263 lines
34 KiB
XML
263 lines
34 KiB
XML
<document id="03E6A5D257E86CF95AFCF54524CB7B48" ID-CLB-Dataset="30862" ID-DOI="10.11646/zootaxa.4377.2.9" ID-GBIF-Dataset="a0cd032c-e3a1-4f69-b7c3-0a20554bb228" ID-ISSN="1175-5326" ID-Zenodo-Dep="1163969" ID-ZooBank="B0EDD476-B035-4DE6-81FA-408CE915DE10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517471985360" checkinUser="plazi" docAuthor="Sehnal, Richard" docDate="2018" docId="3C0087AB29114E1AC5A1FB10F648FD35" docLanguage="en" docName="zootaxa.4377.2.9.pdf" docOrigin="Zootaxa 4377 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Brachylepis turmensis Sehnal 2018, new species" docType="treatment" docVersion="7" lastPageNumber="293" masterDocId="C039FFD329114E18C536FFC6F521FFEC" masterDocTitle="A new species of Brachylepis Kolbe, 1894 from Tanzania (Scarabaeidae: Melolonthinae: Leucopholini)" masterLastPageNumber="295" masterPageNumber="291" pageNumber="291" updateTime="1698518179028" updateUser="ExternalLinkService">
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<mods:mods id="F9B15ECF5F9665FADBD7FFEF5BB28FB9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DDB5535CDF4E31FC2D4B18267551BDA0">
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<mods:title id="018F4AA3D9015E6BD3E47C331B5C0376">A new species of Brachylepis Kolbe, 1894 from Tanzania (Scarabaeidae: Melolonthinae: Leucopholini)</mods:title>
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</mods:titleInfo>
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<mods:name id="229B956DABE32EEAF9BAFBD0C770D369" type="personal">
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<mods:role id="6CC1BA9254F2E174905B9C4BDA7B12A5">
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<mods:roleTerm id="D08FAAB3BAE378E11ECDC14A2DA75BFE">Author</mods:roleTerm>
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<mods:namePart id="FE0D616C8DEBC5D6F9B4C05672D88B09">Sehnal, Richard</mods:namePart>
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<mods:typeOfResource id="58E1BC4F192BB2419AE18BD5A41B6502">text</mods:typeOfResource>
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<mods:titleInfo id="848748A28C0EC690A605B3555EFF515A">
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<mods:title id="448EBBD1ABD4C311BEE3489E463B083D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="466116FD5B3DB3EFA4FDA9E1B748A1DE">
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<mods:date id="663E4C5C1F7345810CBA5E33173B4EB9">2018</mods:date>
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<mods:detail id="E06FB531A8C9CB022A7AA57BF2BA98D4" type="pubDate">
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<mods:number id="3AA6193AC8B4E696AB2942159CC396F4">2018-02-01</mods:number>
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<mods:number id="D07BB0CBE77A4990CCDA1E769B040675">4377</mods:number>
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<mods:detail id="BCA5779D25839AD14C4B768FA1B9229F" type="issue">
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<mods:number id="D7CF0137DA63F523B2E898DF0E7A1D8E">2</mods:number>
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<mods:start id="71009C7C2E73E9187D356C43BFFA9D63">291</mods:start>
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<mods:identifier id="E491F4E3D4EDA293BFC0DEC45C53A40D" type="DOI">10.11646/zootaxa.4377.2.9</mods:identifier>
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<treatment id="3C0087AB29114E1AC5A1FB10F648FD35" ID-DOI="http://doi.org/10.5281/zenodo.5987724" ID-GBIF-Taxon="150777173" ID-Zenodo-Dep="5987724" LSID="urn:lsid:plazi:treatment:3C0087AB29114E1AC5A1FB10F648FD35" httpUri="http://treatment.plazi.org/id/3C0087AB29114E1AC5A1FB10F648FD35" lastPageId="2" lastPageNumber="293" pageId="0" pageNumber="291">
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<subSubSection id="FCB3653629114E18C5A1FB10F140FB1C" box="[151,1121,1238,1264]" pageId="0" pageNumber="291" type="nomenclature">
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<paragraph id="B41636BD29114E18C5A1FB10F140FB1C" blockId="0.[151,1436,1238,1989]" box="[151,1121,1238,1264]" pageId="0" pageNumber="291">
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<heading id="EF5E81D129114E18C5A1FB10F140FB1C" bold="true" box="[151,1121,1238,1264]" fontSize="11" level="1" pageId="0" pageNumber="291" reason="1">
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<emphasis id="86DDEAAF29114E18C5A1FB10F140FB1C" bold="true" box="[151,1121,1238,1264]" pageId="0" pageNumber="291">
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<taxonomicName id="73A94D3E29114E18C5A1FB10F722FB1C" ID-CoL="MWTW" authority="Sehnal" authorityName="Sehnal" authorityYear="2018" box="[151,515,1238,1264]" class="Insecta" family="Scarabaeidae" genus="Brachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="turmensis" status="sp. nov.">
