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<document id="C0C29CD157A17B92CBE3998FC7E75260" ID-DOI="10.11646/zootaxa.5481.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="12749603" ID-ZooBank="17CBED32-6542-4616-8ACE-C32D7258395B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721126335027" checkinUser="plazi" docAuthor="Korai, Shakal Khan &amp; Jäger, Peter" docDate="2024" docId="038A9040D313FFA7868B794D55556AED" docLanguage="en" docName="zootaxa.5481.2.4.pdf" docOrigin="Zootaxa 5481 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heteropoda papilionacea Korai &amp; Jäger, 2024, spec. nov." docType="treatment" docVersion="2" lastPageNumber="252" masterDocId="FFB3E838D31AFFAC861C7D7B54666948" masterDocTitle="Five new species of Heteropoda Latreille, 1804 spiders (Araneae: Sparassidae) from Southeast Asia" masterLastPageNumber="259" masterPageNumber="241" pageNumber="250" updateTime="1721126798175" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5DAE2E155CE1553D120C5ADEF8DA1AC6">Five new species of Heteropoda Latreille, 1804 spiders (Araneae: Sparassidae) from Southeast Asia</mods:title>
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<mods:namePart id="93D3C72D607F54A08EFBEF42770DCB22">Korai, Shakal Khan</mods:namePart>
<mods:affiliation id="FE607472C3C72B8534B3873C47079736">The State Key Laboratory of Biocatalysis and Enzyme Engineering of China, School of Life Sciences, Hubei University, Wuhan 430062, Hubei, China.</mods:affiliation>
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<mods:namePart id="50F33E348788B43BF3A39469EB2B55CA">Jäger, Peter</mods:namePart>
<mods:affiliation id="4688945EB6B8706FAA44AEF4D9869911">Senckenberg Research Institute, Arachnology, Mertonstrasse 17 - 21, 60325 Frankfurt am Main, Germany.</mods:affiliation>
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<mods:date id="5E1A853A59A280B3D38BF054F7227D1C">2024</mods:date>
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<mods:number id="952DE7EBA3DC45A62437497DFAC17A7D">2024-07-16</mods:number>
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<treatment id="038A9040D313FFA7868B794D55556AED" ID-DOI="http://doi.org/10.5281/zenodo.12749595" ID-Zenodo-Dep="12749595" LSID="urn:lsid:plazi:treatment:038A9040D313FFA7868B794D55556AED" httpUri="http://treatment.plazi.org/id/038A9040D313FFA7868B794D55556AED" lastPageId="11" lastPageNumber="252" pageId="9" pageNumber="250">
<subSubSection id="C33972DDD313FFA5868B794D562D6D19" box="[151,587,1078,1105]" pageId="9" pageNumber="250" type="nomenclature">
<paragraph id="8B9C2156D313FFA5868B794D562D6D19" blockId="9.[151,587,1078,1141]" box="[151,587,1078,1105]" pageId="9" pageNumber="250">
<heading id="D0D4963AD313FFA5868B794D562D6D19" bold="true" box="[151,587,1078,1105]" fontSize="11" level="1" pageId="9" pageNumber="250" reason="1">
<emphasis id="B957FD44D313FFA5868B794D562D6D19" bold="true" box="[151,587,1078,1105]" pageId="9" pageNumber="250">
<taxonomicName id="4C235AD5D313FFA5868B794D55AC6D19" authorityName="Korai &amp; Jäger" authorityYear="2024" box="[151,458,1078,1105]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="papilionacea" status="sp. nov.">
<emphasis id="B957FD44D313FFA5868B794D55AC6D19" bold="true" box="[151,458,1078,1105]" italics="true" pageId="9" pageNumber="250">Heteropoda papilionacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264403FD313FFA587CE794C562D6D19" box="[466,587,1079,1105]" pageId="9" pageNumber="250" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33972DDD313FFA5868B7920555C6D3D" box="[151,314,1115,1141]" pageId="9" pageNumber="250" type="description">
<paragraph id="8B9C2156D313FFA5868B7920555C6D3D" blockId="9.[151,587,1078,1141]" box="[151,314,1115,1141]" pageId="9" pageNumber="250">
