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<document id="DDED071B00C0B049339BB72ADF257271" ID-CLB-Dataset="7057" ID-DOI="10.11646/zootaxa.4964.3.8" ID-GBIF-Dataset="b8866544-2d1c-412c-be3c-5f497c7c98f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4714797" ID-ZooBank="A4C79E2B-8555-4351-824E-0B9096C3CD36" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619169124781" checkinUser="plazi" docAuthor="Wang, Lu-Yu, Framenau, Volker W. &amp; Zhang, Zhi-Sheng" docDate="2021" docId="3C488784FFC9E2570CCFFEA0F0BDFDE5" docLanguage="en" docName="zootaxa.4964.3.8.pdf" docOrigin="Zootaxa 4964 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Artoria hamata Wang &amp; Framenau &amp; Zhang 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="578" masterDocId="C071FFFCFFCFE2500C58FF8EF042FF84" masterDocTitle="A further study on the wolf spider subfamily Artoriinae from China (Araneae: Lycosidae)" masterLastPageNumber="584" masterPageNumber="571" pageNumber="577" updateTime="1698937571720" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="41852D5A5137A62E6E46FA0B0D7AFC3D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4A5377385A2FA0AA85602ACCD4033D20">A further study on the wolf spider subfamily Artoriinae from China (Araneae: Lycosidae)</mods:title>
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<mods:namePart id="60FAA7040E9EF42B15BC256BE056C929">Wang, Lu-Yu</mods:namePart>
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<mods:affiliation id="D065141F5ADEBADFE0449A2EDF035077">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China. https: // orcid. org / 0000 - 0002 - 5250 - 3473</mods:affiliation>
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<mods:namePart id="581C213618381469CD8C0AD5738C131A">Framenau, Volker W.</mods:namePart>
<mods:nameIdentifier id="59FF2EE88FE9FA240B6B3105190013D3" type="ORCID">0000-0002-9304-1789</mods:nameIdentifier>
<mods:affiliation id="66B8373DEE5BC1BE17EF57299EA2EF25">Harry Butler Institute, Murdoch University, Murdoch, Western Australia 6150, Australia. &amp; Department of Terrestrial Zoology, Western Australian Museum, Welshpool DC, Western Australia 6986, Australia &amp; Center of Natural History (CeNaK), University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany Corresponding author. zhangzs 327 @ qq. com; https: // orcid. org / 0000 - 0002 - 9304 - 1789</mods:affiliation>
<mods:nameIdentifier id="95C00005BBB5DCBA404FCECCBFF0DBFA" type="email">zhangzs327@qq.com</mods:nameIdentifier>
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<mods:namePart id="49DF610675ECCB3E781471F26C4A31FF">Zhang, Zhi-Sheng</mods:namePart>
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<mods:affiliation id="A00AA110C739B2545C4A7E6220B00773">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China. https: // orcid. org / 0000 - 0002 - 5250 - 3473</mods:affiliation>
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<mods:title id="D7164F209CD8753C06B4A0E23A2CBEF5">Zootaxa</mods:title>
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<mods:date id="840FC98CD24399315DA7CDBABF4580EF">2021</mods:date>
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<mods:number id="A5617CDB22B377DF586DBDF902617B17">4964</mods:number>
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<treatment id="3C488784FFC9E2570CCFFEA0F0BDFDE5" ID-DOI="http://doi.org/10.5281/zenodo.4714812" ID-GBIF-Taxon="181879848" ID-Zenodo-Dep="4714812" LSID="urn:lsid:plazi:treatment:3C488784FFC9E2570CCFFEA0F0BDFDE5" httpUri="http://treatment.plazi.org/id/3C488784FFC9E2570CCFFEA0F0BDFDE5" lastPageId="7" lastPageNumber="578" pageId="6" pageNumber="577">
