treatments-xml/data/3C/E3/B4/3CE3B48E8A328FFAF621E885FDBE1638.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.769.25258" ID-GBIF-Dataset="7b27f7ff-2920-4834-b08c-754a4c0327c8" ID-PMC="PMC6030179" ID-Pensoft-Pub="1313-2970-769-13" ID-Pensoft-UUID="9C3DFFE3003056499148056FFF8CFFBF" ID-PubMed="29988743" ID-Zenodo-Dep="1304503" ID-ZooBank="C31B0F6BD3C242CDBAED8CE9D5769E8A" ModsDocID="1313-2970-769-13" checkinTime="1530090487889" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander &amp; Feher, Zoltan" docDate="2018" docId="3CE3B48E8A328FFAF621E885FDBE1638" docLanguage="en" docName="ZooKeys 769: 13-48" docOrigin="ZooKeys 769" docPubDate="2018-06-26" docSource="http://dx.doi.org/10.3897/zookeys.769.25258" docTitle="Spelaeodiscus virpazarioides Pall-Gergely &amp; Feher 2018, sp. n." docType="treatment" docUuid="6454FD89-D6C1-4607-9992-7F1CC7DBD617" docUuidSource="ZooBank" docVersion="4" id="9C3DFFE3003056499148056FFF8CFFBF" lastPageNumber="13" masterDocId="9C3DFFE3003056499148056FFF8CFFBF" masterDocTitle="Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae)" masterLastPageNumber="48" masterPageNumber="13" pageNumber="13" updateTime="1668165931740" updateUser="ExternalLinkService">
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<mods:title>Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Hungarian Academy of Sciences (MTA), Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deli, Tamas</mods:namePart>
<mods:affiliation>Moricz Zsigmond u. 2, Gyomaendrod, H- 5500, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eross, Zoltan Peter</mods:namePart>
<mods:affiliation>Bem u. 36., Budapest, H- 1151, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Peter L.</mods:namePart>
<mods:affiliation>Puechhaimg. 52, A- 3580, Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimg. 52, A- 3580, Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Feher, Zoltan</mods:namePart>
<mods:affiliation>Department of Zoology, Hungarian Natural History Museum, Baross u. 13, H- 1088, Hungary</mods:affiliation>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-06-26</mods:number>
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<mods:detail type="volume">
<mods:number>769</mods:number>
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<mods:extent unit="page">
<mods:start>13</mods:start>
<mods:end>48</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.769.25258</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.769.25258</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1304503</mods:identifier>
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<subSection pageId="0" pageNumber="13" type="systematics">
<treatment ID-GBIF-Taxon="182229765" LSID="urn:lsid:zoobank.org:act:6454FD89-D6C1-4607-9992-7F1CC7DBD617" httpUri="http://treatment.plazi.org/id/3CE3B48E8A328FFAF621E885FDBE1638" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection pageId="0" pageNumber="13" type="nomenclature">
<paragraph pageId="0" pageNumber="13">
<taxonomicName LSID="http://zoobank.org/6454FD89-D6C1-4607-9992-7F1CC7DBD617" authority="Páll-Gergely &amp; Fehér" authorityName="Pall-Gergely &amp; Feher" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Spelaeodiscus virpazarioides" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="virpazarioides" status="sp. n.">
Spelaeodiscus virpazarioides 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; 
<normalizedToken originalValue="Fehér">Feher</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="13">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Shells of Spelaeodiscus. A-F Spelaeodiscus unidentatus acutus ssp. n., holotype (HNHM 103252) G-L Spelaeodiscus unidentatus acutus ssp. n., Albania, rocks north of Hajmel at the bridge (NHMW 112371) M-R Spelaeodiscus virpazarioides sp. n., holotype (HNHM 103417)." figureDoi="10.3897/zookeys.769.25258.figure14" httpUri="https://binary.pensoft.net/fig/211848" pageId="0" pageNumber="13">Figure 14M-R</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="type material">
<paragraph pageId="0" pageNumber="13">Type material.</paragraph>
<paragraph pageId="0" pageNumber="13">
Albania, 
<normalizedToken originalValue="Malësia">Malesia</normalizedToken>
district, a mountain pass 2 km N of 
<normalizedToken originalValue="Rraps-Starjë">Rraps-Starje</normalizedToken>
, 700 m a.s.l., 
<geoCoordinate degrees="42" direction="north" minutes="24.888" orientation="latitude" precision="1" value="42.4148">42°24.888'N</geoCoordinate>
, 
<geoCoordinate degrees="19" direction="east" minutes="30.240" orientation="longitude" precision="1" value="19.504">19°30.240'E</geoCoordinate>
, leg. 