<figureCitation id="13183DD3D313FFA5868B792055726D3D" box="[151,276,1115,1141]" captionStart="FIGURES 7" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[177,1411,181,1885]" captionTargetId="figure-24@10.[177,1411,181,1885]" captionTargetPageId="10" captionText="FIGURES 7A–E. Heteropoda papilionacea spec. nov., holotype female from Malaysia, Sabah, Maliau Basin Conservation Area. A–C Copulatory organ (A vulva, dorsal; B epigyne, ventral; C schematic course of internal duct system, dorsal); D–E Habitus (D dorsal, E ventral).Abbreviations:AB—anterior bands; CO—copulatory opening; FW—first winding; GP—glandular pores; LL—lateral lobes; MS—median septum. Scale bars: A–C 0.5 mm; D–E 3 mm." figureDoi="http://doi.org/10.5281/zenodo.12749619" httpUri="https://zenodo.org/record/12749619/files/figure.png" pageId="9" pageNumber="250">Figs 7A–E</figureCitation>
,
<figureCitation id="13183DD3D313FFA587037920555C6D3D" box="[287,314,1115,1141]" captionStart="FIGURE 11" captionStartId="16.[151,250,1327,1351]" captionTargetBox="[191,1395,181,1304]" captionTargetId="figure-24@16.[191,1395,181,1304]" captionTargetPageId="16" captionText="FIGURE 11. Locality records of the examined Heteropoda material." figureDoi="http://doi.org/10.5281/zenodo.12749629" httpUri="https://zenodo.org/record/12749629/files/figure.png" pageId="9" pageNumber="250">11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33972DDD313FFA5868B79D8563A6C4D" pageId="9" pageNumber="250" type="materials_examined">
<paragraph id="8B9C2156D313FFA5868B79D8563A6C4D" blockId="9.[151,1437,1186,2041]" pageId="9" pageNumber="250">
<emphasis id="B957FD44D313FFA5868B79D855206DF4" bold="true" box="[151,326,1186,1213]" pageId="9" pageNumber="250">Type material.</emphasis>
<materialsCitation id="3B4B2B0BD313FFA5875179D850C16DA9" collectingDate="2015-10-17" collectionCode="CBEE" collectorName="L. Y. Wang &amp; G. Q. Huang" country="Malaysia" elevation="385" latitude="5.323889" location="Maliau Basin Conservation Area" longLatPrecision="21" longitude="117.01639" municipality="Tongod District" pageId="9" pageNumber="250" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B957FD44D313FFA5875179D8573B6DF5" bold="true" box="[333,861,1186,1213]" pageId="9" pageNumber="250">
<typeStatus id="54989FF4D313FFA5875179D855DD6DF5" box="[333,443,1187,1213]" pageId="9" pageNumber="250" type="holotype">Holotype</typeStatus>
female:
<collectingCountry id="F33461C6D313FFA5843E79D956D86DF4" box="[546,702,1186,1212]" name="Malaysia" pageId="9" pageNumber="250">MALAYSIA</collectingCountry>
:
<emphasis id="B957FD44D313FFA584D679D957326DF5" bold="true" box="[714,852,1186,1213]" italics="true" pageId="9" pageNumber="250">
<collectingRegion id="49E7EFB4D313FFA584D679D957326DF5" box="[714,852,1186,1213]" country="Malaysia" name="Sabah" pageId="9" pageNumber="250">Sabah State</collectingRegion>
</emphasis>
:
</emphasis>
<collectingMunicipality id="6BF8BB2CD313FFA5857879D8507F6DF5" box="[868,1049,1187,1213]" pageId="9" pageNumber="250">Tongod District</collectingMunicipality>
,
<location id="8EFC778DD313FFA5823879D851FE6DF5" LSID="urn:lsid:plazi:treatment:038A9040D313FFA7868B794D55556AED:8EFC778DD313FFA5823879D851FE6DF5" box="[1060,1432,1187,1213]" country="Malaysia" latitude="5.323889" longLatPrecision="21" longitude="117.01639" municipality="Tongod District" name="Maliau Basin Conservation Area" pageId="9" pageNumber="250" stateProvince="Sabah">Maliau Basin Conservation Area</location>
,
<geoCoordinate id="EE174791D313FFA5868B79B355756DAB" box="[151,275,1224,1251]" degrees="5" direction="north" minutes="19" orientation="latitude" pageId="9" pageNumber="250" precision="15" seconds="26" value="5.323889">5°19′26″N</geoCoordinate>
,
<geoCoordinate id="EE174791D313FFA5870379B355D46DAB" box="[287,434,1224,1251]" degrees="117" direction="east" minutes="00" orientation="longitude" pageId="9" pageNumber="250" precision="15" seconds="59" value="117.01639">117°00′59″E</geoCoordinate>
,
<quantity id="4CDB8CB3D313FFA587A379BD566C6DA9" box="[447,522,1222,1249]" metricMagnitude="2" metricUnit="m" metricValue="3.85" pageId="9" pageNumber="250" unit="m" value="385.0">
<elevation id="000EC665D313FFA587A379BD566C6DA9" box="[447,522,1222,1249]" metricMagnitude="2" metricUnit="m" metricValue="3.85" pageId="9" pageNumber="250" unit="m" value="385.0">385 m</elevation>
</quantity>
,
<date id="FF9D0796D313FFA5840A79BC56BE6DA9" box="[534,728,1222,1249]" pageId="9" pageNumber="250" value="2015-10-17">
