageId="45" pageNumber="46">renauleauensis</emphasis>
<bibRefCitation id="387E3CA4FFD8FFF08FACFA22FA1105A6" author="Landau, B. M. &amp; Ceulemans, L. &amp; Van Dingenen, F." box="[937,1432,1420,1447]" pageId="45" pageNumber="46" pagination="135 - 215" refId="ref58790" refString="Landau, B. M., Ceulemans, L. &amp; Van Dingenen, F. (2019) The upper Miocene gastropods of northwestern France, 4. Neogastropoda, Cainozoic Research, 19 / 2, 135 - 215." type="journal article" year="2019">Landau, Ceulemans &amp; Van Dingenen, 2019</bibRefCitation>
</taxonomicName>
, from the Tortonian of
<collectingCountry id="24F801C5FFD8FFF08D8AFA1FFE5705CB" box="[399,478,1457,1483]" name="France" pageId="45" pageNumber="46">France</collectingCountry>
, is superficially similar, but is distinguished by its lower spire, wider apical angle and lower number of axial ribs (see
<bibRefCitation id="387E3CA4FFD8FFF08DFEFA7AFD5E05EE" author="Landau, B. M. &amp; Ceulemans, L. &amp; Van Dingenen, F." box="[507,727,1492,1519]" pageId="45" pageNumber="46" pagination="135 - 215" refId="ref58790" refString="Landau, B. M., Ceulemans, L. &amp; Van Dingenen, F. (2019) The upper Miocene gastropods of northwestern France, 4. Neogastropoda, Cainozoic Research, 19 / 2, 135 - 215." type="journal article" year="2019">
Landau
<emphasis id="6E9B9D47FFD8FFF08E53FA7BFD0E05EE" box="[598,647,1492,1518]" italics="true" pageId="45" pageNumber="46">et al</emphasis>
., 2019
</bibRefCitation>
, pl. 50, figs 24).
</paragraph>
<paragraph id="5C504155FFD8FFF08CC2FA56FDA80637" blockId="45.[151,1437,736,1698]" pageId="45" pageNumber="46">
<emphasis id="6E9B9D47FFD8FFF08CC2FA56FE3F0612" bold="true" box="[199,438,1528,1554]" pageId="45" pageNumber="46">Palaeoenvironment.</emphasis>
The assemblages from Coşteiu de Sus suggests middle to outer neritic palaeoenvironments with soft bottom (own data).
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFD8FFF08CC2F9EEFEBF06A2" pageId="45" pageNumber="46" type="distribution">
<paragraph id="5C504155FFD8FFF08CC2F9EEFEBF06A2" blockId="45.[151,1437,736,1698]" pageId="45" pageNumber="46">
<emphasis id="6E9B9D47FFD8FFF08CC2F9EEFDEF065A" bold="true" box="[199,614,1600,1626]" pageId="45" pageNumber="46">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="6E9B9D47FFD8FFF08FB4F9EFFBEE065A" box="[945,1127,1601,1626]" italics="true" pageId="45" pageNumber="46">Korytnica Basin</emphasis>
: Korytnica (
<collectingCountry id="24F801C5FFD8FFF088F4F9EEFACB065B" box="[1265,1346,1600,1627]" name="Poland" pageId="45" pageNumber="46">Poland</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFD8FFF0895DF9EEFF5A067E" author="Baluk, W." pageId="45" pageNumber="46" pagination="1 - 75" refId="ref54188" refString="Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]" type="journal article" year="1997">Bałuk 1997</bibRefCitation>
);
<emphasis id="6E9B9D47FFD8FFF08CEEF9CAFEFE067E" box="[235,375,1636,1662]" italics="true" pageId="45" pageNumber="46">Făget Basin</emphasis>
: Coşteiu de Sus (
<collectingCountry id="24F801C5FFD8FFF08E4CF9CAFD3B067F" box="[585,690,1636,1663]" name="Romania" pageId="45" pageNumber="46">Romania</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFD8FFF08EC9F9CAFCFC067E" author="Boettger, O." box="[716,885,1636,1662]" pageId="45" pageNumber="46" pagination="1 - 200" refId="ref54619" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger 1902</bibRefCitation>
);
<emphasis id="6E9B9D47FFD8FFF08F89F9CBFB95067E" box="[908,1052,1637,1664]" italics="true" pageId="45" pageNumber="46">Hațeg Basin</emphasis>
: Răhchitova (
<collectingCountry id="24F801C5FFD8FFF088C5F9CAFAA0067F" box="[1216,1321,1636,1663]" name="Romania" pageId="45" pageNumber="46">Romania</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFD8FFF08947F9CAFEA206A2" author="Popa, M. &amp; Ianoliu, C." pageId="45" pageNumber="46" pagination="79 - 92" refId="ref60686" refString="Popa, M. &amp; Ianoliu, C. (2000) Badenian mollusks from Rahchitova (Hațeg Depression, Romania). Studia universitatis Babes- Bolyai, Geologia, 45 (2), 79 - 92. [http: // scholarcommons. usf. edu / cgi / viewcontent. cgi? article = 1123 &amp; context = geologia] https: // doi. org / 10.5038 / 1937 - 8602.45.2.6" type="journal article" year="2000">Popa &amp; Ianoliu 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>