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<emphasis id="86DDEAAF29114E18C5A1FB10F485FB1C" bold="true" box="[151,420,1238,1264]" italics="true" pageId="0" pageNumber="291">Brachylepis turmensis</emphasis>
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Sehnal
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</taxonomicName>
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,
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<taxonomicNameLabel id="9DEE57D429114E18C739FB10F7BEFB1C" box="[527,671,1238,1264]" pageId="0" pageNumber="291" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="2C922A3829114E18C798FB10F664FB1C" box="[686,837,1238,1264]" captionStart="FIGURES 1" captionStartId="1.[151,258,1608,1630]" captionTargetBox="[184,1421,197,1572]" captionTargetId="figure@1.[150,1436,191,1587]" captionTargetPageId="1" captionText="FIGURES 1A–F. Habitus (A–C) and detail of setation on pronotum (D–F), dorsal view. A, D, Brachylepis elephas, male, Tanzania, Voi;B, E, Brachylepis werneri, male, Tanzania, Iringa; C, F, Brachylepis turmensis new species, holotype male.Scale bar = 1 cm (for habitus); scale bar = 2 mm (for detail of pronotum)." httpUri="https://zenodo.org/record/1163971/files/figure.png" pageId="0" pageNumber="291">Figs. 1C, 1F</figureCitation>
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,
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<figureCitation id="2C922A3829114E18C664FB10F6BFFB1C" box="[850,926,1238,1264]" captionStart="FIGURES 2" captionStartId="2.[151,258,1725,1747]" captionTargetBox="[160,1418,797,1684]" captionTargetId="figure@2.[151,1435,781,1705]" captionTargetPageId="2" captionText="FIGURES 2A–I. Aedeagus in dorsal (A, D, G) and lateral (B, E, H) views; apex of aedeagus (C, F, I). A–C, Brachylepis elephas, Tanzania, Voi;D–F,Brachylepis werneri, Tanzania,Iringa; G–I, Brachylepisturmensis new species, holotype.Scale bar= 1 mm." httpUri="https://zenodo.org/record/1163973/files/figure.png" pageId="0" pageNumber="291">2G–H</figureCitation>
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,
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<figureCitation id="2C922A3829114E18C69DFB10F107FB1C" box="[939,1062,1238,1264]" captionStart="FIGURES 3" captionStartId="3.[151,258,1061,1083]" captionTargetBox="[159,1415,198,1026]" captionTargetId="figure@3.[151,1435,191,1040]" captionTargetPageId="3" captionText="FIGURES 3A–I. Mesotibia (A–C), ventral view; protibia (D–F), dorsal view; antenna (G–I), dorsal view. A, D, G, Brachylepis elephas, male, Tanzania, Voi; B, E, H, Brachylepis werneri, male, Tanzania, Iringa; C, F, I, Brachylepis turmensis new species, holotype male.Scale bar= 1 mm." httpUri="https://zenodo.org/record/1163975/files/figure.png" pageId="0" pageNumber="291">
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3C, 3F,
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<date id="C017107D29114E18C13CFB10F107FB1C" box="[1034,1062,1238,1264]" pageId="0" pageNumber="291">3I</date>
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</figureCitation>
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,
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<figureCitation id="2C922A3829114E18C105FB10F176FB1C" box="[1075,1111,1238,1264]" captionStart="FIGURES 4" captionStartId="4.[151,258,1448,1470]" captionTargetBox="[150,1436,191,1427]" captionTargetId="figure@4.[150,1436,191,1427]" captionTargetPageId="4" captionText="FIGURES 4A–C. Map of currently known geographical distributions. A, Brachylepis elephas; B, Brachylepis werneri; C, Brachylepisturmensis newspecies.Reddots indicate new species or new locality." httpUri="https://zenodo.org/record/1163977/files/figure.png" pageId="0" pageNumber="291">4C</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="FCB3653629114E18C5A1FB3FF7FBFA94" pageId="0" pageNumber="291" type="materials_examined">
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<paragraph id="B41636BD29114E18C5A1FB3FF10AFAFC" blockId="0.[151,1436,1238,1989]" box="[151,1067,1273,1296]" pageId="0" pageNumber="291">
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<materialsCitation id="04C13CE029114E18C5A1FB3FF10AFAFC" ID-GBIF-Occurrence="1806316886" box="[151,1067,1273,1296]" country="Ethiopia" location="Peoples' Region" pageId="0" pageNumber="291" specimenCount="1" stateProvince="Southern Nations" typeStatus="holotype">
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<emphasis id="86DDEAAF29114E18C5A1FB3FF40FFAFC" bold="true" box="[151,302,1273,1296]" pageId="0" pageNumber="291">
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<typeStatus id="6B12881F29114E18C5A1FB3FF5EEFAFC" box="[151,207,1273,1296]" pageId="0" pageNumber="291">Type</typeStatus>
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locality.
