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<document id="91E5708EBDF4A20CAC1FE9B31587B7E7" ID-DOI="10.11646/zootaxa.5419.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10791767" ID-ZooBank="39C67C6C-C84F-457E-86AD-03AC5C507979" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709794952182" checkinUser="plazi" docAuthor="Huber, Bernhard A., Meng, Guanliang, García, Jimmy Cabra &amp; Carvalho, Leonardo S." docDate="2024" docId="F5342F12116166627C86F8A7A0FAD61A" docLanguage="en" docName="zootaxa.5419.3.1.pdf" docOrigin="Zootaxa 5419 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5419.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Galapa spiniphila Huber 2020" docType="treatment" docVersion="2" lastPageNumber="330" masterDocId="090D576A1174667F7C11FFEBA04FD04B" masterDocTitle="Thriving in dry conditions: on the Neotropical spider genus Galapa (Araneae: Pholcidae)" masterLastPageNumber="347" masterPageNumber="301" pageNumber="322" updateTime="1709795501417" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5CF422DD9FF4985CD1F16F4669F65646">Thriving in dry conditions: on the Neotropical spider genus Galapa (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="176E306FD2387DDE8C540E7031AB3EFA">Meng, Guanliang</mods:namePart>
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<taxonomicName id="BA9DE5871161666A7C86F8A7A240D72D" ID-CoL="99277" authority="Huber, 2020" authorityName="Huber" authorityYear="2020" box="[151,527,1868,1895]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="spiniphila">
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<emphasis id="4FE942161161666A7C86F8A7A13FD72C" bold="true" box="[151,368,1868,1895]" italics="true" pageId="21" pageNumber="322">Galapa spiniphila</emphasis>
<bibRefCitation id="190CE3F51161666A7D67F8A6A240D72D" author="Huber, B. A. &amp; Villarreal, O." box="[374,527,1868,1895]" pageId="21" pageNumber="322" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber, 2020</bibRefCitation>
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<paragraph id="7D229E041161666A7C86F89AA1C4D7C0" blockId="21.[151,527,1868,1931]" box="[151,395,1905,1931]" pageId="21" pageNumber="322">
<figureCitation id="E5A682811161666A7C86F89AA145D7C0" box="[151,266,1905,1931]" captionStart="FIG" captionStartId="6.[151,198,1834,1858]" captionTargetBox="[151,1436,889,1810]" captionTargetId="figure-204@6.[151,1436,889,1810]" captionTargetPageId="6" captionText="FIG. 2. Galapa Huber, 2000, live specimens. AD. G. murphyi Huber sp. n.; male and female with egg sac from W of Cañas; female with egg-sac from near Playa Iguanita, and female in web (in glass vial) from near Playa Iguanita. EF. G. gabito Huber sp. n.; male and female with egg sac from Santa Marta. GH. G. spiniphila Huber, 2020; male and female with egg sac from S of Riohacha. I. G. spiniphila; male from ESE of Pueblo Bello." figureDoi="http://doi.org/10.5281/zenodo.10791771" httpUri="https://zenodo.org/record/10791771/files/figure.png" pageId="21" pageNumber="322">Figs 2GI</figureCitation>
,
<figureCitation id="E5A682811161666A7D07F89AA176D7C0" box="[278,313,1905,1931]" captionStart="FIG" captionStartId="7.[151,198,1588,1612]" captionTargetBox="[154,1432,743,1562]" captionTargetId="figure-253@7.[151,1437,740,1564]" captionTargetPageId="7" captionText="FIG. 3. Galapa spp., male chelicerae, frontal views. Note unmodified main segments (except for stridulatory files) and processes on fangs (arrows). A. G. murphyi Huber sp. n. from Costa Rica, near Playa Iguanita, ZFMK Ar 24224. B. G. spiniphila Huber, 2020 from Colombia, near Papayal, ZFMK Ar 24221. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10791775" httpUri="https://zenodo.org/record/10791775/files/figure.png" pageId="21" pageNumber="322">3B</figureCitation>
,
<figureCitation id="E5A682811161666A7D54F89AA1C4D7C0" box="[325,395,1905,1931]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStart-7="FIG" captionStartId-0="22.[151,198,956,980]" captionStartId-1="22.[151,198,1882,1906]" captionStartId-2="23.[151,198,1509,1533]" captionStartId-3="24.[151,198,1835,1859]" captionStartId-4="25.[151,198,1548,1572]" captionStartId-5="26.[151,198,1815,1839]" captionStartId-6="27.[151,198,1429,1453]" captionStartId-7="28.[151,198,1819,1843]" captionTargetBox-0="[153,1430,185,927]" captionTargetBox-1="[155,1428,1076,1844]" captionTargetBox-2="[155,1429,745,1486]" captionTargetBox-3="[151,1436,489,1811]" captionTargetBox-4="[151,1436,181,1524]" captionTargetBox-5="[158,1430,487,1792]" captionTargetBox-6="[151,1436,181,1405]" captionTargetBox-7="[151,1436,181,1795]" captionTargetId-0="figure-17@22.[151,1436,181,932]" captionTargetId-1="figure-74@22.[151,1436,1076,1857]" captionTargetId-2="figure-260@23.[151,1436,741,1486]" captionTargetId-3="figure-165@24.[151,1436,489,1811]" captionTargetId-4="figure-17@25.[151,1436,181,1524]" captionTargetId-5="figure-150@26.[158,1430,487,1792]" captionTargetId-6="figure-17@27.[151,1436,181,1405]" captionTargetId-7="figure-17@28.[151,1436,181,1795]" captionTargetPageId-0="22" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionTargetPageId-4="25" captionTargetPageId-5="26" captionTargetPageId-6="27" captionTargetPageId-7="28" captionText-0="FIG. 17. Galapa spiniphila Huber, 2020; Papayal morph from Colombia, Papayal, ZFMK Ar 24221. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Abbreviations: da, distal apophysis of genital bulb; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; va, ventral apophyses of genital bulb. Scale bar: 0.2 mm." captionText-1="FIG. 18. Galapa spiniphila Huber, 2020; Riohacha morph from Colombia, Riohacha, ZFMK Ar 24220. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Arrows point at details that differ among morphs, compare with Figs 17 and 19. Scale bar: 0.2 mm." captionText-2="FIG. 19. Galapa spiniphila Huber, 2020; Pueblo Bello morph from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Arrows point at details that differ among morphs, compare with Figs 17 and 18. Scale bar: 0.2 mm." captionText-3="FIG. 20. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Male prosoma, frontal view. B. Female ocular area, frontal-dorsal view. C. Stridulatory file on right male chelicera. D. Stridulatory pick on male palpal femur. E. Male gonopore. F. Epigynum, ventral view.Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 100 µm (A, F), 20 µm (B, E), 10 µm (C), 2 µm (D)." captionText-4="FIG. 21. Galapa spiniphila Huber, 2020, male from Colombia, near Papayal. AC. Right palp, retrolateral to dorsal views. DF. Left palp, prolateral-ventral to prolateral-dorsal views. Abbreviations: b, genital bulb; da, distal apophysis of genital bulb; dp, dorsal process of procursus; pr, procursus; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; sd, sperm duct opening; to, tarsal organ; va, ventral apophyses of genital bulb. Scale bars: 100 µm." captionText-5="FIG. 22. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A, B. Male ALS and PMS. C. Female spinnerets. D. Female ALS and PMS. E. Tip of right female tarsus 1, prolateral view. F. Tip of right male tarsus 3, prolateral view. Abbreviations:ALS, anterior lateral spinneret; PLS, posterior lateral spinneret; PMS, posterior median spinneret. Scale bars: 10 µm (A, B, DF), 20 µm (C)." captionText-6="FIG. 23. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Right male tibia 2, prolateral view, showing regular mechanoreceptors and sexually dimorphic short vertical hairs. B. Right male tibia 1, prolateral view, showing sexually dimorphic short vertical hairs (and basis of regular mechanoreceptor). C. Trichobothrium on left male metatarsus 2. D. Male palpal tarsal organ. E. Tarsal organ on left female tarsus 3. F. Cuticular plate (and basis of regular mechanoreceptor) on female femur 4. G. Rimmed pore (and basis of regular mechanoreceptor) on left female metatarsus 3. Scale bars: 10 µm (A, C, F), 2 µm (B, D, E, G)." captionText-7="FIG. 24. Galapa spiniphila Huber, 2020; female abdomens, ventral views, and cleared female genitalia, ventral and dorsal views; arrows point at distinctive median membranous processes. A, D, E. Papayal morph, from Colombia, Papayal, ZFMK Ar 24221. B, F, G. Riohacha morph, from Colombia, Riohacha, ZFMK Ar 24220. C, H, I. Pueblo Bello morph, from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10791808" figureDoi-1="http://doi.org/10.5281/zenodo.10791810" figureDoi-2="http://doi.org/10.5281/zenodo.10791812" figureDoi-3="http://doi.org/10.5281/zenodo.10791814" figureDoi-4="http://doi.org/10.5281/zenodo.10791816" figureDoi-5="http://doi.org/10.5281/zenodo.10791818" figureDoi-6="http://doi.org/10.5281/zenodo.10791820" figureDoi-7="http://doi.org/10.5281/zenodo.10791822" httpUri-0="https://zenodo.org/record/10791808/files/figure.png" httpUri-1="https://zenodo.org/record/10791810/files/figure.png" httpUri-2="https://zenodo.org/record/10791812/files/figure.png" httpUri-3="https://zenodo.org/record/10791814/files/figure.png" httpUri-4="https://zenodo.org/record/10791816/files/figure.png" httpUri-5="https://zenodo.org/record/10791818/files/figure.png" httpUri-6="https://zenodo.org/record/10791820/files/figure.png" httpUri-7="https://zenodo.org/record/10791822/files/figure.png" pageId="21" pageNumber="322">1724</figureCitation>
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<taxonomicName id="BA9DE5871161666A7C86F85EA1DBD786" ID-CoL="99277" authority="Huber" authorityName="Huber" authorityYear="2020" box="[151,404,1973,1997]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE942161161666A7C86F85EA100D786" box="[151,335,1973,1997]" italics="true" pageId="21" pageNumber="322">Galapa spiniphila</emphasis>
Huber
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in
<bibRefCitation id="190CE3F51161666A7DA4F85EA29AD786" author="Huber, B. A. &amp; Villarreal, O." box="[437,725,1973,1997]" pageId="21" pageNumber="322" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber &amp; Villarreal 2020: 58</bibRefCitation>
, figs 174175, 178188 (
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).
