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<document id="094F551356CCC10537282B99CA3ACDA2" ID-DOI="10.11646/phytotaxa.663.3.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="14515672" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734535148798" checkinUser="felipe" docAuthor="Negaresh, Kazem" docDate="2024" docId="0399FB4CFF8FFFC4FF70FF294487F950" docLanguage="en" docName="phytotaxa.663.3.157-164.pdf" docOrigin="Phytotaxa 663 (3)" docSource="https://doi.org/10.11646/phytotaxa.663.3.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Centaurea karajensis Negaresh" docType="treatment" docVersion="2" lastPageNumber="158" masterDocId="FFA08334FF8EFFC5FFF8FFDD4107FFFE" masterDocTitle="Taxonomic notes on Centaurea sect. Ammocyanus (Asteraceae, Cardueae- Centaureinae) from Iran" masterLastPageNumber="164" masterPageNumber="157" pageNumber="158" updateTime="1734535638623" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="69E6B28DCF0B2308ABC3556D8C339B03" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="335F871BC77F72FCEFDEBD6B1F84E332">
<mods:title id="5D5355C4F9EE25638F028B686543DE2C">Taxonomic notes on Centaurea sect. Ammocyanus (Asteraceae, Cardueae- Centaureinae) from Iran</mods:title>
</mods:titleInfo>
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<mods:namePart id="C40403DDE629B084DEA8BF5126AA8D26">Negaresh, Kazem</mods:namePart>
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<mods:typeOfResource id="2BC6958CF79230DD03E705CBDF77D2C6">text</mods:typeOfResource>
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<mods:title id="36A08585EB777B2EDAACCC80F24FFC5B">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="E288275674ED11547C502B688E673662">
<mods:date id="4873E3CD17AAFA6042A1EA0B9BC2B431">2024</mods:date>
<mods:detail id="E8EAF1318BDFE1BAAD3DFB9A9E9BEC55" type="pubDate">
<mods:number id="1779F7D0CEE28A315297D68194ED05B7">2024-08-22</mods:number>
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<mods:url id="3C209759566133C62530A5F3C6D7A1DB">https://doi.org/10.11646/phytotaxa.663.3.5</mods:url>
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<mods:classification id="F9949B82EE1006A61FA83085FBDC7160">journal article</mods:classification>
<mods:identifier id="B4A6B5C14D7CC442D942929CAB0B743B" type="DOI">10.11646/phytotaxa.663.3.5</mods:identifier>
<mods:identifier id="CD62CC611956C6BDAA87A44A13ED7B69" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="D9F62C0558E3A29EC2608F381BE9A778" type="Zenodo-Dep">14515672</mods:identifier>
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<treatment id="0399FB4CFF8FFFC4FF70FF294487F950" ID-DOI="http://doi.org/10.5281/zenodo.14515668" ID-Zenodo-Dep="14515668" LSID="urn:lsid:plazi:treatment:0399FB4CFF8FFFC4FF70FF294487F950" httpUri="http://treatment.plazi.org/id/0399FB4CFF8FFFC4FF70FF294487F950" lastPageNumber="158" pageId="1" pageNumber="158">
<subSubSection id="C32A19D1FF8FFFC4FF70FF2943A8FEF0" box="[136,687,244,271]" pageId="1" pageNumber="158" type="nomenclature">
<paragraph id="8B8F4A5AFF8FFFC4FF70FF2943A8FEF0" blockId="1.[136,687,244,271]" box="[136,687,244,271]" pageId="1" pageNumber="158">
<heading id="D0C7FD36FF8FFFC4FF70FF2943A8FEF0" box="[136,687,244,271]" fontSize="11" level="2" pageId="1" pageNumber="158" reason="3">
