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<document ID-DOI="http://doi.org/10.5281/zenodo.5643925" ID-GBIF-Dataset="7399d769-249a-47c9-b099-719ecd66851c" ID-GBIF-Taxon="189980187" ID-Zenodo-Dep="5643925" LSID="urn:lsid:zoobank.org:act:202D3652-BF04-464E-87A2-E1A4611C0CA1" approvedBy="jonas" checkinTime="1635963396380" checkinUser="jonas" docAuthor="Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas &amp; Böhme, Wolfgang" docDate="2021" docId="023E39002430FFF1639BFA682C4EC8AE" docLanguage="en" docName="BonnzoolBull.70.1.141-171.pdf" docOrigin="Bonn zoological Bulletin 70 (1)" docTitle="Calotes vindumbarbatus P. &amp; F. &amp; T. &amp; M. &amp; A. &amp; W. 2021, sp. n." docType="treatment" docUuid="202D3652-BF04-464E-87A2-E1A4611C0CA1" docUuidSource="ZooBank" docVersion="10" lastPageId="25" lastPageNumber="166" masterDocId="FE0741782424FFE86326FFAC2F20CF1D" masterDocTitle="Integrative approach to resolve the Calotes mystaceus Duméril &amp; Bibron, 1837 species complex (Squamata: Agamidae)" masterLastPageNumber="171" masterPageNumber="141" pageId="20" pageNumber="161" updateTime="1636082036242" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Integrative approach to resolve the Calotes mystaceus Duméril &amp; Bibron, 1837 species complex (Squamata: Agamidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wagner, Philipp</mods:namePart>
<mods:nameIdentifier type="LSID">0575CE45-AB5C-4493-A8AD-DD399E18284A</mods:nameIdentifier>
<mods:affiliation>Allwetterzoo Münster, Sentruper Strasse 315, D- 48161 Münster, Germany &amp; Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085, USA</mods:affiliation>
<mods:nameIdentifier type="email">wagner@allwetterzoo.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ihlow, Flora</mods:namePart>
<mods:nameIdentifier type="LSID">10DB565B-DA19-484E-AE50-ED29945658E8</mods:nameIdentifier>
<mods:affiliation>Museum of Zoology Senckenberg, Dresden, Königsbrücker Landstrasse 159, D- 01109 Dresden, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hartmann, Timo</mods:namePart>
<mods:nameIdentifier type="LSID">A911B13A-B364-424E-8579-AE23820F1DC1</mods:nameIdentifier>
<mods:affiliation>ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Flecks, Morris</mods:namePart>
<mods:nameIdentifier type="LSID">974BF839-926B-4039-9617-135F5CEC2596</mods:nameIdentifier>
<mods:affiliation>ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schmitz, Andreas</mods:namePart>
<mods:nameIdentifier type="LSID">5B1146D2-B9A6-48E6-9E4E-EB1C8ED1AE7A</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Geneva, UREC-Herpetology &amp; Ichthyology, Route de Malagnou 1, CH- 1208 Genève, Switzerland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Böhme, Wolfgang</mods:namePart>
<mods:nameIdentifier type="LSID">FFAC2972-9F52-404B-BA9C-489C7793FF8D</mods:nameIdentifier>
<mods:affiliation>ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Bonn zoological Bulletin</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>70</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>141</mods:start>
<mods:end>171</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">https://doi.org/10.20363/BZB-2021.70.1.141</mods:identifier>
<mods:identifier type="GBIF-Dataset">7399d769-249a-47c9-b099-719ecd66851c</mods:identifier>
<mods:identifier type="Zenodo-Dep">5643781</mods:identifier>