<emphasis id="4FE94216116366687C86FF7DA145D0FB" bold="true" box="[151,266,150,176]" pageId="23" pageNumber="324">Diagnosis</emphasis>
(amended; see
<bibRefCitation id="190CE3F5116366687DA8FF7CA280D0FA" author="Huber, B. A. &amp; Villarreal, O." box="[441,719,151,177]" pageId="23" pageNumber="324" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber &amp; Villarreal 2020</bibRefCitation>
). Males are distinguished from known congeners by presence of four distal apophyses on genital bulb (
<figureCitation id="E5A68281116366687E50FF50A2D8D09E" box="[577,663,187,213]" captionStart="FIG" captionStartId="22.[151,198,956,980]" captionTargetBox="[153,1430,185,927]" captionTargetId="figure-17@22.[151,1436,181,932]" captionTargetPageId="22" captionText="FIG. 17. Galapa spiniphila Huber, 2020; Papayal morph from Colombia, Papayal, ZFMK Ar 24221. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Abbreviations: da, distal apophysis of genital bulb; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; va, ventral apophyses of genital bulb. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791808" httpUri="https://zenodo.org/record/10791808/files/figure.png" pageId="23" pageNumber="324">Figs 17</figureCitation>
,
<figureCitation id="E5A68281116366687EB2FF50A2B9D09E" box="[675,758,187,213]" captionStart="FIG" captionStartId="25.[151,198,1548,1572]" captionTargetBox="[151,1436,181,1524]" captionTargetId="figure-17@25.[151,1436,181,1524]" captionTargetPageId="25" captionText="FIG. 21. Galapa spiniphila Huber, 2020, male from Colombia, near Papayal. AC. Right palp, retrolateral to dorsal views. DF. Left palp, prolateral-ventral to prolateral-dorsal views. Abbreviations: b, genital bulb; da, distal apophysis of genital bulb; dp, dorsal process of procursus; pr, procursus; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; sd, sperm duct opening; to, tarsal organ; va, ventral apophyses of genital bulb. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10791816" httpUri="https://zenodo.org/record/10791816/files/figure.png" pageId="23" pageNumber="324">21C, D</figureCitation>
; pair of ventral apophyses, distal apophysis, and retrolateral-dorsal apophysis) and by simple (not bifid) retrolateral process on procursus directed towards retrolateral (rather than towards dorsal). Females without sclerite in ventral wall of uterus externus (as opposed to
<taxonomicName id="BA9DE5871163666878E6FEE8A52AD156" authorityName="Huber" authorityYear="2024" box="[1271,1381,259,285]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="gabito">
<emphasis id="4FE942161163666878E6FEE8A52AD156" box="[1271,1381,259,285]" italics="true" pageId="23" pageNumber="324">G. gabito</emphasis>
</taxonomicName>
) but externally possibly indistinguishable from
<taxonomicName id="BA9DE587116366687E65FECCA2BCD10A" authorityName="Huber" authorityYear="2024" box="[628,755,295,321]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="4FE94216116366687E65FECCA2BCD10A" box="[628,755,295,321]" italics="true" pageId="23" pageNumber="324">G. murphyi</emphasis>
</taxonomicName>
; large membranous structure in internal genitalia not bilobed as in
<taxonomicName id="BA9DE587116366687CC3FEA0A172D12E" authorityName="Huber" authorityYear="2024" box="[210,317,331,357]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="gabito">
<emphasis id="4FE94216116366687CC3FEA0A172D12E" box="[210,317,331,357]" italics="true" pageId="23" pageNumber="324">G. gabito</emphasis>
</taxonomicName>
and
<taxonomicName id="BA9DE587116366687D63FEA0A1BDD12E" authorityName="Huber" authorityYear="2024" box="[370,498,331,357]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="4FE94216116366687D63FEA0A1BDD12E" box="[370,498,331,357]" italics="true" pageId="23" pageNumber="324">G. murphyi</emphasis>
</taxonomicName>
, with distinctive median membranous process (arrows in
<figureCitation id="E5A6828111636668786BFEA0A559D12E" box="[1146,1302,331,357]" captionStart="FIG" captionStartId="28.[151,198,1819,1843]" captionTargetBox="[151,1436,181,1795]" captionTargetId="figure-17@28.[151,1436,181,1795]" captionTargetPageId="28" captionText="FIG. 24. Galapa spiniphila Huber, 2020; female abdomens, ventral views, and cleared female genitalia, ventral and dorsal views; arrows point at distinctive median membranous processes. A, D, E. Papayal morph, from Colombia, Papayal, ZFMK Ar 24221. B, F, G. Riohacha morph, from Colombia, Riohacha, ZFMK Ar 24220. C, H, I. Pueblo Bello morph, from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791822" httpUri="https://zenodo.org/record/10791822/files/figure.png" pageId="23" pageNumber="324">Fig. 24E, G, I</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="29E2CE8C116266697C86FC57A5D6D457" ID-DOI="http://doi.org/10.5281/zenodo.10791808" ID-Zenodo-Dep="10791808" httpUri="https://zenodo.org/record/10791808/files/figure.png" pageId="22" pageNumber="323" startId="22.[151,198,956,980]" targetBox="[153,1430,185,927]" targetPageId="22" targetType="figure">
<paragraph id="7D229E04116266697C86FC57A5D6D457" blockId="22.[151,1436,956,1052]" pageId="22" pageNumber="323">
<emphasis id="4FE94216116266697C86FC57A0BAD39F" bold="true" box="[151,245,956,980]" pageId="22" pageNumber="323">FIG. 17.</emphasis>
<taxonomicName id="BA9DE587116266697CEEFC57A205D39F" authority="Huber, 2020" authorityName="Huber" authorityYear="2020" box="[255,586,956,980]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE94216116266697CEEFC57A1F5D39F" box="[255,442,956,980]" italics="true" pageId="22" pageNumber="323">Galapa spiniphila</emphasis>
<bibRefCitation id="190CE3F5116266697DD4FC57A205D39F" author="Huber, B. A. &amp; Villarreal, O." box="[453,586,956,980]" pageId="22" pageNumber="323" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber, 2020</bibRefCitation>
</taxonomicName>
; Papayal morph from Colombia, Papayal, ZFMK Ar 24221. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Abbreviations: da, distal apophysis of genital bulb; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; va, ventral apophyses of genital bulb. Scale bar: 0.2 mm.
</paragraph>
</caption>
<caption id="29E2CE8C116266697C86F8B1A1E7D7F1" ID-DOI="http://doi.org/10.5281/zenodo.10791810" ID-Zenodo-Dep="10791810" httpUri="https://zenodo.org/record/10791810/files/figure.png" pageId="22" pageNumber="323" startId="22.[151,198,1882,1906]" targetBox="[155,1428,1076,1844]" targetPageId="22" targetType="figure">
<paragraph id="7D229E04116266697C86F8B1A1E7D7F1" blockId="22.[151,1436,1882,1978]" pageId="22" pageNumber="323">
<emphasis id="4FE94216116266697C86F8B1A0BCD739" bold="true" box="[151,243,1882,1906]" pageId="22" pageNumber="323">FIG. 18.</emphasis>
<taxonomicName id="BA9DE587116266697CEAF8B1A20ED739" authority="Huber, 2020" authorityName="Huber" authorityYear="2020" box="[251,577,1882,1906]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE94216116266697CEAF8B1A1FAD739" box="[251,437,1882,1906]" italics="true" pageId="22" pageNumber="323">Galapa spiniphila</emphasis>
<bibRefCitation id="190CE3F5116266697DACF8B1A20ED739" author="Huber, B. A. &amp; Villarreal, O." box="[445,577,1882,1906]" pageId="22" pageNumber="323" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber, 2020</bibRefCitation>
</taxonomicName>
; Riohacha morph from Colombia, Riohacha, ZFMK Ar 24220. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Arrows point at details that differ among morphs, compare with Figs 17 and 19. Scale bar: 0.2 mm.
</paragraph>
</caption>
<subSubSection id="3587CD8F116366677CD6FE84A28AD10A" lastPageId="24" lastPageNumber="325" pageId="23" pageNumber="324" type="discussion">
<paragraph id="7D229E04116366687CD6FE84A226D2CE" blockId="23.[151,1437,150,717]" pageId="23" pageNumber="324">
<emphasis id="4FE94216116366687CD6FE84A17BD1C2" bold="true" box="[199,308,367,393]" pageId="23" pageNumber="324">Remarks</emphasis>
. The Colombian specimens below are assigned tentatively to this species because there are some minor but apparently consistent morphological differences among them and between Colombian and Venezuelan specimens (see Variation below and
<figureCitation id="E5A68281116366687DA9FE5CA275D19A" box="[440,570,439,465]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="22.[151,198,956,980]" captionStartId-1="22.[151,198,1882,1906]" captionStartId-2="23.[151,198,1509,1533]" captionTargetBox-0="[153,1430,185,927]" captionTargetBox-1="[155,1428,1076,1844]" captionTargetBox-2="[155,1429,745,1486]" captionTargetId-0="figure-17@22.[151,1436,181,932]" captionTargetId-1="figure-74@22.[151,1436,1076,1857]" captionTargetId-2="figure-260@23.[151,1436,741,1486]" captionTargetPageId-0="22" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="FIG. 17. Galapa spiniphila Huber, 2020; Papayal morph from Colombia, Papayal, ZFMK Ar 24221. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Abbreviations: da, distal apophysis of genital bulb; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; va, ventral apophyses of genital bulb. Scale bar: 0.2 mm." captionText-1="FIG. 18. Galapa spiniphila Huber, 2020; Riohacha morph from Colombia, Riohacha, ZFMK Ar 24220. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Arrows point at details that differ among morphs, compare with Figs 17 and 19. Scale bar: 0.2 mm." captionText-2="FIG. 19. Galapa spiniphila Huber, 2020; Pueblo Bello morph from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Arrows point at details that differ among morphs, compare with Figs 17 and 18. Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10791808" figureDoi-1="http://doi.org/10.5281/zenodo.10791810" figureDoi-2="http://doi.org/10.5281/zenodo.10791812" httpUri-0="https://zenodo.org/record/10791808/files/figure.png" httpUri-1="https://zenodo.org/record/10791810/files/figure.png" httpUri-2="https://zenodo.org/record/10791812/files/figure.png" pageId="23" pageNumber="324">Figs 1719</figureCitation>
). They are not described as separate species mainly because of the low CO1 distances among the sequenced Colombian specimens (3.34.8%). The distances between the Colombian specimens on one side and the topotypical Venezuelan
<taxonomicName id="BA9DE587116366687E9CFE14A36ED252" authorityName="Huber" authorityYear="2020" box="[653,801,511,537]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE94216116366687E9CFE14A36ED252" box="[653,801,511,537]" italics="true" pageId="23" pageNumber="324">G. spiniphila</emphasis>
</taxonomicName>
specimen on the other side are higher (9.410.1%), but the morphological differences between them are of the same degree as those among Colombian specimens. We thus describe the new Colombian specimens as morphs within
<taxonomicName id="BA9DE587116366687F38FDACA3F2D22A" authorityName="Huber" authorityYear="2020" box="[809,957,583,609]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE94216116366687F38FDACA3F2D22A" box="[809,957,583,609]" italics="true" pageId="23" pageNumber="324">G. spiniphila</emphasis>
</taxonomicName>
, pending further analysis. The topotypical morph is here named Paraguaná morph.
</paragraph>
<paragraph id="7D229E04116366687CD6FD64A187D286" blockId="23.[151,1437,150,717]" pageId="23" pageNumber="324">
The specimens from
<collectingRegion id="BF5950E6116366687DBAFD64A268D2E2" box="[427,551,655,681]" country="Colombia" name="Magdalena" pageId="23" pageNumber="324">Magdalena</collectingRegion>
are extremely bleached and difficult to study.They seem to share the morphology of the Pueblo Bello morph.
</paragraph>
<caption id="29E2CE8C116366687C86FA0EA262D60E" ID-DOI="http://doi.org/10.5281/zenodo.10791812" ID-Zenodo-Dep="10791812" httpUri="https://zenodo.org/record/10791812/files/figure.png" pageId="23" pageNumber="324" startId="23.[151,198,1509,1533]" targetBox="[155,1429,745,1486]" targetPageId="23" targetType="figure">
<paragraph id="7D229E04116366687C86FA0EA262D60E" blockId="23.[151,1436,1509,1606]" pageId="23" pageNumber="324">
<emphasis id="4FE94216116366687C86FA0EA0BDD5B6" bold="true" box="[151,242,1509,1533]" pageId="23" pageNumber="324">FIG. 19.</emphasis>
<taxonomicName id="BA9DE587116366687CE8FA0EA274D5B5" authority="Huber, 2020" authorityName="Huber" authorityYear="2020" box="[249,571,1509,1534]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE94216116366687CE8FA0EA1FED5B6" box="[249,433,1509,1533]" italics="true" pageId="23" pageNumber="324">Galapa spiniphila</emphasis>
<bibRefCitation id="190CE3F5116366687DA8FA0EA274D5B5" author="Huber, B. A. &amp; Villarreal, O." box="[441,571,1509,1534]" pageId="23" pageNumber="324" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber, 2020</bibRefCitation>
</taxonomicName>
; Pueblo Bello morph from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Arrows point at details that differ among morphs, compare with Figs 17 and 18. Scale bar: 0.2 mm.
</paragraph>
</caption>
<paragraph id="7D229E04116366687CD6F97CA1A6D6FA" blockId="23.[151,1437,1687,2038]" box="[199,489,1687,1713]" pageId="23" pageNumber="324">
<emphasis id="4FE94216116366687CD6F97CA1A6D6FA" bold="true" box="[199,489,1687,1713]" pageId="23" pageNumber="324">New material examined.</emphasis>
</paragraph>
<paragraph id="7D229E04116366687CD6F950A0BED72E" blockId="23.[151,1437,1687,2038]" pageId="23" pageNumber="324">
Papayal morph.
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<emphasis id="4FE94216116366687D95F950A2FDD69E" bold="true" box="[388,690,1723,1749]" pageId="23" pageNumber="324">
<collectingCountry id="058ADE94116366687D95F950A268D69E" box="[388,551,1723,1749]" name="Colombia" pageId="23" pageNumber="324">COLOMBIA</collectingCountry>
:
<emphasis id="4FE94216116366687E22F950A2FDD69E" bold="true" box="[563,690,1723,1749]" italics="true" pageId="23" pageNumber="324">
<collectingRegion id="BF5950E6116366687E22F950A2FDD69E" box="[563,690,1723,1749]" country="Colombia" name="La Guajira" pageId="23" pageNumber="324">La Guajira</collectingRegion>
</emphasis>
</emphasis>
<specimenCount id="6B9B558D116366687ED8F950A2BBD69D" box="[713,756,1723,1750]" count="6" pageId="23" pageNumber="324" type="male">6 ♂</specimenCount>
,
<specimenCount id="6B9B558D116366687EEEF950A366D69D" box="[767,809,1723,1750]" count="3" pageId="23" pageNumber="324" type="female">3 ♀</specimenCount>
(
<specimenCount id="6B9B558D116366687F26F950A3D2D69E" box="[823,925,1723,1749]" count="1" pageId="23" pageNumber="324" type="male">one male</specimenCount>
and
<specimenCount id="6B9B558D116366687FC3F957A402D69E" box="[978,1101,1723,1750]" count="1" pageId="23" pageNumber="324" type="female">one female</specimenCount>
used for
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); near
<location id="7842C8DF11636668792DF950A5D7D69E" LSID="urn:lsid:plazi:treatment:F5342F12116166627C86F8A7A0FAD61A:7842C8DF11636668792DF950A5D7D69E" box="[1340,1432,1723,1749]" country="Colombia" latitude="11.0029" longLatPrecision="7" longitude="-72.7708" name="Papayal" pageId="23" pageNumber="324" stateProvince="La Guajira">Papayal</location>
;
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,
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;
<elevation id="F6B07937116366687DA0F934A276D6B2" box="[433,569,1759,1786]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="324" unit="m" value="150.0">
<quantity id="BA6533E1116366687DA0F934A1B5D6B1" box="[433,506,1759,1786]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="324" unit="m" value="150.0">150 m</quantity>
a.s.l.
