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<document id="02B8E51E545EEF31B57795932F57B298" ID-DOI="10.11646/zootaxa.5476.1.29" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681921" ID-ZooBank="DD4EA1FE-A0BE-4BA4-9AD3-E460B131C518" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="felipe" checkinTime="1720375669801" checkinUser="plazi" docAuthor="Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2024" docId="03DD87DAFFAD125128EEF969807318AF" docLanguage="en" docName="zootaxa.5476.1.29.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.29" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyphira linda Rahayu &amp; Ng, 2024, n. sp." docType="treatment" docVersion="4" lastPageNumber="379" masterDocId="FFE4FFA2FFBE12442879FFAF843C1F54" masterDocTitle="On the genus Lyphira Galil, 2009 (Crustacea: Brachyura: Leucosiidae), with descriptions of four new species from the West Pacific" masterLastPageNumber="392" masterPageNumber="358" pageNumber="377" updateTime="1721070200110" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="CA800EFB853D3558F12A72B44B9713A7">On the genus Lyphira Galil, 2009 (Crustacea: Brachyura: Leucosiidae), with descriptions of four new species from the West Pacific</mods:title>
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<mods:namePart id="4E7A7C8B064A7C4C252800DAACF4A511">Rahayu, Dwi Listyo</mods:namePart>
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<taxonomicName id="4C744D4FFFAD125728EEF969850019B5" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[151,316,1734,1761]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="linda" status="sp. nov.">
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(
<figureCitation id="134F2A49FFAD125728E6F94484D31851" box="[159,239,1771,1797]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="19" pageNumber="377">
Figs.
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<figureCitation id="134F2A49FFAD12572882F94485041851" box="[251,312,1771,1797]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="19" pageNumber="377">6E, F</figureCitation>
,
<figureCitation id="134F2A49FFAD1257293AF94485B51851" box="[323,393,1771,1797]" captionStart="FIGURE 8" captionStartId="13.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1714]" captionTargetId="figure-20@13.[151,1435,181,1714]" captionTargetPageId="13" captionText="FIGURE 8. Male thoracic sternum and pleon. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="19" pageNumber="377">8BD</figureCitation>
,
<figureCitation id="134F2A49FFAD125729EDF94485ED1851" box="[404,465,1771,1797]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="19" pageNumber="377">9E, F</figureCitation>
,
<figureCitation id="134F2A49FFAD125729A4F94486111851" box="[477,557,1771,1797]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="19" pageNumber="377">10FH</figureCitation>
,
<figureCitation id="134F2A49FFAD12572A41F94486B41851" box="[568,648,1771,1797]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="19" pageNumber="377">11FH</figureCitation>
,
<figureCitation id="134F2A49FFAD12572AEAF94486DD1851" box="[659,737,1771,1797]" captionStart="FIGURE 12" captionStartId="17.[151,250,1628,1652]" captionTargetBox="[151,1435,181,1605]" captionTargetId="figure-20@17.[151,1435,181,1605]" captionTargetPageId="17" captionText="FIGURE 12. Third maxillipeds. A, Lyphira chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; B, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; C, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore; D, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.97), Malaysia; E, L. linda n. sp., female (17.0 × 16.9 mm) (ZRC 1995.0547), Singapore; F, L. linda n. sp., male (12.1 × 12.4 mm) (ZRC 1995.0547), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681950" httpUri="https://zenodo.org/record/12681950/files/figure.png" pageId="19" pageNumber="377">12DF</figureCitation>
,
<figureCitation id="134F2A49FFAD12572A94F94487031851" box="[749,831,1771,1797]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="19" pageNumber="377">13HL</figureCitation>
,
<figureCitation id="134F2A49FFAD12572B33F94487A91851" box="[842,917,1771,1797]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="19" pageNumber="377">14E, F</figureCitation>
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<figureCitation id="134F2A49FFAD12572BD9F94587CA1851" box="[928,1014,1770,1797]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="19" pageNumber="377">15G, H</figureCitation>
,
<figureCitation id="134F2A49FFAD12572C78F944807A1851" box="[1025,1094,1771,1797]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. AG, IK, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="19" pageNumber="377">16FI</figureCitation>
,
<figureCitation id="134F2A49FFAD12572C2BF94480A01851" box="[1106,1180,1771,1797]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. AE, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; FK, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; LP, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; QV, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: AC, FH, L, QS = 1.0 mm; D, E, IK, MP, TV = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="19" pageNumber="377">18LP</figureCitation>
,
<figureCitation id="134F2A49FFAD12572CDEF94480C21851" box="[1191,1278,1771,1797]" captionStart="FIGURE 20" captionStartId="29.[151,250,1639,1663]" captionTargetBox="[252,1327,191,1613]" captionTargetId="figure-20@29.[239,1349,181,1615]" captionTargetPageId="29" captionText="FIGURE 20. G1. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; C, L. ngankee n. sp., paratype male (19.2 × 19.9 mm) (ZRC 2022.0628), Hong Kong; D, L. ngankee n. sp., paratype male (15.5 × 15.1 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (16.7 × 17.0 mm) (ZRC 2015.0439); Thailand; G, L. linda n. sp., male (18.4 × 19.0 mm) (ZRC 1985.1632), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. linda n. sp., male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; M, L. linda n. sp., male (22.9 × 21.4 mm) (ZRC 2014.0191), Vietnam; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia; O, L. bellagrana n. sp., male (13.2 × 12.6 mm) (ZRC 2022.0745), Papua, Indonesia; P, L. bellagrana n. sp., male (10.6 × 9.9 mm) (ZRC 2022.0750), Papua, Indonesia. A, C, EH, KP, right G1; B, D, I, J, left G1 (laterally inverted)." figureDoi="http://doi.org/10.5281/zenodo.12681975" httpUri="https://zenodo.org/record/12681975/files/figure.png" pageId="19" pageNumber="377">20EM</figureCitation>
,
<figureCitation id="134F2A49FFAD12572D70F94481691851" box="[1289,1365,1771,1797]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="19" pageNumber="377">21E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAD125728EEF88086BE18D1" pageId="19" pageNumber="377" type="reference_group">
<paragraph id="8BCB36CCFFAD125728EEF88080341832" blockId="19.[151,1381,1839,1925]" pageId="19" pageNumber="377">
<emphasis id="B900EADEFFAD125728EEF880856B1813" box="[151,343,1839,1863]" italics="true" pageId="19" pageNumber="377">
<taxonomicName id="4C744D4FFFAD125728EEF88085051813" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[151,313,1839,1863]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="globosa">Philyra globosa</taxonomicName>
.—
</emphasis>
<bibRefCitation id="EFE54B3DFFAD1257292EF880860E1813" author="Lanchester, W. F." box="[343,562,1839,1863]" pageId="19" pageNumber="377" pagination="719 - 770" refId="ref20885" refString="Lanchester, W. F. (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900, 719 - 770." type="book chapter" year="1900">Lanchester 1900: 764</bibRefCitation>
(p.p.). (not
<taxonomicName id="4C744D4FFFAD12572AD2F89F87CD1813" authority="Fabricius, 1793" authorityName="Fabricius" authorityYear="1793" box="[683,1009,1839,1863]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="B900EADEFFAD12572AD2F89F87701813" box="[683,844,1839,1863]" italics="true" pageId="19" pageNumber="377">Cancer globosa</emphasis>
<bibRefCitation id="EFE54B3DFFAD12572B2AF88087CD1813" author="Fabricius, J. C." box="[851,1009,1839,1863]" pageId="19" pageNumber="377" refId="ref19754" refString="Fabricius, J. C. (1793) Entomologia systematica emendata et aucta secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Vol. 2. impensis Christ. Gottl. Proft, Hafniae, viii + 519 pp. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1793">Fabricius, 1793</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C744D4FFFAD12572C72F880815D1813" ID-CoL="76YMG" authority="(Fabricius, 1775))" authorityName="" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[1035,1377,1839,1863]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="globus">
<emphasis id="B900EADEFFAD12572C72F880809D1813" box="[1035,1185,1839,1863]" italics="true" pageId="19" pageNumber="377">Philyra globus</emphasis>
(
<bibRefCitation id="EFE54B3DFFAD12572CD7F880816C1813" author="Fabricius, J. C." box="[1198,1360,1839,1863]" pageId="19" pageNumber="377" refId="ref19698" refString="Fabricius, J. C. (1775) Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, adiectis Synonymis, Locis, Descriptionibus, Observationibus. Flensburgi &amp; Lipsiae, Korte, xxvii + 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510" type="book" year="1775">Fabricius, 1775</bibRefCitation>
))
</taxonomicName>
.
<emphasis id="B900EADEFFAD125728EEF8E185BF1832" box="[151,387,1870,1894]" italics="true" pageId="19" pageNumber="377">
<taxonomicName id="4C744D4FFFAD125728EEF8E185591832" ID-CoL="76YMK" authorityName="Ortmann" authorityYear="1892" box="[151,357,1870,1894]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="heterograna">Philyra heterograna</taxonomicName>
.—
</emphasis>
<bibRefCitation id="EFE54B3DFFAD125729FAF8E1866D1832" author="Wee, Y. C. &amp; Ng, P. K. L." box="[387,593,1870,1894]" pageId="19" pageNumber="377" refId="ref22338" refString="Wee, Y. C. &amp; Ng, P. K. L. (Eds.) (1994) A first look at Biodiversity in Singapore. National Council of the Environment, Singapore, 163 pp." type="book" year="1994">Wee &amp; Ng 1994: 82</bibRefCitation>
. (not
<taxonomicName id="4C744D4FFFAD12572AF3F8E187C11832" ID-CoL="76YMK" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[650,1021,1870,1894]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="heterograna">
<emphasis id="B900EADEFFAD12572AF3F8E187641832" box="[650,856,1870,1894]" italics="true" pageId="19" pageNumber="377">Philyra heterograna</emphasis>
<bibRefCitation id="EFE54B3DFFAD12572B26F8E187C11832" author="Ortmann, A." box="[863,1021,1870,1894]" pageId="19" pageNumber="377" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="8BCB36CCFFAD125728EEF8C386BE18D1" blockId="19.[151,1381,1839,1925]" box="[151,642,1900,1925]" pageId="19" pageNumber="377">
<treatmentCitationGroup id="AB6411E2FFAD125728EEF8C386BE18D1" box="[151,642,1900,1925]" pageId="19" pageNumber="377">
<emphasis id="B900EADEFFAD125728EEF8C385BF18D0" box="[151,387,1900,1924]" italics="true" pageId="19" pageNumber="377">
<taxonomicName id="4C744D4FFFAD125728EEF8C3855918D0" ID-CoL="76YMK" authorityName="Ortmann" authorityYear="1892" box="[151,357,1900,1924]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="heterograna">Philyra heterograna</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0AD510DDFFAD125729FAF8C3861E18D1" author="Galil, B. S." box="[387,546,1900,1925]" page="300" pageId="19" pageNumber="377" year="2009">
<bibRefCitation id="EFE54B3DFFAD125729FAF8C3861E18D1" author="Galil, B. S." box="[387,546,1900,1925]" pageId="19" pageNumber="377" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009: 300</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAD125028EEF81C851319E9" lastPageId="20" lastPageNumber="378" pageId="19" pageNumber="377" type="materials_examined">
<paragraph id="8BCB36CCFFAD125028EEF81C851319E9" blockId="19.[151,1436,1970,2033]" lastBlockId="20.[151,1437,150,2013]" lastPageId="20" lastPageNumber="378" pageId="19" pageNumber="377">
<emphasis id="B900EADEFFAD125728EEF81C85011898" bold="true" box="[151,317,1970,1997]" pageId="19" pageNumber="377">Type material</emphasis>
.
<materialsCitation id="3B1C3C91FFAD12572932F81C860E18A5" collectingDate="1981-03" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Singapore" location="East Coast Lagoon" pageId="19" pageNumber="377" specimenCode="ZRC 1981.9" specimenCount="1" typeStatus="holotype">
<emphasis id="B900EADEFFAD12572932F81C858E1899" box="[331,434,1971,1997]" italics="true" pageId="19" pageNumber="377">
<typeStatus id="54CF886EFFAD12572932F81C858E1899" box="[331,434,1971,1997]" pageId="19" pageNumber="377" type="holotype">Holotype</typeStatus>
</emphasis>
: male (18.4 ×
<quantity id="4C8C9B29FFAD12572A2DF81C86801899" box="[596,700,1971,1997]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="19" pageNumber="377" unit="mm" value="18.2">18.2 mm</quantity>
),
<specimenCode id="DBD29EB7FFAD12572AB7F81C87601899" box="[718,860,1971,1997]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="19" pageNumber="377">ZRC 1981.9</specimenCode>
.2.7,
<location id="8EAB6017FFAD12572BE9F81C80591899" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAD12572BE9F81C80591899" box="[912,1125,1971,1997]" country="Singapore" name="East Coast Lagoon" pageId="19" pageNumber="377">East Coast Lagoon</location>
,
<collectingCountry id="F363765CFFAD12572C09F81C80D81899" box="[1136,1252,1971,1997]" name="Singapore" pageId="19" pageNumber="377">Singapore</collectingCountry>
, from fishermen nets, coll.