<mods:identifier type="ZooBank">4E7B4E14-7682-465D-A2D9-A26EA04AB3FD</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5643925" ID-GBIF-Taxon="189980187" ID-Zenodo-Dep="5643925" LSID="urn:lsid:zoobank.org:act:202D3652-BF04-464E-87A2-E1A4611C0CA1" httpUri="http://treatment.plazi.org/id/023E39002430FFF1639BFA682C4EC8AE" lastPageId="25" lastPageNumber="166" pageId="20" pageNumber="161">
<subSubSection box="[189,520,1476,1500]" pageId="20" pageNumber="161" type="nomenclature">
<paragraph blockId="20.[189,767,1476,1561]" box="[189,520,1476,1500]" pageId="20" pageNumber="161">
<taxonomicName authority="P. &amp; F. &amp; T. &amp; M. &amp; A. &amp; W., 2021" authorityName="P. &amp; F. &amp; T. &amp; M. &amp; A. &amp; W." authorityYear="2021" box="[189,456,1476,1500]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="161" phylum="Chordata" rank="species" species="vindumbarbatus" status="sp. n.">
<emphasis bold="true" box="[189,456,1476,1500]" italics="true" pageId="20" pageNumber="161">Calotes vindumbarbatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[463,520,1477,1500]" pageId="20" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection box="[189,369,1508,1532]" pageId="20" pageNumber="161" type="description">
<paragraph blockId="20.[189,767,1476,1561]" box="[189,369,1508,1532]" pageId="20" pageNumber="161">
(
<figureCitation box="[197,262,1508,1532]" captionStart="Fig" captionStartId="7.[151,186,1855,1877]" captionTargetBox="[162,1361,258,1801]" captionTargetId="figure-15@7.[160,1390,227,1834]" captionTargetPageId="7" captionText="Fig. 2. The Bayesian consensus tree based on 988 bp of mitochondrial DNA (12S rRNA and COI) shows six distinct lineages within Calotes mystaceus. Node support in terms of Bayesian posterior probabilities is indicated by circles at nodes (nodes with a BPP ≥ 0.90 are white, BPP ≥ 0.95 are grey, BPP ≥ 0.99 are black, values &lt;0.90 are not marked). Outgroup (Calotes versicolor) not shown for clarity. Numbers in parentheses behind taxa refer to localities mapped in Fig. 1." figureDoi="http://doi.org/10.5281/zenodo.5643789" httpUri="https://zenodo.org/record/5643789/files/figure.png" pageId="20" pageNumber="161">Fig. 2</figureCitation>
, Clade C)
</paragraph>
</subSubSection>
<subSubSection box="[189,767,1542,1561]" pageId="20" pageNumber="161" type="multiple">
<paragraph blockId="20.[189,767,1476,1561]" box="[189,767,1542,1561]" pageId="20" pageNumber="161">
<emphasis box="[189,767,1542,1561]" italics="true" pageId="20" pageNumber="161">
<uri box="[189,767,1542,1561]" pageId="20" pageNumber="161">
urn:lsid:zoobank.org:act:
<uuid box="[389,767,1542,1561]" pageId="20" pageNumber="161">202D3652-BF04-464E-87A2-E1A4611C0CA1</uuid>
</uri>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="161" type="materials_examined">
<paragraph blockId="20.[189,794,1601,1965]" pageId="20" pageNumber="161">
<materialsCitation ID-GBIF-Occurrence="3397335308" collectingDate="2003-04-08" collectionCode="CAS" collectorName="T. Z. Min" country="Myanmar" latitude="25.37342" location="Gat Shang Yang village" longLatPrecision="1" longitude="97.37475" municipality="Myitkyina Township" pageId="20" pageNumber="161" specimenCode="CAS 232388" specimenCount="1" stateProvince="Kachin State" typeStatus="holotype">
<emphasis bold="true" box="[212,322,1601,1625]" pageId="20" pageNumber="161">
<typeStatus box="[212,318,1601,1625]" pageId="20" pageNumber="161">Holotype</typeStatus>
.
