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<document id="C283B360207EBFE8B8FF5FE3BECFB6D4" ID-CLB-Dataset="218234" ID-DOI="10.1206/0003-0090.458.1.1" ID-GBIF-Dataset="9aaee2d0-a89e-47d4-8578-cdd525f7ebfb" ID-ISSN="0003-0090" ID-Zenodo-Dep="7400026" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1670242945939" checkinUser="felipe" docAuthor="Jones, Lance E. &amp; Grimaldi, David A." docDate="2022" docId="3C6787EDFFB5FFEE60E2FBECFEE4FDAB" docLanguage="en" docName="BAmMusNatHist.2022.458.1-177.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (458)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-458/issue-1/0003-0090.458.1.1/Revision-of-the-Nearctic-Species-of-the-Genus-Amiota-Loew/10.1206/0003-0090.458.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Amiota multiplex Jones &amp; Grimaldi 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="C05EFF95FFF4FFAC6000FFF0FFEDFF88" masterDocTitle="Revision Of The Nearctic Species Of The Genus Amiota Loew (Diptera: Drosophilidae)" masterLastPageNumber="181" masterPageNumber="1" pageNumber="65" updateTime="1699413003214" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D41DAD5A35D9EF3EEC78C0CE96C80468">Revision Of The Nearctic Species Of The Genus Amiota Loew (Diptera: Drosophilidae)</mods:title>
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<mods:namePart id="157D274D122CF9A90F55690492B886B0">Jones, Lance E.</mods:namePart>
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<mods:namePart id="EA3F31AA39CFC758147C0C18676D6D9D">Grimaldi, David A.</mods:namePart>
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<treatment id="3C6787EDFFB5FFEE60E2FBECFEE4FDAB" ID-DOI="http://doi.org/10.5281/zenodo.7506525" ID-GBIF-Taxon="208565084" ID-Zenodo-Dep="7506525" LSID="urn:lsid:plazi:treatment:3C6787EDFFB5FFEE60E2FBECFEE4FDAB" httpUri="http://treatment.plazi.org/id/3C6787EDFFB5FFEE60E2FBECFEE4FDAB" lastPageId="66" lastPageNumber="66" pageId="65" pageNumber="65">
<subSubSection id="FCD46570FFB5FFED60E2FBECFDEAFBBC" box="[226,519,1052,1076]" pageId="65" pageNumber="65" type="nomenclature">
<paragraph id="B47136FBFFB5FFED60E2FBECFDEAFBBC" blockId="65.[226,519,1052,1076]" box="[226,519,1052,1076]" pageId="65" pageNumber="65">
<heading id="EF398197FFB5FFED60E2FBECFDEAFBBC" box="[226,519,1052,1076]" centered="true" fontSize="9" level="2" pageId="65" pageNumber="65" reason="2">
<emphasis id="86BAEAE9FFB5FFED60E2FBECFE40FBBC" bold="true" box="[226,429,1052,1076]" pageId="65" pageNumber="65">
<taxonomicName id="73CE4D78FFB5FFED60E2FBECFE4BFBBC" ID-CoL="BDJJ6" authority="Jones &amp; Grimaldi, 2022" authorityName="Jones &amp; Grimaldi" authorityYear="2022" box="[226,422,1052,1076]" class="Insecta" family="Drosophilidae" genus="Amiota" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="65" phylum="Arthropoda" rank="species" species="multiplex" status="sp. nov.">
<emphasis id="86BAEAE9FFB5FFED60E2FBECFE4BFBBC" bold="true" box="[226,422,1052,1076]" italics="true" pageId="65" pageNumber="65">Amiota multiplex</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="9D895792FFB5FFED61B4FBEDFDEAFBBC" box="[436,519,1053,1076]" pageId="65" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFB5FFED60CEFBBEFDF6FBEE" box="[206,539,1102,1126]" pageId="65" pageNumber="65" type="description">