<subSubSection id="FCFB6519FFC9E2560CCFFEA0F1F5FECC" box="[151,439,301,328]" pageId="6" pageNumber="577" type="nomenclature">
<paragraph id="B45E3692FFC9E2560CCFFEA0F1F5FECC" blockId="6.[151,439,301,365]" box="[151,439,301,328]" pageId="6" pageNumber="577">
<heading id="EF1681FEFFC9E2560CCFFEA0F1F5FECC" bold="true" box="[151,439,301,328]" fontSize="11" level="1" pageId="6" pageNumber="577" reason="1">
<emphasis id="8695EA80FFC9E2560CCFFEA0F1F5FECC" bold="true" box="[151,439,301,328]" pageId="6" pageNumber="577">
<taxonomicName id="73E14D11FFC9E2560CCFFEA0F110FECC" ID-CoL="98YZR" authority="Wang &amp; Framenau &amp; Zhang, 2021" authorityName="Wang &amp; Framenau &amp; Zhang" authorityYear="2021" box="[151,338,301,328]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="hamata" status="sp. nov.">
<emphasis id="8695EA80FFC9E2560CCFFEA0F110FECC" bold="true" box="[151,338,301,328]" italics="true" pageId="6" pageNumber="577">Artoria hamata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9DA657FBFFC9E2560D01FEA0F1F5FECC" box="[345,439,302,328]" pageId="6" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FCFB6519FFC9E2560CCFFEDCF163FEE8" box="[151,289,338,365]" pageId="6" pageNumber="577" type="description">
<paragraph id="B45E3692FFC9E2560CCFFEDCF163FEE8" blockId="6.[151,439,301,365]" box="[151,289,338,365]" pageId="6" pageNumber="577">
<figureCitation id="2CDA2A17FFC9E2560CCFFEDCF0B8FEE9" box="[151,250,338,365]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="6.[151,250,1511,1535]" captionStartId-1="7.[151,250,1637,1661]" captionTargetBox-0="[198,1391,718,1482]" captionTargetBox-1="[151,1435,633,1613]" captionTargetId-0="figure-147@6.[190,1397,712,1488]" captionTargetId-1="figure-193@7.[151,1435,633,1613]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 5. Artoria hamata sp. nov., holotype male (A–B), paratype male (C) and paratype female (D–E). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C.Apical part of bulb, ventral view; D. Epigyne, ventral view; E. Female internal genitalia, dorsal view. BEA, basoembolic apophysis; CO, copulatory opening; Em, embolus; FD, fertilization duct; HS, head of spermathecal; MA, median apophysis; SS, stalk of spermathecal; TA, terminal apophysis (= synembolus sensu Zyuzin 1993); Te, tegulum." captionText-1="FIGURE 6. Artoria hamata sp. nov., holotype male (A, E–F), paratype male (C–D, G) and paratype female (B, H–J). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male pedipalp, bulbus, ventral view; D. Same, retrolateral view; E. Left male pedipalp, ventral view; F. Same, retrolateral view; G. Apical part of bulb, ventral view; H–I. Epigyne, ventral view; J. Female internal genitalia, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.4714854" figureDoi-1="http://doi.org/10.5281/zenodo.4714860" httpUri-0="https://zenodo.org/record/4714854/files/figure.png" httpUri-1="https://zenodo.org/record/4714860/files/figure.png" pageId="6" pageNumber="577">Figs 5–6</figureCitation>
,
<figureCitation id="2CDA2A17FFC9E2560D5EFEDCF163FEE8" box="[262,289,338,364]" captionStart="FIGURE 11" captionStartId="12.[151,250,1665,1689]" captionTargetBox="[151,1435,181,1641]" captionTargetId="figure-23@12.[151,1435,181,1641]" captionTargetPageId="12" captionText="FIGURE 11. Distribution records of the wolf spider subfamily Artoriinae in China." figureDoi="http://doi.org/10.5281/zenodo.4714931" httpUri="https://zenodo.org/record/4714931/files/figure.png" pageId="6" pageNumber="577">11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FCFB6519FFC9E2560CCFFE14F2B7FE78" pageId="6" pageNumber="577" type="materials_examined">