<normalizedToken originalValue="Erőss">Eross</normalizedToken>
, 
<normalizedToken originalValue="Fehér">Feher</normalizedToken>
, Szekeres, Grego, 27.06.2016, HNHM 103417 (holotype, SW = 3.45 mm, SH = 2.0 mm), HNHM 102765/4+1subadult+3fr (paratypes), NHMW 111672/3+2subadult+2fr; JG/3+2subadult+3fr (paratypes); same locality, leg. A. 
<normalizedToken originalValue="Reischütz">Reischuetz</normalizedToken>
, N. 
<normalizedToken originalValue="Reischütz">Reischuetz</normalizedToken>
&amp; P. 
<normalizedToken originalValue="Reischütz">Reischuetz</normalizedToken>
, Apr. 2006, REI/1 juvenile paratype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="type locality">
<paragraph pageId="0" pageNumber="13">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="13">
Albania, 
<normalizedToken originalValue="Malësia">Malesia</normalizedToken>
district, a mountain pass 2 km N of 
<normalizedToken originalValue="Rraps-Starjë">Rraps-Starje</normalizedToken>
, 700 m a.s.l., 
<geoCoordinate degrees="42" direction="north" minutes="24.888" orientation="latitude" precision="1" value="42.4148">42°24.888'N</geoCoordinate>
, 
<geoCoordinate degrees="19" direction="east" minutes="30.240" orientation="longitude" precision="1" value="19.504">19°30.240'E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="diagnosis">
<paragraph pageId="0" pageNumber="13">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="13">A medium sized to large species with elevated spire, strong, very widely spaced ribs, fine spiral lines consisting of series of minute tubercles, hammered protoconch, weak basal thickening, weak parietal tooth, and a thickened parietal callus.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="description">
<paragraph pageId="0" pageNumber="13">Description.</paragraph>
<paragraph pageId="0" pageNumber="13">
Spire elevated; protoconch consists of ca. 1.25-1.75 whorls, roughly sculptured, 
<normalizedToken originalValue="“hammered”">&quot;hammered&quot;</normalizedToken>
; teleoconch with very strong, equidistant, widely spaced ribs (40-70 on the body whorl; addition to the radial ribs very fine, dense spiral striation is visible on the entire teleoconch, consisting of minute tubercles; entire shell with 4-4.75 whorls; aperture subcircular/triangular; basal part with two low swellings (similar to those of 
<taxonomicName authority="albanicus" family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. albanicus subsp. albanicus" order="Stylommatophora" pageId="0" pageNumber="13" rank="subSpecies" species="albanicus" subSpecies="albanicus">
<emphasis italics="true" pageId="0" pageNumber="13">S. albanicus albanicus</emphasis>
</taxonomicName>
), parietal wall with a weak tooth; parietal callus thickened, blunt; umbilicus funnel-shaped, relatively narrow.
</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Measurements.</emphasis>
SW: 3.3-3.6 mm (median = 3.5 mm), SH: 1.95-2.25 mm (median = 2.1 mm), AW: 1.3-1.6 mm (median = 1.4 mm), SW: 1.4-1.5 mm (median = 1.4 mm), (n = 11).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="differential diagnosis">
<paragraph pageId="0" pageNumber="13">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="13">
This new species can be distinguished from all congeners (especially the most similar 
<taxonomicName authority="albanicus" family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. albanicus subsp. albanicus" order="Stylommatophora" pageId="0" pageNumber="13" rank="subSpecies" species="albanicus" subSpecies="albanicus">
<emphasis italics="true" pageId="0" pageNumber="13">S. albanicus albanicus</emphasis>
</taxonomicName>
) by the clearly visible spiral striation, roughly sculptured protoconch and thickened parietal callus. All 
<taxonomicName authorityName="E. Gittenberger" authorityYear="1969" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Virpazaria" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Virpazaria</emphasis>
</taxonomicName>
species possess an elevated parietal callus, but it is sharp in most (all?) species, and the aperture of those species are more slender, crescent-shaped.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="variation">
<paragraph pageId="0" pageNumber="13">Variation among specimens.</paragraph>
<paragraph pageId="0" pageNumber="13">The degree of the thickness of the parietal callus shows some recognisable variability within the single known population, but this trait might be due to the age (degree of development) of the examined shells.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="etymology">
<paragraph pageId="0" pageNumber="13">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="13">
This new species is named after its resemblance to 
<taxonomicName authorityName="E. Gittenberger" authorityYear="1969" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Virpazaria" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Virpazaria</emphasis>
</taxonomicName>
species based on the thickened parietal callus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="distribution">
<paragraph pageId="0" pageNumber="13">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="13">
<taxonomicName authorityName="Pall-Gergely &amp; Feher" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus virpazarioides" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="virpazarioides">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus virpazarioides</emphasis>
</taxonomicName>
sp. n. is known from the type locality only (see also Figure 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution of Spelaeodiscus Brusina, 1886 species." figureDoi="10.3897/zookeys.769.25258.figure7" httpUri="https://binary.pensoft.net/fig/211841" pageId="0" pageNumber="13">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="remarks">
<paragraph pageId="0" pageNumber="13">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="13">
Initially we considered placing this species to the genus 
<taxonomicName authorityName="E. Gittenberger" authorityYear="1969" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Virpazaria" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Virpazaria</emphasis>
</taxonomicName>
due to the thickened parietal callus, which was mentioned in the original description of the genus 
<taxonomicName authorityName="E. Gittenberger" authorityYear="1969" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Virpazaria" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Virpazaria</emphasis>
</taxonomicName>
. However, the subcircular/triangular aperture indicates that it is better to be placed in 
<taxonomicName authorityName="Páll-Gergely &amp; Deli &amp; Erőss &amp; Reischütz &amp; Reischütz &amp; Fehér" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="13">
Formerly, this population was incorrectly referred to as 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. albanicus" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="albanicus">
<emphasis italics="true" pageId="0" pageNumber="13">S. albanicus</emphasis>
</taxonomicName>
and was taken into consideration in that 
<normalizedToken originalValue="species’">species'</normalizedToken>
Red List assessment (
<bibRefCitation DOI="http://dx.doi.org/10.2305/IUCN.UK.2017-3.RLTS.T170947A1319076.en" author="Reischuetz, A" journalOrPublisher="Conchological Section, Academy of Natural Sciences, Philadelphia" pageId="0" pageNumber="13" refId="B32" refString="Reischuetz, A, Feher, Z, 2017. Spelaeodiscus albanicus. The IUCN Red List of Threatened Species 2017: e.T170947A1319076. http://dx.doi.org/10.2305/IUCN.UK.2017-3.RLTS.T170947A1319076.en" title="Spelaeodiscus albanicus. The IUCN Red List of Threatened Species 2017: e. T 170947 A 1319076." url="http://dx.doi.org/10.2305/IUCN.UK.2017-3.RLTS.T170947A1319076.en" year="2017">
<normalizedToken originalValue="Reischütz">Reischuetz</normalizedToken>
and 
<normalizedToken originalValue="Fehér">Feher</normalizedToken>
2017
</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="conservation status">
<paragraph pageId="0" pageNumber="13">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="13">To our present knowledge this species is very rare (currently known from a single location) and thus AOO is smaller than 20 km2. However, there is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number of mature individuals are declining or extremely fluctuating. Therefore, it might be assessed as Near Threatened (NT).</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>