An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea Author Harzhauser, Mathias 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at Author Landau, Bernard 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at text Zootaxa 2021 2021-06-09 4982 1 1 70 journal article 5925 10.11646/zootaxa.4982.1.1 f68b381d-d0dd-44e2-b9fe-6cf3c1aab1fa 1175-5326 4921842 58388FB8-128A-4381-83D1-3C508D0D3873 Pusia vexans ( Boettger, 1902 ) nov. comb. Figs 4L , 15I 1 –I 2 , J 1 –J 2 , K 1 –K 2 * Mitra ( Costellaria ) recticosta Bell. var. vexans n.— Boettger 1902: 13 . Mitra ( Costellaria ) pseudorecticosta n. nom. var. vexans Bttgr. — Boettger 1906: 10 . Vexillum ( Costellaria ) pseudorecticosta vexans (Boettger) — Zilch 1934: 259 , pl. 17, fig. 16. Vexillum ( Costellaria ) vexans (Boettger, 1901) — Bałuk 1997: 37 , pl. 10, figs 1–2. Vexillum ( Costellaria ) pseudorecticosta vexans ( Boettger, 1906 ) — Popa & Ianoliu 2000: 86 , pl. 3, fig. 5. Type material. Lectotype : SMF 360339 (= SMF XII 2213 a), Senckenberg Museum, Frankfurt/Main, Germany , SL: 6 mm , MD: 2.4 mm , Coşteiu de Sus ( Romania ) , illustrated in Zilch (1934 , pl. 17, fig. 16), figs 15J 1 –J 2 . Paralectotypes : SMF XII 2213 b–c, Senckenberg Museum , Frankfurt / Main , Germany , Coşteiu de Sus ( Romania ) . Additional material. NHMW 2020 /0081/0001, SL: 5.4 mm , MD: 2.1 mm , Coşteiu de Sus ( Romania ) , 15I 1 – 15I 2 , 4L; NHMW 2020 /0081/0002, SL: 6.0 mm, MD: 2.5 mm , Coşteiu de Sus ( Romania ) , 15K 1 –K 2 ; NHMW 2018 /0248/0327, 47 specimens , Coşteiu de Sus ( Romania ) . Revised description. Shell small, moderately broad fusiform, with high conical spire. Protoconch (fig. 4L) multispiral, high conical, mammillate, of about 3.5 weakly convex whorls (height: 730 μm, diameter: 550 μm). Teleoconch consisting of five convex whorls, with periphery in abapical third, separated by impressed suture. Last whorl convex, somewhat inflated, constricted at base. Axial sculpture of swollen, convex axial ribs separated by concave interspaces of roughly equal width. Axial ribs becoming slightly more prominent towards abapical suture. Axial ribs and interspaces widening on last whorl, subobsolete towards aperture in some specimens. Aperture moderately narrow, ovoid, posteriorly narrowly angulated, with weak parietal swelling. Columellar callus narrow. Columella with four prominent columellar folds, weakening abapically. Outer lip thin, bearing about five prominent lirae within. Five to seven prominent spiral cords on weak fasciole. Siphonal canal moderately short to moderately long, narrow, straight or bent slightly to left, with very shallow siphonal notch. Shell measurements and ratios. SL: 4.9–6.0 mm, MD: 1.9–2.2 mm ; AA: 38–41°, SL/MD: 2.5–2.6, AL/AW: 3.4–4.3, AH/S: 2.4–2.7. Discussion. This species is characterised by its small size, convex whorls and swollen axial ribs. There are no similar species in Paratethyan faunas. Ebenomitra cf. renauleauensis Landau, Ceulemans & Van Dingenen, 2019 , from the Tortonian of France , is superficially similar, but is distinguished by its lower spire, wider apical angle and lower number of axial ribs (see Landau et al ., 2019 , pl. 50, figs 2–4). Palaeoenvironment. The assemblages from Coşteiu de Sus suggests middle to outer neritic palaeoenvironments with soft bottom (own data). Distribution in Central Paratethys. Badenian (middle Miocene): Korytnica Basin : Korytnica ( Poland ) ( Bałuk 1997 ); Făget Basin : Coşteiu de Sus ( Romania ) ( Boettger 1902 ); Hațeg Basin : Răhchitova ( Romania ) ( Popa & Ianoliu 2000 ).