<collectingDate id="EFD9FE7ED313FFA5840A79BC56BE6DA9" box="[534,728,1222,1249]" pageId="9" pageNumber="250" value="2015-10-17">17 October 2015</collectingDate>
</date>
,
<collectorName id="26D64480D313FFA584F979BC57386DA9" box="[741,862,1223,1249]" pageId="9" pageNumber="250">L.Y. Wang</collectorName>
&amp;
<collectorName id="26D64480D313FFA5859879BC50686DA9" box="[900,1038,1223,1249]" pageId="9" pageNumber="250">G.Q. Huang</collectorName>
leg. (
<collectionCode id="ED32B993D313FFA5824C79BC50F86DA9" box="[1104,1182,1223,1249]" pageId="9" pageNumber="250">CBEE</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B2B0BD313FFA582AF79BC563E6C4D" collectingDate="2015-10-17" collectionCode="CBEE" collectorName="L. Y. Wang &amp; G. Q. Huang" country="Malaysia" elevation="385" latitude="5.323889" location="Maliau Basin Conservation Area" longLatPrecision="21" longitude="117.01639" municipality="Tongod District" pageId="9" pageNumber="250" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B957FD44D313FFA582AF79BC514C6DA9" bold="true" box="[1203,1322,1223,1249]" pageId="9" pageNumber="250">
<typeStatus id="54989FF4D313FFA582AF79BC51436DA9" box="[1203,1317,1223,1249]" pageId="9" pageNumber="250" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D25EADFD313FFA5832E79BC51FE6DA9" box="[1330,1432,1223,1249]" count="1" pageId="9" pageNumber="250" type="female">1 female</specimenCount>
, with same data as for holotype (
<collectionCode id="ED32B993D313FFA5841E799056366C4D" box="[514,592,1259,1285]" pageId="9" pageNumber="250">CBEE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33972DDD313FFA586DB787450D76C05" pageId="9" pageNumber="250" type="etymology">
<paragraph id="8B9C2156D313FFA586DB787450D76C05" blockId="9.[151,1437,1186,2041]" pageId="9" pageNumber="250">
<emphasis id="B957FD44D313FFA586DB787455296C61" bold="true" box="[199,335,1295,1321]" pageId="9" pageNumber="250">Etymology.</emphasis>
The specific name is derived from the Latin adjective
<emphasis id="B957FD44D313FFA585A57874503E6C61" box="[953,1112,1295,1321]" italics="true" pageId="9" pageNumber="250">papilionaceus</emphasis>
,
<emphasis id="B957FD44D313FFA5827A787450C86C60" box="[1126,1198,1295,1320]" italics="true" pageId="9" pageNumber="250">-a, um</emphasis>
, meaning “butterflyshaped”, referring to the median septum and median margins of lateral lobes in ventral view.
</paragraph>
</subSubSection>
<subSubSection id="C33972DDD313FFA586DB782D55996F91" pageId="9" pageNumber="250" type="diagnosis">
<paragraph id="8B9C2156D313FFA586DB782D55996F91" blockId="9.[151,1437,1186,2041]" pageId="9" pageNumber="250">
<emphasis id="B957FD44D313FFA586DB782D55256C38" bold="true" box="[199,323,1366,1392]" pageId="9" pageNumber="250">Diagnosis.</emphasis>
Females of this new species can be distinguished from all other
<taxonomicName id="4C235AD5D313FFA58201782C509A6C39" box="[1053,1276,1367,1393]" pageId="9" pageNumber="250">
<emphasis id="B957FD44D313FFA58201782C50C56C39" box="[1053,1187,1367,1393]" italics="true" pageId="9" pageNumber="250">Heteropoda</emphasis>
species
</taxonomicName>
by the unique combination of long, straight, parallel median margins of the first windings, and anterior part of internal duct system is longer than wide, with just one winding (
<bibRefCitation id="EFB25CA7D313FFA584EF78E457116CF1" author="Jager, P." box="[755,887,1439,1465]" pageId="9" pageNumber="250" pagination="145 - 188" refId="ref8002" refString="Jager, P. (2014) Heteropoda Latreille, 1804: new species, synonymies, transfers and records (Araneae: Sparassidae: Heteropodinae). Arthropoda Selecta, 23, 145 - 188. https: // doi. org / 10.15298 / arthsel. 23.2.06" type="journal article" year="2014">Jäger 2014</bibRefCitation>
: figs 124–135). They are most similar to those of
<taxonomicName id="4C235AD5D313FFA586A478B855B96C95" authority="Strand, 1915" authorityName="Strand" authorityYear="1915" box="[184,479,1474,1501]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="strasseni">
<emphasis id="B957FD44D313FFA586A478B855246C94" box="[184,322,1475,1500]" italics="true" pageId="9" pageNumber="250">H. strasseni</emphasis>