<paragraph id="B47136FBFFB5FFED60CEFBBEFDF6FBEE" blockId="65.[206,539,1102,1126]" box="[206,539,1102,1126]" pageId="65" pageNumber="65">
<heading id="EF398197FFB5FFED60CEFBBEFDF6FBEE" box="[206,539,1102,1126]" centered="true" fontSize="9" level="2" pageId="65" pageNumber="65" reason="2">
<figureCitation id="2CF52A7EFFB5FFED60CEFBBEFE86FBEE" box="[206,363,1102,1126]" captionStart="FIG" captionStartId="67.[109,151,1682,1703]" captionTargetBox="[108,1212,224,1652]" captionTargetId="figure-12@67.[108,1212,224,1652]" captionTargetPageId="67" captionText="FIG. 29. Heads and lateral views, A. cervites species group. AB. A. cervites, sp. nov. (Am 1526, holotype). CD. A. elsaltoensis, sp. nov. (Am 1455, holotype). EF. A. multiplex, sp. nov. (Am 1552, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7400084" httpUri="https://zenodo.org/record/7400084/files/figure.png" pageId="65" pageNumber="65">Figures 29EF</figureCitation>
,
<figureCitation id="2CF52A7EFFB5FFED6176FBBEFE51FBEE" box="[374,444,1102,1126]" captionStart="FIG" captionStartId="68.[109,151,1525,1546]" captionTargetBox="[236,1072,225,1496]" captionTargetId="figure-11@68.[236,1081,224,1496]" captionTargetPageId="68" captionText="FIG. 30. Aristae, A. cervites species group. AB. A. cervites. CD. A. elsaltoensis. EF. A. multiplex. Not the same scale." figureDoi="http://doi.org/10.5281/zenodo.7400086" httpUri="https://zenodo.org/record/7400086/files/figure.png" pageId="65" pageNumber="65">30EF</figureCitation>
,
<figureCitation id="2CF52A7EFFB5FFED61C6FBBEFE0FFBEE" box="[454,482,1102,1126]" captionStart="FIG" captionStartId="71.[108,150,1594,1615]" captionTargetBox="[201,1134,233,1556]" captionTargetId="figure-41@71.[181,1138,224,1565]" captionTargetPageId="71" captionText="FIG. 33. Male terminalia, A. multiplex. A. Epandrium and cerci. B. Lateral view. C. Aedeagal apodeme. D. Paraphyses, lateral view. E. Posterior view. F. Surstylus. (Am 1552, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7400092" httpUri="https://zenodo.org/record/7400092/files/figure.png" pageId="65" pageNumber="65">33</figureCitation>
,
<figureCitation id="2CF52A7EFFB5FFED61EEFBBEFDF6FBEE" box="[494,539,1102,1126]" captionStart="FIG" captionStartId="160.[108,150,1676,1697]" captionTargetBox="[107,1206,224,1647]" captionTargetId="figure-11@160.[107,1206,224,1647]" captionTargetPageId="160" captionText="FIG. 86. Distribution maps, A. cervites species group. A. A. cervites. B. A. elsaltoensis. C. A. multiplex." figureDoi="http://doi.org/10.5281/zenodo.7400209" httpUri="https://zenodo.org/record/7400209/files/figure.png" pageId="65" pageNumber="65">86C</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFB5FFED608CFB70FE1DFAF7" pageId="65" pageNumber="65" type="diagnosis">
<paragraph id="B47136FBFFB5FFED608CFB70FE1DFAF7" blockId="65.[108,637,1151,1738]" pageId="65" pageNumber="65">
DIAGNOSIS: Large fly (ThL
<quantity id="73369B1EFFB5FFED61BFFB8FFDCBFB10" box="[447,550,1151,1176]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="65" pageNumber="65" unit="mm" value="1.65">1.65 mm</quantity>
), black; frons silvery on ventral half, especially ventrolateral corners; epandrium tall, dorsally complete, cerci situated low; outer paraphysis heavily sclerotized, consisting of 2 lobes with 5 pairs of narrow, curved spines; inner paraphyses small, clavate, distal end setose, covered in 68 fine setulae; ejaculatory apodeme small.