<collectorName id="2681531AFFAD12572976F87885AB18A5" box="[271,407,2007,2033]" pageId="19" pageNumber="377">P. K. L. Ng</collectorName>
,
<date id="FFCA100CFFAD125729DCF878860E18A5" box="[421,562,2007,2033]" pageId="19" pageNumber="377" value="1981-03">
<collectingDate id="EF8EE9E4FFAD125729DCF878860E18A5" box="[421,562,2007,2033]" pageId="19" pageNumber="377" value="1981-03">March 1981</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B1C3C91FFAD12572A39F87880A818A5" box="[576,1172,2006,2033]" pageId="19" pageNumber="377" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B900EADEFFAD12572A39F878868E18A4" box="[576,690,2007,2032]" italics="true" pageId="19" pageNumber="377">
<typeStatus id="54CF886EFFAD12572A39F878868E18A4" box="[576,690,2007,2032]" pageId="19" pageNumber="377" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D72FD45FFAD12572ABBF878872718A5" box="[706,795,2007,2033]" count="2" pageId="19" pageNumber="377" type="male">2 males</specimenCount>
(20 ×
<quantity id="4C8C9B29FFAD12572B15F87987E918A5" box="[876,981,2006,2033]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="19" pageNumber="377" unit="mm" value="19.5">19.5 mm</quantity>
, 21 ×
<quantity id="4C8C9B29FFAD12572C58F87880B018A5" box="[1057,1164,2007,2033]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" pageId="19" pageNumber="377" unit="mm" value="21.4">21.4 mm</quantity>
)
</materialsCitation>
,
<materialsCitation id="3B1C3C91FFAD12502CD8F87887DE1F81" collectingDate="1987-03-26" collectionCode="ZRC" collectorName=" Reef Ecology &amp; Research Team" country="Singapore" lastPageId="20" lastPageNumber="378" location="Singapore" pageId="19" pageNumber="377" specimenCode="ZRC 1990.7355, 7356" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAD12572CD8F878813918A5" box="[1185,1285,2007,2033]" count="1" pageId="19" pageNumber="377" type="female">1 female</specimenCount>
(15.5 ×
<quantity id="4C8C9B29FFAD12502D12F87884FB1FE5" lastPageId="20" lastPageNumber="378" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="19" pageNumber="377" unit="mm" value="16.1">16.1 mm</quantity>
),
<collectionCode id="ED65AE09FFAA125028A5FF38852E1FE5" box="[220,274,151,177]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC</collectionCode>
1981.9.2.8-10, same locality as holotype; 2 young females,
<specimenCode id="DBD29EB7FFAA12502BBCFF3880401FE5" box="[965,1148,150,177]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1990.7355</specimenCode>
<specimenCode id="DBD29EB7FFAA12502CF2FF3980F81FE5" box="[1163,1220,150,177]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">7356</specimenCode>
, off
<collectorName id="2681531AFFAA12502C83FF3881A51FE5" box="[1274,1433,151,177]" pageId="20" pageNumber="378">Pulau Tekong</collectorName>
,
<collectingCountry id="F363765CFFAA125028EEFF1485371F81" box="[151,267,187,213]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
, coll. dredge,
<collectorName id="2681531AFFAA125029D6FF1486761F81" box="[431,586,187,213]" pageId="20" pageNumber="378">Reef Ecology</collectorName>
&amp;
<collectorName id="2681531AFFAA12502A0AFF14871F1F81" box="[627,803,187,213]" pageId="20" pageNumber="378">Research Team</collectorName>
,
<date id="FFCA100CFFAA12502B49FF1487DE1F81" box="[816,994,187,213]" pageId="20" pageNumber="378" value="1987-03-26">
<collectingDate id="EF8EE9E4FFAA12502B49FF1487DE1F81" box="[816,994,187,213]" pageId="20" pageNumber="378" value="1987-03-26">26 March 1987</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502B89FF1484F31FAD" pageId="20" pageNumber="378" specimenCount="4" specimenCount-male="4">
<specimenCount id="9D72FD45FFAA12502B89FF1480741F81" box="[1008,1096,187,213]" count="4" pageId="20" pageNumber="378" type="male">4 males</specimenCount>
(7.6 ×
<quantity id="4C8C9B29FFAA12502CD9FF1480C41F81" box="[1184,1272,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="20" pageNumber="378" unit="mm" value="7.2">7.2 mm</quantity>
17.1 ×
<quantity id="4C8C9B29FFAA12502D12FF1484FB1FAD" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="20" pageNumber="378" unit="mm" value="16.7">16.7 mm</quantity>
)
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA125028A5FF7080201E15" collectingDate="2014-01-17" collectionCode="ZRC" collectorName="Comprehensive Marine Biodiversity Survey" country="Singapore" elevation="16" latitude="1.4031833" location="Ketem" longLatPrecision="1" longitude="103.93954" municipality="Pasir Ris" pageId="20" pageNumber="378" specimenCode="ZRC 2022.1001" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA125028A5FF7085031FAD" box="[220,319,223,249]" count="1" pageId="20" pageNumber="378" type="female">1 female</specimenCount>
(14.0 ×
<quantity id="4C8C9B29FFAA125029DAFF7086311FAD" box="[419,525,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="20" pageNumber="378" unit="mm" value="13.9">13.9 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502A58FF7086E61FAD" box="[545,730,223,249]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2022.1001</specimenCode>
, specimen SEA 3978, station DR276, muddy-sandy substrate, between
<location id="8EAB6017FFAA12502978FEAC85701E49" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502978FEAC85701E49" box="[257,332,259,285]" country="Singapore" latitude="1.4031833" longLatPrecision="1" longitude="103.93954" municipality="Pasir Ris" name="Ketem" pageId="20" pageNumber="378">Ketem</location>
and
<collectingMunicipality id="6BAFACB6FFAA125029F5FEAC85C51E49" box="[396,505,259,285]" pageId="20" pageNumber="378">Pasir Ris</collectingMunicipality>
,
<collectingCountry id="F363765CFFAA12502A70FEAC86411E49" box="[521,637,259,285]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
,
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<geoCoordinate id="EE40500BFFAA12502B51FEAD87FC1E49" box="[808,960,258,285]" degrees="103" direction="east" minutes="56.139" orientation="longitude" pageId="20" pageNumber="378" precision="1" value="103.93565">103°56.139'E</geoCoordinate>
01°24.191'N 103°56.372'E,
<quantity id="4C8C9B29FFAA12502D55FEAD84931E15" metricMagnitude="1" metricUnit="m" metricValue="1.61" metricValueMax="1.69" metricValueMin="1.53" pageId="20" pageNumber="378" unit="m" value="16.1" valueMax="16.9" valueMin="15.3">15.316.9 m</quantity>
, coll.
<collectorName id="2681531AFFAA12502881FE8886CC1E15" box="[248,752,295,321]" pageId="20" pageNumber="378">Comprehensive Marine Biodiversity Survey</collectorName>
, dredge,
<date id="FFCA100CFFAA12502B23FE8880201E15" box="[858,1052,295,321]" pageId="20" pageNumber="378" value="2014-01-17">
<collectingDate id="EF8EE9E4FFAA12502B23FE8880201E15" box="[858,1052,295,321]" pageId="20" pageNumber="378" value="2014-01-17">17 January 2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502C53FE8887371EDD" collectingDate="2012-10-30" collectionCode="ZRC" collectorName="Comprehensive Marine Biodiversity Survey" country="Singapore" elevation="22" latitude="1.3179166" location="Johor Strait" longLatPrecision="1" longitude="104.070816" pageId="20" pageNumber="378" specimenCode="ZRC 2022.1002" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502C53FE8880B11E15" box="[1066,1165,295,321]" count="1" pageId="20" pageNumber="378" type="female">1 female</specimenCount>
(10.0 ×
<quantity id="4C8C9B29FFAA12502C8DFE89816C1E15" box="[1268,1360,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="20" pageNumber="378" unit="mm" value="9.5">9.5 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502D1FFE88852D1E31" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2022.1002</specimenCode>
, station DW129,
<location id="8EAB6017FFAA125029AAFEE486641E31" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA125029AAFEE486641E31" box="[467,600,331,357]" country="Singapore" latitude="1.3179166" longLatPrecision="1" longitude="104.070816" name="Johor Strait" pageId="20" pageNumber="378">Johor Strait</location>
,
<collectingCountry id="F363765CFFAA12502A1BFEE486EA1E31" box="[610,726,331,357]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
,
<geoCoordinate id="EE40500BFFAA12502A98FEE587531E31" box="[737,879,330,357]" degrees="01" direction="north" minutes="19.075" orientation="latitude" pageId="20" pageNumber="378" precision="1" value="1.3179166">01°19.075'N</geoCoordinate>
<geoCoordinate id="EE40500BFFAA12502B0CFEE4802C1E31" box="[885,1040,331,357]" degrees="104" direction="east" minutes="04.249" orientation="longitude" pageId="20" pageNumber="378" precision="1" value="104.070816">104°04.249'E</geoCoordinate>
,
<quantity id="4C8C9B29FFAA12502C62FEE580961E31" box="[1051,1194,330,357]" metricMagnitude="1" metricUnit="m" metricValue="2.21" metricValueMax="2.25" metricValueMin="2.17" pageId="20" pageNumber="378" unit="m" value="22.1" valueMax="22.5" valueMin="21.7">21.722.5 m</quantity>
, coll.
<collectorName id="2681531AFFAA12502C95FEE485EB1EDD" pageId="20" pageNumber="378">Comprehensive Marine Biodiversity Survey</collectorName>
, dredge,
<date id="FFCA100CFFAA12502A3DFEC087371EDD" box="[580,779,367,393]" pageId="20" pageNumber="378" value="2012-10-30">
<collectingDate id="EF8EE9E4FFAA12502A3DFEC087371EDD" box="[580,779,367,393]" pageId="20" pageNumber="378" value="2012-10-30">30 October 2012</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502B60FEC081031E85" collectingDate="2012-10-25" collectionCode="ZRC" collectorName="Comprehensive Marine Biodiversity Survey" country="Singapore" elevation="10" latitude="1.3921499" location="Johor Strait" longLatPrecision="1" longitude="103.99983" municipality="Changi Park" pageId="20" pageNumber="378" specimenCode="ZRC 2022.1003" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502B60FEC087421EDD" box="[793,894,367,393]" count="1" pageId="20" pageNumber="378" type="female">1 female</specimenCount>
(14.6 × 14.0 mm),
<specimenCode id="DBD29EB7FFAA12502C10FEC081181EDD" box="[1129,1316,367,393]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2022.1003</specimenCode>
, specimen JS0606, station DW86, on mud,
<location id="8EAB6017FFAA12502A77FE3C86A91EF9" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502A77FE3C86A91EF9" box="[526,661,403,429]" country="Singapore" latitude="1.3921499" longLatPrecision="1" longitude="103.99983" municipality="Changi Park" name="Johor Strait" pageId="20" pageNumber="378">Johor Strait</location>
,
<collectingMunicipality id="6BAFACB6FFAA12502ADBFE3C870E1EF9" box="[674,818,403,429]" pageId="20" pageNumber="378">Changi Park</collectingMunicipality>
,
<collectingCountry id="F363765CFFAA12502B46FE3C878F1EF9" box="[831,947,403,429]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
,
<geoCoordinate id="EE40500BFFAA12502BB9FE3D80721EF9" box="[960,1102,402,429]" degrees="01" direction="north" minutes="23.581" orientation="latitude" pageId="20" pageNumber="378" precision="1" value="1.3930167">01°23.581'N</geoCoordinate>
<geoCoordinate id="EE40500BFFAA12502C2EFE3D80D31EF9" box="[1111,1263,402,429]" degrees="103" direction="east" minutes="59.593" orientation="longitude" pageId="20" pageNumber="378" precision="1" value="103.99322">103°59.593'E</geoCoordinate>
01°23.529'N 103°59.990'E,
<quantity id="4C8C9B29FFAA12502938FE1985FA1E85" box="[321,454,438,465]" metricMagnitude="1" metricUnit="m" metricValue="1.015" metricValueMax="1.47" metricValueMin="0.5599999999999999" pageId="20" pageNumber="378" unit="m" value="10.149999999999999" valueMax="14.7" valueMin="5.6">5.614.7 m</quantity>
, coll.