<bibRefCitation id="EFB25CA7D313FFA5875778B855B96C95" author="Strand, E." box="[331,479,1474,1501]" pageId="9" pageNumber="250" refId="ref8429" refString="Strand, E. (1915) Indoaustralische, papuanische und polynesische Spinnen des Senckenbergischen Museums, gesammelt von Dr E. Wolf, Dr J. Elbert u. a. In: Wissenschaftliche Ergebnisse der Hanseatischen Sudsee-Expedition 1909. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft. Vol. 36. Senckenbergische Naturforschende Gesellschaft, Frankfurt am Main, pp. 179 - 274." type="book" year="1915">Strand, 1915</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C235AD5D313FFA5840678B8569A6C95" authority="Jager, 2014" authorityName="Jager" authorityYear="2014" box="[538,764,1475,1501]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="kusi">
<emphasis id="B957FD44D313FFA5840678B8560B6C95" box="[538,621,1475,1501]" italics="true" pageId="9" pageNumber="250">H. kusi</emphasis>
<bibRefCitation id="EFB25CA7D313FFA5846A78B8569A6C95" author="Jager, P." box="[630,764,1475,1501]" pageId="9" pageNumber="250" pagination="145 - 188" refId="ref8002" refString="Jager, P. (2014) Heteropoda Latreille, 1804: new species, synonymies, transfers and records (Araneae: Sparassidae: Heteropodinae). Arthropoda Selecta, 23, 145 - 188. https: // doi. org / 10.15298 / arthsel. 23.2.06" type="journal article" year="2014">Jäger, 2014</bibRefCitation>
</taxonomicName>
, but can be distinguished from the latter by the following characters: 1. Anterior bands are moderately long (significantly short, and integrated in
<taxonomicName id="4C235AD5D313FFA5829A789C51696F48" authorityName="Strand" authorityYear="1915" box="[1158,1295,1511,1536]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="strasseni">
<emphasis id="B957FD44D313FFA5829A789C51696F48" box="[1158,1295,1511,1536]" italics="true" pageId="9" pageNumber="250">H. strasseni</emphasis>
</taxonomicName>
); 2. Median septum mostly covered, only anteriormost part of septum visible (broad and freely visible in
<taxonomicName id="4C235AD5D313FFA582C57B7051056F6C" authorityName="Strand" authorityYear="1915" box="[1241,1379,1547,1572]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="strasseni">
<emphasis id="B957FD44D313FFA582C57B7051056F6C" box="[1241,1379,1547,1572]" italics="true" pageId="9" pageNumber="250">H. strasseni</emphasis>
</taxonomicName>
, and partly covered in
<taxonomicName id="4C235AD5D313FFA587437B5455D66F01" authorityName="Jager" authorityYear="2014" box="[351,432,1583,1609]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="kusi">
<emphasis id="B957FD44D313FFA587437B5455D66F01" box="[351,432,1583,1609]" italics="true" pageId="9" pageNumber="250">H. kusi</emphasis>
</taxonomicName>
); 3. Lateral lobes touching each other medially (distinctly separated in
<taxonomicName id="4C235AD5D313FFA582C17B5451036F00" authorityName="Strand" authorityYear="1915" box="[1245,1381,1583,1608]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="strasseni">
<emphasis id="B957FD44D313FFA582C17B5451036F00" box="[1245,1381,1583,1608]" italics="true" pageId="9" pageNumber="250">H. strasseni</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C235AD5D313FFA5868B7B28548E6F25" authorityName="Jager" authorityYear="2014" box="[151,232,1619,1645]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="kusi">
<emphasis id="B957FD44D313FFA5868B7B28548E6F25" box="[151,232,1619,1645]" italics="true" pageId="9" pageNumber="250">H. kusi</emphasis>
</taxonomicName>
); 4. Posterior part of internal duct system almost two times longer than wide and roughly kidney-shaped, and extending laterally beyond anterior part (just roughly rounded in
<taxonomicName id="4C235AD5D313FFA585B27B0C50506FD8" authorityName="Strand" authorityYear="1915" box="[942,1078,1655,1680]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="strasseni">
<emphasis id="B957FD44D313FFA585B27B0C50506FD8" box="[942,1078,1655,1680]" italics="true" pageId="9" pageNumber="250">H. strasseni</emphasis>
</taxonomicName>
, and slightly extending beyond anterior part in
<taxonomicName id="4C235AD5D313FFA5875F7BE055F46FFD" authorityName="Jager" authorityYear="2014" box="[323,402,1691,1717]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="kusi">
