250 lines
44 KiB
XML
250 lines
44 KiB
XML
<document id="18587FDE3673EF3705A85670D7E5915F" ID-DOI="10.11646/zootaxa.5563.1.7" ID-GBIF-Dataset="37f7c974-3d42-490a-ba3a-d446310c2a68" ID-ISSN="1175-5326" ID-Zenodo-Dep="14596228" ID-ZooBank="682A7967-5003-4F6B-8B32-9B3D7FC5A9AF" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1735933280982" checkinUser="plazi" docAuthor="Dunlop, Jason A. & Bartel, Christian" docDate="2025" docId="DA2C4500FFE84325E1C9FC6FFAB46CE8" docLanguage="en" docName="zootaxa.5563.1.7.pdf" docOrigin="Zootaxa 5563 (1)" docSource="https://doi.org/10.11646/zootaxa.5563.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phrynus luisdearmasi Dunlop & Bartel, 2025, sp. nov." docType="treatment" docVersion="6" lastPageNumber="68" masterDocId="26153D78FFE94321E15EFB3AFF9E682D" masterDocTitle="A new species of fossil Phrynus Lamarck, 1801, from Dominican Republic amber (Amblypygi: Phrynidae)" masterLastPageNumber="72" masterPageNumber="64" pageNumber="65" updateTime="1736175398253" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="80CDA44E84C941F58FC59F93137273A1">A new species of fossil Phrynus Lamarck, 1801, from Dominican Republic amber (Amblypygi: Phrynidae)</mods:title>
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<mods:name id="699A54591CDFC076281382763F901507" type="personal">
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<mods:namePart id="94930FDFE616BEC8110129C850403675">Dunlop, Jason A.</mods:namePart>
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<mods:affiliation id="A72658EB458D8CB5B4772A8C7D408AAB">Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany</mods:affiliation>
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<mods:namePart id="F604961AA4CE1DD4E74F23B709B6F1D0">Bartel, Christian</mods:namePart>
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<mods:affiliation id="361CC9CB266D18EE47B1032C85BAAFE0">Natural History Museum Bamberg, Staatliche Naturwissenschaftliche Sammlungen Bayerns, Fleischstrasse 2, 96047 Bamberg, Germany</mods:affiliation>
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<mods:title id="879AFCF166C6D0C84791E1AEC8E17C9C">Zootaxa</mods:title>
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<mods:date id="ADC7269F0FE18EE808F59A04ADC01290">2025</mods:date>
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<treatment id="DA2C4500FFE84325E1C9FC6FFAB46CE8" ID-DOI="http://doi.org/10.5281/zenodo.14596970" ID-GBIF-Taxon="251122255" ID-Zenodo-Dep="14596970" LSID="urn:lsid:plazi:treatment:DA2C4500FFE84325E1C9FC6FFAB46CE8" httpUri="http://treatment.plazi.org/id/DA2C4500FFE84325E1C9FC6FFAB46CE8" lastPageId="4" lastPageNumber="68" pageId="1" pageNumber="65" scope_family="Phrynidae" scope_order="Amblypygi">
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<subSubSection id="1A9FA79DFFE84320E1C9FC6FFD9B6F42" box="[151,517,1877,1904]" pageId="1" pageNumber="65" type="nomenclature">
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<paragraph id="523AF416FFE84320E1C9FC6FFD9B6F42" blockId="1.[151,517,1877,1939]" box="[151,517,1877,1904]" pageId="1" pageNumber="65">
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<heading id="0972437AFFE84320E1C9FC6FFD9B6F42" bold="true" box="[151,517,1877,1904]" fontSize="11" level="1" pageId="1" pageNumber="65" reason="1">
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<emphasis id="60F12804FFE84320E1C9FC6FFD9B6F42" bold="true" box="[151,517,1877,1904]" pageId="1" pageNumber="65">
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<taxonomicName id="95858F95FFE84320E1C9FC6FFE3E6F5D" authorityName="Dunlop & Bartel" authorityYear="2025" box="[151,416,1877,1904]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="luisdearmasi" status="sp. nov.">
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<emphasis id="60F12804FFE84320E1C9FC6FFE3E6F5D" bold="true" box="[151,416,1877,1904]" italics="true" pageId="1" pageNumber="65">Phrynus luisdearmasi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7BC2957FFFE84320E0F9FC6FFD9B6F42" box="[423,517,1877,1903]" pageId="1" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1A9FA79DFFE84320E1C9FC43FF666FBE" box="[151,248,1913,1939]" pageId="1" pageNumber="65" type="description">
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<paragraph id="523AF416FFE84320E1C9FC43FF666FBE" blockId="1.[151,517,1877,1939]" box="[151,248,1913,1939]" pageId="1" pageNumber="65">
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<figureCitation id="CABEE893FFE84320E1C9FC43FF666FBE" box="[151,248,1913,1939]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1051,1075]" captionStartId-1="3.[151,250,1100,1124]" captionStartId-2="3.[151,250,1800,1824]" captionTargetBox-0="[151,1434,181,1027]" captionTargetBox-1="[151,1429,181,1076]" captionTargetBox-2="[151,1419,1328,1776]" captionTargetId-0="figure-17@2.[151,1435,181,1027]" captionTargetId-1="figure-18@3.[151,1435,181,1076]" captionTargetId-2="figure-105@3.