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFB5FFED608CFA78FBCCF9AC" pageId="65" pageNumber="65" type="description">
<paragraph id="B47136FBFFB5FFED608CFA78FBCCF9AC" blockId="65.[108,637,1151,1738]" lastBlockId="65.[684,1212,226,1738]" pageId="65" pageNumber="65">
DESCRIPTION: Large fly (ThL
<quantity id="73369B1EFFB5FFED61FFFA78FD83FA28" box="[511,622,1416,1440]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="65" pageNumber="65" unit="mm" value="1.65">1.65 mm</quantity>
), black, legs dark yellow. Frons dark and rather shimmering, especially at ventrolateral corners. Cheek somewhat deep for size (EL/CW 9.25); anterior half dark yellow, posterior half brown. Palp yellow. Arista: Very short, plumose, branches decumbent; longest branch D1; A.R. 0.24; 3 dorsal, 0 ventral branches; branch D3 pointed mediad; arista trunk with medium-length microtrichia nearly to apex. Male genitalia: Epandrium tall, height ~1.25× width; fused at midline, not grading into surrounding membrane; ventral lobes setose, line of setae along medial margin of epandrium. Cercus position low, with large triangularshaped membranous region above; pendulous, not grading into surrounding membrane; slightly narrowed ventrally. Surstylus wide at base, tapered distally; ventral half with very long, fine scattered setulae; 8 short prensisetae, apices blunt. Subepandrial sclerite rounded, U-shaped with an acute, mucronate point; inner margin thickened; subepandrial appendage not well developed. Outer paraphysis heavily sclerotized; consisting of 2 large lobes with a thickened lip on posterior margin; 5 pairs of zygomorphic appendages; most apical pair slender, sinuous, pointed posteriorly; second pair arising medially to apical pair, oriented laterally; third pair ventral to apex, appendages curved, oriented posteriorly; fourth pair arising proximally, curved downward, oriented ventrally; fifth pair most proximal, sinuous, slender, oriented anteriorly; 2 appendages arising between fourth and fifth pairs, each oriented laterally, crossing midline. Inner paraphysis clavate, slender; distal end expanded to rounded pad, setose, with 68 fine setulae. Aedeagal apodeme nearly bent at 90° angle; basal region with widely flared flanges; middle region constricted; distal end round, with a minute medial cleft running to the midpoint, but no significant emargination. Hypandrium U-shaped, width consistent; apex notched, accommodating paraphyses; lateral arm with a small, thickened swelling. Ejaculatory apodeme very small, 0.25× length of the epandrium, barely sclerotized. Head and thorax measurements: (n = 1, Am 1552) FL/FW 0.75, EL/EW 1.14, EL/CW 9.25, FML/FMW 0.31,
<collectingRegion id="760AF819FFB5FFED62EAF9FDFCE6F9AD" box="[746,779,1549,1573]" country="Brazil" name="Para" pageId="65" pageNumber="65">PR</collectingRegion>
/RR 0.60, ThL 1.65 mm.
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFB5FFEE62CCF9DEFED8FE93" lastPageId="66" lastPageNumber="66" pageId="65" pageNumber="65" type="materials_examined">
<paragraph id="B47136FBFFB5FFED62CCF9DEFB55F942" blockId="65.[684,1212,226,1738]" pageId="65" pageNumber="65">
<collectionCode id="D2DFAE3EFFB5FFED62CCF9DEFC90F9CE" box="[716,893,1582,1606]" pageId="65" pageNumber="65">TYPE MATERIAL</collectionCode>
:
<materialsCitation id="04A63CA6FFB5FFED6387F9DEFB55F942" ID-GBIF-Occurrence="4010926761" collectingDate="1964-06-13" collectionCode="CNC" collectorName="J. F. McAlpine" country="Mexico" latitude="23.7851" location="Durango" longLatPrecision="1" longitude="-105.53662" pageId="65" pageNumber="65" specimenCount="1" stateProvince="Durango" typeStatus="holotype">
<emphasis id="86BAEAE9FFB5FFED6387F9DEFC1BF9CE" bold="true" box="[903,1014,1582,1606]" pageId="65" pageNumber="65">
<typeStatus id="6B758859FFB5FFED6387F9DEFC1CF9CE" box="[903,1009,1582,1606]" pageId="65" pageNumber="65" type="holotype">Holotype</typeStatus>
:
</emphasis>
male: 11 mi.
