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<mods:title id="DFA38EEE7DFF32EF6550B18122FA2319">A new species of Lusius Tosquinet (Hymenoptera, Ichneumonidae, Ichneumoninae) from China, with a key to Chinese species</mods:title>
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<mods:namePart id="7EFABD938A2530C9F1FDFA576F6BBA9B">Han, Yuan-Yuan</mods:namePart>
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Genus
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<emphasis id="2395EA836B03A763B390FEF67A24FEB6" bold="true" box="[238,320,295,321]" italics="true" pageId="1" pageNumber="293">Lusius</emphasis>
<bibRefCitation id="75704B606B03A763B239FEF6796FFEB6" author="Tosquinet, J." box="[327,523,295,321]" pageId="1" pageNumber="293" pagination="1 - 403" refId="ref2992" refString="Tosquinet, J. (1903) Ichneumonides nouveaux. Memoires de la Societe Entomologique de Belgique, 10, 1 - 403." type="journal article" year="1903">Tosquinet, 1903</bibRefCitation>
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<treatmentCitation id="904010806B03A763B3E9FEBD7ADDFE73" author="Tosquinet, J." box="[151,441,364,388]" page="384" pageId="1" pageNumber="293" year="1903">
<taxonomicName id="D6E14D126B03A763B3E9FEBD7ADDFE73" ID-CoL="5HCW" authority="Tosquinet, 1903: 384" authorityName="Tosquinet" authorityPageNumber="384" authorityYear="1903" box="[151,441,364,388]" class="Insecta" family="Ichneumonidae" genus="Lusius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2395EA836B03A763B3E9FEBD7BBDFE74" box="[151,217,364,387]" italics="true" pageId="1" pageNumber="293">Lusius</emphasis>
<bibRefCitation id="75704B606B03A763B39EFEBD7ADDFE73" author="Tosquinet, J." box="[224,441,364,388]" pageId="1" pageNumber="293" pagination="1 - 403" refId="ref2992" refString="Tosquinet, J. (1903) Ichneumonides nouveaux. Memoires de la Societe Entomologique de Belgique, 10, 1 - 403." type="journal article" year="1903">Tosquinet, 1903: 384</bibRefCitation>
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.
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<typeStatus id="CE5A88336B03A763B2BCFEBD7A90FE73" box="[450,500,364,388]" pageId="1" pageNumber="293">Type</typeStatus>
species:
<taxonomicName id="D6E14D126B03A763B12FFEBD78D7FE73" ID-CoL="3WFXV" authority="Tosquinet, 1903" authorityName="Tosquinet" authorityYear="1903" box="[593,947,364,388]" class="Insecta" family="Ichneumonidae" genus="Lusius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="macilentus">
<emphasis id="2395EA836B03A763B12FFEBD786CFE73" box="[593,776,364,388]" italics="true" pageId="1" pageNumber="293">Lusius macilentus</emphasis>
<bibRefCitation id="75704B606B03A763B073FEBD78D7FE73" author="Tosquinet, J." box="[781,947,364,388]" pageId="1" pageNumber="293" pagination="1 - 403" refId="ref2992" refString="Tosquinet, J. (1903) Ichneumonides nouveaux. Memoires de la Societe Entomologique de Belgique, 10, 1 - 403." type="journal article" year="1903">Tosquinet, 1903</bibRefCitation>
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, by original designation and monotype.
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<treatmentCitationGroup id="31F111BF6B03A763B3E9FE5B7944FE55" box="[151,544,394,418]" pageId="1" pageNumber="293">
<treatmentCitation id="904010806B03A763B3E9FE5B7978FE55" author="Heinrich, G. H." box="[151,540,394,418]" page="128" pageId="1" pageNumber="293" year="1938">
<taxonomicName id="D6E14D126B03A763B3E9FE5B7978FE55" authority="Heinrich, 1938: 128" authorityName="Heinrich" authorityPageNumber="128" authorityYear="1938" box="[151,540,394,418]" class="Insecta" family="Ichneumonidae" genus="Mesochorischnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2395EA836B03A763B3E9FE5B7A2CFE55" box="[151,328,394,418]" italics="true" pageId="1" pageNumber="293">Mesochorischnus</emphasis>
<bibRefCitation id="75704B606B03A763B230FE5B7978FE55" author="Heinrich, G. H." box="[334,540,394,418]" pageId="1" pageNumber="293" pagination="1 - 139" refId="ref2594" refString="Heinrich, G. H. (1938) Les Ichneumonides de Madagascar. III Ichneumonidae Ichneumoninae. Memoires de l'Academie Malgache Fascicule, 25, 1 - 139." type="journal article" year="1938">Heinrich, 1938: 128</bibRefCitation>
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.
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species:
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<emphasis id="2395EA836B03A763B1CCFE5B7884FE55" box="[690,992,394,418]" italics="true" pageId="1" pageNumber="293">Mesochorischnus tenuissimus</emphasis>
<bibRefCitation id="75704B606B03A763B098FE5B7F1BFE55" author="Heinrich, G. H." box="[998,1151,394,418]" pageId="1" pageNumber="293" pagination="1 - 139" refId="ref2594" refString="Heinrich, G. H. (1938) Les Ichneumonides de Madagascar. III Ichneumonidae Ichneumoninae. Memoires de l'Academie Malgache Fascicule, 25, 1 - 139." type="journal article" year="1938">Heinrich, 1938</bibRefCitation>
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, by original designation and monotype. Synoymized by Townes &amp; Townes (1966: 217).