<paragraph id="B45E3692FFC9E2560CCFFE14F2B7FE78" blockId="6.[151,1436,409,689]" pageId="6" pageNumber="577">
<emphasis id="8695EA80FFC9E2560CCFFE14F10AFE37" bold="true" box="[151,328,409,436]" pageId="6" pageNumber="577">Type material:</emphasis>
<materialsCitation id="04893CCFFFC9E2560D08FE14F4ECFE5C" ID-GBIF-Occurrence="3107827316" collectingDate="2013-05-22" collectorName="D. Wang &amp; X. K. Jiang" country="China" county="Pingwu County" elevation="2949" latitude="33.003418" location="Wanglang National Nature Reserve" longLatPrecision="1" longitude="104.0216" pageId="6" pageNumber="577" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="8695EA80FFC9E2560D08FE14F1FFFE30" bold="true" box="[336,445,410,436]" pageId="6" pageNumber="577">
<typeStatus id="6B5A8830FFC9E2560D08FE14F1FFFE30" box="[336,445,410,436]" pageId="6" pageNumber="577" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="A2E7FD1BFFC9E2560D9DFE14F198FE30" box="[453,474,410,436]" pageId="6" pageNumber="577" type="male">♂</specimenCount>
(SWUC-T-LY-08-01),
<collectingCountry id="CCF67602FFC9E2560EB8FE14F364FE30" box="[736,806,410,436]" name="China" pageId="6" pageNumber="577">China</collectingCountry>
,
<emphasis id="8695EA80FFC9E2560F69FE14F3B5FE30" box="[817,1015,410,436]" italics="true" pageId="6" pageNumber="577">
<collectingRegion id="7625F870FFC9E2560F69FE14F3B5FE30" box="[817,1015,410,436]" country="China" name="Sichuan" pageId="6" pageNumber="577">Sichuan Province</collectingRegion>
</emphasis>
,
<collectingCounty id="5D3F4E1EFFC9E256085EFE14F4FAFE30" box="[1030,1208,410,436]" pageId="6" pageNumber="577">Pingwu County</collectingCounty>
,
<location id="B13E6049FFC9E256089BFE14F105FE5C" LSID="urn:lsid:plazi:treatment:3C488784FFC9E2570CCFFEA0F0BDFDE5:B13E6049FFC9E256089BFE14F105FE5C" country="China" county="Pingwu County" latitude="33.003418" longLatPrecision="1" longitude="104.0216" name="Wanglang National Nature Reserve" pageId="6" pageNumber="577" stateProvince="Sichuan">Wanglang National Nature Reserve</location>
,
<geoCoordinate id="D1D55055FFC9E2560D0AFE31F1A1FE5E" box="[338,483,447,474]" degrees="33" direction="north" minutes="00.205" orientation="latitude" pageId="6" pageNumber="577" precision="1" value="33.003418">33°00.205′N</geoCoordinate>
,
<geoCoordinate id="D1D55055FFC9E2560DB7FE31F2C9FE5E" box="[495,651,447,474]" degrees="104" direction="east" minutes="01.296" orientation="longitude" pageId="6" pageNumber="577" precision="1" value="104.0216">104°01.296′E</geoCoordinate>
,
<quantity id="73199B77FFC9E2560ECEFE30F2AFFE5C" box="[662,749,446,472]" metricMagnitude="3" metricUnit="m" metricValue="2.949" pageId="6" pageNumber="577" unit="m" value="2949.0">
<elevation id="3FCCD1A1FFC9E2560ECEFE30F2AFFE5C" box="[662,749,446,472]" metricMagnitude="3" metricUnit="m" metricValue="2.949" pageId="6" pageNumber="577" unit="m" value="2949.0">2949 m</elevation>
</quantity>
,
<date id="C05F1052FFC9E2560EA0FE30F337FE5C" box="[760,885,446,472]" pageId="6" pageNumber="577" value="2013-05-22">
<collectingDate id="D01BE9BAFFC9E2560EA0FE30F337FE5C" box="[760,885,446,472]" pageId="6" pageNumber="577" value="2013-05-22">22.V. 2013</collectingDate>
</date>
,
<collectorName id="19145344FFC9E2560F27FE30F3A0FE5C" box="[895,994,446,472]" pageId="6" pageNumber="577">D. Wang</collectorName>
&amp;