</elevation>
; semi-arid roadside;
<date id="0923B8C4116366687F3BF934A389D6B2" box="[810,966,1759,1785]" pageId="23" pageNumber="324" value="2022-09-19">
<collectingDate id="1967412C116366687F3BF934A389D6B2" box="[810,966,1759,1785]" pageId="23" pageNumber="324" value="2022-09-19">19 Sep. 2022</collectingDate>
</date>
;
<collectorName id="D068FBD2116366687FC3F934A419D6B2" box="[978,1110,1759,1785]" pageId="23" pageNumber="324">B.A. Huber</collectorName>
leg.; MUSENUV-Ar 2659
</materialsCitation>
<materialsCitation id="CDF59459116366687C86F8E8A4EAD756" box="[151,1189,1795,1822]" collectingDate="2022-09-19" collectionCode="ZFMK" collectorName="Ar" country="Colombia" elevation="150" latitude="11.0029" location="Papayal" longLatPrecision="7" longitude="-72.7708" pageId="23" pageNumber="324" specimenCount="1" specimenCount-male="1" stateProvince="La Guajira">
<specimenCount id="6B9B558D116366687C86F8E8A08AD755" box="[151,197,1795,1822]" count="1" pageId="23" pageNumber="324" type="male">1 ♂</specimenCount>
, one cleared female abdomen; same collection data as for preceding;
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<collectorName id="D068FBD2116366687828F8E8A418D756" box="[1081,1111,1795,1821]" pageId="23" pageNumber="324">Ar</collectorName>
24221
</materialsCitation>
<materialsCitation id="CDF594591163666878D1F8E8A0A2D72E" collectingDate="2022-09-19" collectionCode="ZFMK" collectorName="Col" country="Colombia" elevation="150" latitude="11.0029" location="Papayal" longLatPrecision="7" longitude="-72.7708" pageId="23" pageNumber="324" specimenCount="5" specimenCount-female="4" stateProvince="La Guajira">
<specimenCount id="6B9B558D1163666878D1F8E8A4A3D755" box="[1216,1260,1795,1822]" count="4" pageId="23" pageNumber="324" type="female">4 ♀</specimenCount>
,
<specimenCount id="6B9B558D1163666878EDF8E8A570D756" box="[1276,1343,1795,1821]" count="1" pageId="23" pageNumber="324" type="generic">1 juv.</specimenCount>
, in pure ethanol (one cleared female abdomen transferred to
<collectionCode id="1B8C06C1116366687EC7F8C3A36BD709" box="[726,804,1832,1858]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="324" type="Museum">ZFMK</collectionCode>
Ar 24221); same collection data as for preceding;
<collectionCode id="1B8C06C111636668795FF8C3A5D3D709" box="[1358,1436,1832,1858]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="324" type="Museum">ZFMK</collectionCode>
<collectorName id="D068FBD2116366687C86F8A0A08DD72E" box="[151,194,1867,1893]" pageId="23" pageNumber="324">Col</collectorName>
291
</materialsCitation>
.
</paragraph>
<paragraph id="7D229E04116366687CD6F884A530D7BE" blockId="23.[151,1437,1687,2038]" pageId="23" pageNumber="324">
<emphasis id="4FE94216116366687CD6F884A1DED7C2" bold="true" box="[199,401,1903,1930]" pageId="23" pageNumber="324">
<emphasis id="4FE94216116366687CD6F884A1C5D7C1" bold="true" box="[199,394,1903,1930]" italics="true" pageId="23" pageNumber="324">Riohacha morph</emphasis>
.
</emphasis>
<materialsCitation id="CDF59459116366687D8BF884A44BD7E6" collectingDate="2022-09-19" collectorName="B. A. Huber" country="Colombia" elevation="30" latitude="11.4848" location="Riohacha" longLatPrecision="7" longitude="-72.9051" pageId="23" pageNumber="324" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="La Guajira">
<emphasis id="4FE94216116366687D8BF884A286D7C2" bold="true" box="[410,713,1903,1929]" pageId="23" pageNumber="324">
<collectingCountry id="058ADE94116366687D8BF884A274D7C2" box="[410,571,1903,1929]" name="Colombia" pageId="23" pageNumber="324">COLOMBIA</collectingCountry>
:
<emphasis id="4FE94216116366687E59F884A286D7C2" bold="true" box="[584,713,1903,1929]" italics="true" pageId="23" pageNumber="324">
<collectingRegion id="BF5950E6116366687E59F884A286D7C2" box="[584,713,1903,1929]" country="Colombia" name="La Guajira" pageId="23" pageNumber="324">La Guajira</collectingRegion>
</emphasis>
</emphasis>
<specimenCount id="6B9B558D116366687EF5F884A35ED7C1" box="[740,785,1903,1930]" count="2" pageId="23" pageNumber="324" type="male">2 ♂</specimenCount>
,
<specimenCount id="6B9B558D116366687F0FF884A304D7C1" box="[798,843,1903,1930]" count="2" pageId="23" pageNumber="324" type="female">2 ♀</specimenCount>
;
<quantity id="BA6533E1116366687F4BF884A3DCD7C2" box="[858,915,1903,1930]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="324" unit="km" value="5.0">5 km</quantity>
S of
<location id="7842C8DF116366687FC5F884A40FD7C2" LSID="urn:lsid:plazi:treatment:F5342F12116166627C86F8A7A0FAD61A:7842C8DF116366687FC5F884A40FD7C2" box="[980,1088,1903,1929]" country="Colombia" latitude="11.4848" longLatPrecision="7" longitude="-72.9051" name="Riohacha" pageId="23" pageNumber="324" stateProvince="La Guajira">Riohacha</location>
;
<geoCoordinate id="18A9F8C311636668785DF884A488D7C2" box="[1100,1223,1903,1929]" degrees="11.4848" direction="north" orientation="latitude" pageId="23" pageNumber="324" precision="5" value="11.4848">11.4848°N</geoCoordinate>
,
<geoCoordinate id="18A9F8C31163666878C5F884A519D7C1" box="[1236,1366,1903,1930]" degrees="72.9051" direction="west" orientation="longitude" pageId="23" pageNumber="324" precision="5" value="-72.9051">72.9051°W</geoCoordinate>
;
<elevation id="F6B07937116366687972F884A082D7E6" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="324" unit="m" value="30.0">
<quantity id="BA6533E1116366687972F884A5D2D7C1" box="[1379,1437,1903,1930]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="324" unit="m" value="30.0">30 m</quantity>
a.s.l.
</elevation>
; semi-arid roadside;
<date id="0923B8C4116366687DA6F878A202D7E6" box="[439,589,1939,1965]" pageId="23" pageNumber="324" value="2022-09-19">
<collectingDate id="1967412C116366687DA6F878A202D7E6" box="[439,589,1939,1965]" pageId="23" pageNumber="324" value="2022-09-19">19 Sep. 2022</collectingDate>
</date>
;
<collectorName id="D068FBD2116366687E49F878A296D7E6" box="[600,729,1939,1965]" pageId="23" pageNumber="324">B.A. Huber</collectorName>
leg.; MUSENUV-Ar 2660
</materialsCitation>
<materialsCitation id="CDF5945911636668780CF878A2B9D79A" collectingDate="2022-09-19" collectionCode="ZFMK" collectorName="Ar" country="Colombia" elevation="30" latitude="11.4848" location="Riohacha" longLatPrecision="7" longitude="-72.9051" pageId="23" pageNumber="324" specimenCount="1" specimenCount-male="1" stateProvince="La Guajira">
<specimenCount id="6B9B558D11636668780CF878A406D7E5" box="[1053,1097,1939,1966]" count="1" pageId="23" pageNumber="324" type="male">1 ♂</specimenCount>
, one cleared female abdomen; same collection data as for preceding;
<collectionCode id="1B8C06C1116366687E2DF853A2C6D799" box="[572,649,1976,2002]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="324" type="Museum">ZFMK</collectionCode>
<collectorName id="D068FBD2116366687E9CF85CA2E5D79A" box="[653,682,1975,2001]" pageId="23" pageNumber="324">Ar</collectorName>
24220
</materialsCitation>
<materialsCitation id="CDF59459116366687F1DF85CA533D7BE" collectingDate="2022-09-19" collectionCode="ZFMK" collectorName="Col" country="Colombia" elevation="30" latitude="11.4848" location="Riohacha" longLatPrecision="7" longitude="-72.9051" pageId="23" pageNumber="324" specimenCount="4" specimenCount-female="4" stateProvince="La Guajira">
<specimenCount id="6B9B558D116366687F1DF85CA37BD799" box="[780,820,1975,2002]" count="4" pageId="23" pageNumber="324" type="female">4 ♀</specimenCount>
, in pure ethanol (one prosoma used for molecular work, one abdomen cleared and transferred to
<collectionCode id="1B8C06C1116366687E45F837A2EED7BD" box="[596,673,2012,2038]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="324" type="Museum">ZFMK</collectionCode>
Ar 24220); same collection data as for preceding;
<collectionCode id="1B8C06C11163666878C3F837A550D7BD" box="[1234,1311,2012,2038]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="324" type="Museum">ZFMK</collectionCode>
<collectorName id="D068FBD2116366687937F830A51FD7BE" box="[1318,1360,2011,2037]" pageId="23" pageNumber="324">Col</collectorName>
286
</materialsCitation>
.
</paragraph>
<paragraph id="7D229E04116C66677CD6FF7DA427D156" blockId="24.[151,1436,150,465]" pageId="24" pageNumber="325">
<emphasis id="4FE94216116C66677CD6FF7DA1F7D0FA" bold="true" box="[199,440,150,177]" pageId="24" pageNumber="325">
<emphasis id="4FE94216116C66677CD6FF7DA1FED0FA" bold="true" box="[199,433,150,177]" italics="true" pageId="24" pageNumber="325">Pueblo Bello morph</emphasis>
.
</emphasis>
<materialsCitation id="CDF59459116C66677DD1FF7DA430D09E" collectingDate="2022-09-21" collectorName="B. A. Huber" country="Colombia" elevation="240" latitude="10.3449" location="Pueblo Bello" longLatPrecision="7" longitude="-73.4349" pageId="24" pageNumber="325" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Cesar">
<emphasis id="4FE94216116C66677DD1FF7DA2FCD0FA" bold="true" box="[448,691,150,177]" pageId="24" pageNumber="325">
<collectingCountry id="058ADE94116C66677DD1FF7DA22CD0FB" box="[448,611,150,176]" name="Colombia" pageId="24" pageNumber="325">COLOMBIA</collectingCountry>
:
<emphasis id="4FE94216116C66677E61FF7CA2FCD0FA" bold="true" box="[624,691,151,177]" italics="true" pageId="24" pageNumber="325">
<collectingRegion id="BF5950E6116C66677E61FF7CA2FCD0FA" box="[624,691,151,177]" country="Colombia" name="Cesar" pageId="24" pageNumber="325">Cesar</collectingRegion>
</emphasis>
</emphasis>
<specimenCount id="6B9B558D116C66677EDDFF7CA2B6D0FA" box="[716,761,151,177]" count="3" pageId="24" pageNumber="325" type="male">3 ♂</specimenCount>
,
<specimenCount id="6B9B558D116C66677F17FF7CA37DD0FA" box="[774,818,151,177]" count="1" pageId="24" pageNumber="325" type="female">1 ♀</specimenCount>
;
<quantity id="BA6533E1116C66677F50FF7CA3C6D0FA" box="[833,905,151,177]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="24" pageNumber="325" unit="km" value="18.0">18 km</quantity>
ESE of
<location id="7842C8DF116C66677FF8FF7CA430D0FA" LSID="urn:lsid:plazi:treatment:F5342F12116166627C86F8A7A0FAD61A:7842C8DF116C66677FF8FF7CA430D0FA" box="[1001,1151,151,177]" country="Colombia" latitude="10.3449" longLatPrecision="7" longitude="-73.4349" name="Pueblo Bello" pageId="24" pageNumber="325" stateProvince="Cesar">Pueblo Bello</location>
;
<geoCoordinate id="18A9F8C3116C6667789AFF7CA547D0FA" box="[1163,1288,151,177]" degrees="10.3449" direction="north" orientation="latitude" pageId="24" pageNumber="325" precision="5" value="10.3449">10.3449°N</geoCoordinate>
,
<geoCoordinate id="18A9F8C3116C66677904FF7CA5D7D0FA" box="[1301,1432,151,177]" degrees="73.4349" direction="west" orientation="longitude" pageId="24" pageNumber="325" precision="5" value="-73.4349">73.4349°W</geoCoordinate>
;
<elevation id="F6B07937116C66677C86FF50A156D09E" box="[151,281,187,213]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="24" pageNumber="325" unit="m" value="240.0">
<quantity id="BA6533E1116C66677C86FF50A092D09E" box="[151,221,187,213]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="24" pageNumber="325" unit="m" value="240.0">240 m</quantity>
a.s.l.