</emphasis>
<specimenCode box="[329,468,1601,1625]" pageId="20" pageNumber="161">CAS 232388</specimenCode>
(
<specimenCount box="[482,598,1601,1625]" pageId="20" pageNumber="161" type="adult">adult male</specimenCount>
,
<figureCitation box="[609,672,1601,1625]" captionStart="Fig" captionStartId="19.[151,186,1896,1918]" captionTargetBox="[190,1360,227,1881]" captionTargetId="figure-14@19.[190,1360,227,1881]" captionTargetPageId="19" captionText="Fig. 9. Calotes vindumbarbatus sp. n. A. Holotype (CAS 232388, adult male) from Myanmar, Kachin State, Gat Shang Yang Village, general view from above. B. Holotype, lateral view. C. Living adult male (CAS 232819) from Myanmar, Kachin State, Mohnyin Township, Hepu village [25.094528° N, 96.401833° E, alt. 243 m.]." figureDoi="http://doi.org/10.5281/zenodo.5643809" httpUri="https://zenodo.org/record/5643809/files/figure.png" pageId="20" pageNumber="161">Fig. 9</figureCitation>
A-B, Clade C) from
<collectingCountry box="[287,392,1632,1656]" name="Myanmar" pageId="20" pageNumber="161" value="Myanmar">Myanmar</collectingCountry>
,
<collectingRegion box="[406,549,1631,1656]" country="Myanmar" name="Kachin" pageId="20" pageNumber="161">Kachin State</collectingRegion>
,
<collectingMunicipality box="[562,791,1631,1655]" pageId="20" pageNumber="161">Myitkyina Township</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:023E39002430FFF1639BFA682C4EC8AE:8F48DECD2430FFFC639BF9D32EE4C98B" box="[189,452,1662,1687]" country="Myanmar" latitude="25.37342" longLatPrecision="1" longitude="97.37475" municipality="Myitkyina Township" name="Gat Shang Yang village" pageId="20" pageNumber="161" stateProvince="Kachin State">Gat Shang Yang village</location>
[
<geoCoordinate box="[468,627,1662,1686]" degrees="25.373421" direction="north" orientation="latitude" pageId="20" pageNumber="161" precision="1" value="25.37342">25.373421° N</geoCoordinate>
,
<geoCoordinate box="[641,782,1662,1686]" degrees="97.37475" direction="east" orientation="longitude" pageId="20" pageNumber="161" precision="1" value="97.37475">97.37475° E</geoCoordinate>
], collected by
<collectorName box="[325,419,1693,1717]" pageId="20" pageNumber="161">T.Z. Min</collectorName>
on
<date box="[458,610,1692,1718]" pageId="20" pageNumber="161" value="2003-04-08">
<collectingDate box="[458,610,1692,1718]" pageId="20" pageNumber="161" value="2003-04-08">
April 8
<superScript attach="left" box="[535,547,1692,1706]" fontSize="6" pageId="20" pageNumber="161">th</superScript>
2003
</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph blockId="20.[189,794,1601,1965]" lastBlockId="20.[831,1438,239,1965]" pageId="20" pageNumber="161">
<materialsCitation ID-GBIF-Occurrence="3397335319" collectionCode="CAS" country="Myanmar" latitude="25.371695" location="North of Swekawngaw" longLatPrecision="1" longitude="95.369026" pageId="20" pageNumber="161" specimenCode="CAS 232247" specimenCount="205" specimenCount-male="205" stateProvince="Sagaing Division" typeStatus="paratype">
<emphasis bold="true" box="[212,333,1724,1748]" pageId="20" pageNumber="161">
<typeStatus box="[212,328,1724,1748]" pageId="20" pageNumber="161">Paratypes</typeStatus>
.
</emphasis>
<specimenCode box="[349,497,1724,1748]" pageId="20" pageNumber="161">
<collectionCode box="[349,401,1724,1748]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="20" pageNumber="161">CAS</collectionCode>
232247
</specimenCode>
from
<collectingCountry box="[582,687,1725,1749]" name="Myanmar" pageId="20" pageNumber="161" value="Myanmar">Myanmar</collectingCountry>
,
<collectingRegion pageId="20" pageNumber="161">Sagaing Division</collectingRegion>
,
<collectingRegion box="[301,516,1755,1779]" pageId="20" pageNumber="161">Homalin Township</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:023E39002430FFF1639BFA682C4EC8AE:8F48DECD2430FFFC6133F9772C3AC9EE" box="[533,794,1755,1779]" country="Myanmar" latitude="25.371695" longLatPrecision="1" longitude="95.369026" name="North of Swekawngaw" pageId="20" pageNumber="161" stateProvince="Sagaing Division">North of Swekawngaw</location>
[
<geoCoordinate box="[195,356,1786,1810]" degrees="25.371694" direction="north" orientation="latitude" pageId="20" pageNumber="161" precision="1" value="25.371695">25.371694° N</geoCoordinate>
,
<geoCoordinate box="[372,527,1786,1810]" degrees="95.369028" direction="east" orientation="longitude" pageId="20" pageNumber="161" precision="1" value="95.369026">95.369028° E</geoCoordinate>
,
<specimenCount box="[542,626,1786,1810]" pageId="20" pageNumber="161" type="generic">
<quantity box="[542,621,1786,1810]" metricMagnitude="2" metricUnit="m" metricValue="2.05" pageId="20" pageNumber="161" unit="m" value="205.0">205 m</quantity>
.