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</emphasis>
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||
<collectingCountry id="CCBE762D29114E18C402FB3FF4B0FAFC" box="[308,401,1273,1296]" name="Ethiopia" pageId="0" pageNumber="291">Ethiopia</collectingCountry>
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,
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<collectingRegion id="766DF85F29114E18C4ADFB3FF779FAFC" box="[411,600,1273,1296]" country="Ethiopia" name="Ye Debub Biheroch Bihereseboch na Hizboch" pageId="0" pageNumber="291">Southern Nations</collectingRegion>
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,
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<collectingRegion id="766DF85F29114E18C754FB3FF7CCFAFC" box="[610,749,1273,1296]" country="Ethiopia" name="Nationalities" pageId="0" pageNumber="291">Nationalities</collectingRegion>
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, and
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<location id="B176606629114E18C613FB3FF6F6FAFC" LSID="urn:lsid:plazi:treatment:3C0087AB29114E1AC5A1FB10F648FD35:B176606629114E18C613FB3FF6F6FAFC" box="[805,983,1273,1296]" country="Ethiopia" name="Peoples' Region" pageId="0" pageNumber="291" stateProvince="Southern Nations">Peoples’ Region</location>
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, Turmi.
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</materialsCitation>
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</paragraph>
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<paragraph id="B41636BD29114E18C5F1FADAF7FBFA94" blockId="0.[151,1436,1238,1989]" pageId="0" pageNumber="291">
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<materialsCitation id="04C13CE029114E18C5F1FADAF7F6FA94" ID-GBIF-Occurrence="1806316889" collectingDate="2016-11-04" collectionCode="RSCV" collectorName="Vladimir Major" country="Czech Republic" county="Richard" elevation="950" latitude="36.5" location="Narodni" municipality="Velenice" pageId="0" pageNumber="291" specimenCount="950" stateProvince="Praha" typeStatus="holotype">
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<emphasis id="86DDEAAF29114E18C5F1FADAF44FFADF" bold="true" box="[199,366,1308,1331]" pageId="0" pageNumber="291">
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<typeStatus id="6B12881F29114E18C5F1FADAF5DEFADF" box="[199,255,1308,1331]" pageId="0" pageNumber="291">Type</typeStatus>
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material.
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||
</emphasis>
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||
<typeStatus id="6B12881F29114E18C440FADAF4FCFADF" box="[374,477,1308,1331]" pageId="0" pageNumber="291" type="holotype">Holotype</typeStatus>
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||
: “ETHIOPIA—Hamer or. Turmi near
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<specimenCount id="A2AFFD3429114E18C648FADAF6EBFADF" box="[894,970,1308,1331]" pageId="0" pageNumber="291" type="generic">
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<elevation id="3F84D18E29114E18C648FADAF6E6FADF" box="[894,967,1308,1331]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="0" pageNumber="291" unit="m" value="950.0">
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<quantity id="73519B5829114E18C648FADAF6E6FADF" box="[894,967,1308,1331]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="0" pageNumber="291" unit="m" value="950.0">950 m</quantity>
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</elevation>
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.
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||
</specimenCount>
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||
| 04°58´31˝
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||
<geoCoordinate id="D19D507A29114E18C17CFADAF19CFADF" box="[1098,1213,1308,1331]" direction="north" orientation="latitude" pageId="0" pageNumber="291" precision="925" value="36.5">N 036°30´</geoCoordinate>
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53˝E | 9.-
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||
<collectingDate id="D053E99529114E18C028FADAF0AFFADF" box="[1310,1422,1308,1331]" pageId="0" pageNumber="291" value="2016-11-04">
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||
<date id="C017107D29114E18C028FADAF0AFFADF" box="[1310,1422,1308,1331]" pageId="0" pageNumber="291" value="2016-11-04">11.4.2016</date>
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</collectingDate>
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|
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||
<collectorName id="195C536B29114E18C5A1FAF8F41FFAB9" box="[151,318,1342,1365]" pageId="0" pageNumber="291">Vladimír Major</collectorName>
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leg. [printed] ||
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<collectionCode id="D2B8AE7829114E18C4D0FAF8F70AFAB9" box="[486,555,1342,1365]" pageId="0" pageNumber="291">RSCV</collectionCode>
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|
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<collectingCounty id="5D774E3129114E18C708FAF8F7B3FAB9" box="[574,658,1342,1365]" pageId="0" pageNumber="291">Richard</collectingCounty>
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SEHNAL | collection |
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||
<collectingMunicipality id="5472ACC729114E18C6B8FAF8F6C8FAB9" box="[910,1001,1342,1365]" pageId="0" pageNumber="291">Velenice</collectingMunicipality>
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|
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<collectingCountry id="CCBE762D29114E18C6CDFAF8F185FAB9" box="[1019,1188,1342,1365]" name="Czech Republic" pageId="0" pageNumber="291">Czech Republic</collectingCountry>
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[printed]”.