<taxonomicName id="4C3031D9FF8FFFC4FF70FF2940F4FEF0" ID-CoL="CJYVX" authority="Negaresh" authorityName="Negaresh" box="[136,499,244,271]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="karajensis">
<emphasis id="B9449648FF8FFFC4FF70FF294079FEF1" bold="true" box="[136,382,244,271]" italics="true" pageId="1" pageNumber="158">Centaurea karajensis</emphasis>
Negaresh
</taxonomicName>
,
<emphasis id="B9449648FF8FFFC4FE05FF294353FEF0" bold="true" box="[509,596,244,270]" pageId="1" pageNumber="158">
s
<emphasis id="B9449648FF8FFFC4FDF0FF294353FEF0" bold="true" box="[520,596,244,270]" italics="true" pageId="1" pageNumber="158">p. nov.</emphasis>
</emphasis>
(
<figureCitation id="130B56DFFF8FFFC4FD9CFF2943A3FEF0" box="[612,676,244,270]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[181,1407,187,1925]" captionTargetId="figure-34@2.[181,1407,187,1925]" captionTargetPageId="2" captionText="FIGURE 1. Centaurea karajensis Negaresh (Rechinger 638, W)." figureDoi="http://doi.org/10.5281/zenodo.14515674" httpUri="https://zenodo.org/record/14515674/files/figure.png" pageId="1" pageNumber="158">Fig. 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A19D1FF8FFFC4FF70FE9D433EFE87" pageId="1" pageNumber="158" type="materials_examined">
<paragraph id="8B8F4A5AFF8FFFC4FF70FE9D433EFE87" blockId="1.[136,1452,320,630]" pageId="1" pageNumber="158">
<emphasis id="B9449648FF8FFFC4FF70FE9D41C5FEAB" bold="true" box="[136,194,320,341]" pageId="1" pageNumber="158">Type:</emphasis>
—
<collectingCountry id="F3270ACAFF8FFFC4FF23FE9D401FFEAB" box="[219,280,320,341]" name="Iran" pageId="1" pageNumber="158">IRAN</collectingCountry>
.
<collectingRegion id="49F484B8FF8FFFC4FEDAFE9D40C1FEA8" box="[290,454,320,342]" country="Iran" name="Alborz" pageId="1" pageNumber="158">Alborz Province</collectingRegion>
: in ditione oppidi Karaj, in montibus cdc. Kuh-e Nemar,
<quantity id="4CC8E7BFFF8FFFC4FBF8FE9D458EFEAB" box="[1024,1161,320,342]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="1" pageNumber="158" unit="m" value="1800.0" valueMax="2000.0" valueMin="1600.0">1600–2000 m</quantity>
,
<date id="FF8E6C9AFF8FFFC4FB6DFE9D4417FEAB" box="[1173,1296,320,341]" pageId="1" pageNumber="158" value="1937-06-03">3 June 1937</date>
,
<emphasis id="B9449648FF8FFFC4FAE4FE9D44ACFEAB" box="[1308,1451,320,341]" italics="true" pageId="1" pageNumber="158">Rechinger 638</emphasis>
(
<typeStatus id="548BF4F8FF8FFFC4FF3CFEB9401DFE84" box="[196,282,356,378]" pageId="1" pageNumber="158" type="holotype">holotype</typeStatus>
<collectionCode id="ED21D29FFF8FFFC4FEE7FEB9403BFE87" box="[287,316,356,377]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="158" type="Herbarium">W</collectionCode>
!,
<typeStatus id="548BF4F8FF8FFFC4FEB2FEB9409EFE84" box="[330,409,356,378]" pageId="1" pageNumber="158" type="isotype">isotypes</typeStatus>
<collectingCountry id="F3270ACAFF8FFFC4FE66FEB940E7FE87" box="[414,480,356,377]" name="Iran" pageId="1" pageNumber="158">IRAN</collectingCountry>
!,
<collectionCode id="ED21D29FFF8FFFC4FE14FEB9432CFE87" box="[492,555,356,377]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="1" pageNumber="158">TARI</collectionCode>
!).