[151,1435,1328,1776]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURE 1. Phrynus luisdearmasi sp. nov., MB.A. 1747 (holotype and only known specimen), from Dominican Republic amber (Miocene: Burdigalian). A. Close-up of the anterior body in dorsal view. B. Camera lucida drawing of the dorsal body and legs. C. Frontal view of the chelicerae and pedipalps. D. Camera lucida drawing of the ventral body and legs. Abbreviations: ch, chelicerae; fe, femur; pp, pedipalp; pt, patella; ti, tibia; legs numbered from I–IV. Scale bars equal 2 mm (A), 5 mm (B, D) and 1 mm (C)." captionText-1="FIGURE 2. Phrynus luisdearmasi sp. nov. A. Details of pedipalps in dorsal view. B. Camera lucida drawing of A; dorsal tibial spine 1 arrowed, being larger than the corresponding spine in the previous amber species P. resinae Schawaller, 1979 and its putative extant relative P. marginemaculatus C.L. Koch, 1840. C. Details of pedipalps in ventral view. D. Camera lucida drawing of B. Abbreviations: ch, chelicerae; fe, femur; ley, lateral eyes; mey, median eyes; pp, pedipalp; pt, patella; ti, tibia. Scale bars equal 1 mm." captionText-2="FIGURE 3. Phrynus luisdearmasi sp. nov. A. Detail of the of leg I tibia; note the high number of tibial articles (34). B. Schematic, annotated drawing of the pedipalp’s dorsal spines; large tibial spine 1 differentiating it from the previous amber species again arrowed. Abbreviations: fe, femur; pt, patella; ti, tibia. Scale bars equal 2 mm (A) and 1 mm (B)." figureDoi-0="http://doi.org/10.5281/zenodo.14596232" figureDoi-1="http://doi.org/10.5281/zenodo.14596238" figureDoi-2="http://doi.org/10.5281/zenodo.14596242" httpUri-0="https://zenodo.org/record/14596232/files/figure.png" httpUri-1="https://zenodo.org/record/14596238/files/figure.png" httpUri-2="https://zenodo.org/record/14596242/files/figure.png" pageId="1" pageNumber="65">Figs 1–3</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1A9FA79DFFE84320E1C9FC87FBF56FD6" pageId="1" pageNumber="65" type="etymology">
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<paragraph id="523AF416FFE84320E1C9FC87FBF56FD6" blockId="1.[151,1436,1981,2043]" pageId="1" pageNumber="65">
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<emphasis id="60F12804FFE84320E1C9FC87FE806FFA" bold="true" box="[151,286,1981,2007]" pageId="1" pageNumber="65">Etymology.</emphasis>
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The new species-group name is proposed in honour of Prof. Luis F. de Armas for his extensive contributions towards the study of whip spiders, and other arachnids, in the Neotropics.
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</paragraph>
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</subSubSection>
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<caption id="06FAA49EFFEB4323E1C9FF21FEB56CE9" ID-DOI="http://doi.org/10.5281/zenodo.14596232" ID-Zenodo-Dep="14596232" httpUri="https://zenodo.org/record/14596232/files/figure.png" pageId="2" pageNumber="66" startId="2.[151,250,1051,1075]" targetBox="[151,1434,181,1027]" targetPageId="2" targetType="figure">
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<paragraph id="523AF416FFEB4323E1C9FF21FEB56CE9" blockId="2.[151,1437,1051,1220]" pageId="2" pageNumber="66">
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<emphasis id="60F12804FFEB4323E1C9FF21FE886C1E" bold="true" box="[151,278,1051,1075]" pageId="2" pageNumber="66">FIGURE 1.</emphasis>
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<taxonomicName id="95858F95FFEB4323E041FF21FD9E6C1E" authorityName="Dunlop & Bartel" authorityYear="2025" box="[287,512,1051,1075]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="luisdearmasi" status="sp. nov.">
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<emphasis id="60F12804FFEB4323E041FF21FD9E6C1E" box="[287,512,1051,1075]" italics="true" pageId="2" pageNumber="66">Phrynus luisdearmasi</emphasis>
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</taxonomicName>
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<emphasis id="60F12804FFEB4323E357FF26FDC26C19" bold="true" box="[521,604,1052,1076]" pageId="2" pageNumber="66">
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<taxonomicNameLabel id="7BC2957FFFEB4323E357FF26FDC26C19" box="[521,604,1052,1076]" pageId="2" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, MB.A. 1747 (holotype and only known specimen), from Dominican Republic amber (Miocene: Burdigalian). A. Close-up of the anterior body in dorsal view. B. Camera lucida drawing of the dorsal body and legs. C. Frontal view of the chelicerae and pedipalps. D. Camera lucida drawing of the ventral body and legs. Abbreviations: ch, chelicerae; fe, femur; pp, pedipalp; pt, patella; ti, tibia; legs numbered from I–IV. Scale bars equal 2 mm (A), 5 mm (B, D) and 1 mm (C).
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</paragraph>
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</caption>
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<subSubSection id="1A9FA79DFFEB4323E199FE2CFDA66D79" pageId="2" pageNumber="66" type="materials_examined">
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<paragraph id="523AF416FFEB4323E199FE2CFDA66D79" blockId="2.[151,1437,1301,1976]" pageId="2" pageNumber="66">
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<emphasis id="60F12804FFEB4323E199FE2CFECA6D02" bold="true" box="[199,340,1301,1328]" pageId="2" pageNumber="66">Type series.</emphasis>
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<materialsCitation id="E2EDFE4BFFEB4323E003FE2CFDA66D79" ID-GBIF-Occurrence="5000541301" collectionCode="MB" country="Dominican Republic" pageId="2" pageNumber="66" specimenCode="MB.A. 1747" specimenCount="1" typeStatus="holotype">
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<specimenCode id="02235C6DFFEB4323E003FE2CFE716D1D" box="[349,495,1302,1328]" pageId="2" pageNumber="66">MB.A. 1747</specimenCode>
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.
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<typeStatus id="8D3E4AB4FFEB4323E0A5FE2CFDFD6D1D" box="[507,611,1302,1328]" pageId="2" pageNumber="66" type="holotype">Holotype</typeStatus>
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and only known specimen. From
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<collectingCountry id="2A92B486FFEB4323E2B5FE2CFB486D1D" box="[1003,1238,1302,1328]" name="Dominican Republic" pageId="2" pageNumber="66">Dominican Republic</collectingCountry>
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amber.