</elevation>
; semi-arid hill near road;
<date id="0923B8C4116C66677E27FF50A284D09E" box="[566,715,187,213]" pageId="24" pageNumber="325" value="2022-09-21">
<collectingDate id="1967412C116C66677E27FF50A284D09E" box="[566,715,187,213]" pageId="24" pageNumber="325" value="2022-09-21">21 Sep. 2022</collectingDate>
</date>
;
<collectorName id="D068FBD2116C66677EC5FF50A31AD09E" box="[724,853,187,213]" pageId="24" pageNumber="325">B.A. Huber</collectorName>
leg.; MUSENUV-Ar 2661
</materialsCitation>
<materialsCitation id="CDF59459116C66677884FF50A337D0B2" collectingDate="2022-09-21" collectionCode="ZFMK" collectorName="Ar" country="Colombia" elevation="240" latitude="10.3449" location="Pueblo Bello" longLatPrecision="7" longitude="-73.4349" pageId="24" pageNumber="325" specimenCount="1" specimenCount-male="1" stateProvince="Cesar">
<specimenCount id="6B9B558D116C66677884FF50A48FD09E" box="[1173,1216,187,213]" count="1" pageId="24" pageNumber="325" type="male">1 ♂</specimenCount>
, one cleared female abdomen; same collection data as for preceding;
<collectionCode id="1B8C06C1116C66677EABFF34A347D0B2" box="[698,776,223,249]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="24" pageNumber="325" type="Museum">ZFMK</collectionCode>
<collectorName id="D068FBD2116C66677F1FFF34A363D0B2" box="[782,812,223,249]" pageId="24" pageNumber="325">Ar</collectorName>
24222
</materialsCitation>
<materialsCitation id="CDF59459116C66677F81FF34A42BD156" collectingDate="2022-09-21" collectionCode="ZFMK" collectorName="Col" country="Colombia" elevation="240" latitude="10.3449" location="Pueblo Bello" longLatPrecision="7" longitude="-73.4349" pageId="24" pageNumber="325" specimenCount="2" specimenCount-female="2" stateProvince="Cesar">
<specimenCount id="6B9B558D116C66677F81FF34A3F6D0B2" box="[912,953,223,249]" count="2" pageId="24" pageNumber="325" type="female">2 ♀</specimenCount>
, in pure ethanol (one abdomen cleared and transferred to
<collectionCode id="1B8C06C1116C66677D25FEE8A1CDD156" box="[308,386,259,285]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="24" pageNumber="325" type="Museum">ZFMK</collectionCode>
Ar 24222); same collection data as for preceding;
<collectionCode id="1B8C06C1116C66677FA8FEE8A448D156" box="[953,1031,259,285]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="24" pageNumber="325" type="Museum">ZFMK</collectionCode>
<collectorName id="D068FBD2116C6667781FFEE8A476D156" box="[1038,1081,259,285]" pageId="24" pageNumber="325">Col</collectorName>
307
</materialsCitation>
.
</paragraph>
<paragraph id="7D229E04116C66677CD6FECDA28AD10A" blockId="24.[151,1436,150,465]" box="[199,709,294,321]" pageId="24" pageNumber="325">
<emphasis id="4FE94216116C66677CD6FECDA183D10B" bold="true" box="[199,460,294,320]" pageId="24" pageNumber="325">Uncertain assignment</emphasis>
(see Remarks above).
</paragraph>
</subSubSection>
<subSubSection id="3587CD8F116C66677CD6FEA1A1CFD19A" pageId="24" pageNumber="325" type="materials_examined">
<paragraph id="7D229E04116C66677CD6FEA1A1CFD19A" blockId="24.[151,1436,150,465]" pageId="24" pageNumber="325">
<materialsCitation id="CDF59459116C66677CD6FEA1A3F9D1C2" collectingDate="1985-10" collectionCode="MHNG" collectorName="H. - G. Muller" country="Colombia" latitude="11.23" location="Villa Culebra" longLatPrecision="777" longitude="-74.12" municipality="Bonda" pageId="24" pageNumber="325" specimenCount="22" specimenCount-female="16" specimenCount-male="6" stateProvince="Magdalena">
<emphasis id="4FE94216116C66677CD6FEA1A1B7D12E" bold="true" box="[199,504,330,357]" pageId="24" pageNumber="325">
<collectingCountry id="058ADE94116C66677CD6FEA1A125D12F" box="[199,362,330,356]" name="Colombia" pageId="24" pageNumber="325">COLOMBIA</collectingCountry>
:
<emphasis id="4FE94216116C66677D64FEA1A1B7D12E" bold="true" box="[373,504,330,357]" italics="true" pageId="24" pageNumber="325">
<collectingRegion id="BF5950E6116C66677D64FEA1A1B7D12E" box="[373,504,330,357]" country="Colombia" name="Magdalena" pageId="24" pageNumber="325">Magdalena</collectingRegion>
</emphasis>
</emphasis>
<specimenCount id="6B9B558D116C66677E1CFEA0A277D12E" box="[525,568,331,357]" count="6" pageId="24" pageNumber="325" type="male">6 ♂</specimenCount>
,
<specimenCount id="6B9B558D116C66677E52FEA0A236D12E" box="[579,633,331,357]" count="16" pageId="24" pageNumber="325" type="female">16 ♀</specimenCount>
,
<locationDeviation id="6C908726116C66677E97FEA0A3C0D12E" box="[646,911,331,357]" location="Santa Marta" pageId="24" pageNumber="325">
<quantity id="BA6533E1116C66677E97FEA0A283D12E" box="[646,716,331,357]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="325" unit="km" value="10.0">10 km</quantity>
E of Santa Marta
</locationDeviation>
, near
<collectingMunicipality id="9D46047E116C66677FDEFEA0A454D12E" box="[975,1051,331,357]" pageId="24" pageNumber="325">Bonda</collectingMunicipality>
,
<location id="7842C8DF116C66677835FEA0A4F3D12E" LSID="urn:lsid:plazi:treatment:F5342F12116166627C86F8A7A0FAD61A:7842C8DF116C66677835FEA0A4F3D12E" box="[1060,1212,331,357]" country="Colombia" latitude="11.23" longLatPrecision="777" longitude="-74.12" municipality="Bonda" name="Villa Culebra" pageId="24" pageNumber="325" stateProvince="Magdalena">Villa Culebra</location>
;
<geoCoordinate id="18A9F8C3116C666778D6FEA0A568D12E" box="[1223,1319,331,357]" degrees="11.23" direction="north" orientation="latitude" pageId="24" pageNumber="325" precision="555" value="11.23">11.23°N</geoCoordinate>
,
<geoCoordinate id="18A9F8C3116C66677920FEA0A5D7D12E" box="[1329,1432,331,357]" degrees="74.12" direction="west" orientation="longitude" pageId="24" pageNumber="325" precision="555" value="-74.12">74.12°W</geoCoordinate>
; among hollow blocks of concrete;
<date id="0923B8C4116C66677E07FE84A2C9D1C2" box="[534,646,366,393]" pageId="24" pageNumber="325" value="1985-10">
<collectingDate id="1967412C116C66677E07FE84A2C9D1C2" box="[534,646,366,393]" pageId="24" pageNumber="325" value="1985-10">Oct. 1985</collectingDate>
</date>
;
<collectorName id="D068FBD2116C66677E80FE84A36CD1C2" box="[657,803,367,393]" pageId="24" pageNumber="325">H.-G. Müller</collectorName>
leg.;
<collectionCode id="1B8C06C1116C66677F71FE84A3F9D1C2" box="[864,950,367,393]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="24" pageNumber="325" type="Museum">MHNG</collectionCode>
</materialsCitation>
<materialsCitation id="CDF59459116C66677FDAFE84A1E7D1E6" collectingDate="1985-10" collectionCode="MHNG" collectorName="H. - G. Muller" country="Colombia" latitude="11.23" location="Villa Culebra" longLatPrecision="777" longitude="-74.12" municipality="Bonda" pageId="24" pageNumber="325" specimenCount="2" specimenCount-male="2" stateProvince="Magdalena">
<specimenCount id="6B9B558D116C66677FDAFE84A3B9D1C2" box="[971,1014,367,393]" count="2" pageId="24" pageNumber="325" type="male">2 ♂</specimenCount>
, same collection data as for preceding but
<collectingMethod id="A4DCE613116C66677CD3FE78A14AD1E6" box="[194,261,403,429]" pageId="24" pageNumber="325">pitfall</collectingMethod>
traps;
<collectionCode id="1B8C06C1116C66677D43FE78A1E7D1E6" box="[338,424,403,429]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="24" pageNumber="325" type="Museum">MHNG</collectionCode>
</materialsCitation>
<materialsCitation id="CDF59459116C66677DAEFE78A134D19A" collectingDate="1985-07-12" collectionCode="MHNG" collectorName="de Gairaca &amp; H. G. Muller" country="Colombia" latitude="11.316" location="Bahia de Gairaca" longLatPrecision="77" longitude="-74.108" municipality="Tayrona National Park" pageId="24" pageNumber="325" specimenCount="1" specimenCount-female="1" stateProvince="Magdalena">
<specimenCount id="6B9B558D116C66677DAEFE78A1A6D1E6" box="[447,489,403,429]" count="1" pageId="24" pageNumber="325" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="9D46047E116C66677DE6FE78A2B8D1E6" box="[503,759,403,429]" pageId="24" pageNumber="325">Tayrona National Park</collectingMunicipality>
,
<location id="7842C8DF116C66677F12FE78A387D1E6" LSID="urn:lsid:plazi:treatment:F5342F12116166627C86F8A7A0FAD61A:7842C8DF116C66677F12FE78A387D1E6" box="[771,968,403,429]" country="Colombia" latitude="11.316" longLatPrecision="77" longitude="-74.108" municipality="Tayrona National Park" name="Bahia de Gairaca" pageId="24" pageNumber="325" stateProvince="Magdalena">
Bahia
<collectorName id="D068FBD2116C66677F5AFE78A387D1E6" box="[843,968,403,429]" pageId="24" pageNumber="325">de Gairaca</collectorName>
</location>
;
<geoCoordinate id="18A9F8C3116C66677FC2FE78A40ED1E6" box="[979,1089,403,429]" degrees="11.316" direction="north" orientation="latitude" pageId="24" pageNumber="325" precision="55" value="11.316">11.316°N</geoCoordinate>
,
<geoCoordinate id="18A9F8C3116C6667785DFE78A48ED1E6" box="[1100,1217,403,429]" degrees="74.108" direction="west" orientation="longitude" pageId="24" pageNumber="325" precision="55" value="-74.108">74.108°W</geoCoordinate>
;
<date id="0923B8C4116C666778DCFE78A515D1E6" box="[1229,1370,402,429]" pageId="24" pageNumber="325" value="1985-07-12">
<collectingDate id="1967412C116C666778DCFE78A515D1E6" box="[1229,1370,402,429]" pageId="24" pageNumber="325" value="1985-07-12">12 Jul. 1985</collectingDate>
</date>
;
<collectorName id="D068FBD2116C66677977FE78A0ACD19A" pageId="24" pageNumber="325">H.G. Müller</collectorName>
leg.;
<collectionCode id="1B8C06C1116C66677D33FE5CA134D19A" box="[290,379,439,465]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="24" pageNumber="325" type="Museum">MHNG</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="29E2CE8C116C66677C86F8C0A21ED7FB" ID-DOI="http://doi.org/10.5281/zenodo.10791814" ID-Zenodo-Dep="10791814" httpUri="https://zenodo.org/record/10791814/files/figure.png" pageId="24" pageNumber="325" startId="24.[151,198,1835,1859]" targetBox="[151,1436,489,1811]" targetPageId="24" targetType="figure">
<paragraph id="7D229E04116C66677C86F8C0A21ED7FB" blockId="24.[151,1437,1835,1968]" pageId="24" pageNumber="325">
<emphasis id="4FE94216116C66677C86F8C0A0BED708" bold="true" box="[151,241,1835,1859]" pageId="24" pageNumber="325">FIG. 20.</emphasis>
<taxonomicName id="BA9DE587116C66677CE6F8C0A27BD70F" authority="Huber, 2020" authorityName="Huber" authorityYear="2020" box="[247,564,1835,1860]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE94216116C66677CE6F8C0A1E1D708" box="[247,430,1835,1859]" italics="true" pageId="24" pageNumber="325">Galapa spiniphila</emphasis>
<bibRefCitation id="190CE3F5116C66677DA5F8C0A27BD70F" author="Huber, B. A. &amp; Villarreal, O." box="[436,564,1835,1860]" pageId="24" pageNumber="325" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber, 2020</bibRefCitation>
</taxonomicName>
, from Colombia, near Papayal.