</paragraph>
</subSubSection>
<subSubSection id="C32A19D1FF8FFFC4FF70FE5A4243FB24" pageId="1" pageNumber="158" type="diagnosis">
<paragraph id="8B8F4A5AFF8FFFC4FF70FE5A4050FD88" blockId="1.[136,1452,320,630]" pageId="1" pageNumber="158">
<emphasis id="B9449648FF8FFFC4FF70FE5A41F4FE63" bold="true" box="[136,243,391,413]" pageId="1" pageNumber="158">Diagnosis:</emphasis>
—
<taxonomicName id="4C3031D9FF8FFFC4FEF3FE5540DDFE63" box="[267,474,392,413]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="karajensis">
<emphasis id="B9449648FF8FFFC4FEF3FE5540DDFE63" box="[267,474,392,413]" italics="true" pageId="1" pageNumber="158">Centaurea karajensis</emphasis>
</taxonomicName>
is related to
<taxonomicName id="4C3031D9FF8FFFC4FDA7FE5543FCFE63" authorityName="Boissier" authorityYear="1849" box="[607,763,392,413]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="ammocyanus">
<emphasis id="B9449648FF8FFFC4FDA7FE5543FCFE63" box="[607,763,392,413]" italics="true" pageId="1" pageNumber="158">C. ammocyanus</emphasis>
</taxonomicName>
, from which it differs mainly in its plant never with capitula at base, main stem elongated,
<quantity id="4CC8E7BFFF8FFFC4FE6CFE7140F3FE3F" box="[404,500,428,450]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="158" unit="cm" value="55.0" valueMax="60.0" valueMin="50.0">50–60 cm</quantity>
long (
<emphasis id="B9449648FF8FFFC4FDCEFE71434DFE3F" box="[566,586,428,449]" italics="true" pageId="1" pageNumber="158">vs</emphasis>
. central capitulum nearly sessile, main stem very short,
<quantity id="4CC8E7BFFF8FFFC4FB89FE7145BEFE3F" box="[1137,1209,428,449]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="1" pageNumber="158" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
long), stem with several branches divaricated and unequal in length (
<emphasis id="B9449648FF8FFFC4FD93FE0D4378FE1B" box="[619,639,464,485]" italics="true" pageId="1" pageNumber="158">vs</emphasis>
. with several prostrate or ascending branches arising below the terminal capitulum, often further branched), median cauline leaves narrowly linear (
<emphasis id="B9449648FF8FFFC4FCCEFE29424DFDF7" box="[822,842,500,521]" italics="true" pageId="1" pageNumber="158">vs</emphasis>
. lanceolate), upper cauline leaves narrowly linear to acicular, shorter than involucre (
<emphasis id="B9449648FF8FFFC4FE59FDC540B2FDD3" box="[417,437,536,557]" italics="true" pageId="1" pageNumber="158">vs</emphasis>
. linear-lanceolate, inserted directly below capitulum, usually longer than involucre), cilia
<quantity id="4CC8E7BFFF8FFFC4FAD8FDC54485FDD3" box="[1312,1410,536,558]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="158" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
(
<emphasis id="B9449648FF8FFFC4FA69FDC544A2FDD3" box="[1425,1445,536,557]" italics="true" pageId="1" pageNumber="158">vs</emphasis>
.
<quantity id="4CC8E7BFFF8FFFC4FF45FDE14033FDAF" box="[189,308,572,594]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="1" pageNumber="158" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
) long, spine erect,
<quantity id="4CC8E7BFFF8FFFC4FE17FDE14356FDAF" box="[495,593,572,594]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="158" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long (
<emphasis id="B9449648FF8FFFC4FD6BFDE143A0FDAF" box="[659,679,572,593]" italics="true" pageId="1" pageNumber="158">vs</emphasis>
. patent, sometimes squarrose,
<quantity id="4CC8E7BFFF8FFFC4FC2CFDE14523FDAF" box="[980,1060,572,593]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="158" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long), central florets 5–8 (
<emphasis id="B9449648FF8FFFC4FAD4FDE14447FDAF" box="[1324,1344,572,593]" italics="true" pageId="1" pageNumber="158">vs</emphasis>
. 15–20) in each capitulum.
</paragraph>
<paragraph id="8B8F4A5AFF8FFFC4FF70FD794243FB24" blockId="1.[136,1452,676,1711]" pageId="1" pageNumber="158">
<emphasis id="B9449648FF8FFFC4FF70FD794020FD40" bold="true" box="[136,295,676,702]" pageId="1" pageNumber="158">Descriptions:</emphasis>
—Annual plant, never with capitula at base, whole plant usually green, densely covered with arachnoid-tomentose hairs; main stem very long,
<quantity id="4CC8E7BFFF8FFFC4FDB3FD1543BAFD1D" box="[587,701,712,739]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="158" unit="cm" value="55.0" valueMax="60.0" valueMin="50.0">50–60 cm</quantity>
long, up to
<quantity id="4CC8E7BFFF8FFFC4FCB7FD154295FD1D" box="[847,914,712,739]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="158" unit="mm" value="4.0">4 mm</quantity>
in diameter at the base, with several branches divaricated and unequal in length, ribbed-sulcate. Leaves thin, loosely covered with arachnoid-tomentose hairs, denser along midrib. Basal and lower cauline leaves petiolate,
<quantity id="4CC8E7BFFF8FFFC4FCFCFCCD425EFCD5" box="[772,857,784,811]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="1" pageNumber="158" unit="cm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 cm</quantity>
long, pinnatisect; segments in 4–6 pairs,
<quantity id="4CC8E7BFFF8FFFC4FAC0FCCD44ABFCD5" box="[1336,1452,784,811]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="158" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
wide, linear, acute at apex, terminal segment longer. Median cauline leaves sessile, pinnatifid or sublyrate; segments in 1–3 pairs,
<quantity id="4CC8E7BFFF8FFFC4FF03FC854086FC8D" box="[251,385,856,883]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="1" pageNumber="158" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 mm</quantity>
wide, narrowly linear, acuminate at apex, terminal segment longer. Upper cauline leaves sessile, simple,
<quantity id="4CC8E7BFFF8FFFC4FF18FCA14060FC69" box="[224,359,892,919]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="158" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 mm</quantity>
wide, narrowly linear to acicular, shorter than involucre, acuminate or mucronate at apex, smooth at margins. Capitula numerous, persistent, solitary at tips of many branches, with short peduncles (
<quantity id="4CC8E7BFFF8FFFC4FB1AFC7D4466FC45" box="[1250,1377,928,955]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="1" pageNumber="158" unit="cm" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 cm</quantity>
long). Involucres ovoid, 10–12 ×
<quantity id="4CC8E7BFFF8FFFC4FE4DFC194313FC21" box="[437,532,964,991]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="158" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
. Phyllaries multiseriate, herbaceous, imbricate, yellowish-green, glabrous; median and inner ones with prominent longitudinal or elevated nerves. Appendages small, concealing part of phyllaries only, narrowly triangular, brown or dark brown,
<quantity id="4CC8E7BFFF8FFFC4FD86FBD1420FFBD9" box="[638,776,1036,1063]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="1" pageNumber="158" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
wide at base (excluding cilia), not decurrent, ciliate; cilia several, scabrous, 4–5(–6) on each side,
<quantity id="4CC8E7BFFF8FFFC4FDABFBED43C1FBB5" box="[595,710,1072,1099]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="158" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long, brownish or whitish; spine erect,
<quantity id="4CC8E7BFFF8FFFC4FB71FBED45FBFBB5" box="[1161,1276,1072,1099]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="158" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long, equal the adjacent cilia. Flowers pale pink; central florets hermaphroditic,
<quantity id="4CC8E7BFFF8FFFC4FC9DFB8942A9FB90" box="[869,942,1108,1135]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.032" metricValueMin="1.27" pageId="1" pageNumber="158" unit="in" value="6.5" valueMax="8.0" valueMin="5.0">5–8 in</quantity>
each capitulum,
<quantity id="4CC8E7BFFF8FFFC4FB88FB8945C9FB91" box="[1136,1230,1108,1135]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="1" pageNumber="158" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
long, corolla
<quantity id="4CC8E7BFFF8FFFC4FA94FB8941B3FB6D" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="1" pageNumber="158" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 mm</quantity>
long, 5-lobed; peripheral florets sterile, inconspicuous, finely dissected, not radiant, 5-lobed, limb lobe narrowly linear, ca.
<quantity id="4CC8E7BFFF8FFFC4FF06FB414047FB49" box="[254,320,1180,1207]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="158" unit="mm" value="3.0">3 mm</quantity>
long. Achenes oblong, glabrous,
<quantity id="4CC8E7BFFF8FFFC4FD44FB414228FB49" box="[700,815,1180,1207]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="1" pageNumber="158" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
long,
<quantity id="4CC8E7BFFF8FFFC4FC80FB414507FB49" box="[888,1024,1180,1207]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="1" pageNumber="158" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
wide; insertion areole lateral, yellow, ca.
<quantity id="4CC8E7BFFF8FFFC4FF57FB1D4002FB25" box="[175,261,1216,1243]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="158" unit="mm" value="0.5">0.5 mm</quantity>
long. Pappus persistent, whitish,
<quantity id="4CC8E7BFFF8FFFC4FD85FB1D4203FB25" box="[637,772,1216,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="1" pageNumber="158" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.8–3.5 mm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="C32A19D1FF8FFFC4FF45FB394487F950" pageId="1" pageNumber="158" type="etymology">
<paragraph id="8B8F4A5AFF8FFFC4FF45FB3945A3FB00" blockId="1.[136,1452,676,1711]" box="[189,1188,1252,1279]" pageId="1" pageNumber="158">
<emphasis id="B9449648FF8FFFC4FF45FB39404FFB00" bold="true" box="[189,328,1252,1278]" pageId="1" pageNumber="158">Etymology:</emphasis>
—The specific epithet refers to Karaj city, where the new species was found.