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<typeStatus id="6B12881F29114E18C016FAF8F0A2FAB9" box="[1312,1411,1342,1365]" pageId="0" pageNumber="291" type="holotype">Holotype</typeStatus>
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is deposited in
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||
<location id="B176606629114E18C429FAA7F457FA94" LSID="urn:lsid:plazi:treatment:3C0087AB29114E1AC5A1FB10F648FD35:B176606629114E18C429FAA7F457FA94" box="[287,374,1377,1400]" country="Czech Republic" county="Richard" latitude="36.5" municipality="Velenice" name="Narodni" pageId="0" pageNumber="291" stateProvince="Praha">Národní</location>
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muzeum
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<collectingRegion id="766DF85F29114E18C4E9FAA7F73EFA94" box="[479,543,1377,1400]" country="Czech Republic" name="Praha" pageId="0" pageNumber="291">Praha</collectingRegion>
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,
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||
<collectingCountry id="CCBE762D29114E18C71FFAA7F7F6FA94" box="[553,727,1377,1400]" name="Czech Republic" pageId="0" pageNumber="291">Czech Republic</collectingCountry>
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</materialsCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="FCB3653629114E1AC5F1FA42F729FDC0" lastPageId="2" lastPageNumber="293" pageId="0" pageNumber="291" type="description">
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<paragraph id="B41636BD29114E18C5F1FA42F022FA51" blockId="0.[151,1436,1238,1989]" pageId="0" pageNumber="291">
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<emphasis id="86DDEAAF29114E18C5F1FA42F4FCFA77" bold="true" box="[199,477,1412,1435]" pageId="0" pageNumber="291">
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||
Description of
|
||
<typeStatus id="6B12881F29114E18C442FA42F4F9FA77" box="[372,472,1412,1435]" pageId="0" pageNumber="291" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Body length
|
||
<quantity id="73519B5829114E18C741FA42F7F8FA77" box="[631,729,1412,1435]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="0" pageNumber="291" unit="mm" value="24.5">24.5 mm</quantity>
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||
(without pygidium). Body elongate, posteriorly slightly dilated. Dorsal surface and abdomen dark brown, legs partly lighter, antenna bicolorous, elytra pruinose (
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<figureCitation id="2C922A3829114E18C194FA60F1D4FA51" box="[1186,1269,1446,1469]" captionStart="FIGURES 1" captionStartId="1.[151,258,1608,1630]" captionTargetBox="[184,1421,197,1572]" captionTargetId="figure@1.[150,1436,191,1587]" captionTargetPageId="1" captionText="FIGURES 1A–F. Habitus (A–C) and detail of setation on pronotum (D–F), dorsal view. A, D, Brachylepis elephas, male, Tanzania, Voi;B, E, Brachylepis werneri, male, Tanzania, Iringa; C, F, Brachylepis turmensis new species, holotype male.Scale bar = 1 cm (for habitus); scale bar = 2 mm (for detail of pronotum)." httpUri="https://zenodo.org/record/1163971/files/figure.png" pageId="0" pageNumber="291">Fig. 1C</figureCitation>
|
||
).
|
||
</paragraph>
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||
<paragraph id="B41636BD29114E18C5A1FA0FF045F8D7" blockId="0.[151,1436,1238,1989]" pageId="0" pageNumber="291">
|
||
Clypeus 2.8x wider than long, with anterior angles broadly rounded and anterior margin strongly upturned and bisinuate; surface deeply and densely punctate, with smooth facets between punctures; posterolaterally oriented, pointed, semierect scale originating from centre of each puncture; some medial scales narrower than others, frontoclypeal suture weakly indicated. Frons deeply and densely punctate, with a scale originating from each puncture; scales near midline point laterally toward a longitudinal bulge in each half of frons, forming a bisinuate pattern (
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||
<figureCitation id="2C922A3829114E18C11DF992F15FF987" box="[1067,1150,1620,1643]" captionStart="FIGURES 1" captionStartId="1.[151,258,1608,1630]" captionTargetBox="[184,1421,197,1572]" captionTargetId="figure@1.[150,1436,191,1587]" captionTargetPageId="1" captionText="FIGURES 1A–F. Habitus (A–C) and detail of setation on pronotum (D–F), dorsal view. A, D, Brachylepis elephas, male, Tanzania, Voi;B, E, Brachylepis werneri, male, Tanzania, Iringa; C, F, Brachylepis turmensis new species, holotype male.Scale bar = 1 cm (for habitus); scale bar = 2 mm (for detail of pronotum)." httpUri="https://zenodo.org/record/1163971/files/figure.png" pageId="0" pageNumber="291">Fig. 1C</figureCitation>
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). Labrum bilobed, anterior angles obtuse but distinct, macrosetae longest at edge of clypeus and around eye canthus. Eye canthus long, with macrosetae in two orderly rows; anterior conical setae short, posterior conical setae
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<date id="C017107D29114E18C17FF95FF145F95C" box="[1097,1124,1689,1712]" pageId="0" pageNumber="291">2x</date>
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||
shorter than clypeal scales. Antenna bicolorous, with nine antennomeres; antennal shaft reddish brown, antennal club yellow. Club with three antennomeres, straight, same length as antennal shaft (antennomeres 1–4 combined), apex of club with irregular, fine, short macrosetae with sensilla pits (
|
||
<figureCitation id="2C922A3829114E18C725F8C7F77FF8F4" box="[531,606,1793,1816]" captionStart="FIGURES 3" captionStartId="3.[151,258,1061,1083]" captionTargetBox="[159,1415,198,1026]" captionTargetId="figure@3.[151,1435,191,1040]" captionTargetPageId="3" captionText="FIGURES 3A–I. Mesotibia (A–C), ventral view; protibia (D–F), dorsal view; antenna (G–I), dorsal view. A, D, G, Brachylepis elephas, male, Tanzania, Voi; B, E, H, Brachylepis werneri, male, Tanzania, Iringa; C, F, I, Brachylepis turmensis new species, holotype male.Scale bar= 1 mm." httpUri="https://zenodo.org/record/1163975/files/figure.png" pageId="0" pageNumber="291">
|
||
Fig.