<emphasis id="B957FD44D313FFA5875F7BE055F46FFD" box="[323,402,1691,1717]" italics="true" pageId="9" pageNumber="250">H. kusi</emphasis>
</taxonomicName>
); 5. Glandular pores postero-laterad (posteriad in
<taxonomicName id="4C235AD5D313FFA585DD7BE050216FFC" authorityName="Strand" authorityYear="1915" box="[961,1095,1691,1716]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="strasseni">
<emphasis id="B957FD44D313FFA585DD7BE050216FFC" box="[961,1095,1691,1716]" italics="true" pageId="9" pageNumber="250">H. strasseni</emphasis>
</taxonomicName>
, and laterad to slightly anterolaterad in
<taxonomicName id="4C235AD5D313FFA587157BC4553F6F91" authorityName="Jager" authorityYear="2014" box="[265,345,1727,1753]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="kusi">
<emphasis id="B957FD44D313FFA587157BC4553F6F91" box="[265,345,1727,1753]" italics="true" pageId="9" pageNumber="250">H. kusi</emphasis>
</taxonomicName>
) (
<figureCitation id="13183DD3D313FFA5876D7BC455956F91" box="[369,499,1727,1753]" captionStart="FIGURES 7" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[177,1411,181,1885]" captionTargetId="figure-24@10.[177,1411,181,1885]" captionTargetPageId="10" captionText="FIGURES 7A–E. Heteropoda papilionacea spec. nov., holotype female from Malaysia, Sabah, Maliau Basin Conservation Area. A–C Copulatory organ (A vulva, dorsal; B epigyne, ventral; C schematic course of internal duct system, dorsal); D–E Habitus (D dorsal, E ventral).Abbreviations:AB—anterior bands; CO—copulatory opening; FW—first winding; GP—glandular pores; LL—lateral lobes; MS—median septum. Scale bars: A–C 0.5 mm; D–E 3 mm." figureDoi="http://doi.org/10.5281/zenodo.12749619" httpUri="https://zenodo.org/record/12749619/files/figure.png" pageId="9" pageNumber="250">Figs 7A–C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33972DDD313FFA786DB7B9956AD6B75" lastPageId="11" lastPageNumber="252" pageId="9" pageNumber="250" type="description">
<paragraph id="8B9C2156D313FFA586DB7B9956406FB5" blockId="9.[151,1437,1186,2041]" box="[199,550,1762,1789]" pageId="9" pageNumber="250">
<emphasis id="B957FD44D313FFA586DB7B9955CE6FB5" bold="true" box="[199,424,1762,1789]" pageId="9" pageNumber="250">Description. Male:</emphasis>
Unknown.
</paragraph>
<paragraph id="8B9C2156D313FFA586DB7A7C55416EB1" blockId="9.[151,1437,1186,2041]" pageId="9" pageNumber="250">
<emphasis id="B957FD44D313FFA586DB7A7C55CF6E68" bold="true" box="[199,425,1798,1825]" pageId="9" pageNumber="250">
Female (
<typeStatus id="54989FF4D313FFA587317A7D55FA6E68" box="[301,412,1798,1824]" pageId="9" pageNumber="250" type="holotype">holotype</typeStatus>
):
</emphasis>
Measurements: BL 15.6; PL 6.7, PW 6.2; AW 0.4; OL 8.9, OW 5.2. Eyes: AME 0.31, ALE 0.54, PME 0.36, PLE 0.58, AME–AME 0.28, AME–ALE 0.16, PME–PME 0.44, PME–PLE 0.68, AME–PME 0.54, ALE–PLE 0.55,
<collectingCountry id="F33461C6D313FFA587887A3455DD6E21" box="[404,443,1871,1897]" name="Switzerland" pageId="9" pageNumber="250">CH</collectingCountry>
AME 0.41,
<collectingCountry id="F33461C6D313FFA5845A7A34560B6E21" box="[582,621,1871,1897]" name="Switzerland" pageId="9" pageNumber="250">CH</collectingCountry>
ALE 0.36. Spination: Palp 120, 000, 2111; Fe I–II 212, III 302, IV 211; Pa I–IV 021; Ti
<date id="FF9D0796D313FFA5874F7A0855A76EC5" box="[339,449,1907,1933]" pageId="9" pageNumber="250" value="1302-01" valueMax="1302-02-31" valueMin="1302-01-00">I–II 1302</date>
,
<date id="FF9D0796D313FFA587D27A08563F6EC5" box="[462,601,1907,1933]" pageId="9" pageNumber="250" value="2411-03" valueMax="2411-04-31" valueMin="2411-03-00">III–IV 2411</date>
; Mt
<date id="FF9D0796D313FFA584927A0857616EC5" box="[654,775,1907,1933]" pageId="9" pageNumber="250" value="1010-01" valueMax="1010-04-31" valueMin="1010-01-00">I–IV 1010</date>
. Measurements of palp and legs: Palp 7.6 (2.3, 1.1, 1.9,–, 2.2), I 22.4 (7.1, 2.3, 7.0, 4.4, 1.5), II 24.5 (7.5, 2.3, 7.6, 5.5, 1.4), III 18.3 (5.4, 1.9, 5.9, 3.4, 1.5), IV 21.8 (5.4, 1.7, 6.5, 5.9, 2.1). Leg formula: II–I–IV–III. Cheliceral furrow with 3 promarginal, 4 retromarginal teeth and ca. 35 denticles.