</paragraph>
<paragraph id="8B8F4A5AFF8FFFC4FF45FAD54341F9E1" blockId="1.[136,1452,676,1711]" pageId="1" pageNumber="158">
<emphasis id="B9449648FF8FFFC4FF45FAD5438CFADC" bold="true" box="[189,651,1288,1314]" pageId="1" pageNumber="158">Taxonomic and distribution remarks:</emphasis>
—
<taxonomicName id="4C3031D9FF8FFFC4FD5FFAD5429CFADC" box="[679,923,1288,1314]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="karajensis">
<emphasis id="B9449648FF8FFFC4FD5FFAD5429CFADC" box="[679,923,1288,1314]" italics="true" pageId="1" pageNumber="158">Centaurea karajensis</emphasis>
</taxonomicName>
is endemic to
<collectingRegion id="49F484B8FF8FFFC4FBB6FAD5440AFADC" box="[1102,1293,1288,1314]" country="Iran" name="Alborz" pageId="1" pageNumber="158">Alborz Province</collectingRegion>
, N
<collectingCountry id="F3270ACAFF8FFFC4FAC5FAD44469FADD" box="[1341,1390,1289,1315]" name="Iran" pageId="1" pageNumber="158">Iran</collectingCountry>
. It is an Irano-Turanian element, and grows on sandy gravel and stony plains, at elevations of
<quantity id="4CC8E7BFFF8FFFC4FB71FAF1442EFAB9" box="[1161,1321,1324,1351]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="1" pageNumber="158" unit="m" value="1800.0" valueMax="2000.0" valueMin="1600.0">1600–2000 m</quantity>
.
<taxonomicName id="4C3031D9FF8FFFC4FACEFAF141FCFA94" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="karajensis">
<emphasis id="B9449648FF8FFFC4FACEFAF141FCFA94" italics="true" pageId="1" pageNumber="158">Centaurea karajensis</emphasis>
</taxonomicName>
belongs to
<emphasis id="B9449648FF8FFFC4FE85FA8D4097FA94" box="[381,400,1360,1386]" italics="true" pageId="1" pageNumber="158">C</emphasis>
. sect.
<taxonomicName id="4C3031D9FF8FFFC4FE2FFA8D436BFA94" authorityName="Boissier" authorityYear="1849" box="[471,620,1360,1386]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="ammocyanus">
<emphasis id="B9449648FF8FFFC4FE2FFA8D436BFA94" box="[471,620,1360,1386]" italics="true" pageId="1" pageNumber="158">Ammocyanus</emphasis>
</taxonomicName>
, which includes taxa with annual habit, lyrate or more often pinnatipartite lower cauline leaves, small-sized and ovoid to fusiform involucres, with triangular and ciliate appendages ending in a mucro or short spinule, and often pink-purple flowers (
<bibRefCitation id="EFA137ABFF8FFFC4FCC0FA4542EFFA4C" author="Wagenitz, G." box="[824,1000,1432,1458]" pageId="1" pageNumber="158" pagination="313 - 420" refId="ref5454" refString="Wagenitz, G. (1980) Centaurea L. In: Rechinger, K. H. (Ed.) Flora Iranica, vol. 139 b. Akademische Druck-und Verlagsanstalt, Graz, pp. 313 - 420." type="book chapter" year="1980">Wagenitz 1980</bibRefCitation>
). It is similar to
<taxonomicName id="4C3031D9FF8FFFC4FB53FA454467FA4C" authorityName="Boissier" authorityYear="1849" box="[1195,1376,1432,1458]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="ammocyanus">
<emphasis id="B9449648FF8FFFC4FB53FA454467FA4C" box="[1195,1376,1432,1458]" italics="true" pageId="1" pageNumber="158">C. ammocyanus</emphasis>
</taxonomicName>
in the indumentum of habit, shape of involucres, color of flowers and shape of achenes (
<figureCitation id="130B56DFFF8FFFC4FBEAFA614552FA28" box="[1042,1109,1468,1494]" captionStart="FIGURE 2" captionStartId="3.[136,229,1953,1975]" captionTargetBox="[176,1412,190,1928]" captionTargetId="figure-15@3.[176,1412,190,1928]" captionTargetPageId="3" captionText="FIGURE 2. Centaurea ammocyanus Boissier (Boissier s.n., G-BOIS)." figureDoi="http://doi.org/10.5281/zenodo.14515676" httpUri="https://zenodo.org/record/14515676/files/figure.png" pageId="1" pageNumber="158">Fig. 2</figureCitation>
), but differs in some important characters (see