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<geologicalTimeScale id="20A574BDFFEB4323E46CFE2CFDAA6D79" pageId="2" pageNumber="66">Miocene: Burdigalian (ca. 20.43 – 15.97 Mya)</geologicalTimeScale>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1A9FA79DFFEB4323E199FE67FC096F29" pageId="2" pageNumber="66" type="diagnosis">
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<paragraph id="523AF416FFEB4323E199FE67FC096F29" blockId="2.[151,1437,1301,1976]" pageId="2" pageNumber="66">
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<emphasis id="60F12804FFEB4323E199FE67FEDC6D5A" bold="true" box="[199,322,1373,1399]" pageId="2" pageNumber="66">Diagnosis.</emphasis>
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Fossil
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<taxonomicName id="95858F95FFEB4323E0CEFE64FDDA6D55" box="[400,580,1374,1400]" pageId="2" pageNumber="66">
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<emphasis id="60F12804FFEB4323E0CEFE64FE736D55" box="[400,493,1374,1400]" italics="true" pageId="2" pageNumber="66">Phrynus</emphasis>
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species
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</taxonomicName>
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from the ‘A’ group
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<emphasis id="60F12804FFEB4323E243FE64FCC56D5A" box="[797,859,1374,1399]" italics="true" pageId="2" pageNumber="66">sensu</emphasis>
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<bibRefCitation id="361489E7FFEB4323E23FFE64FB896D55" author="Quintero Jr, D." box="[865,1047,1374,1400]" pageId="2" pageNumber="66" pagination="1 - 54" refId="ref7168" refString="Quintero Jr, D. (1983) Revision of the amblypygid spiders of Cuba and their relationships with the Caribbean and continental American amblypygid fauna. Studies on the Fauna of Curacao and other Caribbean Islands, 65, 1 - 54." type="journal article" year="1983">Quintero (1983)</bibRefCitation>
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, body length of
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<typeStatus id="8D3E4AB4FFEB4323E596FE64FAB46D55" box="[1224,1322,1374,1400]" pageId="2" pageNumber="66" type="holotype">holotype</typeStatus>
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ca.
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<quantity id="957D59F3FFEB4323E408FE64FA066D55" box="[1366,1432,1374,1400]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="66" unit="mm" value="8.0">8 mm</quantity>
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, with leg I tibia subdivided into 34 articles. Ventral pedipalp femoral spination pattern, FvI–II> FvV> FvIII> FvVI> 5vIV; dorsal pedipalp patella spination pattern Pd5> Pd3> Pd2> Pd4> Pd6> Pd1 = Pd7, with Pd6 almost the same size as Pd4; both dorsal pedipalp tibial spine 1 and proximal ventral tibial spine pronounced. Closely-related living (and fossil) species have less than 34 leg I tibial articles. Relatively large dorsal tibial spine
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<quantity id="957D59F3FFEB4323E452FED4FAA76E25" box="[1292,1337,1518,1544]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="66" unit="in" value="1.0">1 in</quantity>
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the new species also differentiates it from both the amber species
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<taxonomicName id="95858F95FFEB4323E246FD28FC086E06" baseAuthorityName="Schawaller" baseAuthorityYear="1979" box="[792,918,1554,1580]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="resinae">
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<emphasis id="60F12804FFEB4323E246FD28FC086E06" box="[792,918,1554,1580]" italics="true" pageId="2" pageNumber="66">Ph. resinae</emphasis>
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</taxonomicName>
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and the extant species
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<taxonomicName id="95858F95FFEB4323E5C7FD28FA0B6E01" authorityName="C. L. Koch" authorityYear="1840" box="[1177,1429,1554,1580]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="marginemaculatus">
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<emphasis id="60F12804FFEB4323E5C7FD28FA0B6E01" box="[1177,1429,1554,1580]" italics="true" pageId="2" pageNumber="66">Ph. marginemaculatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="95858F95FFEB4323E1C9FD0CFDD06E7D" authority="Armas & Teruel, 2010" authorityName="Armas & Teruel" authorityYear="2010" box="[151,590,1590,1616]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="alejandroi">
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<emphasis id="60F12804FFEB4323E1C9FD0CFEA26E7D" box="[151,316,1590,1616]" italics="true" pageId="2" pageNumber="66">Ph. alejandroi</emphasis>
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<bibRefCitation id="361489E7FFEB4323E01BFD0CFDD06E7D" author="Armas, L. F. de & Teruel, R." box="[325,590,1590,1616]" pageId="2" pageNumber="66" pagination="127 - 130" refId="ref6166" refString="Armas, L. F. de & Teruel, R. (2010) Nueva especie de Phrynus Lamarck, 1801 (Amblypygi: Phrynidae) de Puerto Rico. Boletin de la Sociedad Entomologica Aragonesa, 47, 127 - 130" type="journal article" year="2010">Armas & Teruel, 2010</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="95858F95FFEB4323E303FD0CFBF16E7D" authority="Armas & Perez Gonzalez, 2001" authorityName="Armas & Perez Gonzalez" authorityYear="2001" box="[605,1135,1590,1616]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="eucharis">
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<emphasis id="60F12804FFEB4323E303FD0CFD706E7D" box="[605,750,1590,1616]" italics="true" pageId="2" pageNumber="66">Ph. eucharis</emphasis>
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<bibRefCitation id="361489E7FFEB4323E3A7FD0CFBF16E7D" author="Armas, L. F. de & Perez Gonzalez, A." box="[761,1135,1590,1616]" pageId="2" pageNumber="66" pagination="47 - 66" refId="ref6128" refString="Armas, L. F. de & Perez Gonzalez, A. (2001) Los amblipigidos de Republica Dominicana (Arachnida: Amblypygi). Revista Iberica de Aracnologia, 3, 47 - 66." type="journal article" year="2001">Armas & Pérez González, 2001</bibRefCitation>