<emphasis id="4FE94216116C66677F64F8C0A3C1D708" bold="true" box="[885,910,1835,1859]" pageId="24" pageNumber="325">A.</emphasis>
Male prosoma, frontal view.
<emphasis id="4FE94216116C666778A9F8C7A480D70F" bold="true" box="[1208,1231,1836,1860]" pageId="24" pageNumber="325">B.</emphasis>
Female ocular area, frontal-dorsal view.
<emphasis id="4FE94216116C66677D74F8A4A132D72C" bold="true" box="[357,381,1871,1895]" pageId="24" pageNumber="325">C.</emphasis>
Stridulatory file on right male chelicera.
<emphasis id="4FE94216116C66677F34F8BBA371D723" bold="true" box="[805,830,1872,1896]" pageId="24" pageNumber="325">D.</emphasis>
Stridulatory pick on male palpal femur.
<emphasis id="4FE94216116C666778CCF8BBA4BBD723" bold="true" box="[1245,1268,1872,1896]" pageId="24" pageNumber="325">E.</emphasis>
Male gonopore.
<emphasis id="4FE94216116C66677C86F89FA0E5D7C7" bold="true" box="[151,170,1908,1932]" pageId="24" pageNumber="325">F.</emphasis>
Epigynum, ventral view. Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 100 µm (A, F), 20 µm (B, E), 10 µm (C), 2 µm (D).
</paragraph>
</caption>
<caption id="29E2CE8C116D66667C86F9E7A4BFD6DA" ID-DOI="http://doi.org/10.5281/zenodo.10791816" ID-Zenodo-Dep="10791816" httpUri="https://zenodo.org/record/10791816/files/figure.png" pageId="25" pageNumber="326" startId="25.[151,198,1548,1572]" targetBox="[151,1436,181,1524]" targetPageId="25" targetType="figure">
<paragraph id="7D229E04116D66667C86F9E7A4BFD6DA" blockId="25.[151,1437,1548,1681]" pageId="25" pageNumber="326">
<emphasis id="4FE94216116D66667C86F9E7A0BCD66F" bold="true" box="[151,243,1548,1572]" pageId="25" pageNumber="326">FIG. 21.</emphasis>
<taxonomicName id="BA9DE587116D66667CEAF9E7A20FD66F" authority="Huber, 2020" authorityName="Huber" authorityYear="2020" box="[251,576,1548,1572]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE94216116D66667CEAF9E7A1FAD66F" box="[251,437,1548,1572]" italics="true" pageId="25" pageNumber="326">Galapa spiniphila</emphasis>
<bibRefCitation id="190CE3F5116D66667DACF9E7A20FD66F" author="Huber, B. A. &amp; Villarreal, O." box="[445,576,1548,1572]" pageId="25" pageNumber="326" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber, 2020</bibRefCitation>
</taxonomicName>
, male from Colombia, near Papayal.
<emphasis id="4FE94216116D66667FD4F9E7A3B2D66F" bold="true" box="[965,1021,1548,1572]" pageId="25" pageNumber="326">AC.</emphasis>
Right palp, retrolateral to dorsal views.
<emphasis id="4FE94216116D66667C86F9DBA086D603" bold="true" box="[151,201,1584,1608]" pageId="25" pageNumber="326">DF.</emphasis>
Left palp, prolateral-ventral to prolateral-dorsal views. Abbreviations: b, genital bulb; da, distal apophysis of genital bulb; dp, dorsal process of procursus; pr, procursus; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; sd, sperm duct opening; to, tarsal organ; va, ventral apophyses of genital bulb. Scale bars: 100 µm.
</paragraph>
</caption>
<subSubSection id="3587CD8F116D66627CD6F909A3D1D642" lastPageId="29" lastPageNumber="330" pageId="25" pageNumber="326" type="description">
<paragraph id="7D229E04116D66667CD6F909A13AD6B6" blockId="25.[151,1437,1762,2038]" box="[199,373,1762,1789]" pageId="25" pageNumber="326">
<emphasis id="4FE94216116D66667CD6F909A122D6B7" bold="true" box="[199,365,1762,1788]" pageId="25" pageNumber="326">Redescription</emphasis>
.
</paragraph>
<paragraph id="7D229E04116D66667CD6F8EDA1E5D76B" blockId="25.[151,1437,1762,2038]" box="[199,426,1798,1824]" pageId="25" pageNumber="326">Male (near Papayal)</paragraph>
<paragraph id="7D229E04116D66667CD6F8C1A164D7FB" blockId="25.[151,1437,1762,2038]" pageId="25" pageNumber="326">MEASUREMENTS. Total body length 0.98, carapace width 0.43. Distance PME-PME 35 µm; diameter PME 35 µm; distance PME-ALE 10 µm; Distance AME-AME 10 µm; diameter AME 15 µm. Leg 1: 1.90 (0.56 + 0.12 + 0.50 + 0.44+ 0.28), tibia 2: 0.40, tibia 3: 0.36, tibia 4: 0.58; tibia 1 L/d: 10; diameters of leg femora 0.090, of leg tibiae: 0.050.</paragraph>
<paragraph id="7D229E04116D66667CD6F852A2CFD7BD" blockId="25.[151,1437,1762,2038]" pageId="25" pageNumber="326">COLOUR (in ethanol). Prosoma and legs pale ochre-yellow, carapace without pattern, legs without darker rings; abdomen monochromous ochre-grey.</paragraph>
<paragraph id="7D229E04116E66657CD6FF7CA248D128" blockId="26.[151,1437,151,462]" pageId="26" pageNumber="327">
BODY. Habitus as in
<figureCitation id="E5A68281116E66657DD2FF7CA278D0FA" box="[451,567,151,177]" captionStart="FIG" captionStartId="6.[151,198,1834,1858]" captionTargetBox="[151,1436,889,1810]" captionTargetId="figure-204@6.[151,1436,889,1810]" captionTargetPageId="6" captionText="FIG. 2. Galapa Huber, 2000, live specimens. AD. G. murphyi Huber sp. n.; male and female with egg sac from W of Cañas; female with egg-sac from near Playa Iguanita, and female in web (in glass vial) from near Playa Iguanita. EF. G. gabito Huber sp. n.; male and female with egg sac from Santa Marta. GH. G. spiniphila Huber, 2020; male and female with egg sac from S of Riohacha. I. G. spiniphila; male from ESE of Pueblo Bello." figureDoi="http://doi.org/10.5281/zenodo.10791771" httpUri="https://zenodo.org/record/10791771/files/figure.png" pageId="26" pageNumber="327">Fig. 2G, I</figureCitation>
. Ocular area not raised. Carapace without thoracic groove. Clypeus slightly more protruding than in female, with sclerotized rim. Sternum slightly wider than long (0.30/0.28), with pair of indistinct low anterior processes near coxae 1 (10 µm long, 20 µm diameter at basis). Abdomen globular. Gonopore apparently without epiandrous spigots (
<figureCitation id="E5A68281116E66657E42FEE9A2F6D157" box="[595,697,258,284]" captionStart="FIG" captionStartId="24.[151,198,1835,1859]" captionTargetBox="[151,1436,489,1811]" captionTargetId="figure-165@24.[151,1436,489,1811]" captionTargetPageId="24" captionText="FIG. 20. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Male prosoma, frontal view. B. Female ocular area, frontal-dorsal view. C. Stridulatory file on right male chelicera. D. Stridulatory pick on male palpal femur. E. Male gonopore. F. Epigynum, ventral view.Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 100 µm (A, F), 20 µm (B, E), 10 µm (C), 2 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10791814" httpUri="https://zenodo.org/record/10791814/files/figure.png" pageId="26" pageNumber="327">Fig. 20E</figureCitation>
). ALS with one strongly widened spigot, one long pointed spigot, and five cylindrical spigots, of which one is much wider than the others (
<figureCitation id="E5A68281116E66657FA9FECEA472D10B" box="[952,1085,293,320]" captionStart="FIG" captionStartId="26.[151,198,1815,1839]" captionTargetBox="[158,1430,487,1792]" captionTargetId="figure-150@26.[158,1430,487,1792]" captionTargetPageId="26" captionText="FIG. 22. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A, B. Male ALS and PMS. C. Female spinnerets. D. Female ALS and PMS. E. Tip of right female tarsus 1, prolateral view. F. Tip of right male tarsus 3, prolateral view. Abbreviations:ALS, anterior lateral spinneret; PLS, posterior lateral spinneret; PMS, posterior median spinneret. Scale bars: 10 µm (A, B, DF), 20 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.10791818" httpUri="https://zenodo.org/record/10791818/files/figure.png" pageId="26" pageNumber="327">Fig. 22A, B</figureCitation>
); PMS with two conical spigots (
<figureCitation id="E5A68281116E66657C8EFEA2A147D128" box="[159,264,329,355]" captionStart="FIG" captionStartId="26.[151,198,1815,1839]" captionTargetBox="[158,1430,487,1792]" captionTargetId="figure-150@26.[158,1430,487,1792]" captionTargetPageId="26" captionText="FIG. 22. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A, B. Male ALS and PMS. C. Female spinnerets. D. Female ALS and PMS. E. Tip of right female tarsus 1, prolateral view. F. Tip of right male tarsus 3, prolateral view. Abbreviations:ALS, anterior lateral spinneret; PLS, posterior lateral spinneret; PMS, posterior median spinneret. Scale bars: 10 µm (A, B, DF), 20 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.10791818" httpUri="https://zenodo.org/record/10791818/files/figure.png" pageId="26" pageNumber="327">Fig. 22A</figureCitation>
); PLS without spigots.
</paragraph>
<paragraph id="7D229E04116E66657CD6FE86A2D9D185" blockId="26.[151,1437,151,462]" pageId="26" pageNumber="327">
CHELICERAE. In general, very similar to congeners (
<figureCitation id="E5A68281116E66657F3EFE86A3C8D1CC" box="[815,903,365,391]" captionStart="FIG" captionStartId="7.[151,198,1588,1612]" captionTargetBox="[154,1432,743,1562]" captionTargetId="figure-253@7.[151,1437,740,1564]" captionTargetPageId="7" captionText="FIG. 3. Galapa spp., male chelicerae, frontal views. Note unmodified main segments (except for stridulatory files) and processes on fangs (arrows). A. G. murphyi Huber sp. n. from Costa Rica, near Playa Iguanita, ZFMK Ar 24224. B. G. spiniphila Huber, 2020 from Colombia, near Papayal, ZFMK Ar 24221. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10791775" httpUri="https://zenodo.org/record/10791775/files/figure.png" pageId="26" pageNumber="327">Fig. 3B</figureCitation>
), with curved process proximally on fang; with stridulatory files consisting of ~30 ridges, distances between ridges proximally 2.0 µm, distally 1.1 µm (
<figureCitation id="E5A68281116E66657938FE7AA5DFD1E0" box="[1321,1424,401,427]" captionStart="FIG" captionStartId="24.[151,198,1835,1859]" captionTargetBox="[151,1436,489,1811]" captionTargetId="figure-165@24.[151,1436,489,1811]" captionTargetPageId="24" captionText="FIG. 20. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Male prosoma, frontal view. B. Female ocular area, frontal-dorsal view. C. Stridulatory file on right male chelicera. D. Stridulatory pick on male palpal femur. E. Male gonopore. F. Epigynum, ventral view.Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 100 µm (A, F), 20 µm (B, E), 10 µm (C), 2 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10791814" httpUri="https://zenodo.org/record/10791814/files/figure.png" pageId="26" pageNumber="327">Fig. 20C</figureCitation>
), proximal ridges with further fine subdivision.
</paragraph>
<caption id="29E2CE8C116E66657C86F8FCA1F7D7D0" ID-DOI="http://doi.org/10.5281/zenodo.10791818" ID-Zenodo-Dep="10791818" httpUri="https://zenodo.org/record/10791818/files/figure.png" pageId="26" pageNumber="327" startId="26.[151,198,1815,1839]" targetBox="[158,1430,487,1792]" targetPageId="26" targetType="figure">
<paragraph id="7D229E04116E66657C86F8FCA1F7D7D0" blockId="26.[151,1437,1815,1948]" pageId="26" pageNumber="327">
<emphasis id="4FE94216116E66657C86F8FCA0BBD764" bold="true" box="[151,244,1815,1839]" pageId="26" pageNumber="327">FIG. 22.</emphasis>
<taxonomicName id="BA9DE587116E66657CECF8FCA20CD764" authority="Huber, 2020" authorityName="Huber" authorityYear="2020" box="[253,579,1815,1839]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE94216116E66657CECF8FCA1F8D764" box="[253,439,1815,1839]" italics="true" pageId="26" pageNumber="327">Galapa spiniphila</emphasis>
<bibRefCitation id="190CE3F5116E66657DD1F8FCA20CD764" author="Huber, B. A. &amp; Villarreal, O." box="[448,579,1815,1839]" pageId="26" pageNumber="327" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber, 2020</bibRefCitation>
</taxonomicName>
, from Colombia, near Papayal.
<emphasis id="4FE94216116E66657F82F8FCA384D764" bold="true" box="[915,971,1815,1839]" pageId="26" pageNumber="327">A, B.</emphasis>
Male ALS and PMS.
<emphasis id="4FE94216116E666578ABF8FCA49DD764" bold="true" box="[1210,1234,1815,1839]" pageId="26" pageNumber="327">C.</emphasis>
Female spinnerets.
<emphasis id="4FE94216116E66657C86F8D0A0FFD718" bold="true" box="[151,176,1851,1875]" pageId="26" pageNumber="327">D.</emphasis>
Female ALS and PMS.
<emphasis id="4FE94216116E66657DA8F8D0A19FD718" bold="true" box="[441,464,1851,1875]" pageId="26" pageNumber="327">E.</emphasis>
Tip of right female tarsus 1, prolateral view.
<emphasis id="4FE94216116E66657FAAF8D0A381D718" bold="true" box="[955,974,1851,1875]" pageId="26" pageNumber="327">F.</emphasis>
Tip of right male tarsus 3, prolateral view. Abbreviations: ALS, anterior lateral spinneret; PLS, posterior lateral spinneret; PMS, posterior median spinneret. Scale bars: 10 µm (A, B, DF), 20 µm (C).
</paragraph>
</caption>
<caption id="29E2CE8C116F66647C86FA7EA10DD629" ID-DOI="http://doi.org/10.5281/zenodo.10791820" ID-Zenodo-Dep="10791820" httpUri="https://zenodo.org/record/10791820/files/figure.png" pageId="27" pageNumber="328" startId="27.[151,198,1429,1453]" targetBox="[151,1436,181,1405]" targetPageId="27" targetType="figure">
<paragraph id="7D229E04116F66647C86FA7EA10DD629" blockId="27.[151,1437,1429,1634]" pageId="27" pageNumber="328">
<emphasis id="4FE94216116F66647C86FA7EA0BED5E6" bold="true" box="[151,241,1429,1453]" pageId="27" pageNumber="328">FIG. 23.</emphasis>
<taxonomicName id="BA9DE587116F66647CE6FA7DA27BD5E5" authority="Huber, 2020" authorityName="Huber" authorityYear="2020" box="[247,564,1430,1454]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE94216116F66647CE6FA7DA1E1D5E5" box="[247,430,1430,1454]" italics="true" pageId="27" pageNumber="328">Galapa spiniphila</emphasis>
<bibRefCitation id="190CE3F5116F66647DA5FA7DA27BD5E5" author="Huber, B. A. &amp; Villarreal, O." box="[436,564,1430,1454]" pageId="27" pageNumber="328" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber, 2020</bibRefCitation>
</taxonomicName>
, from Colombia, near Papayal.
<emphasis id="4FE94216116F66647F65FA7EA3C2D5E6" bold="true" box="[884,909,1429,1453]" pageId="27" pageNumber="328">A.</emphasis>
Right male tibia 2, prolateral view, showing regular mechanoreceptors and sexually dimorphic short vertical hairs.
<emphasis id="4FE94216116F66647F52FA51A315D599" bold="true" box="[835,858,1466,1490]" pageId="27" pageNumber="328">B.</emphasis>
Right male tibia 1, prolateral view, showing sexually dimorphic short vertical hairs (and basis of regular mechanoreceptor).
<emphasis id="4FE94216116F66647F64FA36A3C1D5BE" bold="true" box="[885,910,1501,1525]" pageId="27" pageNumber="328">C.</emphasis>
Trichobothrium on left male metatarsus 2.
<emphasis id="4FE94216116F66647958FA35A52ED5BD" bold="true" box="[1353,1377,1502,1526]" pageId="27" pageNumber="328">D.</emphasis>
Male palpal tarsal organ.
<emphasis id="4FE94216116F66647D4EF9E9A139D651" bold="true" box="[351,374,1538,1562]" pageId="27" pageNumber="328">E.</emphasis>
Tarsal organ on left female tarsus 3.
<emphasis id="4FE94216116F66647EFFF9E9A34ED651" bold="true" box="[750,769,1538,1562]" pageId="27" pageNumber="328">F.</emphasis>
Cuticular plate (and basis of regular mechanoreceptor) on female femur 4.
<emphasis id="4FE94216116F66647CE4F9CEA140D676" bold="true" box="[245,271,1573,1597]" pageId="27" pageNumber="328">G.</emphasis>
Rimmed pore (and basis of regular mechanoreceptor) on left female metatarsus 3. Scale bars: 10 µm (A, C, F), 2 µm (B, D, E, G).
</paragraph>
</caption>
<paragraph id="7D229E04116F66647CD6F95FA285D781" blockId="27.[151,1436,1716,1994]" pageId="27" pageNumber="328">
PALPS. As in
<figureCitation id="E5A68281116F66647D66F95FA181D685" box="[375,462,1716,1742]" captionStart="FIG" captionStartId="22.[151,198,956,980]" captionTargetBox="[153,1430,185,927]" captionTargetId="figure-17@22.[151,1436,181,932]" captionTargetPageId="22" captionText="FIG. 17. Galapa spiniphila Huber, 2020; Papayal morph from Colombia, Papayal, ZFMK Ar 24221. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Abbreviations: da, distal apophysis of genital bulb; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; va, ventral apophyses of genital bulb. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791808" httpUri="https://zenodo.org/record/10791808/files/figure.png" pageId="27" pageNumber="328">Fig. 17</figureCitation>
; coxa unmodified; trochanter with very short ventral protrusion; femur proximally with prolateral stridulatory pick (
<figureCitation id="E5A68281116F66647E0EF933A2C2D6B9" box="[543,653,1752,1778]" captionStart="FIG" captionStartId="24.[151,198,1835,1859]" captionTargetBox="[151,1436,489,1811]" captionTargetId="figure-165@24.[151,1436,489,1811]" captionTargetPageId="24" captionText="FIG. 20. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Male prosoma, frontal view. B. Female ocular area, frontal-dorsal view. C. Stridulatory file on right male chelicera. D. Stridulatory pick on male palpal femur. E. Male gonopore. F. Epigynum, ventral view.Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 100 µm (A, F), 20 µm (B, E), 10 µm (C), 2 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10791814" httpUri="https://zenodo.org/record/10791814/files/figure.png" pageId="27" pageNumber="328">Fig. 20D</figureCitation>
), distally slightly widened but without modification; femur-patella joints slightly shifted towards prolateral side; tibia oval, with two trichobothria; tibia-tarsus joints barely shifted towards retrolateral side; palpal tarsal organ weakly raised (
<figureCitation id="E5A68281116F66647F50F8CBA3E5D771" box="[833,938,1824,1850]" captionStart="FIG" captionStartId="27.[151,198,1429,1453]" captionTargetBox="[151,1436,181,1405]" captionTargetId="figure-17@27.[151,1436,181,1405]" captionTargetPageId="27" captionText="FIG. 23. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Right male tibia 2, prolateral view, showing regular mechanoreceptors and sexually dimorphic short vertical hairs. B. Right male tibia 1, prolateral view, showing sexually dimorphic short vertical hairs (and basis of regular mechanoreceptor). C. Trichobothrium on left male metatarsus 2. D. Male palpal tarsal organ. E. Tarsal organ on left female tarsus 3. F. Cuticular plate (and basis of regular mechanoreceptor) on female femur 4. G. Rimmed pore (and basis of regular mechanoreceptor) on left female metatarsus 3. Scale bars: 10 µm (A, C, F), 2 µm (B, D, E, G)." figureDoi="http://doi.org/10.5281/zenodo.10791820" httpUri="https://zenodo.org/record/10791820/files/figure.png" pageId="27" pageNumber="328">Fig. 23D</figureCitation>
), diameter 4.1 µm, diameter of opening 1.0 µm; procursus with pointed retrolateral-distal process directed towards retrolateral (and slightly ventral) and with dorsal process curved towards prolateral and lodged in dorsal pocket of genital bulb; genital bulb with strong distal apophysis, straight retrolateral-dorsal process, and pair of pointed ventral processes; sperm duct opening on prolateral-ventral side directly on bulb (
<figureCitation id="E5A68281116F66647E44F85BA2F1D781" box="[597,702,1968,1994]" captionStart="FIG" captionStartId="25.[151,198,1548,1572]" captionTargetBox="[151,1436,181,1524]" captionTargetId="figure-17@25.[151,1436,181,1524]" captionTargetPageId="25" captionText="FIG. 21. Galapa spiniphila Huber, 2020, male from Colombia, near Papayal. AC. Right palp, retrolateral to dorsal views. DF. Left palp, prolateral-ventral to prolateral-dorsal views. Abbreviations: b, genital bulb; da, distal apophysis of genital bulb; dp, dorsal process of procursus; pr, procursus; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; sd, sperm duct opening; to, tarsal organ; va, ventral apophyses of genital bulb. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10791816" httpUri="https://zenodo.org/record/10791816/files/figure.png" pageId="27" pageNumber="328">Fig. 21D</figureCitation>
).
</paragraph>
<caption id="29E2CE8C116866637C86F8F0A16CD789" ID-DOI="http://doi.org/10.5281/zenodo.10791822" ID-Zenodo-Dep="10791822" httpUri="https://zenodo.org/record/10791822/files/figure.png" pageId="28" pageNumber="329" startId="28.[151,198,1819,1843]" targetBox="[151,1436,181,1795]" targetPageId="28" targetType="figure">
<paragraph id="7D229E04116866637C86F8F0A16CD789" blockId="28.[151,1437,1819,1987]" pageId="28" pageNumber="329">
<emphasis id="4FE94216116866637C86F8F0A0BCD778" bold="true" box="[151,243,1819,1843]" pageId="28" pageNumber="329">FIG. 24.</emphasis>
<taxonomicName id="BA9DE587116866637CEDF8F0A20CD778" authority="Huber, 2020" authorityName="Huber" authorityYear="2020" box="[252,579,1819,1843]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="329" phylum="Arthropoda" rank="species" species="spiniphila">
<emphasis id="4FE94216116866637CEDF8F0A1F9D778" box="[252,438,1819,1843]" italics="true" pageId="28" pageNumber="329">Galapa spiniphila</emphasis>
<bibRefCitation id="190CE3F5116866637DAEF8F0A20CD778" author="Huber, B. A. &amp; Villarreal, O." box="[447,579,1819,1843]" pageId="28" pageNumber="329" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber, 2020</bibRefCitation>
</taxonomicName>
; female abdomens, ventral views, and cleared female genitalia, ventral and dorsal views; arrows point at distinctive median membranous processes.
<emphasis id="4FE94216116866637F22F8D4A3C8D71C" bold="true" box="[819,903,1855,1879]" pageId="28" pageNumber="329">A, D, E.</emphasis>
Papayal morph, from Colombia, Papayal, ZFMK Ar 24221.
<emphasis id="4FE94216116866637CF5F888A178D730" bold="true" box="[228,311,1891,1915]" pageId="28" pageNumber="329">B, F, G.</emphasis>
Riohacha morph, from Colombia, Riohacha, ZFMK Ar 24220.
<emphasis id="4FE94216116866637FD7F888A458D730" bold="true" box="[966,1047,1891,1915]" pageId="28" pageNumber="329">C, H, I.</emphasis>
Pueblo Bello morph, from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="7D229E04116966627CD6FF7CA47FD1C2" blockId="29.[151,1437,151,1617]" pageId="29" pageNumber="330">
LEGS. Without spines and curved hairs. With round cuticular plates (cf.
<figureCitation id="E5A68281116966627FF3FF7CA40DD0FA" box="[994,1090,151,177]" captionStart="FIG" captionStartId="27.[151,198,1429,1453]" captionTargetBox="[151,1436,181,1405]" captionTargetId="figure-17@27.[151,1436,181,1405]" captionTargetPageId="27" captionText="FIG. 23. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Right male tibia 2, prolateral view, showing regular mechanoreceptors and sexually dimorphic short vertical hairs. B. Right male tibia 1, prolateral view, showing sexually dimorphic short vertical hairs (and basis of regular mechanoreceptor). C. Trichobothrium on left male metatarsus 2. D. Male palpal tarsal organ. E. Tarsal organ on left female tarsus 3. F. Cuticular plate (and basis of regular mechanoreceptor) on female femur 4. G. Rimmed pore (and basis of regular mechanoreceptor) on left female metatarsus 3. Scale bars: 10 µm (A, C, F), 2 µm (B, D, E, G)." figureDoi="http://doi.org/10.5281/zenodo.10791820" httpUri="https://zenodo.org/record/10791820/files/figure.png" pageId="29" pageNumber="330">Fig. 23F</figureCitation>
; diameter 67 µm) and rimmed pores (cf.
<figureCitation id="E5A68281116966627D18FF50A13ED09E" box="[265,369,187,213]" captionStart="FIG" captionStartId="27.[151,198,1429,1453]" captionTargetBox="[151,1436,181,1405]" captionTargetId="figure-17@27.[151,1436,181,1405]" captionTargetPageId="27" captionText="FIG. 23. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Right male tibia 2, prolateral view, showing regular mechanoreceptors and sexually dimorphic short vertical hairs. B. Right male tibia 1, prolateral view, showing sexually dimorphic short vertical hairs (and basis of regular mechanoreceptor). C. Trichobothrium on left male metatarsus 2. D. Male palpal tarsal organ. E. Tarsal organ on left female tarsus 3. F. Cuticular plate (and basis of regular mechanoreceptor) on female femur 4. G. Rimmed pore (and basis of regular mechanoreceptor) on left female metatarsus 3. Scale bars: 10 µm (A, C, F), 2 µm (B, D, E, G)." figureDoi="http://doi.org/10.5281/zenodo.10791820" httpUri="https://zenodo.org/record/10791820/files/figure.png" pageId="29" pageNumber="330">Fig. 23G</figureCitation>
; outer diameter 3.5 µm, diameter of opening 0.3 µm) apparently on all leg segments. Sexually dimorphic short vertical hairs present on tibiae 1 and 2 (
<figureCitation id="E5A68281116966627F11FF34A3C9D0B2" box="[768,902,223,249]" captionStart="FIG" captionStartId="27.[151,198,1429,1453]" captionTargetBox="[151,1436,181,1405]" captionTargetId="figure-17@27.[151,1436,181,1405]" captionTargetPageId="27" captionText="FIG. 23. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Right male tibia 2, prolateral view, showing regular mechanoreceptors and sexually dimorphic short vertical hairs. B. Right male tibia 1, prolateral view, showing sexually dimorphic short vertical hairs (and basis of regular mechanoreceptor). C. Trichobothrium on left male metatarsus 2. D. Male palpal tarsal organ. E. Tarsal organ on left female tarsus 3. F. Cuticular plate (and basis of regular mechanoreceptor) on female femur 4. G. Rimmed pore (and basis of regular mechanoreceptor) on left female metatarsus 3. Scale bars: 10 µm (A, C, F), 2 µm (B, D, E, G)." figureDoi="http://doi.org/10.5281/zenodo.10791820" httpUri="https://zenodo.org/record/10791820/files/figure.png" pageId="29" pageNumber="330">Fig. 23A, B</figureCitation>
), barely visible in dissecting microscope, length ~17 µm, diameter at basis 1.1 µm. Chemoreceptive hairs similar in size (length ~12 µm, diameter at basis 1.5 µm) but with side branch (
<figureCitation id="E5A68281116966627D9FFECCA25CD10A" box="[398,531,295,321]" captionStart="FIG" captionStartId="26.[151,198,1815,1839]" captionTargetBox="[158,1430,487,1792]" captionTargetId="figure-150@26.[158,1430,487,1792]" captionTargetPageId="26" captionText="FIG. 22. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A, B. Male ALS and PMS. C. Female spinnerets. D. Female ALS and PMS. E. Tip of right female tarsus 1, prolateral view. F. Tip of right male tarsus 3, prolateral view. Abbreviations:ALS, anterior lateral spinneret; PLS, posterior lateral spinneret; PMS, posterior median spinneret. Scale bars: 10 µm (A, B, DF), 20 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.10791818" httpUri="https://zenodo.org/record/10791818/files/figure.png" pageId="29" pageNumber="330">Fig. 22E, F</figureCitation>
). Retrolateral trichobothrium of tibia 1 at 64%; prolateral trichobothrium absent on tibia 1. Tarsus 1 with five pseudosegments, fairly distinct; leg tarsal organs capsulate (cf.
<figureCitation id="E5A682811169666278B0FEA0A54AD12E" box="[1185,1285,331,357]" captionStart="FIG" captionStartId="27.[151,198,1429,1453]" captionTargetBox="[151,1436,181,1405]" captionTargetId="figure-17@27.[151,1436,181,1405]" captionTargetPageId="27" captionText="FIG. 23. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Right male tibia 2, prolateral view, showing regular mechanoreceptors and sexually dimorphic short vertical hairs. B. Right male tibia 1, prolateral view, showing sexually dimorphic short vertical hairs (and basis of regular mechanoreceptor). C. Trichobothrium on left male metatarsus 2. D. Male palpal tarsal organ. E. Tarsal organ on left female tarsus 3. F. Cuticular plate (and basis of regular mechanoreceptor) on female femur 4. G. Rimmed pore (and basis of regular mechanoreceptor) on left female metatarsus 3. Scale bars: 10 µm (A, C, F), 2 µm (B, D, E, G)." figureDoi="http://doi.org/10.5281/zenodo.10791820" httpUri="https://zenodo.org/record/10791820/files/figure.png" pageId="29" pageNumber="330">Fig. 23E</figureCitation>
; diameter 3.7 µm, diameter of opening 0.7 µm). Main tarsal claws with ~1011 tines (
<figureCitation id="E5A68281116966627FAEFE84A46CD1C2" box="[959,1059,367,393]" captionStart="FIG" captionStartId="26.[151,198,1815,1839]" captionTargetBox="[158,1430,487,1792]" captionTargetId="figure-150@26.[158,1430,487,1792]" captionTargetPageId="26" captionText="FIG. 22. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A, B. Male ALS and PMS. C. Female spinnerets. D. Female ALS and PMS. E. Tip of right female tarsus 1, prolateral view. F. Tip of right male tarsus 3, prolateral view. Abbreviations:ALS, anterior lateral spinneret; PLS, posterior lateral spinneret; PMS, posterior median spinneret. Scale bars: 10 µm (A, B, DF), 20 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.10791818" httpUri="https://zenodo.org/record/10791818/files/figure.png" pageId="29" pageNumber="330">Fig. 22F</figureCitation>
).
</paragraph>
<paragraph id="7D229E04116966627CD6FE78A18CD1E6" blockId="29.[151,1437,151,1617]" box="[199,451,403,429]" pageId="29" pageNumber="330">Female (near Papayal)</paragraph>
<paragraph id="7D229E04116966627CD6FE5CA49ED2CE" blockId="29.[151,1437,151,1617]" pageId="29" pageNumber="330">
In general, similar to male (
<figureCitation id="E5A68281116966627DEDFE5CA21CD19A" box="[508,595,439,465]" captionStart="FIG" captionStartId="6.[151,198,1834,1858]" captionTargetBox="[151,1436,889,1810]" captionTargetId="figure-204@6.[151,1436,889,1810]" captionTargetPageId="6" captionText="FIG. 2. Galapa Huber, 2000, live specimens. AD. G. murphyi Huber sp. n.; male and female with egg sac from W of Cañas; female with egg-sac from near Playa Iguanita, and female in web (in glass vial) from near Playa Iguanita. EF. G. gabito Huber sp. n.; male and female with egg sac from Santa Marta. GH. G. spiniphila Huber, 2020; male and female with egg sac from S of Riohacha. I. G. spiniphila; male from ESE of Pueblo Bello." figureDoi="http://doi.org/10.5281/zenodo.10791771" httpUri="https://zenodo.org/record/10791771/files/figure.png" pageId="29" pageNumber="330">Fig. 2H</figureCitation>
) but sternum without pair of anterior humps, chelicerae apparently without stridulatory files, cheliceral fangs unmodified, clypeus less protruding and without sclerotized rim. Tibia
<quantity id="BA6533E111696662792CFE30A526D1BE" box="[1341,1385,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="29" pageNumber="330" unit="in" value="1.0">1 in</quantity>
<specimenCount id="6B9B558D116966627960FE30A0BCD252" count="5" pageId="29" pageNumber="330" type="female">five females</specimenCount>
: 0.40
<emphasis id="4FE94216116966627D23FE14A10FD253" box="[306,320,511,536]" italics="true" pageId="29" pageNumber="330"></emphasis>
0.44 (mean 0.42). Spinnerets as in male (
<figureCitation id="E5A68281116966627F39FE14A3F6D252" box="[808,953,511,537]" captionStart="FIG" captionStartId="26.[151,198,1815,1839]" captionTargetBox="[158,1430,487,1792]" captionTargetId="figure-150@26.[158,1430,487,1792]" captionTargetPageId="26" captionText="FIG. 22. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A, B. Male ALS and PMS. C. Female spinnerets. D. Female ALS and PMS. E. Tip of right female tarsus 1, prolateral view. F. Tip of right male tarsus 3, prolateral view. Abbreviations:ALS, anterior lateral spinneret; PLS, posterior lateral spinneret; PMS, posterior median spinneret. Scale bars: 10 µm (A, B, DF), 20 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.10791818" httpUri="https://zenodo.org/record/10791818/files/figure.png" pageId="29" pageNumber="330">Fig. 22C, D</figureCitation>
). Epigynum (
<figureCitation id="E5A68281116966627849FE14A48ED252" box="[1112,1217,511,537]" captionStart="FIG" captionStartId="24.[151,198,1835,1859]" captionTargetBox="[151,1436,489,1811]" captionTargetId="figure-165@24.[151,1436,489,1811]" captionTargetPageId="24" captionText="FIG. 20. Galapa spiniphila Huber, 2020, from Colombia, near Papayal. A. Male prosoma, frontal view. B. Female ocular area, frontal-dorsal view. C. Stridulatory file on right male chelicera. D. Stridulatory pick on male palpal femur. E. Male gonopore. F. Epigynum, ventral view.Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 100 µm (A, F), 20 µm (B, E), 10 µm (C), 2 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10791814" httpUri="https://zenodo.org/record/10791814/files/figure.png" pageId="29" pageNumber="330">Figs 20F</figureCitation>
,
<figureCitation id="E5A682811169666278C1FE14A54DD252" box="[1232,1282,511,537]" captionStart="FIG" captionStartId="28.[151,198,1819,1843]" captionTargetBox="[151,1436,181,1795]" captionTargetId="figure-17@28.[151,1436,181,1795]" captionTargetPageId="28" captionText="FIG. 24. Galapa spiniphila Huber, 2020; female abdomens, ventral views, and cleared female genitalia, ventral and dorsal views; arrows point at distinctive median membranous processes. A, D, E. Papayal morph, from Colombia, Papayal, ZFMK Ar 24221. B, F, G. Riohacha morph, from Colombia, Riohacha, ZFMK Ar 24220. C, H, I. Pueblo Bello morph, from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791822" httpUri="https://zenodo.org/record/10791822/files/figure.png" pageId="29" pageNumber="330">24A</figureCitation>
) with simple semicircular anterior plate, weakly protruding, posteriorly indented medially; posterior plate short and simple, with indistinct median division. Internal genitalia (
<figureCitation id="E5A68281116966627E9EFDACA35ED22A" box="[655,785,583,609]" captionStart="FIG" captionStartId="28.[151,198,1819,1843]" captionTargetBox="[151,1436,181,1795]" captionTargetId="figure-17@28.[151,1436,181,1795]" captionTargetPageId="28" captionText="FIG. 24. Galapa spiniphila Huber, 2020; female abdomens, ventral views, and cleared female genitalia, ventral and dorsal views; arrows point at distinctive median membranous processes. A, D, E. Papayal morph, from Colombia, Papayal, ZFMK Ar 24221. B, F, G. Riohacha morph, from Colombia, Riohacha, ZFMK Ar 24220. C, H, I. Pueblo Bello morph, from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791822" httpUri="https://zenodo.org/record/10791822/files/figure.png" pageId="29" pageNumber="330">Fig. 24D, E</figureCitation>
) very simple, apparently without pore plates; membranous structure with long median process (twisted in
<figureCitation id="E5A68281116966627EB5FD80A365D2CE" box="[676,810,619,645]" captionStart="FIG" captionStartId="28.[151,198,1819,1843]" captionTargetBox="[151,1436,181,1795]" captionTargetId="figure-17@28.[151,1436,181,1795]" captionTargetPageId="28" captionText="FIG. 24. Galapa spiniphila Huber, 2020; female abdomens, ventral views, and cleared female genitalia, ventral and dorsal views; arrows point at distinctive median membranous processes. A, D, E. Papayal morph, from Colombia, Papayal, ZFMK Ar 24221. B, F, G. Riohacha morph, from Colombia, Riohacha, ZFMK Ar 24220. C, H, I. Pueblo Bello morph, from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791822" httpUri="https://zenodo.org/record/10791822/files/figure.png" pageId="29" pageNumber="330">Fig. 24D, E</figureCitation>
), covered by posterior epigynal plate.
</paragraph>
<paragraph id="7D229E04116966627CD6FD65A1F7D382" blockId="29.[151,1437,151,1617]" pageId="29" pageNumber="330">
<emphasis id="4FE94216116966627CD6FD65A177D2E3" bold="true" box="[199,312,654,680]" pageId="29" pageNumber="330">Variation</emphasis>
. Males from different localities show variation in at least three palpal structures (
<figureCitation id="E5A682811169666278F0FD64A529D2E2" box="[1249,1382,655,681]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="22.[151,198,956,980]" captionStartId-1="22.[151,198,1882,1906]" captionStartId-2="23.[151,198,1509,1533]" captionTargetBox-0="[153,1430,185,927]" captionTargetBox-1="[155,1428,1076,1844]" captionTargetBox-2="[155,1429,745,1486]" captionTargetId-0="figure-17@22.[151,1436,181,932]" captionTargetId-1="figure-74@22.[151,1436,1076,1857]" captionTargetId-2="figure-260@23.[151,1436,741,1486]" captionTargetPageId-0="22" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="FIG. 17. Galapa spiniphila Huber, 2020; Papayal morph from Colombia, Papayal, ZFMK Ar 24221. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Abbreviations: da, distal apophysis of genital bulb; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; va, ventral apophyses of genital bulb. Scale bar: 0.2 mm." captionText-1="FIG. 18. Galapa spiniphila Huber, 2020; Riohacha morph from Colombia, Riohacha, ZFMK Ar 24220. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Arrows point at details that differ among morphs, compare with Figs 17 and 19. Scale bar: 0.2 mm." captionText-2="FIG. 19. Galapa spiniphila Huber, 2020; Pueblo Bello morph from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Arrows point at details that differ among morphs, compare with Figs 17 and 18. Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10791808" figureDoi-1="http://doi.org/10.5281/zenodo.10791810" figureDoi-2="http://doi.org/10.5281/zenodo.10791812" httpUri-0="https://zenodo.org/record/10791808/files/figure.png" httpUri-1="https://zenodo.org/record/10791810/files/figure.png" httpUri-2="https://zenodo.org/record/10791812/files/figure.png" pageId="29" pageNumber="330">Figs 1719</figureCitation>
): (1) retrolateral-dorsal bulbal process, (2) distal bulbal apophysis, and (3) retrolateral-distal process of procursus. The retrolateral-dorsal bulbal apophysis (ra in
<figureCitation id="E5A68281116966627E7BFD3CA2F6D2BA" box="[618,697,727,753]" captionStart="FIG" captionStartId="22.[151,198,956,980]" captionTargetBox="[153,1430,185,927]" captionTargetId="figure-17@22.[151,1436,181,932]" captionTargetPageId="22" captionText="FIG. 17. Galapa spiniphila Huber, 2020; Papayal morph from Colombia, Papayal, ZFMK Ar 24221. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Abbreviations: da, distal apophysis of genital bulb; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; va, ventral apophyses of genital bulb. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791808" httpUri="https://zenodo.org/record/10791808/files/figure.png" pageId="29" pageNumber="330">Fig. 17</figureCitation>
) is curved towards the distal bulbal apophysis in the Riohacha and Pueblo Bello morphs, but straight in the Paraguaná and Papayal morphs. In the Riohacha morph, this apophysis is curved towards the proximal section of the distal bulbal apophysis, while in the Pueblo Bello morph it is curved towards the mid-section of this apophysis. The distal bulbal apophysis (da in
<figureCitation id="E5A68281116966627805FCA8A429D316" box="[1044,1126,835,861]" captionStart="FIG" captionStartId="22.[151,198,956,980]" captionTargetBox="[153,1430,185,927]" captionTargetId="figure-17@22.[151,1436,181,932]" captionTargetPageId="22" captionText="FIG. 17. Galapa spiniphila Huber, 2020; Papayal morph from Colombia, Papayal, ZFMK Ar 24221. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Abbreviations: da, distal apophysis of genital bulb; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; va, ventral apophyses of genital bulb. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791808" httpUri="https://zenodo.org/record/10791808/files/figure.png" pageId="29" pageNumber="330">Fig. 17</figureCitation>
) is long and slender in the Paraguaná and Riohacha morphs, wider and stronger in the Papayal morph, shorter in the Pueblo Bello morph. The retrolateral-distal process of the procursus (rp in
<figureCitation id="E5A68281116966627EACFC60A341D3EE" box="[701,782,907,933]" captionStart="FIG" captionStartId="22.[151,198,956,980]" captionTargetBox="[153,1430,185,927]" captionTargetId="figure-17@22.[151,1436,181,932]" captionTargetPageId="22" captionText="FIG. 17. Galapa spiniphila Huber, 2020; Papayal morph from Colombia, Papayal, ZFMK Ar 24221. Left male palp in prolateral (A), dorsal (B), and retrolateral (C) views. Abbreviations: da, distal apophysis of genital bulb; ra, retrolateral-dorsal apophysis of genital bulb; rp, retrolateral-distal process of procursus; va, ventral apophyses of genital bulb. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791808" httpUri="https://zenodo.org/record/10791808/files/figure.png" pageId="29" pageNumber="330">Fig. 17</figureCitation>
) is wider (in lateral view) in the Pueblo Bello morph than in the three other morphs.
</paragraph>
<paragraph id="7D229E04116966627CD6FC38A3B4D48E" blockId="29.[151,1437,151,1617]" pageId="29" pageNumber="330">
Females of the four morphs are apparently indistinguishable. The posterior margin of the anterior epigynal plate is straight in the Pueblo Bello morph (
<figureCitation id="E5A68281116966627E9BFC1CA2A0D45A" box="[650,751,1015,1041]" captionStart="FIG" captionStartId="28.[151,198,1819,1843]" captionTargetBox="[151,1436,181,1795]" captionTargetId="figure-17@28.[151,1436,181,1795]" captionTargetPageId="28" captionText="FIG. 24. Galapa spiniphila Huber, 2020; female abdomens, ventral views, and cleared female genitalia, ventral and dorsal views; arrows point at distinctive median membranous processes. A, D, E. Papayal morph, from Colombia, Papayal, ZFMK Ar 24221. B, F, G. Riohacha morph, from Colombia, Riohacha, ZFMK Ar 24220. C, H, I. Pueblo Bello morph, from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791822" httpUri="https://zenodo.org/record/10791822/files/figure.png" pageId="29" pageNumber="330">Fig. 24C</figureCitation>
) but undulating in the other morphs (
<figureCitation id="E5A6828111696662788BFC1CA56BD45A" box="[1178,1316,1015,1041]" captionStart="FIG" captionStartId="28.[151,198,1819,1843]" captionTargetBox="[151,1436,181,1795]" captionTargetId="figure-17@28.[151,1436,181,1795]" captionTargetPageId="28" captionText="FIG. 24. Galapa spiniphila Huber, 2020; female abdomens, ventral views, and cleared female genitalia, ventral and dorsal views; arrows point at distinctive median membranous processes. A, D, E. Papayal morph, from Colombia, Papayal, ZFMK Ar 24221. B, F, G. Riohacha morph, from Colombia, Riohacha, ZFMK Ar 24220. C, H, I. Pueblo Bello morph, from Colombia, ESE of Pueblo Bello, ZFMK Ar 24222. Abbreviations: aep, anterior (main) epigynal plate; pep, posterior epigynal plate. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10791822" httpUri="https://zenodo.org/record/10791822/files/figure.png" pageId="29" pageNumber="330">Fig. 24A, B</figureCitation>
; cf.
<bibRefCitation id="190CE3F5116966627947FC1CA12BD47E" author="Huber, B. A. &amp; Villarreal, O." pageId="29" pageNumber="330" pagination="1 - 317" refId="ref19980" refString="Huber, B. A. &amp; Villarreal, O. (2020) On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy, 718, 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101" type="journal article" year="2020">Huber &amp; Villarreal 2020</bibRefCitation>
: figs 183, 186); however, this seems to vary within populations and is also slightly dependent on the angle of view and on the degree of opening of the epigynal cleft. The median posterior process in the internal genitalia appears variably long (e.g. longer in the Paraguaná morph than in the Pueblo Bello morph) but this observation rests on a very small sample of cleared specimens and may partly be an artifact (e.g., the process appears short in the Papayal morph but apparently only because it is twisted).
</paragraph>
<paragraph id="7D229E04116966627CD6FB24A31FD546" blockId="29.[151,1437,151,1617]" pageId="29" pageNumber="330">
Tibia 1 lengths among the four morphs are widely overlapping and thus pooled here. Tibia
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<specimenCount id="6B9B558D11696662793CFB24A5D6D4A2" box="[1325,1433,1231,1257]" count="17" pageId="29" pageNumber="330" type="male">17 males</specimenCount>
: 0.470.54 (mean 0.50); in
<specimenCount id="6B9B558D116966627DD0FB18A20FD546" box="[449,576,1267,1293]" count="29" pageId="29" pageNumber="330" type="female">29 females</specimenCount>
: 0.400.50 (mean 0.44).
</paragraph>
<paragraph id="7D229E04116966627CD6FAFCA3D1D642" blockId="29.[151,1437,151,1617]" pageId="29" pageNumber="330">
<emphasis id="4FE94216116966627CD6FAFCA1CED57B" bold="true" box="[199,385,1302,1329]" pageId="29" pageNumber="330">Natural history</emphasis>
. At Riohacha, all specimens were beaten out of dead cactus branches lying on the ground in a degraded roadside environment with bushes and a few small trees. At Papayal, most specimens were also beaten out of dead cacti lying on the ground in a similar roadside environment (
<figureCitation id="E5A68281116966627FF4FAB4A40ED532" box="[997,1089,1374,1401]" captionStart="FIG" captionStartId="9.[151,198,1684,1708]" captionTargetBox="[151,1436,181,1660]" captionTargetId="figure-17@9.[151,1436,181,1660]" captionTargetPageId="9" captionText="FIG. 5. Typical habitats of Galapa spp. AB. Galápagos, Santa Cruz, near Garrapatero beach and near Canal de Itabaca (G. bella). C. Colombia, Magdalena, near Santa Marta (G. gabito). D. Colombia, La Guajira, near Papayal (G. spiniphila). EF. Costa Rica, Guanacaste, NW of Cañas and S of La Cruz (G. murphyi). Photos BAH." figureDoi="http://doi.org/10.5281/zenodo.10791781" httpUri="https://zenodo.org/record/10791781/files/figure.png" pageId="29" pageNumber="330">Fig. 5D</figureCitation>
). At Pueblo Bello, the spiders were found in a low, dry forest on a roadside hill. They were beaten from old dry branches lying on the ground and hollowed by termites. At all three localities, they shared the used microhabitats with another species of
<taxonomicName id="BA9DE587116966627939FA4CA5D7D58A" authorityName="Simon" authorityYear="1890" box="[1320,1432,1447,1473]" class="Arachnida" family="Pholcidae" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
, an undescribed representative of
<taxonomicName id="BA9DE587116966627E03FA20A2E5D5AE" authorityName="Mello-Leitao" authorityYear="1944" box="[530,682,1483,1509]" class="Arachnida" family="Pholcidae" genus="Ibotyporanga" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="genus">
<emphasis id="4FE94216116966627E03FA20A2E5D5AE" box="[530,682,1483,1509]" italics="true" pageId="29" pageNumber="330">Ibotyporanga</emphasis>
</taxonomicName>
; at Riohacha, also with
<taxonomicName id="BA9DE587116966627FD4FA20A511D5AE" authority="(Simon, 1893)" authorityName="Huber &amp; Acurio" authorityYear="2022" baseAuthorityName="Simon" baseAuthorityYear="1893" box="[965,1374,1483,1509]" class="Arachnida" family="Pholcidae" genus="Modisimus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="culicinus">
<emphasis id="4FE94216116966627FD4FA20A4E1D5AE" box="[965,1198,1483,1509]" italics="true" pageId="29" pageNumber="330">Modisimus culicinus</emphasis>
(Simon, 1893)
</taxonomicName>
. Two egg-sacs contained
<specimenCount id="6B9B558D116966627D63FA04A182D642" box="[370,461,1519,1545]" count="6" pageId="29" pageNumber="330" type="egg">six eggs</specimenCount>
each, with an egg diameter of 0.380.40.
</paragraph>
</subSubSection>
<subSubSection id="3587CD8F116966627CD6F9F9A0FAD61A" pageId="29" pageNumber="330" type="distribution">
<paragraph id="7D229E04116966627CD6F9F9A0FAD61A" blockId="29.[151,1437,151,1617]" pageId="29" pageNumber="330">
<emphasis id="4FE94216116966627CD6F9F9A116D667" bold="true" box="[199,345,1554,1580]" pageId="29" pageNumber="330">Distribution</emphasis>
. This species seems to be widespread in north-western
<collectingCountry id="058ADE94116966627FC8F9F8A403D666" box="[985,1100,1555,1581]" name="Venezuela" pageId="29" pageNumber="330">Venezuela</collectingCountry>
and northern
<collectingCountry id="058ADE941169666278FEF9F8A512D666" box="[1263,1373,1555,1581]" name="Colombia" pageId="29" pageNumber="330">Colombia</collectingCountry>
(
<figureCitation id="E5A6828111696662797EF9F8A0E7D61A" captionStart="FIG" captionStartId="8.[151,198,1986,2010]" captionTargetBox="[151,1435,849,1963]" captionTargetId="figure-334@8.[151,1435,849,1963]" captionTargetPageId="8" captionText="FIG. 4. Known distribution of Galapa Huber, 2000." figureDoi="http://doi.org/10.5281/zenodo.10791777" httpUri="https://zenodo.org/record/10791777/files/figure.png" pageId="29" pageNumber="330">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>