</specimenCount>
]
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3397335306" collectionCode="CAS" country="Myanmar" location="Gat Shang Yang village" municipality="Myitkyina" pageId="20" pageNumber="161" specimenCode="CAS 232387" specimenCount="1" stateProvince="Kachin State">
<specimenCode box="[650,794,1786,1810]" pageId="20" pageNumber="161">
<collectionCode box="[650,702,1786,1810]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="20" pageNumber="161">CAS</collectionCode>
232387
</specimenCode>
from
<collectingCountry box="[255,360,1818,1842]" name="Myanmar" pageId="20" pageNumber="161" value="Myanmar">Myanmar</collectingCountry>
,
<collectingRegion box="[378,526,1817,1841]" country="Myanmar" name="Kachin" pageId="20" pageNumber="161">Kachin State</collectingRegion>
:
<collectingMunicipality box="[543,657,1817,1841]" pageId="20" pageNumber="161">Myitkyina</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:023E39002430FFF1639BFA682C4EC8AE:8F48DECD2430FFFC6185F8B52E68C84D" country="Myanmar" municipality="Myitkyina" name="Gat Shang Yang village" pageId="20" pageNumber="161" stateProvince="Kachin State">Gat Shang Yang village</location>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3397335312" collectionCode="CAS" country="Myanmar" elevation="155" latitude="25.803389" location="Linpha village" longLatPrecision="1" longitude="95.52878" pageId="20" pageNumber="161" specimenCode="CAS 239206" specimenCount="1" stateProvince="Sagaing Division">
<specimenCode box="[340,480,1848,1872]" pageId="20" pageNumber="161">
<collectionCode box="[340,392,1848,1872]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="20" pageNumber="161">CAS</collectionCode>
239206
</specimenCode>
from
<collectingCountry box="[547,652,1849,1873]" name="Myanmar" pageId="20" pageNumber="161" value="Myanmar">Myanmar</collectingCountry>
,
<collectingRegion pageId="20" pageNumber="161">Sagaing Division</collectingRegion>
,
<collectingRegion box="[268,430,1879,1903]" pageId="20" pageNumber="161">Hkanti District</collectingRegion>
,
<collectingRegion box="[441,628,1879,1903]" pageId="20" pageNumber="161">Hkanti Township</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:023E39002430FFF1639BFA682C4EC8AE:8F48DECD2430FFFC6159F8FB2C3AC872" box="[639,794,1879,1903]" country="Myanmar" latitude="25.803389" longLatPrecision="1" longitude="95.52878" name="Linpha village" pageId="20" pageNumber="161" stateProvince="Sagaing Division">Linpha village</location>
[
<geoCoordinate box="[195,356,1910,1934]" degrees="25.803389" direction="north" orientation="latitude" pageId="20" pageNumber="161" precision="1" value="25.803389">25.803389° N</geoCoordinate>
,
<geoCoordinate box="[372,527,1910,1934]" degrees="95.528778" direction="east" orientation="longitude" pageId="20" pageNumber="161" precision="1" value="95.52878">95.528778° E</geoCoordinate>
,
<specimenCount box="[543,626,1910,1934]" pageId="20" pageNumber="161" type="generic">
<quantity box="[543,622,1910,1934]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="20" pageNumber="161" unit="m" value="155.0">
<elevation box="[543,622,1910,1934]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="20" pageNumber="161" unit="m" value="155.0">155 m</elevation>
</quantity>
.