<collectorName id="2681531AFFAA12502A69FE1880371E85" box="[528,1035,439,465]" pageId="20" pageNumber="378">Comprehensive Marine Biodiversity Survey</collectorName>
, dredge,
<date id="FFCA100CFFAA12502C01FE1981031E85" box="[1144,1343,438,465]" pageId="20" pageNumber="378" value="2012-10-25">
<collectingDate id="EF8EE9E4FFAA12502C01FE1981031E85" box="[1144,1343,438,465]" pageId="20" pageNumber="378" value="2012-10-25">25 October 2012</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502D34FE18862E1D69" collectingDate="2014-01-17" collectionCode="ZRC" collectorName="Comprehensive Marine Biodiversity Survey" country="Singapore" elevation="15" latitude="1.4040333" location="Ketem" longLatPrecision="1" longitude="103.94" municipality="Pasir Ris" pageId="20" pageNumber="378" specimenCode="ZRC 2022.1004" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502D34FE1881A01E85" box="[1357,1436,439,465]" count="1" pageId="20" pageNumber="378" type="male">1 male</specimenCount>
(14.2 ×
<quantity id="4C8C9B29FFAA12502896FE74856B1EA1" box="[239,343,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="20" pageNumber="378" unit="mm" value="13.9">13.9 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502913FE74861D1EA1" box="[362,545,475,501]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2022.1004</specimenCode>
, specimen SEA 3930, station DR277, between
<location id="8EAB6017FFAA12502C48FE7480401EA1" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502C48FE7480401EA1" box="[1073,1148,475,501]" country="Singapore" latitude="1.4040333" longLatPrecision="1" longitude="103.94" municipality="Pasir Ris" name="Ketem" pageId="20" pageNumber="378">Ketem</location>
and
<collectingMunicipality id="6BAFACB6FFAA12502CCBFE7481241EA1" box="[1202,1304,475,501]" pageId="20" pageNumber="378">Pasir Ris</collectingMunicipality>
,
<collectingCountry id="F363765CFFAA12502D5DFE7481A41EA1" box="[1316,1432,475,501]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
,
<geoCoordinate id="EE40500BFFAA125028EEFE5085181D4D" box="[151,292,511,537]" degrees="01" direction="north" minutes="24.126" orientation="latitude" pageId="20" pageNumber="378" precision="1" value="1.4021">01°24.126'N</geoCoordinate>
<geoCoordinate id="EE40500BFFAA12502957FE5185FA1D4D" box="[302,454,510,537]" degrees="103" direction="east" minutes="56.568" orientation="longitude" pageId="20" pageNumber="378" precision="1" value="103.9428">103°56.568'E</geoCoordinate>
01°24.242'N 103°56.400'E,
<quantity id="4C8C9B29FFAA12502B5FFE50878B1D4D" box="[806,951,511,537]" metricMagnitude="1" metricUnit="m" metricValue="1.46" metricValueMax="1.48" metricValueMin="1.44" pageId="20" pageNumber="378" unit="m" value="14.6" valueMax="14.8" valueMin="14.4">14.414.8 m</quantity>
, coll.
<collectorName id="2681531AFFAA12502B86FE5084DB1D69" pageId="20" pageNumber="378">Comprehensive Marine Biodiversity Survey</collectorName>
, dredge,
<date id="FFCA100CFFAA12502928FD8C862E1D69" box="[337,530,547,573]" pageId="20" pageNumber="378" value="2014-01-17">
<collectingDate id="EF8EE9E4FFAA12502928FD8C862E1D69" box="[337,530,547,573]" pageId="20" pageNumber="378" value="2014-01-17">17 January 2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502A66FD8C85611DD1" collectingDate="2012-10-12" collectionCode="ZRC" collectorName="Comprehensive Marine Biodiversity Survey" country="Singapore" elevation="9" latitude="1.4215833" location="Chek Jawa" longLatPrecision="1" longitude="103.99408" pageId="20" pageNumber="378" specimenCode="ZRC 2022.1005" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502A66FD8C86571D69" box="[543,619,547,573]" count="1" pageId="20" pageNumber="378" type="male">1 male</specimenCount>
(17.3 ×
<quantity id="4C8C9B29FFAA12502AA8FD8C87071D69" box="[721,827,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="20" pageNumber="378" unit="mm" value="17.2">17.2 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502B29FD8C80351D69" box="[848,1033,546,573]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2022.1005</specimenCode>
, station DW28, east of
<location id="8EAB6017FFAA12502D60FD8C81A41D69" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502D60FD8C81A41D69" box="[1305,1432,547,573]" country="Singapore" latitude="1.4215833" longLatPrecision="1" longitude="103.99408" name="Chek Jawa" pageId="20" pageNumber="378">Chek Jawa</location>
,
<collectingCountry id="F363765CFFAA125028EEFDE885371D35" box="[151,267,583,609]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
,
<geoCoordinate id="EE40500BFFAA12502967FDE985971D35" box="[286,427,582,609]" degrees="01" direction="north" minutes="25.295" orientation="latitude" pageId="20" pageNumber="378" precision="1" value="1.4215833">01°25.295'N</geoCoordinate>
<geoCoordinate id="EE40500BFFAA125029C0FDE986681D35" box="[441,596,582,609]" degrees="103" direction="east" minutes="59.645" orientation="longitude" pageId="20" pageNumber="378" precision="1" value="103.99408">103°59.645'E</geoCoordinate>
,
<quantity id="4C8C9B29FFAA12502A1FFDE886DC1D35" box="[614,736,583,609]" metricMagnitude="0" metricUnit="m" metricValue="8.899999999999999" metricValueMax="9.6" metricValueMin="8.2" pageId="20" pageNumber="378" unit="m" value="8.899999999999999" valueMax="9.6" valueMin="8.2">8.29.6 m</quantity>
, coll.
<collectorName id="2681531AFFAA12502B4BFDE8810B1D35" box="[818,1335,583,609]" pageId="20" pageNumber="378">Comprehensive Marine Biodiversity Survey</collectorName>
, dredge,
<date id="FFCA100CFFAA125028EEFDC485611DD1" box="[151,349,619,645]" pageId="20" pageNumber="378" value="2012-10-12">
<collectingDate id="EF8EE9E4FFAA125028EEFDC485611DD1" box="[151,349,619,645]" pageId="20" pageNumber="378" value="2012-10-12">12 October 2012</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502912FDC484EF1D99" collectingDate="2012-10-19" collectionCode="ZRC" collectorName="Comprehensive Marine Biodiversity Survey" country="Singapore" elevation="18" latitude="1.4090834" location="Pulau Serangoon" longLatPrecision="1" longitude="103.9332" pageId="20" pageNumber="378" specimenCode="ZRC 2022.1006" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502912FDC485F21DD1" box="[363,462,619,645]" count="1" pageId="20" pageNumber="378" type="female">1 female</specimenCount>
(14.5 ×
<quantity id="4C8C9B29FFAA12502A4FFDC486A21DD1" box="[566,670,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="20" pageNumber="378" unit="mm" value="14.3">14.3 mm</quantity>
)
<specimenCode id="DBD29EB7FFAA12502AC9FDC487561DD1" box="[688,874,619,645]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2022.1006</specimenCode>
, station DW36, off
<location id="8EAB6017FFAA12502C28FDC4812A1DD1" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502C28FDC4812A1DD1" box="[1105,1302,619,645]" country="Singapore" latitude="1.4090834" longLatPrecision="1" longitude="103.9332" name="Pulau Serangoon" pageId="20" pageNumber="378">Pulau Serangoon</location>
,
<collectingCountry id="F363765CFFAA12502D5DFDC481A41DD1" box="[1316,1432,619,645]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
,
<geoCoordinate id="EE40500BFFAA125028EEFD2185181DFD" box="[151,292,654,681]" degrees="01" direction="north" minutes="24.545" orientation="latitude" pageId="20" pageNumber="378" precision="1" value="1.4090834">01°24.545'N</geoCoordinate>
<geoCoordinate id="EE40500BFFAA12502956FD2185F61DFD" box="[303,458,654,681]" degrees="103" direction="east" minutes="55.992" orientation="longitude" pageId="20" pageNumber="378" precision="1" value="103.9332">103°55.992'E</geoCoordinate>
,
<quantity id="4C8C9B29FFAA125029A0FD2086571DFD" box="[473,619,655,681]" metricMagnitude="1" metricUnit="m" metricValue="1.77" metricValueMax="1.86" metricValueMin="1.68" pageId="20" pageNumber="378" unit="m" value="17.7" valueMax="18.6" valueMin="16.8">16.818.6 m</quantity>
, coll.
<collectorName id="2681531AFFAA12502ACCFD2080931DFD" box="[693,1199,655,681]" pageId="20" pageNumber="378">Comprehensive Marine Biodiversity Survey</collectorName>
, dredge,
<date id="FFCA100CFFAA12502D65FD2084EF1D99" pageId="20" pageNumber="378" value="2012-10-19">
<collectingDate id="EF8EE9E4FFAA12502D65FD2084EF1D99" pageId="20" pageNumber="378" value="2012-10-19">19 October 2012</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA125028A4FD1C86CE1C41" collectingDate="2014-03-20" collectionCode="ZRC" collectorName="Comprehensive Marine Biodiversity Survey" country="Singapore" elevation="9" latitude="1.42145" location="Noordin Beach" longLatPrecision="1" longitude="103.96066" municipality="Pulau Ubin" pageId="20" pageNumber="378" specimenCode="ZRC 2022.1007" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA125028A4FD1C85021D99" box="[221,318,691,717]" count="1" pageId="20" pageNumber="378" type="female">1 female</specimenCount>
(15.5 ×
<quantity id="4C8C9B29FFAA125029E2FD1D863F1D99" box="[411,515,690,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="20" pageNumber="378" unit="mm" value="15.3">15.3 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502A6CFD1C86F01D99" box="[533,716,691,717]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2022.1007</specimenCode>
, specimen SEA 5374, station DR333, in front of
<location id="8EAB6017FFAA12502C95FD1C81A41D99" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502C95FD1C81A41D99" box="[1260,1432,691,717]" country="Singapore" latitude="1.42145" longLatPrecision="1" longitude="103.96066" municipality="Pulau Ubin" name="Noordin Beach" pageId="20" pageNumber="378">Noordin Beach</location>
,
<collectingMunicipality id="6BAFACB6FFAA125028EEFD7885261DA5" box="[151,282,727,753]" pageId="20" pageNumber="378">Pulau Ubin</collectingMunicipality>
,
<collectingCountry id="F363765CFFAA1250295EFD7885A71DA5" box="[295,411,727,753]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
,
<geoCoordinate id="EE40500BFFAA125029DEFD7986091DA5" box="[423,565,726,753]" degrees="01" direction="north" minutes="25.248" orientation="latitude" pageId="20" pageNumber="378" precision="1" value="1.4208">01°25.248'N</geoCoordinate>
<geoCoordinate id="EE40500BFFAA12502A44FD7986EA1DA5" box="[573,726,726,753]" degrees="103" direction="east" minutes="57.898" orientation="longitude" pageId="20" pageNumber="378" precision="1" value="103.964966">103°57.898'E</geoCoordinate>
01°25.287'N 103°57.640'E,
<quantity id="4C8C9B29FFAA12502C4BFD78809A1DA5" box="[1074,1190,727,753]" metricMagnitude="0" metricUnit="m" metricValue="8.6" metricValueMax="9.3" metricValueMin="7.9" pageId="20" pageNumber="378" unit="m" value="8.6" valueMax="9.3" valueMin="7.9">7.99.3 m</quantity>
, coll.
<collectorName id="2681531AFFAA12502C95FD7885E91C41" pageId="20" pageNumber="378">Comprehensive Marine Biodiversity Survey</collectorName>
, dredge,
<date id="FFCA100CFFAA12502A46FD5486CE1C41" box="[575,754,763,789]" pageId="20" pageNumber="378" value="2014-03-20">
<collectingDate id="EF8EE9E4FFAA12502A46FD5486CE1C41" box="[575,754,763,789]" pageId="20" pageNumber="378" value="2014-03-20">20 March 2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502A86FD5486AF1C09" collectingDate="2012-10-19" collectionCode="ZRC" collectorName="Comprehensive Marine Biodiversity Survey" country="Singapore" elevation="19" latitude="1.3966167" location="Changi Radar station" longLatPrecision="1" longitude="103.97919" pageId="20" pageNumber="378" specimenCode="ZRC 2022.1008" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502A86FD5487711C41" box="[767,845,763,789]" count="1" pageId="20" pageNumber="378" type="male">1 male</specimenCount>
(11.4 × 11.0 mm),
<specimenCode id="DBD29EB7FFAA12502C49FD5480D51C41" box="[1072,1257,763,789]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2022.1008</specimenCode>
, station DW40, opposite
<location id="8EAB6017FFAA12502886FCB085CC1C6D" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502886FCB085CC1C6D" box="[255,496,799,825]" country="Singapore" latitude="1.3966167" longLatPrecision="1" longitude="103.97919" name="Changi Radar station" pageId="20" pageNumber="378">Changi Radar station</location>
,
<collectingCountry id="F363765CFFAA12502984FCB0864D1C6D" box="[509,625,799,825]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
,
<geoCoordinate id="EE40500BFFAA12502A07FCB087371C6D" box="[638,779,799,825]" degrees="01" direction="north" minutes="23.797" orientation="latitude" pageId="20" pageNumber="378" precision="1" value="1.3966167">01°23.797'N</geoCoordinate>
<geoCoordinate id="EE40500BFFAA12502B6DFCB1878C1C6D" box="[788,944,798,825]" degrees="103" direction="east" minutes="58.751" orientation="longitude" pageId="20" pageNumber="378" precision="1" value="103.97919">103°58.751'E</geoCoordinate>
,
<quantity id="4C8C9B29FFAA12502BC5FCB080701C6D" box="[956,1100,799,825]" metricMagnitude="1" metricUnit="m" metricValue="1.905" metricValueMax="2.4699999999999998" metricValueMin="1.34" pageId="20" pageNumber="378" unit="m" value="19.05" valueMax="24.7" valueMin="13.4">
<elevation id="0059D1FFFFAA12502BC5FCB080701C6D" box="[956,1100,799,825]" metricMagnitude="1" metricUnit="m" metricValue="1.905" metricValueMax="2.4699999999999998" metricValueMin="1.34" pageId="20" pageNumber="378" unit="m" value="19.05" valueMax="24.7" valueMin="13.4">13.424.7 m</elevation>
</quantity>
, coll.