<collectorName id="19145344FFC9E256085DFE30F43FFE5C" box="[1029,1149,446,472]" pageId="6" pageNumber="577">X.K. Jiang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="04893CCFFFC9E25608ECFE30F2B0FE78" ID-GBIF-Occurrence="3107827318" collectingDate="2013-05-22" collectorName="D. Wang &amp; X. K. Jiang" country="China" county="Pingwu County" elevation="2949" latitude="33.003418" location="Wanglang National Nature Reserve" longLatPrecision="1" longitude="104.0216" pageId="6" pageNumber="577" specimenCount="47" specimenCount-female="30" specimenCount-male="17" stateProvince="Sichuan" typeStatus="paratype">
<emphasis id="8695EA80FFC9E25608ECFE30F577FE5C" bold="true" box="[1204,1333,446,472]" pageId="6" pageNumber="577">
<typeStatus id="6B5A8830FFC9E25608ECFE30F572FE5C" box="[1204,1328,446,472]" pageId="6" pageNumber="577" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="A2E7FD1BFFC9E2560964FE30F52FFE5C" box="[1340,1389,446,472]" pageId="6" pageNumber="577" type="male">17♂</specimenCount>
and
<specimenCount id="A2E7FD1BFFC9E2560CCFFE6CF08AFE78" box="[151,200,482,508]" pageId="6" pageNumber="577" type="female">30♀</specimenCount>
(SWUC-T-LY-08-02~48), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FCFB6519FFC9E2560C9FFD88F2E4FDC0" pageId="6" pageNumber="577" type="etymology">
<paragraph id="B45E3692FFC9E2560C9FFD88F2E4FDC0" blockId="6.[151,1436,409,689]" pageId="6" pageNumber="577">
<emphasis id="8695EA80FFC9E2560C9FFD88F10CFDA4" bold="true" box="[199,334,518,544]" pageId="6" pageNumber="577">Etymology.</emphasis>
The specific name is an adjective from the Latin word ‘
<emphasis id="8695EA80FFC9E2560FEFFD88F45DFDA4" box="[951,1055,518,544]" italics="true" pageId="6" pageNumber="577">hamatus’</emphasis>
, meaning ‘hamate’ or ‘hook-like’, referring to the shape of the median apophysis.
</paragraph>
</subSubSection>
<subSubSection id="FCFB6519FFC9E2560C9FFDC3F2F9FD35" pageId="6" pageNumber="577" type="diagnosis">
<paragraph id="B45E3692FFC9E2560C9FFDC3F2F9FD35" blockId="6.[151,1436,409,689]" pageId="6" pageNumber="577">
<emphasis id="8695EA80FFC9E2560C9FFDC3F100FDE3" bold="true" box="[199,322,589,615]" pageId="6" pageNumber="577">Diagnosis.</emphasis>
The species can be easily distinguished from other
<taxonomicName id="73E14D11FFC9E2560FFCFDC0F410FDEC" box="[932,1106,590,616]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8695EA80FFC9E2560FFCFDC0F3B4FDEC" box="[932,1014,590,616]" italics="true" pageId="6" pageNumber="577">Artoria</emphasis>
species
</taxonomicName>
by the hook-shaped median apophysis and the arched embolus, with bifurcated end (
<figureCitation id="2CDA2A17FFC9E2560F5FFDFCF3C7FD09" box="[775,901,626,653]" captionStart="FIGURE 5" captionStartId="6.[151,250,1511,1535]" captionTargetBox="[198,1391,718,1482]" captionTargetId="figure-147@6.[190,1397,712,1488]" captionTargetPageId="6" captionText="FIGURE 5. Artoria hamata sp. nov., holotype male (A–B), paratype male (C) and paratype female (D–E). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C.Apical part of bulb, ventral view; D. Epigyne, ventral view; E. Female internal genitalia, dorsal view. BEA, basoembolic apophysis; CO, copulatory opening; Em, embolus; FD, fertilization duct; HS, head of spermathecal; MA, median apophysis; SS, stalk of spermathecal; TA, terminal apophysis (= synembolus sensu Zyuzin 1993); Te, tegulum." figureDoi="http://doi.org/10.5281/zenodo.4714854" httpUri="https://zenodo.org/record/4714854/files/figure.png" pageId="6" pageNumber="577">Figs 5A–C</figureCitation>
,
