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<document id="6688AD558936B582410676109F968E7D" ID-CLB-Dataset="32199" ID-DOI="10.5852/ejt.2017.261" ID-GBIF-Dataset="708e152e-4450-4439-84a9-3c56148cd082" ID-Zenodo-Dep="885318" ID-ZooBank="C59239D1-0BF0-446A-AF6E-71234D01B23D" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1504621962129" checkinUser="plazi" docAuthor="Reboleira, Ana Sofa P. S., Fresneda, Javier &amp; Salgado, José Maria" docDate="2017" docId="03FB5E336669AB5DB75061A2FC11393F" docLanguage="en" docName="ejt-261.pdf.imf" docOrigin="European Journal of Taxonomy 261" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Speonemadus breuili Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado 2017" docType="treatment" docVersion="18" lastPageNumber="15" masterDocId="FFC2264B6665AB53B56C656DFFD63B4C" masterDocTitle="A new species of Speonemadus from Portugal, with the revision of the escalerai-group (Coleoptera, Leiodidae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="13" updateTime="1734103965070" updateUser="ExternalLinkService">
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<mods:title id="EFA5DAAEAFB966BB4A29AD63CF17D9FA">A new species of Speonemadus from Portugal, with the revision of the escalerai-group (Coleoptera, Leiodidae)</mods:title>
</mods:titleInfo>
<mods:name id="A5BF3BBE3FA104F066E21AAF5E2E1B1C" type="personal">
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</mods:role>
<mods:namePart id="AB1726F27E98BC7F479BB9DF087BB91C">Reboleira, Ana Sofa P. S.</mods:namePart>
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<mods:namePart id="A752414962827073CF93718F3ED5FBF8">Fresneda, Javier</mods:namePart>
</mods:name>
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<mods:role id="0AEAE5F12CB13A9C836F8B4DBB24A535">
<mods:roleTerm id="4B8D165AC50400165D1D9C9BD5E0FEDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AE077426472C3C77E19F23B1D8EB2BF0">Salgado, José Maria</mods:namePart>
</mods:name>
<mods:typeOfResource id="32DD6BC7797BF4BF4D3CFFE2989C44D2">text</mods:typeOfResource>
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<mods:title id="A5D5CFCCE3E5B10FA18D1BFE2B89C5C8">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="6FF69E824F29E16C3AC416E125EF6F7A">
<mods:date id="6A8CE902627ACC9A7AC48AC097EF9AFA">2017</mods:date>
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<mods:number id="D8415E5CDFDCF180CA81CD575FE45D60">2017-01-19</mods:number>
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<mods:detail id="4E0CF1B30B9CBFB650B91FD268121880" type="volume">
<mods:number id="7A9E147497F82A8692D8931DA6C3349F">261</mods:number>
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<mods:start id="18A324DA3F6184E61304996C32FC266F">1</mods:start>
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<mods:classification id="ED34AE5069B4E8FED8B05F7198E6195F">journal article</mods:classification>
<mods:identifier id="3AAB5D725490C5B9F77B9C2C482C9121" type="CLB-Dataset">32199</mods:identifier>
<mods:identifier id="4C1C713D9A27BC510214B56C0DFF623B" type="DOI">10.5852/ejt.2017.261</mods:identifier>
<mods:identifier id="FEE1246BF1CA817C041C936DD7D61AAB" type="GBIF-Dataset">708e152e-4450-4439-84a9-3c56148cd082</mods:identifier>
<mods:identifier id="DC9EAD56CF3949E369BFEC77C58FA8FD" type="Zenodo-Dep">885318</mods:identifier>
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<treatment id="03FB5E336669AB5DB75061A2FC11393F" ID-DOI="http://doi.org/10.5281/zenodo.5689572" ID-GBIF-Taxon="133183539" ID-Zenodo-Dep="5689572" LSID="urn:lsid:plazi:treatment:03FB5E336669AB5DB75061A2FC11393F" httpUri="http://treatment.plazi.org/id/03FB5E336669AB5DB75061A2FC11393F" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" scope_family="Leiodidae" scope_order="Coleoptera" scope_subFamily="Cholevinae" scope_tribe="Anemadini">
<subSubSection id="C348BCAE6669AB5FB75061A2FC213FA6" box="[572,1015,1231,1258]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BEDEF256669AB5FB75061A2FC213FA6" blockId="12.[572,1015,1231,1258]" box="[572,1015,1231,1258]" pageId="12" pageNumber="13">
<heading id="D0A558496669AB5FB75061A2FC213FA6" box="[572,1015,1231,1258]" centered="true" fontSize="11" level="3" pageId="12" pageNumber="13" reason="4">
<treatmentCitation id="0AF3C9346669AB5FB75061A2FC213FA6" author="Jeannel" authority="(Jeannel, 1922)" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[572,1015,1231,1258]" class="Insecta" family="Leiodidae" firstAuthor="Jeannel" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" pagination="1 - 98" phylum="Arthropoda" rank="species" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." species="breuili" treatmentTaxon="Speonemadus breuili" type="journal article" year="1922">
<taxonomicName id="4C5294A66669AB5FB75061A2FC213FA6" ID-CoL="4YHYX" authority="(Jeannel, 1922)" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[572,1015,1231,1258]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B92633376669AB5FB75061A2FCED3FA6" bold="true" box="[572,827,1231,1258]" italics="true" pageId="12" pageNumber="13">Speonemadus breuili</emphasis>
(
<bibRefCitation id="EFC392D46669AB5FB62761BDFC393FA6" author="Jeannel" box="[843,1007,1232,1258]" firstAuthor="Jeannel" pageId="12" pageNumber="13" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel, 1922</bibRefCitation>
)
</taxonomicName>
</treatmentCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE6669AB5FB5D1607CFC313EFF" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="8BEDEF256669AB5FB5D1607CFCF33E67" blockId="12.[189,805,1297,1324]" box="[189,805,1297,1324]" pageId="12" pageNumber="13">