</paragraph>
<caption id="DF5C71DED310FFA6868B7A0F579A6EB0" ID-DOI="http://doi.org/10.5281/zenodo.12749619" ID-Zenodo-Dep="12749619" httpUri="https://zenodo.org/record/12749619/files/figure.png" pageId="10" pageNumber="251" startId="10.[151,264,1908,1932]" targetBox="[177,1411,181,1885]" targetPageId="10" targetType="figure">
<paragraph id="8B9C2156D310FFA6868B7A0F579A6EB0" blockId="10.[151,1436,1908,2041]" pageId="10" pageNumber="251">
<emphasis id="B957FD44D310FFA6868B7A0F55326EC4" bold="true" box="[151,340,1908,1932]" pageId="10" pageNumber="251">FIGURES 7A–E.</emphasis>
<taxonomicName id="4C235AD5D310FFA687407A0E56066EC5" authorityName="Korai &amp; Jäger" authorityYear="2024" box="[348,608,1909,1933]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="papilionacea" status="sp. nov.">
<emphasis id="B957FD44D310FFA687407A0E56066EC5" box="[348,608,1909,1933]" italics="true" pageId="10" pageNumber="251">Heteropoda papilionacea</emphasis>
</taxonomicName>
<emphasis id="B957FD44D310FFA684757A0E56B46EC5" bold="true" box="[617,722,1909,1933]" pageId="10" pageNumber="251">
<taxonomicNameLabel id="A264403FD310FFA684757A0E56B46EC5" box="[617,722,1909,1933]" pageId="10" pageNumber="251" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, holotype female from Malaysia, Sabah, Maliau Basin Conservation Area. A–C Copulatory organ (A vulva, dorsal; B epigyne, ventral; C schematic course of internal duct system, dorsal); D–E Habitus (D dorsal, E ventral).Abbreviations:AB—anterior bands; CO—copulatory opening; FW—first winding; GP—glandular pores; LL—lateral lobes; MS—median septum. Scale bars: A–C 0.5 mm; D–E 3 mm.
</paragraph>
</caption>
<paragraph id="8B9C2156D311FFA786DB7DEC55C9682D" blockId="11.[151,1437,151,933]" pageId="11" pageNumber="252">
Copulatory organ as in diagnosis. Epigynal field roughly as long as wide, posteriorly distinctly wider than anteriorly, with short triangular anterior bands and one slit sensillum on each side separated from the field. Median margins of lateral lobes strongly curved in anterior half when almost touching in ventral view. Median margins of first windings of both functional entities of the internal duct system very close to each other and running parallel. Posterior part of internal duct system large, elongated and roughly kidney-shaped. Fertilization ducts located posteriorly (
<figureCitation id="13183DD3D311FFA7873D7C3055C5682D" box="[289,419,331,357]" captionStart="FIGURES 7" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[177,1411,181,1885]" captionTargetId="figure-24@10.[177,1411,181,1885]" captionTargetPageId="10" captionText="FIGURES 7A–E. Heteropoda papilionacea spec. nov., holotype female from Malaysia, Sabah, Maliau Basin Conservation Area. A–C Copulatory organ (A vulva, dorsal; B epigyne, ventral; C schematic course of internal duct system, dorsal); D–E Habitus (D dorsal, E ventral).Abbreviations:AB—anterior bands; CO—copulatory opening; FW—first winding; GP—glandular pores; LL—lateral lobes; MS—median septum. Scale bars: A–C 0.5 mm; D–E 3 mm." figureDoi="http://doi.org/10.5281/zenodo.12749619" httpUri="https://zenodo.org/record/12749619/files/figure.png" pageId="11" pageNumber="252">Figs 7A–C</figureCitation>
).
</paragraph>
<paragraph id="8B9C2156D311FFA786DB7C1456AD6B75" blockId="11.[151,1437,151,933]" pageId="11" pageNumber="252">
Colouration in ethanol. Dorsal shield of prosoma yellowish-brown to reddish-brown, with bright transversal crescent submarginally at posterior margin and large area covered with dark setae and few lighter patches especially submarginally and close to eyes. Chelicerae deep reddish-brown. Labium, sternum and gnathocoxae deep yellowish-brown. Opisthosoma dorsally brown, with pairs of dark patches and dark inverse V-shaped pattern in posterior half; ventrally yellowish-brown, with slightly darker elongated V-shaped lines and a smaller dark Vshaped pattern in front of spinnerets (
<figureCitation id="13183DD3D311FFA784237F5856D96B75" box="[575,703,547,573]" captionStart="FIGURES 7" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[177,1411,181,1885]" captionTargetId="figure-24@10.[177,1411,181,1885]" captionTargetPageId="10" captionText="FIGURES 7A–E. Heteropoda papilionacea spec. nov., holotype female from Malaysia, Sabah, Maliau Basin Conservation Area. A–C Copulatory organ (A vulva, dorsal; B epigyne, ventral; C schematic course of internal duct system, dorsal); D–E Habitus (D dorsal, E ventral).Abbreviations:AB—anterior bands; CO—copulatory opening; FW—first winding; GP—glandular pores; LL—lateral lobes; MS—median septum. Scale bars: A–C 0.5 mm; D–E 3 mm." figureDoi="http://doi.org/10.5281/zenodo.12749619" httpUri="https://zenodo.org/record/12749619/files/figure.png" pageId="11" pageNumber="252">Figs 7D–E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33972DDD311FFA786DB7F3D55556AED" pageId="11" pageNumber="252" type="distribution">
<paragraph id="8B9C2156D311FFA786DB7F3D56BF6B29" blockId="11.[151,1437,151,933]" box="[199,729,582,609]" pageId="11" pageNumber="252">
<emphasis id="B957FD44D311FFA786DB7F3D55046B28" bold="true" box="[199,354,582,608]" pageId="11" pageNumber="252">Distribution.</emphasis>
<collectingCountry id="F33461C6D311FFA787757F3C55B76B29" box="[361,465,583,609]" name="Malaysia" pageId="11" pageNumber="252">Malaysia</collectingCountry>
(
<collectingRegion id="49E7EFB4D311FFA787FC7F3C560E6B29" box="[480,616,583,609]" country="Malaysia" name="Sabah" pageId="11" pageNumber="252">Sabah State</collectingRegion>
) (
<figureCitation id="13183DD3D311FFA7849C7F3C56B76B29" box="[640,721,583,609]" captionStart="FIGURE 11" captionStartId="16.[151,250,1327,1351]" captionTargetBox="[191,1395,181,1304]" captionTargetId="figure-24@16.[191,1395,181,1304]" captionTargetPageId="16" captionText="FIGURE 11. Locality records of the examined Heteropoda material." figureDoi="http://doi.org/10.5281/zenodo.12749629" httpUri="https://zenodo.org/record/12749629/files/figure.png" pageId="11" pageNumber="252">Fig. 11</figureCitation>
)
</paragraph>
<paragraph id="8B9C2156D311FFA786DB7F1055556AED" blockId="11.[151,1437,151,933]" pageId="11" pageNumber="252">
<emphasis id="B957FD44D311FFA786DB7F1055746BCD" bold="true" box="[199,274,619,645]" pageId="11" pageNumber="252">Notes.</emphasis>
Although the male of
<taxonomicName id="4C235AD5D311FFA7840C7F1056FD6BCD" authorityName="Korai &amp; Jäger" authorityYear="2024" box="[528,667,619,645]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="verticalis" status="spec. nov.">
<emphasis id="B957FD44D311FFA7840C7F1056FD6BCD" box="[528,667,619,645]" italics="true" pageId="11" pageNumber="252">H. verticalis</emphasis>
</taxonomicName>
<emphasis id="B957FD44D311FFA784BF7F1057716BCD" bold="true" box="[675,791,619,645]" pageId="11" pageNumber="252">
<taxonomicNameLabel id="A264403FD311FFA784BF7F1057716BCD" box="[675,791,619,645]" pageId="11" pageNumber="252" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is occurring syntopically it is considered not conspecific with the present species due to considerable differences in size and colouration pattern. The body length of
<taxonomicName id="4C235AD5D311FFA7839D7FF455636B85" authorityName="Korai &amp; Jäger" authorityYear="2024" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="cuspidata" status="spec. nov.">
<emphasis id="B957FD44D311FFA7839D7FF455636B85" italics="true" pageId="11" pageNumber="252">H. cuspidata</emphasis>
</taxonomicName>
<emphasis id="B957FD44D311FFA7870D7FC855EC6B85" bold="true" box="[273,394,691,717]" pageId="11" pageNumber="252">
<taxonomicNameLabel id="A264403FD311FFA7870D7FC855EC6B85" box="[273,394,691,717]" pageId="11" pageNumber="252" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is considerably larger than