<figureCitation id="2CDA2A17FFC9E2560FD7FDFCF395FD09" box="[911,983,626,653]" captionStart="FIGURE 6" captionStartId="7.[151,250,1637,1661]" captionTargetBox="[151,1435,633,1613]" captionTargetId="figure-193@7.[151,1435,633,1613]" captionTargetPageId="7" captionText="FIGURE 6. Artoria hamata sp. nov., holotype male (A, E–F), paratype male (C–D, G) and paratype female (B, H–J). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male pedipalp, bulbus, ventral view; D. Same, retrolateral view; E. Left male pedipalp, ventral view; F. Same, retrolateral view; G. Apical part of bulb, ventral view; H–I. Epigyne, ventral view; J. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4714860" httpUri="https://zenodo.org/record/4714860/files/figure.png" pageId="6" pageNumber="577">6C–G</figureCitation>
). Females can be recognized by the wide copulatory openings of epigyne (
<figureCitation id="2CDA2A17FFC9E2560E50FD18F227FD35" box="[520,613,662,689]" captionStart="FIGURE 5" captionStartId="6.[151,250,1511,1535]" captionTargetBox="[198,1391,718,1482]" captionTargetId="figure-147@6.[190,1397,712,1488]" captionTargetPageId="6" captionText="FIGURE 5. Artoria hamata sp. nov., holotype male (A–B), paratype male (C) and paratype female (D–E). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C.Apical part of bulb, ventral view; D. Epigyne, ventral view; E. Female internal genitalia, dorsal view. BEA, basoembolic apophysis; CO, copulatory opening; Em, embolus; FD, fertilization duct; HS, head of spermathecal; MA, median apophysis; SS, stalk of spermathecal; TA, terminal apophysis (= synembolus sensu Zyuzin 1993); Te, tegulum." figureDoi="http://doi.org/10.5281/zenodo.4714854" httpUri="https://zenodo.org/record/4714854/files/figure.png" pageId="6" pageNumber="577">Figs 5D</figureCitation>
,
<figureCitation id="2CDA2A17FFC9E2560E29FD18F2EDFD35" box="[625,687,662,689]" captionStart="FIGURE 6" captionStartId="7.[151,250,1637,1661]" captionTargetBox="[151,1435,633,1613]" captionTargetId="figure-193@7.[151,1435,633,1613]" captionTargetPageId="7" captionText="FIGURE 6. Artoria hamata sp. nov., holotype male (A, E–F), paratype male (C–D, G) and paratype female (B, H–J). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male pedipalp, bulbus, ventral view; D. Same, retrolateral view; E. Left male pedipalp, ventral view; F. Same, retrolateral view; G. Apical part of bulb, ventral view; H–I. Epigyne, ventral view; J. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4714860" httpUri="https://zenodo.org/record/4714860/files/figure.png" pageId="6" pageNumber="577">6H–I</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="E09E661AFFC9E2560CCFFA69F155F914" ID-DOI="http://doi.org/10.5281/zenodo.4714854" ID-Zenodo-Dep="4714854" httpUri="https://zenodo.org/record/4714854/files/figure.png" pageId="6" pageNumber="577" startId="6.[151,250,1511,1535]" targetBox="[198,1391,718,1482]" targetPageId="6">
<paragraph id="B45E3692FFC9E2560CCFFA69F155F914" blockId="6.[151,1436,1511,1680]" pageId="6" pageNumber="577">
<emphasis id="8695EA80FFC9E2560CCFFA69F151F984" bold="true" box="[151,275,1511,1536]" pageId="6" pageNumber="577">FIGURE 5.</emphasis>
<taxonomicName id="73E14D11FFC9E2560D42FA66F1F5F984" authority="Wang &amp; Framenau &amp; Zhang, 2021" authorityName="Wang &amp; Framenau &amp; Zhang" authorityYear="2021" box="[282,439,1512,1536]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="hamata" status="sp. nov.">
<emphasis id="8695EA80FFC9E2560D42FA66F1F5F984" box="[282,439,1512,1536]" italics="true" pageId="6" pageNumber="577">Artoria hamata</emphasis>
</taxonomicName>
<emphasis id="8695EA80FFC9E2560DE6FA66F24DF984" bold="true" box="[446,527,1512,1536]" pageId="6" pageNumber="577">
<taxonomicNameLabel id="9DA657FBFFC9E2560DE6FA66F24DF984" box="[446,527,1512,1536]" pageId="6" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (A
<emphasis id="8695EA80FFC9E2560E97FA66F29EF984" bold="true" box="[719,732,1512,1536]" pageId="6" pageNumber="577">–</emphasis>
B), paratype male (C) and paratype female (D
<emphasis id="8695EA80FFC9E25608E8FA66F4FFF984" bold="true" box="[1200,1213,1512,1536]" pageId="6" pageNumber="577">–</emphasis>
E). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C. Apical part of bulb, ventral view; D. Epigyne, ventral view; E. Female internal genitalia, dorsal view. BEA, basoembolic apophysis; CO, copulatory opening; Em, embolus; FD, fertilization duct; HS, head of spermathecal; MA, median apophysis; SS, stalk of spermathecal; TA, terminal apophysis (= synembolus
<emphasis id="8695EA80FFC9E2560894F9DAF546F9EF" box="[1228,1284,1620,1643]" italics="true" pageId="6" pageNumber="577">sensu</emphasis>
<bibRefCitation id="D0704B63FFC9E2560953F9DAF5D2F9E8" author="Zyuzin, A. A." box="[1291,1424,1620,1644]" pageId="6" pageNumber="577" pagination="693 - 700" refId="ref7241" refString="Zyuzin, A. A. (1993) Studies on the wolf spiders (Araneae: Lycosidae). I. A new genus and species from Kazakhstan, with comments on the Lycosinae. Memoirs of the Queensland Museum, 33, 693 - 700." type="journal article" year="1993">Zyuzin 1993</bibRefCitation>
); Te, tegulum.
</paragraph>
</caption>
<subSubSection id="FCFB6519FFC9E2570C9FF933F143FD9D" lastPageId="7" lastPageNumber="578" pageId="6" pageNumber="577" type="description">
<paragraph id="B45E3692FFC9E2560C9FF933F3BBF87C" blockId="6.[151,1437,1725,2040]" pageId="6" pageNumber="577">
<emphasis id="8695EA80FFC9E2560C9FF933F11BF953" bold="true" box="[199,345,1725,1751]" pageId="6" pageNumber="577">Description.</emphasis>
Male. Total length 5.20–5.90.
<typeStatus id="6B5A8830FFC9E2560E9BF930F369F95C" box="[707,811,1726,1752]" pageId="6" pageNumber="577" type="holotype">Holotype</typeStatus>
(
<figureCitation id="2CDA2A17FFC9E2560F66F930F3DBF95D" box="[830,921,1726,1753]" captionStart="FIGURE 6" captionStartId="7.[151,250,1637,1661]" captionTargetBox="[151,1435,633,1613]" captionTargetId="figure-193@7.[151,1435,633,1613]" captionTargetPageId="7" captionText="FIGURE 6. Artoria hamata sp. nov., holotype male (A, E–F), paratype male (C–D, G) and paratype female (B, H–J). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male pedipalp, bulbus, ventral view; D. Same, retrolateral view; E. Left male pedipalp, ventral view; F. Same, retrolateral view; G. Apical part of bulb, ventral view; H–I. Epigyne, ventral view; J. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4714860" httpUri="https://zenodo.org/record/4714860/files/figure.png" pageId="6" pageNumber="577">Fig. 6A</figureCitation>
) total length 5.48. Carapace 2.97 long, 2.12 wide; opisthosoma 2.57 long, 1.99 wide. Carapace brown. Eye region black. Radial furrows distinct. Eye sizes and interdistances: AME 0.09, ALE 0.13, PME 0.32, PLE 0.27; AME–AME 0.08, AME–ALE 0.07, PME–PME 0.25, PME–PLE 0.30. Clypeus 0.13 high. Chelicerae brown, with three teeth on both margins. Labium yellowish brown, with dark base, longer than wide. Endites yellowish brown, longer than wide. Sternum dark brown and scutellate, with sparse brown setae. Legs yellowish brown, with dark brown pigmentation. Leg measurements: I 6.70 (1.84, 2.33, 1.50, 1.03); II 6.40 (1.79, 2.15, 1.47, 0.99); III 6.12 (1.72, 1.92, 1.55, 0.93); IV 8.60 (2.22, 2.63, 2.46, 1.29). Abdomen yellowish brown, dorsally with lanceolate cardiac mark in anterior half followed by irregular black markings extending posteriorly. Venter of abdomen brown, with brown spinnerets.