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<emphasis id="2395EA836B03A763B3E9FE3E7A76FDFE" bold="true" box="[151,274,495,521]" pageId="1" pageNumber="293">Diagnosis.</emphasis>
See diagnosis in (
<bibRefCitation id="75704B606B03A763B2A3FE3E7983FDFE" author="Nascimento, A. C. &amp; Padua, D. &amp; Araujo, R. &amp; Mazariegos, L. &amp; Fernandes, D. R. R." box="[477,743,495,522]" pageId="1" pageNumber="293" pagination="289 - 296" refId="ref2739" refString="Nascimento, A. C., Padua, D., Araujo, R., Mazariegos, L. &amp; Fernandes, D. R. R. (2024) Synopsis of Lusius Tosquinet, 1903 (Hymenoptera: Ichneumonidae: Phaeogenini) from South America: a new species, new records, and key to Neotropical species. Zootaxa, 5410 (2), 289 - 296. https: // doi. org / 10.11646 / zootaxa. 5410.2.9" type="journal article" year="2024">
Nascimento
<emphasis id="2395EA836B03A763B115FE2179F9FDFE" box="[619,669,495,521]" italics="true" pageId="1" pageNumber="293">et al</emphasis>
. 2024
</bibRefCitation>
).
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Key to Chinese species of
<taxonomicName id="D6E14D126B03A763B2A7FD8A794FFD82" ID-CoL="5HCW" ID-ENA="144393" authorityName="Tosquinet" authorityYear="1903" box="[473,555,603,629]" class="Insecta" family="Ichneumonidae" genus="Lusius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2395EA836B03A763B2A7FD8A794FFD82" bold="true" box="[473,555,603,629]" italics="true" pageId="1" pageNumber="293">Lusius</emphasis>
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<keyLead id="AA1095A46B03A763B3E9FD717EF8FD07" pageId="1" pageNumber="293">
1. Apical margin of clypeus thin, not emarginated; frons not vertically striate; malar space 1.01.3× longer than basal width of mandible; hind basitarsus as long as segments 25; tarsal claws pectinate; second metasomal tergite 2.5× longer than wide apex; gonostyle 0.45× as long as hind tibia.................................................................
<emphasis id="2395EA836B03A763B6EEFD0A7EF8FD07" bold="true" box="[1424,1436,731,752]" pageId="1" pageNumber="293">2</emphasis>
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<paragraph id="115E36916B03A763B3E9FD287EF8FCBE" blockId="1.[151,1437,672,958]" pageId="1" pageNumber="293">
<keyLead id="AA1095A46B03A763B3E9FD287EF8FCBE" pageId="1" pageNumber="293">
-. Apical margin of clypeus thick, emarginated; frons vertically striate laterally; malar space 0.7× as long as basal width of mandible; hind basitarsus 1.3× longer than segments 25; tarsal claws simple; second abdominal tergite 4.5× longer than wide at apex; gonostyle 0.6× as long as hind tibia..........................................
<taxonomicName id="D6E14D126B03A763B7F7FCE27EF8FCBE" ID-CoL="6R239" authority="Kusigemati, 1986" authorityName="Kusigemati" authorityYear="1986" box="[1161,1436,819,841]" class="Insecta" family="Ichneumonidae" genus="Lusius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="2395EA836B03A763B7F7FCE27F8FFCBF" box="[1161,1259,819,840]" italics="true" pageId="1" pageNumber="293">L. gracilis</emphasis>
<bibRefCitation id="75704B606B03A763B78CFCE27EF8FCBE" author="Kusigemati, K." box="[1266,1436,819,841]" pageId="1" pageNumber="293" pagination="22 - 27" refId="ref2624" refString="Kusigemati, K. (1986) On the Formosan species of the genus Lusius Tosquinet, with description of a new species (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, 7, 22 - 27." type="journal article" year="1986">Kusigemati, 1986</bibRefCitation>
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<keyLead id="AA1095A46B03A763B3E9FC817EF8FC74" pageId="1" pageNumber="293">
2. Propodeum reticulate-rugose, with strong punctures; first tergite fulvous brown posteriorly, outer side of hind coxa sometimes fusco-rufous......................................................................
<taxonomicName id="D6E14D126B03A763B7DBFCBF7EF8FC74" ID-CoL="3WFXS" authority="(Morley, 1913)" baseAuthorityName="Morley" baseAuthorityYear="1913" box="[1189,1436,877,899]" class="Insecta" family="Ichneumonidae" genus="Lusius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="apollos">
<emphasis id="2395EA836B03A763B7DBFCBF7E61FC75" box="[1189,1285,877,899]" italics="true" pageId="1" pageNumber="293">L. apollos</emphasis>
(
<bibRefCitation id="75704B606B03A763B66CFCBC7EF1FC74" author="Morley, C." box="[1298,1429,877,899]" pageId="1" pageNumber="293" refId="ref2665" refString="Morley, C. (1913) The fauna of British India including Ceylon and Burma. Hymenoptera, Vol. 3. Ichneumonidae. London, British Museum, 531 pp." type="book" year="1913">Morley, 1913</bibRefCitation>
)
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-. Propodeum weakly rugose with transverse-striation (
<figureCitation id="89DA2A146B03A763B19EFC5A7848FC57" box="[736,812,907,929]" captionStart="FIGURE 2" captionStartId="3.[152,255,1707,1731]" captionTargetBox="[300,1288,237,1684]" captionTargetId="figure-55@3.[300,1288,237,1684]" captionTargetPageId="3" captionText="FIGURE 2. Lusius hunanensis sp. nov., holotype, female. A fore wing B mesosoma, lateral view C propodeum, dorsal view D first metasomal segment, lateral view E head, anterior view F head, dorsal view G mesoscutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14594710" httpUri="https://zenodo.org/record/14594710/files/figure.png" pageId="1" pageNumber="293">Fig. 2C</figureCitation>
), without punctures; first metasomal tergite entirely yellow (
<figureCitation id="89DA2A146B03A763B60BFC5A7A66FC4A" captionStart="FIGURE 2" captionStartId="3.[152,255,1707,1731]" captionTargetBox="[300,1288,237,1684]" captionTargetId="figure-55@3.[300,1288,237,1684]" captionTargetPageId="3" captionText="FIGURE 2. Lusius hunanensis sp. nov., holotype, female. A fore wing B mesosoma, lateral view C propodeum, dorsal view D first metasomal segment, lateral view E head, anterior view F head, dorsal view G mesoscutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14594710" httpUri="https://zenodo.org/record/14594710/files/figure.png" pageId="1" pageNumber="293">Fig. 2D</figureCitation>
); hind coxa entirely pale yellow (
<figureCitation id="89DA2A146B03A763B13AFC7979EBFC4A" box="[580,655,936,958]" captionStart="FIGURE 2" captionStartId="3.[152,255,1707,1731]" captionTargetBox="[300,1288,237,1684]" captionTargetId="figure-55@3.[300,1288,237,1684]" captionTargetPageId="3" captionText="FIGURE 2. Lusius hunanensis sp. nov., holotype, female. A fore wing B mesosoma, lateral view C propodeum, dorsal view D first metasomal segment, lateral view E head, anterior view F head, dorsal view G mesoscutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14594710" httpUri="https://zenodo.org/record/14594710/files/figure.png" pageId="1" pageNumber="293">Fig. 2D</figureCitation>
)..............................................
<taxonomicName id="D6E14D126B03A763B7BAFC787E2EFC4A" authorityName="Han &amp; Chen &amp; Dong" authorityYear="2024" box="[1220,1354,936,958]" class="Insecta" family="Ichneumonidae" genus="Lusius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="hunanensis" status="sp. nov.">
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<emphasis id="2395EA836B03A763B62EFC797EF8FC4A" bold="true" box="[1360,1436,936,957]" pageId="1" pageNumber="293">
<taxonomicNameLabel id="38A657F86B03A763B62EFC797EF8FC4A" box="[1360,1436,936,957]" pageId="1" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
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View file

@ -1,56 +1,55 @@
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@ -89,18 +88,15 @@ The new species-group name is proposed in honour of Prof. Luis F. de Armas for h
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<emphasis id="60F12804FFEB4323E199FE2CFECA6D02" bold="true" box="[199,340,1301,1328]" pageId="2" pageNumber="66">Type series.</emphasis>
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.A. 1747.
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and only known specimen.
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<treatment id="DA2C4500FFED4325E1C9FE2CFDBF6F84" ID-DOI="http://doi.org/10.5281/zenodo.14596972" ID-GBIF-Taxon="251122250" ID-Zenodo-Dep="14596972" LSID="urn:lsid:plazi:treatment:DA2C4500FFED4325E1C9FE2CFDBF6F84" httpUri="http://treatment.plazi.org/id/DA2C4500FFED4325E1C9FE2CFDBF6F84" lastPageNumber="68" pageId="4" pageNumber="68" scope_family="Phrynidae" scope_order="Amblypygi">
<subSubSection id="1A9FA79DFFED4325E1C9FE2CFE406D1C" box="[151,478,1302,1329]" pageId="4" pageNumber="68" type="nomenclature">
<paragraph id="523AF416FFED4325E1C9FE2CFE406D1C" blockId="4.[151,478,1302,1329]" box="[151,478,1302,1329]" pageId="4" pageNumber="68">
<heading id="0972437AFFED4325E1C9FE2CFE406D1C" bold="true" box="[151,478,1302,1329]" fontSize="11" level="1" pageId="4" pageNumber="68" reason="1">
<emphasis id="60F12804FFED4325E1C9FE2CFE406D1C" bold="true" box="[151,478,1302,1329]" pageId="4" pageNumber="68">
<taxonomicName id="95858F95FFED4325E1C9FE2CFEC36D1C" authorityName="Dunlop &amp; Bartel" authorityYear="2025" box="[151,349,1302,1329]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="poinari" status="nom. nov.">
<emphasis id="60F12804FFED4325E1C9FE2CFEC36D1C" bold="true" box="[151,349,1302,1329]" italics="true" pageId="4" pageNumber="68">Phrynus poinari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BC2957FFFED4325E03BFE2DFE406D1C" box="[357,478,1303,1329]" pageId="4" pageNumber="68">nom. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="1A9FA79DFFED4325E1C9FE61FC2D6D9D" pageId="4" pageNumber="68" type="reference_group">
<paragraph id="523AF416FFED4325E1C9FE61FEF66DBF" blockId="4.[151,1436,1371,1457]" pageId="4" pageNumber="68">
<taxonomicName id="95858F95FFED4325E1C9FE61FECF6D5E" authorityName="Poinar &amp; Brown" authorityYear="2004" box="[151,337,1371,1395]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="60F12804FFED4325E1C9FE61FECF6D5E" box="[151,337,1371,1395]" italics="true" pageId="4" pageNumber="68">Phrynus mexicana</emphasis>
</taxonomicName>
[
<emphasis id="60F12804FFED4325E000FE66FEE46D5E" box="[350,378,1372,1395]" italics="true" pageId="4" pageNumber="68">sic</emphasis>
]
<treatmentCitation id="D324D207FFED4325E0D9FE61FD776D5E" author="Poinar Jr., G. O. &amp; Brown, A. E." box="[391,745,1371,1395]" page="1882" pageId="4" pageNumber="68" year="2004">
<bibRefCitation id="361489E7FFED4325E0D9FE61FD776D5E" author="Poinar Jr., G. O. &amp; Brown, A. E." box="[391,745,1371,1395]" pageId="4" pageNumber="68" pagination="1881 - 1885" refId="ref7090" refString="Poinar Jr., G. O. &amp; Brown, A. E. (2004) A new whip spider (Arachnida: Amblypygi), Phrynus mexicana, is described from Mexican amber. Beitrage zur Araneologie, 3, 1881 - 1885." type="journal article" year="2004">Poinar &amp; Brown, 2004: 18821883</bibRefCitation>
</treatmentCitation>
, figs 19, photo 696 (junior primary homonym of
<taxonomicName id="95858F95FFED4325E586FE61FEC26DBF" authority="Bilimek, 1867" authorityName="Bilimek" authorityYear="1867" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="60F12804FFED4325E586FE61FA026D5E" box="[1240,1436,1371,1395]" italics="true" pageId="4" pageNumber="68">Phrynus mexicanus</emphasis>
Bilimek, 1867
</taxonomicName>
).