<collectorName id="2681531AFFAA12502CEAFCB0854A1C09" pageId="20" pageNumber="378">Comprehensive Marine Biodiversity Survey</collectorName>
, dredge,
<date id="FFCA100CFFAA125029AEFCEC86AF1C09" box="[471,659,835,861]" pageId="20" pageNumber="378" value="2012-10-19">
<collectingDate id="EF8EE9E4FFAA125029AEFCEC86AF1C09" box="[471,659,835,861]" pageId="20" pageNumber="378" value="2012-10-19">19 October 2012</collectingDate>
</date>
</materialsCitation>
.
<location id="8EAB6017FFAA12502AE3FCEC86D51C09" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502AE3FCEC86D51C09" box="[666,745,835,861]" country="Singapore" name="Others" pageId="20" pageNumber="378">Others</location>
:
<materialsCitation id="3B1C3C91FFAA12502A88FCEC86711CD5" collectionCode="ZRC" collectorName="D. Vandenspiegel" country="Singapore" location="East Coast" pageId="20" pageNumber="378" specimenCode="ZRC 1995.0547" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502A88FCEC87011C09" box="[753,829,835,861]" count="1" pageId="20" pageNumber="378" type="male">1 male</specimenCount>
,
<specimenCount id="9D72FD45FFAA12502B3FFCEC879B1C09" box="[838,935,835,861]" count="1" pageId="20" pageNumber="378" type="female">1 female</specimenCount>
,
<specimenCode id="DBD29EB7FFAA12502BC9FCEC80591C09" box="[944,1125,834,861]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1995.0547</specimenCode>
,
<location id="8EAB6017FFAA12502C17FCEC80DA1C09" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502C17FCEC80DA1C09" box="[1134,1254,835,861]" country="Singapore" name="East Coast" pageId="20" pageNumber="378">East Coast</location>
,
<collectingCountry id="F363765CFFAA12502C97FCEC815E1C09" box="[1262,1378,835,861]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
, coll. dredge,
<collectorName id="2681531AFFAA1250288BFCC8858B1CD5" box="[242,439,871,897]" pageId="20" pageNumber="378">D. Vandenspiegel</collectorName>
<emphasis id="B900EADEFFAA125029C6FCC885CD1CD5" box="[447,497,871,897]" italics="true" pageId="20" pageNumber="378">et al</emphasis>
., 1990s
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502A20FCC886BB1CF1" collectionCode="ZRC" collectorName="D. Vandenspiegel" country="Singapore" location="East Coast" pageId="20" pageNumber="378" specimenCode="ZRC 1995.0546" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502A20FCC886F41CD5" box="[601,712,871,897]" count="1" pageId="20" pageNumber="378" type="juvenile">1 juvenile</specimenCount>
male (badly damaged),
<specimenCode id="DBD29EB7FFAA12502BA3FCC880AE1CD5" box="[986,1170,870,897]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1995.0546</specimenCode>
,
<location id="8EAB6017FFAA12502CE7FCC881241CD5" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502CE7FCC881241CD5" box="[1182,1304,871,897]" country="Singapore" name="East Coast" pageId="20" pageNumber="378">East Coast</location>
,
<collectingCountry id="F363765CFFAA12502D5DFCC881A41CD5" box="[1316,1432,871,897]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
, coll. dredge,
<collectorName id="2681531AFFAA12502952FC2485CC1CF1" box="[299,496,907,933]" pageId="20" pageNumber="378">D. Vandenspiegel</collectorName>
<emphasis id="B900EADEFFAA12502980FC2486171CF1" box="[505,555,907,933]" italics="true" pageId="20" pageNumber="378">et al</emphasis>
., 1990s
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502AEAFC2485E51C9D" collectingDate="1982-05-09" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Singapore" location="Changi Point" pageId="20" pageNumber="378" specimenCode="ZRC 1985.1631" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502AEAFC2486E31CF1" box="[659,735,907,933]" count="1" pageId="20" pageNumber="378" type="male">1 male</specimenCount>
(18.4 × 19.0 mm),
<specimenCode id="DBD29EB7FFAA12502BC4FC2480491CF1" box="[957,1141,906,933]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1985.1631</specimenCode>
,
<location id="8EAB6017FFAA12502CF8FC2481241CF1" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502CF8FC2481241CF1" box="[1153,1304,907,933]" country="Singapore" name="Changi Point" pageId="20" pageNumber="378">Changi Point</location>
,
<collectingCountry id="F363765CFFAA12502D5DFC2481A41CF1" box="[1316,1432,907,933]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
, coll.
<collectorName id="2681531AFFAA125028A9FC00857F1C9D" box="[208,323,943,969]" pageId="20" pageNumber="378">P.K.L. Ng</collectorName>
,
<date id="FFCA100CFFAA12502937FC0085E51C9D" box="[334,473,943,969]" pageId="20" pageNumber="378" value="1982-05-09">
<collectingDate id="EF8EE9E4FFAA12502937FC0085E51C9D" box="[334,473,943,969]" pageId="20" pageNumber="378" value="1982-05-09">9 May 1982</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA1250299CFC0087A91CB9" collectionCode="ZRC" collectorName="D. S. Johnson" country="Singapore" elevation="17" location="Bedok" pageId="20" pageNumber="378" specimenCode="ZRC 1985.0090" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA1250299CFC0086691C9D" box="[485,597,943,969]" count="1" pageId="20" pageNumber="378" type="juvenile">1 juvenile</specimenCount>
male (12.7 ×
<quantity id="4C8C9B29FFAA12502A8AFC0087671C9D" box="[755,859,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="20" pageNumber="378" unit="mm" value="12.4">12.4 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502B16FC00801B1C9D" box="[879,1063,942,969]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1985.0090</specimenCode>
, station B77, on mud, off
<location id="8EAB6017FFAA12502D35FC0081A41C9D" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502D35FC0081A41C9D" box="[1356,1432,943,969]" country="Singapore" name="Bedok" pageId="20" pageNumber="378">Bedok</location>
,
<collectingCountry id="F363765CFFAA125028EEFC7C85371CB9" box="[151,267,979,1005]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
,
<quantity id="4C8C9B29FFAA12502963FC7D85501CB9" box="[282,364,978,1005]" metricMagnitude="1" metricUnit="m" metricValue="1.65" pageId="20" pageNumber="378" unit="m" value="16.5">
<elevation id="0059D1FFFFAA12502963FC7D85501CB9" box="[282,364,978,1005]" metricMagnitude="1" metricUnit="m" metricValue="1.65" pageId="20" pageNumber="378" unit="m" value="16.5">16.5 m</elevation>
</quantity>
(9 fathoms), coll. dredge,
<collectorName id="2681531AFFAA12502ADCFC7C877D1CB9" box="[677,833,979,1005]" pageId="20" pageNumber="378">D.S. Johnson</collectorName>
, 1960s
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502BDDFC7C80511B45" collectingDate="1989-02-23" collectionCode="ZRC" collectorName="Reef Ecology &amp; Study Team" country="Singapore" location="Kallang Basin" pageId="20" pageNumber="378" specimenCode="ZRC 1990.8405" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502BDDFC7C802A1CB9" box="[932,1046,979,1005]" count="1" pageId="20" pageNumber="378" type="juvenile">1 juvenile</specimenCount>
male,
<specimenCode id="DBD29EB7FFAA12502C10FC7C81181CB9" box="[1129,1316,978,1005]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1990.8405</specimenCode>
, station 6,
<location id="8EAB6017FFAA125028EEFC5885061B45" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA125028EEFC5885061B45" box="[151,314,1015,1041]" country="Singapore" name="Kallang Basin" pageId="20" pageNumber="378">Kallang Basin</location>
,
<collectingCountry id="F363765CFFAA1250293DFC5885841B45" box="[324,440,1015,1041]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
, coll.
<collectorName id="2681531AFFAA12502982FC5886AF1B45" box="[507,659,1015,1041]" pageId="20" pageNumber="378">Reef Ecology</collectorName>
&amp;
<collectorName id="2681531AFFAA12502ACEFC58877C1B45" box="[695,832,1015,1041]" pageId="20" pageNumber="378">Study Team</collectorName>
, dredge,
<date id="FFCA100CFFAA12502BDDFC5880511B45" box="[932,1133,1015,1041]" pageId="20" pageNumber="378" value="1989-02-23">
<collectingDate id="EF8EE9E4FFAA12502BDDFC5880511B45" box="[932,1133,1015,1041]" pageId="20" pageNumber="378" value="1989-02-23">23 February 1989</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502C00FC5884EE1B0D" collectingDate="1989-02-22" collectionCode="ZRC" collectorName="Reef Ecology &amp; Study Team" country="Singapore" location="Kallang Basin" pageId="20" pageNumber="378" specimenCode="ZRC 1990.8329" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502C00FC5880DB1B45" box="[1145,1255,1015,1041]" count="1" pageId="20" pageNumber="378" type="juvenile">1 juvenile</specimenCount>
male (crushed),
<specimenCode id="DBD29EB7FFAA125028EEFBB4856F1B61" box="[151,339,1051,1077]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1990.8329</specimenCode>
, station 6,
<location id="8EAB6017FFAA125029A1FBB486421B61" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA125029A1FBB486421B61" box="[472,638,1051,1077]" country="Singapore" name="Kallang Basin" pageId="20" pageNumber="378">Kallang Basin</location>
,
<collectingCountry id="F363765CFFAA12502AF4FBB4873D1B61" box="[653,769,1051,1077]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
, coll.