</paragraph>
<paragraph id="B45E3692FFC8E2570C9FFF19F428FE99" blockId="7.[151,1437,150,609]" pageId="7" pageNumber="578">
Pedipalp (
<figureCitation id="2CDA2A17FFC8E2570D63FF19F1FEFF35" box="[315,444,150,177]" captionStart="FIGURE 5" captionStartId="6.[151,250,1511,1535]" captionTargetBox="[198,1391,718,1482]" captionTargetId="figure-147@6.[190,1397,712,1488]" captionTargetPageId="6" captionText="FIGURE 5. Artoria hamata sp. nov., holotype male (A–B), paratype male (C) and paratype female (D–E). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C.Apical part of bulb, ventral view; D. Epigyne, ventral view; E. Female internal genitalia, dorsal view. BEA, basoembolic apophysis; CO, copulatory opening; Em, embolus; FD, fertilization duct; HS, head of spermathecal; MA, median apophysis; SS, stalk of spermathecal; TA, terminal apophysis (= synembolus sensu Zyuzin 1993); Te, tegulum." figureDoi="http://doi.org/10.5281/zenodo.4714854" httpUri="https://zenodo.org/record/4714854/files/figure.png" pageId="7" pageNumber="578">Figs 5A–C</figureCitation>
,
<figureCitation id="2CDA2A17FFC8E2570D93FF19F250FF36" box="[459,530,151,178]" captionStart="FIGURE 6" captionStartId="7.[151,250,1637,1661]" captionTargetBox="[151,1435,633,1613]" captionTargetId="figure-193@7.[151,1435,633,1613]" captionTargetPageId="7" captionText="FIGURE 6. Artoria hamata sp. nov., holotype male (A, E–F), paratype male (C–D, G) and paratype female (B, H–J). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male pedipalp, bulbus, ventral view; D. Same, retrolateral view; E. Left male pedipalp, ventral view; F. Same, retrolateral view; G. Apical part of bulb, ventral view; H–I. Epigyne, ventral view; J. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4714860" httpUri="https://zenodo.org/record/4714860/files/figure.png" pageId="7" pageNumber="578">6C–G</figureCitation>
). Median apophysis sclerotized, hook-shaped anteriorly, longer than wide, and with pointed end. Basoembolic apophysis highly sclerotized, slightly curved with blunt end. Terminal apophysis with two arms, one of them needle-like as the functional conductor (TA1); the other one sclerotized, relatively broad at base, curved distally (TA2). Embolus longer than wide, arched, distal end bifurcated.
</paragraph>
<paragraph id="B45E3692FFC8E2570C9FFEA9F48CFE29" blockId="7.[151,1437,150,609]" pageId="7" pageNumber="578">
Female (
<figureCitation id="2CDA2A17FFC8E2570D70FEA9F13DFEC6" box="[296,383,295,322]" captionStart="FIGURE 6" captionStartId="7.[151,250,1637,1661]" captionTargetBox="[151,1435,633,1613]" captionTargetId="figure-193@7.[151,1435,633,1613]" captionTargetPageId="7" captionText="FIGURE 6. Artoria hamata sp. nov., holotype male (A, E–F), paratype male (C–D, G) and paratype female (B, H–J). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male pedipalp, bulbus, ventral view; D. Same, retrolateral view; E. Left male pedipalp, ventral view; F. Same, retrolateral view; G. Apical part of bulb, ventral view; H–I. Epigyne, ventral view; J. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4714860" httpUri="https://zenodo.org/record/4714860/files/figure.png" pageId="7" pageNumber="578">Fig. 6B</figureCitation>
). Total length: 7.85. Carapace 3.34 long, 2.35 wide; opisthosoma 4.26 long, 3.31 wide. Eye sizes and interdistances: AME 0.09, ALE 0.12, PME 0.35, PLE 0.27; AME–AME 0.12, AME–ALE 0.12, PME–PME 0.28, PME–PLE 0.36. Clypeus 0.17 high. Leg measurements: I 7.03 (2.05, 2.48, 1.51, 0.99); II 6.74 (1.96, 2.35, 1.46, 0.97); III 6.72 (1.82, 2.07, 1.73, 1.10); IV 9.37 (2.42, 2.83, 2.72, 1.40). Leg formula: 4123.