<treatmentCitation id="0AF3C9346669AB5FB5D1607CFCF73E67" author="Jeannel" authority="Jeannel, 1922: 60" authorityName="Jeannel" authorityPageNumber="60" authorityYear="1922" box="[189,801,1297,1324]" class="Insecta" family="Leiodidae" firstAuthor="Jeannel" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" pagination="1 - 98" phylum="Arthropoda" rank="species" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." species="breuili" subGenus="Speonemadus" treatmentTaxon="Anemadus (Speonemadus) breuili" type="journal article" year="1922">
<taxonomicName id="4C5294A66669AB5FB5D1607CFCF73E67" ID-CoL="DWR5" authority="Jeannel, 1922: 60" authorityName="Jeannel" authorityPageNumber="60" authorityYear="1922" box="[189,801,1297,1324]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="breuili" subGenus="Speonemadus">
<emphasis id="B92633376669AB5FB5D1607CFEE13E67" box="[189,311,1297,1323]" italics="true" pageId="12" pageNumber="13">Anemadus</emphasis>
(
<emphasis id="B92633376669AB5FB424607CFE3F3E67" box="[328,489,1297,1323]" italics="true" pageId="12" pageNumber="13">Speonemadus</emphasis>
)
<emphasis id="B92633376669AB5FB497607CFD9E3E67" box="[507,584,1297,1323]" italics="true" pageId="12" pageNumber="13">breuili</emphasis>
<bibRefCitation id="EFC392D46669AB5FB723607CFCF73E67" author="Jeannel" box="[591,801,1297,1323]" firstAuthor="Jeannel" pageId="12" pageNumber="13" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel, 1922: 60</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D1603EFCED3E21" blockId="12.[189,999,1363,1460]" box="[189,827,1363,1389]" pageId="12" pageNumber="13">
<taxonomicName id="4C5294A66669AB5FB5D1603EFD863E21" authority="Jeannel 1922: 61" box="[189,592,1363,1389]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="verneri" subGenus="Speonemadus">
<emphasis id="B92633376669AB5FB5D1603EFEE13E21" box="[189,311,1363,1389]" italics="true" pageId="12" pageNumber="13">Anemadus</emphasis>
(
<emphasis id="B92633376669AB5FB424603EFE3F3E21" box="[328,489,1363,1389]" italics="true" pageId="12" pageNumber="13">Speonemadus</emphasis>
)
<emphasis id="B92633376669AB5FB4976039FD863E21" box="[507,592,1364,1389]" italics="true" pageId="12" pageNumber="13">verneri</emphasis>
</taxonomicName>
–
<bibRefCitation id="EFC392D46669AB5FB701603EFCE03E21" author="Jeannel" box="[621,822,1363,1389]" firstAuthor="Jeannel" pageId="12" pageNumber="13" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel 1922: 61</bibRefCitation>
.
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D1601BFC093EDC" blockId="12.[189,999,1363,1460]" box="[189,991,1398,1424]" pageId="12" pageNumber="13">
<taxonomicName id="4C5294A66669AB5FB5D1601BFD333EDC" ID-CoL="DWR5" authority="Jeannel, 1922" box="[189,741,1398,1424]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="angusticollis" subGenus="Speonemadus" subSpecies="breuili">
<emphasis id="B92633376669AB5FB5D1601BFEE13EDC" box="[189,311,1398,1424]" italics="true" pageId="12" pageNumber="13">Anemadus</emphasis>
(
<emphasis id="B92633376669AB5FB424601BFE3F3EDC" box="[328,489,1398,1424]" italics="true" pageId="12" pageNumber="13">Speonemadus</emphasis>
)
<emphasis id="B92633376669AB5FB497601BFD333EDC" box="[507,741,1398,1424]" italics="true" pageId="12" pageNumber="13">angusticollis breuili</emphasis>
</taxonomicName>
–
<bibRefCitation id="EFC392D46669AB5FB66E601BFC0D3EDC" author="Jeannel" box="[770,987,1398,1424]" firstAuthor="Jeannel" pageId="12" pageNumber="13" pagination="1" refId="ref12306" refString="Jeannel R. 1936. Monographie des Catopides. Memoires du Museum national d'Histoire naturelle 1 (1). Museum national d'Histoire naturelle, Paris." type="journal article" year="1936">Jeannel 1936: 221</bibRefCitation>
.
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D160F4FC313EFF" blockId="12.[189,999,1363,1460]" box="[189,999,1433,1460]" pageId="12" pageNumber="13">
<taxonomicName id="4C5294A66669AB5FB5D160F4FD3B3EFF" authority="(Jeannel 1922: 61)" box="[189,749,1433,1460]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="angusticollis" subGenus="Speonemadus" subSpecies="verneri">
<emphasis id="B92633376669AB5FB5D160F4FEE13EFF" box="[189,311,1433,1459]" italics="true" pageId="12" pageNumber="13">Anemadus</emphasis>
(
<emphasis id="B92633376669AB5FB42460F4FE3F3EFF" box="[328,489,1433,1459]" italics="true" pageId="12" pageNumber="13">Speonemadus</emphasis>
)
<emphasis id="B92633376669AB5FB49760F4FD3B3EFF" box="[507,749,1433,1459]" italics="true" pageId="12" pageNumber="13">angusticollis verneri</emphasis>
</taxonomicName>
–
<bibRefCitation id="EFC392D46669AB5FB66660F4FC343EFF" author="Jeannel" box="[778,994,1433,1459]" firstAuthor="Jeannel" pageId="12" pageNumber="13" pagination="1" refId="ref12306" refString="Jeannel R. 1936. Monographie des Catopides. Memoires du Museum national d'Histoire naturelle 1 (1). Museum national d'Histoire naturelle, Paris." type="journal article" year="1936">Jeannel 1936: 221</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE6669AB5EB5D160B6FDEE3922" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BEDEF256669AB5FB5D160B6FE8B3EB8" blockId="12.[189,1140,1498,1572]" box="[189,349,1498,1525]" pageId="12" pageNumber="13">
<heading id="D0A558496669AB5FB5D160B6FE8B3EB8" bold="true" box="[189,349,1498,1525]" fontSize="11" level="2" pageId="12" pageNumber="13" reason="6">
<emphasis id="B92633376669AB5FB5D160B6FE8B3EB8" bold="true" box="[189,349,1498,1525]" pageId="12" pageNumber="13">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D16364FBA23D6F" blockId="12.[189,1140,1498,1572]" box="[189,1140,1545,1572]" pageId="12" pageNumber="13">
“Cueva de las Motillas, Jerez de la Frontera, provincia de Cádiz” (
<bibRefCitation id="EFC392D46669AB5FB6AB6364FBB13D6F" author="Jeannel" box="[967,1127,1545,1571]" firstAuthor="Jeannel" pageId="12" pageNumber="13" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel 1922</bibRefCitation>
).
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D16327FE7C3D28" blockId="12.[189,426,1610,1636]" box="[189,426,1610,1636]" pageId="12" pageNumber="13">
<heading id="D0A558496669AB5FB5D16327FE7C3D28" bold="true" box="[189,426,1610,1636]" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<emphasis id="B92633376669AB5FB5D16327FE7C3D28" bold="true" box="[189,426,1610,1636]" pageId="12" pageNumber="13">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BEDEF256669AB5FB5D16314FADE3CC6" blockId="12.[189,1399,1655,1930]" pageId="12" pageNumber="13">
<collectingCountry id="F345AFB56669AB5FB5D16314FEC23DDF" box="[189,276,1657,1683]" name="Spain" pageId="12" pageNumber="13">SPAIN</collectingCountry>
:
<collectorName id="26A78AF36669AB5FB44D6314FEBC3DDF" box="[289,362,1657,1683]" pageId="12" pageNumber="13">Cádiz</collectorName>
: 3 ♂♂,
<specimenCount id="9D5424AC6669AB5FB4A16314FDC53DD8" box="[461,531,1657,1684]" pageId="12" pageNumber="13" type="female">2 ♀♀</specimenCount>
,
<collectorName id="26A78AF36669AB5FB74F6314FD7C3DDF" box="[547,682,1657,1683]" pageId="12" pageNumber="13">Villaluenga</collectorName>
del Rosario, Cueva Aljíbez (=
<collectorName id="26A78AF36669AB5FB1496314FBA63DDF" box="[1061,1136,1657,1683]" pageId="12" pageNumber="13">Cueva</collectorName>
del
<collectorName id="26A78AF36669AB5FB1CA6314FB243DDF" box="[1190,1266,1657,1683]" pageId="12" pageNumber="13">Aljíbe</collectorName>
,
<date id="FFECC9E56669AB5FB06D6314FAA73DDF" box="[1281,1393,1657,1683]" pageId="12" pageNumber="13" value="1970-07">
<collectingDate id="EFA8300D6669AB5FB06D6314FAA73DDF" box="[1281,1393,1657,1683]" pageId="12" pageNumber="13" value="1970-07">Jul. 1970</collectingDate>
</date>
,
<collectorName id="26A78AF36669AB5FB5D163F1FEDD3DFA" box="[189,267,1692,1718]" pageId="12" pageNumber="13">Ribera</collectorName>
&amp;
<collectorName id="26A78AF36669AB5FB44163F1FEA63DFA" box="[301,368,1692,1718]" pageId="12" pageNumber="13">Viñas</collectorName>
leg. (
<collectionCode id="ED4377E06669AB5FB4C363F1FDDA3DFA" box="[431,524,1692,1718]" pageId="12" pageNumber="13">CMCN</collectionCode>
). — Málaga: many ♂♂ and ♀♀, Cortés de la Frontera, Cueva del Berrueco,
<date id="FFECC9E56669AB5FB5D163ADFE8B3D96" box="[189,349,1728,1754]" pageId="12" pageNumber="13" value="2009-09-12">12 Sep. 2009</date>
and
<date id="FFECC9E56669AB5FB4F063D2FDFB3D96" box="[412,557,1727,1754]" pageId="12" pageNumber="13" value="2010-06-06">6 Jun. 2010</date>
, GIEX leg. (CFL, CPB, CJMS, CZULE); 15 ♂♂, 9 ♀♀ Cortés de la Frontera, Peñón del Berrueco, Cueva del Humo,
<date id="FFECC9E56669AB5FB662638EFC633DB1" box="[782,949,1763,1789]" pageId="12" pageNumber="13" value="2012-12-22">22 Dec. 2012</date>
, GIEX leg. (CFL, CJMS); many ♂♂ and ♀♀, Cortés de la Frontera, Complejo Motillas, Cueva Motilla fósil,
<date id="FFECC9E56669AB5FB160626BFB713C6C" box="[1036,1191,1798,1824]" pageId="12" pageNumber="13" value="2009-09-13">13 Sep. 2009</date>
and
<date id="FFECC9E56669AB5FB1B3626BFAA43C6C" box="[1247,1394,1798,1824]" pageId="12" pageNumber="13" value="2009-11-08">8 Nov. 2009</date>
, GIEX leg. (CFL; CPB; CJMS); many ♂♂ and ♀♀, Cortés de la Frontera, Complejo Motillas, Cueva Motilla Parralejo,
<date id="FFECC9E56669AB5FB4FF6221FDE63C2A" box="[403,560,1868,1894]" pageId="12" pageNumber="13" value="2009-09-13">13 Sep. 2009</date>
,
<date id="FFECC9E56669AB5FB7576221FD1D3C2A" box="[571,715,1868,1895]" pageId="12" pageNumber="13" value="2009-11-08">8 Nov. 2009</date>
and
<date id="FFECC9E56669AB5FB6686221FC593C2A" box="[772,911,1868,1895]" pageId="12" pageNumber="13" value="2010-06-05">5 Jun. 2010</date>
, GIEX leg. (CFL, CPB, CJMS, CZULE); 1 ♂, 2 ♀♀, Montejaque, El Pozuelo, Cueva Sima Nueva,
<date id="FFECC9E56669AB5FB60F621DFBD43CC5" box="[867,1026,1903,1930]" pageId="12" pageNumber="13" value="1986-03-29">29 Mar. 1986</date>
, Escolà leg. (CMCN).