</paragraph>
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<taxonomicName id="95858F95FFED4325E1C9FEA2FEC36D9D" authorityName="Bilimek" authorityYear="1867" box="[151,349,1432,1456]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="60F12804FFED4325E1C9FEA2FEC36D9D" box="[151,349,1432,1456]" italics="true" pageId="4" pageNumber="68">Phrynus mexicanus</emphasis>
</taxonomicName>
: Dunlop et al. 2015: 201203; Opinion 2453 2020: 6162.
</paragraph>
</subSubSection>
<subSubSection id="1A9FA79DFFED4325E1C9FEE5FBB46E30" pageId="4" pageNumber="68" type="etymology">
<paragraph id="523AF416FFED4325E1C9FEE5FBB46E30" blockId="4.[151,1437,1503,1961]" pageId="4" pageNumber="68">
<emphasis id="60F12804FFED4325E1C9FEE5FE806DD4" bold="true" box="[151,286,1503,1529]" pageId="4" pageNumber="68">Etymology.</emphasis>
The new species-group name is proposed in honour of Prof. George O. Poinar jr. for his extensive contributions towards the study of amber inclusions; especially in the Neotropics.
</paragraph>
</subSubSection>
<subSubSection id="1A9FA79DFFED4325E199FD1DFCF56E48" pageId="4" pageNumber="68" type="materials_examined">
<paragraph id="523AF416FFED4325E199FD1DFCF56E48" blockId="4.[151,1437,1503,1961]" pageId="4" pageNumber="68">
<emphasis id="60F12804FFED4325E199FD1DFED06E6D" bold="true" box="[199,334,1574,1601]" pageId="4" pageNumber="68">Type series</emphasis>
.
<materialsCitation id="E2EDFE4BFFED4325E003FD1DFCF56E48" ID-GBIF-Occurrence="5000541302" country="Mexico" pageId="4" pageNumber="68" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<typeStatus id="8D3E4AB4FFED4325E003FD1DFE5B6E6C" box="[349,453,1575,1601]" pageId="4" pageNumber="68" type="holotype">Holotype</typeStatus>
in the Poinar amber collection (Accession no. A-10-264), maintained at Oregon State University, USA.
<collectingRegion id="90413AF4FFED4325E03EFD71FE246E48" box="[352,442,1611,1637]" country="Mexico" name="Chiapas" pageId="4" pageNumber="68">Chiapas</collectingRegion>
amber,
<collectingCountry id="2A92B486FFED4325E34AFD71FDF36E48" box="[532,621,1611,1637]" name="Mexico" pageId="4" pageNumber="68">Mexico</collectingCountry>
.
<geologicalTimeScale id="20A574BDFFED4325E329FD71FCF86E48" box="[631,870,1611,1637]" pageId="4" pageNumber="68">Miocene: Aquitanian</geologicalTimeScale>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1A9FA79DFFED4325E199FD55FDBF6F84" pageId="4" pageNumber="68" type="discussion">
<paragraph id="523AF416FFED4325E199FD55FDBF6F84" blockId="4.[151,1437,1503,1961]" pageId="4" pageNumber="68">
<emphasis id="60F12804FFED4325E199FD55FEA56EA4" bold="true" box="[199,315,1647,1673]" pageId="4" pageNumber="68">Remarks.</emphasis>
This second amber
<taxonomicName id="95858F95FFED4325E340FD55FD4D6EA4" box="[542,723,1647,1673]" pageId="4" pageNumber="68">
<emphasis id="60F12804FFED4325E340FD55FDE56EA4" box="[542,635,1647,1673]" italics="true" pageId="4" pageNumber="68">Phrynus</emphasis>
species
</taxonomicName>
was initially named
<taxonomicName id="95858F95FFED4325E2E0FD55FA066EA4" authority="Poinar &amp; Brown, 2004" authorityName="Poinar &amp; Brown" authorityYear="2004" box="[958,1432,1647,1673]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="60F12804FFED4325E2E0FD55FB106EA5" box="[958,1166,1647,1673]" italics="true" pageId="4" pageNumber="68">Phrynus mexicana</emphasis>
<bibRefCitation id="361489E7FFED4325E5CBFD55FA066EA4" author="Poinar Jr., G. O. &amp; Brown, A. E." box="[1173,1432,1647,1673]" pageId="4" pageNumber="68" pagination="1881 - 1885" refId="ref7090" refString="Poinar Jr., G. O. &amp; Brown, A. E. (2004) A new whip spider (Arachnida: Amblypygi), Phrynus mexicana, is described from Mexican amber. Beitrage zur Araneologie, 3, 1881 - 1885." type="journal article" year="2004">Poinar &amp; Brown, 2004</bibRefCitation>
</taxonomicName>
, and comes from the probably slightly older (
<bibRefCitation id="361489E7FFED4325E3DFFDA9FCF26E80" author="Riquelme, F. &amp; Ortega-Flores, B. &amp; Estrada-Ruiz, E. &amp; Cordova-Tabares, V." box="[641,876,1683,1709]" pageId="4" pageNumber="68" refId="ref7212" refString="Riquelme, F., Ortega-Flores, B. Estrada-Ruiz, E. &amp; Cordova-Tabares, V. (2024) Zircon U - Pb Ages of the Chiapas Amber- Lagerstatte (Mexico: Chiapas: Simojovel). SSRN. http: // doi. org / 10.2139 / ssrn. 4778409" type="book" year="2024">
Riquelme
<emphasis id="60F12804FFED4325E3A8FDA9FCB36E80" box="[758,813,1683,1709]" italics="true" pageId="4" pageNumber="68">et al.</emphasis>
2024
</bibRefCitation>
)
<collectingRegion id="90413AF4FFED4325E224FDA9FC4A6E80" box="[890,980,1683,1709]" country="Mexico" name="Chiapas" pageId="4" pageNumber="68">Chiapas</collectingRegion>
amber of
<collectingCountry id="2A92B486FFED4325E51CFDA9FB056E80" box="[1090,1179,1683,1709]" name="Mexico" pageId="4" pageNumber="68">Mexico</collectingCountry>
.As noted by Dunlop et al. (2015), it should have been named
<taxonomicName id="95858F95FFED4325E31BFD8DFD5A6EFD" authorityName="Bilimek" authorityYear="1867" box="[581,708,1719,1745]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="60F12804FFED4325E310FD8DFD5A6EFD" box="[590,708,1719,1744]" italics="true" pageId="4" pageNumber="68">mexicanus</emphasis>
</taxonomicName>
which renders it a junior primary homonym of a Recent species
<taxonomicName id="95858F95FFED4325E1C9FDE1FF6A6ED8" authorityName="Lamarck" authorityYear="1801" box="[151,244,1755,1781]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="60F12804FFED4325E1C9FDE1FF6A6ED8" box="[151,244,1755,1781]" italics="true" pageId="4" pageNumber="68">Phrynus</emphasis>
</taxonomicName>
(now
<taxonomicName id="95858F95FFED4325E01DFDE1FE4B6ED8" authorityName="Moreno" authorityYear="1940" box="[323,469,1755,1781]" class="Arachnida" family="Phrynidae" genus="Paraphrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="60F12804FFED4325E01DFDE1FE4B6ED8" box="[323,469,1755,1781]" italics="true" pageId="4" pageNumber="68">Paraphrynus</emphasis>
</taxonomicName>
)
<taxonomicName id="95858F95FFED4325E0B7FDE1FC8E6ED8" authority="Bilimek, 1867" authorityName="Bilimek" authorityYear="1867" box="[489,784,1755,1781]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="60F12804FFED4325E0B7FDE1FDC16ED9" box="[489,607,1755,1780]" italics="true" pageId="4" pageNumber="68">mexicanus</emphasis>
Bilimek, 1867
</taxonomicName>
. An appeal by Dunlop et al. (2015) to the
<emphasis id="60F12804FFED4325E456FDE1FDEC6F34" italics="true" pageId="4" pageNumber="68">International Commission on Zoological Nomenclature</emphasis>
to conserve
<taxonomicName id="95858F95FFED4325E256FDC5FB5C6F34" authority="Poinar &amp; Brown, 2004" authorityName="Poinar &amp; Brown" authorityYear="2004" box="[776,1218,1791,1817]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="60F12804FFED4325E256FDC5FC306F35" box="[776,942,1791,1817]" italics="true" pageId="4" pageNumber="68">Ph. mexicanus</emphasis>
<bibRefCitation id="361489E7FFED4325E2E6FDC5FB5C6F34" author="Poinar Jr., G. O. &amp; Brown, A. E." box="[952,1218,1791,1817]" pageId="4" pageNumber="68" pagination="1881 - 1885" refId="ref7090" refString="Poinar Jr., G. O. &amp; Brown, A. E. (2004) A new whip spider (Arachnida: Amblypygi), Phrynus mexicana, is described from Mexican amber. Beitrage zur Araneologie, 3, 1881 - 1885." type="journal article" year="2004">Poinar &amp; Brown, 2004</bibRefCitation>
</taxonomicName>
, on the grounds of Bilimeks species having been transferred to another genus was unsuccessful (Opinion 2453). The commissions ruling suggested that the simplest solution would be a replacement name for the amber fossil. In this context, we take the opportunity to rename the
<collectingRegion id="90413AF4FFED4325E345FC51FDEB6FA8" box="[539,629,1899,1925]" country="Mexico" name="Chiapas" pageId="4" pageNumber="68">Chiapas</collectingRegion>
whip spider
<taxonomicName id="95858F95FFED4325E25BFC51FC256FA9" authorityName="Dunlop &amp; Bartel" authorityYear="2025" box="[773,955,1899,1925]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="poinari" status="nom. nov.">
<emphasis id="60F12804FFED4325E25BFC51FC256FA9" box="[773,955,1899,1925]" italics="true" pageId="4" pageNumber="68">Phrynus poinari</emphasis>
</taxonomicName>
<emphasis id="60F12804FFED4325E29FFC51FBAB6FA8" bold="true" box="[961,1077,1899,1925]" pageId="4" pageNumber="68">
<taxonomicNameLabel id="7BC2957FFFED4325E29FFC51FBAB6FA8" box="[961,1077,1899,1925]" pageId="4" pageNumber="68">nom. nov.</taxonomicNameLabel>
</emphasis>
in recognition of George Poinar who initially described the species.
</paragraph>
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<mods:role id="91A9D96EF52192586AB6E108AD34C32E">
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<mods:namePart id="D09741DCB5FFF4E7CFA5D305B069D907">Delgado-Santa, Leonardo</mods:namePart>
<mods:affiliation id="1D58D2DC0C99F65EF5542F54749160B8">Grupo de Investigación ECDYSIS, Programa de Biología, Facultad de Ciencias Básicas y Tecnologías, Universidad del Quindío. Armenia, Quindío, Colombia.</mods:affiliation>
<mods:namePart id="5A77143F6E0554358166263D0B09C8D5">Delgado-Santa, Leonardo</mods:namePart>
<mods:affiliation id="6D386ACFF9DE71555445B0E46055CDD8">Grupo de Investigación ECDYSIS, Programa de Biología, Facultad de Ciencias Básicas y Tecnologías, Universidad del Quindío. Armenia, Quindío, Colombia.</mods:affiliation>
</mods:name>
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<mods:role id="767FED98E664F44822C1C719B146BC88">
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<mods:role id="F16E7F20EE851C11847D4C1678E7F82D">
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<mods:namePart id="717567F27056BEB9E9943C938D8288D8">Villarreal, Osvaldo</mods:namePart>
<mods:affiliation id="66B9396E56BFC154F7DA07DEE72AC988">Centro de Ecología, Instituto Venezolano de Investigaciones Científicas (IVIC), km 11 carretera Panamericana, Altos de Pipe, Miranda 1204 - A, Venezuela. &amp; Instituto y Museo del Instituto de Zoología Agrícola, Facultad de Agronomía, Universidad Central de Venezuela, Maracay, Aragua, Venezuela</mods:affiliation>
<mods:namePart id="9A506D49D46BEA3053BE8015FAA65132">Villarreal, Osvaldo</mods:namePart>
<mods:affiliation id="BD220C5FA11E363152222E9213723054">Centro de Ecología, Instituto Venezolano de Investigaciones Científicas (IVIC), km 11 carretera Panamericana, Altos de Pipe, Miranda 1204 - A, Venezuela. &amp; Instituto y Museo del Instituto de Zoología Agrícola, Facultad de Agronomía, Universidad Central de Venezuela, Maracay, Aragua, Venezuela</mods:affiliation>
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<paragraph id="791936B21B2EC90BFF20F8EB62902B54" blockId="1.[151,952,1806,1905]" box="[151,248,1879,1905]" pageId="1" pageNumber="74">
<figureCitation id="E19D2A371B2EC90BFF20F8EB62902B54" box="[151,248,1879,1905]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,250,568,592]" captionStartId-1="3.[151,250,1859,1883]" captionStartId-2="4.[151,250,1982,2006]" captionStartId-3="5.[151,250,1348,1372]" captionStartId-4="6.[151,250,1416,1440]" captionStartId-5="7.[151,250,1943,1967]" captionTargetBox-0="[152,1433,181,540]" captionTargetBox-1="[151,1436,689,1836]" captionTargetBox-2="[151,1436,1023,1955]" captionTargetBox-3="[155,1429,181,1317]" captionTargetBox-4="[151,1432,182,1381]" captionTargetBox-5="[182,1410,191,1910]" captionTargetId-0="figure-21@3.[151,1436,181,544]" captionTargetId-1="figure-46@3.[151,1436,689,1836]" captionTargetId-2="figure-437@4.[151,1436,1022,1958]" captionTargetId-3="figure-21@5.[151,1435,181,1324]" captionTargetId-4="figure-26@6.[151,1436,181,1392]" captionTargetId-5="figure-21@7.[172,1415,181,1919]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. Calima dearmasi sp. nov., male holotype (CIUQ-25563): (A) Lateral habitus; (B) dorsal habitus. Scale bars = 1.0 mm." captionText-1="FIGURE 2. Calima dearmasi sp. nov., male paratype (CIUQ-25565), right chelicerae: (A) ectal view; (B) Movable finger, mesal view, arrow shows the lamellated edge (T); (C) fixed finger; (D) Serrula and setal group G1 with spicules (Sp) on the stalk. G17 = Setal groups; GT = guard tooth; S = serrula;. Figures not scaled." captionText-2="FIGURE 3. Calima dearmasi sp. nov., male holotype (CIUQ-25563) and female (CIUQ-25566): (A, B) Right pedipalp, ectal (A) and mesal (B) views; (CE) Flagellum, dorsal (C), ventral (D), and lateral (E) views. Scale bars = 0.5 mm." captionText-3="FIGURE 4. Calima dearmasi sp. nov., (A-F) male holotype (CIUQ-25563): (A) Cheliceral fixed finger; (B) Cheliceral movable finger; (C) Right pedipalp, ectal view; (D) Flagellum, dorsal view; (E) Flagellum, ventral view; (F) Flagellum, lateral view. (G) Female (CIUQ25566), Spermathecae, dorsal view. Scale bars = 0.5 mm. Figures A, B and G without scales." captionText-4="FIGURE 5. Male and female genitalia. (A-B) Calima dearmasi sp. nov., (A) female, spermathecae, dorsal view (CIUQ-25566); (B) male, genitalia, dorsal view (paratype, CIUQ-25565); (C) Colombiazomus sp., male, genitalia, dorsal view; (D) Piaroa sp., male, genitalia, dorsal view. B= bulb; De= dorsal eminence; Br= bridge; D= diverticules; DAFu= dorsal arm of the furcula; FGp= fold of gonopods; LBFu= lateral bar of the furcula; MS= median septum; Pt= pterapohysis; S= stalk; Sd= sclerotized duct; VAFu= Ventral arm of the furcula. Figures not scaled." captionText-5="FIGURE 6. Distribution map of the genus Calima Moreno-Gonzalez &amp; Villarreal. The area highlighted in gray represents the ecoregion Cauca Valley montane forests." figureDoi-0="http://doi.org/10.5281/zenodo.14596262" figureDoi-1="http://doi.org/10.5281/zenodo.14596266" figureDoi-2="http://doi.org/10.5281/zenodo.14596270" figureDoi-3="http://doi.org/10.5281/zenodo.14596276" figureDoi-4="http://doi.org/10.5281/zenodo.14596282" figureDoi-5="http://doi.org/10.5281/zenodo.14596288" httpUri-0="https://zenodo.org/record/14596262/files/figure.png" httpUri-1="https://zenodo.org/record/14596266/files/figure.png" httpUri-2="https://zenodo.org/record/14596270/files/figure.png" httpUri-3="https://zenodo.org/record/14596276/files/figure.png" httpUri-4="https://zenodo.org/record/14596282/files/figure.png" httpUri-5="https://zenodo.org/record/14596288/files/figure.png" pageId="1" pageNumber="74">Figs 16</figureCitation>
</paragraph>
@ -75,10 +75,10 @@ urn:lsid:zoobank.org:act:
<subSubSection id="31BC65391B2EC908FF20F82363C62CF0" lastPageId="2" lastPageNumber="75" pageId="1" pageNumber="74" type="materials_examined">
<paragraph id="791936B21B2EC908FF20F82363C62CF0" blockId="1.[151,1437,1950,2013]" lastBlockId="2.[151,1436,150,573]" lastPageId="2" lastPageNumber="75" pageId="1" pageNumber="74">
<emphasis id="4BD2EAA01B2EC90BFF20F82363342B9D" bold="true" box="[151,348,1950,1977]" pageId="1" pageNumber="74">Type material—</emphasis>
<materialsCitation id="C9CE3CEF1B2EC908FEEBF82362862C94" collectingDate="2024-02-04" collectionCode="CIUQ" collectorName="L. Delgado-Santa" country="Colombia" elevation="2153" lastPageId="2" lastPageNumber="75" latitude="5.0561113" location="Manizales" longLatPrecision="21" longitude="-75.492775" pageId="1" pageNumber="74" specimenCode="CIUQ-25563" specimenCount="1" stateProvince="Caldas" typeStatus="holotype">
<specimenCount id="6FA0FD3B1B2EC90BFEEBF82363192B9C" box="[348,369,1951,1977]" count="1" pageId="1" pageNumber="74" type="male"></specimenCount>
<typeStatus id="A61D88101B2EC90BFECDF82363B72B9C" box="[378,479,1951,1977]" pageId="1" pageNumber="74" type="holotype">holotype</typeStatus>
,
<materialsCitation id="C9CE3CEF1B2EC908FE5BF823630D2C94" collectingDate="2024-02-04" collectorName="L. Delgado-Santa &amp; University" country="Colombia" elevation="2153" lastPageId="2" lastPageNumber="75" latitude="5.0561113" location="Manizales" longLatPrecision="21" longitude="-75.492775" pageId="1" pageNumber="74" specimenCount="1" stateProvince="Caldas" typeStatus="paratype">
<collectingCountry id="01B176221B2EC90BFE5BF82360EE2B9C" box="[492,646,1951,1977]" name="Colombia" pageId="1" pageNumber="74">COLOMBIA</collectingCountry>
:
<collectingRegion id="BB62F8501B2EC90BFD25F823608B2B9C" box="[658,739,1951,1977]" country="Colombia" name="Caldas" pageId="1" pageNumber="74">Caldas</collectingRegion>
@ -98,23 +98,26 @@ a.s.l.
</date>
; (
<collectorName id="D45353641B2EC90BFF50F87F63C42BF8" box="[231,428,1987,2013]" pageId="1" pageNumber="74">L. Delgado-Santa</collectorName>
leg.); secondary forest inside the campus of
<collectorName id="D45353641B2EC90BFC14F87F66712BF8" box="[931,1049,1987,2013]" pageId="1" pageNumber="74">University</collectorName>
of
<collectingRegion id="BB62F8501B2EC90BFB89F87F66E52BF8" box="[1086,1165,1987,2013]" country="Colombia" name="Caldas" pageId="1" pageNumber="74">Caldas</collectingRegion>
, under leaflitter (CIUQ-25563).
leg.); secondary forest inside the campus of University of Caldas, under leaflitter (
<specimenCode id="29009EC91B2EC908FAFAF87F628A2C94" lastPageId="2" lastPageNumber="75" pageId="1" pageNumber="74">CIUQ-25563</specimenCode>
).
</materialsCitation>
<materialsCitation id="C9CE3CEF1B2DC908FF43FF2B61EB2C94" box="[244,899,150,177]" collectingDate="2024-02-04" collectionCode="CIUQ" collectorName="L. Delgado-Santa &amp; University" country="Colombia" elevation="2153" latitude="5.0561113" location="Manizales" longitude="-75.492775" pageId="2" pageNumber="75" specimenCode="CIUQ-25564, CIUQ-25565" specimenCount="2" specimenCount-male="2" stateProvince="Caldas" typeStatus="paratype">
<typeStatus id="A61D88101B2DC908FF43FF2B630D2C94" box="[244,357,151,177]" pageId="2" pageNumber="75" type="paratype">Paratypes</typeStatus>
</materialsCitation>
:
<materialsCitation id="C9CE3CEF1B2DC908FEC7FF2B66CB2C94" box="[368,1187,150,177]" collectingDate="2024-02-04" collectorName="L. Delgado-Santa &amp; University" country="Colombia" elevation="2153" latitude="5.0561113" location="Manizales" longLatPrecision="21" longitude="-75.492775" pageId="2" pageNumber="75" specimenCount="2" specimenCount-male="2" stateProvince="Caldas">
<specimenCount id="6FA0FD3B1B2DC908FEC7FF2B63D92C94" box="[368,433,151,177]" count="2" pageId="2" pageNumber="75" type="male">2 ♂♂</specimenCount>
, same data (CIUQ-25564, CIUQ-25565).
<typeStatus id="A61D88101B2DC908FC3EFF2B66CB2C94" box="[905,1187,151,177]" isOtherTypeStatus="true" pageId="2" pageNumber="75">Other material examined</typeStatus>
, same data (
<specimenCode id="29009EC91B2DC908FD8CFF2A60BA2C94" box="[571,722,150,177]" pageId="2" pageNumber="75">CIUQ-25564</specimenCode>
,
<specimenCode id="29009EC91B2DC908FD69FF2A611F2C94" box="[734,887,150,177]" pageId="2" pageNumber="75">CIUQ-25565</specimenCode>
).
</materialsCitation>
:
<materialsCitation id="C9CE3CEF1B2DC908FB19FF2B63C32CF0" collectingDate="2024-02-04" collectorName="L. Delgado-Santa &amp; University" country="Colombia" elevation="2153" latitude="5.0561113" location="Manizales" longLatPrecision="21" longitude="-75.492775" pageId="2" pageNumber="75" specimenCount="1" specimenCount-female="1" stateProvince="Caldas">
<materialsCitation id="C9CE3CEF1B2DC908FC3EFF2B63C32CF0" collectingDate="2024-02-04" collectionCode="CIUQ" collectorName="L. Delgado-Santa &amp; University" country="Colombia" elevation="2153" latitude="5.0561113" location="Manizales" longLatPrecision="21" longitude="-75.492775" pageId="2" pageNumber="75" specimenCode="CIUQ-25566" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Caldas">
Other material examined:
<specimenCount id="6FA0FD3B1B2DC908FB19FF2B66B02C94" box="[1198,1240,151,177]" count="1" pageId="2" pageNumber="75" type="female">1 ♀</specimenCount>
, same data as the holotype (CIUQ-25566)
, same data as the holotype (
<specimenCode id="29009EC91B2DC908FEBFFF0663CA2CF0" box="[264,418,186,213]" pageId="2" pageNumber="75">CIUQ-25566</specimenCode>
)
</materialsCitation>
.
</paragraph>
@ -626,7 +629,7 @@ locality (
The specimen was collected in an Andean evergreen forest remnant, a secondary urban forest patch impacted by anthropogenic activities. Specimens were found by actively looking under leaf litter and under decaying logs during the day.
</paragraph>
</subSubSection>
<subSubSection id="31BC65391B27C902FF70FE2F61652914" pageId="8" pageNumber="81" type="etymology">
<subSubSection id="31BC65391B27C902FF70FE2F60802DD0" pageId="8" pageNumber="81" type="etymology">
<paragraph id="791936B21B27C902FF70FE2F60802DD0" blockId="8.[151,1437,151,1329]" pageId="8" pageNumber="81">
<emphasis id="4BD2EAA01B27C902FF70FE2F630C2D88" bold="true" box="[199,356,403,429]" pageId="8" pageNumber="81">Etymology—</emphasis>
<taxonomicName id="BEA64D311B27C902FED3FE2F63A42D88" authorityName="Delgado-Santa &amp; Villarreal" authorityYear="2025" box="[356,460,403,429]" class="Arachnida" family="Hubbardiidae" genus="Calima" kingdom="Animalia" order="Schizomida" pageId="8" pageNumber="81" phylum="Arthropoda" rank="species" species="dearmasi">
@ -634,6 +637,8 @@ The specimen was collected in an Andean evergreen forest remnant, a secondary ur
</taxonomicName>
is a patronym in honor of the Cuban arachnologist Luis de Armas, in recognition of his work training numerous Latin American Arachnologists, as well as his contribution to the taxonomy of various taxa, including primarily the schizomids of the Caribbean.
</paragraph>
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<subSubSection id="31BC65391B27C902FF70FE4361652914" pageId="8" pageNumber="81" type="description">
<paragraph id="791936B21B27C902FF70FE43606D2E3D" blockId="8.[151,1437,151,1329]" box="[199,517,510,537]" pageId="8" pageNumber="81">
<emphasis id="4BD2EAA01B27C902FF70FE43606D2E3D" bold="true" box="[199,517,510,537]" pageId="8" pageNumber="81">Male genitalia comparison</emphasis>
</paragraph>