198 lines
18 KiB
XML
198 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.84.67749" ID-GBIF-Dataset="0f428317-2ed5-4b7d-8144-e07e12f9cae9" ID-Pensoft-Pub="1314-2607-84-29" ID-Pensoft-UUID="BC4EE689C5DA528AA9599B83CC2489FC" ID-Zenodo-Dep="5349477" ID-ZooBank="23615AA22FA040718B26C5336CC42878" ModsDocID="1314-2607-84-29" checkinTime="1629847253858" checkinUser="pensoft" docAuthor="Belokobylskij, Sergey A., Dubovikoff, Dmitry A., Manukyan, Andranik R. & Zharkov, Dmitry M." docDate="2021" docId="3C22B366AD2C5B3F9F7239261FD15775" docLanguage="en" docName="JourHymenoptRes 84: 29-43" docOrigin="Journal of Hymenoptera Research 84" docPubDate="2021-08-24" docSource="http://dx.doi.org/10.3897/jhr.84.67749" docTitle="Elasmosomites arkadyleleji Belokobylskij 2021, sp. nov." docType="treatment" docUuid="A2326A01-D24D-4E23-91CF-7807AF9F6F30" docUuidSource="ZooBank" docVersion="4" id="BC4EE689C5DA528AA9599B83CC2489FC" lastPageNumber="29" masterDocId="BC4EE689C5DA528AA9599B83CC2489FC" masterDocTitle="Braconid parasitoids of ants (Hymenoptera, Braconidae, Euphorinae, Neoneurini) from Baltic amber with a discussion of records of fossil larvae parasitizing ant workers" masterLastPageNumber="43" masterPageNumber="29" pageNumber="29" updateTime="1678755999230" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Braconid parasitoids of ants (Hymenoptera, Braconidae, Euphorinae, Neoneurini) from Baltic amber with a discussion of records of fossil larvae parasitizing ant workers</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Belokobylskij, Sergey A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3646-3459</mods:nameIdentifier>
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<mods:affiliation>Zoological Institute of the Russian Academy of Sciences, St Petersburg 199034, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">doryctes@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dubovikoff, Dmitry A.</mods:namePart>
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<mods:affiliation>Saint Petersburg State University, 29 16 th line of Vasilievskiy Island, St Petersburg 199178, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Manukyan, Andranik R.</mods:namePart>
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<mods:affiliation>Kaliningrad Amber Museum, 1 Marshal Vasilevskiy Square, Kaliningrad 236016, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zharkov, Dmitry M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1204-1304</mods:nameIdentifier>
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<mods:affiliation>Saint Petersburg State University, 29 16 th line of Vasilievskiy Island, St Petersburg 199178, Russia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-08-24</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>84</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>43</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.84.67749</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.84.67749</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-84-29</mods:identifier>
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<mods:identifier type="ZooBank">23615AA22FA040718B26C5336CC42878</mods:identifier>
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<mods:identifier type="Pensoft-UUID">BC4EE689C5DA528AA9599B83CC2489FC</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5349477</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="184941485" LSID="urn:lsid:zoobank.org:act:A2326A01-D24D-4E23-91CF-7807AF9F6F30" httpUri="http://treatment.plazi.org/id/3C22B366AD2C5B3F9F7239261FD15775" lastPageNumber="29" pageId="0" pageNumber="29">
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<subSubSection pageId="0" pageNumber="29" type="nomenclature">
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<paragraph pageId="0" pageNumber="29">
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<taxonomicName LSID="http://zoobank.org/A2326A01-D24D-4E23-91CF-7807AF9F6F30" authority="Belokobylskij" authorityName="Belokobylskij" authorityYear="2021" class="Insecta" family="Braconidae" genus="Elasmosomites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elasmosomites arkadyleleji" order="Hymenoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="arkadyleleji" status="sp. nov.">Elasmosomites arkadyleleji Belokobylskij</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="29">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="description">
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<paragraph pageId="0" pageNumber="29">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Elasmosomites arkadyleleji sp. nov. (female, holotype, Baltic amber, # BX 9 / 21.1) A habitus, dorsal view B habitus, ventral view C antenna D head and mesosoma, dorsal view E head, mesosoma and legs, ventral view F metasoma and distal part of fore wing, dorsal view J metasoma, middle and hind legs, ventral view H fore wing." figureDoi="10.3897/jhr.84.67749.figure1" httpUri="https://binary.pensoft.net/fig/578898" pageId="0" pageNumber="29">Fig. 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="type material">
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<paragraph pageId="0" pageNumber="29">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="29">Holotype</emphasis>
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: female, Eocene Baltic amber, # BX 9/21.1 (KAM).