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||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="95858F95FFEB4323E5EFFD0CFE2B6E59" authority="Armas & Perez Gonzalez, 2001" authorityName="Armas & Perez Gonzalez" authorityYear="2001" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="kennidae">
|
||
<emphasis id="60F12804FFEB4323E5EFFD0CFAD96E7D" box="[1201,1351,1590,1616]" italics="true" pageId="2" pageNumber="66">Ph. kennidae</emphasis>
|
||
<bibRefCitation id="361489E7FFEB4323E40CFD0CFE2B6E59" author="Armas, L. F. de & Perez Gonzalez, A." pageId="2" pageNumber="66" pagination="47 - 66" refId="ref6128" refString="Armas, L. F. de & Perez Gonzalez, A. (2001) Los amblipigidos de Republica Dominicana (Arachnida: Amblypygi). Revista Iberica de Aracnologia, 3, 47 - 66." type="journal article" year="2001">Armas & Pérez González, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
, in which this tibial spine is noticeably smaller. Pedipalp spination of the new species resembles that of
|
||
<taxonomicName id="95858F95FFEB4323E001FD44FC1E6EB5" authority="Armas & Perez Gonzalez, 2001" authorityName="Armas & Perez Gonzalez" authorityYear="2001" box="[351,896,1662,1688]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="hispaniolae">
|
||
<emphasis id="60F12804FFEB4323E001FD44FD8E6EB5" box="[351,528,1662,1688]" italics="true" pageId="2" pageNumber="66">Ph. hispaniolae</emphasis>
|
||
<bibRefCitation id="361489E7FFEB4323E349FD44FC1E6EB5" author="Armas, L. F. de & Perez Gonzalez, A." box="[535,896,1662,1688]" pageId="2" pageNumber="66" pagination="47 - 66" refId="ref6128" refString="Armas, L. F. de & Perez Gonzalez, A. (2001) Los amblipigidos de Republica Dominicana (Arachnida: Amblypygi). Revista Iberica de Aracnologia, 3, 47 - 66." type="journal article" year="2001">Armas & Pérez González, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="95858F95FFEB4323E2E2FD44FAC16EB5" authority="Teruel & Armas, 2005" authorityName="Teruel & Armas" authorityYear="2005" box="[956,1375,1661,1688]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="decoratus">
|
||
<emphasis id="60F12804FFEB4323E2E2FD44FBC66EB5" box="[956,1112,1662,1688]" italics="true" pageId="2" pageNumber="66">Ph. decoratus</emphasis>
|
||
<bibRefCitation id="361489E7FFEB4323E53EFD44FAC16EB5" author="Teruel, R. & Armas, L. F. de" box="[1120,1375,1661,1688]" pageId="2" pageNumber="66" pagination="129 - 133" refId="ref7549" refString="Teruel, R. & Armas, L. F. de (2005) Novedades aracnologicas de la Republica Dominicana (Arachnida: Amblypygi, Schizomida, Solpugida, Uropygi). Boletin de la Sociedad Entomologica Aragonesa, 37, 129 - 133." type="journal article" year="2005">Teruel & Armas, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
, both of which also have a more pronounced dorsal tibial spine 1, but in the new species Pd6 and Pd4 are similar in size whereas in
|
||
<taxonomicName id="95858F95FFEB4323E048FDFCFE2F6ECD" authorityName="Teruel & Armas" authorityYear="2005" box="[278,433,1734,1760]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="decoratus">
|
||
<emphasis id="60F12804FFEB4323E048FDFCFE2F6ECD" box="[278,433,1734,1760]" italics="true" pageId="2" pageNumber="66">Ph. decoratus</emphasis>
|
||
</taxonomicName>
|
||
P6 is noticeably shorter than P4; while the number of tibial articles (34) differentiates the new fossil from
|
||
<taxonomicName id="95858F95FFEB4323E010FDD0FE606F29" authorityName="Armas & Perez Gonzalez" authorityYear="2001" box="[334,510,1770,1796]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="hispaniolae">
|
||
<emphasis id="60F12804FFEB4323E010FDD0FE606F29" box="[334,510,1770,1796]" italics="true" pageId="2" pageNumber="66">Ph. hispaniolae</emphasis>
|
||
</taxonomicName>
|
||
which has, at most, only 32 articles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1A9FA79DFFEB4325E199FC37FAB46CE8" lastPageId="4" lastPageNumber="68" pageId="2" pageNumber="66" type="description">
|
||
<paragraph id="523AF416FFEB4323E199FC37FE166F95" blockId="2.[151,1437,1301,1976]" pageId="2" pageNumber="66">
|
||
<emphasis id="60F12804FFEB4323E199FC37FEC76F0A" bold="true" box="[199,345,1805,1831]" pageId="2" pageNumber="66">Description.</emphasis>
|
||
Almost complete inclusion, visible in both dorsal and ventral views (
|
||
<figureCitation id="CABEE893FFEB4323E52FFC34FB2A6F05" box="[1137,1204,1806,1832]" captionStart="FIGURE 1" captionStartId="2.[151,250,1051,1075]" captionTargetBox="[151,1434,181,1027]" captionTargetId="figure-17@2.[151,1435,181,1027]" captionTargetPageId="2" captionText="FIGURE 1. Phrynus luisdearmasi sp. nov., MB.A. 1747 (holotype and only known specimen), from Dominican Republic amber (Miocene: Burdigalian). A. Close-up of the anterior body in dorsal view. B. Camera lucida drawing of the dorsal body and legs. C. Frontal view of the chelicerae and pedipalps. D. Camera lucida drawing of the ventral body and legs. Abbreviations: ch, chelicerae; fe, femur; pp, pedipalp; pt, patella; ti, tibia; legs numbered from I–IV. Scale bars equal 2 mm (A), 5 mm (B, D) and 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.14596232" httpUri="https://zenodo.org/record/14596232/files/figure.png" pageId="2" pageNumber="66">Fig. 1</figureCitation>
|
||
); limb series almost complete, better preserved on right side. Total preserved length ca. 8. Carapace reniform, wider than long; length 2.9, maximum width 4.2. Median and lateral eyes present (
|
||
<figureCitation id="CABEE893FFEB4323E27EFC6CFCFD6F5D" box="[800,867,1878,1904]" captionStart="FIGURE 2" captionStartId="3.[151,250,1100,1124]" captionTargetBox="[151,1429,181,1076]" captionTargetId="figure-18@3.[151,1435,181,1076]" captionTargetPageId="3" captionText="FIGURE 2. Phrynus luisdearmasi sp. nov. A. Details of pedipalps in dorsal view. B. Camera lucida drawing of A; dorsal tibial spine 1 arrowed, being larger than the corresponding spine in the previous amber species P. resinae Schawaller, 1979 and its putative extant relative P. marginemaculatus C.L. Koch, 1840. C. Details of pedipalps in ventral view. D. Camera lucida drawing of B. Abbreviations: ch, chelicerae; fe, femur; ley, lateral eyes; mey, median eyes; pp, pedipalp; pt, patella; ti, tibia. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14596238" httpUri="https://zenodo.org/record/14596238/files/figure.png" pageId="2" pageNumber="66">Fig. 2</figureCitation>
|
||
), but lateral eyes inconspicuous. Chelicerae largely tucked beneath carapace, total length ca. 1.5; pattern of cheliceral dentition equivocal. Sternal elements between leg coxae also equivocal.