|
||
<date id="C017107D29114E18C773F8C7F77FF8F4" box="[581,606,1793,1816]" pageId="0" pageNumber="291">3I</date>
|
||
</figureCitation>
|
||
). Terminal maxillary palpomere widest at midlength, apically truncate, with a terminal tubercle on inner, more angular side, and dorsally with a large oval alutaceous area widening toward apex.
|
||
</paragraph>
|
||
<paragraph id="B41636BD29114E18C5F1F880F484F829" blockId="0.[151,1436,1238,1989]" pageId="0" pageNumber="291">
|
||
Pronotum weekly convex, approximately octagonal (
|
||
<figureCitation id="2C922A3829114E18C7C9F880F673F8B1" box="[767,850,1862,1885]" captionStart="FIGURES 1" captionStartId="1.[151,258,1608,1630]" captionTargetBox="[184,1421,197,1572]" captionTargetId="figure@1.[150,1436,191,1587]" captionTargetPageId="1" captionText="FIGURES 1A–F. Habitus (A–C) and detail of setation on pronotum (D–F), dorsal view. A, D, Brachylepis elephas, male, Tanzania, Voi;B, E, Brachylepis werneri, male, Tanzania, Iringa; C, F, Brachylepis turmensis new species, holotype male.Scale bar = 1 cm (for habitus); scale bar = 2 mm (for detail of pronotum)." httpUri="https://zenodo.org/record/1163971/files/figure.png" pageId="0" pageNumber="291">Fig. 1C</figureCitation>
|
||
),
|
||
<date id="C017107D29114E18C651F880F6A3F8B1" box="[871,898,1862,1885]" pageId="0" pageNumber="291">2x</date>
|
||
wider than long, widest in posterior third. Lateral margins crenulate, with a pale brown straight seta issuing from each crenulation. Surface deeply and densely punctate; spaces between them smooth and shiny; each puncture bearing a pointed scale. Scales forming groups oriented mostly toward midline (
|
||
<figureCitation id="2C922A3829114E18C471F868F4B9F829" box="[327,408,1966,1989]" captionStart="FIGURES 1" captionStartId="1.[151,258,1608,1630]" captionTargetBox="[184,1421,197,1572]" captionTargetId="figure@1.[150,1436,191,1587]" captionTargetPageId="1" captionText="FIGURES 1A–F. Habitus (A–C) and detail of setation on pronotum (D–F), dorsal view. A, D, Brachylepis elephas, male, Tanzania, Voi;B, E, Brachylepis werneri, male, Tanzania, Iringa; C, F, Brachylepis turmensis new species, holotype male.Scale bar = 1 cm (for habitus); scale bar = 2 mm (for detail of pronotum)." httpUri="https://zenodo.org/record/1163971/files/figure.png" pageId="0" pageNumber="291">Fig. 1F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="E0D6663529104E19C5A1F98EF7A6F974" httpUri="https://zenodo.org/record/1163971/files/figure.png" pageId="1" pageNumber="292" targetBox="[184,1421,197,1572]" targetPageId="1">
|
||
<paragraph id="B41636BD29104E19C5A1F98EF7A6F974" blockId="1.[151,1436,1608,1688]" pageId="1" pageNumber="292">
|
||
<emphasis id="86DDEAAF29104E19C5A1F98EF465F9B2" bold="true" box="[151,324,1608,1630]" pageId="1" pageNumber="292">FIGURES 1A–F.</emphasis>
|
||
Habitus (A–C) and detail of setation on pronotum (D–F), dorsal view. A, D,
|
||
<emphasis id="86DDEAAF29104E19C103F98EF1DFF9B2" box="[1077,1278,1608,1630]" italics="true" pageId="1" pageNumber="292">
|
||
<taxonomicName id="73A94D3E29104E19C103F98EF1DBF9B2" baseAuthorityName="Gerstaecker" baseAuthorityYear="1867" box="[1077,1274,1608,1630]" class="Insecta" family="Scarabaeidae" genus="Brachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="elephas">Brachylepis elephas</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
male, Tanzania, Voi; B, E,
|
||
<taxonomicName id="73A94D3E29104E19C5CDF9A3F49EF997" authorityName="Lacroix" authorityYear="2009" box="[251,447,1637,1659]" class="Insecta" family="Scarabaeidae" genus="Brachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="werneri">
|
||
<emphasis id="86DDEAAF29104E19C5CDF9A3F49EF997" box="[251,447,1637,1659]" italics="true" pageId="1" pageNumber="292">Brachylepis werneri</emphasis>
|
||
</taxonomicName>
|
||
, male, Tanzania, Iringa; C, F,
|
||
<taxonomicName id="73A94D3E29104E19C7D7F9A3F697F997" authority="Sehnal, 2018" authorityName="Sehnal" authorityYear="2018" box="[737,950,1637,1659]" class="Insecta" family="Scarabaeidae" genus="Brachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="turmensis" status="sp. nov.">
|
||