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D16466FE593A69" blockId="13.[189,399,267,293]" box="[189,399,267,293]" pageId="13" pageNumber="14">
<heading id="D0A558496668AB5EB5D16466FE593A69" bold="true" box="[189,399,267,293]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="B92633376668AB5EB5D16466FE593A69" bold="true" box="[189,399,267,293]" pageId="13" pageNumber="14">Previous records</emphasis>
</heading>
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D16457FDEE3922" blockId="13.[189,1399,313,622]" pageId="13" pageNumber="14">
<collectingCountry id="F345AFB56668AB5EB5D16457FEC23A18" box="[189,276,314,340]" name="Spain" pageId="13" pageNumber="14">SPAIN</collectingCountry>
: Cádiz, Algeciras (
<bibRefCitation id="EFC392D46668AB5EB4946454FCF03A18" author="Giachino P. M. &amp; Vailati D." box="[504,806,313,340]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="13" pageNumber="14" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
); Algeciras, Los Barrios (
<bibRefCitation id="EFC392D46668AB5EB1316454FB0D3A18" author="Blas M." box="[1117,1243,313,340]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="13" pageNumber="14" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas 1989</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB1866454FE983A3B" author="Giachino P. M. &amp; Vailati D." firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="13" pageNumber="14" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
); Villaluenga del Rosario (
<bibRefCitation id="EFC392D46668AB5EB7EC6431FC763A3B" author="Giachino P. M. &amp; Vailati D." box="[640,928,348,375]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="13" pageNumber="14" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
); Villaluenga del Rosario, Cueva Aljíbez (= Cueva del Aljibe) (
<bibRefCitation id="EFC392D46668AB5EB4D364EDFDEC3AD6" author="Blas M." box="[447,570,384,410]" firstAuthor="Blas" journalOrPublisher="Congreso Nacional de Espeleologia" pageId="13" pageNumber="14" pagination="157 - 160" part="4" refId="ref11503" refString="Blas M. 1976. Coleopteros cavernicolas del distrito andaluz (Catopidae). Actas 4 Congreso Nacional de Espeleologia: 157 - 160." title="Coleopteros cavernicolas del distrito andaluz (Catopidae). Actas" type="journal article" year="1976">Blas 1976</bibRefCitation>
,
<bibRefCitation id="EFC392D46668AB5EB72A64EDFD553AD6" author="Blas M." box="[582,643,384,410]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="13" pageNumber="14" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">1977</bibRefCitation>
,
<bibRefCitation id="EFC392D46668AB5EB7FC64EDFD183AD6" author="Blas M." box="[656,718,384,410]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="13" pageNumber="14" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">1979</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB7B664EDFCBA3AD6" author="Tinaut" box="[730,876,384,410]" firstAuthor="Tinaut" pageId="13" pageNumber="14" pagination="3 - 28" refId="ref13778" refString="Tinaut A. 1998. Artropodos terrestres de las cavidades andaluzas. Zoologica Baetica 9: 3 - 28." type="journal article" year="1998">Tinaut 1998</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB61464EDFBD63AD7" author="Perez" box="[888,1024,384,411]" firstAuthor="Perez" pageId="13" pageNumber="14" pagination="1 - 97" refId="ref12561" refString="Perez T. 2015. Catalogo de invertebrados citados en cuevas, simas y minas de Andalucia. Monografias Bioespeleologicas 9: 1 - 97." type="journal article" year="2015">Pérez 2015</bibRefCitation>
); Vilalluenga del Rosario, Sima Aljibe nº 2, (
<bibRefCitation id="EFC392D46668AB5EB43E64CEFE3F3AF1" author="Ribera" box="[338,489,419,445]" firstAuthor="Ribera" pageId="13" pageNumber="14" pagination="123 - 129" refId="ref13567" refString="Ribera C. 1970. Resultados faunisticos de la campana Serrania de Ronda. In: Actas I Congreso Nacional de Espeleologia: 123 - 129. Editorial Ayuntamiento, Barcelona." type="book chapter" year="1970">Ribera 1970</bibRefCitation>
). — Málaga, Cortés de la Frontera, Cueva del Berrueco (
<bibRefCitation id="EFC392D46668AB5EB1E564CEFAF13AF1" author="Jeannel" box="[1161,1319,419,445]" firstAuthor="Jeannel" pageId="13" pageNumber="14" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel 1922</bibRefCitation>
,
<bibRefCitation id="EFC392D46668AB5EB05864CEFAA43AF1" author="Jeannel" box="[1332,1394,419,445]" firstAuthor="Jeannel" pageId="13" pageNumber="14" pagination="1" refId="ref12306" refString="Jeannel R. 1936. Monographie des Catopides. Memoires du Museum national d'Histoire naturelle 1 (1). Museum national d'Histoire naturelle, Paris." type="journal article" year="1936">1936</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB5D164ABFE853AAC" author="Ribera" box="[189,339,454,480]" firstAuthor="Ribera" pageId="13" pageNumber="14" pagination="123 - 129" refId="ref13567" refString="Ribera C. 1970. Resultados faunisticos de la campana Serrania de Ronda. In: Actas I Congreso Nacional de Espeleologia: 123 - 129. Editorial Ayuntamiento, Barcelona." type="book chapter" year="1970">Ribera 1970</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB40E64ABFE0B3AAC" author="Blas M." box="[354,477,454,480]" firstAuthor="Blas" journalOrPublisher="Congreso Nacional de Espeleologia" pageId="13" pageNumber="14" pagination="157 - 160" part="4" refId="ref11503" refString="Blas M. 1976. Coleopteros cavernicolas del distrito andaluz (Catopidae). Actas 4 Congreso Nacional de Espeleologia: 157 - 160." title="Coleopteros cavernicolas del distrito andaluz (Catopidae). Actas" type="journal article" year="1976">Blas 1976</bibRefCitation>
,
<bibRefCitation id="EFC392D46668AB5EB48764ABFDFF3AAC" author="Blas M." box="[491,553,454,480]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="13" pageNumber="14" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">1979</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB75464ABFCB43AAC" author="Giachino P. M. &amp; Vailati D." box="[568,866,454,480]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="13" pageNumber="14" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB61C64ABFBD23AAC" author="Tinaut" box="[880,1028,454,480]" firstAuthor="Tinaut" pageId="13" pageNumber="14" pagination="3 - 28" refId="ref13778" refString="Tinaut A. 1998. Artropodos terrestres de las cavidades andaluzas. Zoologica Baetica 9: 3 - 28." type="journal article" year="1998">Tinaut 1998</bibRefCitation>
); Cortés de la Frontera, Cueva de las Motillas (
<bibRefCitation id="EFC392D46668AB5EB4106484FDC43948" author="Ribera" box="[380,530,489,516]" firstAuthor="Ribera" pageId="13" pageNumber="14" pagination="123 - 129" refId="ref13567" refString="Ribera C. 1970. Resultados faunisticos de la campana Serrania de Ronda. In: Actas I Congreso Nacional de Espeleologia: 123 - 129. Editorial Ayuntamiento, Barcelona." type="book chapter" year="1970">Ribera 1970</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB7736484FD4F3948" author="Blas M." box="[543,665,489,516]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="13" pageNumber="14" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">Blas 1977</bibRefCitation>
,
<bibRefCitation id="EFC392D46668AB5EB7CA6487FD353948" author="Blas M." box="[678,739,490,516]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="13" pageNumber="14" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">1979</bibRefCitation>
,
<bibRefCitation id="EFC392D46668AB5EB79C6487FCF83948" author="Blas M." box="[752,814,490,516]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="13" pageNumber="14" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">1989</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB6506484FBB73948" author="Giachino P. M. &amp; Vailati D." box="[828,1121,489,516]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="13" pageNumber="14" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB1016484FB293948" author="Tinaut" box="[1133,1279,489,516]" firstAuthor="Tinaut" pageId="13" pageNumber="14" pagination="3 - 28" refId="ref13778" refString="Tinaut A. 1998. Artropodos terrestres de las cavidades andaluzas. Zoologica Baetica 9: 3 - 28." type="journal article" year="1998">Tinaut 1998</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB0606484FEE9396B" author="Fresneda J. &amp; Cardenas A. M. &amp; Castro A. &amp; Lencina J. L. &amp; Lopez-Colon J. I. &amp; Baena. M." etAl="et al." firstAuthor="Fresneda" journalOrPublisher="Boletin de la Asociacion espanola de entomologia" pageId="13" pageNumber="14" pagination="187 - 214" part="31" refId="ref11915" refString="Fresneda J., Cardenas A. M., Castro A., Lencina J. L., Lopez-Colon J. I. &amp; Baena. M. 2007. Nuevos datos de los Cholevidae en la Peninsula Iberica (Coleoptera). Boletin de la Asociacion espanola de entomologia 31 (3 - 4): 187 - 214." title="Nuevos datos de los Cholevidae en la Peninsula Iberica (Coleoptera)" type="journal article" year="2007">
Fresneda
<emphasis id="B92633376668AB5EB5D16760FF2E396B" box="[189,248,525,551]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2007
</bibRefCitation>
); Montejaque, El Pozuelo, Cueva Sima Nueva (
<bibRefCitation id="EFC392D46668AB5EB61C6761FC3A396B" author="Blas M." box="[880,1004,524,551]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="13" pageNumber="14" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas 1989</bibRefCitation>
); Montejaque, El Pozuelo, Cueva del Requejo (
<bibRefCitation id="EFC392D46668AB5EB436675DFE023906" author="Blas M." box="[346,468,560,586]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="13" pageNumber="14" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">Blas 1979</bibRefCitation>
,
<bibRefCitation id="EFC392D46668AB5EB48C675DFDC83906" author="Blas M." box="[480,542,560,586]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="13" pageNumber="14" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">1989</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB746675DFD6D3906" author="Tinaut" box="[554,699,560,586]" firstAuthor="Tinaut" pageId="13" pageNumber="14" pagination="3 - 28" refId="ref13778" refString="Tinaut A. 1998. Artropodos terrestres de las cavidades andaluzas. Zoologica Baetica 9: 3 - 28." type="journal article" year="1998">Tinaut 1998</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB7AB675DFC993907" author="Perez" box="[711,847,560,587]" firstAuthor="Perez" pageId="13" pageNumber="14" pagination="1 - 97" refId="ref12561" refString="Perez T. 2015. Catalogo de invertebrados citados en cuevas, simas y minas de Andalucia. Monografias Bioespeleologicas 9: 1 - 97." type="journal article" year="2015">Pérez 2015</bibRefCitation>
); Montejaque, El Pozuelo, Cueva Sivieja (
<bibRefCitation id="EFC392D46668AB5EB02D675DFF2D3921" author="Blas M." firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="13" pageNumber="14" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas 1989</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB46B673EFE413921" author="Tinaut" box="[263,407,595,621]" firstAuthor="Tinaut" pageId="13" pageNumber="14" pagination="3 - 28" refId="ref13778" refString="Tinaut A. 1998. Artropodos terrestres de las cavidades andaluzas. Zoologica Baetica 9: 3 - 28." type="journal article" year="1998">Tinaut 1998</bibRefCitation>
;
<bibRefCitation id="EFC392D46668AB5EB4C8673EFDFA3922" author="Perez" box="[420,556,595,622]" firstAuthor="Perez" pageId="13" pageNumber="14" pagination="1 - 97" refId="ref12561" refString="Perez T. 2015. Catalogo de invertebrados citados en cuevas, simas y minas de Andalucia. Monografias Bioespeleologicas 9: 1 - 97." type="journal article" year="2015">Pérez 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE6668AB5EB5D167F9FAA13E87" pageId="13" pageNumber="14" type="description">
<paragraph id="8BEDEF256668AB5EB5D167F9FEBD39E2" blockId="13.[189,363,660,686]" box="[189,363,660,686]" pageId="13" pageNumber="14">
<heading id="D0A558496668AB5EB5D167F9FEBD39E2" bold="true" box="[189,363,660,686]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="B92633376668AB5EB5D167F9FEBD39E2" bold="true" box="[189,363,660,686]" pageId="13" pageNumber="14">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D167AEFAA6384D" blockId="13.[189,1399,707,769]" pageId="13" pageNumber="14">
BODY. Length 4.0–4.5 (males) and
<quantity id="4CAA42C06668AB5EB73467AEFD303991" box="[600,742,707,733]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.7" metricValueMin="3.9" pageId="13" pageNumber="14" unit="mm" value="4.3" valueMax="4.7" valueMin="3.9">3.9–4.7 mm</quantity>
(females), body width 1.4–1.5 (males) and
<quantity id="4CAA42C06668AB5EB18567AEFAA13991" box="[1257,1399,707,734]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="13" pageNumber="14" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
(females) (
<figureCitation id="1369F3A06668AB5EB450678BFE53384C" box="[316,389,742,768]" captionStart="Figs 1 – 5" captionStartId="3.[189,241,1892,1918]" captionTargetBox="[189,1395,1323,1850]" captionTargetId="figure@3.[189,1398,1323,1850]" captionTargetPageId="3" captionText="Figs 1 – 5. Speonemadus of the escalera i- group habitus. 1. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 2. S. angusticollis (Kraatz, 1870) (Rute, Spain). 3. S. bolivari (Jeannel, 1922) (Ardales, Spain). 4. S. breuili (Jeannel, 1922) (Motillas, Spain). 5. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 1 mm." httpUri="https://zenodo.org/record/889348/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
). Body color, pubescence and punctuation similar to that observed in
<taxonomicName id="4C5294A66668AB5EB1DA678AFABF384C" baseAuthorityName="Blas" baseAuthorityYear="1989" box="[1206,1385,742,769]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B92633376668AB5EB1DA678AFABF384C" box="[1206,1385,742,769]" italics="true" pageId="13" pageNumber="14">S. angusticollis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D16645FD7738C4" blockId="13.[189,1399,806,905]" pageId="13" pageNumber="14">
ANTENNAE. Reaching the basal 4th part of elytra; the antennomeres 1
<taxonomicName id="4C5294A66668AB5EB6B9664BFBC7380E" box="[981,1041,806,834]" pageId="13" pageNumber="14" rank="subSpecies" subSpecies="and">st and</taxonomicName>
2nd are almost equal in length, also equal in length are the 4th, 5th and 7th ones being the 3rd ones clearly longer than the 2nd one and the 8th nearly twice as long as wide (
<tableCitation id="C6D0DA9E6668AB5EB7546603FD4638C4" box="[568,656,878,904]" captionStart="Table 1" captionStartId="4.[188,257,267,293]" captionTargetBox="[193,1384,385,1378]" captionTargetPageId="4" captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width." httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04" pageId="13" pageNumber="14" tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D166DDFD9C3F5C" blockId="13.[189,1398,943,1040]" pageId="13" pageNumber="14">
PRONOTUM. Slightly transverse, maximum width at middle third part; the angles converge, smoothly from the middle area to the base in a straight line; posterior angles obtuse and rounded at vertex; sculpture and granularity well defined (
<figureCitation id="1369F3A06668AB5EB489669BFDEB3F5C" box="[485,573,1014,1040]" captionStart="Figs 6–15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6–15. Speonemadus of the escalerai-group pronotum. 6–7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 8–9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 10–11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 12–13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 14–15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="13" pageNumber="14">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D16155FEDF3FD4" blockId="13.[189,1399,1079,1176]" pageId="13" pageNumber="14">
ELYTRA. Elliptical and elongated, maximum length/width ratio = 1.92–2.10 (males) and
<quantity id="4CAA42C06668AB5EB1A7615AFAA13F1E" box="[1227,1399,1079,1106]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.04" metricValueMin="1.86" pageId="13" pageNumber="14" unit="mm" value="1.95" valueMax="2.04" valueMin="1.86">1.86–2.04 mm</quantity>
(females), with overall shape of the species group and sculpture of “C”
<typeStatus id="54E951876668AB5EB1676136FBE83F39" box="[1035,1086,1115,1141]" pageId="13" pageNumber="14">type</typeStatus>
(
<emphasis id="B92633376668AB5EB1236136FB463F38" box="[1103,1168,1115,1140]" italics="true" pageId="13" pageNumber="14">sensu</emphasis>
<bibRefCitation id="EFC392D46668AB5EB1F46137FF2A3FD4" author="Giachino P. M. &amp; Vailati D." firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="13" pageNumber="14" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
).
</paragraph>
<paragraph id="8BEDEF256668AB5EB58661D2FE413F95" blockId="13.[189,1306,1215,1277]" box="[234,407,1215,1241]" pageId="13" pageNumber="14">
<emphasis id="B92633376668AB5EB58661D2FEFC3F95" bold="true" box="[234,298,1215,1241]" pageId="13" pageNumber="14">Male</emphasis>
(
<figureCitation id="1369F3A06668AB5EB45661D2FE463F95" box="[314,400,1215,1241]" captionStart="Figs 21 – 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 – 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="13" pageNumber="14">Fig. 24</figureCitation>
)
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D1618EFACC3FB1" blockId="13.[189,1306,1215,1277]" box="[189,1306,1250,1277]" pageId="13" pageNumber="14">
Features of the genital segment and aedeagus are similar to those described for
<taxonomicName id="4C5294A66668AB5EB10C618EFAC53FB0" baseAuthorityName="Blas" baseAuthorityYear="1989" box="[1120,1299,1250,1277]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B92633376668AB5EB10C618EFAC53FB0" box="[1120,1299,1250,1277]" italics="true" pageId="13" pageNumber="14">S. angusticollis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEDEF256668AB5EB5866049FE933E72" blockId="13.[189,1399,1316,1483]" box="[234,325,1316,1342]" pageId="13" pageNumber="14">
<heading id="D0A558496668AB5EB5866049FE933E72" bold="true" box="[234,325,1316,1342]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="6">
<emphasis id="B92633376668AB5EB5866049FE933E72" bold="true" box="[234,325,1316,1342]" pageId="13" pageNumber="14">Female</emphasis>
</heading>
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D1602AFAA13E87" blockId="13.[189,1399,1316,1483]" pageId="13" pageNumber="14">
The antennae are similar to those of the male, but the antennomeres are more transverse; 2nd, 4th, 5th and 7th antennomeres having similar length (
<tableCitation id="C6D0DA9E6668AB5EB7F26007FD2E3EC8" box="[670,760,1386,1412]" captionStart="Table 1" captionStartId="4.[188,257,267,293]" captionTargetBox="[193,1384,385,1378]" captionTargetPageId="4" captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width." httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04" pageId="13" pageNumber="14" tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
). Pronotum less elongated; the overall shape and the ratio of width/ length ratio similar to that described for males (
<figureCitation id="1369F3A06668AB5EB6C860E0FC2A3EEB" box="[932,1020,1421,1447]" captionStart="Figs 6–15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6–15. Speonemadus of the escalerai-group pronotum. 6–7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♂ and ♀. 8–9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♂ and ♀. 10–11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♂ and ♀. 12–13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♂ and ♀. 14–15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♂ and ♀. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.889350" httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="13" pageNumber="14">Fig. 13</figureCitation>
). Elytra elliptical and elongated with apical parts notched and serrated. 7th and 8th uroventrites, and genitalia typical of the species group.
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE6668AB5EB5D1609FFC713CF5" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BEDEF256668AB5EB5D1609FFEF93D40" blockId="13.[189,303,1522,1548]" box="[189,303,1522,1548]" pageId="13" pageNumber="14">
<heading id="D0A558496668AB5EB5D1609FFEF93D40" bold="true" box="[189,303,1522,1548]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="B92633376668AB5EB5D1609FFEF93D40" bold="true" box="[189,303,1522,1548]" pageId="13" pageNumber="14">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D1634CFB983C7D" blockId="13.[189,1399,1568,1841]" pageId="13" pageNumber="14">
<taxonomicName id="4C5294A66668AB5EB5D1634CFE623D77" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[189,436,1569,1595]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B92633376668AB5EB5D1634CFE623D77" box="[189,436,1569,1595]" italics="true" pageId="13" pageNumber="14">Speonemadus breuili</emphasis>
</taxonomicName>
is re-established as a valid species due to the clear morphological characters that differentiate it from
<taxonomicName id="4C5294A66668AB5EB4DD6329FDB13D12" baseAuthorityName="Blas" baseAuthorityYear="1989" box="[433,615,1604,1630]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B92633376668AB5EB4DD6329FDB13D12" box="[433,615,1604,1630]" italics="true" pageId="13" pageNumber="14">S. angusticollis</emphasis>
</taxonomicName>
, which was synonymized by
<treatmentCitation id="0AF3C9346668AB5EB6A16329FB8A3D12" author="Blas M." box="[973,1116,1604,1630]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="13" pageNumber="14" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" treatmentTaxon="unknown" type="journal article" year="1989">
<bibRefCitation id="EFC392D46668AB5EB6A16329FB8A3D12" author="Blas M." box="[973,1116,1604,1630]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="13" pageNumber="14" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas (1989)</bibRefCitation>
</treatmentCitation>
. This classification was subsequently followed by several authors (
<treatmentCitation id="0AF3C9346668AB5EB7C2630AFC063DCD" author="Giachino P. M. &amp; Vailati D." box="[686,976,1639,1665]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="13" pageNumber="14" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" treatmentTaxon="unknown" type="journal article" year="1993">
<bibRefCitation id="EFC392D46668AB5EB7C2630AFC063DCD" author="Giachino P. M. &amp; Vailati D." box="[686,976,1639,1665]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana" pageId="13" pageNumber="14" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. &amp; Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di &quot; Natura Bresciana &quot; 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF3C9346668AB5EB6B7630AFBAC3DCD" author="Perreau" box="[987,1146,1639,1665]" firstAuthor="Perreau" pageId="13" pageNumber="14" pagination="4" refId="ref12692" refString="Perreau M. 2000. Catalogue des Coleopteres Leiodidae Cholevinae et Platypsyllinae. Memoires de la SEF 4, Societe entomologique de France, Paris." treatmentTaxon="unknown" type="book chapter" year="2000">
<bibRefCitation id="EFC392D46668AB5EB6B7630AFBAC3DCD" author="Perreau" box="[987,1146,1639,1665]" firstAuthor="Perreau" pageId="13" pageNumber="14" pagination="4" refId="ref12692" refString="Perreau M. 2000. Catalogue des Coleopteres Leiodidae Cholevinae et Platypsyllinae. Memoires de la SEF 4, Societe entomologique de France, Paris." type="book chapter" year="2000">Perreau 2000</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF3C9346668AB5EB1E9630AFABD3DCD" author="Salgado" box="[1157,1387,1639,1665]" etAl="et al." firstAuthor="Salgado" pageId="13" pageNumber="14" refId="ref13714" refString="Salgado J. M., Blas M. &amp; Fresneda J. 2008. Coleoptera, Cholevidae. Fauna Iberica, vol. 31, Museo Nacional de Ciencias Naturales, CSIC, Madrid." treatmentTaxon="unknown" type="journal volume" year="2008">
<bibRefCitation id="EFC392D46668AB5EB1E9630AFABD3DCD" author="Salgado" box="[1157,1387,1639,1665]" etAl="et al." firstAuthor="Salgado" pageId="13" pageNumber="14" refId="ref13714" refString="Salgado J. M., Blas M. &amp; Fresneda J. 2008. Coleoptera, Cholevidae. Fauna Iberica, vol. 31, Museo Nacional de Ciencias Naturales, CSIC, Madrid." type="journal volume" year="2008">
Salgado
<emphasis id="B92633376668AB5EB1866305FAF23DCD" box="[1258,1316,1639,1665]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2008
</bibRefCitation>
</treatmentCitation>
). The main character used to re-evaluate its specific status is the shape of the male protibial keel, which is shorter and with a more rounded ridge, but also because in
<taxonomicName id="4C5294A66668AB5EB63063C3FC123D8B" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[860,964,1709,1736]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B92633376668AB5EB63063C3FC123D8B" box="[860,964,1709,1736]" italics="true" pageId="13" pageNumber="14">S. breuili</emphasis>
</taxonomicName>
the 8th antennomere is less transverse, the elytra are proportionally shorter, and the female pronotum is more hexagonal and clearly transverse. In addition,
<taxonomicName id="4C5294A66668AB5EB4226399FD983C42" box="[334,590,1780,1806]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="verneri">
<emphasis id="B92633376668AB5EB4226399FD983C42" box="[334,590,1780,1806]" italics="true" pageId="13" pageNumber="14">Speonemadus verneri</emphasis>
</taxonomicName>
is established as a synonym of
<taxonomicName id="4C5294A66668AB5EB6A66399FBE03C42" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[970,1078,1780,1806]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B92633376668AB5EB6A66399FBE03C42" box="[970,1078,1780,1806]" italics="true" pageId="13" pageNumber="14">S. breuili</emphasis>
</taxonomicName>
and not of
<treatmentCitation id="0AF3C9346668AB5EB1AE6399FE9C3C7D" author="Blas M." authority="(Blas 1989)" authorityName="Blas" authorityYear="1989" baseAuthorityName="Blas" baseAuthorityYear="1989" class="Insecta" family="Leiodidae" firstAuthor="Blas" genus="Anemadus" higherTaxonomySource="GBIF" journalOrPublisher="Memoires de Biospeologie" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" pagination="135 - 140" part="16" phylum="Arthropoda" rank="species" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." species="angusticollis" title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" treatmentTaxon="S. angusticollis" type="journal article" year="1989">
<taxonomicName id="4C5294A66668AB5EB1AE6399FE9C3C7D" authority="(Blas 1989)" baseAuthorityName="Blas" baseAuthorityYear="1989" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B92633376668AB5EB1AE6399FAA03C42" box="[1218,1398,1780,1806]" italics="true" pageId="13" pageNumber="14">S. angusticollis</emphasis>
(
<bibRefCitation id="EFC392D46668AB5EB5A9627AFE963C7D" author="Blas M." box="[197,320,1815,1841]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="13" pageNumber="14" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas 1989</bibRefCitation>
)
</taxonomicName>
</treatmentCitation>
, due to the absence of relevant differences compared to
<taxonomicName id="4C5294A66668AB5EB6B0627AFB913C7D" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[988,1095,1815,1841]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B92633376668AB5EB6B0627AFB913C7D" box="[988,1095,1815,1841]" italics="true" pageId="13" pageNumber="14">S. breuili</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEDEF256668AB5EB5D16234FC713CF5" blockId="13.[189,1399,1880,1977]" pageId="13" pageNumber="14">
Apart from the morphological differences pointed out, there is an obvious geographical isolation.