<collectorName id="193B532DFFB5FFED6489F9DEFD0BF9EF" pageId="65" pageNumber="65">E. El Salto</collectorName>
, Dgo. [
<collectingRegion id="760AF819FFB5FFED6352F9A0FC53F9E0" box="[850,958,1616,1640]" country="Mexico" name="Durango" pageId="65" pageNumber="65">Durango</collectingRegion>
]
<collectingCountry id="CCD9766BFFB5FFED63DAF9A0FBF6F9EF" box="[986,1051,1616,1639]" name="Mexico" pageId="65" pageNumber="65">MEX</collectingCountry>
., [
<geoCoordinate id="D1FA503CFFB5FFED643EF9BFFB5AF9EF" box="[1086,1207,1615,1639]" degrees="23.785100" direction="north" orientation="latitude" pageId="65" pageNumber="65" precision="1" value="23.7851">23.785100</geoCoordinate>
,
<geoCoordinate id="D1FA503CFFB5FFED62ACF980FCD9F900" box="[684,820,1648,1672]" degrees="105.536622" direction="west" orientation="longitude" pageId="65" pageNumber="65" precision="1" value="-105.53662">-105.536622</geoCoordinate>
],
<date id="C070103BFFB5FFED6344F981FC38F900" box="[836,981,1648,1672]" pageId="65" pageNumber="65" value="1964-06-13">
<collectingDate id="D034E9D3FFB5FFED6344F981FC38F900" box="[836,981,1648,1672]" pageId="65" pageNumber="65" value="1964-06-13">June 13, 1964</collectingDate>
</date>
,
<collectorName id="193B532DFFB5FFED63DFF981FB9DF900" box="[991,1136,1648,1672]" pageId="65" pageNumber="65">J.F. McAlpine</collectorName>
, 8000,
<collectorName id="193B532DFFB5FFED62ACF961FD38F921" box="[684,725,1681,1705]" pageId="65" pageNumber="65">Am</collectorName>
1552, [pinned specimen, dissected].
<collectorName id="193B532DFFB5FFED646DF962FD39F942" pageId="65" pageNumber="65">Deposited</collectorName>
in the
<collectorName id="193B532DFFB5FFED6321F942FB8CF942" box="[801,1121,1714,1738]" pageId="65" pageNumber="65">Canadian National Collection</collectorName>
(
<collectionCode id="D2DFAE3EFFB5FFED646FF942FB41F942" box="[1135,1196,1714,1738]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="65" pageNumber="65">CNC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="B47136FBFFB6FFEE609CFF11FED8FE93" blockId="66.[108,636,225,548]" pageId="66" pageNumber="66">
<collectionCode id="D2DFAE3EFFB6FFEE609CFF11FE05FF71" box="[156,488,225,249]" pageId="66" pageNumber="66">OTHER MATERIAL EXAMINED</collectionCode>
: Known only from the
<typeStatus id="6B758859FFB6FFEE60D1FEF3FEDCFE93" box="[209,305,259,283]" pageId="66" pageNumber="66" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFB6FFEE608CFED5FF0BFEF7" pageId="66" pageNumber="66" type="etymology">
<paragraph id="B47136FBFFB6FFEE608CFED5FF0BFEF7" blockId="66.[108,636,225,548]" pageId="66" pageNumber="66">ETYMOLOGY: From Latin, meaning “complex, multilayered,” in reference to the outer paraphysis.</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFB6FFEE608CFE78FE14FE49" pageId="66" pageNumber="66" type="distribution">
<paragraph id="B47136FBFFB6FFEE608CFE78FE14FE49" blockId="66.[108,636,225,548]" pageId="66" pageNumber="66">
DISTRIBUTION:
<taxonomicName id="73CE4D78FFB6FFEE6141FE78FDE8FE28" authorityName="Jones &amp; Grimaldi" authorityYear="2022" box="[321,517,392,416]" class="Insecta" family="Drosophilidae" genus="Amiota" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="66" phylum="Arthropoda" rank="species" species="multiplex">
<emphasis id="86BAEAE9FFB6FFEE6141FE78FDE8FE28" box="[321,517,392,416]" italics="true" pageId="66" pageNumber="66">Amiota multiplex</emphasis>
</taxonomicName>
is known only from
<collectingRegion id="760AF819FFB6FFEE60E0FE59FE90FE49" box="[224,381,425,449]" country="Mexico" name="Durango" pageId="66" pageNumber="66">Durango State</collectingRegion>
in
<collectingRegion id="760AF819FFB6FFEE61A2FE59FE18FE49" box="[418,501,425,449]" country="Mexico" name="Mexico" pageId="66" pageNumber="66">Mexico</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFB6FFEE608CFE3AFEE4FDAB" pageId="66" pageNumber="66" type="discussion">
<paragraph id="B47136FBFFB6FFEE608CFE3AFEE4FDAB" blockId="66.[108,636,225,548]" pageId="66" pageNumber="66">
COMMENTS: It is unknown whether this species is attracted to the face and eyes as other
<taxonomicName id="73CE4D78FFB6FFEE60B1FDFCFEEEFDAB" box="[177,259,524,547]" class="Insecta" family="Drosophilidae" genus="Amiota" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="86BAEAE9FFB6FFEE60B1FDFCFEEEFDAB" box="[177,259,524,547]" italics="true" pageId="66" pageNumber="66">Amiota</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>