</specimenCount>
]
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3397335301" collectionCode="CAS" country="Myanmar" elevation="254" latitude="25.094528" location="Hepu village" longLatPrecision="1" longitude="96.40183" pageId="20" pageNumber="161" specimenCode="CAS 232819" specimenCount="1" stateProvince="Kachin State">
<specimenCode box="[650,794,1910,1934]" pageId="20" pageNumber="161">
<collectionCode box="[650,702,1910,1934]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="20" pageNumber="161">CAS</collectionCode>
232819
</specimenCode>
from
<collectingCountry box="[247,352,1941,1965]" name="Myanmar" pageId="20" pageNumber="161" value="Myanmar">Myanmar</collectingCountry>
,
<collectingRegion box="[363,503,1941,1965]" country="Myanmar" name="Kachin" pageId="20" pageNumber="161">Kachin State</collectingRegion>
,
<collectingRegion box="[514,726,1941,1965]" pageId="20" pageNumber="161">Mohnyin Township</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:023E39002430FFF1639BFA682C4EC8AE:8F48DECD2430FFFC61C6F8392CA8CE1A" box="[736,904,239,1964]" country="Myanmar" latitude="25.094528" longLatPrecision="1" longitude="96.40183" name="Hepu village" pageId="20" pageNumber="161" stateProvince="Kachin State">Hepu village</location>
[
<geoCoordinate box="[920,1079,239,263]" degrees="25.094528" direction="north" orientation="latitude" pageId="20" pageNumber="161" precision="1" value="25.094528">25.094528° N</geoCoordinate>
,
<geoCoordinate box="[1092,1245,239,263]" degrees="96.401833" direction="east" orientation="longitude" pageId="20" pageNumber="161" precision="1" value="96.40183">96.401833° E</geoCoordinate>
,
<specimenCount box="[1259,1340,239,263]" pageId="20" pageNumber="161" type="generic">
<quantity box="[1259,1336,239,263]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="161" unit="m" value="254.0">
<elevation box="[1259,1336,239,263]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="161" unit="m" value="254.0">254 m</elevation>
</quantity>
.
</specimenCount>
]
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3397335302" collectionCode="ZFMK" country="Myanmar" location="Gat Shang Yang village" municipality="Myitkyina" pageId="20" pageNumber="161" specimenCode="ZFMK 97990, CAS 232389" specimenCount="1" stateProvince="Kachin State">
<specimenCode pageId="20" pageNumber="161">
<collectionCode box="[1362,1436,240,263]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" pageId="20" pageNumber="161">ZFMK</collectionCode>
97990
</specimenCode>
(formerly
<specimenCode box="[1019,1163,271,295]" pageId="20" pageNumber="161">
<collectionCode box="[1019,1071,271,295]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="20" pageNumber="161">CAS</collectionCode>
232389
</specimenCode>
) from
<collectingCountry box="[1240,1345,271,295]" name="Myanmar" pageId="20" pageNumber="161" value="Myanmar">Myanmar</collectingCountry>
,
<collectingRegion country="Myanmar" name="Kachin" pageId="20" pageNumber="161">Kachin State</collectingRegion>
:
<collectingMunicipality box="[899,1013,301,325]" pageId="20" pageNumber="161">Myitkyina</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:023E39002430FFF1639BFA682C4EC8AE:8F48DECD2430FFFC60D9FE822BDFCE58" box="[1023,1279,301,326]" country="Myanmar" municipality="Myitkyina" name="Gat Shang Yang village" pageId="20" pageNumber="161" stateProvince="Kachin State">Gat Shang Yang village</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="161" type="diagnosis">
<paragraph blockId="20.[831,1438,239,1965]" pageId="20" pageNumber="161">
<emphasis bold="true" box="[854,971,332,356]" pageId="20" pageNumber="161">Diagnosis.</emphasis>
Asmall
<taxonomicName box="[1079,1160,332,356]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="161" phylum="Chordata" rank="species" species="undetermined">
<emphasis box="[1079,1160,332,356]" italics="true" pageId="20" pageNumber="161">Calotes</emphasis>
</taxonomicName>
species of the complex, males with a known maximum SVL of
<quantity box="[1262,1341,364,387]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="20" pageNumber="161" unit="mm" value="77.0">77 mm</quantity>
, females with a SVL of
<quantity box="[988,1066,394,418]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="20" pageNumber="161" unit="mm" value="73.0">73 mm</quantity>
. Tail length short, up to
<quantity box="[1321,1412,394,418]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="20" pageNumber="161" unit="mm" value="140.0">140 mm</quantity>
. It can be distinguished from the other species of the complex by the combination of the following characters: 1) head slender, not as distinctfrom the body as in other species of the complex; 2) body scales small and homogeneous, smooth, and arranged in regular rows around the body. 3) Upper dorsolateral scales pointing backwards; 4) body scales arranged in 50–54 rows around midbody; 5) 40–47 vertebral spines and scales in males,
<quantity box="[1341,1437,642,666]" metricMagnitude="0" metricUnit="m" metricValue="1.1684" metricValueMax="1.2192" metricValueMin="1.1176" pageId="20" pageNumber="161" unit="in" value="46.0" valueMax="48.0" valueMin="44.0">44–48 in</quantity>
females; two short and well separated spines above the tympanum; 6) low nuchal and dorsal crest continuous to the midpoint between the limbs, composed of erect compressed scales, which are larger on the nuchal and smaller on the dorsal crest, gradually decreasing towards the end of the crest; 7) oblique skinfold in front of the fore limb; 8) extremities and tail long, but shorter than in
<taxonomicName authorityName="Dumeril &amp; Bibron" authorityYear="1837" box="[831,971,890,913]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="161" phylum="Chordata" rank="species" species="mystaceus">
<emphasis box="[831,971,890,913]" italics="true" pageId="20" pageNumber="161">C. mystaceus</emphasis>
</taxonomicName>
; 9) bluish head and chest; 10) whitish stripe from about the nostril, along the upper lip to about the insertion of the fore limb, posteriorly the whitish stripe is laterally intersected by a dark reticulate pattern across vertebrae; 11) brownish blotches missing.
</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="163" pageId="20" pageNumber="161" type="description">
<paragraph blockId="20.[831,1438,239,1965]" pageId="20" pageNumber="161">
<emphasis bold="true" box="[854,1206,1044,1068]" pageId="20" pageNumber="161">
Description of the
<typeStatus box="[1102,1202,1044,1068]" pageId="20" pageNumber="161">holotype</typeStatus>
.
</emphasis>
Small male (SVL
<quantity box="[831,915,1075,1099]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="20" pageNumber="161" unit="mm" value="77.0">77 mm</quantity>
). Extremities relatively slender with the fourth finger and toe longest, tail incomplete. Head slightly distinct from the neck, area posterior of jaw angle slightly swollen. Tip of the snout blunt, rostral small, nostril in a single scale, separated from the rostral and the first supralabial scale by two scales in a row. Canthus rostralis of five scales, distinct and straight, canthus scales continuous with supraciliary scales. Lateral sides of the head flat with 9/9 supralabial scales separated from the orbit by three rows of small scales. Five feebly ridged scales from the orbit to above the tympanum. Tympanum distinct, one spiny scale posteriorly above the tympanum. Mental scale small, bordered by two postmental scales which are separated from each other, only the first pair is in contact with the infralabial scales; 10/10 infralabial scales. Lateral scales on the chin and throat smooth, becoming keeled towards the ventral part. Acontinuous vertebral crest of elevated spiny scales from above the tympanum to about mid-body, spine height gradually decreasing posterior of the neck, in total 40 vertebral scales from the nape to above the cloaca. Dorsal and lateral scales keeled, pointing back- and upwards, in 50 rows around midbody. Caudal scales parallel keeled, directed backwards. Ventral scales parallel keeled.