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</paragraph>
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<caption ID-Zenodo-Dep="5349481" doi="10.3897/jhr.84.67749.figure1" httpUri="https://binary.pensoft.net/fig/578898" pageId="0" pageNumber="29" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Figure 1.</emphasis>
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<taxonomicName authorityName="Belokobylskij" authorityYear="2021" class="Insecta" family="Braconidae" genus="Elasmosomites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elasmosomites arkadyleleji" order="Hymenoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="arkadyleleji">
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<emphasis italics="true" pageId="0" pageNumber="29">Elasmosomites arkadyleleji</emphasis>
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</taxonomicName>
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sp. nov. (female, holotype, Baltic amber, # BX 9/21.1)
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<emphasis bold="true" pageId="0" pageNumber="29">A</emphasis>
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habitus, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="29">B</emphasis>
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habitus, ventral view
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<emphasis bold="true" pageId="0" pageNumber="29">C</emphasis>
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antenna
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<emphasis bold="true" pageId="0" pageNumber="29">D</emphasis>
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head and mesosoma, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="29">E</emphasis>
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head, mesosoma and legs, ventral view
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<emphasis bold="true" pageId="0" pageNumber="29">F</emphasis>
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metasoma and distal part of fore wing, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="29">J</emphasis>
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metasoma, middle and hind legs, ventral view
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<emphasis bold="true" pageId="0" pageNumber="29">H</emphasis>
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fore wing.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="description">
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<paragraph pageId="0" pageNumber="29">Description.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Female</emphasis>
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. Body length 2.0 mm, fore wing length 1.6 mm.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="29">Head</emphasis>
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: Relatively short and transverse in dorsal view, about 2.5 times wider than medial length. Vertex densely transverse striate. Ocelli slightly enlarged, arranged in obtuse triangle with base 1.7 times its sides. POL about 1.6 times OD. Eye oval, approximately 1.4 times as high as broad, below slightly converging to each other. Transverse diameter of eye about 2.5 times length of temple. Malar suture invisible. Malar space short, about 0.3 times eye height. Face slightly convex, without facial tubercles and spines, its minimum width 0.8 times medial height with clypeus, 0.7 times eye height. Clypeus distinctly curvedly emarginated on lower margin, short, its width about 6.0 times maximum height medially, almost equal to width of face. Mandible and palps not visible. Face seemingly entirely smooth.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="29">Antenna</emphasis>
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: Slightly thickened, 14-segmented, with short and rather sparse dark setosity, about 0.7 times as long as body. Scape 1.8 times longer than maximum width, about 2.2 times longer than pedicel. Length of first flagellar segment 3.7 times its maximum apical width, 1.3 times length of scape, 1.3 times length of second segment. Length of penultimate segment 1.9 times its width, 0.7 times length of acuminate apical segment.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="29">Mesosoma</emphasis>
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: Length of mesosoma 1.6 times its maximum width. Mesoscutum 1.4 times wider than its median length, seemingly densely punctate with submedian longitudinal carina in posterior third. Prescutellar depression (scutal sulcus) relatively short, with at least two distinct submedian carinae and slightly sparsely and finely crenulate. Scutellum slightly convex, about as long as wide anteriorly, seemingly slightly punctate. Mesopleuron not visible. Prepectal carina seemingly absent. Mesosoma densely and finely granulate below.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="29">Wings</emphasis>
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: Fore wing 2.7 times longer than its maximum width, its membrane entirely and densely setose. Costal vein (C+SC+R) distinctly thickened, clearly widened towards apex. Short and vestigial costal cell present distally. Pterostigma wide and relatively short, triangular, 2.3 times longer than its maximum width. Metacarp (1-R1) distinctly and entirely sclerotised, 0.8 times as long as pterostigma. Radial vein (r) arising from basal 0.3 of pterostigma. Radial (marginal) cell distinctly shortened, 3.7 times longer than its maximum width, without additional transverse vein. First radial abscissa (r) long and vertical to pterostigma, 0.6 times as long as maximum width of pterostigma. Second radial abscissa (3-SR) short, 0.8 times as long as first abscissa (r), 0.12 times as long as almost straight third abscissa (SR1), 0.7 times as long as first radiomedial vein (2-SR). Third radial abscissa (SR1) almost complete, but well sclerotised only in basal 0.5 and desclerotised in remaining apical part. Second radiomedial (submarginal) cell rather small, completely delineated by almost entirely sclerotised veins, 1.4 times longer than wide. Second radiomedial vein (r-m) straight, subvertical, desclerotised medially, 0.9 times as long as first radiomedial vein (2-SR). Discoidal (discal) cell with relatively long petiole (1-SR) anteriorly, 1.4 times longer than maximum height; petiole (1-SR) not thickened; parastigma enlarged. First medial abscissa (1-SR+M) almost straight. Recurrent vein (m-cu) short, strongly postfurcal, about 1.5 times longer than second medial abscissa (2-SR+M), 0.9 times as long as first radiomedial vein (2-SR) and second radial abscissa (3-SR). Nervulus (cu-a) strongly postfurcal, distance (1-CU1) between nervulus (cu-a) and basal vein (1-M) equal to nervulus (cu-a) length. Parallel vein (CU1a) distinctly broken basally. Hind wing not visible.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="29">Legs</emphasis>