<taxonomicName id="4C5294A66668AB5EB5D16211FE633CDA" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[189,437,1916,1942]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B92633376668AB5EB5D16211FE633CDA" box="[189,437,1916,1942]" italics="true" pageId="13" pageNumber="14">Speonemadus breuili</emphasis>
</taxonomicName>
colonizes the southernmost caves in Spanish
<collectingRegion id="499621C76668AB5EB6B26216FB8E3CD9" box="[990,1112,1915,1941]" country="Spain" name="Andalucia" pageId="13" pageNumber="14">Andalusia</collectingRegion>
, the provinces of Cadiz and Málaga, away from the distribution area of
<taxonomicName id="4C5294A66668AB5EB78162F2FC763CF5" baseAuthorityName="Blas" baseAuthorityYear="1989" box="[749,928,1951,1977]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B92633376668AB5EB78162F2FC763CF5" box="[749,928,1951,1977]" italics="true" pageId="13" pageNumber="14">S. angusticollis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE666BAB5DB5D16466FA973A3A" pageId="14" pageNumber="15" type="biology_ecology">
<paragraph id="8BEDEF25666BAB5DB5D16466FE6A3A69" blockId="14.[189,1399,267,375]" box="[189,444,267,293]" pageId="14" pageNumber="15">
<heading id="D0A55849666BAB5DB5D16466FE6A3A69" bold="true" box="[189,444,267,293]" fontSize="11" level="2" pageId="14" pageNumber="15" reason="6">
<emphasis id="B9263337666BAB5DB5D16466FE6A3A69" bold="true" box="[189,444,267,293]" pageId="14" pageNumber="15">Biology and Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEDEF25666BAB5DB5D16454FA973A3A" blockId="14.[189,1399,267,375]" pageId="14" pageNumber="15">
<taxonomicName id="4C5294A6666BAB5DB5D16454FE653A1F" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[189,435,313,339]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B9263337666BAB5DB5D16454FE653A1F" box="[189,435,313,339]" italics="true" pageId="14" pageNumber="15">Speonemadus breuili</emphasis>
</taxonomicName>
is commonly found in the subterranean ecosystem, as are the other species of the group. Inside the caves, it is found mainly in parts rich in organic matter and with a high humidity.
</paragraph>
</subSubSection>
<subSubSection id="C348BCAE666BAB5DB5D164F3FC11393F" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BEDEF25666BAB5DB5D164F3FE803AF4" blockId="14.[189,342,414,440]" box="[189,342,414,440]" pageId="14" pageNumber="15">
<heading id="D0A55849666BAB5DB5D164F3FE803AF4" bold="true" box="[189,342,414,440]" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<emphasis id="B9263337666BAB5DB5D164F3FE803AF4" bold="true" box="[189,342,414,440]" pageId="14" pageNumber="15">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEDEF25666BAB5DB5D164A0FC11393F" blockId="14.[189,1399,460,628]" pageId="14" pageNumber="15">
<taxonomicName id="4C5294A6666BAB5DB5D164A0FE653AAB" authorityName="Ana Sofa P.S. Reboleira &amp; Javier Fresneda &amp; José Maria Salgado" authorityYear="2017" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[189,435,461,487]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B9263337666BAB5DB5D164A0FE653AAB" box="[189,435,461,487]" italics="true" pageId="14" pageNumber="15">Speonemadus breuili</emphasis>
</taxonomicName>
is the southernmost species of the
<taxonomicName id="4C5294A6666BAB5DB62164A0FC6F3AAB" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[845,953,461,487]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="escalerai">
<emphasis id="B9263337666BAB5DB62164A0FC6F3AAB" box="[845,953,461,487]" italics="true" pageId="14" pageNumber="15">escalerai</emphasis>
</taxonomicName>
-group in the Iberian Peninsula, found in the provinces of Cadiz and Málaga. The limit of its northern distribution is established by the mountain ridge of Ronda. It is distributed in the Sierra de Grazalema and related foothills, while the distribution of
<taxonomicName id="4C5294A6666BAB5DB58C675AFE8B391C" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[224,349,566,593]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B9263337666BAB5DB58C675AFE8B391C" box="[224,349,566,593]" italics="true" pageId="14" pageNumber="15">S. bolivari</emphasis>
</taxonomicName>
is linked to the Sierra de
<collectingCountry id="F345AFB5666BAB5DB7F1675BFD38391C" box="[669,750,566,592]" name="Saint Kitts and Nevis" pageId="14" pageNumber="15">Nieves</collectingCountry>
and its foothills. Both mountain ranges are separated geographically by a wide valley of sedimentary material (
<figureCitation id="1369F3A0666BAB5DB60E6734FC6C393F" box="[866,954,601,627]" captionStart="Fig. 32" captionStartId="18.[189,232,1955,1981]" captionTargetBox="[189,1398,986,1883]" captionTargetId="figure@18.[189,1398,986,1914]" captionTargetPageId="18" captionText="Fig. 32. Distribution of Speonemadus of the escalerai - group: S. algarvensis sp. nov., S. angusticollis (Kraatz, 1870), S. bolivari (Jeannel, 1922), S. breuili (Jeannel, 1922) and S. escalerai (Uhagón, 1898)." httpUri="https://zenodo.org/record/889358/files/figure.png" pageId="14" pageNumber="15">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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