<collectorName id="2681531AFFAA12502B34FBB487D61B61" box="[845,1002,1051,1077]" pageId="20" pageNumber="378">Reef Ecology</collectorName>
&amp;
<collectorName id="2681531AFFAA12502C6FFBB4809F1B61" box="[1046,1187,1051,1077]" pageId="20" pageNumber="378">Study Team</collectorName>
, dredge,
<date id="FFCA100CFFAA12502D69FBB484EE1B0D" pageId="20" pageNumber="378" value="1989-02-22">
<collectingDate id="EF8EE9E4FFAA12502D69FBB484EE1B0D" pageId="20" pageNumber="378" value="1989-02-22">22 February 1989</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA125028A6FB9085541B29" collectingDate="1987-05-12" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Singapore" location="Southern Islands" pageId="20" pageNumber="378" specimenCode="ZRC 1989.2943, 2944" specimenCount="2" specimenCount-juvenile="2" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA125028A6FB9085721B0D" box="[223,334,1087,1113]" count="1" pageId="20" pageNumber="378" type="juvenile">1 juvenile</specimenCount>
male,
<specimenCount id="9D72FD45FFAA125029E2FB9086371B0D" box="[411,523,1087,1113]" count="1" pageId="20" pageNumber="378" type="juvenile">1 juvenile</specimenCount>
female,
<specimenCode id="DBD29EB7FFAA12502A14FB9087181B0D" box="[621,804,1087,1113]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1989.2943</specimenCode>
<specimenCode id="DBD29EB7FFAA12502B4BFB9087571B0D" box="[818,875,1087,1113]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">2944</specimenCode>
,
<location id="8EAB6017FFAA12502B0EFB90800A1B0D" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502B0EFB90800A1B0D" box="[887,1078,1087,1113]" country="Singapore" name="Southern Islands" pageId="20" pageNumber="378">Southern Islands</location>
,
<collectingCountry id="F363765CFFAA12502C3BFB90808A1B0D" box="[1090,1206,1087,1113]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
, coll. dredge,
<collectorName id="2681531AFFAA12502D2CFB9084801B29" pageId="20" pageNumber="378">P.K.L. Ng</collectorName>
,
<date id="FFCA100CFFAA125028B3FBCC85541B29" box="[202,360,1123,1149]" pageId="20" pageNumber="378" value="1987-05-12">
<collectingDate id="EF8EE9E4FFAA125028B3FBCC85541B29" box="[202,360,1123,1149]" pageId="20" pageNumber="378" value="1987-05-12">12 May 1987</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA1250290FFBCC871A1B29" box="[374,806,1123,1149]" pageId="20" pageNumber="378" specimenCount="1" specimenCount-male="1" typeStatus="type">
<specimenCount id="9D72FD45FFAA1250290FFBCC85F81B29" box="[374,452,1123,1149]" count="1" pageId="20" pageNumber="378" type="male">1 male</specimenCount>
(spiny type, 18.9 ×
<quantity id="4C8C9B29FFAA12502ACAFBCC87221B29" box="[691,798,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="20" pageNumber="378" unit="mm" value="18.7">18.7 mm</quantity>
)
</materialsCitation>
,
<materialsCitation id="3B1C3C91FFAA12502B4DFBCC81BD1BF5" collectingDate="1997-08-05" collectionCode="ZRC" collectorName="L. W. Loh" country="Singapore" elevation="15" latitude="1.40365" location="Pulau Sajahat" longLatPrecision="1" longitude="104.01273" pageId="20" pageNumber="378" specimenCode="ZRC 1999.0507" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502B4DFBCC879E1B29" box="[820,930,1123,1149]" count="4" pageId="20" pageNumber="378" type="female">4 females</specimenCount>
(largest 16.8 ×
<quantity id="4C8C9B29FFAA12502C27FBCC80F41B29" box="[1118,1224,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="20" pageNumber="378" unit="mm" value="16.4">16.4 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502CA7FBCC81A41B29" box="[1246,1432,1122,1149]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1999.0507</specimenCode>
, near
<location id="8EAB6017FFAA125028A9FB2885521BF5" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA125028A9FB2885521BF5" box="[208,366,1159,1185]" country="Singapore" latitude="1.40365" longLatPrecision="1" longitude="104.01273" name="Pulau Sajahat" pageId="20" pageNumber="378">Pulau Sajahat</location>
,
<collectingCountry id="F363765CFFAA12502902FB2885D31BF5" box="[379,495,1159,1185]" name="Singapore" pageId="20" pageNumber="378">Singapore</collectingCountry>
,
<geoCoordinate id="EE40500BFFAA12502984FB2886BC1BF5" box="[509,640,1159,1185]" degrees="1" direction="north" minutes="24.219" orientation="latitude" pageId="20" pageNumber="378" precision="1" value="1.40365">1°24.219N</geoCoordinate>
<geoCoordinate id="EE40500BFFAA12502AF0FB2887141BF5" box="[649,808,1159,1185]" degrees="104" direction="east" minutes="00.764" orientation="longitude" pageId="20" pageNumber="378" precision="1" value="104.01273">104°00.764E</geoCoordinate>
,
<quantity id="4C8C9B29FFAA12502B4FFB29874F1BF5" box="[822,883,1158,1185]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="378" unit="m" value="15.0">
<elevation id="0059D1FFFFAA12502B4FFB29874F1BF5" box="[822,883,1158,1185]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="378" unit="m" value="15.0">15 m</elevation>
</quantity>
, coll. dredge,
<collectorName id="2681531AFFAA12502C6EFB2880BF1BF5" box="[1047,1155,1159,1185]" pageId="20" pageNumber="378">L.W. Loh</collectorName>
<emphasis id="B900EADEFFAA12502CF4FB2880FC1BF5" box="[1165,1216,1159,1185]" italics="true" pageId="20" pageNumber="378">et al</emphasis>
.,
<date id="FFCA100CFFAA12502CAFFB2981BD1BF5" box="[1238,1409,1158,1185]" pageId="20" pageNumber="378" value="1997-08-05">
<collectingDate id="EF8EE9E4FFAA12502CAFFB2981BD1BF5" box="[1238,1409,1158,1185]" pageId="20" pageNumber="378" value="1997-08-05">5 August 1997</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502DF7FB2885181BBD" collectingDate="1969-03-05" collectionCode="ZRC" collectorName="C. Leong" country="Malaysia" location="Mersing" municipality="Peninsular" pageId="20" pageNumber="378" specimenCode="ZRC 1985.1442, 1444" specimenCount="1" specimenCount-male="1" stateProvince="Johor" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502DF7FB2884EC1B91" count="1" pageId="20" pageNumber="378" type="male">1 male</specimenCount>
, 2 ovigerous females,
<specimenCode id="DBD29EB7FFAA125029B6FB0486B41B91" box="[463,648,1194,1221]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1985.1442</specimenCode>
<specimenCode id="DBD29EB7FFAA12502AEFFB0586F11B91" box="[662,717,1194,1221]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">1444</specimenCode>
,
<location id="8EAB6017FFAA12502AA0FB0487051B91" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502AA0FB0487051B91" box="[729,825,1195,1221]" country="Malaysia" municipality="Peninsular" name="Mersing" pageId="20" pageNumber="378" stateProvince="Johor">Mersing</location>
, eastern
<collectingRegion id="49B0F82EFFAA12502BE4FB0487E11B91" box="[925,989,1195,1221]" country="Malaysia" name="Johor" pageId="20" pageNumber="378">Johor</collectingRegion>
, Peninsular
<collectingCountry id="F363765CFFAA12502C1EFB0480EC1B91" box="[1127,1232,1195,1221]" name="Malaysia" pageId="20" pageNumber="378">Malaysia</collectingCountry>
, coll.
<collectorName id="2681531AFFAA12502D6FFB0481BE1B91" box="[1302,1410,1195,1221]" pageId="20" pageNumber="378">C. Leong</collectorName>
,
<date id="FFCA100CFFAA12502DF7FB0585181BBD" pageId="20" pageNumber="378" value="1969-03-05">
<collectingDate id="EF8EE9E4FFAA12502DF7FB0585181BBD" pageId="20" pageNumber="378" value="1969-03-05">5 March 1969</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA1250294BFB6085BF1A65" collectionCode="ZRC" collectorName="D. S. Johnson" country="Malaysia" county="Peninsular" elevation="31" latitude="1.35" location="Tanjung Stapa" longLatPrecision="1" longitude="104.13333" municipality="Pengerang" pageId="20" pageNumber="378" specimenCode="ZRC 1985.0097" specimenCount="1" specimenCount-juvenile="1" stateProvince="Johor" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA1250294BFB60859E1BBD" box="[306,418,1231,1257]" count="1" pageId="20" pageNumber="378" type="juvenile">1 juvenile</specimenCount>
male (12.6 ×
<quantity id="4C8C9B29FFAA12502A3FFB60868C1BBD" box="[582,688,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="378" unit="mm" value="12.7">12.7 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502ABCFB6087421BBD" box="[709,894,1230,1257]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1985.0097</specimenCode>
, station B78, mud substrate, off
<location id="8EAB6017FFAA12502C8AFB6081A41BBD" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502C8AFB6081A41BBD" box="[1267,1432,1231,1257]" country="Malaysia" county="Peninsular" latitude="1.35" longLatPrecision="1" longitude="104.13333" municipality="Pengerang" name="Tanjung Stapa" pageId="20" pageNumber="378" stateProvince="Johor">Tanjung Stapa</location>
,
<collectingMunicipality id="6BAFACB6FFAA125028EEFB5C852C1A59" box="[151,272,1267,1293]" pageId="20" pageNumber="378">Pengerang</collectingMunicipality>
, southeastern
<collectingRegion id="49B0F82EFFAA125029D6FB5C85D31A59" box="[431,495,1267,1293]" country="Malaysia" name="Johor" pageId="20" pageNumber="378">Johor</collectingRegion>
,
<collectingCounty id="62AA4E40FFAA12502982FB5C864D1A59" box="[507,625,1267,1293]" pageId="20" pageNumber="378">Peninsular</collectingCounty>
<collectingCountry id="F363765CFFAA12502A01FB5C86DD1A59" box="[632,737,1267,1293]" name="Malaysia" pageId="20" pageNumber="378">Malaysia</collectingCountry>
,
<geoCoordinate id="EE40500BFFAA12502A94FB5C87441A59" box="[749,888,1267,1293]" degrees="1" direction="north" minutes="21" orientation="latitude" pageId="20" pageNumber="378" precision="1" seconds="00.0" value="1.35">1°21'00.0&quot;N</geoCoordinate>
<geoCoordinate id="EE40500BFFAA12502B06FB5C801A1A59" box="[895,1062,1267,1293]" degrees="104" direction="east" minutes="08" orientation="longitude" pageId="20" pageNumber="378" precision="1" seconds="00.0" value="104.13333">104°08'00.0&quot;E</geoCoordinate>
,
<quantity id="4C8C9B29FFAA12502C4BFB5C80571A59" box="[1074,1131,1267,1293]" metricMagnitude="1" metricUnit="m" metricValue="3.1" pageId="20" pageNumber="378" unit="m" value="31.0">
<elevation id="0059D1FFFFAA12502C4BFB5C80571A59" box="[1074,1131,1267,1293]" metricMagnitude="1" metricUnit="m" metricValue="3.1" pageId="20" pageNumber="378" unit="m" value="31.0">31 m</elevation>
</quantity>
(17 fathoms), coll. dredge,
<collectorName id="2681531AFFAA125028EEFAB8850C1A65" box="[151,304,1303,1329]" pageId="20" pageNumber="378">D.S. Johnson</collectorName>
, 1960s
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA125029F6FAB884EE1A01" collectingDate="2003-08-28" collectionCode="ZRC" country="Thailand" location="Pattani" pageId="20" pageNumber="378" specimenCode="ZRC 2011.0646" specimenCount="1" specimenCount-male="1" stateProvince="Pattani" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA125029F6FAB885E71A65" box="[399,475,1303,1329]" count="1" pageId="20" pageNumber="378" type="male">1 male</specimenCount>
(18.00 ×
<quantity id="4C8C9B29FFAA12502A34FAB8868A1A65" box="[589,694,1303,1329]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="20" pageNumber="378" unit="mm" value="17.7">17.7 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502AB3FAB887BE1A65" box="[714,898,1303,1329]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2011.0646</specimenCode>
, fishing port,
<collectingRegion id="49B0F82EFFAA12502C59FAB8804D1A65" box="[1056,1137,1303,1329]" country="Thailand" name="Pattani" pageId="20" pageNumber="378">Pattani</collectingRegion>
,
<collectingCountry id="F363765CFFAA12502C04FAB880DF1A65" box="[1149,1251,1303,1329]" name="Thailand" pageId="20" pageNumber="378">Thailand</collectingCountry>
, coll.
<date id="FFCA100CFFAA12502D50FAB884EE1A01" pageId="20" pageNumber="378" value="2003-08-28">
<collectingDate id="EF8EE9E4FFAA12502D50FAB884EE1A01" pageId="20" pageNumber="378" value="2003-08-28">28 August 2003</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA125028A4FA95877C1A01" box="[221,832,1338,1365]" pageId="20" pageNumber="378" specimenCount="25" specimenCount-male="25" typeStatus="type">
<specimenCount id="9D72FD45FFAA125028A4FA95857C1A01" box="[221,320,1338,1365]" count="25" pageId="20" pageNumber="378" type="male">25 males</specimenCount>
(smooth type, 18.4 ×
<quantity id="4C8C9B29FFAA12502A4BFA9486A91A01" box="[562,661,1339,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="20" pageNumber="378" unit="mm" value="18.6">18.6 mm</quantity>
photographed)
</materialsCitation>
,
<materialsCitation id="3B1C3C91FFAA12502B33FA9485961A2D" pageId="20" pageNumber="378" specimenCount="26" specimenCount-female="7" specimenCount-male="19">
<specimenCount id="9D72FD45FFAA12502B33FA9487911A01" box="[842,941,1339,1365]" count="19" pageId="20" pageNumber="378" type="male">19 males</specimenCount>
(spiny type, 17.4 ×
<quantity id="4C8C9B29FFAA12502CF3FA9480D01A01" box="[1162,1260,1339,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="20" pageNumber="378" unit="mm" value="17.1">17.1 mm</quantity>
photographed),
<specimenCount id="9D72FD45FFAA125028EEFAF085381A2D" box="[151,260,1375,1401]" count="7" pageId="20" pageNumber="378" type="female">7 females</specimenCount>
(smooth type)
</materialsCitation>
,
<materialsCitation id="3B1C3C91FFAA125029CFFAF086F01AC9" collectingDate="2003-02-20" collectionCode="ZRC" collectorName="D. C. J. Yeo" country="Thailand" location="Fishing Port" municipality="Gulf" pageId="20" pageNumber="378" specimenCode="ZRC 2003.0107" specimenCount="3" specimenCount-female="3" stateProvince="Pattani" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA125029CFFAF0861F1A2D" box="[438,547,1375,1401]" count="3" pageId="20" pageNumber="378" type="female">3 females</specimenCount>
(spiny type),
<specimenCode id="DBD29EB7FFAA12502AB9FAF087441A2D" box="[704,888,1375,1401]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2003.0107</specimenCode>
,
<collectingRegion id="49B0F82EFFAA12502BFAFAF087ED1A2D" box="[899,977,1375,1401]" country="Thailand" name="Pattani" pageId="20" pageNumber="378">Pattani</collectingRegion>
<location id="8EAB6017FFAA12502BA1FAF0805A1A2D" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502BA1FAF0805A1A2D" box="[984,1126,1375,1401]" country="Thailand" municipality="Gulf" name="Fishing Port" pageId="20" pageNumber="378" stateProvince="Pattani">Fishing Port</location>
, Gulf of Thailand, southern
<collectingCountry id="F363765CFFAA125028EEFA2C84C11AC9" box="[151,253,1411,1437]" name="Thailand" pageId="20" pageNumber="378">Thailand</collectingCountry>
, coll.
<collectorName id="2681531AFFAA12502939FA2C85841AC9" box="[320,440,1411,1437]" pageId="20" pageNumber="378">D.C.J. Yeo</collectorName>
<emphasis id="B900EADEFFAA125029C7FA2C85D31AC9" box="[446,495,1411,1437]" italics="true" pageId="20" pageNumber="378">et al</emphasis>
.,
<date id="FFCA100CFFAA12502A7AFA2C86F01AC9" box="[515,716,1411,1437]" pageId="20" pageNumber="378" value="2003-02-20">
<collectingDate id="EF8EE9E4FFAA12502A7AFA2C86F01AC9" box="[515,716,1411,1437]" pageId="20" pageNumber="378" value="2003-02-20">20 February 2003</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502AAEFA2C80DD1AC9" box="[727,1249,1410,1437]" pageId="20" pageNumber="378" specimenCount="4" specimenCount-male="4">
<specimenCount id="9D72FD45FFAA12502AAEFA2C87111AC9" box="[727,813,1411,1437]" count="4" pageId="20" pageNumber="378" type="male">4 males</specimenCount>
(smooth type, largest 21.5 ×
<quantity id="4C8C9B29FFAA12502C0BFA2C80E61AC9" box="[1138,1242,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="20" pageNumber="378" unit="mm" value="22.1">22.1 mm</quantity>
)
</materialsCitation>
,
<materialsCitation id="3B1C3C91FFAA12502C95FA2C80371AB1" collectingDate="1998-10-25" collectionCode="ZRC" collectorName="H. H. Tan" country="Thailand" location="Fishing Port" municipality="Gulf" pageId="20" pageNumber="378" specimenCode="ZRC 1999.0317" specimenCount="14" specimenCount-male="14" stateProvince="Pattani" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502C95FA2C81731AC9" box="[1260,1359,1411,1437]" count="14" pageId="20" pageNumber="378" type="male">14 males</specimenCount>
(spiny type), 3 ovigerous females (spiny type, 17.7 ×
<quantity id="4C8C9B29FFAA12502ADCFA0887361A95" box="[677,778,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="20" pageNumber="378" unit="mm" value="17.7">17.7 mm</quantity>
18.6 ×
<quantity id="4C8C9B29FFAA12502B0FFA0887E31A95" box="[886,991,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="20" pageNumber="378" unit="mm" value="18.1">18.1 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502B8BFA0880961A95" box="[1010,1194,1447,1473]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 1999.0317</specimenCode>
,
<collectingRegion id="49B0F82EFFAA12502CCFFA0881381A95" box="[1206,1284,1447,1473]" country="Thailand" name="Pattani" pageId="20" pageNumber="378">Pattani</collectingRegion>
<location id="8EAB6017FFAA12502D72FA0881A41A95" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502D72FA0881A41A95" box="[1291,1432,1447,1473]" country="Thailand" municipality="Gulf" name="Fishing Port" pageId="20" pageNumber="378" stateProvince="Pattani">Fishing Port</location>
, Gulf of Thailand, southern
<collectingCountry id="F363765CFFAA125029ACFA6486071AB1" box="[469,571,1483,1509]" name="Thailand" pageId="20" pageNumber="378">Thailand</collectingCountry>
, coll.
<collectorName id="2681531AFFAA12502AFCFA6486D31AB1" box="[645,751,1483,1509]" pageId="20" pageNumber="378">H.H. Tan</collectorName>
<emphasis id="B900EADEFFAA12502A80FA6487121AB1" box="[761,814,1483,1509]" italics="true" pageId="20" pageNumber="378">et al</emphasis>
.,
<date id="FFCA100CFFAA12502B3FFA6580371AB1" box="[838,1035,1482,1509]" pageId="20" pageNumber="378" value="1998-10-25">
<collectingDate id="EF8EE9E4FFAA12502B3FFA6580371AB1" box="[838,1035,1482,1509]" pageId="20" pageNumber="378" value="1998-10-25">25 October 1998</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502C62FA64806F195D" collectingDate="2003-10-16" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" location="Fishing Port" municipality="Gulf" pageId="20" pageNumber="378" specimenCode="ZRC 2003.0623" specimenCount="2" specimenCount-male="2" stateProvince="Pattani" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502C62FA64804B1AB1" box="[1051,1143,1483,1509]" count="2" pageId="20" pageNumber="378" type="male">2 males</specimenCount>
,
<specimenCode id="DBD29EB7FFAA12502CFCFA64817C1AB1" box="[1157,1344,1483,1509]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2003.0623</specimenCode>
,
<collectingRegion id="49B0F82EFFAA12502D37FA6481A01AB1" box="[1358,1436,1483,1509]" country="Thailand" name="Pattani" pageId="20" pageNumber="378">Pattani</collectingRegion>
<location id="8EAB6017FFAA125028EEFA408519195D" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA125028EEFA408519195D" box="[151,293,1519,1545]" country="Thailand" municipality="Gulf" name="Fishing Port" pageId="20" pageNumber="378" stateProvince="Pattani">Fishing Port</location>
,
<locality id="8EAF02C1FFAA12502948FA4085CD195D" box="[305,497,1519,1545]" pageId="20" pageNumber="378">Gulf of Thailand</locality>
, southern
<collectingCountry id="F363765CFFAA12502A1FFA4086F0195D" box="[614,716,1519,1545]" name="Thailand" pageId="20" pageNumber="378">Thailand</collectingCountry>
, coll.
<collectorName id="2681531AFFAA12502B68FA4087B9195D" box="[785,901,1519,1545]" pageId="20" pageNumber="378">P.K.L. Ng</collectorName>
,
<date id="FFCA100CFFAA12502BE8FA40806F195D" box="[913,1107,1519,1545]" pageId="20" pageNumber="378" value="2003-10-16">
<collectingDate id="EF8EE9E4FFAA12502BE8FA40806F195D" box="[913,1107,1519,1545]" pageId="20" pageNumber="378" value="2003-10-16">16 October 2003</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D72FD45FFAA12502C26FA41808A195D" box="[1119,1206,1518,1545]" count="5" pageId="20" pageNumber="378" type="male">5 males</specimenCount>
(largest 19.7 ×
<quantity id="4C8C9B29FFAA12502D12FA4084FB1979" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="20" pageNumber="378" unit="mm" value="19.9">19.9 mm</quantity>
),
<materialsCitation id="3B1C3C91FFAA125028A3F9BC87071905" collectingDate="2003-10-15" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" location="Sakom Crab Landing Port" municipality="Gulf" pageId="20" pageNumber="378" specimenCode="ZRC 2003.0609" specimenCount="7" specimenCount-female="7" stateProvince="Songkhla" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA125028A3F9BC85791979" box="[218,325,1555,1581]" count="7" pageId="20" pageNumber="378" type="female">7 females</specimenCount>
(largest 15.5 ×
<quantity id="4C8C9B29FFAA1250298CF9BD86601979" box="[501,604,1554,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="20" pageNumber="378" unit="mm" value="15.5">15.5 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502A16F9BC871A1979" box="[623,806,1555,1581]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2003.0609</specimenCode>
,
<location id="8EAB6017FFAA12502B48F9BC80641979" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502B48F9BC80641979" box="[817,1112,1555,1581]" country="Thailand" municipality="Gulf" name="Sakom Crab Landing Port" pageId="20" pageNumber="378" stateProvince="Songkhla">Sakom Crab Landing Port</location>
,
<collectingRegion id="49B0F82EFFAA12502C1AF9BC80EC1979" box="[1123,1232,1555,1581]" country="Thailand" name="Songkhla" pageId="20" pageNumber="378">Songkhla</collectingRegion>
, Gulf of Thailand, southern
<collectingCountry id="F363765CFFAA12502978F998855B1905" box="[257,359,1591,1617]" name="Thailand" pageId="20" pageNumber="378">Thailand</collectingCountry>
, coll.
<collectorName id="2681531AFFAA125029D6F998861E1905" box="[431,546,1591,1617]" pageId="20" pageNumber="378">P.K.L. Ng</collectorName>
<emphasis id="B900EADEFFAA12502A55F998865C1905" box="[556,608,1591,1617]" italics="true" pageId="20" pageNumber="378">et al</emphasis>
.,
<date id="FFCA100CFFAA12502A0EF99987071905" box="[631,827,1590,1617]" pageId="20" pageNumber="378" value="2003-10-15">
<collectingDate id="EF8EE9E4FFAA12502A0EF99987071905" box="[631,827,1590,1617]" pageId="20" pageNumber="378" value="2003-10-15">15 October 2003</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502B33F99880471921" collectingDate="2003-11-21" collectionCode="ZRC" collectorName="Z. Jaafar" country="Thailand" location="Angsila Fishing Port" municipality="Gulf" pageId="20" pageNumber="378" specimenCode="ZRC 2015.0439" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA12502B33F99887AB1905" box="[842,919,1591,1617]" count="1" pageId="20" pageNumber="378" type="male">1 male</specimenCount>
(16.7 × 17.0 mm),
<specimenCode id="DBD29EB7FFAA12502C06F99881051905" box="[1151,1337,1590,1617]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2015.0439</specimenCode>
,
<location id="8EAB6017FFAA12502D3CF99885181921" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502D3CF99885181921" country="Thailand" municipality="Gulf" name="Angsila Fishing Port" pageId="20" pageNumber="378">Angsila Fishing Port</location>
, Gulf of Thailand, eastern
<collectingCountry id="F363765CFFAA12502A32F9F4868D1921" box="[587,689,1627,1653]" name="Thailand" pageId="20" pageNumber="378">Thailand</collectingCountry>
, coll.
<collectorName id="2681531AFFAA12502A8DF9F487691921" box="[756,853,1627,1653]" pageId="20" pageNumber="378">Z. Jaafar</collectorName>
<emphasis id="B900EADEFFAA12502B25F9F487B01921" box="[860,908,1627,1653]" italics="true" pageId="20" pageNumber="378">et al</emphasis>
.,
<date id="FFCA100CFFAA12502BD9F9F480471921" box="[928,1147,1627,1653]" pageId="20" pageNumber="378" value="2003-11-21">
<collectingDate id="EF8EE9E4FFAA12502BD9F9F480471921" box="[928,1147,1627,1653]" pageId="20" pageNumber="378" value="2003-11-21">21 November 2003</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1C3C91FFAA12502CFFF9F4855019CD" pageId="20" pageNumber="378" specimenCount="4" specimenCount-male="4">
<specimenCount id="9D72FD45FFAA12502CFFF9F480E71921" box="[1158,1243,1627,1653]" count="4" pageId="20" pageNumber="378" type="male">4 males</specimenCount>
(20.1 × 20.0 mm 22.9 ×
<quantity id="4C8C9B29FFAA12502885F9D0855819CD" box="[252,356,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" pageId="20" pageNumber="378" unit="mm" value="21.4">21.4 mm</quantity>
)
</materialsCitation>
,
<materialsCitation id="3B1C3C91FFAA1250290EF9D0851019E9" collectingDate="2007-07-27" collectionCode="ZRC" country="Vietnam" location="Cua Lo Beach" pageId="20" pageNumber="378" specimenCode="ZRC 2014.0191" specimenCount="1" specimenCount-female="1" stateProvince="Nghe An" typeStatus="paratype">
<specimenCount id="9D72FD45FFAA1250290EF9D085E419CD" box="[375,472,1663,1689]" count="1" pageId="20" pageNumber="378" type="female">1 female</specimenCount>
(16.4 ×
<quantity id="4C8C9B29FFAA12502A41F9D1869D19CD" box="[568,673,1662,1689]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="20" pageNumber="378" unit="mm" value="15.6">15.6 mm</quantity>
),
<specimenCode id="DBD29EB7FFAA12502ACDF9D0875019CD" box="[692,876,1663,1689]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="378">ZRC 2014.0191</specimenCode>
,
<collectingRegion id="49B0F82EFFAA12502B0EF9D087E219CD" box="[887,990,1663,1689]" country="Vietnam" name="Nghe An" pageId="20" pageNumber="378">Nghe An</collectingRegion>
,
<location id="8EAB6017FFAA12502B93F9D080B119CD" LSID="urn:lsid:plazi:treatment:03DD87DAFFAD125128EEF969807318AF:8EAB6017FFAA12502B93F9D080B119CD" box="[1002,1165,1663,1689]" country="Vietnam" name="Cua Lo Beach" pageId="20" pageNumber="378" stateProvince="Nghe An">Cua Lo Beach</location>
, northern
<collectingCountry id="F363765CFFAA12502C87F9D0815C19CD" box="[1278,1376,1663,1689]" name="Vietnam" pageId="20" pageNumber="378">Vietnam</collectingCountry>
, coll.
<date id="FFCA100CFFAA125028EEF90C851019E9" box="[151,300,1699,1725]" pageId="20" pageNumber="378" value="2007-07-27">
<collectingDate id="EF8EE9E4FFAA125028EEF90C851019E9" box="[151,300,1699,1725]" pageId="20" pageNumber="378" value="2007-07-27">27 July 2007</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAA125128BEF96987A01C2A" lastPageId="21" lastPageNumber="379" pageId="20" pageNumber="378" type="diagnosis">
<paragraph id="8BCB36CCFFAA125128BEF96981201C0E" blockId="20.[151,1437,150,2013]" lastBlockId="21.[151,1437,151,2043]" lastPageId="21" lastPageNumber="379" pageId="20" pageNumber="378">
<emphasis id="B900EADEFFAA125028BEF969850619B4" bold="true" box="[199,314,1734,1760]" pageId="20" pageNumber="378">Diagnosis</emphasis>
. Carapace approximately as broad as long or slightly broader than long, more or less rounded (
<figureCitation id="134F2A49FFAA125028E6F94484CE1851" box="[159,242,1771,1797]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="20" pageNumber="378">
Figs.
<geoCoordinate id="EE40500BFFAA1250289BF94484CE1851" box="[226,242,1771,1797]" degrees="4, 6" direction="east" orientation="longitude" pageId="20" pageNumber="378" precision="5555" value="4.6">4</geoCoordinate>
</figureCitation>
,
<figureCitation id="134F2A49FFAA1250297BF94485791851" box="[258,325,1771,1797]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="20" pageNumber="378">6E, F</figureCitation>
); dorsal surface of carapace covered with closely spaced minute granules or corneous spinules, hepatic, branchial and intestinal regions with slightly larger, more prominent granules. Frontal margin granulated; postfrontal lobe slightly convex with slight concave medially (
<figureCitation id="134F2A49FFAA12502B0AF89C80341819" box="[883,1032,1843,1869]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="20" pageNumber="378">Figs. 11FH</figureCitation>
). Endostomial spine broad, strong, produced, visible from dorsal view (
<figureCitation id="134F2A49FFAA12502A4BF8F886F91825" box="[562,709,1879,1905]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="20" pageNumber="378">Figs. 13HL</figureCitation>
,
<figureCitation id="134F2A49FFAA12502AA8F8F8871C1825" box="[721,800,1879,1905]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="20" pageNumber="378">14E, F</figureCitation>
). Lateral margin of carapace with row of small granules interspersed by larger granules; posterolateral margin gently convex with large granule medially (
<figureCitation id="134F2A49FFAA12502C96F8D4817C18C1" box="[1263,1344,1915,1941]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="20" pageNumber="378">
Figs.
<geoCoordinate id="EE40500BFFAA12502D49F8D4817C18C1" box="[1328,1344,1915,1941]" degrees="4, 6" direction="east" orientation="longitude" pageId="20" pageNumber="378" precision="5555" value="4.6">4</geoCoordinate>
</figureCitation>
,
<figureCitation id="134F2A49FFAA12502D37F8D481B318C1" box="[1358,1423,1915,1941]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="20" pageNumber="378">6E, F</figureCitation>
). Posterior carapace margin gently convex, with row of small granules, slightly larger granule medially (
<figureCitation id="134F2A49FFAA12502D70F830816418ED" box="[1289,1368,1951,1977]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="20" pageNumber="378">Figs. 4</figureCitation>
,
<figureCitation id="134F2A49FFAA12502D1AF83084931889" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="20" pageNumber="378">10F H</figureCitation>
). Third maxilliped exopod 1.51.8 times as long as broad, broader than basal part of endopod, forming petaliform structure with strongly convex outer margin, with submarginal row of low to very low granules along inner margin; endopod with completely fused basis-ischium, distinctly shorter than merus, with shallow submarginal sulcus near inner margin (
<figureCitation id="134F2A49FFAB1251294FFF7185F81FAC" box="[310,452,222,248]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="21" pageNumber="379">Figs. 11FH</figureCitation>
,
<figureCitation id="134F2A49FFAB125129B6FF71861E1FAC" box="[463,546,222,248]" captionStart="FIGURE 12" captionStartId="17.[151,250,1628,1652]" captionTargetBox="[151,1435,181,1605]" captionTargetId="figure-20@17.[151,1435,181,1605]" captionTargetPageId="17" captionText="FIGURE 12. Third maxillipeds. A, Lyphira chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; B, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; C, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore; D, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.97), Malaysia; E, L. linda n. sp., female (17.0 × 16.9 mm) (ZRC 1995.0547), Singapore; F, L. linda n. sp., male (12.1 × 12.4 mm) (ZRC 1995.0547), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681950" httpUri="https://zenodo.org/record/12681950/files/figure.png" pageId="21" pageNumber="379">12DF</figureCitation>
). Chelipeds subequal; merus, carpus and chela surfaces smooth or covered with low, small rounded granules (
<figureCitation id="134F2A49FFAB1251299CFEAD86091E48" box="[485,565,258,284]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="21" pageNumber="379">
Figs.
<geoCoordinate id="EE40500BFFAB12512A5CFEAD86091E48" box="[549,565,258,284]" degrees="4, 6" direction="east" orientation="longitude" pageId="21" pageNumber="379" precision="5555" value="4.6">4</geoCoordinate>
</figureCitation>
,
<figureCitation id="134F2A49FFAB12512A38FEAD86BE1E48" box="[577,642,258,284]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="21" pageNumber="379">6E, F</figureCitation>
); merus surface granulated, large granules on proximal area and along margins, minutes granules distally; carpus smooth, unarmed; chela stout, palm surface smooth, upper and lower margins with row of granules; fingers longer than length of palm, surface with longitudinal sulcus; pollex bent, cutting edge with broad, large subproximal tooth lined with denticles, remaining of edge with small denticles; dactylus 1.6 times as long as palm, with row of small granules on upper margin, cutting edge with subdistal large tooth lined with denticles, remaining of edge with small denticles (
<figureCitation id="134F2A49FFAB12512BD5FE1A80031E84" box="[940,1087,437,464]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="21" pageNumber="379">Fig. 15G, H</figureCitation>
). P2P5 slender, short; merus distinctly longer than carpus and propodus; dactylus longer than propodus, lanceolate, terminating in corneous tips; fourth leg shortest, with merus 4 times as long as broad, margins lined with small granules, more prominent on ventral margin (
<figureCitation id="134F2A49FFAB12512917FD8E85D61D6F" box="[366,490,545,571]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. AG, IK, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="21" pageNumber="379">Fig. 16FI</figureCitation>
). Thoracic sternites transversely narrow, surface granulated, larger granules laterally (
<figureCitation id="134F2A49FFAB125128E6FDEA85201D0B" box="[159,284,581,607]" captionStart="FIGURE 8" captionStartId="13.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1714]" captionTargetId="figure-20@13.[151,1435,181,1714]" captionTargetPageId="13" captionText="FIGURE 8. Male thoracic sternum and pleon. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="21" pageNumber="379">Fig. 8C, D</figureCitation>
). Male sternopleonal cavity deep, reaching to mid distance between fused thoracic sternites 13. Male pleon narrow, slender, long, edge granulated; somites 26 fused, shallow suture just visible between somites 5 and 6, surface smooth; somite 6 longitudinally subrectangular, surface with one low granule medially, low, short sulcus subproximally, lateral margins straight; telson triangular, longer than wide (
<figureCitation id="134F2A49FFAB12512C4DFD1E80861D9F" box="[1076,1210,689,715]" captionStart="FIGURE 8" captionStartId="13.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1714]" captionTargetId="figure-20@13.[151,1435,181,1714]" captionTargetPageId="13" captionText="FIGURE 8. Male thoracic sternum and pleon. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="21" pageNumber="379">Figs. 8C, D</figureCitation>
,
<figureCitation id="134F2A49FFAB12512CBCFD1E80C51D9F" box="[1221,1273,689,715]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. AE, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; FK, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; LP, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; QV, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: AC, FH, L, QS = 1.0 mm; D, E, IK, MP, TV = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="21" pageNumber="379">18G</figureCitation>
). G1 elongate, slender, same size from proximal to distal area; distal process, stout, bilobed, with bent, rounded tip (
<figureCitation id="134F2A49FFAB12512D7EFD7B81A41DBB" box="[1287,1432,724,751]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. AE, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; FK, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; LP, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; QV, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: AC, FH, L, QS = 1.0 mm; D, E, IK, MP, TV = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="21" pageNumber="379">Figs. 18HV</figureCitation>
,
<figureCitation id="134F2A49FFAB125128EEFD5784CC1C46" box="[151,240,760,786]" captionStart="FIGURE 20" captionStartId="29.[151,250,1639,1663]" captionTargetBox="[252,1327,191,1613]" captionTargetId="figure-20@29.[239,1349,181,1615]" captionTargetPageId="29" captionText="FIGURE 20. G1. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; C, L. ngankee n. sp., paratype male (19.2 × 19.9 mm) (ZRC 2022.0628), Hong Kong; D, L. ngankee n. sp., paratype male (15.5 × 15.1 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (16.7 × 17.0 mm) (ZRC 2015.0439); Thailand; G, L. linda n. sp., male (18.4 × 19.0 mm) (ZRC 1985.1632), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. linda n. sp., male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; M, L. linda n. sp., male (22.9 × 21.4 mm) (ZRC 2014.0191), Vietnam; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia; O, L. bellagrana n. sp., male (13.2 × 12.6 mm) (ZRC 2022.0745), Papua, Indonesia; P, L. bellagrana n. sp., male (10.6 × 9.9 mm) (ZRC 2022.0750), Papua, Indonesia. A, C, EH, KP, right G1; B, D, I, J, left G1 (laterally inverted)." figureDoi="http://doi.org/10.5281/zenodo.12681975" httpUri="https://zenodo.org/record/12681975/files/figure.png" pageId="21" pageNumber="379">20EM</figureCitation>
). Female sternopleonal cavity reaches distal part of sternite 1; female pleon broad, convex, smooth, somites 26 fused, suture between somites 2 and 3; telson shorter than its basal width (
<figureCitation id="134F2A49FFAB12512C69FCB380B91C62" box="[1040,1157,796,822]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="21" pageNumber="379">Fig. 9E, F</figureCitation>
); vulvae spaced apart on sternite 6, each opening weakly crescent-shaped, relatively long, without opercular cover (
<figureCitation id="134F2A49FFAB12512CF5FCEF81331C0E" box="[1164,1295,832,858]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="21" pageNumber="379">Fig. 21E, F</figureCitation>
).
</paragraph>
<paragraph id="8BCB36CCFFAB125128BEFCCB87A01C2A" blockId="21.[151,1437,151,2043]" box="[199,924,868,895]" pageId="21" pageNumber="379">
<emphasis id="B900EADEFFAB125128BEFCCB85271C2A" bold="true" box="[199,283,868,894]" pageId="21" pageNumber="379">Colour</emphasis>
. Brownish grey to dirty white, merus of cheliped yellow.
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAB125128BEFC2786F71B65" pageId="21" pageNumber="379" type="etymology">
<paragraph id="8BCB36CCFFAB125128BEFC2786F71B65" blockId="21.[151,1437,151,2043]" pageId="21" pageNumber="379">
<emphasis id="B900EADEFFAB125128BEFC2785741CF6" bold="true" box="[199,328,904,930]" pageId="21" pageNumber="379">Etymology</emphasis>
. The species is named after the late Linda Goh. She was an Assistant Director at the National Parks Board and was overseeing the Comprehensive Marine Biodiversity Survey for the government. She was instrumental in making sure the project progressed smoothly and ensured its success. Beyond this, she was a good friend and a lovely person to work with. Her sudden passing in 2021 was a tragedy for both the agency and conservation in general. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAB125128BEFB94877818E6" pageId="21" pageNumber="379" type="discussion">
<paragraph id="8BCB36CCFFAB125128BEFB9480BE1973" blockId="21.[151,1437,151,2043]" pageId="21" pageNumber="379">
<emphasis id="B900EADEFFAB125128BEFB9485081B01" bold="true" box="[199,308,1083,1109]" pageId="21" pageNumber="379">Remarks</emphasis>
. The variation in the smoothness of the carapace and pereopods of
<taxonomicName id="4C744D4FFFAB12512C2DFB93808C1B01" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[1108,1200,1083,1109]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
<emphasis id="B900EADEFFAB12512C2DFB93808C1B01" box="[1108,1200,1083,1109]" italics="true" pageId="21" pageNumber="379">L. linda</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB12512CC3FB9480C11B01" bold="true" box="[1210,1277,1083,1109]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB12512CC3FB9480C11B01" box="[1210,1277,1083,1109]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is surprising. While, most specimens have smooth and unarmed carapaces, chelipeds and legs, with the granules present rounded and smooth (e.g.,
<figureCitation id="134F2A49FFAB1251291BFB2C85E11BC9" box="[354,477,1155,1181]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="21" pageNumber="379">Fig. 4AC</figureCitation>
); the same surfaces of several specimens are covered with short corneous spinules to various degrees, even on the thoracic sternum and pleon (
<figureCitation id="134F2A49FFAB12512B63FB0887A51B95" box="[794,921,1191,1217]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="21" pageNumber="379">Figs. 4DF</figureCitation>
,
<figureCitation id="134F2A49FFAB12512BDDFB0887FE1B95" box="[932,962,1191,1217]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="21" pageNumber="379">6F</figureCitation>
,
<figureCitation id="134F2A49FFAB12512BB4FB08802F1B95" box="[973,1043,1191,1217]" captionStart="FIGURE 8" captionStartId="13.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1714]" captionTargetId="figure-20@13.[151,1435,181,1714]" captionTargetPageId="13" captionText="FIGURE 8. Male thoracic sternum and pleon. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="21" pageNumber="379">8C, D</figureCitation>
). These spiny specimens, however, agree with the smooth ones in all other diagnostic characters, including the structures of their endostomial spines, third maxillipeds and G1s. In addition, there are many specimens with intermediate characters, e.g., only some parts of the carapace are spinous, only some legs or some articles of the legs are spiny, only one cheliped is spinous with the other smooth etc. (e.g., see
<figureCitation id="134F2A49FFAB12512A56FA9986951A04" box="[559,681,1334,1360]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="21" pageNumber="379">Fig. 4D, E</figureCitation>
). In some specimens, there are only a few scattered spinules on the carapace or pereopods, with remaining of the surfaces smooth. These variations are independent of sex. All juvenile and subadult specimens are of the smooth
<typeStatus id="54CF886EFFAB12512AF3FAD086811ACD" box="[650,701,1407,1433]" pageId="21" pageNumber="379">type</typeStatus>
, suggesting spination is associated with adulthood. These adult specimens with varying degrees of spination can be collected together from the same location. For example, in lot ZRC 2003.0107 from
<collectingRegion id="49B0F82EFFAB125129F7FA6985E01AB4" box="[398,476,1478,1504]" country="Thailand" name="Pattani" pageId="21" pageNumber="379">Pattani</collectingRegion>
in the Gulf of
<collectingCountry id="F363765CFFAB12512A05FA6986DE1AB4" box="[636,738,1478,1504]" name="Thailand" pageId="21" pageNumber="379">Thailand</collectingCountry>
,
<specimenCount id="9D72FD45FFAB12512A94FA6987B81AB4" box="[749,900,1478,1504]" count="32" pageId="21" pageNumber="379" type="generic">32 specimens</specimenCount>
were of the smooth
<typeStatus id="54CF886EFFAB12512C1CFA6980A41AB4" box="[1125,1176,1478,1504]" pageId="21" pageNumber="379">type</typeStatus>
, with 22 having at least some some spinules on some part of the body. This substantial variation is not known for any other known leucosiid species, with the possible exception of
<taxonomicName id="4C744D4FFFAB12512A34F9A186DA1973" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[589,742,1549,1575]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
<emphasis id="B900EADEFFAB12512A34F9A186DA1973" box="[589,742,1549,1575]" italics="true" pageId="21" pageNumber="379">L. bellagrana</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB12512A95F9A18717197C" bold="true" box="[748,811,1550,1576]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB12512A95F9A18717197C" box="[748,811,1550,1576]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(see remarks for that species).
</paragraph>
<paragraph id="8BCB36CCFFAB125128BEF99E877818E6" blockId="21.[151,1437,151,2043]" pageId="21" pageNumber="379">
Spination aside,
<taxonomicName id="4C744D4FFFAB125129F4F99D85D7191F" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[397,491,1585,1611]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
<emphasis id="B900EADEFFAB125129F4F99D85D7191F" box="[397,491,1585,1611]" italics="true" pageId="21" pageNumber="379">L. linda</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB12512981F99E8601191F" bold="true" box="[504,573,1585,1611]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB12512981F99E8601191F" box="[504,573,1585,1611]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C744D4FFFAB12512AF9F99D873C191F" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[640,768,1585,1611]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAB12512AF9F99D873C191F" box="[640,768,1585,1611]" italics="true" pageId="21" pageNumber="379">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB12512B77F99E876F191F" bold="true" box="[782,851,1585,1611]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB12512B77F99E876F191F" box="[782,851,1585,1611]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are morphologically close; possessing prominent endostomial spines, a wide third maxilliped exopod and a slightly convex posterior carapace margin. The carapace of
<taxonomicName id="4C744D4FFFAB125128CEF9D5852C19C7" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[183,272,1657,1683]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
<emphasis id="B900EADEFFAB125128CEF9D5852C19C7" box="[183,272,1657,1683]" italics="true" pageId="21" pageNumber="379">L. linda</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB1251296EF9D6856B19C7" bold="true" box="[279,343,1657,1683]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB1251296EF9D6856B19C7" box="[279,343,1657,1683]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, however, is as long as broad (
<figureCitation id="134F2A49FFAB12512AC3F9D6873619C7" box="[698,778,1657,1683]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="21" pageNumber="379">
Figs.
<geoCoordinate id="EE40500BFFAB12512A83F9D6873619C7" box="[762,778,1657,1683]" degrees="4, 6" direction="east" orientation="longitude" pageId="21" pageNumber="379" precision="5555" value="4.6">4</geoCoordinate>
</figureCitation>
,
<figureCitation id="134F2A49FFAB12512B61F9D6876B19C7" box="[792,855,1657,1683]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="21" pageNumber="379">6E, F</figureCitation>
) (broader than long in
<taxonomicName id="4C744D4FFFAB12512C26F9D580E719C7" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[1119,1243,1657,1683]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAB12512C26F9D580E719C7" box="[1119,1243,1657,1683]" italics="true" pageId="21" pageNumber="379">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB12512C9AF9D6811F19C7" bold="true" box="[1251,1315,1657,1683]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB12512C9AF9D6811F19C7" box="[1251,1315,1657,1683]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="134F2A49FFAB12512D4AF9D6849019E3" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="21" pageNumber="379">Figs. 5A, B</figureCitation>
,
<figureCitation id="134F2A49FFAB125128C1F93284E119E3" box="[184,221,1693,1719]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="21" pageNumber="379">6D</figureCitation>
), the gastrocardiac grooves very shallow and barely visible (Fig. E, F) (grooves clearly visible and relatively deeper in
<taxonomicName id="4C744D4FFFAB12512970F96D85BA198F" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[265,390,1729,1755]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAB12512970F96D85BA198F" box="[265,390,1729,1755]" italics="true" pageId="21" pageNumber="379">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB125129F6F96E85F3198F" bold="true" box="[399,463,1729,1755]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB125129F6F96E85F3198F" box="[399,463,1729,1755]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="134F2A49FFAB12512999F96E8660198F" box="[480,604,1729,1756]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="21" pageNumber="379">Fig. 5A, B</figureCitation>
); the endostomial spines are slightly weaker and positioned further apart (
<figureCitation id="134F2A49FFAB125128E6F94A851819AB" box="[159,292,1765,1791]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="21" pageNumber="379">Fig. 13HL</figureCitation>
) (spines stronger and positioned closer together in
<taxonomicName id="4C744D4FFFAB12512B24F94A87EB19AB" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[861,983,1765,1791]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAB12512B24F94A87EB19AB" box="[861,983,1765,1791]" italics="true" pageId="21" pageNumber="379">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB12512BA7F94A802019AB" bold="true" box="[990,1052,1765,1791]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB12512BA7F94A802019AB" box="[990,1052,1765,1791]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="134F2A49FFAB12512C52F94A809019AB" box="[1067,1196,1765,1791]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="21" pageNumber="379">Fig. 13F, G</figureCitation>
); the third maxilliped exopod is somewhat more rounded, being 1.5 to 1.8 as long as broad (
<figureCitation id="134F2A49FFAB12512BCCF8A780041877" box="[949,1080,1800,1827]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. AE, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; FK, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; LP, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; QV, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: AC, FH, L, QS = 1.0 mm; D, E, IK, MP, TV = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="21" pageNumber="379">Fig. 18F, Q</figureCitation>
) (slightly longer, being 2 times as long as broad in
<taxonomicName id="4C744D4FFFAB1251290EF88285CF1812" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[375,499,1836,1862]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAB1251290EF88285CF1812" box="[375,499,1836,1862]" italics="true" pageId="21" pageNumber="379">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB12512982F88386001812" bold="true" box="[507,572,1836,1862]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB12512982F88386001812" box="[507,572,1836,1862]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="134F2A49FFAB12512A35F88386881812" box="[588,692,1836,1862]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. AE, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; FK, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; LP, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; QV, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: AC, FH, L, QS = 1.0 mm; D, E, IK, MP, TV = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="21" pageNumber="379">Fig. 18A</figureCitation>
); the palm of the male cheliped is distinctly stouter with a longer dactylus, 1.6 times as long as the palm (
<figureCitation id="134F2A49FFAB12512A37F8FF86E8183E" box="[590,724,1872,1899]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="21" pageNumber="379">Fig. 15G, H</figureCitation>
) (palm distinctly more slender with a shorter dactylus, 1.4 times as long as palm in
<taxonomicName id="4C744D4FFFAB12512913F8DA85D918DA" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[362,485,1908,1934]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAB12512913F8DA85D918DA" box="[362,485,1908,1934]" italics="true" pageId="21" pageNumber="379">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB12512994F8DB861018DA" bold="true" box="[493,556,1908,1934]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB12512994F8DB861018DA" box="[493,556,1908,1934]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="134F2A49FFAB12512A42F8DB86A218DA" box="[571,670,1908,1934]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="21" pageNumber="379">Fig. 14F</figureCitation>
); and the G1 apical process is longer and bilobed (
<figureCitation id="134F2A49FFAB12512CAEF8DB84F118E6" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. AE, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; FK, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; LP, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; QV, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: AC, FH, L, QS = 1.0 mm; D, E, IK, MP, TV = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="21" pageNumber="379">Fig. 18IK, MP, TV</figureCitation>
) (shorter and broader in
<taxonomicName id="4C744D4FFFAB1251299BF836866018E6" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[482,604,1944,1970]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAB1251299BF836866018E6" box="[482,604,1944,1970]" italics="true" pageId="21" pageNumber="379">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAB12512A1AF837869E18E6" bold="true" box="[611,674,1944,1970]" pageId="21" pageNumber="379">
<taxonomicNameLabel id="A23357A5FFAB12512A1AF837869E18E6" box="[611,674,1944,1970]" pageId="21" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="134F2A49FFAB12512AC8F837870418E6" box="[689,824,1944,1970]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. AE, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; FK, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; LP, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; QV, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: AC, FH, L, QS = 1.0 mm; D, E, IK, MP, TV = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="21" pageNumber="379">Fig. 18D, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAB125128BEF81385DB1882" box="[199,487,1980,2007]" pageId="21" pageNumber="379" type="materials_examined">
<paragraph id="8BCB36CCFFAB125128BEF81385DB1882" blockId="21.[151,1437,151,2043]" box="[199,487,1980,2007]" pageId="21" pageNumber="379">
<materialsCitation id="3B1C3C91FFAB125128BEF81385DF1882" box="[199,483,1980,2007]" country="Singapore" location="Singapore" pageId="21" pageNumber="379" specimenCount="1" typeStatus="holotype">
<emphasis id="B900EADEFFAB125128BEF813855C1882" bold="true" box="[199,352,1980,2006]" pageId="21" pageNumber="379">
<typeStatus id="54CF886EFFAB125128BEF813853E1882" box="[199,258,1980,2006]" pageId="21" pageNumber="379">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F363765CFFAB12512917F81385DF1882" box="[366,483,1980,2006]" name="Singapore" pageId="21" pageNumber="379">Singapore</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAB125128BEF84F807318AF" box="[199,1103,2016,2043]" pageId="21" pageNumber="379" type="distribution">
<paragraph id="8BCB36CCFFAB125128BEF84F807318AF" blockId="21.[151,1437,151,2043]" box="[199,1103,2016,2043]" pageId="21" pageNumber="379">
<emphasis id="B900EADEFFAB125128BEF84F856518AE" bold="true" box="[199,345,2016,2042]" pageId="21" pageNumber="379">Distribution</emphasis>
.
<collectingCountry id="F363765CFFAB1251291EF84F85E718AE" box="[359,475,2016,2042]" name="Singapore" pageId="21" pageNumber="379">Singapore</collectingCountry>
,
<collectingCountry id="F363765CFFAB1251299EF84F866C18AE" box="[487,592,2016,2042]" name="Malaysia" pageId="21" pageNumber="379">Malaysia</collectingCountry>
,
<collectingCountry id="F363765CFFAB12512A22F84F868318AE" box="[603,703,2016,2042]" name="Thailand" pageId="21" pageNumber="379">Thailand</collectingCountry>
and southern
<collectingCountry id="F363765CFFAB12512B25F84F878218AE" box="[860,958,2016,2042]" name="Vietnam" pageId="21" pageNumber="379">Vietnam</collectingCountry>
,
<quantity id="4C8C9B29FFAB12512BB0F84F807718AF" box="[969,1099,2016,2043]" metricMagnitude="1" metricUnit="m" metricValue="1.535" metricValueMax="2.25" metricValueMin="0.82" pageId="21" pageNumber="379" unit="m" value="15.35" valueMax="22.5" valueMin="8.2">8.222.5 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>