<tableCitation id="C6B27FE1FF8FFFC4FECBFA3D4083FA04" box="[307,388,1504,1530]" captionStart="TABLE 1" captionStartId="4.[136,224,159,183]" captionText="TABLE 1. Diagnostic morphological characters of Centaurea karajensis, C. ammocyanus and C. patula." pageId="1" pageNumber="158">Table 1</tableCitation>
). Furthermore,
<taxonomicName id="4C3031D9FF8FFFC4FDCDFA3D43CFFA04" box="[565,712,1504,1530]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="karajensis">
<emphasis id="B9449648FF8FFFC4FDCDFA3D43CFFA04" box="[565,712,1504,1530]" italics="true" pageId="1" pageNumber="158">C. karajensis</emphasis>
</taxonomicName>
occurs at elevations higher than
<quantity id="4CC8E7BFFF8FFFC4FBCDFA3D458BFA05" box="[1077,1164,1504,1531]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="158" unit="m" value="1600.0">1600 m</quantity>
, whereas
<taxonomicName id="4C3031D9FF8FFFC4FB01FA3D44ACFA04" authorityName="Boissier" authorityYear="1849" box="[1273,1451,1504,1530]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="ammocyanus">
<emphasis id="B9449648FF8FFFC4FB01FA3D44ACFA04" box="[1273,1451,1504,1530]" italics="true" pageId="1" pageNumber="158">C. ammocyanus</emphasis>
</taxonomicName>
occurs at elevations lower than
<quantity id="4CC8E7BFFF8FFFC4FE12F9D94345F9E1" box="[490,578,1540,1567]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="158" unit="m" value="1300.0">1300 m</quantity>
.
</paragraph>
<paragraph id="8B8F4A5AFF8FFFC4FF45F9F54487F950" blockId="1.[136,1452,676,1711]" pageId="1" pageNumber="158">
<taxonomicName id="4C3031D9FF8FFFC4FF45F9F540ACF9BC" box="[189,427,1576,1602]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="karajensis">
<emphasis id="B9449648FF8FFFC4FF45F9F540ACF9BC" box="[189,427,1576,1602]" italics="true" pageId="1" pageNumber="158">Centaurea karajensis</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="4C3031D9FF8FFFC4FD85F9F543E1F9BC" authorityName="Candolle" authorityYear="1838" box="[637,742,1576,1602]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="patula">
<emphasis id="B9449648FF8FFFC4FD85F9F543E1F9BC" box="[637,742,1576,1602]" italics="true" pageId="1" pageNumber="158">C. patula</emphasis>
</taxonomicName>
, from
<collectingCountry id="F3270ACAFF8FFFC4FCCFF9F44264F9BD" box="[823,867,1577,1603]" name="Iran" pageId="1" pageNumber="158">Iran</collectingCountry>
and
<collectingCountry id="F3270ACAFF8FFFC4FC66F9F542EAF9BC" box="[926,1005,1576,1602]" name="Turkey" pageId="1" pageNumber="158">Turkey</collectingCountry>
, especially because of the indumentum of leaves, size of involucres, shape of appendages and length of achenes. However, it differs from
<taxonomicName id="4C3031D9FF8FFFC4FB19F991444DF998" authorityName="Candolle" authorityYear="1838" box="[1249,1354,1612,1638]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="158" phylum="Tracheophyta" rank="species" species="patula">
<emphasis id="B9449648FF8FFFC4FB19F991444DF998" box="[1249,1354,1612,1638]" italics="true" pageId="1" pageNumber="158">C. patula</emphasis>
</taxonomicName>
in some characters like life form, ramification of stems, dissection of lower cauline leaves, shape of median cauline leaves, color of appendages, length of cilia and spine, number of central florets in each capitulum, and length of pappus (
<tableCitation id="C6B27FE1FF8FFFC4FAE6F9494477F950" box="[1310,1392,1684,1710]" captionStart="TABLE 1" captionStartId="4.[136,224,159,183]" captionText="TABLE 1. Diagnostic morphological characters of Centaurea karajensis, C. ammocyanus and C. patula." pageId="1" pageNumber="158">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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