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: Fore coxa subglobular. Trochantelli short and distinctly separated from femora on all legs. Fore femur rather wide, almost 3.0 times longer than wide, without tooth. Fore tibia without protuberances or processes, distinctly widened towards apex, its length 4.7 times maximum width. Fore tarsus slightly longer than fore tibia. Fore tibial spur almost straight, about 0.8 times as long as fore basitarsus. Tarsal segments of middle leg elongate. Hind femur wide, elongate-oval, about 2.8 times longer than its maximum width. Hind tibia distinctly widened towards apex, about 6.0 times longer than maximum width, 1.1 times longer than femur, 0.9 times as long as hind tarsus. Both spurs of hind tibia acute apically, longest spur about as long as hind basitarsus. Hind tarsus relatively slender, all its segments of similar width and not narrowed towards apex.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="29">Metasoma</emphasis>
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: Almost twice longer than maximum width, 1.3 times longer than mesosoma; its sculpture and pubescence not visible (dorsally covered by wings). First tergite distinctly widened towards apex; its median length about 0.8 times maximum distal width. Segments behind third one distinctly projected; any sutures, if present, not visible. Sternites with distinct median keel. Ovipositor relatively short, its sheath with dense short setae, 0.4 times as long as hind tibia, about 0.2 times as long as metasoma.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="29">Colour</emphasis>
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: Body reddish brown, with a few dark spots, metasoma in posterior half pale brown. Fore and middle legs yellowish brown, hind leg reddish brown. Wings faintly and evenly infuscate; pterostigma dark brown medially and paler basally and apically.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Male.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="comparative diagnosis">
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<paragraph pageId="0" pageNumber="29">Comparative diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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The new species differs from the type species
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<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. primordialis" order="Hymenoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="primordialis">
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<emphasis italics="true" pageId="0" pageNumber="29">E. primordialis</emphasis>
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</taxonomicName>
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Brues, 1933, by having the body mainly reddish brown and metasoma in posterior half pale brown (entirely black in
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<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. primordialis" order="Hymenoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="primordialis">
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<emphasis italics="true" pageId="0" pageNumber="29">E. primordialis</emphasis>
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</taxonomicName>
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), antenna basally distinctly pale (entirely black in
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<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. primordialis" order="Hymenoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="primordialis">
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<emphasis italics="true" pageId="0" pageNumber="29">E. primordialis</emphasis>
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</taxonomicName>
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), fore and middle legs yellowish brown (almost black or dark brown in
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<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. primordialis" order="Hymenoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="primordialis">
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<emphasis italics="true" pageId="0" pageNumber="29">E. primordialis</emphasis>
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</taxonomicName>
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), length of first flagellar segment 3.7 times its maximum width (2.8 times in
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<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. primordialis" order="Hymenoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="primordialis">
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<emphasis italics="true" pageId="0" pageNumber="29">E. primordialis</emphasis>
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</taxonomicName>
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), recurrent vein (m-cu) of fore wing longer, about 1.5 times longer than second medial abscissa (2-SR+M) (shorter, not longer than second medial abscissa (2-SR+M) in
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<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. primordialis" order="Hymenoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="primordialis">
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<emphasis italics="true" pageId="0" pageNumber="29">E. primordialis</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="etymology">
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<paragraph pageId="0" pageNumber="29">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="29">This species is named in honour of Professor Arkady Stepanovich Lelej, the well-known Russian hymenopterist, dedicated in celebration of his 75th birthday.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="remarks">
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<paragraph pageId="0" pageNumber="29">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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Most extant species of the tribe
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<taxonomicName baseAuthorityName="Bengtsson" baseAuthorityYear="1918" lsidName="" pageId="0" pageNumber="29" rank="tribe" tribe="Neoneurini">Neoneurini</taxonomicName>
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are predominantly black in colour, only sometimes legs entirely pale or with a contrasting black and white pattern.
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<taxonomicName authorityName="Belokobylskij" authorityYear="2021" class="Insecta" family="Braconidae" genus="Elasmosomites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elasmosomites arkadyleleji" order="Hymenoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="arkadyleleji">
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<emphasis italics="true" pageId="0" pageNumber="29">Elasmosomites arkadyleleji</emphasis>
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</taxonomicName>
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sp. nov. is the first species of the
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<taxonomicName baseAuthorityName="Bengtsson" baseAuthorityYear="1918" lsidName="" pageId="0" pageNumber="29" rank="tribe" tribe="Neoneurini">Neoneurini</taxonomicName>
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with mainly reddish brown body.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |