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<mods:title id="9A857B855123F0140E9C32411ECAEB2E">On the genus Lyphira Galil, 2009 (Crustacea: Brachyura: Leucosiidae), with descriptions of four new species from the West Pacific</mods:title>
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<mods:namePart id="6AA6D05117620072E5873E8DD407122A">Rahayu, Dwi Listyo</mods:namePart>
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<mods:affiliation id="65A0800AF24BA11A491F484853FE944C">Research Center for Marine and Land Bioindustry, National Research and Innovation Agency, Dusun Teluk Kodek, Pemenang, Lombok Utara 83756, NTB, Indonesia.</mods:affiliation>
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<mods:namePart id="6AFB2E2C07AB70E6E1D0F1D152EE823A">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="EA388102494499427BA02C9FBE0874DF">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore.</mods:affiliation>
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<mods:date id="A8E976A27A1C3E1E91229A87E9F44552">2024</mods:date>
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<mods:number id="5B6795545278749E689189E9A2B10325">2024-07-04</mods:number>
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<taxonomicName id="4C744D4FFFAC125628EEFCB185591C6D" authorityName="Rahayu & Ng" authorityYear="2024" box="[151,357,798,825]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="ngankee" status="sp. nov.">
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<emphasis id="B900EADEFFAC125628EEFCB185591C6D" bold="true" box="[151,357,798,825]" italics="true" pageId="18" pageNumber="376">Lyphira ngankee</emphasis>
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<taxonomicNameLabel id="A23357A5FFAC12562915FCB085921C6D" box="[364,430,799,825]" pageId="18" pageNumber="376" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="134F2A49FFAC125628E6FCEC85251C09" box="[159,281,835,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="18" pageNumber="376">Figs. 1E–J</figureCitation>
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<figureCitation id="134F2A49FFAC1256295CFCED85561C09" box="[293,362,834,861]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="18" pageNumber="376">5A, B</figureCitation>
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,
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<figureCitation id="134F2A49FFAC1256290FFCEC85A61C09" box="[374,410,835,861]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="18" pageNumber="376">6D</figureCitation>
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,
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<figureCitation id="134F2A49FFAC125629DFFCEC85F61C09" box="[422,458,835,861]" captionStart="FIGURE 8" captionStartId="13.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1714]" captionTargetId="figure-20@13.[151,1435,181,1714]" captionTargetPageId="13" captionText="FIGURE 8. Male thoracic sternum and pleon. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="18" pageNumber="376">8A</figureCitation>
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,
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<figureCitation id="134F2A49FFAC125629AFFCEC85C61C09" box="[470,506,835,861]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="18" pageNumber="376">9D</figureCitation>
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<figureCitation id="134F2A49FFAC12562A7FFCEC860A1C09" box="[518,566,835,861]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="18" pageNumber="376">10E</figureCitation>
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,
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<figureCitation id="134F2A49FFAC12562A38FCEC864C1C09" box="[577,624,835,861]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="18" pageNumber="376">11E</figureCitation>
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<figureCitation id="134F2A49FFAC12562A02FCEC86F61C09" box="[635,714,835,861]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="18" pageNumber="376">13F, G</figureCitation>
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<figureCitation id="134F2A49FFAC12562AACFCEC873B1C09" box="[725,775,835,861]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="18" pageNumber="376">14D</figureCitation>
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|
||||
<figureCitation id="134F2A49FFAC12562B6AFCED87031C09" box="[787,831,834,861]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="18" pageNumber="376">15F</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562B32FCEC87461C09" box="[843,890,835,861]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="18" pageNumber="376">16E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562BFFFCEC87EB1C09" box="[902,983,835,861]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="18" pageNumber="376">18A‒E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562B9AFCEC80041C09" box="[995,1080,835,861]" captionStart="FIGURE 20" captionStartId="29.[151,250,1639,1663]" captionTargetBox="[252,1327,191,1613]" captionTargetId="figure-20@29.[239,1349,181,1615]" captionTargetPageId="29" captionText="FIGURE 20. G1. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; C, L. ngankee n. sp., paratype male (19.2 × 19.9 mm) (ZRC 2022.0628), Hong Kong; D, L. ngankee n. sp., paratype male (15.5 × 15.1 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (16.7 × 17.0 mm) (ZRC 2015.0439); Thailand; G, L. linda n. sp., male (18.4 × 19.0 mm) (ZRC 1985.1632), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. linda n. sp., male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; M, L. linda n. sp., male (22.9 × 21.4 mm) (ZRC 2014.0191), Vietnam; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia; O, L. bellagrana n. sp., male (13.2 × 12.6 mm) (ZRC 2022.0745), Papua, Indonesia; P, L. bellagrana n. sp., male (10.6 × 9.9 mm) (ZRC 2022.0750), Papua, Indonesia. A, C, E–H, K–P, right G1; B, D, I, J, left G1 (laterally inverted)." figureDoi="http://doi.org/10.5281/zenodo.12681975" httpUri="https://zenodo.org/record/12681975/files/figure.png" pageId="18" pageNumber="376">20A‒D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562C3AFCEC80491C09" box="[1091,1141,835,861]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="18" pageNumber="376">21D</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFAC125628EEFC2880ED1B6D" pageId="18" pageNumber="376" type="reference_group">
|
||||
<paragraph id="8BCB36CCFFAC125628EEFC2885D31CEA" blockId="18.[151,1436,903,1081]" pageId="18" pageNumber="376">
|
||||
<treatmentCitationGroup id="AB6411E2FFAC125628EEFC2885D31CEA" pageId="18" pageNumber="376">
|
||||
<emphasis id="B900EADEFFAC125628EEFC2885411CCB" box="[151,381,903,927]" italics="true" pageId="18" pageNumber="376">
|
||||
<taxonomicName id="4C744D4FFFAC125628EEFC2885631CCB" ID-CoL="4GDVK" authorityName="Stimpson" authorityYear="1858" box="[151,351,903,927]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="tuberculosa">Philyra tuberculosa</taxonomicName>
|
||||
.—
|
||||
</emphasis>
|
||||
<treatmentCitation id="0AD510DDFFAC12562904FC2886221CCB" author="Shen, C. - J." box="[381,542,903,927]" page="185" pageId="18" pageNumber="376" year="1931">
|
||||
<bibRefCitation id="EFE54B3DFFAC12562904FC2886221CCB" author="Shen, C. - J." box="[381,542,903,927]" pageId="18" pageNumber="376" pagination="185 - 197" refId="ref21785" refString="Shen, C. - J. (1931) The crabs of Hong Kong. Part 2. Hong Kong Naturalist, 2 (3), 185 - 197." type="journal article" year="1931">Shen 1931: 185</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
, text figs. 1, 2, 4;
|
||||
<bibRefCitation id="EFE54B3DFFAC12562AACFC2887781CCB" author="Shen, C. - J." box="[725,836,903,927]" pageId="18" pageNumber="376" pagination="1 - 300" refId="ref21817" refString="Shen, C. - J. (1932) The brachyuran Crustacea of North China. Zoologia Sinica, 9 (1), 1 - 300, pls. 1 - 10." type="journal article" year="1932">Shen 1932</bibRefCitation>
|
||||
: text fig. 11e, f. (not
|
||||
<taxonomicName id="4C744D4FFFAC12562C60FC2881B41CCB" ID-CoL="4GDVK" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[1049,1416,903,927]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="tuberculosa">
|
||||
<emphasis id="B900EADEFFAC12562C60FC2880DE1CCB" box="[1049,1250,903,927]" italics="true" pageId="18" pageNumber="376">Philyra tuberculosa</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFAC12562C91FC2881B41CCB" author="Stimpson, W." box="[1256,1416,903,927]" pageId="18" pageNumber="376" pagination="159 - 163" refId="ref21886" refString="Stimpson, W. (1858) Crustacea Oxystomata. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VI. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 159 - 163. https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||||
</taxonomicName>
|
||||
=
|
||||
<taxonomicName id="4C744D4FFFAC125628BEFC0985DF1CEA" ID-CoL="76YLX" authority="Bell, 1855" authorityName="Bell" authorityYear="1855" box="[199,483,934,958]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="carinata">
|
||||
<emphasis id="B900EADEFFAC125628BEFC09854C1CE9" box="[199,368,934,958]" italics="true" pageId="18" pageNumber="376">Philyra carinata</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFAC1256290FFC0985DF1CEA" author="Bell, T." box="[374,483,934,958]" pageId="18" pageNumber="376" pagination="361 - 367" refId="ref19289" refString="Bell, T. (1855) Horae carcinologicae, or Notices of Crustacea. I. A Monograph of the Leucosiidae, with observations on the relations, structure, habits and distribution of the family; a revision of the generic characters; and descriptions of new genera and species. Annals and Magazine of Natural History, Series 2, 16 (45), 361 - 367. https: // doi. org / 10.5962 / bhl. title. 9957" type="journal article" year="1855">Bell, 1855</bibRefCitation>
|
||||
</taxonomicName>
|
||||
).
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFAC125628EEFC6B80E11C88" blockId="18.[151,1436,903,1081]" box="[151,1245,964,989]" pageId="18" pageNumber="376">
|
||||
<treatmentCitationGroup id="AB6411E2FFAC125628EEFC6B80E11C88" box="[151,1245,964,989]" pageId="18" pageNumber="376">
|
||||
<emphasis id="B900EADEFFAC125628EEFC6B856B1C88" box="[151,343,964,988]" italics="true" pageId="18" pageNumber="376">
|
||||
<taxonomicName id="4C744D4FFFAC125628EEFC6B85051C88" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[151,313,964,988]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="globosa">Philyra globosa</taxonomicName>
|
||||
.—
|
||||
</emphasis>
|
||||
<treatmentCitation id="0AD510DDFFAC1256292EFC6B85CA1C88" author="Shen, C. - J." box="[343,502,964,989]" page="216" pageId="18" pageNumber="376" year="1940">
|
||||
<bibRefCitation id="EFE54B3DFFAC1256292EFC6B85CA1C88" author="Shen, C. - J." box="[343,502,964,989]" pageId="18" pageNumber="376" pagination="211 - 242" refId="ref21852" refString="Shen, C. - J. (1940) The brachyuran fauna of Hongkong. Journal of the Hong Kong Fisheries Research Station, 1 (2), 211 - 242." type="journal article" year="1940">Shen 1940: 216</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
(not
|
||||
<taxonomicName id="4C744D4FFFAC12562A52FC6A87461C89" authority="Fabricius, 1793" authorityName="Fabricius" authorityYear="1793" box="[555,890,964,989]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="globosus">
|
||||
<emphasis id="B900EADEFFAC12562A52FC6A86EA1C88" box="[555,726,964,988]" italics="true" pageId="18" pageNumber="376">Cancer globosus</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFAC12562AA5FC6B87461C89" author="Fabricius, J. C." box="[732,890,964,989]" pageId="18" pageNumber="376" refId="ref19754" refString="Fabricius, J. C. (1793) Entomologia systematica emendata et aucta secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Vol. 2. impensis Christ. Gottl. Proft, Hafniae, viii + 519 pp. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1793">Fabricius, 1793</bibRefCitation>
|
||||
</taxonomicName>
|
||||
=
|
||||
<taxonomicName id="4C744D4FFFAC12562BECFC6A80ED1C88" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[917,1233,964,988]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="globus">
|
||||
<emphasis id="B900EADEFFAC12562BECFC6A80161C88" box="[917,1066,964,988]" italics="true" pageId="18" pageNumber="376">Cancer globus</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFAC12562C49FC6B80ED1C88" author="Fabricius, J. C." box="[1072,1233,964,988]" pageId="18" pageNumber="376" refId="ref19698" refString="Fabricius, J. C. (1775) Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, adiectis Synonymis, Locis, Descriptionibus, Observationibus. Flensburgi & Lipsiae, Korte, xxvii + 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510" type="book" year="1775">Fabricius, 1775</bibRefCitation>
|
||||
</taxonomicName>
|
||||
).
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFAC125628EEFC4C81B61CAF" blockId="18.[151,1436,903,1081]" box="[151,1418,995,1019]" pageId="18" pageNumber="376">
|
||||
<treatmentCitationGroup id="AB6411E2FFAC125628EEFC4C81B61CAF" box="[151,1418,995,1019]" pageId="18" pageNumber="376">
|
||||
<taxonomicName id="4C744D4FFFAC125628EEFC4C855B1CAF" ID-CoL="76YMQ" authorityName="Doflein" authorityYear="1904" box="[151,359,995,1019]" class="Malacostraca" family="Leucosiidae" genus="Philyra" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="laminata">
|
||||
<emphasis id="B900EADEFFAC125628EEFC4C84DF1CAF" box="[151,227,995,1019]" italics="true" pageId="18" pageNumber="376">Philyra</emphasis>
|
||||
cf.
|
||||
<emphasis id="B900EADEFFAC12562973FC4C855B1CAF" box="[266,359,995,1019]" italics="true" pageId="18" pageNumber="376">laminata</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAC1256291EFC4C85571CAF" box="[359,363,995,1019]" italics="true" pageId="18" pageNumber="376">.</emphasis>
|
||||
—
|
||||
<treatmentCitation id="0AD510DDFFAC125629FDFC4C86251CAF" author="Hill, D. S." box="[388,537,995,1019]" page="199" pageId="18" pageNumber="376" year="1982">
|
||||
<bibRefCitation id="EFE54B3DFFAC125629FDFC4C86251CAF" author="Hill, D. S." box="[388,537,995,1019]" pageId="18" pageNumber="376" pagination="195 - 205" refId="ref20600" refString="Hill, D. S. (1982) The Leucosiidae (Crustacea: Decapoda) of Hong Kong. In: Morton, B. S. & Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong University Press, Hong Kong, pp. 195 - 205." type="book chapter" year="1982">Hill 1982: 199</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
, pl. 3C. (not
|
||||
<taxonomicName id="4C744D4FFFAC12562AE7FC4C87E21CAF" ID-CoL="76YMQ" authority="Doflein, 1904" authorityName="Doflein" authorityYear="1904" box="[670,990,995,1019]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="laminata">
|
||||
<emphasis id="B900EADEFFAC12562AE7FC4C87761CAF" box="[670,842,995,1019]" italics="true" pageId="18" pageNumber="376">Philyra laminata</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFAC12562B28FC4C87E21CAF" author="Doflein, F." box="[849,990,995,1019]" pageId="18" pageNumber="376" pagination="1 - 58" refId="ref19632" refString="Doflein, F. (1904) Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer " Valdivia " 1898 - 1899. Vol. 6. Verlag von Gustav Fischer, Jena, pp. 1 - 6 + i - xiv + 1 - 314 pp., figs. I - LVIII, atlas pls. 1 - 58." type="book chapter" year="1904">Doflein, 1904</bibRefCitation>
|
||||
</taxonomicName>
|
||||
=
|
||||
<taxonomicName id="4C744D4FFFAC12562B81FC4B81BA1CAF" ID-CoL="6LSP4" authority="(Doflein, 1904))" authorityName="" baseAuthorityName="Doflein" baseAuthorityYear="1904" box="[1016,1414,995,1019]" class="Malacostraca" family="Leucosiidae" genus="Heteronucia" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="laminata">
|
||||
<emphasis id="B900EADEFFAC12562B81FC4B80EA1CAF" box="[1016,1238,995,1019]" italics="true" pageId="18" pageNumber="376">Heteronucia laminata</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFAC12562C9DFC4C81491CAF" author="Doflein, F." box="[1252,1397,995,1019]" pageId="18" pageNumber="376" pagination="1 - 58" refId="ref19632" refString="Doflein, F. (1904) Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer " Valdivia " 1898 - 1899. Vol. 6. Verlag von Gustav Fischer, Jena, pp. 1 - 6 + i - xiv + 1 - 314 pp., figs. I - LVIII, atlas pls. 1 - 58." type="book chapter" year="1904">Doflein, 1904</bibRefCitation>
|
||||
))
|
||||
</taxonomicName>
|
||||
.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFAC125628EEFBAD80E21B4E" blockId="18.[151,1436,903,1081]" box="[151,1246,1026,1050]" pageId="18" pageNumber="376">
|
||||
<emphasis id="B900EADEFFAC125628EEFBAD85BF1B4E" box="[151,387,1026,1050]" italics="true" pageId="18" pageNumber="376">
|
||||
<taxonomicName id="4C744D4FFFAC125628EEFBAD85591B4E" ID-CoL="76YMK" authorityName="Ortmann" authorityYear="1892" box="[151,357,1026,1050]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="heterograna">Philyra heterograna</taxonomicName>
|
||||
.—
|
||||
</emphasis>
|
||||
<treatmentCitation id="0AD510DDFFAC125629FAFBAD86971B4E" author="Blackmore & Rainbow" box="[387,683,1026,1050]" pageId="18" pageNumber="376" year="2000">
|
||||
<bibRefCitation id="EFE54B3DFFAC125629FAFBAD86971B4E" author="Blackmore, G. & Rainbow, P. S." box="[387,683,1026,1050]" pageId="18" pageNumber="376" pagination="517 - 529" refId="ref19375" refString="Blackmore, G. & Rainbow, P. S. (2000) Epibenthic crab (Crustacea: Brachyura) assemblages of the southeastern waters of Hong Kong: the 1998 Trawl Programme. In: Morton, B. (Ed.), Proceedings of the Tenth International marine Biological Workshop: The marine Flora and Fauna of Hong Kong and southern China. Hong Kong University Press, Hong Kong, pp. 517 - 529." type="book chapter" year="2000">Blackmore & Rainbow 2000</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
: appendix 1. (not
|
||||
<taxonomicName id="4C744D4FFFAC12562B19FBAD80EE1B4E" ID-CoL="76YMK" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[864,1234,1026,1050]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFAC12562B19FBAD80121B4E" box="[864,1070,1026,1050]" italics="true" pageId="18" pageNumber="376">Philyra heterograna</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFAC12562C4CFBAD80EE1B4E" author="Ortmann, A." box="[1077,1234,1026,1050]" pageId="18" pageNumber="376" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFAC125628EEFB8F80ED1B6D" blockId="18.[151,1436,903,1081]" box="[151,1233,1056,1081]" pageId="18" pageNumber="376">
|
||||
<treatmentCitationGroup id="AB6411E2FFAC125628EEFB8F80ED1B6D" box="[151,1233,1056,1081]" pageId="18" pageNumber="376">
|
||||
<emphasis id="B900EADEFFAC125628EEFB8F85BA1B6C" box="[151,390,1056,1080]" italics="true" pageId="18" pageNumber="376">
|
||||
<taxonomicName id="4C744D4FFFAC125628EEFB8F85541B6C" ID-CoL="3WTBK" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,360,1056,1080]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="heterograna">Lyphira heterograna</taxonomicName>
|
||||
.—
|
||||
</emphasis>
|
||||
<treatmentCitation id="0AD510DDFFAC125629FFFB8E86611B6D" author="Wong, K. J. H. & Tao, S. - R. & Leung, K. M. Y." box="[390,605,1056,1081]" page="20" pageId="18" pageNumber="376" year="2021">
|
||||
<bibRefCitation id="EFE54B3DFFAC125629FFFB8E86611B6D" author="Wong, K. J. H. & Tao, S. - R. & Leung, K. M. Y." box="[390,605,1056,1081]" pageId="18" pageNumber="376" pagination="1 - 74" refId="ref22379" refString="Wong, K. J. H., Tao, S. - R. & Leung, K. M. Y. (2021) Subtidal crabs of Hong Kong: Brachyura (Crustacea: Decapoda) from benthic trawl surveys conducted by the University of Hong Kong, 2012 to 2018. Regional Studies in Marine Science, 48, 1 - 74. 102013. https: // doi. org / 10.1016 / j. rsma. 2021.102013" type="journal article" year="2021">
|
||||
Wong
|
||||
<emphasis id="B900EADEFFAC125629B1FB8E85C01B6C" box="[456,508,1056,1080]" italics="true" pageId="18" pageNumber="376">et al.</emphasis>
|
||||
2021: 20
|
||||
</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
, fig. 30a–c, pl. 6C. (not
|
||||
<taxonomicName id="4C744D4FFFAC12562B2AFB8F80FA1B6D" ID-CoL="76YMK" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[851,1222,1056,1081]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFAC12562B2AFB8F801D1B6C" box="[851,1057,1056,1080]" italics="true" pageId="18" pageNumber="376">Philyra heterograna</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFAC12562C51FB8E80FA1B6D" author="Ortmann, A." box="[1064,1222,1057,1081]" pageId="18" pageNumber="376" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
</taxonomicName>
|
||||
).
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFAC125628EEFBC880F41BB9" pageId="18" pageNumber="376" type="materials_examined">
|
||||
<paragraph id="8BCB36CCFFAC125628EEFBC880F41BB9" blockId="18.[151,1437,1126,2017]" pageId="18" pageNumber="376">
|
||||
<emphasis id="B900EADEFFAC125628EEFBC885021BD4" bold="true" box="[151,318,1126,1153]" pageId="18" pageNumber="376">Type material</emphasis>
|
||||
.
|
||||
<materialsCitation id="3B1C3C91FFAC12562934FBC886B71BF1" collectingDate="1998-11-12" collectionCode="ZRC" collectorName="N. K. Ng & Y. Cai" location="Nanao Island" pageId="18" pageNumber="376" specimenCode="ZRC 1999.0456" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="B900EADEFFAC12562934FBC885881BD5" box="[333,436,1127,1153]" italics="true" pageId="18" pageNumber="376">
|
||||
<typeStatus id="54CF886EFFAC12562934FBC885881BD5" box="[333,436,1127,1153]" pageId="18" pageNumber="376" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: male (20.8 × 21.8 mm),
|
||||
<specimenCode id="DBD29EB7FFAC12562AACFBC887B11BD5" box="[725,909,1126,1153]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="376">ZRC 1999.0456</specimenCode>
|
||||
,
|
||||
<location id="8EAB6017FFAC12562BE0FBC8800C1BD5" LSID="urn:lsid:plazi:treatment:03DD87DAFFAC125728EEFCB187691921:8EAB6017FFAC12562BE0FBC8800C1BD5" box="[921,1072,1127,1153]" name="Nanao Island" pageId="18" pageNumber="376">Nanao Island</location>
|
||||
, Gianjing fish port, Guangdong, coll.
|
||||
<collectorName id="2681531AFFAC125628A9FB24850D1BF1" box="[208,305,1163,1189]" pageId="18" pageNumber="376">N.K. Ng</collectorName>
|
||||
&
|
||||
<collectorName id="2681531AFFAC1256292EFB24859D1BF1" box="[343,417,1163,1189]" pageId="18" pageNumber="376">Y. Cai</collectorName>
|
||||
,
|
||||
<date id="FFCA100CFFAC125629D4FB2486B71BF1" box="[429,651,1163,1189]" pageId="18" pageNumber="376" value="1998-11-12">
|
||||
<collectingDate id="EF8EE9E4FFAC125629D4FB2486B71BF1" box="[429,651,1163,1189]" pageId="18" pageNumber="376" value="1998-11-12">12 November 1998</collectingDate>
|
||||
</date>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B1C3C91FFAC12562AEEFB2480AB1BF1" box="[663,1175,1162,1189]" pageId="18" pageNumber="376">
|
||||
<emphasis id="B900EADEFFAC12562AEEFB2487351BF0" box="[663,777,1163,1188]" italics="true" pageId="18" pageNumber="376">
|
||||
<typeStatus id="54CF886EFFAC12562AEEFB2487351BF0" box="[663,777,1163,1188]" pageId="18" pageNumber="376" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="9D72FD45FFAC12562B60FB25874C1BF1" box="[793,880,1162,1189]" count="5" pageId="18" pageNumber="376" type="male">5 males</specimenCount>
|
||||
(largest 20.7 × 21.3 mm)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B1C3C91FFAC12562CDAFB2480F81BB9" collectingDate="2012-08-27" collectionCode="ZRC" country="Hong Kong" elevation="12" latitude="22.215433" location="Lantau Island" longLatPrecision="1" longitude="113.9095" municipality="University Trawl" pageId="18" pageNumber="376" specimenCode="ZRC 2022.0628" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||||
<specimenCount id="9D72FD45FFAC12562CDAFB24812C1BF1" box="[1187,1296,1163,1189]" count="2" pageId="18" pageNumber="376" type="female">2 females</specimenCount>
|
||||
(larger 18.8 × 18.8 mm),
|
||||
<specimenCode id="DBD29EB7FFAC1256294CFB0085CC1B9D" box="[309,496,1199,1225]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="376">ZRC 2022.0628</specimenCode>
|
||||
, station W2, southeast of
|
||||
<location id="8EAB6017FFAC12562B5CFB0087F91B9D" LSID="urn:lsid:plazi:treatment:03DD87DAFFAC125728EEFCB187691921:8EAB6017FFAC12562B5CFB0087F91B9D" box="[805,965,1199,1225]" country="Hong Kong" latitude="22.215433" longLatPrecision="1" longitude="113.9095" municipality="University Trawl" name="Lantau Island" pageId="18" pageNumber="376">Lantau Island</location>
|
||||
,
|
||||
<collectingCountry id="F363765CFFAC12562BADFB0080621B9D" box="[980,1118,1199,1225]" name="Hong Kong" pageId="18" pageNumber="376">Hong Kong</collectingCountry>
|
||||
, 22°11.264'N 113°51.358'E –
|
||||
<geoCoordinate id="EE40500BFFAC125628D5FB7C85051BB9" box="[172,313,1235,1261]" degrees="22" direction="north" minutes="12.926" orientation="latitude" pageId="18" pageNumber="376" precision="1" value="22.215433">22°12.926'N</geoCoordinate>
|
||||
<geoCoordinate id="EE40500BFFAC12562938FB7D85E71BB9" box="[321,475,1234,1261]" degrees="113" direction="east" minutes="54.570" orientation="longitude" pageId="18" pageNumber="376" precision="1" value="113.9095">113°54.570'E</geoCoordinate>
|
||||
, 10.0–11.6 m, coll. Hong Kong
|
||||
<collectingMunicipality id="6BAFACB6FFAC12562B3BFB7C80381BB9" box="[834,1028,1235,1261]" pageId="18" pageNumber="376">University Trawl</collectingMunicipality>
|
||||
,
|
||||
<date id="FFCA100CFFAC12562C76FB7C80F81BB9" box="[1039,1220,1235,1261]" pageId="18" pageNumber="376" value="2012-08-27">
|
||||
<collectingDate id="EF8EE9E4FFAC12562C76FB7C80F81BB9" box="[1039,1220,1235,1261]" pageId="18" pageNumber="376" value="2012-08-27">27 August 2012</collectingDate>
|
||||
</date>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFAC125728BEFB59864B1EF9" lastPageId="19" lastPageNumber="377" pageId="18" pageNumber="376" type="diagnosis">
|
||||
<paragraph id="8BCB36CCFFAC125728BEFB5980641EDD" blockId="18.[151,1437,1126,2017]" lastBlockId="19.[151,1437,150,1653]" lastPageId="19" lastPageNumber="377" pageId="18" pageNumber="376">
|
||||
<emphasis id="B900EADEFFAC125628BEFB5985061A44" bold="true" box="[199,314,1270,1296]" pageId="18" pageNumber="376">Diagnosis</emphasis>
|
||||
. Carapace approximately as broad as long, rounded (
|
||||
<figureCitation id="134F2A49FFAC12562BF1FB5887DB1A45" box="[904,999,1271,1297]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="18" pageNumber="376">Figs. 1E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562B8BFB59800B1A45" box="[1010,1079,1270,1297]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="18" pageNumber="376">5A, B</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562C3BFB5880541A45" box="[1090,1128,1271,1297]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="18" pageNumber="376">6D</figureCitation>
|
||||
); dorsal surface of carapace covered with closely spaced minute granules, hepatic, branchial and intestinal regions with slightly larger, more prominent granules;. Frontal margin granulated; postfrontal lobe slightly convex with slight concave medially (
|
||||
<figureCitation id="134F2A49FFAC12562D1DFA9084FA1A29" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="18" pageNumber="376">Figs. 11E</figureCitation>
|
||||
). Endostomial spine strong, produced, visible from dorsal view (
|
||||
<figureCitation id="134F2A49FFAC12562BC0FACC80241A29" box="[953,1048,1379,1405]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="18" pageNumber="376">Figs. 1F</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562C51FACC80451A29" box="[1064,1145,1379,1405]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="18" pageNumber="376">13F, G</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562CF1FACC80801A29" box="[1160,1212,1379,1405]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="18" pageNumber="376">14D</figureCitation>
|
||||
). Lateral margin of carapace with row of small granules interspersed by larger granule; posterior margin gently convex (
|
||||
<figureCitation id="134F2A49FFAC12562C99FA28815C1AF5" box="[1248,1376,1414,1441]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="18" pageNumber="376">Figs. 5A, B</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562D13FA2881AC1AF5" box="[1386,1424,1415,1441]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="18" pageNumber="376">6D</figureCitation>
|
||||
). Posterior carapace margin with row of granules, slightly larger granule medially and on each posterolateral corner, with margin between them gently convex (
|
||||
<figureCitation id="134F2A49FFAC12562A0BFA6086DE1ABD" box="[626,738,1487,1513]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="18" pageNumber="376">Figs. 10E</figureCitation>
|
||||
). Third maxilliped exopod 2 times as long as broad, wider than basal part of endopod, forming petaliform structure with strongly convex outer margin, with submarginal row of low to very low granules along inner margin; endopod with completely fused basis-ischium, distinctly longer than merus, with shallow submarginal sulcus near inner margin (
|
||||
<figureCitation id="134F2A49FFAC12562B54F99487B31901" box="[813,911,1595,1621]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="18" pageNumber="376">Figs. 1G</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562BE3F99487F51901" box="[922,969,1595,1621]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="18" pageNumber="376">11E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562BADF99480341901" box="[980,1032,1595,1621]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="18" pageNumber="376">18A</figureCitation>
|
||||
). Chelipeds subequal; merus, carpus and chela surfaces smooth or covered with low, small rounded granules (
|
||||
<figureCitation id="134F2A49FFAC12562BA1F9F0805C192D" box="[984,1120,1630,1657]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="18" pageNumber="376">Figs. 5A, B</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562C17F9F080AF192D" box="[1134,1171,1631,1657]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="18" pageNumber="376">6D</figureCitation>
|
||||
); merus subcylindrical, symmetrical along entire length, surface granulated, large granules on proximal area and along margins, minutes granules distally; carpus smooth, unarmed; chela stout, palm surface smooth, upper and lower margins with row of granules, longitudinal row of relatively large granules on lower outer surface of palm; pollex slightly bent, cutting edge lined with denticles; dactylus 1.4 times as long as palm along upper margin, row of small granules on upper margin, cutting edge with median large tooth lined with denticles, remaining of edge with small denticles (
|
||||
<figureCitation id="134F2A49FFAC12562D54F8BC81B31879" box="[1325,1423,1810,1837]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="18" pageNumber="376">Fig. 15F</figureCitation>
|
||||
). P2–P5 slender, short; merus distinctly longer than carpus and propodus, dactylus longer than propodus, lanceolate, terminating in corneous tips (
|
||||
<figureCitation id="134F2A49FFAC1256299FF8F486511821" box="[486,621,1882,1909]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="18" pageNumber="376">Figs. 5A, B</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAC12562A03F8F4869C1821" box="[634,672,1883,1909]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="18" pageNumber="376">6D</figureCitation>
|
||||
); fourth leg shortest with merus 3.8 times as long as broad, margin lined with small granules, more prominent on ventral margin (
|
||||
<figureCitation id="134F2A49FFAC12562B1FF8D087F218CD" box="[870,974,1919,1945]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="18" pageNumber="376">Fig. 16E</figureCitation>
|
||||
). Thoracic sternites transversely narrow, surface finely granulated, larger granules laterally (
|
||||
<figureCitation id="134F2A49FFAC12562AAFF80C870C18E9" box="[726,816,1955,1981]" captionStart="FIGURE 8" captionStartId="13.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1714]" captionTargetId="figure-20@13.[151,1435,181,1714]" captionTargetPageId="13" captionText="FIGURE 8. Male thoracic sternum and pleon. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="18" pageNumber="376">Fig. 8A</figureCitation>
|
||||
). Male sternopleonal cavity deep, reaching to proximal distance between fused thoracic sternites 1–3. Male pleon narrow, slender, long; somite 1 longitudinally narrow, wide; somites 2–6 fused, forming elongate trapezoidal plate, shallow suture just visible between somites 5 and 6, surface smooth; somite 6 longitudinally subrectangular, surface with one granule submedially, lateral margins gently convex; telson triangular, longer than wide (
|
||||
<figureCitation id="134F2A49FFAD12572A07FF7086E31FAD" box="[638,735,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1H</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572A92FF7087331FAD" box="[747,783,223,249]" captionStart="FIGURE 8" captionStartId="13.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1714]" captionTargetId="figure-20@13.[151,1435,181,1714]" captionTargetPageId="13" captionText="FIGURE 8. Male thoracic sternum and pleon. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="19" pageNumber="377">8A</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572B63FF7087771FAD" box="[794,843,223,249]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="19" pageNumber="377">18B</figureCitation>
|
||||
). G1 elongate, slender, slightly dilated distally; apical process short, bent at an angle of 60º along the longitudinal axis, with blunt tip (
|
||||
<figureCitation id="134F2A49FFAD12572C64FEAC804B1E49" box="[1053,1143,259,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1J</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572CFDFEAC80E81E49" box="[1156,1236,259,285]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="19" pageNumber="377">18C‒E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572C99FEAC810B1E49" box="[1248,1335,259,285]" captionStart="FIGURE 20" captionStartId="29.[151,250,1639,1663]" captionTargetBox="[252,1327,191,1613]" captionTargetId="figure-20@29.[239,1349,181,1615]" captionTargetPageId="29" captionText="FIGURE 20. G1. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; C, L. ngankee n. sp., paratype male (19.2 × 19.9 mm) (ZRC 2022.0628), Hong Kong; D, L. ngankee n. sp., paratype male (15.5 × 15.1 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (16.7 × 17.0 mm) (ZRC 2015.0439); Thailand; G, L. linda n. sp., male (18.4 × 19.0 mm) (ZRC 1985.1632), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. linda n. sp., male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; M, L. linda n. sp., male (22.9 × 21.4 mm) (ZRC 2014.0191), Vietnam; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia; O, L. bellagrana n. sp., male (13.2 × 12.6 mm) (ZRC 2022.0745), Papua, Indonesia; P, L. bellagrana n. sp., male (10.6 × 9.9 mm) (ZRC 2022.0750), Papua, Indonesia. A, C, E–H, K–P, right G1; B, D, I, J, left G1 (laterally inverted)." figureDoi="http://doi.org/10.5281/zenodo.12681975" httpUri="https://zenodo.org/record/12681975/files/figure.png" pageId="19" pageNumber="377">20A‒D</figureCitation>
|
||||
). Female sternopleonal cavity reaches distal part of sternite 1; female pleon broad, convex, smooth, somites 2‒6 fused, suture between somites 2 and 3; telson shorter than its basal width (
|
||||
<figureCitation id="134F2A49FFAD12572B31FEE4879C1E31" box="[840,928,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1I</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572BD7FEE487EF1E31" box="[942,979,331,357]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="19" pageNumber="377">9D</figureCitation>
|
||||
); vulvae spaced apart on sternite 6, each opening weakly crescent-shaped, relatively short, without opercular cover (
|
||||
<figureCitation id="134F2A49FFAD12572B9AFEC080771EDD" box="[995,1099,367,393]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="19" pageNumber="377">Fig. 21D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFAD125728BEFE3D864B1EF9" blockId="19.[151,1437,150,1653]" box="[199,631,402,429]" pageId="19" pageNumber="377">
|
||||
<emphasis id="B900EADEFFAD125728BEFE3D85271EF8" bold="true" box="[199,283,402,428]" pageId="19" pageNumber="377">Colour</emphasis>
|
||||
. Light grey on dorsal surfaces.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFAD125728BEFE1885E71D69" pageId="19" pageNumber="377" type="etymology">
|
||||
<paragraph id="8BCB36CCFFAD125728BEFE1885E71D69" blockId="19.[151,1437,150,1653]" pageId="19" pageNumber="377">
|
||||
<emphasis id="B900EADEFFAD125728BEFE1885741E85" bold="true" box="[199,328,439,465]" pageId="19" pageNumber="377">Etymology</emphasis>
|
||||
. The species is named after the late Ng Ngan Kee, an esteemed colleague and very good friend, departed all too soon in 2022. She was a leader in varunid systematics and also an avid educator and field scientist; the
|
||||
<typeStatus id="54CF886EFFAD125728B8FE50851F1D4D" box="[193,291,511,537]" pageId="19" pageNumber="377" type="holotype">holotype</typeStatus>
|
||||
specimen was actually collected by her during one of her many forays throughout Asia. The name is used as a noun in apposition.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFAD125728BEFDE886791979" pageId="19" pageNumber="377" type="discussion">
|
||||
<paragraph id="8BCB36CCFFAD125728BEFDE8852A1CD5" blockId="19.[151,1437,150,1653]" pageId="19" pageNumber="377">
|
||||
<emphasis id="B900EADEFFAD125728BEFDE885081D35" bold="true" box="[199,308,583,609]" pageId="19" pageNumber="377">Remarks</emphasis>
|
||||
. The difference in the shape of carapace, the P5 merus and the G1 between
|
||||
<taxonomicName id="4C744D4FFFAD12572CF5FDE8813B1D35" authorityName="Rahayu & Ng" authorityYear="2024" box="[1164,1287,583,609]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD12572CF5FDE8813B1D35" box="[1164,1287,583,609]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572D76FDE881721D35" bold="true" box="[1295,1358,583,609]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572D76FDE881721D35" box="[1295,1358,583,609]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAD12572DFFFDE8851C1DD1" authorityName="Ortmann" authorityYear="1892" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFAD12572DFFFDE8851C1DD1" italics="true" pageId="19" pageNumber="377">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
are well defined. In
|
||||
<taxonomicName id="4C744D4FFFAD12572A77FDC486B61DD1" authorityName="Rahayu & Ng" authorityYear="2024" box="[526,650,619,645]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD12572A77FDC486B61DD1" box="[526,650,619,645]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572AEBFDC486EE1DD1" bold="true" box="[658,722,619,645]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572AEBFDC486EE1DD1" box="[658,722,619,645]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, the carapace is relatively more rounded and more convex, the endostomial spine is strongly produced, clearly visible from dorsal view, the P5 merus leg is short, only 3.8 times as long as broad, the exopod of the third maxilliped is broad, strongly convex, and the apical process of G1 short, broad and bent at an angle of 60° along the longitudinal axis (
|
||||
<figureCitation id="134F2A49FFAD12572B2EFD7887C91DA5" box="[855,1013,727,753]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1E–G, J</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572C7BFD7980741DA5" box="[1026,1096,726,753]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="19" pageNumber="377">5A, B</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572C2CFD7880451DA5" box="[1109,1145,727,753]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="19" pageNumber="377">6D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572CFFFD7880E11DA5" box="[1158,1245,727,753]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="19" pageNumber="377">13 F, G</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572C90FD7881201DA5" box="[1257,1308,727,753]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="19" pageNumber="377">14D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572D50FD7881641DA5" box="[1321,1368,727,753]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="19" pageNumber="377">16E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572D1CFD7884F21C41" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="19" pageNumber="377">18A, C‒E</figureCitation>
|
||||
).
|
||||
<taxonomicName id="4C744D4FFFAD1257289BFD5485F01C41" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[226,460,763,789]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" sensu="stricto" species="heterograna">
|
||||
<emphasis id="B900EADEFFAD1257289BFD5485F01C41" box="[226,460,763,789]" italics="true" pageId="19" pageNumber="377">Lyphira heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23357A5FFAD125729ACFD54862E1C41" box="[469,530,763,789]" pageId="19" pageNumber="377" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
has the carapace usually slightly longer than broad, the endostomial spines are short, the P5 merus is relatively longer 4.3 times as long as broad, the exopod of the third maxilliped is narrower, not as convex, and the G1 with apical process long, very slightly bent (
|
||||
<figureCitation id="134F2A49FFAD12572BBEFCEC80711C09" box="[967,1101,835,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1A–C</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572C23FCEC809D1C09" box="[1114,1185,835,861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1924,1948]" captionTargetBox="[151,1435,813,1901]" captionTargetId="figure-285@4.[151,1435,813,1901]" captionTargetPageId="4" captionText="FIGURE 2. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191) (specimen number Ortmann 658), Japan.A, overall dorsal habitus; B, overall dorsal habitus (carapace slightly tilted posteriorly); C, overall ventral habitus; D, anterior thoracic sternum, buccal cavity and third maxillipeds." figureDoi="http://doi.org/10.5281/zenodo.12681930" httpUri="https://zenodo.org/record/12681930/files/figure.png" pageId="19" pageNumber="377">2A, B</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572CD6FCEC80C91C09" box="[1199,1269,835,861]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="19" pageNumber="377">3A, B</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572D7AFCEC81091C09" box="[1283,1333,835,861]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="19" pageNumber="377">13A</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572D3AFCEC81A41C09" box="[1347,1432,835,861]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="19" pageNumber="377">16A, B</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD125728EEFCC885361CD5" box="[151,266,871,897]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="19" pageNumber="377">17A, C‒F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFAD125728BEFC2485BD1CB9" blockId="19.[151,1437,150,1653]" pageId="19" pageNumber="377">
|
||||
<taxonomicName id="4C744D4FFFAD125728BEFC2485BF1CF1" authorityName="Rahayu & Ng" authorityYear="2024" box="[199,387,907,933]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="sp. nov.">
|
||||
<emphasis id="B900EADEFFAD125728BEFC2485BF1CF1" box="[199,387,907,933]" italics="true" pageId="19" pageNumber="377">Lyphira ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD125729F0FC2485F41CF1" bold="true" box="[393,456,907,933]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD125729F0FC2485F41CF1" box="[393,456,907,933]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is perhaps most similar to
|
||||
<taxonomicName id="4C744D4FFFAD12572A8AFC2487761CF1" authorityName="Rahayu & Ng" authorityYear="2024" box="[755,842,907,933]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD12572A8AFC2487761CF1" box="[755,842,907,933]" italics="true" pageId="19" pageNumber="377">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572B29FC2487B21CF1" bold="true" box="[848,910,907,933]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572B29FC2487B21CF1" box="[848,910,907,933]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, sharing the characters of a strong endostomial spine and a wide exopod of the third maxilliped. The differences between the two are discussed under the remarks for the latter species.