<taxonomicName id="4C235AD5D311FFA784C07FC857FE6B85" authorityName="Korai &amp; Jäger" authorityYear="2024" box="[732,920,691,717]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="papilionacea" status="spec. nov.">
<emphasis id="B957FD44D311FFA784C07FC857FE6B85" box="[732,920,691,717]" italics="true" pageId="11" pageNumber="252">H. papilionacea</emphasis>
</taxonomicName>
<emphasis id="B957FD44D311FFA785B87FC8507A6B85" bold="true" box="[932,1052,691,717]" pageId="11" pageNumber="252">
<taxonomicNameLabel id="A264403FD311FFA785B87FC8507A6B85" box="[932,1052,691,717]" pageId="11" pageNumber="252" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(usually males are smaller than conspecific females), and
<taxonomicName id="4C235AD5D311FFA787A37FAC56296BB9" authorityName="Korai &amp; Jäger" authorityYear="2024" box="[447,591,727,753]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="cuspidata" status="spec. nov.">
<emphasis id="B957FD44D311FFA787A37FAC56296BB9" box="[447,591,727,753]" italics="true" pageId="11" pageNumber="252">H. cuspidata</emphasis>
</taxonomicName>
<emphasis id="B957FD44D311FFA7844B7FAC56AC6BB9" bold="true" box="[599,714,727,753]" pageId="11" pageNumber="252">
<taxonomicNameLabel id="A264403FD311FFA7844B7FAC56AC6BB9" box="[599,714,727,753]" pageId="11" pageNumber="252" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
exhibits a dark ventral side on both, prosoma and opisthosoma, whereas
<taxonomicName id="4C235AD5D311FFA786E07F8055D26A5D" authorityName="Korai &amp; Jäger" authorityYear="2024" box="[252,436,763,789]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="papilionacea" status="spec. nov.">
<emphasis id="B957FD44D311FFA786E07F8055D26A5D" box="[252,436,763,789]" italics="true" pageId="11" pageNumber="252">H. papilionacea</emphasis>
</taxonomicName>
<emphasis id="B957FD44D311FFA787A07F8056566A5D" bold="true" box="[444,560,763,789]" pageId="11" pageNumber="252">
<taxonomicNameLabel id="A264403FD311FFA787A07F8056566A5D" box="[444,560,763,789]" pageId="11" pageNumber="252" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is light yellowish-brown. Sexual colour dimorphisms known to occur in the genus
<taxonomicName id="4C235AD5D311FFA786FF7E64550F6A71" authorityName="Latreille" authorityYear="1804" box="[227,361,799,825]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="B957FD44D311FFA786FF7E64550F6A71" box="[227,361,799,825]" italics="true" pageId="11" pageNumber="252">Heteropoda</emphasis>
</taxonomicName>
are restricted to dorsal appearances, especially the dorsal prosoma. It is known that several
<taxonomicName id="4C235AD5D311FFA7868B7E38551E6A15" box="[151,376,835,861]" pageId="11" pageNumber="252">
<emphasis id="B957FD44D311FFA7868B7E38557B6A15" box="[151,285,835,861]" italics="true" pageId="11" pageNumber="252">Heteropoda</emphasis>
species
</taxonomicName>
as larger invertebrate predators can occur syntopically in one habitat (e.g. in southern
<collectingCountry id="F33461C6D311FFA783797E3851FA6A15" box="[1381,1436,835,861]" name="Laos" pageId="11" pageNumber="252">Laos</collectingCountry>
four
<taxonomicName id="4C235AD5D311FFA786C87E1C55FF6AC9" box="[212,409,871,897]" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B957FD44D311FFA786C87E1C553C6AC9" box="[212,346,871,897]" italics="true" pageId="11" pageNumber="252">Heteropoda</emphasis>
spp.
</taxonomicName>
, together with additional
<taxonomicName id="4C235AD5D311FFA784C87E1C573C6AC9" authorityName="Bertkau" authorityYear="1872" box="[724,858,871,897]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="252" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
like
<taxonomicName id="4C235AD5D311FFA785837E1C503D6AC9" box="[927,1115,871,897]" class="Arachnida" family="Sparassidae" genus="Pseudopoda" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B957FD44D311FFA785837E1C504D6AC9" box="[927,1067,871,897]" italics="true" pageId="11" pageNumber="252">Pseudopoda</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="4C235AD5D311FFA782727E1C51286AC9" box="[1134,1358,871,897]" class="Arachnida" family="Sparassidae" genus="Gnathopalystes" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B957FD44D311FFA782727E1C51786AC9" box="[1134,1310,871,897]" italics="true" pageId="11" pageNumber="252">Gnathopalystes</emphasis>
sp.
</taxonomicName>
, Jäger unpublished).
</paragraph>
</subSubSection>
</treatment>
</document>
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