|
||
</paragraph>
|
||
<caption id="06FAA49EFFEA4322E1C9FF76FE1A6CD9" ID-DOI="http://doi.org/10.5281/zenodo.14596238" ID-Zenodo-Dep="14596238" httpUri="https://zenodo.org/record/14596238/files/figure.png" pageId="3" pageNumber="67" startId="3.[151,250,1100,1124]" targetBox="[151,1429,181,1076]" targetPageId="3" targetType="figure">
|
||
<paragraph id="523AF416FFEA4322E1C9FF76FE1A6CD9" blockId="3.[151,1437,1100,1268]" pageId="3" pageNumber="67">
|
||
<emphasis id="60F12804FFEA4322E1C9FF76FE886C49" bold="true" box="[151,278,1100,1124]" pageId="3" pageNumber="67">FIGURE 2.</emphasis>
|
||
<taxonomicName id="95858F95FFEA4322E041FF76FD9E6C49" authorityName="Dunlop & Bartel" authorityYear="2025" box="[287,512,1100,1124]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="luisdearmasi" status="sp. nov.">
|
||
<emphasis id="60F12804FFEA4322E041FF76FD9E6C49" box="[287,512,1100,1124]" italics="true" pageId="3" pageNumber="67">Phrynus luisdearmasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="60F12804FFEA4322E357FF76FDC26C49" bold="true" box="[521,604,1100,1124]" pageId="3" pageNumber="67">
|
||
<taxonomicNameLabel id="7BC2957FFFEA4322E357FF76FDC26C49" box="[521,604,1100,1124]" pageId="3" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Details of pedipalps in dorsal view. B. Camera lucida drawing of A; dorsal tibial spine 1 arrowed, being larger than the corresponding spine in the previous amber species
|
||
<taxonomicName id="95858F95FFEA4322E504FF4BFAEC6CA5" authority="Schawaller, 1979" authorityName="Schawaller" authorityYear="1979" box="[1114,1394,1136,1160]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="resinae">
|
||
<emphasis id="60F12804FFEA4322E504FF4BFB236CA5" box="[1114,1213,1137,1160]" italics="true" pageId="3" pageNumber="67">P. resinae</emphasis>
|
||
<bibRefCitation id="361489E7FFEA4322E59DFF4AFAEC6CA5" author="Schawaller, W." box="[1219,1394,1136,1160]" pageId="3" pageNumber="67" pagination="1 - 12" refId="ref7265" refString="Schawaller, W. (1979) Erstnachweis der Ordnung Geisselspinnen in Dominikanischem Bernstein (Stuttgarter Bernsteinsammlung: Arachnida, Amblypygi). Stuttgarter Beitrage zur Naturkunde (B), 50, 1 - 12." type="journal article" year="1979">Schawaller, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
and its putative extant relative
|
||
<taxonomicName id="95858F95FFEA4322E0F2FFAFFCA26C81" authority="C. L. Koch, 1840" authorityName="C. L. Koch" authorityYear="1840" box="[428,828,1172,1196]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="marginemaculatus">
|
||
<emphasis id="60F12804FFEA4322E0F2FFAFFD1D6C81" box="[428,643,1172,1196]" italics="true" pageId="3" pageNumber="67">P. marginemaculatus</emphasis>
|
||
C.L. Koch, 1840
|
||
</taxonomicName>
|
||
. C. Details of pedipalps in ventral view. D. Camera lucida drawing of B. Abbreviations: ch, chelicerae; fe, femur; ley, lateral eyes; mey, median eyes; pp, pedipalp; pt, patella; ti, tibia. Scale bars equal 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="06FAA49EFFEA4322E1C9FC32FB666F45" ID-DOI="http://doi.org/10.5281/zenodo.14596242" ID-Zenodo-Dep="14596242" httpUri="https://zenodo.org/record/14596242/files/figure.png" pageId="3" pageNumber="67" startId="3.[151,250,1800,1824]" targetBox="[151,1419,1328,1776]" targetPageId="3" targetType="figure">
|
||
<paragraph id="523AF416FFEA4322E1C9FC32FB666F45" blockId="3.[151,1436,1800,1896]" pageId="3" pageNumber="67">
|
||
<emphasis id="60F12804FFEA4322E1C9FC32FE886F0D" bold="true" box="[151,278,1800,1824]" pageId="3" pageNumber="67">FIGURE 3.</emphasis>
|
||
<taxonomicName id="95858F95FFEA4322E07EFC32FD9F6F0D" authorityName="Dunlop & Bartel" authorityYear="2025" box="[288,513,1800,1824]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="luisdearmasi" status="sp. nov.">
|
||
<emphasis id="60F12804FFEA4322E07EFC32FD9F6F0D" box="[288,513,1800,1824]" italics="true" pageId="3" pageNumber="67">Phrynus luisdearmasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="60F12804FFEA4322E355FC32FDC16F0D" bold="true" box="[523,607,1800,1824]" pageId="3" pageNumber="67">
|
||
<taxonomicNameLabel id="7BC2957FFFEA4322E355FC32FDC16F0D" box="[523,607,1800,1824]" pageId="3" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Detail of the of leg I tibia; note the high number of tibial articles (34). B. Schematic, annotated drawing of the pedipalp’s dorsal spines; large tibial spine 1 differentiating it from the previous amber species again arrowed. Abbreviations: fe, femur; pt, patella; ti, tibia. Scale bars equal 2 mm (A) and 1 mm (B).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="523AF416FFED4325E199FBADFCBA6AA8" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="68">
|
||
Pedipalps robust and spinous (
|
||
<figureCitation id="CABEE893FFED4325E37BFBADFDF2689C" box="[549,620,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1100,1124]" captionTargetBox="[151,1429,181,1076]" captionTargetId="figure-18@3.[151,1435,181,1076]" captionTargetPageId="3" captionText="FIGURE 2. Phrynus luisdearmasi sp. nov. A. Details of pedipalps in dorsal view. B. Camera lucida drawing of A; dorsal tibial spine 1 arrowed, being larger than the corresponding spine in the previous amber species P. resinae Schawaller, 1979 and its putative extant relative P. marginemaculatus C.L. Koch, 1840. C. Details of pedipalps in ventral view. D. Camera lucida drawing of B. Abbreviations: ch, chelicerae; fe, femur; ley, lateral eyes; mey, median eyes; pp, pedipalp; pt, patella; ti, tibia. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14596238" httpUri="https://zenodo.org/record/14596238/files/figure.png" pageId="4" pageNumber="68">Fig. 2</figureCitation>
|
||
). Article lengths: trochanter 0.8, femur 1.4, patella 2.4, tibia 1.4, tarsus 1.2. Trochanter with 4 spines, from proximal to distal second spine largest; trochanter with at least three long setae. Femur with six dorsal spines (
|
||
<figureCitation id="CABEE893FFED4325E0A9FBE5FDCF68D4" box="[503,593,223,249]" captionStart="FIGURE 3" captionStartId="3.[151,250,1800,1824]" captionTargetBox="[151,1419,1328,1776]" captionTargetId="figure-105@3.[151,1435,1328,1776]" captionTargetPageId="3" captionText="FIGURE 3. Phrynus luisdearmasi sp. nov. A. Detail of the of leg I tibia; note the high number of tibial articles (34). B. Schematic, annotated drawing of the pedipalp’s dorsal spines; large tibial spine 1 differentiating it from the previous amber species again arrowed. Abbreviations: fe, femur; pt, patella; ti, tibia. Scale bars equal 2 mm (A) and 1 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14596242" httpUri="https://zenodo.org/record/14596242/files/figure.png" pageId="4" pageNumber="68">Fig. 3B</figureCitation>
|
||
), numbered from proximal to distal; Fd1–3 larger than 4–6, but angle of preservation makes resolving whether there was a common F1+F2 base
|
||
<emphasis id="60F12804FFED4325E282FA39FB846931" box="[988,1050,259,284]" italics="true" pageId="4" pageNumber="68">sensu</emphasis>
|
||
<bibRefCitation id="361489E7FFED4325E57CFA39FA026930" author="Armas, L. F. de & Perez Gonzalez, A." box="[1058,1436,259,285]" pageId="4" pageNumber="68" pagination="47 - 66" refId="ref6128" refString="Armas, L. F. de & Perez Gonzalez, A. (2001) Los amblipigidos de Republica Dominicana (Arachnida: Amblypygi). Revista Iberica de Aracnologia, 3, 47 - 66." type="journal article" year="2001">Armas & Pérez González (2001)</bibRefCitation>
|
||
equivocal, and also makes it difficult to accurately determine the relative heights of 4–6. Femur also ventrally with six spines (designated by Roman numerals
|
||
<emphasis id="60F12804FFED4325E39AFA71FC9C6949" box="[708,770,331,356]" italics="true" pageId="4" pageNumber="68">sensu</emphasis>
|
||
<bibRefCitation id="361489E7FFED4325E255FA71FC4B6948" author="Seiter, M. & Strobl, L. & Schwaha, T. & Prendini, L. & Schramm, F. D." box="[779,981,331,357]" pageId="4" pageNumber="68" pagination="10 - 28" refId="ref7394" refString="Seiter, M., Strobl, L., Schwaha, T., Prendini, L. & Schramm, F. D. (2022) Morphometry of the pedipalp patella provides new characters for species-level taxonomy in whip spiders (Arachnida, Amblypygi): A test case with description of a new species of Phrynus. Zoologischer Anzeiger, 298, 10 - 28." type="journal article" year="2022">
|
||
Seiter
|
||
<emphasis id="60F12804FFED4325E208FA71FC0E6948" box="[854,912,331,357]" italics="true" pageId="4" pageNumber="68">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
), FvI–II longest, followed by FvV, then FvIII and FvVI, with 5vIV shortest. Patella wide with eight (left,
|
||
<figureCitation id="CABEE893FFED4325E2D7FA55FC7E69A4" box="[905,992,367,393]" captionStart="FIGURE 3" captionStartId="3.[151,250,1800,1824]" captionTargetBox="[151,1419,1328,1776]" captionTargetId="figure-105@3.[151,1435,1328,1776]" captionTargetPageId="3" captionText="FIGURE 3. Phrynus luisdearmasi sp. nov. A. Detail of the of leg I tibia; note the high number of tibial articles (34). B. Schematic, annotated drawing of the pedipalp’s dorsal spines; large tibial spine 1 differentiating it from the previous amber species again arrowed. Abbreviations: fe, femur; pt, patella; ti, tibia. Scale bars equal 2 mm (A) and 1 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14596242" httpUri="https://zenodo.org/record/14596242/files/figure.png" pageId="4" pageNumber="68">Fig. 3B</figureCitation>
|
||
) and seven (right) dorsal spines, again numbered from proximal to distal. From longest to shortest: Pd5> Pd3> Pd2> Pd4> Pd6> Pd1 = Pd7. Additional (eighth) spine on left side shorter than Pd7 (
|
||
<figureCitation id="CABEE893FFED4325E3CAFA8DFD7069FC" box="[660,750,439,465]" captionStart="FIGURE 3" captionStartId="3.[151,250,1800,1824]" captionTargetBox="[151,1419,1328,1776]" captionTargetId="figure-105@3.[151,1435,1328,1776]" captionTargetPageId="3" captionText="FIGURE 3. Phrynus luisdearmasi sp. nov. A. Detail of the of leg I tibia; note the high number of tibial articles (34). B. Schematic, annotated drawing of the pedipalp’s dorsal spines; large tibial spine 1 differentiating it from the previous amber species again arrowed. Abbreviations: fe, femur; pt, patella; ti, tibia. Scale bars equal 2 mm (A) and 1 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14596242" httpUri="https://zenodo.org/record/14596242/files/figure.png" pageId="4" pageNumber="68">Fig. 3B</figureCitation>
|
||
); not usually included in notation scheme. Ventrally with at least 5 subequal spines, but details lacking. Tibia with three dorsal spines (
|
||
<figureCitation id="CABEE893FFED4325E2BCFAE1FBA569D8" box="[994,1083,475,501]" captionStart="FIGURE 3" captionStartId="3.[151,250,1800,1824]" captionTargetBox="[151,1419,1328,1776]" captionTargetId="figure-105@3.[151,1435,1328,1776]" captionTargetPageId="3" captionText="FIGURE 3. Phrynus luisdearmasi sp. nov. A. Detail of the of leg I tibia; note the high number of tibial articles (34). B. Schematic, annotated drawing of the pedipalp’s dorsal spines; large tibial spine 1 differentiating it from the previous amber species again arrowed. Abbreviations: fe, femur; pt, patella; ti, tibia. Scale bars equal 2 mm (A) and 1 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14596242" httpUri="https://zenodo.org/record/14596242/files/figure.png" pageId="4" pageNumber="68">Fig. 3B</figureCitation>
|
||
); from proximal to distal 1 and 3 approximately equal in length, spine 2 about twice as long as 1 and 3. Tibia with at least two ventral spines (
|
||
<figureCitation id="CABEE893FFED4325E431FAC5FF226A10" captionStart="FIGURE 2" captionStartId="3.[151,250,1100,1124]" captionTargetBox="[151,1429,181,1076]" captionTargetId="figure-18@3.[151,1435,181,1076]" captionTargetPageId="3" captionText="FIGURE 2. Phrynus luisdearmasi sp. nov. A. Details of pedipalps in dorsal view. B. Camera lucida drawing of A; dorsal tibial spine 1 arrowed, being larger than the corresponding spine in the previous amber species P. resinae Schawaller, 1979 and its putative extant relative P. marginemaculatus C.L. Koch, 1840. C. Details of pedipalps in ventral view. D. Camera lucida drawing of B. Abbreviations: ch, chelicerae; fe, femur; ley, lateral eyes; mey, median eyes; pp, pedipalp; pt, patella; ti, tibia. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14596238" httpUri="https://zenodo.org/record/14596238/files/figure.png" pageId="4" pageNumber="68">Fig. 2D</figureCitation>
|
||
), proximal one shorter than distal one. Details equivocal, but presence of a well-developed first ventral tibial spine here distinctive. Tarsus sickle-shaped, bearing a tuft of seta (the cleaning organ) about midway along its length (
|
||
<figureCitation id="CABEE893FFED4325E1C1F951FE836AA8" box="[159,285,619,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1100,1124]" captionTargetBox="[151,1429,181,1076]" captionTargetId="figure-18@3.[151,1435,181,1076]" captionTargetPageId="3" captionText="FIGURE 2. Phrynus luisdearmasi sp. nov. A. Details of pedipalps in dorsal view. B. Camera lucida drawing of A; dorsal tibial spine 1 arrowed, being larger than the corresponding spine in the previous amber species P. resinae Schawaller, 1979 and its putative extant relative P. marginemaculatus C.L. Koch, 1840. C. Details of pedipalps in ventral view. D. Camera lucida drawing of B. Abbreviations: ch, chelicerae; fe, femur; ley, lateral eyes; mey, median eyes; pp, pedipalp; pt, patella; ti, tibia. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14596238" httpUri="https://zenodo.org/record/14596238/files/figure.png" pageId="4" pageNumber="68">Figs 2B, D</figureCitation>
|
||
); no additional spine on the prodorsal surface.
|
||
</paragraph>
|
||
<paragraph id="523AF416FFED4325E199F9B5FE316BE4" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="68">
|
||
Legs gracile (
|
||
<figureCitation id="CABEE893FFED4325E03AF9B5FE396A84" box="[356,423,655,681]" captionStart="FIGURE 1" captionStartId="2.[151,250,1051,1075]" captionTargetBox="[151,1434,181,1027]" captionTargetId="figure-17@2.[151,1435,181,1027]" captionTargetPageId="2" captionText="FIGURE 1. Phrynus luisdearmasi sp. nov., MB.A. 1747 (holotype and only known specimen), from Dominican Republic amber (Miocene: Burdigalian). A. Close-up of the anterior body in dorsal view. B. Camera lucida drawing of the dorsal body and legs. C. Frontal view of the chelicerae and pedipalps. D. Camera lucida drawing of the ventral body and legs. Abbreviations: ch, chelicerae; fe, femur; pp, pedipalp; pt, patella; ti, tibia; legs numbered from I–IV. Scale bars equal 2 mm (A), 5 mm (B, D) and 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.14596232" httpUri="https://zenodo.org/record/14596232/files/figure.png" pageId="4" pageNumber="68">Fig. 1</figureCitation>
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), leg I especially elongate and slender. Leg I article lengths: femur 6.9, patella 0.5, tibia 13.3, tarsus at least 8.5, but distal tip equivocal. Both tibia and metatarsus with multiple short articles, 34 the tibia (
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<figureCitation id="CABEE893FFED4325E1C1F9EDFF666ADC" box="[159,248,727,753]" captionStart="FIGURE 3" captionStartId="3.[151,250,1800,1824]" captionTargetBox="[151,1419,1328,1776]" captionTargetId="figure-105@3.[151,1435,1328,1776]" captionTargetPageId="3" captionText="FIGURE 3. Phrynus luisdearmasi sp. nov. A. Detail of the of leg I tibia; note the high number of tibial articles (34). B. Schematic, annotated drawing of the pedipalp’s dorsal spines; large tibial spine 1 differentiating it from the previous amber species again arrowed. Abbreviations: fe, femur; pt, patella; ti, tibia. Scale bars equal 2 mm (A) and 1 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14596242" httpUri="https://zenodo.org/record/14596242/files/figure.png" pageId="4" pageNumber="68">Fig. 3A</figureCitation>
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) and at least 50 tarsal elements. Leg II article lengths: femur 5.1, patella, 0.5, basitibia 4.2, distitibia 2.5, basitarsus 0.6, tarsus 0.7. Leg III article lengths: femur 6.0, patella 0.5, basitibia 3.7, remaining articles ca. 4.4, but details equivocal. Leg IV article lengths: femur 5.5, patella 0.5, basitibia 4.4, distitibia 2.9, basitarsus 0.7, tarsus 0.8. Basitibiae of legs II–IV with weakly expressed subdivision into three articles, distal basitibiae and distitibiae with several long trichobothria, but details equivocal. Tarsi of at least legs II and IV subdivided into three tarsomeres; tarsus terminates in a pair of small claws. This tarsal configuration was probably present in leg III too, but structure of leg tip here equivocal.
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</paragraph>
|
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<paragraph id="523AF416FFED4325E199F8E9FAB46CE8" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="68">
|
||
Opisthosoma broadly oval (
|
||
<figureCitation id="CABEE893FFED4325E348F8E9FDFE6BC0" box="[534,608,979,1005]" captionStart="FIGURE 1" captionStartId="2.[151,250,1051,1075]" captionTargetBox="[151,1434,181,1027]" captionTargetId="figure-17@2.[151,1435,181,1027]" captionTargetPageId="2" captionText="FIGURE 1. Phrynus luisdearmasi sp. nov., MB.A. 1747 (holotype and only known specimen), from Dominican Republic amber (Miocene: Burdigalian). A. Close-up of the anterior body in dorsal view. B. Camera lucida drawing of the dorsal body and legs. C. Frontal view of the chelicerae and pedipalps. D. Camera lucida drawing of the ventral body and legs. Abbreviations: ch, chelicerae; fe, femur; pp, pedipalp; pt, patella; ti, tibia; legs numbered from I–IV. Scale bars equal 2 mm (A), 5 mm (B, D) and 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.14596232" httpUri="https://zenodo.org/record/14596232/files/figure.png" pageId="4" pageNumber="68">Fig. 1</figureCitation>
|
||
), anteriormost part slightly tucked under posterior carapace margin, opisthosoma also tapering somewhat posteriorly; length 5.4, maximum width 3.9. At least 8 tergites visible, anteriormost elements slightly recurved, otherwise all with straight posterior margins; at least 5 tergites with pairs of indented muscle apodemes towards centre of the plate. Ventrally (
|
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<figureCitation id="CABEE893FFED4325E2D3FF05FB996C74" box="[909,1031,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[151,250,1051,1075]" captionTargetBox="[151,1434,181,1027]" captionTargetId="figure-17@2.[151,1435,181,1027]" captionTargetPageId="2" captionText="FIGURE 1. Phrynus luisdearmasi sp. nov., MB.A. 1747 (holotype and only known specimen), from Dominican Republic amber (Miocene: Burdigalian). A. Close-up of the anterior body in dorsal view. B. Camera lucida drawing of the dorsal body and legs. C. Frontal view of the chelicerae and pedipalps. D. Camera lucida drawing of the ventral body and legs. Abbreviations: ch, chelicerae; fe, femur; pp, pedipalp; pt, patella; ti, tibia; legs numbered from I–IV. Scale bars equal 2 mm (A), 5 mm (B, D) and 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.14596232" httpUri="https://zenodo.org/record/14596232/files/figure.png" pageId="4" pageNumber="68">Fig. 1C–D</figureCitation>
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), genital operculum, and succeeding sternite, with notably recurved posterior margins. Genital structures associated with genital plate equivocal. Other sternites poorly preserved, but ca. five elements demarcated by weak transverse lines. Posteriormost opisthosomal segments converge towards a blunt pygidium, but details of individual elements here difficult to resolve.
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</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
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</document> |