</paragraph>
<paragraph id="B45E3692FFC8E2570C9FFE39F143FD9D" blockId="7.[151,1437,150,609]" pageId="7" pageNumber="578">
Female genitalia (
<figureCitation id="2CDA2A17FFC8E2570DD7FE39F24EFE55" box="[399,524,438,465]" captionStart="FIGURE 5" captionStartId="6.[151,250,1511,1535]" captionTargetBox="[198,1391,718,1482]" captionTargetId="figure-147@6.[190,1397,712,1488]" captionTargetPageId="6" captionText="FIGURE 5. Artoria hamata sp. nov., holotype male (A–B), paratype male (C) and paratype female (D–E). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C.Apical part of bulb, ventral view; D. Epigyne, ventral view; E. Female internal genitalia, dorsal view. BEA, basoembolic apophysis; CO, copulatory opening; Em, embolus; FD, fertilization duct; HS, head of spermathecal; MA, median apophysis; SS, stalk of spermathecal; TA, terminal apophysis (= synembolus sensu Zyuzin 1993); Te, tegulum." figureDoi="http://doi.org/10.5281/zenodo.4714854" httpUri="https://zenodo.org/record/4714854/files/figure.png" pageId="7" pageNumber="578">Figs 5D–E</figureCitation>
,
<figureCitation id="2CDA2A17FFC8E2570E4FFE39F215FE56" box="[535,599,439,466]" captionStart="FIGURE 6" captionStartId="7.[151,250,1637,1661]" captionTargetBox="[151,1435,633,1613]" captionTargetId="figure-193@7.[151,1435,633,1613]" captionTargetPageId="7" captionText="FIGURE 6. Artoria hamata sp. nov., holotype male (A, E–F), paratype male (C–D, G) and paratype female (B, H–J). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male pedipalp, bulbus, ventral view; D. Same, retrolateral view; E. Left male pedipalp, ventral view; F. Same, retrolateral view; G. Apical part of bulb, ventral view; H–I. Epigyne, ventral view; J. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4714860" httpUri="https://zenodo.org/record/4714860/files/figure.png" pageId="7" pageNumber="578">6H–J</figureCitation>
). Epigyne septum wide, oval. Copulatory openings wide. Copulatory ducts thick and long. Head of spermathecal small and club-like, extending forward. Fertilization ducts wide and slightly distorted.
</paragraph>
</subSubSection>
<subSubSection id="FCFB6519FFC8E2570C9FFDACF0BDFDE5" pageId="7" pageNumber="578" type="distribution">
<paragraph id="B45E3692FFC8E2570C9FFDACF0BDFDE5" blockId="7.[151,1437,150,609]" pageId="7" pageNumber="578">
<emphasis id="8695EA80FFC8E2570C9FFDACF123FDB8" bold="true" box="[199,353,546,572]" pageId="7" pageNumber="578">Distribution.</emphasis>
Currently known only from the
<typeStatus id="6B5A8830FFC8E2570E9DFDADF2B7FDB9" box="[709,757,547,573]" pageId="7" pageNumber="578">type</typeStatus>
locality, Wanglang Nature Reserve,
<collectingRegion id="7625F870FFC8E25708D1FDADF50DFDB9" box="[1161,1359,547,573]" country="China" name="Sichuan" pageId="7" pageNumber="578">Sichuan Province</collectingRegion>
,
<collectingCountry id="CCF67602FFC8E2570901FDADF5DEFDB9" box="[1369,1436,547,573]" name="China" pageId="7" pageNumber="578">China</collectingCountry>
(
<figureCitation id="2CDA2A17FFC8E2570CF8FDC9F0B3FDE5" box="[160,241,583,609]" captionStart="FIGURE 11" captionStartId="12.[151,250,1665,1689]" captionTargetBox="[151,1435,181,1641]" captionTargetId="figure-23@12.[151,1435,181,1641]" captionTargetPageId="12" captionText="FIGURE 11. Distribution records of the wolf spider subfamily Artoriinae in China." figureDoi="http://doi.org/10.5281/zenodo.4714931" httpUri="https://zenodo.org/record/4714931/files/figure.png" pageId="7" pageNumber="578">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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