<emphasis id="86DDEAAF29104E19C7D7F9A3F697F997" box="[737,950,1637,1659]" italics="true" pageId="1" pageNumber="292">Brachylepis turmensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9DEE57D429104E19C68AF9A0F112F997" box="[956,1075,1638,1659]" pageId="1" pageNumber="292" rank="species">new species</taxonomicNameLabel>
|
||
, holotype male. Scale bar = 1 cm (for habitus); scale bar = 2 mm (for detail of pronotum).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B41636BD29104E19C5F1F902F5E4F911" blockId="1.[151,1436,1732,2032]" pageId="1" pageNumber="292">
|
||
Scutellum wider than long, triangular, evenly punctate in anterior half, with scales similar to those on pronotum (
|
||
<figureCitation id="2C922A3829104E19C046F902F599F911" captionStart="FIGURES 1" captionStartId="1.[151,258,1608,1630]" captionTargetBox="[184,1421,197,1572]" captionTargetId="figure@1.[150,1436,191,1587]" captionTargetPageId="1" captionText="FIGURES 1A–F. Habitus (A–C) and detail of setation on pronotum (D–F), dorsal view. A, D, Brachylepis elephas, male, Tanzania, Voi;B, E, Brachylepis werneri, male, Tanzania, Iringa; C, F, Brachylepis turmensis new species, holotype male.Scale bar = 1 cm (for habitus); scale bar = 2 mm (for detail of pronotum)." httpUri="https://zenodo.org/record/1163971/files/figure.png" pageId="1" pageNumber="292">Fig. 1C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B41636BD29104E19C5F1F8CFF747F864" blockId="1.[151,1436,1732,2032]" pageId="1" pageNumber="292">Elytra convex, elytron 1.25x longer than wide, without true striae and intervals, instead only indistinct broad, slightly elevated or lowered strips indicated by relative density of squamation, with oblique crests nearly glabrous. Part of elytron between terminal bulge and apex short, steeply inclined. Spaces between punctures and scales finely rugate and shiny. Lateral margins with short setae.</paragraph>
|
||
<paragraph id="B41636BD29104E1AC5F1F852F63AFF18" blockId="1.[151,1436,1732,2032]" lastBlockId="2.[151,1436,152,729]" lastPageId="2" lastPageNumber="293" pageId="1" pageNumber="292">
|
||
<emphasis id="86DDEAAF29104E19C5F1F852F44FF847" bold="true" box="[199,366,1940,1963]" pageId="1" pageNumber="292">Macropterous.</emphasis>
|
||
All femora shiny, irregularly punctate, with yellowish-white scales and reddish-brown long setae. Protibia tridentate; inner terminal spur at level of proximal tooth (
|
||
<figureCitation id="2C922A3829104E19C643F870F6E8F821" box="[885,969,1974,1997]" captionStart="FIGURES 3" captionStartId="3.[151,258,1061,1083]" captionTargetBox="[159,1415,198,1026]" captionTargetId="figure@3.[151,1435,191,1040]" captionTargetPageId="3" captionText="FIGURES 3A–I. Mesotibia (A–C), ventral view; protibia (D–F), dorsal view; antenna (G–I), dorsal view. A, D, G, Brachylepis elephas, male, Tanzania, Voi; B, E, H, Brachylepis werneri, male, Tanzania, Iringa; C, F, I, Brachylepis turmensis new species, holotype male.Scale bar= 1 mm." httpUri="https://zenodo.org/record/1163975/files/figure.png" pageId="1" pageNumber="292">Fig. 3F</figureCitation>
|
||
). Meso- and metatibiae expanded distally, densely covered by broad scales and long setae, with one curved spur apically and one oblique interrupted carina externally (
|
||
<figureCitation id="2C922A3829134E1AC426FF5EF442FF43" box="[272,355,152,175]" captionStart="FIGURES 3" captionStartId="3.[151,258,1061,1083]" captionTargetBox="[159,1415,198,1026]" captionTargetId="figure@3.[151,1435,191,1040]" captionTargetPageId="3" captionText="FIGURES 3A–I. Mesotibia (A–C), ventral view; protibia (D–F), dorsal view; antenna (G–I), dorsal view. A, D, G, Brachylepis elephas, male, Tanzania, Voi; B, E, H, Brachylepis werneri, male, Tanzania, Iringa; C, F, I, Brachylepis turmensis new species, holotype male.Scale bar= 1 mm." httpUri="https://zenodo.org/record/1163975/files/figure.png" pageId="2" pageNumber="293">Fig. 3C</figureCitation>
|
||
). Distal edge with a row of short, stout setae of equal length, terminal calcars stout, long, lower calcar little shorter than upper; pro-, meso- and metatarsomeres without patches of short, dense setae; all tarsomeres ventrally covered by sparse setae. Claws bifid, with ventrobasal teeth.