</paragraph>
<paragraph blockId="20.[831,1438,239,1965]" lastBlockId="22.[188,796,239,1222]" lastPageId="22" lastPageNumber="163" pageId="20" pageNumber="161">
<emphasis bold="true" box="[854,1061,1817,1841]" pageId="20" pageNumber="161">Male coloration.</emphasis>
Brilliant coloration unknown (
<figureCitation box="[839,911,1848,1872]" captionStart="Fig" captionStartId="19.[151,186,1896,1918]" captionTargetBox="[190,1360,227,1881]" captionTargetId="figure-14@19.[190,1360,227,1881]" captionTargetPageId="19" captionText="Fig. 9. Calotes vindumbarbatus sp. n. A. Holotype (CAS 232388, adult male) from Myanmar, Kachin State, Gat Shang Yang Village, general view from above. B. Holotype, lateral view. C. Living adult male (CAS 232819) from Myanmar, Kachin State, Mohnyin Township, Hepu village [25.094528° N, 96.401833° E, alt. 243 m.]." figureDoi="http://doi.org/10.5281/zenodo.5643809" httpUri="https://zenodo.org/record/5643809/files/figure.png" pageId="20" pageNumber="161">Fig. 9</figureCitation>
C). Head most probably blue, with a whitish stripe, less broad than in the other species of the
<taxonomicName authorityName="Dumeril &amp; Bibron" authorityYear="1837" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="161" phylum="Chordata" rank="species" species="mystaceus">
<emphasis italics="true" pageId="20" pageNumber="161">C. mystaceus</emphasis>
</taxonomicName>
complex, from behind the nasal scale along the upper lips and the tympanum to the shoulders. Followed by one distinct and one or more indistinct lateral blotches of the same color.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="163" type="materials_examined">
<paragraph blockId="22.[188,796,239,1222]" pageId="22" pageNumber="163">
<emphasis bold="true" box="[212,326,301,325]" pageId="22" pageNumber="163">Variation.</emphasis>
Body measurements and meristic characters for adult individuals are given in
<tableCitation box="[587,668,332,356]" captionStart="Table 4" captionStartId="11.[151,207,757,778]" captionTargetBox="[181,1368,845,1030]" captionTargetPageId="11" captionText="Table 4. Eigenvalues and percent of explained variance per principal component of mensural and meristic data of males and females as shown in Figs 3–4." httpUri="http://table.plazi.org/id/DEE8D89E242FFFE363B1FD592E87CC38" pageId="22" pageNumber="163" tableUuid="DEE8D89E242FFFE363B1FD592E87CC38">Table 4</tableCitation>
. Specimens ranged in size from the smallest female with a SVL of
<quantity box="[189,263,394,418]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="22" pageNumber="163" unit="mm" value="47.0">47 mm</quantity>
(CAS 232247) and the largest female with a SVL of
<quantity box="[218,293,425,449]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="22" pageNumber="163" unit="mm" value="73.0">73 mm</quantity>
(CAS 232387) to the largest male with a SVL of
<quantity box="[219,293,457,480]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="22" pageNumber="163" unit="mm" value="77.0">77 mm</quantity>
(CAS 232388). Adult males are slightly larger than females, because of longer SVLs and tail lengths, and have fewer vertebral scales including the crest scales. However, proportionally males and females are equal. Dorsal coloration of females unknown, but most probably similar to other taxa of the species group with females having lighter coloration characters (lateral stripe, blotches) than males. Most probably, both sexes are able to change their coloration.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="163" type="etymology">
<paragraph blockId="22.[188,796,239,1222]" pageId="22" pageNumber="163">
<emphasis bold="true" box="[212,340,735,759]" pageId="22" pageNumber="163">Etymology.</emphasis>
The specific epithet is a patronym honoring Jens Vindum, retired staff of the California Academy of Sciences, in respect of his outstanding contributions to the herpetology of Myanmar and SE Asia. The patronym of his family name was used, in recognition of his impressive beard which has a direct link to “mystaceus” [Greek for bearded], together with the Latin word “barbatus” for bearded.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="163" type="distribution">
<paragraph blockId="22.[188,796,239,1222]" pageId="22" pageNumber="163">