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFAD125728BEFC5886BD1A59" blockId="19.[151,1437,150,1653]" pageId="19" pageNumber="377">
|
||||
<taxonomicName id="4C744D4FFFAD125728BEFC5885BB1B45" authorityName="Rahayu & Ng" authorityYear="2024" box="[199,391,1015,1041]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="sp. nov.">
|
||||
<emphasis id="B900EADEFFAD125728BEFC5885BB1B45" box="[199,391,1015,1041]" italics="true" pageId="19" pageNumber="377">Lyphira ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD125729EAFC5885EA1B45" bold="true" box="[403,470,1015,1041]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD125729EAFC5885EA1B45" box="[403,470,1015,1041]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAD12572A6CFC58864C1B45" authorityName="Rahayu & Ng" authorityYear="2024" box="[533,624,1015,1041]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD12572A6CFC58864C1B45" box="[533,624,1015,1041]" italics="true" pageId="19" pageNumber="377">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572A02FC5886821B45" bold="true" box="[635,702,1015,1041]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572A02FC5886821B45" box="[635,702,1015,1041]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
are superficially similar to
|
||||
<taxonomicName id="4C744D4FFFAD12572C71FC5880541B44" authorityName="Galil" authorityYear="2009" box="[1032,1128,1015,1040]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ovata">
|
||||
<emphasis id="B900EADEFFAD12572C71FC5880541B44" box="[1032,1128,1015,1040]" italics="true" pageId="19" pageNumber="377">L. ovata</emphasis>
|
||||
</taxonomicName>
|
||||
, described from
|
||||
<specimenCount id="9D72FD45FFAD12572D4BFC5881A01B45" box="[1330,1436,1015,1041]" count="1" pageId="19" pageNumber="377" type="male">one male</specimenCount>
|
||||
specimen from
|
||||
<collectingCountry id="F363765CFFAD12572930FBB485811B61" box="[329,445,1051,1077]" name="Sri Lanka" pageId="19" pageNumber="377">Sri Lanka</collectingCountry>
|
||||
.
|
||||
<taxonomicName id="4C744D4FFFAD125729B0FBB486541B60" authorityName="Galil" authorityYear="2009" box="[457,616,1051,1077]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ovata">
|
||||
<emphasis id="B900EADEFFAD125729B0FBB486541B60" box="[457,616,1051,1077]" italics="true" pageId="19" pageNumber="377">Lyphira ovata</emphasis>
|
||||
</taxonomicName>
|
||||
, however, can easily be separated from these two species in having the dorsal carapace surface more densely covered with granules and the anterolateral margin is almost straight in dorsal view (cf.
|
||||
<bibRefCitation id="EFE54B3DFFAD1257297DFBCC85BC1B29" author="Galil, B. S." box="[260,384,1123,1149]" pageId="19" pageNumber="377" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
|
||||
: fig. 16A). In
|
||||
<taxonomicName id="4C744D4FFFAD12572A58FBCC86A11B29" authorityName="Rahayu & Ng" authorityYear="2024" box="[545,669,1123,1149]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD12572A58FBCC86A11B29" box="[545,669,1123,1149]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572ADCFBCC86D91B29" bold="true" box="[677,741,1123,1149]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572ADCFBCC86D91B29" box="[677,741,1123,1149]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAD12572B67FBCC874A1B29" authorityName="Rahayu & Ng" authorityYear="2024" box="[798,886,1123,1149]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD12572B67FBCC874A1B29" box="[798,886,1123,1149]" italics="true" pageId="19" pageNumber="377">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572B07FBCC87821B29" bold="true" box="[894,958,1123,1149]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572B07FBCC87821B29" box="[894,958,1123,1149]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, there are distinctly fewer granules on the dorsal carapace surface and the anterolateral margin is distinctly convex in dorsal view (
|
||||
<figureCitation id="134F2A49FFAD12572C1DFB28808F1BF5" box="[1124,1203,1159,1185]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="19" pageNumber="377">Figs. 4</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572CC4FB29813D1BF5" box="[1213,1281,1158,1185]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="19" pageNumber="377">5A, B</figureCitation>
|
||||
).
|
||||
<bibRefCitation id="EFE54B3DFFAD12572D6CFB2884F71B91" author="Galil, B. S." pageId="19" pageNumber="377" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil (2009: 303)</bibRefCitation>
|
||||
did not figure the G1, only describing it as “with apical process digitate, squat”, suggesting it is similar to the condition observed for
|
||||
<taxonomicName id="4C744D4FFFAD125729E3FB6086281BBD" authorityName="Rahayu & Ng" authorityYear="2024" box="[410,532,1231,1257]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD125729E3FB6086281BBD" box="[410,532,1231,1257]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572A63FB6086641BBD" bold="true" box="[538,600,1231,1257]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572A63FB6086641BBD" box="[538,600,1231,1257]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAD12572AF7FB6086D91BBD" authorityName="Rahayu & Ng" authorityYear="2024" box="[654,741,1231,1257]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD12572AF7FB6086D91BBD" box="[654,741,1231,1257]" italics="true" pageId="19" pageNumber="377">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572A92FB6087151BBD" bold="true" box="[747,809,1231,1257]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572A92FB6087151BBD" box="[747,809,1231,1257]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, in which the distal process is short and not prominently elongate (
|
||||
<figureCitation id="134F2A49FFAD1257297CFB5C85F01A59" box="[261,460,1267,1293]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="19" pageNumber="377">Figs.18D, E, O, P</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD125729A1FB5C86491A59" box="[472,629,1267,1293]" captionStart="FIGURE 20" captionStartId="29.[151,250,1639,1663]" captionTargetBox="[252,1327,191,1613]" captionTargetId="figure-20@29.[239,1349,181,1615]" captionTargetPageId="29" captionText="FIGURE 20. G1. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; C, L. ngankee n. sp., paratype male (19.2 × 19.9 mm) (ZRC 2022.0628), Hong Kong; D, L. ngankee n. sp., paratype male (15.5 × 15.1 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (16.7 × 17.0 mm) (ZRC 2015.0439); Thailand; G, L. linda n. sp., male (18.4 × 19.0 mm) (ZRC 1985.1632), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. linda n. sp., male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; M, L. linda n. sp., male (22.9 × 21.4 mm) (ZRC 2014.0191), Vietnam; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia; O, L. bellagrana n. sp., male (13.2 × 12.6 mm) (ZRC 2022.0745), Papua, Indonesia; P, L. bellagrana n. sp., male (10.6 × 9.9 mm) (ZRC 2022.0750), Papua, Indonesia. A, C, E–H, K–P, right G1; B, D, I, J, left G1 (laterally inverted)." figureDoi="http://doi.org/10.5281/zenodo.12681975" httpUri="https://zenodo.org/record/12681975/files/figure.png" pageId="19" pageNumber="377">20A‒D, E‒M</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFAD125728BEFAB886791979" blockId="19.[151,1437,150,1653]" pageId="19" pageNumber="377">
|
||||
The wide exopod of the third maxilliped of
|
||||
<taxonomicName id="4C744D4FFFAD12572AD7FAB887141A65" authorityName="Rahayu & Ng" authorityYear="2024" box="[686,808,1303,1329]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD12572AD7FAB887141A65" box="[686,808,1303,1329]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572B56FAB887511A65" bold="true" box="[815,877,1303,1329]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572B56FAB887511A65" box="[815,877,1303,1329]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
similar to those of
|
||||
<taxonomicName id="4C744D4FFFAD12572C3DFAB8808B1A64" authorityName="Trivedi, Soni & Vachhrajani" authorityYear="2016" box="[1092,1207,1303,1328]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="georgei">
|
||||
<emphasis id="B900EADEFFAD12572C3DFAB8808B1A64" box="[1092,1207,1303,1328]" italics="true" pageId="19" pageNumber="377">L. georgei</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAD12572C95FAB881571A65" authorityName="Galil" authorityYear="2009" box="[1260,1387,1303,1329]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="perplexa">
|
||||
<emphasis id="B900EADEFFAD12572C95FAB881571A65" box="[1260,1387,1303,1329]" italics="true" pageId="19" pageNumber="377">L. perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
, but differences can be seen in the more rounded carapace in
|
||||
<taxonomicName id="4C744D4FFFAD12572B76FA9487B61A01" authorityName="Rahayu & Ng" authorityYear="2024" box="[783,906,1339,1365]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD12572B76FA9487B61A01" box="[783,906,1339,1365]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572BE8FA9487F31A01" bold="true" box="[913,975,1339,1365]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572BE8FA9487F31A01" box="[913,975,1339,1365]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="134F2A49FFAD12572BA6FA9480031A01" box="[991,1087,1339,1365]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572C32FA9580AC1A01" box="[1099,1168,1338,1365]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="19" pageNumber="377">5A, B</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572CE2FA9480821A01" box="[1179,1214,1339,1365]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="19" pageNumber="377">6D</figureCitation>
|
||||
) (longer than broad in
|
||||
<taxonomicName id="4C744D4FFFAD125728CDFAF0851A1A2C" authorityName="Trivedi, Soni & Vachhrajani" authorityYear="2016" box="[180,294,1375,1400]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="georgei">
|
||||
<emphasis id="B900EADEFFAD125728CDFAF0851A1A2C" box="[180,294,1375,1400]" italics="true" pageId="19" pageNumber="377">L. georgei</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAD12572925FAF085E71A2D" authorityName="Galil" authorityYear="2009" box="[348,475,1375,1401]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="perplexa">
|
||||
<emphasis id="B900EADEFFAD12572925FAF085E71A2D" box="[348,475,1375,1401]" italics="true" pageId="19" pageNumber="377">L. perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
), the presence of strong endostomial spines in
|
||||
<taxonomicName id="4C744D4FFFAD12572B90FAF0805F1A2D" authorityName="Rahayu & Ng" authorityYear="2024" box="[1001,1123,1375,1401]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee">
|
||||
<emphasis id="B900EADEFFAD12572B90FAF0805F1A2D" box="[1001,1123,1375,1401]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="134F2A49FFAD12572C0AFAF0813C1A2D" box="[1139,1280,1375,1401]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="19" pageNumber="377">Figs. 13F, G</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFAD12572D72FAF081011A2D" box="[1291,1341,1375,1401]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="19" pageNumber="377">14D</figureCitation>
|
||||
) (absent in
|
||||
<taxonomicName id="4C744D4FFFAD125728CAFA2C851A1AC8" authorityName="Trivedi, Soni & Vachhrajani" authorityYear="2016" box="[179,294,1411,1436]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="georgei">
|
||||
<emphasis id="B900EADEFFAD125728CAFA2C851A1AC8" box="[179,294,1411,1436]" italics="true" pageId="19" pageNumber="377">L. georgei</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAD12572922FA2C85E61AC9" authorityName="Galil" authorityYear="2009" box="[347,474,1411,1437]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="perplexa">
|
||||
<emphasis id="B900EADEFFAD12572922FA2C85E61AC9" box="[347,474,1411,1437]" italics="true" pageId="19" pageNumber="377">L. perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
), and the P5 merus is proportionately shorter in
|
||||
<taxonomicName id="4C744D4FFFAD12572B8DFA2C80521AC9" authorityName="Rahayu & Ng" authorityYear="2024" box="[1012,1134,1411,1437]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee">
|
||||
<emphasis id="B900EADEFFAD12572B8DFA2C80521AC9" box="[1012,1134,1411,1437]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
, 3.8 times as long as broad (
|
||||
<figureCitation id="134F2A49FFAD125728E6FA08853E1A95" box="[159,258,1447,1473]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="19" pageNumber="377">Fig. 16E</figureCitation>
|
||||
) (proportionately longer, being 4 times in
|
||||
<taxonomicName id="4C744D4FFFAD12572AA2FA0887661A95" authorityName="Galil" authorityYear="2009" box="[731,858,1447,1473]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="perplexa">
|
||||
<emphasis id="B900EADEFFAD12572AA2FA0887661A95" box="[731,858,1447,1473]" italics="true" pageId="19" pageNumber="377">L. perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
but much shorter in
|
||||
<emphasis id="B900EADEFFAD12572C3BFA0880861A94" box="[1090,1210,1447,1472]" italics="true" pageId="19" pageNumber="377">
|
||||
<taxonomicName id="4C744D4FFFAD12572C3BFA08808A1A94" authorityName="Trivedi, Soni & Vachhrajani" authorityYear="2016" box="[1090,1206,1447,1472]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="georgei">L. georgei</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
3.2 times as long as broad) (cf.
|
||||
<bibRefCitation id="EFE54B3DFFAD1257296FFA6485AE1AB1" author="Galil, B. S." box="[278,402,1483,1509]" pageId="19" pageNumber="377" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
|
||||
: fig. 17A, B;
|
||||
<bibRefCitation id="EFE54B3DFFAD12572A55FA64873E1AB1" author="Trivedi, J. N. & Soni, G. M. & Vachhrajani, K. D." box="[556,770,1483,1509]" pageId="19" pageNumber="377" pagination="148 - 154" refId="ref22214" refString="Trivedi, J. N., Soni, G. M. & Vachhrajani, K. D. (2016) A new species of Lyphira Galil, 2009 (Crustacea, Decapoda, Leucosiidae) from Gujarat, India. Tropical Zoology, 29 (3), 148 - 154. https: // doi. org / 10.1080 / 03946975.2016.1181954" type="journal article" year="2016">
|
||||
Trivedi
|
||||
<emphasis id="B900EADEFFAD12572AFCFA6486821AB1" box="[645,702,1483,1509]" italics="true" pageId="19" pageNumber="377">et al.</emphasis>
|
||||
2016
|
||||
</bibRefCitation>
|
||||
: figs. 1a, b). Their colours in life are also different, with
|
||||
<taxonomicName id="4C744D4FFFAD12572DFFFA6484C8195D" authorityName="Rahayu & Ng" authorityYear="2024" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAD12572DFFFA6484C8195D" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAD12572882FA408505195D" bold="true" box="[251,313,1519,1545]" pageId="19" pageNumber="377">
|
||||
<taxonomicNameLabel id="A23357A5FFAD12572882FA408505195D" box="[251,313,1519,1545]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
being greyish white overall while the carapace of
|
||||
<taxonomicName id="4C744D4FFFAD12572B1EFA4087E5195C" authorityName="Trivedi, Soni & Vachhrajani" authorityYear="2016" box="[871,985,1519,1544]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="georgei">
|
||||
<emphasis id="B900EADEFFAD12572B1EFA4087E5195C" box="[871,985,1519,1544]" italics="true" pageId="19" pageNumber="377">L. georgei</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAD12572C74FA4080B0195D" authorityName="Galil" authorityYear="2009" box="[1037,1164,1519,1545]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="perplexa">
|
||||
<emphasis id="B900EADEFFAD12572C74FA4080B0195D" box="[1037,1164,1519,1545]" italics="true" pageId="19" pageNumber="377">L. perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
are orange to purple (cf.
|
||||
<bibRefCitation id="EFE54B3DFFAD125728EEF9BC85571979" author="Trivedi, J. N. & Soni, G. M. & Vachhrajani, K. D." box="[151,363,1555,1581]" pageId="19" pageNumber="377" pagination="148 - 154" refId="ref22214" refString="Trivedi, J. N., Soni, G. M. & Vachhrajani, K. D. (2016) A new species of Lyphira Galil, 2009 (Crustacea, Decapoda, Leucosiidae) from Gujarat, India. Tropical Zoology, 29 (3), 148 - 154. https: // doi. org / 10.1080 / 03946975.2016.1181954" type="journal article" year="2016">
|
||||
Trivedi
|
||||
<emphasis id="B900EADEFFAD12572889F9BC85141979" box="[240,296,1555,1581]" italics="true" pageId="19" pageNumber="377">et al.</emphasis>
|
||||
2016
|
||||
</bibRefCitation>
|
||||
; unpublished data).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
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<paragraph id="8BCB36CCFFAD125728BEF99886D01905" blockId="19.[151,1437,150,1653]" box="[199,748,1590,1617]" pageId="19" pageNumber="377">
|
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<materialsCitation id="3B1C3C91FFAD125728BEF99886D41905" box="[199,744,1590,1617]" country="China" location="Nanao Island" pageId="19" pageNumber="377" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
|
||||
<emphasis id="B900EADEFFAD125728BEF998855C1904" bold="true" box="[199,352,1590,1617]" pageId="19" pageNumber="377">
|
||||
<typeStatus id="54CF886EFFAD125728BEF998853E1905" box="[199,258,1591,1617]" pageId="19" pageNumber="377">Type</typeStatus>
|
||||
locality
|
||||
</emphasis>
|
||||
.
|
||||
<location id="8EAB6017FFAD12572917F99886391905" LSID="urn:lsid:plazi:treatment:03DD87DAFFAC125728EEFCB187691921:8EAB6017FFAD12572917F99886391905" box="[366,517,1591,1617]" country="China" name="Nanao Island" pageId="19" pageNumber="377" stateProvince="Guangdong">Nanao Island</location>
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,
|
||||
<collectingRegion id="49B0F82EFFAD12572A69F99886AA1905" box="[528,662,1591,1617]" country="China" name="Guangdong" pageId="19" pageNumber="377">Guangdong</collectingRegion>
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||||
,
|
||||
<collectingCountry id="F363765CFFAD12572AD8F99886D41905" box="[673,744,1591,1617]" name="China" pageId="19" pageNumber="377">China</collectingCountry>
|
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</materialsCitation>
|
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.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFAD125728BEF9F587691921" box="[199,853,1626,1653]" pageId="19" pageNumber="377" type="distribution">
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<paragraph id="8BCB36CCFFAD125728BEF9F587691921" blockId="19.[151,1437,150,1653]" box="[199,853,1626,1653]" pageId="19" pageNumber="377">
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<emphasis id="B900EADEFFAD125728BEF9F585651920" bold="true" box="[199,345,1626,1652]" pageId="19" pageNumber="377">Distribution</emphasis>
|
||||
. Nanao Island, Guangdong, and
|
||||
<collectingCountry id="F363765CFFAD12572AB3F9F4876D1921" box="[714,849,1627,1653]" name="Hong Kong" pageId="19" pageNumber="377">Hong Kong</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
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</document>
|
||||
1009
data/03/DD/87/03DD87DAFFAD125128EEF969807318AF.xml
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866
data/03/DD/87/03DD87DAFFB4125628EEFDE9875B1D99.xml
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866
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@ -0,0 +1,866 @@
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<taxonomicName id="4C744D4FFFB4124E28EEFDE9856A1D35" authorityName="Rahayu & Ng" authorityYear="2024" box="[151,342,582,609]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="chomel" status="sp. nov.">
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<emphasis id="B900EADEFFB4124E28EEFDE9856A1D35" bold="true" box="[151,342,582,609]" italics="true" pageId="10" pageNumber="368">Lyphira chomel</emphasis>
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<taxonomicNameLabel id="A23357A5FFB4124E2924FDE885A31D35" box="[349,415,583,609]" pageId="10" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BCB36CCFFB4124E28EEFDC480051DD1" blockId="10.[151,1081,582,645]" box="[151,1081,618,645]" pageId="10" pageNumber="368">
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(
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<figureCitation id="134F2A49FFB4124E28E6FDC4851E1DD1" box="[159,290,619,645]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="10" pageNumber="368">Figs. 3E‒G</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2957FDC4856D1DD1" box="[302,337,619,645]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="10" pageNumber="368">6C</figureCitation>
|
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,
|
||||
<figureCitation id="134F2A49FFB4124E2924FDC485421DD1" box="[349,382,619,645]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="10" pageNumber="368">7E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E29F3FDC485901DD1" box="[394,428,619,645]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="10" pageNumber="368">9C</figureCitation>
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,
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<figureCitation id="134F2A49FFB4124E29C0FDC485D71DD1" box="[441,491,619,645]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="10" pageNumber="368">10D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E298EFDC486141DD1" box="[503,552,619,645]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="10" pageNumber="368">11D</figureCitation>
|
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,
|
||||
<figureCitation id="134F2A49FFB4124E2A54FDC486431DD1" box="[557,639,619,645]" captionStart="FIGURE 12" captionStartId="17.[151,250,1628,1652]" captionTargetBox="[151,1435,181,1605]" captionTargetId="figure-20@17.[151,1435,181,1605]" captionTargetPageId="17" captionText="FIGURE 12. Third maxillipeds. A, Lyphira chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; B, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; C, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore; D, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.97), Malaysia; E, L. linda n. sp., female (17.0 × 16.9 mm) (ZRC 1995.0547), Singapore; F, L. linda n. sp., male (12.1 × 12.4 mm) (ZRC 1995.0547), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681950" httpUri="https://zenodo.org/record/12681950/files/figure.png" pageId="10" pageNumber="368">12B, C</figureCitation>
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,
|
||||
<figureCitation id="134F2A49FFB4124E2AF3FDC486E71DD1" box="[650,731,619,645]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="10" pageNumber="368">13D, E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2A91FDC487241DD1" box="[744,792,619,645]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="10" pageNumber="368">14C</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2B5DFDC587681DD1" box="[804,852,618,645]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="10" pageNumber="368">15E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2B26FDC487AE1DD1" box="[863,914,619,645]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="10" pageNumber="368">16D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2BE7FDC487CA1DD1" box="[926,1014,619,645]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="10" pageNumber="368">17M‒R</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2C78FDC4800D1DD1" box="[1025,1073,619,645]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="10" pageNumber="368">21C</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFB4124E28EEFD0080891D93" box="[151,1205,687,711]" pageId="10" pageNumber="368" type="reference_group">
|
||||
<paragraph id="8BCB36CCFFB4124E28EEFD0080891D93" blockId="10.[151,1205,687,711]" box="[151,1205,687,711]" pageId="10" pageNumber="368">
|
||||
<treatmentCitationGroup id="AB6411E2FFB4124E28EEFD0080891D93" box="[151,1205,687,711]" pageId="10" pageNumber="368">
|
||||
<emphasis id="B900EADEFFB4124E28EEFD00854D1D93" box="[151,369,687,711]" italics="true" pageId="10" pageNumber="368">
|
||||
<taxonomicName id="4C744D4FFFB4124E28EEFD0085511D93" ID-CoL="3WTBK" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,365,687,711]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="heterograna">Lyphira heterograna</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
—
|
||||
<treatmentCitation id="0AD510DDFFB4124E29F2FD0086121D93" author="Galil, B. S." box="[395,558,687,711]" httpUri="http://treatment.plazi.org/id/039D87BCFFBF4A26FD4BC4ADFDE1ECDD" page="300" pageId="10" pageNumber="368" year="2009">
|
||||
<bibRefCitation id="EFE54B3DFFB4124E29F2FD0086121D93" author="Galil, B. S." box="[395,558,687,711]" pageId="10" pageNumber="368" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil, 2009: 300</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
(part), figs. 12B, 14. (Not
|
||||
<taxonomicName id="4C744D4FFFB4124E2B47FD0080921D93" ID-CoL="76YMK" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[830,1198,687,711]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB4124E2B47FD0080301D93" box="[830,1036,687,711]" italics="true" pageId="10" pageNumber="368">Philyra heterograna</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFB4124E2C6AFD0080921D93" author="Ortmann, A." box="[1043,1198,687,711]" pageId="10" pageNumber="368" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
</taxonomicName>
|
||||
)
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFB4124E28EEFD5A86621B07" pageId="10" pageNumber="368" type="materials_examined">
|
||||
<paragraph id="8BCB36CCFFB4124E28EEFD5A86621B07" blockId="10.[151,1437,757,2008]" pageId="10" pageNumber="368">
|
||||
<emphasis id="B900EADEFFB4124E28EEFD5A857D1C5B" bold="true" box="[151,321,757,783]" pageId="10" pageNumber="368">Type material</emphasis>
|
||||
.
|
||||
<materialsCitation id="3B1C3C91FFB4124E292BFD5A86951C67" collectionCode="ZRC" collectorName="T. H. T. Tan & D. Vandenspiegel" country="Malaysia" location="Pontian" pageId="10" pageNumber="368" specimenCode="ZRC 1999.0798" specimenCount="1" specimenCount-male="1" stateProvince="Johor" typeStatus="holotype">
|
||||
<emphasis id="B900EADEFFB4124E292BFD5A85851C5B" box="[338,441,757,783]" italics="true" pageId="10" pageNumber="368">
|
||||
<typeStatus id="54CF886EFFB4124E292BFD5A85851C5B" box="[338,441,757,783]" pageId="10" pageNumber="368" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="9D72FD45FFB4124E29B2FD5A863D1C5B" box="[459,513,757,783]" pageId="10" pageNumber="368" type="male">male</specimenCount>
|
||||
(10.1 ×
|
||||
<quantity id="4C8C9B29FFB4124E2A10FD5A86E91C5B" box="[617,725,757,783]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="10" pageNumber="368" unit="mm" value="10.3">10.3 mm</quantity>
|
||||
),
|
||||
<specimenCode id="DBD29EB7FFB4124E2A92FD5A879E1C5B" box="[747,930,757,783]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="368">ZRC 1999.0798</specimenCode>
|
||||
, off
|
||||
<location id="8EAB6017FFB4124E2BA4FD5A80091C5B" LSID="urn:lsid:plazi:treatment:03DD87DAFFB4125628EEFDE9875B1D99:8EAB6017FFB4124E2BA4FD5A80091C5B" box="[989,1077,757,783]" country="Malaysia" name="Pontian" pageId="10" pageNumber="368" stateProvince="Johor">Pontian</location>
|
||||
, western
|
||||
<collectingRegion id="49B0F82EFFB4124E2CDCFD5A80D91C5B" box="[1189,1253,757,783]" country="Malaysia" name="Johor" pageId="10" pageNumber="368">Johor</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="F363765CFFB4124E2C8DFD5A81611C5B" box="[1268,1373,757,783]" name="Malaysia" pageId="10" pageNumber="368">Malaysia</collectingCountry>
|
||||
, coll. dredge,
|
||||
<collectorName id="2681531AFFB4124E288AFCB585511C60" box="[243,365,794,820]" pageId="10" pageNumber="368">T.H.T. Tan</collectorName>
|
||||
&
|
||||
<collectorName id="2681531AFFB4124E29EFFCB5865C1C67" box="[406,608,793,820]" pageId="10" pageNumber="368">D. Vandenspiegel</collectorName>
|
||||
, 1992
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B1C3C91FFB4124E2ACFFCB581A11C03" collectionCode="ZRC" collectorName="D. S. Johnson" country="Singapore" elevation="17" location="Sig" municipality="Bedok" pageId="10" pageNumber="368" specimenCode="ZRC 1985.0099" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paratype">
|
||||
<emphasis id="B900EADEFFB4124E2ACFFCB587141C67" box="[694,808,794,819]" italics="true" pageId="10" pageNumber="368">
|
||||
<typeStatus id="54CF886EFFB4124E2ACFFCB587141C67" box="[694,808,794,819]" pageId="10" pageNumber="368" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
: 1 ovigerous
|
||||
<specimenCount id="9D72FD45FFB4124E2BB1FCB680281C67" box="[968,1044,793,819]" pageId="10" pageNumber="368" type="female">female</specimenCount>
|
||||
(9.6 × 9.7 mm),
|
||||
<specimenCode id="DBD29EB7FFB4124E2CA7FCB681A41C60" box="[1246,1432,793,820]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="368">ZRC 1985.0099</specimenCode>
|
||||
, station
|
||||
<location id="8EAB6017FFB4124E2890FC92852D1C03" LSID="urn:lsid:plazi:treatment:03DD87DAFFB4125628EEFDE9875B1D99:8EAB6017FFB4124E2890FC92852D1C03" box="[233,273,829,855]" country="Singapore" municipality="Bedok" name="Sig" pageId="10" pageNumber="368">Sig</location>
|
||||
3/11, on muddy substrate, off
|
||||
<collectingMunicipality id="6BAFACB6FFB4124E2A26FC9286971C03" box="[607,683,829,855]" pageId="10" pageNumber="368">Bedok</collectingMunicipality>
|
||||
,
|
||||
<collectingCountry id="F363765CFFB4124E2ACFFC9287161C03" box="[694,810,829,855]" name="Singapore" pageId="10" pageNumber="368">Singapore</collectingCountry>
|
||||
, 16.5 m (9 fathoms), coll. dredge,
|
||||
<collectorName id="2681531AFFB4124E2CD7FC92817A1C03" box="[1198,1350,829,855]" pageId="10" pageNumber="368">D.S. Johnson</collectorName>
|
||||
, 1960s;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B1C3C91FFB4124E28EEFCCE87D81C2F" box="[151,996,865,892]" collectionCode="ZRC" collectorName="D. S. Johnson" country="Singapore" elevation="17" location="Sig" municipality="Bedok" pageId="10" pageNumber="368" specimenCode="ZRC 1985.0099" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paratype">
|
||||
<specimenCount id="9D72FD45FFB4124E28EEFCCE84D11C2F" box="[151,237,865,891]" count="4" pageId="10" pageNumber="368" type="male">4 males</specimenCount>
|
||||
(8.8 × 9.0 mm, 10.4 × 10.3 mm, 10.5 × 10.6 mm, 12.6 × 12.8 mm)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B1C3C91FFB4124E2B89FCCE811E1C2F" box="[1008,1314,865,892]" pageId="10" pageNumber="368" specimenCount="1" specimenCount-female="1">
|
||||
<specimenCount id="9D72FD45FFB4124E2B89FCCE806D1C2F" box="[1008,1105,865,891]" count="1" pageId="10" pageNumber="368" type="female">1 female</specimenCount>
|
||||
(11.2 × 11.2 mm),
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B1C3C91FFB4124E2D53FCCE86B11C97" collectionCode="ZRC" collectorName="D. S. Johnson" country="Singapore" elevation="17" location="Bedok" pageId="10" pageNumber="368" specimenCode="ZRC 1985.0091, 0096" specimenCount="1" specimenCount-female="1">
|
||||
1 partially crushed
|
||||
<specimenCount id="9D72FD45FFB4124E288EFC2A857F1CCB" box="[247,323,901,927]" pageId="10" pageNumber="368" type="female">female</specimenCount>
|
||||
(13.5 × 13.4 mm),
|
||||
<specimenCode id="DBD29EB7FFB4124E2A5FFC2A86E01CF4" box="[550,732,901,928]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="368">ZRC 1985.0091</specimenCode>
|
||||
–
|
||||
<specimenCode id="DBD29EB7FFB4124E2A92FC2A87191CF4" box="[747,805,901,928]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="368">0096</specimenCode>
|
||||
, station B77, on mud, off
|
||||
<location id="8EAB6017FFB4124E2C2AFC2A80A31CCB" LSID="urn:lsid:plazi:treatment:03DD87DAFFB4125628EEFDE9875B1D99:8EAB6017FFB4124E2C2AFC2A80A31CCB" box="[1107,1183,901,927]" country="Singapore" name="Bedok" pageId="10" pageNumber="368">Bedok</location>
|
||||
,
|
||||
<collectingCountry id="F363765CFFB4124E2CD2FC2A81231CCB" box="[1195,1311,901,927]" name="Singapore" pageId="10" pageNumber="368">Singapore</collectingCountry>
|
||||
, 16.5 m (9 fathoms), coll. dredge,
|
||||
<collectorName id="2681531AFFB4124E29D9FC0686061C97" box="[416,570,937,963]" pageId="10" pageNumber="368">D.S. Johnson</collectorName>
|
||||
, 1960s
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B1C3C91FFB4124E2AE3FC0686CE1B5F" collectionCode="ZRC" collectorName="D. S. Johnson" country="Malaysia" county="Peninsular" elevation="31" latitude="1.35" location="Tanjung Stapa" longLatPrecision="1" longitude="104.13333" municipality="Pengerang" pageId="10" pageNumber="368" specimenCode="ZRC 1985.0098" specimenCount="1" specimenCount-female="1" stateProvince="Johor">
|
||||
1 ovigerous
|
||||
<specimenCount id="9D72FD45FFB4124E2B5EFC06874F1C97" box="[807,883,937,963]" pageId="10" pageNumber="368" type="female">female</specimenCount>
|
||||
(11.4 × 11.1 mm)
|
||||
<specimenCode id="DBD29EB7FFB4124E2C34FC06813A1C90" box="[1101,1286,937,964]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="368">ZRC 1985.0098</specimenCode>
|
||||
, station B78, mud substrate, off
|
||||
<location id="8EAB6017FFB4124E291CFC62863B1CB3" LSID="urn:lsid:plazi:treatment:03DD87DAFFB4125628EEFDE9875B1D99:8EAB6017FFB4124E291CFC62863B1CB3" box="[357,519,973,999]" country="Malaysia" county="Peninsular" latitude="1.35" longLatPrecision="1" longitude="104.13333" municipality="Pengerang" name="Tanjung Stapa" pageId="10" pageNumber="368" stateProvince="Johor">Tanjung Stapa</location>
|
||||
,
|
||||
<collectingMunicipality id="6BAFACB6FFB4124E2A6BFC6186B71CBC" box="[530,651,974,1000]" pageId="10" pageNumber="368">Pengerang</collectingMunicipality>
|
||||
, southeastern
|
||||
<collectingRegion id="49B0F82EFFB4124E2B5EFC62875B1CB3" box="[807,871,973,999]" country="Malaysia" name="Johor" pageId="10" pageNumber="368">Johor</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="62AA4E40FFB4124E2B08FC6287DB1CB3" box="[881,999,973,999]" pageId="10" pageNumber="368">Peninsular</collectingCounty>
|
||||
<collectingCountry id="F363765CFFB4124E2B94FC62806A1CB3" box="[1005,1110,973,999]" name="Malaysia" pageId="10" pageNumber="368">Malaysia</collectingCountry>
|
||||
,
|
||||
<geoCoordinate id="EE40500BFFB4124E2C18FC6280D71CB3" box="[1121,1259,973,999]" degrees="1" direction="north" minutes="21" orientation="latitude" pageId="10" pageNumber="368" precision="1" seconds="00.0" value="1.35">1°21'00.0"N</geoCoordinate>
|
||||
<geoCoordinate id="EE40500BFFB4124E2C88FC6281A41CB3" box="[1265,1432,973,999]" degrees="104" direction="east" minutes="08" orientation="longitude" pageId="10" pageNumber="368" precision="1" seconds="00.0" value="104.13333">104°08'00.0"E</geoCoordinate>
|
||||
, 31 m (17 fathoms), coll. dredge,
|
||||
<collectorName id="2681531AFFB4124E2A7CFC5E86A01B5F" box="[517,668,1009,1035]" pageId="10" pageNumber="368">D.S. Johnson</collectorName>
|
||||
, 1960s;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B1C3C91FFB4124E2A81FC5E81A41B5F" box="[760,1432,1009,1036]" pageId="10" pageNumber="368" specimenCount="3" specimenCount-male="3">
|
||||
<specimenCount id="9D72FD45FFB4124E2A81FC5E87721B5F" box="[760,846,1009,1035]" count="3" pageId="10" pageNumber="368" type="male">3 males</specimenCount>
|
||||
(12.5 × 11.3 mm, 12.7 × 12.7 mm, 13.0 × 12.9 mm)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B1C3C91FFB4124E28EEFBBA86621B07" collectionCode="NHM" collectorName="F. P. Bedford & W. F. Lanchester" country="Malaysia" elevation="2" location="Peninsular" municipality="Peninsular" pageId="10" pageNumber="368" specimenCount="2" specimenCount-female="2">
|
||||
<specimenCount id="9D72FD45FFB4124E28EEFBBA84C51B7B" box="[151,249,1045,1071]" count="2" pageId="10" pageNumber="368" type="female">2 female</specimenCount>
|
||||
(11.6 × 11.4 mm),
|
||||
<collectionCode id="ED65AE09FFB4124E29AFFBB9862B1B64" box="[470,535,1046,1072]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="10" pageNumber="368">NHM</collectionCode>
|
||||
1900.10.22.340–343, on mud,
|
||||
<collectingCountry id="F363765CFFB4124E2B02FBBA87E11B7B" box="[891,989,1045,1071]" name="Malaysia" pageId="10" pageNumber="368">Malacca</collectingCountry>
|
||||
,
|
||||
<collectingMunicipality id="6BAFACB6FFB4124E2B90FBBA80631B7B" box="[1001,1119,1045,1071]" country="Malaysia" name="Peninsular" pageId="10" pageNumber="368">Peninsular</collectingMunicipality>
|
||||
Malaysia, 1–2 m, coll.
|
||||
<collectorName id="2681531AFFB4124E2D15FBB984CF1B07" pageId="10" pageNumber="368">F. P. Bedford</collectorName>
|
||||
&
|
||||
<collectorName id="2681531AFFB4124E296FFB9585F11B07" box="[278,461,1081,1108]" pageId="10" pageNumber="368">W.F. Lanchester</collectorName>
|
||||
, 1899–1900.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFB4124E28BEFBF2859218DB" pageId="10" pageNumber="368" type="diagnosis">
|
||||
<paragraph id="8BCB36CCFFB4124E28BEFBF2861A183F" blockId="10.[151,1437,757,2008]" pageId="10" pageNumber="368">
|
||||
<emphasis id="B900EADEFFB4124E28BEFBF285061B23" bold="true" box="[199,314,1117,1143]" pageId="10" pageNumber="368">Diagnosis</emphasis>
|
||||
. Carapace approximately as broad as long or slightly longer than broad, dorsal surface of carapace covered with closely spaced minute granules, hepatic and intestinal regions with larger, more prominent granules (
|
||||
<figureCitation id="134F2A49FFB4124E28E6FB0A851C1BEB" box="[159,288,1189,1215]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="10" pageNumber="368">Figs. 3E‒G</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2953FB0A85721BEB" box="[298,334,1189,1215]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="10" pageNumber="368">6C</figureCitation>
|
||||
). Frontal margin granulated, postfrontal lobe slightly convex (
|
||||
<figureCitation id="134F2A49FFB4124E2B8CFB0A805F1BEB" box="[1013,1123,1189,1215]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="10" pageNumber="368">Figs. 11D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2C14FB0A80821BEB" box="[1133,1214,1189,1215]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="10" pageNumber="368">13D, E</figureCitation>
|
||||
). Endostomial spine absent (
|
||||
<figureCitation id="134F2A49FFB4124E2896FB6685BD1BB0" box="[239,385,1225,1252]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="10" pageNumber="368">Figs. 13D, E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E29F7FB6685FD1BB7" box="[398,449,1225,1251]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="10" pageNumber="368">14C</figureCitation>
|
||||
). Lateral and posterolateral margin of carapace with row of small granules interspersed by larger granules; posterior carapace margin gently convex with row of larger granules (
|
||||
<figureCitation id="134F2A49FFB4124E2CF5FB4281351A53" box="[1164,1289,1261,1287]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="10" pageNumber="368">Figs. 3E‒F</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2D6FFB4281761A53" box="[1302,1354,1261,1287]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="10" pageNumber="368">10D</figureCitation>
|
||||
). Third maxilliped exopod 2.5 times as long as broad, slightly narrower than basal part of endopod, forming petaliform structure with convex outer margin, endopod with completely fused basis-ischium, longer than merus, with shallow submarginal sulcus near inner margin (
|
||||
<figureCitation id="134F2A49FFB4124E2A26FAF686EF1A27" box="[607,723,1369,1395]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="10" pageNumber="368">Figs. 11D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2A9AFAF687201A27" box="[739,796,1369,1395]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="10" pageNumber="368">17M</figureCitation>
|
||||
). Chelipeds subequal; merus covered by large rounded granules (
|
||||
<figureCitation id="134F2A49FFB4124E297AFAD2855E1AC3" box="[259,354,1405,1431]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="10" pageNumber="368">Figs. 3E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2915FAD285B21AC3" box="[364,398,1405,1431]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="10" pageNumber="368">6C</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E29E1FAD285801AC3" box="[408,444,1405,1431]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="10" pageNumber="368">9C</figureCitation>
|
||||
); surfaces of carpus and propodus covered by closely spaced small granules; chela slender, fingers longer than length of palm, surface with longitudinal sulcus flanked by row of granules; pollex slightly bent, cutting edge lined with denticles, large tooth subproximally; dactylus 1.5 times as long as palm along upper margin, row of small granules on upper margin, cutting edge with median large tooth lined with denticles, remaining of edge with small denticles (
|
||||
<figureCitation id="134F2A49FFB4124E2995F9A2866F197C" box="[492,595,1549,1576]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="10" pageNumber="368">Fig. 15E</figureCitation>
|
||||
). P2–P5 slender, long; merus distinctly longer than carpus and propodus; dactylus longer than propodus, lanceolate, terminating in corneous tips (
|
||||
<figureCitation id="134F2A49FFB4124E2BB4F99E800C191F" box="[973,1072,1585,1611]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="10" pageNumber="368">Figs. 3E</figureCitation>
|
||||
); fourth leg shortest with merus 5.6 times as long as broad, margins almost smooth, lined barely with small granules (
|
||||
<figureCitation id="134F2A49FFB4124E2C21F9FA80FD193B" box="[1112,1217,1621,1647]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="10" pageNumber="368">Fig. 16D</figureCitation>
|
||||
). Thoracic sternites transversely narrow, surface granulated, larger granules laterally (
|
||||
<figureCitation id="134F2A49FFB4124E2B04F9D687E919C7" box="[893,981,1657,1683]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="10" pageNumber="368">Fig. 7E</figureCitation>
|
||||
). Sternopleonal cavity deep, reaching to proximal distance between fused thoracic sternites 1–3. Pleon narrow, somites 2–6 fused, somite 6 longitudinally subrectangular, surface with one low granule subdistally, lateral margins gently convex; telson triangular (
|
||||
<figureCitation id="134F2A49FFB4124E2D4EF96E81AB1988" box="[1335,1431,1729,1756]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="10" pageNumber="368">Figs. 7E</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E28EEF94A84F019AB" box="[151,204,1765,1791]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="10" pageNumber="368">17N</figureCitation>
|
||||
). G1 elongate, slender, same size from proximal to distal area; distal process gently curved, with blunt tip (
|
||||
<figureCitation id="134F2A49FFB4124E2D16F94A84D11877" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="10" pageNumber="368">Fig. 17O‒R</figureCitation>
|
||||
). Female pleon broad, convex, smooth, somites 2‒6 fused, suture between somites 2 and 3; telson as long as its basal width (
|
||||
<figureCitation id="134F2A49FFB4124E293EF88285A21813" box="[327,414,1837,1863]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="10" pageNumber="368">Fig. 9C</figureCitation>
|
||||
); vulvae positioned closer to median part of sternite 6, each opening relatively broad, ovate, without opercular cover (
|
||||
<figureCitation id="134F2A49FFB4124E29CBF8FE8625183F" box="[434,537,1873,1899]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="10" pageNumber="368">Fig. 21C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFB4124E28BEF8DA859218DB" blockId="10.[151,1437,757,2008]" box="[199,430,1909,1936]" pageId="10" pageNumber="368">
|
||||
<emphasis id="B900EADEFFB4124E28BEF8DA852718DB" bold="true" box="[199,283,1909,1935]" pageId="10" pageNumber="368">Colour</emphasis>
|
||||
. Not known.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFB4124E28BEF836858D1883" pageId="10" pageNumber="368" type="etymology">
|
||||
<paragraph id="8BCB36CCFFB4124E28BEF836858D1883" blockId="10.[151,1437,757,2008]" pageId="10" pageNumber="368">
|
||||
<emphasis id="B900EADEFFB4124E28BEF836857418E7" bold="true" box="[199,328,1945,1971]" pageId="10" pageNumber="368">Etymology</emphasis>
|
||||
. The named is derived from the Malay word for small and cute,
|
||||
<taxonomicName id="4C744D4FFFB4124E2C50F83680B118E7" authorityName="Rahayu & Ng" authorityYear="2024" box="[1065,1165,1945,1971]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="chomel">“chomel</taxonomicName>
|
||||
”. The name is used as a Latin noun in apposition.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF0B6644FFB5124F28EEF8B185F018BE" ID-DOI="http://doi.org/10.5281/zenodo.12681938" ID-Zenodo-Dep="12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="11" pageNumber="369" startId="11.[151,250,1822,1846]" targetBox="[242,1346,181,1798]" targetPageId="11" targetType="figure">
|
||||
<paragraph id="8BCB36CCFFB5124F28EEF8B185F018BE" blockId="11.[151,1436,1822,2026]" pageId="11" pageNumber="369">
|
||||
<emphasis id="B900EADEFFB5124F28EEF8B185281862" bold="true" box="[151,276,1822,1846]" pageId="11" pageNumber="369">FIGURE 6.</emphasis>
|
||||
Overall dorsal habitus. A,
|
||||
<taxonomicName id="4C744D4FFFB5124F2A51F8B187931862" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[552,943,1822,1846]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="369" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB5124F2A51F8B186C71862" box="[552,763,1822,1846]" italics="true" pageId="11" pageNumber="369">Lyphira heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB5124F2B70F8B187941862" author="Ortmann, A." box="[777,936,1822,1846]" pageId="11" pageNumber="369" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B,
|
||||
<taxonomicName id="4C744D4FFFB5124F2886F8EC8633180E" authority="(Chen, 1987)" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[255,527,1858,1882]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="369" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB5124F2886F8EC8541180E" box="[255,381,1858,1882]" italics="true" pageId="11" pageNumber="369">L. acutidens</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB5124F29F5F8ED863B180E" author="Chen, H." box="[396,519,1858,1882]" pageId="11" pageNumber="369" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C,
|
||||
<taxonomicName id="4C744D4FFFB5124F2CF7F8EC80C8180E" authorityName="Rahayu & Ng" authorityYear="2024" box="[1166,1268,1858,1882]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="369" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB5124F2CF7F8EC80C8180E" box="[1166,1268,1858,1882]" italics="true" pageId="11" pageNumber="369">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB5124F2C85F8ED810B180E" bold="true" box="[1276,1335,1858,1882]" pageId="11" pageNumber="369">
|
||||
<taxonomicNameLabel id="A23357A5FFB5124F2C85F8ED810B180E" box="[1276,1335,1858,1882]" pageId="11" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D,
|
||||
<taxonomicName id="4C744D4FFFB5124F2ABEF8C88709182A" authorityName="Rahayu & Ng" authorityYear="2024" box="[711,821,1894,1918]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="369" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB5124F2ABEF8C88709182A" box="[711,821,1894,1918]" italics="true" pageId="11" pageNumber="369">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB5124F2B42F8C98746182A" bold="true" box="[827,890,1894,1918]" pageId="11" pageNumber="369">
|
||||
<taxonomicNameLabel id="A23357A5FFB5124F2B42F8C98749182A" box="[827,885,1894,1918]" pageId="11" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
|
||||
,
|
||||
</emphasis>
|
||||
Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E,
|
||||
<taxonomicName id="4C744D4FFFB5124F29B9F824862C18F6" authorityName="Rahayu & Ng" authorityYear="2024" box="[448,528,1930,1954]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="369" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB5124F29B9F824862C18F6" box="[448,528,1930,1954]" italics="true" pageId="11" pageNumber="369">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB5124F2A61F825866F18F6" bold="true" box="[536,595,1930,1954]" pageId="11" pageNumber="369">
|
||||
<taxonomicNameLabel id="A23357A5FFB5124F2A61F825866F18F6" box="[536,595,1930,1954]" pageId="11" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F,
|
||||
<taxonomicName id="4C744D4FFFB5124F2D56F82481BC18F6" authorityName="Rahayu & Ng" authorityYear="2024" box="[1327,1408,1930,1954]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="369" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB5124F2D56F82481BC18F6" box="[1327,1408,1930,1954]" italics="true" pageId="11" pageNumber="369">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB5124F2DF1F825848A1892" bold="true" pageId="11" pageNumber="369">
|
||||
<taxonomicNameLabel id="A23357A5FFB5124F2DF1F825848A1892" pageId="11" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G,
|
||||
<taxonomicName id="4C744D4FFFB5124F2B65F800879B1892" authorityName="Rahayu & Ng" authorityYear="2024" box="[796,935,1966,1990]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="369" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB5124F2B65F800879B1892" box="[796,935,1966,1990]" italics="true" pageId="11" pageNumber="369">L. bellagrana</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB5124F2BD6F80187D61892" bold="true" box="[943,1002,1966,1990]" pageId="11" pageNumber="369">
|
||||
<taxonomicNameLabel id="A23357A5FFB5124F2BD6F80187D61892" box="[943,1002,1966,1990]" pageId="11" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF0B6644FFB2124828EEF97A80E4180D" ID-DOI="http://doi.org/10.5281/zenodo.12681940" ID-Zenodo-Dep="12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="12" pageNumber="370" startId="12.[151,250,1749,1773]" targetBox="[151,1435,181,1725]" targetPageId="12" targetType="figure">
|
||||
<paragraph id="8BCB36CCFFB2124828EEF97A80E4180D" blockId="12.[151,1437,1749,1881]" pageId="12" pageNumber="370">
|
||||
<emphasis id="B900EADEFFB2124828EEF97A852919B9" bold="true" box="[151,277,1749,1773]" pageId="12" pageNumber="370">FIGURE 7.</emphasis>
|
||||
Male thoracic sternum and pleon. A,
|
||||
<taxonomicName id="4C744D4FFFB212482AD9F97A801619B9" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[672,1066,1749,1773]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB212482AD9F97A874819B9" box="[672,884,1749,1773]" italics="true" pageId="12" pageNumber="370">Lyphira heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB212482BFAF97A801F19B9" author="Ortmann, A." box="[899,1059,1749,1773]" pageId="12" pageNumber="370" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B,
|
||||
<taxonomicName id="4C744D4FFFB212482909F95586FD1845" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[368,705,1785,1809]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB212482909F95586351845" box="[368,521,1785,1809]" italics="true" pageId="12" pageNumber="370">L. heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB212482A60F95686861845" author="Ortmann, A." box="[537,698,1785,1809]" pageId="12" pageNumber="370" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C,
|
||||
<taxonomicName id="4C744D4FFFB212482D7AF95585781861" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB212482D7AF95581A01845" box="[1283,1436,1785,1809]" italics="true" pageId="12" pageNumber="370">L. heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB2124828E7F8B285011861" author="Ortmann, A." box="[158,317,1821,1845]" pageId="12" pageNumber="370" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D,
|
||||
<taxonomicName id="4C744D4FFFB212482B09F8B180411861" authority="(Chen, 1987)" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[880,1149,1821,1845]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB212482B09F8B187D01861" box="[880,1004,1821,1845]" italics="true" pageId="12" pageNumber="370">L. acutidens</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB212482B82F8B280491861" author="Chen, H." box="[1019,1141,1821,1845]" pageId="12" pageNumber="370" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E,
|
||||
<taxonomicName id="4C744D4FFFB2124829CAF8ED862B180D" authorityName="Rahayu & Ng" authorityYear="2024" box="[435,535,1857,1881]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB2124829CAF8ED862B180D" box="[435,535,1857,1881]" italics="true" pageId="12" pageNumber="370">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB212482A67F8EE866B180D" bold="true" box="[542,599,1857,1881]" pageId="12" pageNumber="370">
|
||||
<taxonomicNameLabel id="A23357A5FFB212482A67F8EE866B180D" box="[542,599,1857,1881]" pageId="12" pageNumber="370" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF0B6644FFB3124928EEF96586301826" ID-DOI="http://doi.org/10.5281/zenodo.12681942" ID-Zenodo-Dep="12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="13" pageNumber="371" startId="13.[151,250,1738,1762]" targetBox="[151,1435,181,1714]" targetPageId="13" targetType="figure">
|
||||
<paragraph id="8BCB36CCFFB3124928EEF96586301826" blockId="13.[151,1436,1738,1907]" pageId="13" pageNumber="371">
|
||||
<emphasis id="B900EADEFFB3124928EEF965852F19B6" bold="true" box="[151,275,1738,1762]" pageId="13" pageNumber="371">FIGURE 8.</emphasis>
|
||||
Male thoracic sternum and pleon. A,
|
||||
<taxonomicName id="4C744D4FFFB312492AEFF965877C19B6" authorityName="Rahayu & Ng" authorityYear="2024" box="[662,832,1738,1762]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="371" phylum="Arthropoda" rank="species" species="ngankee" status="sp. nov.">
|
||||
<emphasis id="B900EADEFFB312492AEFF965877C19B6" box="[662,832,1738,1762]" italics="true" pageId="13" pageNumber="371">Lyphira ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB312492B31F96487BD19B7" bold="true" box="[840,897,1739,1763]" pageId="13" pageNumber="371">
|
||||
<taxonomicNameLabel id="A23357A5FFB312492B31F96487BD19B7" box="[840,897,1739,1763]" pageId="13" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B,
|
||||
<taxonomicName id="4C744D4FFFB312492935F94085A71852" authorityName="Rahayu & Ng" authorityYear="2024" box="[332,411,1774,1798]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="371" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB312492935F94085A71852" box="[332,411,1774,1798]" italics="true" pageId="13" pageNumber="371">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB3124929D8F94085E51853" bold="true" box="[417,473,1775,1799]" pageId="13" pageNumber="371">
|
||||
<taxonomicNameLabel id="A23357A5FFB3124929D8F94085E51853" box="[417,473,1775,1799]" pageId="13" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C,
|
||||
<taxonomicName id="4C744D4FFFB312492CFAF94080ED1852" authorityName="Rahayu & Ng" authorityYear="2024" box="[1155,1233,1774,1798]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="371" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB312492CFAF94080ED1852" box="[1155,1233,1774,1798]" italics="true" pageId="13" pageNumber="371">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB312492CAEF940812C1853" bold="true" box="[1239,1296,1775,1799]" pageId="13" pageNumber="371">
|
||||
<taxonomicNameLabel id="A23357A5FFB312492CAEF940812C1853" box="[1239,1296,1775,1799]" pageId="13" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D,
|
||||
<taxonomicName id="4C744D4FFFB312492A36F8BC86A1187E" authorityName="Rahayu & Ng" authorityYear="2024" box="[591,669,1810,1834]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="371" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB312492A36F8BC86A1187E" box="[591,669,1810,1834]" italics="true" pageId="13" pageNumber="371">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB312492ADAF8BC86E7187F" bold="true" box="[675,731,1811,1835]" pageId="13" pageNumber="371">
|
||||
<taxonomicNameLabel id="A23357A5FFB312492ADAF8BC86E7187F" box="[675,731,1811,1835]" pageId="13" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E,
|
||||
<taxonomicName id="4C744D4FFFB312492D6AF8BC81A0187E" authorityName="Rahayu & Ng" authorityYear="2024" box="[1299,1436,1810,1834]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="371" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB312492D6AF8BC81A0187E" box="[1299,1436,1810,1834]" italics="true" pageId="13" pageNumber="371">L. bellagrana</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB3124928EEF89884EE181B" bold="true" box="[151,210,1847,1871]" pageId="13" pageNumber="371">
|
||||
<taxonomicNameLabel id="A23357A5FFB3124928EEF89884EE181B" box="[151,210,1847,1871]" pageId="13" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F,
|
||||
<taxonomicName id="4C744D4FFFB312492BA9F8988060181A" authorityName="Rahayu & Ng" authorityYear="2024" box="[976,1116,1846,1870]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="371" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB312492BA9F8988060181A" box="[976,1116,1846,1870]" italics="true" pageId="13" pageNumber="371">L. bellagrana</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB312492C1DF89880A3181B" bold="true" box="[1124,1183,1847,1871]" pageId="13" pageNumber="371">
|
||||
<taxonomicNameLabel id="A23357A5FFB312492C1DF89880A3181B" box="[1124,1183,1847,1871]" pageId="13" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF0B6644FFB0124A28EEF8B1863018BE" ID-DOI="http://doi.org/10.5281/zenodo.12681944" ID-Zenodo-Dep="12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="14" pageNumber="372" startId="14.[151,250,1822,1846]" targetBox="[256,1332,181,1798]" targetPageId="14" targetType="figure">
|
||||
<paragraph id="8BCB36CCFFB0124A28EEF8B1863018BE" blockId="14.[151,1437,1822,2026]" pageId="14" pageNumber="372">
|
||||
<emphasis id="B900EADEFFB0124A28EEF8B1852E1862" bold="true" box="[151,274,1822,1846]" pageId="14" pageNumber="372">FIGURE 9.</emphasis>
|
||||
Female thoracic sternum and pleon. A,
|
||||
<taxonomicName id="4C744D4FFFB0124A2ADDF8B1801B1862" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[676,1063,1822,1846]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="372" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB0124A2ADDF8B187491862" box="[676,885,1822,1846]" italics="true" pageId="14" pageNumber="372">Lyphira heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB0124A2BFBF8B1801C1862" author="Ortmann, A." box="[898,1056,1822,1846]" pageId="14" pageNumber="372" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B,
|
||||
<taxonomicName id="4C744D4FFFB0124A2915F8EC8646180E" authority="(Chen, 1987)" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[364,634,1858,1882]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="372" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB0124A2915F8EC85D5180E" box="[364,489,1858,1882]" italics="true" pageId="14" pageNumber="372">L. acutidens</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB0124A2981F8ED864E180E" author="Chen, H." box="[504,626,1858,1882]" pageId="14" pageNumber="372" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C,
|
||||
<taxonomicName id="4C744D4FFFB0124A2C89F8EC8168180E" authorityName="Rahayu & Ng" authorityYear="2024" box="[1264,1364,1858,1882]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="372" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB0124A2C89F8EC8168180E" box="[1264,1364,1858,1882]" italics="true" pageId="14" pageNumber="372">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB0124A2D25F8ED81AA180E" bold="true" box="[1372,1430,1858,1882]" pageId="14" pageNumber="372">
|
||||
<taxonomicNameLabel id="A23357A5FFB0124A2D25F8ED81AA180E" box="[1372,1430,1858,1882]" pageId="14" pageNumber="372" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D,
|
||||
<taxonomicName id="4C744D4FFFB0124A2B49F8C8879C182A" authorityName="Rahayu & Ng" authorityYear="2024" box="[816,928,1894,1918]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="372" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB0124A2B49F8C8879C182A" box="[816,928,1894,1918]" italics="true" pageId="14" pageNumber="372">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB0124A2BDEF8C987DD182A" bold="true" box="[935,993,1894,1918]" pageId="14" pageNumber="372">
|
||||
<taxonomicNameLabel id="A23357A5FFB0124A2BDEF8C987DD182A" box="[935,993,1894,1918]" pageId="14" pageNumber="372" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E,
|
||||
<taxonomicName id="4C744D4FFFB0124A2A56F824864218F6" authorityName="Rahayu & Ng" authorityYear="2024" box="[559,638,1930,1954]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="372" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB0124A2A56F824864218F6" box="[559,638,1930,1954]" italics="true" pageId="14" pageNumber="372">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB0124A2AFCF825868218F6" bold="true" box="[645,702,1930,1954]" pageId="14" pageNumber="372">
|
||||
<taxonomicNameLabel id="A23357A5FFB0124A2AFCF825868218F6" box="[645,702,1930,1954]" pageId="14" pageNumber="372" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F,
|
||||
<taxonomicName id="4C744D4FFFB0124A2DF1F82484F71892" authorityName="Rahayu & Ng" authorityYear="2024" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="372" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB0124A2DF1F82484F71892" italics="true" pageId="14" pageNumber="372">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB0124A28ABF80185301892" bold="true" box="[210,268,1966,1990]" pageId="14" pageNumber="372">
|
||||
<taxonomicNameLabel id="A23357A5FFB0124A28ABF80185301892" box="[210,268,1966,1990]" pageId="14" pageNumber="372" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G,
|
||||
<taxonomicName id="4C744D4FFFB0124A2B1EF80087CD1892" authorityName="Rahayu & Ng" authorityYear="2024" box="[871,1009,1966,1990]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="372" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB0124A2B1EF80087CD1892" box="[871,1009,1966,1990]" italics="true" pageId="14" pageNumber="372">L. bellagrana</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB0124A2B81F801800E1892" bold="true" box="[1016,1074,1966,1990]" pageId="14" pageNumber="372">
|
||||
<taxonomicNameLabel id="A23357A5FFB0124A2B81F801800E1892" box="[1016,1074,1966,1990]" pageId="14" pageNumber="372" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF0B6644FFB1124B28EEF9FF853D1830" ID-DOI="http://doi.org/10.5281/zenodo.12681946" ID-Zenodo-Dep="12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="15" pageNumber="373" startId="15.[151,250,1616,1640]" targetBox="[151,1435,181,1593]" targetPageId="15" targetType="figure">
|
||||
<paragraph id="8BCB36CCFFB1124B28EEF9FF853D1830" blockId="15.[151,1437,1616,1892]" pageId="15" pageNumber="373">
|
||||
<emphasis id="B900EADEFFB1124B28EEF9FF851E193C" bold="true" box="[151,290,1616,1640]" pageId="15" pageNumber="373">FIGURE 10.</emphasis>
|
||||
Posterior carapace margin. A,
|
||||
<taxonomicName id="4C744D4FFFB1124B2A1FF9FF87CE193D" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[614,1010,1616,1641]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="373" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB1124B2A1FF9FF8706193C" box="[614,826,1616,1640]" italics="true" pageId="15" pageNumber="373">Lyphira heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB1124B2B33F9FE87D7193D" author="Ortmann, A." box="[842,1003,1617,1641]" pageId="15" pageNumber="373" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B,
|
||||
<taxonomicName id="4C744D4FFFB1124B2900F9DA86F519D9" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[377,713,1652,1677]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="373" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB1124B2900F9DA862D19D8" box="[377,529,1652,1676]" italics="true" pageId="15" pageNumber="373">L. heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB1124B2A58F9DA86FE19D9" author="Ortmann, A." box="[545,706,1653,1677]" pageId="15" pageNumber="373" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C,
|
||||
<taxonomicName id="4C744D4FFFB1124B2D67F9DA851C19E5" authority="(Chen, 1987)" baseAuthorityName="Chen" baseAuthorityYear="1987" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="373" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB1124B2D67F9DA81A019D8" box="[1310,1436,1652,1676]" italics="true" pageId="15" pageNumber="373">L. acutidens</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB1124B28E7F937852419E5" author="Chen, H." box="[158,280,1688,1713]" pageId="15" pageNumber="373" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D,
|
||||
<taxonomicName id="4C744D4FFFB1124B2B1BF93687FA19E4" authorityName="Rahayu & Ng" authorityYear="2024" box="[866,966,1688,1712]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="373" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB1124B2B1BF93687FA19E4" box="[866,966,1688,1712]" italics="true" pageId="15" pageNumber="373">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB1124B2BB5F936803A19E5" bold="true" box="[972,1030,1689,1713]" pageId="15" pageNumber="373">
|
||||
<taxonomicNameLabel id="A23357A5FFB1124B2BB5F936803A19E5" box="[972,1030,1689,1713]" pageId="15" pageNumber="373" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E,
|
||||
<taxonomicName id="4C744D4FFFB1124B29DBF912862E1980" authorityName="Rahayu & Ng" authorityYear="2024" box="[418,530,1724,1748]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="373" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB1124B29DBF912862E1980" box="[418,530,1724,1748]" italics="true" pageId="15" pageNumber="373">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB1124B2A63F91286681981" bold="true" box="[538,596,1725,1749]" pageId="15" pageNumber="373">
|
||||
<taxonomicNameLabel id="A23357A5FFB1124B2A63F91286681981" box="[538,596,1725,1749]" pageId="15" pageNumber="373" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F,
|
||||
<taxonomicName id="4C744D4FFFB1124B2D49F91281BC1980" authorityName="Rahayu & Ng" authorityYear="2024" box="[1328,1408,1724,1748]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="373" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB1124B2D49F91281BC1980" box="[1328,1408,1724,1748]" italics="true" pageId="15" pageNumber="373">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB1124B2DF1F912848A19AD" bold="true" pageId="15" pageNumber="373">
|
||||
<taxonomicNameLabel id="A23357A5FFB1124B2DF1F912848A19AD" pageId="15" pageNumber="373" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G,
|
||||
<taxonomicName id="4C744D4FFFB1124B2B27F94E879019AC" authorityName="Rahayu & Ng" authorityYear="2024" box="[862,940,1760,1784]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="373" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB1124B2B27F94E879019AC" box="[862,940,1760,1784]" italics="true" pageId="15" pageNumber="373">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB1124B2BC8F94E87D619AD" bold="true" box="[945,1002,1761,1785]" pageId="15" pageNumber="373">
|
||||
<taxonomicNameLabel id="A23357A5FFB1124B2BC8F94E87D619AD" box="[945,1002,1761,1785]" pageId="15" pageNumber="373" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H,
|
||||
<taxonomicName id="4C744D4FFFB1124B2952F8AA85461848" authorityName="Rahayu & Ng" authorityYear="2024" box="[299,378,1796,1820]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="373" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB1124B2952F8AA85461848" box="[299,378,1796,1820]" italics="true" pageId="15" pageNumber="373">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB1124B29F8F8AA85871849" bold="true" box="[385,443,1797,1821]" pageId="15" pageNumber="373">
|
||||
<taxonomicNameLabel id="A23357A5FFB1124B29F8F8AA85871849" box="[385,443,1797,1821]" pageId="15" pageNumber="373" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I,
|
||||
<taxonomicName id="4C744D4FFFB1124B2B81F8AA80BE1848" authorityName="Rahayu & Ng" authorityYear="2024" box="[1016,1154,1796,1820]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="373" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB1124B2B81F8AA80BE1848" box="[1016,1154,1796,1820]" italics="true" pageId="15" pageNumber="373">L. bellagrana</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB1124B2CF0F8AA80FF1849" bold="true" box="[1161,1219,1797,1821]" pageId="15" pageNumber="373">
|
||||
<taxonomicNameLabel id="A23357A5FFB1124B2CF0F8AA80FF1849" box="[1161,1219,1797,1821]" pageId="15" pageNumber="373" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J,
|
||||
<taxonomicName id="4C744D4FFFB1124B2ACFF886877E1814" authorityName="Rahayu & Ng" authorityYear="2024" box="[694,834,1832,1856]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="373" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB1124B2ACFF886877E1814" box="[694,834,1832,1856]" italics="true" pageId="15" pageNumber="373">L. bellagrana</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB1124B2B32F88687BB1815" bold="true" box="[843,903,1833,1857]" pageId="15" pageNumber="373">
|
||||
<taxonomicNameLabel id="A23357A5FFB1124B2B32F88687BB1815" box="[843,903,1833,1857]" pageId="15" pageNumber="373" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF0B6644FFAE125428EEF972857018A5" ID-DOI="http://doi.org/10.5281/zenodo.12681948" ID-Zenodo-Dep="12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="16" pageNumber="374" startId="16.[151,250,1757,1781]" targetBox="[290,1298,181,1733]" targetPageId="16" targetType="figure">
|
||||
<paragraph id="8BCB36CCFFAE125428EEF972857018A5" blockId="16.[151,1437,1757,2033]" pageId="16" pageNumber="374">
|
||||
<emphasis id="B900EADEFFAE125428EEF972852319A1" bold="true" box="[151,287,1757,1781]" pageId="16" pageNumber="374">FIGURE 11.</emphasis>
|
||||
Frontal view of cephalothorax and third maxilliped. A,
|
||||
<taxonomicName id="4C744D4FFFAE12542B2EF97280E119A1" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[855,1245,1757,1781]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="374" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFAE12542B2EF972801519A1" box="[855,1065,1757,1781]" italics="true" pageId="16" pageNumber="374">Lyphira heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFAE12542C4EF97280EA19A1" author="Ortmann, A." box="[1079,1238,1757,1781]" pageId="16" pageNumber="374" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B,
|
||||
<taxonomicName id="4C744D4FFFAE12542A64F8AD8751184D" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[541,877,1793,1817]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="374" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFAE12542A64F8AD868A184D" box="[541,694,1793,1817]" italics="true" pageId="16" pageNumber="374">L. heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFAE12542ABFF8AE875A184D" author="Ortmann, A." box="[710,870,1793,1817]" pageId="16" pageNumber="374" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C,
|
||||
<taxonomicName id="4C744D4FFFAE125428EEF88985941869" authority="(Chen, 1987)" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[151,424,1829,1853]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="374" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFAE125428EEF88985291869" box="[151,277,1829,1853]" italics="true" pageId="16" pageNumber="374">L. acutidens</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFAE1254295CF88A859C1869" author="Chen, H." box="[293,416,1829,1853]" pageId="16" pageNumber="374" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D,
|
||||
<taxonomicName id="4C744D4FFFAE12542B87F88980581869" authorityName="Rahayu & Ng" authorityYear="2024" box="[1022,1124,1829,1853]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="374" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAE12542B87F88980581869" box="[1022,1124,1829,1853]" italics="true" pageId="16" pageNumber="374">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAE12542C15F88A80941869" bold="true" box="[1132,1192,1829,1853]" pageId="16" pageNumber="374">
|
||||
<taxonomicNameLabel id="A23357A5FFAE12542C15F88A80941869" box="[1132,1192,1829,1853]" pageId="16" pageNumber="374" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E,
|
||||
<taxonomicName id="4C744D4FFFAE12542A32F8E586871835" authorityName="Rahayu & Ng" authorityYear="2024" box="[587,699,1865,1889]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="374" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAE12542A32F8E586871835" box="[587,699,1865,1889]" italics="true" pageId="16" pageNumber="374">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAE12542ABBF8E686C01835" bold="true" box="[706,764,1865,1889]" pageId="16" pageNumber="374">
|
||||
<taxonomicNameLabel id="A23357A5FFAE12542ABBF8E686C01835" box="[706,764,1865,1889]" pageId="16" pageNumber="374" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F,
|
||||
<taxonomicName id="4C744D4FFFAE12542DF1F8E584F718D1" authorityName="Rahayu & Ng" authorityYear="2024" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="374" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAE12542DF1F8E584F718D1" italics="true" pageId="16" pageNumber="374">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAE125428AAF8C2853118D1" bold="true" box="[211,269,1901,1925]" pageId="16" pageNumber="374">
|
||||
<taxonomicNameLabel id="A23357A5FFAE125428AAF8C2853118D1" box="[211,269,1901,1925]" pageId="16" pageNumber="374" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G,
|
||||
<taxonomicName id="4C744D4FFFAE12542BB5F8C1802018D1" authorityName="Rahayu & Ng" authorityYear="2024" box="[972,1052,1901,1925]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="374" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAE12542BB5F8C1802018D1" box="[972,1052,1901,1925]" italics="true" pageId="16" pageNumber="374">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAE12542C5AF8C2806218D1" bold="true" box="[1059,1118,1901,1925]" pageId="16" pageNumber="374">
|
||||
<taxonomicNameLabel id="A23357A5FFAE12542C5AF8C2806218D1" box="[1059,1118,1901,1925]" pageId="16" pageNumber="374" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H,
|
||||
<taxonomicName id="4C744D4FFFAE125429D3F83D85C518FD" authorityName="Rahayu & Ng" authorityYear="2024" box="[426,505,1937,1961]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="374" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAE125429D3F83D85C518FD" box="[426,505,1937,1961]" italics="true" pageId="16" pageNumber="374">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAE12542986F83E860418FD" bold="true" box="[511,568,1937,1961]" pageId="16" pageNumber="374">
|
||||
<taxonomicNameLabel id="A23357A5FFAE12542986F83E860418FD" box="[511,568,1937,1961]" pageId="16" pageNumber="374" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I,
|
||||
<taxonomicName id="4C744D4FFFAE12542C17F83D80C418FD" authorityName="Rahayu & Ng" authorityYear="2024" box="[1134,1272,1937,1961]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="374" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAE12542C17F83D80C418FD" box="[1134,1272,1937,1961]" italics="true" pageId="16" pageNumber="374">L. bellagrana</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAE12542C87F83E810B18FD" bold="true" box="[1278,1335,1937,1961]" pageId="16" pageNumber="374">
|
||||
<taxonomicNameLabel id="A23357A5FFAE12542C87F83E810B18FD" box="[1278,1335,1937,1961]" pageId="16" pageNumber="374" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J,
|
||||
<taxonomicName id="4C744D4FFFAE12542B61F819879E1899" authorityName="Rahayu & Ng" authorityYear="2024" box="[792,930,1973,1997]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="374" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAE12542B61F819879E1899" box="[792,930,1973,1997]" italics="true" pageId="16" pageNumber="374">L. bellagrana</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAE12542BD0F81A87DF1899" bold="true" box="[937,995,1973,1997]" pageId="16" pageNumber="374">
|
||||
<taxonomicNameLabel id="A23357A5FFAE12542BD0F81A87DF1899" box="[937,995,1973,1997]" pageId="16" pageNumber="374" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF0B6644FFAF125528EEF9F3816B19B5" ID-DOI="http://doi.org/10.5281/zenodo.12681950" ID-Zenodo-Dep="12681950" httpUri="https://zenodo.org/record/12681950/files/figure.png" pageId="17" pageNumber="375" startId="17.[151,250,1628,1652]" targetBox="[151,1435,181,1605]" targetPageId="17" targetType="figure">
|
||||
<paragraph id="8BCB36CCFFAF125528EEF9F3816B19B5" blockId="17.[151,1437,1628,1761]" pageId="17" pageNumber="375">
|
||||
<emphasis id="B900EADEFFAF125528EEF9F3851C1921" bold="true" box="[151,288,1628,1653]" pageId="17" pageNumber="375">FIGURE 12.</emphasis>
|
||||
Third maxillipeds. A,
|
||||
<taxonomicName id="4C744D4FFFAF12552A7CF9F286981921" authorityName="Rahayu & Ng" authorityYear="2024" box="[517,676,1629,1653]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="chomel" status="sp. nov.">
|
||||
<emphasis id="B900EADEFFAF12552A7CF9F286981921" box="[517,676,1629,1653]" italics="true" pageId="17" pageNumber="375">Lyphira chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAF12552AD3F9F286DF1921" bold="true" box="[682,739,1629,1653]" pageId="17" pageNumber="375">
|
||||
<taxonomicNameLabel id="A23357A5FFAF12552AD3F9F286DF1921" box="[682,739,1629,1653]" pageId="17" pageNumber="375" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; B,
|
||||
<taxonomicName id="4C744D4FFFAF12552DF1F9F284DC19CD" authorityName="Rahayu & Ng" authorityYear="2024" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAF12552DF1F9F284DC19CD" italics="true" pageId="17" pageNumber="375">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAF1255289FF92E852219CD" bold="true" box="[230,286,1665,1689]" pageId="17" pageNumber="375">
|
||||
<taxonomicNameLabel id="A23357A5FFAF1255289FF92E852219CD" box="[230,286,1665,1689]" pageId="17" pageNumber="375" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; C,
|
||||
<taxonomicName id="4C744D4FFFAF12552BD7F92E802D19CD" authorityName="Rahayu & Ng" authorityYear="2024" box="[942,1041,1665,1689]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAF12552BD7F92E802D19CD" box="[942,1041,1665,1689]" italics="true" pageId="17" pageNumber="375">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAF12552C6FF92E807219CD" bold="true" box="[1046,1102,1665,1689]" pageId="17" pageNumber="375">
|
||||
<taxonomicNameLabel id="A23357A5FFAF12552C6FF92E807219CD" box="[1046,1102,1665,1689]" pageId="17" pageNumber="375" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore; D,
|
||||
<taxonomicName id="4C744D4FFFAF12552989F90A867C19E9" authorityName="Rahayu & Ng" authorityYear="2024" box="[496,576,1701,1725]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAF12552989F90A867C19E9" box="[496,576,1701,1725]" italics="true" pageId="17" pageNumber="375">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAF12552A3EF90A86BE19E9" bold="true" box="[583,642,1701,1725]" pageId="17" pageNumber="375">
|
||||
<taxonomicNameLabel id="A23357A5FFAF12552A3EF90A86BE19E9" box="[583,642,1701,1725]" pageId="17" pageNumber="375" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (12.6 × 12.7 mm) (ZRC 1985.97), Malaysia; E,
|
||||
<taxonomicName id="4C744D4FFFAF12552CC1F90A813419E9" authorityName="Rahayu & Ng" authorityYear="2024" box="[1208,1288,1701,1725]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAF12552CC1F90A813419E9" box="[1208,1288,1701,1725]" italics="true" pageId="17" pageNumber="375">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAF12552D76F90A817619E9" bold="true" box="[1295,1354,1701,1725]" pageId="17" pageNumber="375">
|
||||
<taxonomicNameLabel id="A23357A5FFAF12552D76F90A817619E9" box="[1295,1354,1701,1725]" pageId="17" pageNumber="375" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, female (17.0 × 16.9 mm) (ZRC 1995.0547), Singapore; F,
|
||||
<taxonomicName id="4C744D4FFFAF12552AE4F96686D019B5" authorityName="Rahayu & Ng" authorityYear="2024" box="[669,748,1737,1761]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAF12552AE4F96686D019B5" box="[669,748,1737,1761]" italics="true" pageId="17" pageNumber="375">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAF12552A8AF966871019B5" bold="true" box="[755,812,1737,1761]" pageId="17" pageNumber="375">
|
||||
<taxonomicNameLabel id="A23357A5FFAF12552A8AF966871019B5" box="[755,812,1737,1761]" pageId="17" pageNumber="375" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (12.1 × 12.4 mm) (ZRC 1995.0547), Singapore.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C36E6547FFAF125628BEF8A086441DD1" lastPageId="18" lastPageNumber="376" pageId="17" pageNumber="375" type="discussion">
|
||||
<paragraph id="8BCB36CCFFAF125628BEF8A087311E15" blockId="17.[151,1437,1807,2013]" lastBlockId="18.[151,1437,151,717]" lastPageId="18" lastPageNumber="376" pageId="17" pageNumber="375">
|
||||
<emphasis id="B900EADEFFAF125528BEF8A08508187D" bold="true" box="[199,308,1807,1833]" pageId="17" pageNumber="375">Remarks</emphasis>
|
||||
.
|
||||
<taxonomicName id="4C744D4FFFAF1255293BF8A085CF187D" authorityName="Rahayu & Ng" authorityYear="2024" box="[322,499,1807,1833]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="chomel" status="sp. nov.">
|
||||
<emphasis id="B900EADEFFAF1255293BF8A085CF187D" box="[322,499,1807,1833]" italics="true" pageId="17" pageNumber="375">Lyphira chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAF12552983F8A08605187D" bold="true" box="[506,569,1807,1833]" pageId="17" pageNumber="375">
|
||||
<taxonomicNameLabel id="A23357A5FFAF12552983F8A08605187D" box="[506,569,1807,1833]" pageId="17" pageNumber="375" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is a very distinct species. Its short and poorly developed endostomial spines, and the relatively narrow third maxilliped exopod allies it with
|
||||
<taxonomicName id="4C744D4FFFAF12552B2DF89B87C61819" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[852,1018,1843,1869]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFAF12552B2DF89B87C61819" box="[852,1018,1843,1869]" italics="true" pageId="17" pageNumber="375">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23357A5FFAF12552C78F89C80071819" box="[1025,1083,1843,1869]" pageId="17" pageNumber="375" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAF12552C16F89B80CB1819" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[1135,1271,1843,1869]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFAF12552C16F89B80CB1819" box="[1135,1271,1843,1869]" italics="true" pageId="17" pageNumber="375">L. acutidens</emphasis>
|
||||
</taxonomicName>
|
||||
, but
|
||||
<taxonomicName id="4C744D4FFFAF12552D56F89B81A01819" authorityName="Rahayu & Ng" authorityYear="2024" box="[1327,1436,1843,1869]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAF12552D56F89B81A01819" box="[1327,1436,1843,1869]" italics="true" pageId="17" pageNumber="375">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAF125528EEF8F884EA1825" bold="true" box="[151,214,1879,1905]" pageId="17" pageNumber="375">
|
||||
<taxonomicNameLabel id="A23357A5FFAF125528EEF8F884EA1825" box="[151,214,1879,1905]" pageId="17" pageNumber="375" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
can immediately be distinguished by its prominently slenderer and elongate ambulatory meri (
|
||||
<figureCitation id="134F2A49FFAF12552C86F8F881591825" box="[1279,1381,1879,1905]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="17" pageNumber="375">Fig. 16D</figureCitation>
|
||||
) and the granules on the merus of the cheliped being evenly spread out, reaching to the distal surfaces (
|
||||
<figureCitation id="134F2A49FFAF12552C81F8D4816D18C1" box="[1272,1361,1915,1941]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="17" pageNumber="375">Fig. 3E</figureCitation>
|
||||
). In
|
||||
<taxonomicName id="4C744D4FFFAF12552DFFF8D3851C18ED" baseAuthorityName="Ortmann" baseAuthorityYear="1892" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFAF12552DFFF8D3851C18ED" italics="true" pageId="17" pageNumber="375">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23357A5FFAF12552953F830855B18ED" box="[298,359,1951,1977]" pageId="17" pageNumber="375" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAF125529D8F80F861718ED" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[417,555,1951,1977]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="375" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFAF125529D8F80F861718ED" box="[417,555,1951,1977]" italics="true" pageId="17" pageNumber="375">L. acutidens</emphasis>
|
||||
</taxonomicName>
|
||||
, the ambulatory meri are distinctly shorter and stouter (
|
||||
<figureCitation id="134F2A49FFAF12552CD6F830810418ED" box="[1199,1336,1951,1977]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="17" pageNumber="375">Fig. 16A‒C</figureCitation>
|
||||
) and the distal surfaces of the merus of the cheliped are relatively smoother with only scattered granules present (
|
||||
<figureCitation id="134F2A49FFAF12552D6DF86C81AD1889" box="[1300,1425,1987,2013]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="17" pageNumber="375">Fig. 3A‒D</figureCitation>
|
||||
). The G1s of
|
||||
<taxonomicName id="4C744D4FFFAC12562961FF3885B91FE5" authorityName="Rahayu & Ng" authorityYear="2024" box="[280,389,151,177]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAC12562961FF3885B91FE5" box="[280,389,151,177]" italics="true" pageId="18" pageNumber="376">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAC125629F2FF3885F41FE5" bold="true" box="[395,456,151,177]" pageId="18" pageNumber="376">
|
||||
<taxonomicNameLabel id="A23357A5FFAC125629F2FF3885F41FE5" box="[395,456,151,177]" pageId="18" pageNumber="376" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAC12562985FF38869D1FE5" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[508,673,151,177]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFAC12562985FF38869D1FE5" box="[508,673,151,177]" italics="true" pageId="18" pageNumber="376">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23357A5FFAC12562ADEFF3886DF1FE5" box="[679,739,151,177]" pageId="18" pageNumber="376" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
, however, are similar, with the apical process longer and gently curved (
|
||||
<figureCitation id="134F2A49FFAC1256288AFF1485871F81" box="[243,443,187,213]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="18" pageNumber="376">Fig. 17C‒F, O‒R</figureCitation>
|
||||
). On the basis of published data,
|
||||
<taxonomicName id="4C744D4FFFAC12562B4BFF14879D1F81" authorityName="Rahayu & Ng" authorityYear="2024" box="[818,929,187,213]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAC12562B4BFF14879D1F81" box="[818,929,187,213]" italics="true" pageId="18" pageNumber="376">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAC12562BD0FF1487D41F81" bold="true" box="[937,1000,187,213]" pageId="18" pageNumber="376">
|
||||
<taxonomicNameLabel id="A23357A5FFAC12562BD0FF1487D41F81" box="[937,1000,187,213]" pageId="18" pageNumber="376" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is also the smallest species, with male specimens as small as
|
||||
<quantity id="4C8C9B29FFAC125629E2FF7085D01FAD" box="[411,492,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="376" unit="mm" value="10.0">10 mm</quantity>
|
||||
in carapace width already mature and females of a similar size already ovigerous.
|
||||
<taxonomicName id="4C744D4FFFAC125628EEFEAC85421E49" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,382,259,285]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" sensu="stricto" species="heterograna">
|
||||
<emphasis id="B900EADEFFAC125628EEFEAC85421E49" box="[151,382,259,285]" italics="true" pageId="18" pageNumber="376">Lyphira heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23357A5FFAC125629FCFEAC85831E49" box="[389,447,259,285]" pageId="18" pageNumber="376" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C744D4FFFAC1256298DFEAC86401E49" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[500,636,259,285]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFAC1256298DFEAC86401E49" box="[500,636,259,285]" italics="true" pageId="18" pageNumber="376">L. acutidens</emphasis>
|
||||
</taxonomicName>
|
||||
both reach larger sizes (smallest adult female averages
|
||||
<quantity id="4C8C9B29FFAC12562C9FFEAC81081E49" box="[1254,1332,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="376" unit="mm" value="12.0">12 mm</quantity>
|
||||
carapace width) (cf.
|
||||
<bibRefCitation id="EFE54B3DFFAC1256296DFE8885AD1E15" author="Chen, H." box="[276,401,295,321]" pageId="18" pageNumber="376" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen 1987</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFE54B3DFFAC125629E4FE8886571E15" author="Chen, H. & Sun, H." box="[413,619,295,321]" pageId="18" pageNumber="376" refId="ref19489" refString="Chen, H. & Sun, H. (2002) Brachyura marine primitive crabs. Fauna Sinica Invertebrata Arthropoda Crustacea. Vol. 30. Science Press, Beijing, xiii + 597 pp., 237 figs., 6 pls." type="book" year="2002">Chen & Sun 2002</bibRefCitation>
|
||||
; present data).
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFAC125628BEFEE486621D4D" blockId="18.[151,1437,151,717]" pageId="18" pageNumber="376">
|
||||
<bibRefCitation id="EFE54B3DFFAC125628BEFEE485A01E31" author="Galil, B. S." box="[199,412,331,357]" pageId="18" pageNumber="376" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil’s (2009: 300)</bibRefCitation>
|
||||
stated she examined several specimens of “
|
||||
<taxonomicName id="4C744D4FFFAC12562BFCFEE480171E31" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[901,1067,331,357]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFAC12562BFCFEE480171E31" box="[901,1067,331,357]" italics="true" pageId="18" pageNumber="376">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
” from Peninsular
|
||||
<collectingCountry id="F363765CFFAC12562C80FEE481581E31" box="[1273,1380,331,357]" name="Malaysia" pageId="18" pageNumber="376">Malaysia</collectingCountry>
|
||||
:
|
||||
<specimenCount id="9D72FD45FFAC12562D17FEE484E41EDD" count="4" pageId="18" pageNumber="376" type="male">four males</specimenCount>
|
||||
and
|
||||
<specimenCount id="9D72FD45FFAC12562968FEC085A61EDD" box="[273,410,367,393]" count="2" pageId="18" pageNumber="376" type="female">two females</specimenCount>
|
||||
from
|
||||
<collectingCountry id="F363765CFFAC12562999FEC086031EDD" box="[480,575,367,393]" name="Malaysia" pageId="18" pageNumber="376">Malacca</collectingCountry>
|
||||
(NHM 1900.10.22.340–345) and
|
||||
<specimenCount id="9D72FD45FFAC12562BBBFEC080151EDD" box="[962,1065,367,393]" count="1" pageId="18" pageNumber="376" type="male">one male</specimenCount>
|
||||
from Pontian (ZRC 1999.0798).
|
||||
<bibRefCitation id="EFE54B3DFFAC125628EEFE3C851D1EF9" author="Galil, B. S." box="[151,289,403,429]" pageId="18" pageNumber="376" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil (2009</bibRefCitation>
|
||||
: figs. 12B, 14) figured a male specimen (
|
||||
<quantity id="4C8C9B29FFAC12562B67FE3D87B51EF9" box="[798,905,402,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="18" pageNumber="376" unit="mm" value="15.1">15.1 mm</quantity>
|
||||
carapace length, NHM 1900.10.22.340–345) (incorrectly stated as from
|
||||
<collectingCountry id="F363765CFFAC125629AAFE1886791E85" box="[467,581,439,465]" name="Indonesia" pageId="18" pageNumber="376">Indonesia</collectingCountry>
|
||||
). Her figures agree very well with the
|
||||
<typeStatus id="54CF886EFFAC12562C77FE18804C1E85" box="[1038,1136,439,465]" pageId="18" pageNumber="376" type="holotype">holotype</typeStatus>
|
||||
male of
|
||||
<taxonomicName id="4C744D4FFFAC12562CA4FE1881731E85" authorityName="Rahayu & Ng" authorityYear="2024" box="[1245,1359,439,465]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAC12562CA4FE1881731E85" box="[1245,1359,439,465]" italics="true" pageId="18" pageNumber="376">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAC12562D23FE1881A01E85" bold="true" box="[1370,1436,439,465]" pageId="18" pageNumber="376">
|
||||
<taxonomicNameLabel id="A23357A5FFAC12562D23FE1881A01E85" box="[1370,1436,439,465]" pageId="18" pageNumber="376" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(which is here based on her Pontian male), including the pattern of granules on the merus of the cheliped, relatively slenderer P2–P5 and structure of the G1.
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFAC125628BEFD8C86441DD1" blockId="18.[151,1437,151,717]" pageId="18" pageNumber="376">
|
||||
In
|
||||
<collectingCountry id="F363765CFFAC12562890FD8C85611D69" box="[233,349,547,573]" name="Singapore" pageId="18" pageNumber="376">Singapore</collectingCountry>
|
||||
,
|
||||
<taxonomicName id="4C744D4FFFAC12562915FD8C85E21D69" authorityName="Rahayu & Ng" authorityYear="2024" box="[364,478,547,573]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAC12562915FD8C85E21D69" box="[364,478,547,573]" italics="true" pageId="18" pageNumber="376">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAC12562991FD8C86161D69" bold="true" box="[488,554,547,573]" pageId="18" pageNumber="376">
|
||||
<taxonomicNameLabel id="A23357A5FFAC12562991FD8C86161D69" box="[488,554,547,573]" pageId="18" pageNumber="376" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is sympatric with
|
||||
<taxonomicName id="4C744D4FFFAC12562B7EFD8C875E1D69" authorityName="Rahayu & Ng" authorityYear="2024" box="[775,866,547,573]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAC12562B7EFD8C875E1D69" box="[775,866,547,573]" italics="true" pageId="18" pageNumber="376">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAC12562B15FD8C87931D69" bold="true" box="[876,943,547,573]" pageId="18" pageNumber="376">
|
||||
<taxonomicNameLabel id="A23357A5FFAC12562B15FD8C87931D69" box="[876,943,547,573]" pageId="18" pageNumber="376" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, but they appear to have slightly different habitats. Although both are intertidal to subtidal,
|
||||
<taxonomicName id="4C744D4FFFAC12562AC1FDE8871A1D35" authorityName="Rahayu & Ng" authorityYear="2024" box="[696,806,583,609]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAC12562AC1FDE8871A1D35" box="[696,806,583,609]" italics="true" pageId="18" pageNumber="376">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAC12562B55FDE887561D35" bold="true" box="[812,874,583,609]" pageId="18" pageNumber="376">
|
||||
<taxonomicNameLabel id="A23357A5FFAC12562B55FDE887561D35" box="[812,874,583,609]" pageId="18" pageNumber="376" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
appears to prefer more muddy substrates, whereas
|
||||
<taxonomicName id="4C744D4FFFAC125628EEFDC484D31DD1" authorityName="Rahayu & Ng" authorityYear="2024" box="[151,239,619,645]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
|
||||
<emphasis id="B900EADEFFAC125628EEFDC484D31DD1" box="[151,239,619,645]" italics="true" pageId="18" pageNumber="376">L. linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFAC1256288CFDC485081DD1" bold="true" box="[245,308,619,645]" pageId="18" pageNumber="376">
|
||||
<taxonomicNameLabel id="A23357A5FFAC1256288CFDC485081DD1" box="[245,308,619,645]" pageId="18" pageNumber="376" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
likes area which are sandier.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFAC125628BEFD2086811DFD" box="[199,701,654,681]" pageId="18" pageNumber="376" type="materials_examined">
|
||||
<paragraph id="8BCB36CCFFAC125628BEFD2086811DFD" blockId="18.[151,1437,151,717]" box="[199,701,654,681]" pageId="18" pageNumber="376">
|
||||
<materialsCitation id="3B1C3C91FFAC125628BEFD2086851DFD" box="[199,697,654,681]" country="Malaysia" location="Pontian" municipality="Peninsular" pageId="18" pageNumber="376" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="B900EADEFFAC125628BEFD20855C1DFC" bold="true" box="[199,352,654,681]" pageId="18" pageNumber="376">
|
||||
<typeStatus id="54CF886EFFAC125628BEFD20853E1DFD" box="[199,258,655,681]" pageId="18" pageNumber="376">Type</typeStatus>
|
||||
locality
|
||||
</emphasis>
|
||||
.
|
||||
<location id="8EAB6017FFAC12562917FD2085FA1DFD" LSID="urn:lsid:plazi:treatment:03DD87DAFFB4125628EEFDE9875B1D99:8EAB6017FFAC12562917FD2085FA1DFD" box="[366,454,655,681]" country="Malaysia" municipality="Peninsular" name="Pontian" pageId="18" pageNumber="376">Pontian</location>
|
||||
,
|
||||
<collectingMunicipality id="6BAFACB6FFAC125629A8FD20867B1DFD" box="[465,583,655,681]" pageId="18" pageNumber="376">Peninsular</collectingMunicipality>
|
||||
<collectingCountry id="F363765CFFAC12562A36FD2086851DFD" box="[591,697,655,681]" name="Malaysia" pageId="18" pageNumber="376">Malaysia</collectingCountry>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFAC125628BEFD1D875B1D99" box="[199,871,690,717]" pageId="18" pageNumber="376" type="distribution">
|
||||
<paragraph id="8BCB36CCFFAC125628BEFD1D875B1D99" blockId="18.[151,1437,151,717]" box="[199,871,690,717]" pageId="18" pageNumber="376">
|
||||
<emphasis id="B900EADEFFAC125628BEFD1D85651D98" bold="true" box="[199,345,690,716]" pageId="18" pageNumber="376">Distribution</emphasis>
|
||||
. Southern Peninsular
|
||||
<collectingCountry id="F363765CFFAC12562A28FD1C86841D99" box="[593,696,691,717]" name="Malaysia" pageId="18" pageNumber="376">Malaysia</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="F363765CFFAC12562A97FD1C875F1D99" box="[750,867,691,717]" name="Singapore" pageId="18" pageNumber="376">Singapore</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
469
data/03/DD/87/03DD87DAFFB8124E28EEFBBC87BE1EA1.xml
Normal file
469
data/03/DD/87/03DD87DAFFB8124E28EEFBBC87BE1EA1.xml
Normal file
|
|
@ -0,0 +1,469 @@
|
|||
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<mods:title id="4710E450AFE0E9CDC454A646FAE27824">On the genus Lyphira Galil, 2009 (Crustacea: Brachyura: Leucosiidae), with descriptions of four new species from the West Pacific</mods:title>
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<taxonomicName id="4C744D4FFFB8124228EEFBBC86241B7A" ID-CoL="4GDTY" authority="(Chen, 1987)" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[151,536,1043,1070]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="acutidens">
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<emphasis id="B900EADEFFB8124228EEFBBC854E1B7A" bold="true" box="[151,370,1043,1070]" italics="true" pageId="6" pageNumber="364">Lyphira acutidens</emphasis>
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(
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<bibRefCitation id="EFE54B3DFFB8124229F8FBBB862C1B7A" author="Chen, H." box="[385,528,1044,1070]" pageId="6" pageNumber="364" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987</bibRefCitation>
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<paragraph id="8BCB36CCFFB8124228EEFB9787831B06" blockId="6.[151,959,1043,1107]" box="[151,959,1080,1107]" pageId="6" pageNumber="364">
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(
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<figureCitation id="134F2A49FFB8124228E6FB97853F1B06" box="[159,259,1080,1106]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Figs. 3D</figureCitation>
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<figureCitation id="134F2A49FFB812422976FB97850E1B06" box="[271,306,1080,1106]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="6" pageNumber="364">6B</figureCitation>
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<figureCitation id="134F2A49FFB812422947FB96855E1B07" box="[318,354,1081,1107]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="6" pageNumber="364">7D</figureCitation>
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<figureCitation id="134F2A49FFB812422917FB9785AD1B06" box="[366,401,1080,1106]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="6" pageNumber="364">9B</figureCitation>
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<figureCitation id="134F2A49FFB8124229E4FB9785F11B06" box="[413,461,1080,1106]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="6" pageNumber="364">10C</figureCitation>
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<figureCitation id="134F2A49FFB8124229A0FB9786351B06" box="[473,521,1080,1106]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="6" pageNumber="364">11C</figureCitation>
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<figureCitation id="134F2A49FFB812422A6CFB97865B1B06" box="[533,615,1080,1106]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="6" pageNumber="364">13B, C</figureCitation>
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<figureCitation id="134F2A49FFB812422A0BFB9786981B06" box="[626,676,1080,1106]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="6" pageNumber="364">14B</figureCitation>
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<figureCitation id="134F2A49FFB812422AD6FB9786DE1B07" box="[687,738,1080,1107]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="6" pageNumber="364">15D</figureCitation>
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|
||||
<figureCitation id="134F2A49FFB812422A94FB9787221B06" box="[749,798,1080,1106]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="6" pageNumber="364">16C</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB812422B53FB9787471B06" box="[810,891,1080,1106]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="6" pageNumber="364">17H–L</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB812422BFEFB97878B1B06" box="[903,951,1080,1106]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="6" pageNumber="364">21B</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFB8124228EEFBD880181BFA" pageId="6" pageNumber="364" type="reference_group">
|
||||
<paragraph id="8BCB36CCFFB8124228EEFBD880E31BDB" blockId="6.[151,1247,1143,1198]" box="[151,1247,1143,1168]" pageId="6" pageNumber="364">
|
||||
<treatmentCitationGroup id="AB6411E2FFB8124228EEFBD880E31BDB" box="[151,1247,1143,1168]" pageId="6" pageNumber="364">
|
||||
<treatmentCitation id="0AD510DDFFB8124228EEFBD885C71BDB" author="Chen, H." box="[151,507,1143,1167]" page="201" pageId="6" pageNumber="364" year="1987">
|
||||
<taxonomicName id="4C744D4FFFB8124228EEFBD885C71BDB" authority="Chen, 1987: 201" authorityName="Chen" authorityPageNumber="201" authorityYear="1987" box="[151,507,1143,1167]" class="Malacostraca" family="Leucosiidae" genus="Phylira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB8124228EEFBD885771BDB" box="[151,331,1143,1167]" italics="true" pageId="6" pageNumber="364">Phylira acutidens</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFB812422928FBD885C71BDB" author="Chen, H." box="[337,507,1143,1167]" pageId="6" pageNumber="364" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987: 201</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
, fig. 1;
|
||||
<treatmentCitation id="0AD510DDFFB812422A30FBD887081BDB" author="Chen, H. & Sun, H." box="[585,820,1143,1167]" page="381" pageId="6" pageNumber="364" year="2002">
|
||||
<bibRefCitation id="EFE54B3DFFB812422A30FBD887081BDB" author="Chen, H. & Sun, H." box="[585,820,1143,1167]" pageId="6" pageNumber="364" refId="ref19489" refString="Chen, H. & Sun, H. (2002) Brachyura marine primitive crabs. Fauna Sinica Invertebrata Arthropoda Crustacea. Vol. 30. Science Press, Beijing, xiii + 597 pp., 237 figs., 6 pls." type="book" year="2002">Chen & Sun 2002: 381</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
, text fig. 170, pl. 15.1;
|
||||
<treatmentCitation id="0AD510DDFFB812422C58FBD780E71BDB" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[1057,1243,1143,1168]" page="93" pageId="6" pageNumber="364" year="2008">
|
||||
<bibRefCitation id="EFE54B3DFFB812422C58FBD780E71BDB" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[1057,1243,1143,1168]" pageId="6" pageNumber="364" pagination="1 - 286" refId="ref21130" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
|
||||
Ng
|
||||
<emphasis id="B900EADEFFB812422C3FFBD780451BDB" box="[1094,1145,1143,1167]" italics="true" pageId="6" pageNumber="364">et al.</emphasis>
|
||||
2008: 93
|
||||
</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFB8124228EEFB3980181BFA" blockId="6.[151,1247,1143,1198]" box="[151,1060,1174,1198]" pageId="6" pageNumber="364">
|
||||
<treatmentCitationGroup id="AB6411E2FFB8124228EEFB3980181BFA" box="[151,1060,1174,1198]" pageId="6" pageNumber="364">
|
||||
<emphasis id="B900EADEFFB8124228EEFB3985BA1BFA" box="[151,390,1174,1198]" italics="true" pageId="6" pageNumber="364">
|
||||
<taxonomicName id="4C744D4FFFB8124228EEFB3985541BFA" ID-CoL="3WTBK" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,360,1174,1198]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="heterograna">Lyphira heterograna</taxonomicName>
|
||||
.—
|
||||
</emphasis>
|
||||
<treatmentCitation id="0AD510DDFFB8124229FFFB3986101BFA" author="Galil, B. S." box="[390,556,1174,1198]" httpUri="http://treatment.plazi.org/id/039D87BCFFBF4A26FD4BC4ADFDE1ECDD" page="300" pageId="6" pageNumber="364" year="2009">
|
||||
<bibRefCitation id="EFE54B3DFFB8124229FFFB3986101BFA" author="Galil, B. S." box="[390,556,1174,1198]" pageId="6" pageNumber="364" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil, 2009: 300</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
(part). (not
|
||||
<taxonomicName id="4C744D4FFFB812422ADFFB3980251BFA" ID-CoL="76YMK" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[678,1049,1174,1198]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB812422ADFFB3987481BFA" box="[678,884,1174,1198]" italics="true" pageId="6" pageNumber="364">Philyra heterograna</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFB812422B02FB3980251BFA" author="Ortmann, A." box="[891,1049,1174,1198]" pageId="6" pageNumber="364" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
</taxonomicName>
|
||||
).
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFB8124228EEFB79801F1A41" pageId="6" pageNumber="364" type="materials_examined">
|
||||
<paragraph id="8BCB36CCFFB8124228EEFB79801F1A41" blockId="6.[151,1437,1238,2021]" pageId="6" pageNumber="364">
|
||||
<emphasis id="B900EADEFFB8124228EEFB7985451BA4" bold="true" box="[151,377,1238,1264]" pageId="6" pageNumber="364">Material examined</emphasis>
|
||||
.
|
||||
<specimenCount id="9D72FD45FFB8124229FFFB7885EC1BA5" box="[390,464,1239,1265]" count="1" pageId="6" pageNumber="364" type="male">1 male</specimenCount>
|
||||
(8.9 ×
|
||||
<quantity id="4C8C9B29FFB812422A66FB7886441BA5" box="[543,632,1239,1265]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="6" pageNumber="364" unit="mm" value="8.6">8.6 mm</quantity>
|
||||
),
|
||||
<materialsCitation id="3B1C3C91FFB812422AF2FB7880231A41" collectingDate="1958-12-09" collectionCode="ZRC" location="Institute of Oceanography" municipality="Chinese Academy of Science" pageId="6" pageNumber="364" specimenCode="ZRC 2002.0504" specimenCount="1" specimenCount-female="1">
|
||||
<specimenCount id="9D72FD45FFB812422AF2FB7886D71BA5" box="[651,747,1239,1265]" count="1" pageId="6" pageNumber="364" type="female">1 female</specimenCount>
|
||||
(13.4 ×
|
||||
<quantity id="4C8C9B29FFB812422B31FB7887931BA5" box="[840,943,1239,1265]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="6" pageNumber="364" unit="mm" value="12.9">12.9 mm</quantity>
|
||||
),
|
||||
<specimenCode id="DBD29EB7FFB812422BB8FB7880441BA5" box="[961,1144,1238,1265]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="364">ZRC 2002.0504</specimenCode>
|
||||
, station 4027, Yellow Sea, coll.
|
||||
<location id="8EAB6017FFB8124228B6FB5485C81A41" LSID="urn:lsid:plazi:treatment:03DD87DAFFB8124E28EEFBBC87BE1EA1:8EAB6017FFB8124228B6FB5485C81A41" box="[207,500,1275,1301]" municipality="Chinese Academy of Science" name="Institute of Oceanography" pageId="6" pageNumber="364">Institute of Oceanography</location>
|
||||
,
|
||||
<collectingMunicipality id="6BAFACB6FFB812422A79FB5487761A41" box="[512,842,1275,1301]" pageId="6" pageNumber="364">Chinese Academy of Science</collectingMunicipality>
|
||||
,
|
||||
<date id="FFCA100CFFB812422B2CFB5480231A41" box="[853,1055,1274,1301]" pageId="6" pageNumber="364" value="1958-12-09">
|
||||
<collectingDate id="EF8EE9E4FFB812422B2CFB5480231A41" box="[853,1055,1274,1301]" pageId="6" pageNumber="364" value="1958-12-09">9 December 1958</collectingDate>
|
||||
</date>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFB8124328BEFAB185921FAD" lastPageId="7" lastPageNumber="365" pageId="6" pageNumber="364" type="diagnosis">
|
||||
<paragraph id="8BCB36CCFFB8124328BEFAB186591F81" blockId="6.[151,1437,1238,2021]" lastBlockId="7.[151,1436,151,249]" lastPageId="7" lastPageNumber="365" pageId="6" pageNumber="364">
|
||||
<emphasis id="B900EADEFFB8124228BEFAB185061A6C" bold="true" box="[199,314,1310,1336]" pageId="6" pageNumber="364">Diagnosis</emphasis>
|
||||
. Carapace slightly broader than long, dorsal surface of carapace covered with closely spaced minute granules, hepatic and intestinal regions with larger, more prominent granules (
|
||||
<figureCitation id="134F2A49FFB812422C41FAEC809D1A09" box="[1080,1185,1347,1373]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Figs. 3D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB812422CCAFAEC80EB1A09" box="[1203,1239,1347,1373]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="6" pageNumber="364">6B</figureCitation>
|
||||
). Frontal margin granulated, postfrontal lobe slightly convex (
|
||||
<figureCitation id="134F2A49FFB812422AE3FAC887371AD5" box="[666,779,1383,1409]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="6" pageNumber="364">Figs. 11C</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB812422B63FAC887531AD5" box="[794,879,1383,1409]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="6" pageNumber="364">13B, C</figureCitation>
|
||||
). Endostomial spine absent (
|
||||
<figureCitation id="134F2A49FFB812422CC0FAC881731AD5" box="[1209,1359,1383,1409]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="6" pageNumber="364">Figs. 13B, C</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB812422D24FAC881B31AD5" box="[1373,1423,1383,1409]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="6" pageNumber="364">14B</figureCitation>
|
||||
). Lateral and posterolateral margin of carapace with row of small granules interspersed by several larger granules (
|
||||
<figureCitation id="134F2A49FFB8124228E6FA0084C71A9D" box="[159,251,1455,1481]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Fig. 3D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB812422970FA0085111A9D" box="[265,301,1455,1481]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="6" pageNumber="364">6B</figureCitation>
|
||||
); posterior carapace margin gently concave with row of larger granule, medially slightly produced (
|
||||
<figureCitation id="134F2A49FFB8124228E6FA7C85391AB9" box="[159,261,1491,1517]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Figs. 3D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB81242296BFA7C85791AB9" box="[274,325,1491,1517]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="6" pageNumber="364">10C</figureCitation>
|
||||
). Third maxilliped exopod 2.9 times as long as broad, as broad as basal part of endopod, forming petaliform structure with convex outer margin, endopod with completely fused basis-ischium, as long as merus, with shallow submarginal sulcus near inner margin (
|
||||
<figureCitation id="134F2A49FFB812422A9AF9B4876E1961" box="[739,850,1563,1589]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="6" pageNumber="364">Figs. 11C</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB812422B27F9B487AD1961" box="[862,913,1563,1589]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="6" pageNumber="364">17H</figureCitation>
|
||||
). Chelipeds subequal; merus, carpus and chela surfaces smooth or covered by closely spaced, small rounded granules (
|
||||
<figureCitation id="134F2A49FFB812422BB1F9908011190D" box="[968,1069,1599,1625]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Figs. 3D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB812422C42F9908062190D" box="[1083,1118,1599,1625]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="6" pageNumber="364">6B</figureCitation>
|
||||
); chela stout, fingers longer than length of palm, surface with longitudinal sulcus flanked by row of granules; pollex slightly bent, cutting edge lined with denticles; dactylus 1.2 times as long as palm along upper margin, row of small granules on upper margin, cutting edge with median large tooth lined with denticles, remaining of edge with small denticles (
|
||||
<figureCitation id="134F2A49FFB812422D5AF90481B31991" box="[1315,1423,1706,1733]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="6" pageNumber="364">Fig. 15D</figureCitation>
|
||||
). P2–P5 slender, short; merus distinctly longer than carpus and propodus; dactylus longer than propodus, lanceolate, terminating in corneous tips (
|
||||
<figureCitation id="134F2A49FFB8124229A4F95C867C1859" box="[477,576,1779,1805]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Figs. 3D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB812422A32F95C86531859" box="[587,623,1779,1805]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="6" pageNumber="364">6B</figureCitation>
|
||||
); fourth leg shortest with merus 4.1 times as long as broad, margins lined with small granules, more prominent on ventral margin (
|
||||
<figureCitation id="134F2A49FFB812422B6FF8B887411865" box="[790,893,1815,1841]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="6" pageNumber="364">Fig. 16C</figureCitation>
|
||||
). Thoracic sternites transversely narrow, surface finely granulated, larger granules laterally (
|
||||
<figureCitation id="134F2A49FFB812422AE8F89486D31801" box="[657,751,1851,1877]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="6" pageNumber="364">Fig. 7D</figureCitation>
|
||||
). Sternopleonal cavity deep, reaching to proximal distance between fused thoracic sternites 1–3. Pleon narrow, somites 2–6 fused, somite 6 longitudinally subrectangular, surface with one low granule medially, lateral margins gently convex; telson triangular (
|
||||
<figureCitation id="134F2A49FFB812422C1CF82C80F418C9" box="[1125,1224,1923,1949]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="6" pageNumber="364">Figs. 7D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB812422CAAF82C80C018C9" box="[1235,1276,1923,1949]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="6" pageNumber="364">17I</figureCitation>
|
||||
). G1 elongate, slender, same size from proximal to distal area; distal process bilobed, tip rounded (
|
||||
<figureCitation id="134F2A49FFB812422C10F80880D71895" box="[1129,1259,1959,1985]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="6" pageNumber="364">Fig. 17J‒L</figureCitation>
|
||||
). Female pleon broad, convex, smooth, somites 2‒6 fused, suture between somites 2 and 3; telson as long as its basal width (
|
||||
<figureCitation id="134F2A49FFB812432D16F86484861FE5" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" lastPageId="7" lastPageNumber="365" pageId="6" pageNumber="364">Fig. 9B</figureCitation>
|
||||
); vulvae positioned closer to median part of sternite 6, each opening prominently crescent-shaped, relatively long, without opercular cover (
|
||||
<figureCitation id="134F2A49FFB91243298BFF1486641F81" box="[498,600,187,213]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="7" pageNumber="365">Fig. 21B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFB9124328BEFF7185921FAD" blockId="7.[151,1436,151,249]" box="[199,430,222,249]" pageId="7" pageNumber="365">
|
||||
<emphasis id="B900EADEFFB9124328BEFF7185271FAC" bold="true" box="[199,283,222,248]" pageId="7" pageNumber="365">Colour</emphasis>
|
||||
. Not known.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF0B6644FFB9124328EEF8B5864318B2" ID-DOI="http://doi.org/10.5281/zenodo.12681932" ID-Zenodo-Dep="12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="7" pageNumber="365" startId="7.[151,250,1818,1842]" targetBox="[151,1435,273,1794]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="8BCB36CCFFB9124328EEF8B5864318B2" blockId="7.[151,1436,1818,2022]" pageId="7" pageNumber="365">
|
||||
<emphasis id="B900EADEFFB9124328EEF8B585291866" bold="true" box="[151,277,1818,1842]" pageId="7" pageNumber="365">FIGURE 3.</emphasis>
|
||||
Overall dorsal habitus. A,
|
||||
<taxonomicName id="4C744D4FFFB912432A56F8B587861866" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[559,954,1818,1842]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB912432A56F8B5873F1866" box="[559,771,1818,1842]" italics="true" pageId="7" pageNumber="365">Lyphira heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB912432B6BF8B5878F1866" author="Ortmann, A." box="[786,947,1818,1842]" pageId="7" pageNumber="365" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B,
|
||||
<taxonomicName id="4C744D4FFFB912432885F89086791802" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[252,581,1854,1878]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB912432885F89085AF1802" box="[252,403,1854,1878]" italics="true" pageId="7" pageNumber="365">L. heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB9124329D9F89186021802" author="Ortmann, A." box="[416,574,1854,1878]" pageId="7" pageNumber="365" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of
|
||||
<taxonomicName id="4C744D4FFFB912432CFCF89184F2182E" authority="Shen, 1932" authorityName="Shen" authorityYear="1932" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="peitaihoensis">
|
||||
<emphasis id="B900EADEFFB912432CFCF89181611802" box="[1157,1373,1854,1878]" italics="true" pageId="7" pageNumber="365">Philyra peitaihoensis</emphasis>
|
||||
<bibRefCitation id="EFE54B3DFFB912432D1AF89184F2182E" author="Shen, C. - J." pageId="7" pageNumber="365" pagination="1 - 300" refId="ref21817" refString="Shen, C. - J. (1932) The brachyuran Crustacea of North China. Zoologia Sinica, 9 (1), 1 - 300, pls. 1 - 10." type="journal article" year="1932">Shen, 1932</bibRefCitation>
|
||||
</taxonomicName>
|
||||
], China; C,
|
||||
<taxonomicName id="4C744D4FFFB912432938F8CC86B8182E" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[321,644,1890,1914]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB912432938F8CC85E9182E" box="[321,469,1890,1914]" italics="true" pageId="7" pageNumber="365">L. heterograna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB912432998F8CD8641182E" author="Ortmann, A." box="[481,637,1890,1914]" pageId="7" pageNumber="365" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D,
|
||||
<taxonomicName id="4C744D4FFFB912432CEDF8CC81A5182E" authority="(Chen, 1987)" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[1172,1433,1890,1914]" class="Malacostraca" family="Leucosiidae" genus="Phylira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB912432CEDF8CC8132182E" box="[1172,1294,1890,1914]" italics="true" pageId="7" pageNumber="365">L. acutidens</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB912432D60F8CD81AD182E" author="Chen, H." box="[1305,1425,1890,1914]" pageId="7" pageNumber="365" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E,
|
||||
<taxonomicName id="4C744D4FFFB912432AAFF828870718CA" authorityName="Rahayu & Ng" authorityYear="2024" box="[726,827,1926,1950]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB912432AAFF828870718CA" box="[726,827,1926,1950]" italics="true" pageId="7" pageNumber="365">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB912432B3BF829874118CA" bold="true" box="[834,893,1926,1950]" pageId="7" pageNumber="365">
|
||||
<taxonomicNameLabel id="A23357A5FFB912432B3BF829874118CA" box="[834,893,1926,1950]" pageId="7" pageNumber="365" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F,
|
||||
<taxonomicName id="4C744D4FFFB912432962F80485431896" authorityName="Rahayu & Ng" authorityYear="2024" box="[283,383,1962,1986]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB912432962F80485431896" box="[283,383,1962,1986]" italics="true" pageId="7" pageNumber="365">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB9124329FEF80585FC1896" bold="true" box="[391,448,1962,1986]" pageId="7" pageNumber="365">
|
||||
<taxonomicNameLabel id="A23357A5FFB9124329FEF80585FC1896" box="[391,448,1962,1986]" pageId="7" pageNumber="365" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G,
|
||||
<taxonomicName id="4C744D4FFFB912432C22F80480831896" authorityName="Rahayu & Ng" authorityYear="2024" box="[1115,1215,1962,1986]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB912432C22F80480831896" box="[1115,1215,1962,1986]" italics="true" pageId="7" pageNumber="365">L. chomel</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB912432CBFF805813C1896" bold="true" box="[1222,1280,1962,1986]" pageId="7" pageNumber="365">
|
||||
<taxonomicNameLabel id="A23357A5FFB912432CBFF805813C1896" box="[1222,1280,1962,1986]" pageId="7" pageNumber="365" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF0B6644FFB6124C28EEF8BE854418C1" ID-DOI="http://doi.org/10.5281/zenodo.12681934" ID-Zenodo-Dep="12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="8" pageNumber="366" startId="8.[151,250,1809,1833]" targetBox="[151,1435,181,1785]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="8BCB36CCFFB6124C28EEF8BE854418C1" blockId="8.[151,1437,1809,1941]" pageId="8" pageNumber="366">
|
||||
<emphasis id="B900EADEFFB6124C28EEF8BE8528187D" bold="true" box="[151,276,1809,1833]" pageId="8" pageNumber="366">FIGURE 4.</emphasis>
|
||||
Overall dorsal habitus,
|
||||
<taxonomicName id="4C744D4FFFB6124C2A70F8BE86A8187D" authorityName="Rahayu & Ng" authorityYear="2024" box="[521,660,1809,1833]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="366" phylum="Arthropoda" rank="species" species="linda" status="sp. nov.">
|
||||
<emphasis id="B900EADEFFB6124C2A70F8BE86A8187D" box="[521,660,1809,1833]" italics="true" pageId="8" pageNumber="366">Lyphira linda</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB6124C2AE2F8BE86E9187D" bold="true" box="[667,725,1809,1833]" pageId="8" pageNumber="366">
|
||||
<taxonomicNameLabel id="A23357A5FFB6124C2AE2F8BE86E9187D" box="[667,725,1809,1833]" pageId="8" pageNumber="366" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF0B6644FFB7124D28EEFB7185701A36" ID-DOI="http://doi.org/10.5281/zenodo.12681936" ID-Zenodo-Dep="12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="9" pageNumber="367" startId="9.[151,250,1246,1270]" targetBox="[151,1435,181,1222]" targetPageId="9" targetType="figure">
|
||||
<paragraph id="8BCB36CCFFB7124D28EEFB7185701A36" blockId="9.[151,1437,1246,1378]" pageId="9" pageNumber="367">
|
||||
<emphasis id="B900EADEFFB7124D28EEFB7185291BA2" bold="true" box="[151,277,1246,1270]" pageId="9" pageNumber="367">FIGURE 5.</emphasis>
|
||||
Overall dorsal habitus. A,
|
||||
<taxonomicName id="4C744D4FFFB7124D2A48FB7186E21BA2" authorityName="Rahayu & Ng" authorityYear="2024" box="[561,734,1246,1270]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="ngankee" status="sp. nov.">
|
||||
<emphasis id="B900EADEFFB7124D2A48FB7186E21BA2" box="[561,734,1246,1270]" italics="true" pageId="9" pageNumber="367">Lyphira ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB7124D2A9EFB7187141BA2" bold="true" box="[743,808,1246,1270]" pageId="9" pageNumber="367">
|
||||
<taxonomicNameLabel id="A23357A5FFB7124D2A9EFB71871F1BA2" box="[743,803,1246,1270]" pageId="9" pageNumber="367" rank="species">n. sp.</taxonomicNameLabel>
|
||||
,
|
||||
</emphasis>
|
||||
holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B,
|
||||
<taxonomicName id="4C744D4FFFB7124D2886FAAC85521A4E" authorityName="Rahayu & Ng" authorityYear="2024" box="[255,366,1282,1306]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB7124D2886FAAC85521A4E" box="[255,366,1282,1306]" italics="true" pageId="9" pageNumber="367">L. ngankee</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23357A5FFB7124D290CFAAD85921A4D" box="[373,430,1282,1306]" pageId="9" pageNumber="367" rank="species">
|
||||
<emphasis id="B900EADEFFB7124D290CFAAD85941A4E" bold="true" box="[373,424,1282,1306]" pageId="9" pageNumber="367">n. sp</emphasis>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
, paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C,
|
||||
<taxonomicName id="4C744D4FFFB7124D2C14FAAC80CB1A4E" authorityName="Rahayu & Ng" authorityYear="2024" box="[1133,1271,1282,1306]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB7124D2C14FAAC80CB1A4E" box="[1133,1271,1282,1306]" italics="true" pageId="9" pageNumber="367">L. bellagrana</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB7124D2C87FAAD810B1A4E" bold="true" box="[1278,1335,1282,1306]" pageId="9" pageNumber="367">
|
||||
<taxonomicNameLabel id="A23357A5FFB7124D2C87FAAD810B1A4E" box="[1278,1335,1282,1306]" pageId="9" pageNumber="367" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D,
|
||||
<taxonomicName id="4C744D4FFFB7124D2B65FA8887991A6A" authorityName="Rahayu & Ng" authorityYear="2024" box="[796,933,1318,1342]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||||
<emphasis id="B900EADEFFB7124D2B65FA8887991A6A" box="[796,933,1318,1342]" italics="true" pageId="9" pageNumber="367">L. bellagrana</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B900EADEFFB7124D2BD5FA8987D91A6A" bold="true" box="[940,997,1318,1342]" pageId="9" pageNumber="367">
|
||||
<taxonomicNameLabel id="A23357A5FFB7124D2BD5FA8987D91A6A" box="[940,997,1318,1342]" pageId="9" pageNumber="367" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C36E6547FFB7124E28BEFA3F800C1EF9" lastPageId="10" lastPageNumber="368" pageId="9" pageNumber="367" type="discussion">
|
||||
<paragraph id="8BCB36CCFFB7124D28BEFA3F807A19F2" blockId="9.[151,1437,1424,2027]" pageId="9" pageNumber="367">
|
||||
<emphasis id="B900EADEFFB7124D28BEFA3F85081AFE" bold="true" box="[199,308,1424,1450]" pageId="9" pageNumber="367">Remarks</emphasis>
|
||||
.
|
||||
<bibRefCitation id="EFE54B3DFFB7124D293FFA3F85EA1AFE" author="Chen, H." box="[326,470,1424,1450]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen (1987)</bibRefCitation>
|
||||
described
|
||||
<taxonomicName id="4C744D4FFFB7124D2A2EFA3F871C1AFE" authorityName="Chen" authorityYear="1987" box="[599,800,1424,1450]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB7124D2A2EFA3F871C1AFE" box="[599,800,1424,1450]" italics="true" pageId="9" pageNumber="367">Philyra acutidens</emphasis>
|
||||
</taxonomicName>
|
||||
from a large series of specimens from the East
|
||||
<collectingCountry id="F363765CFFB7124D2D20FA3F81A01AFE" box="[1369,1436,1424,1450]" name="China" pageId="9" pageNumber="367">China</collectingCountry>
|
||||
Sea and Yellow Sea (
|
||||
<specimenCount id="9D72FD45FFB7124D29F5FA1B85CF1A9A" box="[396,499,1460,1486]" count="87" pageId="9" pageNumber="367" type="male">87 males</specimenCount>
|
||||
and
|
||||
<specimenCount id="9D72FD45FFB7124D2A57FA1B86821A9A" box="[558,702,1460,1486]" count="101" pageId="9" pageNumber="367" type="female">101 females</specimenCount>
|
||||
), and she compared it with
|
||||
<taxonomicName id="4C744D4FFFB7124D2B87FA1A80981A9A" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[1022,1188,1460,1486]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB7124D2B87FA1A80981A9A" box="[1022,1188,1460,1486]" italics="true" pageId="9" pageNumber="367">P. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
, noting that it can be distinguished by its slightly smaller adult size (largest carapace length
|
||||
<quantity id="4C8C9B29FFB7124D2BC4FA77801E1AA7" box="[957,1058,1496,1523]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="9" pageNumber="367" unit="mm" value="16.5">16.5 mm</quantity>
|
||||
versus
|
||||
<quantity id="4C8C9B29FFB7124D2C03FA7780D81AA6" box="[1146,1252,1496,1523]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="9" pageNumber="367" unit="mm" value="18.5">18.5 mm</quantity>
|
||||
), slightly longer carapace, the presence of more granules on the carapace, the third maxilliped exopod is slightly narrower, the proportionately shorter fingers of the chela, the cutting margin of the fingers lacking a large tubercle, male pleonal somite 6 with median tooth, and the female telson is longer than broad. She figured a specimen of what she regarded was
|
||||
<taxonomicName id="4C744D4FFFB7124D28B3F9C6854D19D6" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[202,369,1640,1666]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB7124D28B3F9C6854D19D6" box="[202,369,1640,1666]" italics="true" pageId="9" pageNumber="367">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23357A5FFB7124D2900F9C6858819D7" box="[377,436,1641,1667]" pageId="9" pageNumber="367" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
(
|
||||
<bibRefCitation id="EFE54B3DFFB7124D29BAF9C7867F19D6" author="Chen, H." box="[451,579,1640,1666]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen 1987</bibRefCitation>
|
||||
: fig. 2) but did not state where it was from. From her figures,
|
||||
<bibRefCitation id="EFE54B3DFFB7124D2C87F9C781A519D6" author="Chen, H." box="[1278,1433,1640,1666]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen’s (1987</bibRefCitation>
|
||||
: fig. 2) “
|
||||
<taxonomicName id="4C744D4FFFB7124D2896F92385EF19F2" authorityName="Ortmann" authorityYear="1892" box="[239,467,1676,1702]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB7124D2896F92385EF19F2" box="[239,467,1676,1702]" italics="true" pageId="9" pageNumber="367">Philyra heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
” agrees well with
|
||||
<taxonomicName id="4C744D4FFFB7124D2ADCF922877019F2" authorityName="Ortmann" authorityYear="1892" box="[677,844,1676,1702]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB7124D2ADCF922877019F2" box="[677,844,1676,1702]" italics="true" pageId="9" pageNumber="367">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23357A5FFB7124D2B2AF92287B219F3" box="[851,910,1677,1703]" pageId="9" pageNumber="367" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
as defined here.
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFB7124D28BEF91F8636182A" blockId="9.[151,1437,1424,2027]" pageId="9" pageNumber="367">
|
||||
<bibRefCitation id="EFE54B3DFFB7124D28BEF91F8569199E" author="Galil, B. S." box="[199,341,1712,1738]" pageId="9" pageNumber="367" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil (2009)</bibRefCitation>
|
||||
commented that
|
||||
<taxonomicName id="4C744D4FFFB7124D2A5DF91E868D199E" authorityName="Chen" authorityYear="1987" box="[548,689,1712,1738]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB7124D2A5DF91E868D199E" box="[548,689,1712,1738]" italics="true" pageId="9" pageNumber="367">L. acutidens</emphasis>
|
||||
</taxonomicName>
|
||||
is a junior synonym of
|
||||
<taxonomicName id="4C744D4FFFB7124D2BACF91E80BC199E" authorityName="Ortmann" authorityYear="1892" box="[981,1152,1712,1738]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB7124D2BACF91E80BC199E" box="[981,1152,1712,1738]" italics="true" pageId="9" pageNumber="367">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
, arguing the differences observed were only due to their subadult nature. This is not the case; the species is not small, with the
|
||||
<typeStatus id="54CF886EFFB7124D2C87F97B815C19BA" box="[1278,1376,1748,1774]" pageId="9" pageNumber="367" type="holotype">holotype</typeStatus>
|
||||
male measuring 15.0 × 16.0 mm and
|
||||
<typeStatus id="54CF886EFFB7124D2A7BF95786601846" box="[514,604,1784,1810]" pageId="9" pageNumber="367" type="allotype">allotype</typeStatus>
|
||||
female 11.9 × 12.0 mm. The present male specimen (ZRC 2002.0504) is relatively small (8.9 ×
|
||||
<quantity id="4C8C9B29FFB7124D29ECF8B385D01862" box="[405,492,1820,1846]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="9" pageNumber="367" unit="mm" value="8.6">8.6 mm</quantity>
|
||||
) and does not appear fully adult; but the slightly larger female (13.4 ×
|
||||
<quantity id="4C8C9B29FFB7124D2D7AF8B381541862" box="[1283,1384,1820,1846]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="9" pageNumber="367" unit="mm" value="12.9">12.9 mm</quantity>
|
||||
) has the pleon round and domed and is clearly adult. Their characters, however, agree in most aspects with
|
||||
<bibRefCitation id="EFE54B3DFFB7124D2C87F8EF81A0180E" author="Chen, H." box="[1278,1436,1856,1882]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen’s (1987)</bibRefCitation>
|
||||
detailed descriptions and figures.
|
||||
</paragraph>
|
||||
<paragraph id="8BCB36CCFFB7124E28BEF826800C1EF9" blockId="9.[151,1437,1424,2027]" lastBlockId="10.[151,1437,151,501]" lastPageId="10" lastPageNumber="368" pageId="9" pageNumber="367">
|
||||
We are of the opinion the two species are distinct. The differences in the adult size observed by
|
||||
<bibRefCitation id="EFE54B3DFFB7124D2C91F827814818F6" author="Chen, H." box="[1256,1396,1928,1954]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen (1987)</bibRefCitation>
|
||||
are actually not significant, and carapace proportions alone is not enough to justify recognizing two species. The authors have on hand two non-type topotypic specimens from the Yellow Sea but are part of
|
||||
<bibRefCitation id="EFE54B3DFFB7124D2C29F87F80CC18BE" author="Chen, H." box="[1104,1264,2000,2026]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen’s (1987)</bibRefCitation>
|
||||
original series, and they do show the other differences highlighted by her. Most marked are the denser concentration of granules on the dorsal surface. The median tubercle on male pleonal somite 6 of
|
||||
<taxonomicName id="4C744D4FFFB4124E2BDCFF1480131F81" authorityName="Chen" authorityYear="1987" box="[933,1071,187,213]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB4124E2BDCFF1480131F81" box="[933,1071,187,213]" italics="true" pageId="10" pageNumber="368">L. acutidens</emphasis>
|
||||
</taxonomicName>
|
||||
is slightly sharper than those of
|
||||
<taxonomicName id="4C744D4FFFB4124E28EEFF7085011FAD" authorityName="Ortmann" authorityYear="1892" box="[151,317,223,249]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB4124E28EEFF7085011FAD" box="[151,317,223,249]" italics="true" pageId="10" pageNumber="368">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
and is slightly more subdistal in position, just behind the distal margin (
|
||||
<figureCitation id="134F2A49FFB4124E2C1AFF7080FA1FAD" box="[1123,1222,223,249]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="10" pageNumber="368">Figs. 7D</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2CA8FF7080C41FAD" box="[1233,1272,223,249]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="10" pageNumber="368">17I</figureCitation>
|
||||
) whereas in
|
||||
<taxonomicName id="4C744D4FFFB4124E2DFFFF70851C1E49" authorityName="Ortmann" authorityYear="1892" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB4124E2DFFFF70851C1E49" italics="true" pageId="10" pageNumber="368">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
, it is positioned a bit further back (
|
||||
<figureCitation id="134F2A49FFB4124E2ADCFEAC87151E49" box="[677,809,259,285]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="10" pageNumber="368">Figs. 7A‒C</figureCitation>
|
||||
,
|
||||
<figureCitation id="134F2A49FFB4124E2B4DFEAC87591E49" box="[820,869,259,285]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="10" pageNumber="368">17B</figureCitation>
|
||||
).
|
||||
<bibRefCitation id="EFE54B3DFFB4124E2B00FEAC80241E49" author="Chen, H." box="[889,1048,259,285]" pageId="10" pageNumber="368" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen’s (1987)</bibRefCitation>
|
||||
figure of the male pleon is slightly inaccurate, the median tubercle on somite 6 is not distal in position (
|
||||
<bibRefCitation id="EFE54B3DFFB4124E2BE4FE8880221E15" author="Chen, H." box="[925,1054,295,321]" pageId="10" pageNumber="368" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen 1987</bibRefCitation>
|
||||
: fig. 1.3) but is actually subdistal. Also significant is that the vulvae are differently structured, with the opening in
|
||||
<taxonomicName id="4C744D4FFFB4124E2C6FFEE480811E31" authorityName="Ortmann" authorityYear="1892" box="[1046,1213,331,357]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="heterograna">
|
||||
<emphasis id="B900EADEFFB4124E2C6FFEE480811E31" box="[1046,1213,331,357]" italics="true" pageId="10" pageNumber="368">L. heterograna</emphasis>
|
||||
</taxonomicName>
|
||||
is short and weakly crescent-shaped (
|
||||
<figureCitation id="134F2A49FFB4124E2920FEC085821EDD" box="[345,446,367,393]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="10" pageNumber="368">Fig. 21A</figureCitation>
|
||||
) whereas it is slit-like and distinctly more crescent-shaped in
|
||||
<taxonomicName id="4C744D4FFFB4124E2C17FEC080CA1EDD" authorityName="Chen" authorityYear="1987" box="[1134,1270,367,393]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB4124E2C17FEC080CA1EDD" box="[1134,1270,367,393]" italics="true" pageId="10" pageNumber="368">L. acutidens</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="134F2A49FFB4124E2D7CFEC081571EDD" box="[1285,1387,367,393]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="10" pageNumber="368">Fig. 21B</figureCitation>
|
||||
). As such, we believe it is better to recognize
|
||||
<taxonomicName id="4C744D4FFFB4124E2A19FE3C86D41EF9" authorityName="Chen" authorityYear="1987" box="[608,744,403,429]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="acutidens">
|
||||
<emphasis id="B900EADEFFB4124E2A19FE3C86D41EF9" box="[608,744,403,429]" italics="true" pageId="10" pageNumber="368">L. acutidens</emphasis>
|
||||
</taxonomicName>
|
||||
as a distinct species for now.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFB4124E28BEFE1886801E85" box="[199,700,438,465]" pageId="10" pageNumber="368" type="materials_examined">
|
||||
<paragraph id="8BCB36CCFFB4124E28BEFE1886801E85" blockId="10.[151,1437,151,501]" box="[199,700,438,465]" pageId="10" pageNumber="368">
|
||||
<materialsCitation id="3B1C3C91FFB4124E28BEFE1886841E85" box="[199,696,438,465]" country="China" county="Sea" location="Yellow Sea" municipality="North" pageId="10" pageNumber="368" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="B900EADEFFB4124E28BEFE18855C1E84" bold="true" box="[199,352,438,465]" pageId="10" pageNumber="368">
|
||||
<typeStatus id="54CF886EFFB4124E28BEFE18853E1E85" box="[199,258,439,465]" pageId="10" pageNumber="368">Type</typeStatus>
|
||||
locality
|
||||
</emphasis>
|
||||
.
|
||||
<location id="8EAB6017FFB4124E2914FE1885D21E85" LSID="urn:lsid:plazi:treatment:03DD87DAFFB8124E28EEFBBC87BE1EA1:8EAB6017FFB4124E2914FE1885D21E85" box="[365,494,439,465]" country="China" county="Sea" municipality="North" name="Yellow Sea" pageId="10" pageNumber="368">Yellow Sea</location>
|
||||
,
|
||||
<collectingMunicipality id="6BAFACB6FFB4124E2983FE1886071E85" box="[506,571,439,465]" pageId="10" pageNumber="368">North</collectingMunicipality>
|
||||
<collectingCountry id="F363765CFFB4124E2A3BFE1886B91E85" box="[578,645,439,465]" name="China" pageId="10" pageNumber="368">China</collectingCountry>
|
||||
<collectingCounty id="62AA4E40FFB4124E2AF5FE1886841E85" box="[652,696,439,465]" pageId="10" pageNumber="368">Sea</collectingCounty>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C36E6547FFB4124E28BEFE7587BE1EA1" box="[199,898,474,501]" pageId="10" pageNumber="368" type="distribution">
|
||||
<paragraph id="8BCB36CCFFB4124E28BEFE7587BE1EA1" blockId="10.[151,1437,151,501]" box="[199,898,474,501]" pageId="10" pageNumber="368">
|
||||
<emphasis id="B900EADEFFB4124E28BEFE7585651EA0" bold="true" box="[199,345,474,500]" pageId="10" pageNumber="368">Distribution</emphasis>
|
||||
. Bohai Sea, Yellow Sea and Northern
|
||||
<collectingCountry id="F363765CFFB4124E2B71FE7487771EA1" box="[776,843,475,501]" name="China" pageId="10" pageNumber="368">China</collectingCountry>
|
||||
Sea.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
1009
data/03/DD/87/03DD87DAFFBC124228EEFEE5803F1CEB.xml
Normal file
1009
data/03/DD/87/03DD87DAFFBC124228EEFEE5803F1CEB.xml
Normal file
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Reference in a new issue