|
||
</paragraph>
|
||
<paragraph id="B41636BD29134E1AC5F1FEC6F1F0FEFB" blockId="2.[151,1436,152,729]" box="[199,1233,256,279]" pageId="2" pageNumber="293">Abdominal ventrites reddish brown, shiny, finely punctate, densely covered by oval white scales.</paragraph>
|
||
<paragraph id="B41636BD29134E1AC5F1FEE4F4BEFEB0" blockId="2.[151,1436,152,729]" pageId="2" pageNumber="293">Pygidium slightly concave, shallowly punctate, with yellowish-white, caudally oriented scales becoming finer toward posterior margin.</paragraph>
|
||
<paragraph id="B41636BD29134E1AC5F1FEAEF131FE28" blockId="2.[151,1436,152,729]" pageId="2" pageNumber="293">
|
||
Male genitalia (
|
||
<figureCitation id="2C922A3829134E1AC445FEAEF4CDFE93" box="[371,492,360,383]" captionStart="FIGURES 2" captionStartId="2.[151,258,1725,1747]" captionTargetBox="[160,1418,797,1684]" captionTargetId="figure@2.[151,1435,781,1705]" captionTargetPageId="2" captionText="FIGURES 2A–I. Aedeagus in dorsal (A, D, G) and lateral (B, E, H) views; apex of aedeagus (C, F, I). A–C, Brachylepis elephas, Tanzania, Voi;D–F,Brachylepis werneri, Tanzania,Iringa; G–I, Brachylepisturmensis new species, holotype.Scale bar= 1 mm." httpUri="https://zenodo.org/record/1163973/files/figure.png" pageId="2" pageNumber="293">Figs. 2G–I</figureCitation>
|
||
). Aedeagus symmetrical, conical in dorsal view. Phallobase in lateral view dorsally as well as ventrally doubly undulate, dorsally at apex terminating in a distinct hook. Each paramere with a double tooth, medially teeth meet and in frontoventral view form a horizontal number 8 (
|
||
<figureCitation id="2C922A3829134E1AC68EFE6BF122FE28" box="[952,1027,429,452]" captionStart="FIGURES 2" captionStartId="2.[151,258,1725,1747]" captionTargetBox="[160,1418,797,1684]" captionTargetId="figure@2.[151,1435,781,1705]" captionTargetPageId="2" captionText="FIGURES 2A–I. Aedeagus in dorsal (A, D, G) and lateral (B, E, H) views; apex of aedeagus (C, F, I). A–C, Brachylepis elephas, Tanzania, Voi;D–F,Brachylepis werneri, Tanzania,Iringa; G–I, Brachylepisturmensis new species, holotype.Scale bar= 1 mm." httpUri="https://zenodo.org/record/1163973/files/figure.png" pageId="2" pageNumber="293">
|
||
Fig.
|
||
<date id="C017107D29134E1AC6DCFE6BF122FE28" box="[1002,1027,429,452]" pageId="2" pageNumber="293">2I</date>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B41636BD29134E1AC5F1FE16F753FE0B" blockId="2.[151,1436,152,729]" box="[199,626,464,487]" pageId="2" pageNumber="293">
|
||
<emphasis id="86DDEAAF29134E1AC5F1FE16F485FE0B" bold="true" box="[199,420,464,487]" pageId="2" pageNumber="293">Sexual dimorphism</emphasis>
|
||
. Female unknown.
|
||
</paragraph>
|
||
<paragraph id="B41636BD29134E1AC5F1FE34F729FDC0" blockId="2.[151,1436,152,729]" pageId="2" pageNumber="293">
|
||
<materialsCitation id="04C13CE029134E1AC5F1FE34F729FDC0" ID-GBIF-Occurrence="1806316890" country="Ethiopia" location="Peoples' Region" municipality="Geographic" pageId="2" pageNumber="293" specimenCount="1" stateProvince="Southern Nations" typeStatus="holotype">
|
||
<emphasis id="86DDEAAF29134E1AC5F1FE34F4FCFDE5" bold="true" box="[199,477,498,521]" pageId="2" pageNumber="293">
|
||
<collectingMunicipality id="5472ACC729134E1AC5F1FE34F46DFDE5" box="[199,332,498,521]" pageId="2" pageNumber="293">Geographic</collectingMunicipality>
|
||
distribution
|
||
</emphasis>
|
||
.