<emphasis bold="true" box="[212,358,982,1006]" pageId="22" pageNumber="163">Distribution.</emphasis>
This new species is so far only known from northern Myanmar.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="163" type="biology_ecology">
<paragraph blockId="22.[188,796,239,1222]" pageId="22" pageNumber="163">
<emphasis bold="true" box="[212,308,1044,1068]" pageId="22" pageNumber="163">Ecology.</emphasis>
<taxonomicName authority="P. &amp; F. &amp; T. &amp; M. &amp; A. &amp; W., 2021" authorityName="P. &amp; F. &amp; T. &amp; M. &amp; A. &amp; W." authorityYear="2021" box="[314,576,1044,1068]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="163" phylum="Chordata" rank="species" species="vindumbarbatus" status="sp. n.">
<emphasis box="[314,576,1044,1068]" italics="true" pageId="22" pageNumber="163">Calotes vindumbarbatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[583,641,1044,1067]" pageId="22" pageNumber="163" rank="species">sp. n.</taxonomicNameLabel>
is diurnal and arboreal to semi-arboreal. It inhabits forests and more open habitats with a dense canopy, but detailed habitat preference is unknown. As in other
<taxonomicName box="[584,665,1137,1161]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="163" phylum="Chordata" rank="species" species="undetermined">
<emphasis box="[584,665,1137,1161]" italics="true" pageId="22" pageNumber="163">Calotes</emphasis>
</taxonomicName>
species the diet most likely consists predominantly of arthropods, but details are not known.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="166" type="materials_examined">
<paragraph blockId="25.[790,1399,1297,1971]" pageId="25" pageNumber="166">
<emphasis bold="true" pageId="25" pageNumber="166">
<taxonomicName authority="P. &amp; F. &amp; T. &amp; M. &amp; A. &amp; W., 2021" authorityName="P. &amp; F. &amp; T. &amp; M. &amp; A. &amp; W." authorityYear="2021" box="[1124,1398,1849,1873]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="166" phylum="Chordata" rank="species" species="vindumbarbatus" status="sp. n.">
<emphasis bold="true" box="[1124,1398,1849,1873]" italics="true" pageId="25" pageNumber="166">Calotes vindumbarbatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[791,851,1882,1906]" pageId="25" pageNumber="166" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<emphasis box="[867,880,1882,1905]" italics="true" pageId="25" pageNumber="166">n</emphasis>
=
<specimenCount box="[910,926,1882,1906]" pageId="25" pageNumber="166">6</specimenCount>
):
<materialsCitation ID-GBIF-Occurrence="3398324320" box="[947,1093,1882,1906]" collectionCode="CAS" pageId="25" pageNumber="166" specimenCode="CAS 232247" specimenCount="1">
<specimenCode box="[947,1089,1882,1906]" pageId="25" pageNumber="166">CAS 232247</specimenCode>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3398324323" box="[1100,1246,1882,1906]" collectionCode="CAS" pageId="25" pageNumber="166" specimenCode="CAS 232387" specimenCount="1">
<specimenCode box="[1100,1242,1882,1906]" pageId="25" pageNumber="166">CAS 232387</specimenCode>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3398324324" box="[1253,1398,1882,1906]" collectionCode="CAS" pageId="25" pageNumber="166" specimenCode="CAS 232388" specimenCount="1">
<specimenCode box="[1253,1395,1882,1906]" pageId="25" pageNumber="166">CAS 232388</specimenCode>
,
</materialsCitation>
<specimenCode box="[791,934,1914,1938]" pageId="25" pageNumber="166">CAS 232389</specimenCode>
(
<materialsCitation ID-GBIF-Occurrence="3398326304" box="[952,1177,1914,1938]" collectionCode="ZFMK" pageId="25" pageNumber="166" specimenCode="ZFMK 97990" specimenCount="1">
now
<specimenCode box="[1010,1165,1914,1938]" pageId="25" pageNumber="166">ZFMK 97990</specimenCode>
),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3398324307" box="[1187,1337,1914,1938]" collectionCode="CAS" pageId="25" pageNumber="166" specimenCode="CAS 232819" specimenCount="1">
<specimenCode box="[1187,1333,1914,1938]" pageId="25" pageNumber="166">CAS 232819</specimenCode>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3398324302" collectionCode="CAS" pageId="25" pageNumber="166" specimenCode="CAS 239206" specimenCount="1">
<specimenCode pageId="25" pageNumber="166">CAS 239206</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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