|
||
<collectingCountry id="CCBE762D29134E1AC4D8FE34F76AFDE5" box="[494,587,498,521]" name="Ethiopia" pageId="2" pageNumber="293">Ethiopia</collectingCountry>
|
||
,
|
||
<collectingRegion id="766DF85F29134E1AC76FFE34F638FDE5" box="[601,793,498,521]" country="Ethiopia" name="Ye Debub Biheroch Bihereseboch na Hizboch" pageId="2" pageNumber="293">Southern Nations</collectingRegion>
|
||
,
|
||
<collectingRegion id="766DF85F29134E1AC611FE34F68EFDE5" box="[807,943,498,521]" country="Ethiopia" name="Nationalities" pageId="2" pageNumber="293">Nationalities</collectingRegion>
|
||
and
|
||
<location id="B176606629134E1AC6DFFE34F1BEFDE5" LSID="urn:lsid:plazi:treatment:3C0087AB29114E1AC5A1FB10F648FD35:B176606629134E1AC6DFFE34F1BEFDE5" box="[1001,1183,498,521]" country="Ethiopia" municipality="Geographic" name="Peoples' Region" pageId="2" pageNumber="293" stateProvince="Southern Nations">Peoples’ Region</location>
|
||
, Turmi. The species is known only from the
|
||
<typeStatus id="6B12881F29134E1AC448FDD3F48DFDC0" box="[382,428,533,556]" pageId="2" pageNumber="293">type</typeStatus>
|
||
locality.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FCB3653629134E1AC5F1FDFEF60AFD5B" pageId="2" pageNumber="293" type="diagnosis">
|
||
<paragraph id="B41636BD29134E1AC5F1FDFEF60AFD5B" blockId="2.[151,1436,152,729]" pageId="2" pageNumber="293">
|
||
<emphasis id="86DDEAAF29134E1AC5F1FDFEF49DFDA3" bold="true" box="[199,444,568,591]" pageId="2" pageNumber="293">Differential diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="73A94D3E29134E1AC4FAFDF1F79AFDA3" authority="Sehnal, 2018" authorityName="Sehnal" authorityYear="2018" box="[460,699,567,591]" class="Insecta" family="Scarabaeidae" genus="Brachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="turmensis" status="sp. nov.">
|
||
<emphasis id="86DDEAAF29134E1AC4FAFDF1F79AFDA3" box="[460,699,567,591]" italics="true" pageId="2" pageNumber="293">Brachylepis turmensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9DEE57D429134E1AC7F2FDFEF666FDA3" box="[708,839,568,591]" pageId="2" pageNumber="293" rank="species">new species</taxonomicNameLabel>
|
||
is the third representative of the genus
|
||
<taxonomicName id="73A94D3E29134E1AC1CBFDF1F05DFDA3" authorityName="Kolbe" authorityYear="1894" box="[1277,1404,567,591]" class="Insecta" family="Scarabaeidae" genus="Brachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="86DDEAAF29134E1AC1CBFDF1F05DFDA3" box="[1277,1404,567,591]" italics="true" pageId="2" pageNumber="293">Brachylepis</emphasis>
|
||
</taxonomicName>
|
||
. It can be distinguished from the previously described species by the following characters: length <25.0 mm, pronotal scales oval, toward base weakly overlapping margins of punctures (
|
||
<figureCitation id="2C922A3829134E1AC6BDFDBBF6FFFD78" box="[907,990,637,660]" captionStart="FIGURES 1" captionStartId="1.[151,258,1608,1630]" captionTargetBox="[184,1421,197,1572]" captionTargetId="figure@1.[150,1436,191,1587]" captionTargetPageId="1" captionText="FIGURES 1A–F. Habitus (A–C) and detail of setation on pronotum (D–F), dorsal view. A, D, Brachylepis elephas, male, Tanzania, Voi;B, E, Brachylepis werneri, male, Tanzania, Iringa; C, F, Brachylepis turmensis new species, holotype male.Scale bar = 1 cm (for habitus); scale bar = 2 mm (for detail of pronotum)." httpUri="https://zenodo.org/record/1163971/files/figure.png" pageId="2" pageNumber="293">Fig. 1F</figureCitation>
|
||
); antennal club yellow and shorter than reddish-brown antennal shaft; shape of the aedeagus (
|
||
<figureCitation id="2C922A3829134E1AC7E5FD66F63FFD5B" box="[723,798,672,695]" captionStart="FIGURES 3" captionStartId="3.[151,258,1061,1083]" captionTargetBox="[159,1415,198,1026]" captionTargetId="figure@3.[151,1435,191,1040]" captionTargetPageId="3" captionText="FIGURES 3A–I. Mesotibia (A–C), ventral view; protibia (D–F), dorsal view; antenna (G–I), dorsal view. A, D, G, Brachylepis elephas, male, Tanzania, Voi; B, E, H, Brachylepis werneri, male, Tanzania, Iringa; C, F, I, Brachylepis turmensis new species, holotype male.Scale bar= 1 mm." httpUri="https://zenodo.org/record/1163975/files/figure.png" pageId="2" pageNumber="293">
|
||
Fig.
|
||
<date id="C017107D29134E1AC633FD66F63FFD5B" box="[773,798,672,695]" pageId="2" pageNumber="293">3I</date>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FCB3653629134E1AC5F1FD04F648FD35" box="[199,873,706,729]" pageId="2" pageNumber="293" type="etymology">
|
||
<paragraph id="B41636BD29134E1AC5F1FD04F648FD35" blockId="2.[151,1436,152,729]" box="[199,873,706,729]" pageId="2" pageNumber="293">
|
||
<emphasis id="86DDEAAF29134E1AC5F1FD04F461FD35" bold="true" box="[199,320,706,729]" pageId="2" pageNumber="293">Etymology</emphasis>
|
||
. Derived from the name of the
|
||
<typeStatus id="6B12881F29134E1AC7A6FD04F79FFD35" box="[656,702,706,729]" pageId="2" pageNumber="293">type</typeStatus>
|
||
locality, Turmi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |