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<document id="65279FC459CA162711C397C1A4173E86" ID-CLB-Dataset="3916" ID-DOI="10.11646/zootaxa.3879.1.1" ID-GBIF-Dataset="4b976ba3-1a7b-4dd5-8afb-630b7cc8a695" ID-ISSN="1175-5326" ID-Zenodo-Dep="4949051" ID-ZooBank="6423894B-97D9-4286-ABB9-D4AF072B57FD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623702926238" checkinUser="felipe" docAuthor="Borkent, Art" docDate="2014" docId="027587C9BD79302FFD521C6A4F11E679" docLanguage="en" docName="zt03879p327.pdf.imf" docOrigin="Zootaxa 3879 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Nilobezzia Kieffer" docType="treatment" docVersion="10" lastPageNumber="90" masterDocId="FE4CFFB1BD2E3076FFF81A154D70E204" masterDocTitle="The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera" masterLastPageNumber="327" masterPageNumber="1" pageNumber="88" updateTime="1699015668130" updateUser="ExternalLinkService">
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<mods:title id="79B81F3060D4A81234F8542C497AD6CC">The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera</mods:title>
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<mods:namePart id="C480EFB72588D28AB18EB78B1E77808E">Borkent, Art</mods:namePart>
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<mods:date id="7DAFB06379A42E5848A274F88EADE043">2014</mods:date>
<mods:detail id="B237B1D700D77C00D9ABE22A53976C28" type="pubDate">
<mods:number id="C2E66DBE0B28AA66A0049DEBE941B942">2014-10-31</mods:number>
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<mods:number id="777364641427DBC2F9553A60432F5272">3879</mods:number>
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<mods:classification id="C707EAEF1D524A40952A50498447D597">journal article</mods:classification>
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<mods:identifier id="BB9666F8582766C48CE007724559F160" type="DOI">10.11646/zootaxa.3879.1.1</mods:identifier>
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<mods:identifier id="DC488BBAA1778976CF57857F2DE440C8" type="ISSN">1175-5326</mods:identifier>
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<treatment id="027587C9BD79302FFD521C6A4F11E679" ID-DOI="http://doi.org/10.5281/zenodo.5593040" ID-GBIF-Taxon="189161026" ID-Zenodo-Dep="5593040" LSID="urn:lsid:plazi:treatment:027587C9BD79302FFD521C6A4F11E679" httpUri="http://treatment.plazi.org/id/027587C9BD79302FFD521C6A4F11E679" lastPageId="89" lastPageNumber="90" pageId="87" pageNumber="88">
<subSubSection id="C2C66554BD793021FD521C6A4EF8E49E" box="[682,904,1663,1690]" pageId="87" pageNumber="88" type="nomenclature">
<paragraph id="8A6336DFBD793021FD521C6A4EF8E49E" blockId="87.[457,1130,1663,1723]" box="[682,904,1663,1690]" pageId="87" pageNumber="88">
<taxonomicName id="4DDC4D5CBD793021FD521C6A4EF8E49E" ID-CoL="6469" ID-ENA="1874307" authority="Kieffer" authorityName="Kieffer" box="[682,904,1663,1690]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD793021FD521C6A4EF8E49E" bold="true" box="[682,904,1663,1690]" pageId="87" pageNumber="88">
<emphasis id="B8A8EACDBD793021FD521C6A4E57E49D" bold="true" box="[682,807,1663,1689]" italics="true" pageId="87" pageNumber="88">Nilobezzia</emphasis>
Kieffer
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD793021FE311CB7491AE4BF" box="[457,1130,1698,1723]" pageId="87" pageNumber="88" type="description">
<paragraph id="8A6336DFBD793021FE311CB7491AE4BF" blockId="87.[457,1130,1663,1723]" box="[457,1130,1698,1723]" pageId="87" pageNumber="88">
(
<figureCitation id="12E72A5ABD793021FE291CB74F2AE4BF" box="[465,602,1698,1723]" captionStart="FIGURE 21" captionStartId="266.[151,250,1837,1859]" captionTargetBox="[182,1405,192,1792]" captionTargetId="figure-122@266.[160,1419,186,1818]" captionTargetPageId="266" captionText="FIGURE 21. Dorsal apotomes, anterior view. A. Dibezzia prominens (from Mayer 1934c). B. Hebetula tonnoiri (from Elson- Harris 1987). C. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). D. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). E. Mallochohelea termophila (from Spinelli 1984, modified). F. Probezzia pallida (from Wirth &amp; Grogan 1979). G. Neobezzia fittkaui (from Ronderos et al. 2011, modified). H. Nilobezzia basispinigera (from Elson-Harris 1987). I. Nilobezzia virago (from Elson-Harris 1987). J. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). K. Lanatomyia electra (from Elson-Harris 1987). L. Anebomyia atripes (from Wirth &amp; Grogan 1979, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949136" httpUri="https://zenodo.org/record/4949136/files/figure.png" pageId="87" pageNumber="88">Figs. 21H–I</figureCitation>
,
<figureCitation id="12E72A5ABD793021FD9E1CB74FE8E4BF" box="[614,664,1698,1723]" captionStart="FIGURE 27" captionStartId="272.[151,250,1950,1972]" captionTargetBox="[198,1418,210,1940]" captionTargetId="figure-69@272.[190,1421,205,1944]" captionTargetPageId="272" captionText="FIGURE 27. Posterior portion of heads, in ventral view. A. Mallochohelea caudellii. B. Probezzia sabroskyi. C. Neobezzia fittkaui. D. Nilobezzia curticornis. E. Johannsenomyia argentata. F. Lanatomyia electra. G. Anebomyia atripes. H. Homohelea delanoe." figureDoi="http://doi.org/10.5281/zenodo.4949164" httpUri="https://zenodo.org/record/4949164/files/figure.png" pageId="87" pageNumber="88">27D</figureCitation>
,
<figureCitation id="12E72A5ABD793021FD5C1CB74FA7E4BF" box="[676,727,1698,1723]" captionStart="FIGURE 30" captionStartId="275.[151,250,1839,1861]" captionTargetBox="[161,1425,193,1797]" captionTargetId="figure-119@275.[161,1426,193,1813]" captionTargetPageId="275" captionText="FIGURE 30. Dorsal setae from left side of thoraces. A. Serromyia sp. B. Allohelea japonica. C. Monohelea obscura. D. Atyphohelea macroneura. E. Parabezzia huberti. F. Echinohelea lanei. G. Clinohelea curriei. H. Heteromyia clavata. I. Pellucidomyia leei. J. Dibezzia debenhamae (dirty specimen, D-5-T perhaps broken setal socket). K. Hebetula tonnoiri. L. Jenkinshelea magnipennis. M. Macropeza natalensis. N. Mallochohelea caudellii. O. Probezzia sabroskyi. P. Neobezzia fittkaui (from Ronderos et al. 2011, modified). Q. Nilobezzia curticornis. R. Johannsenomyia argentata. S. Lanatomyia miles. T. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949174" httpUri="https://zenodo.org/record/4949174/files/figure.png" pageId="87" pageNumber="88">30Q</figureCitation>
,
<figureCitation id="12E72A5ABD793021FD1A1CB74E65E4BF" box="[738,789,1698,1723]" captionStart="FIGURE 33" captionStartId="278.[151,250,1883,1905]" captionTargetBox="[176,1420,177,1860]" captionTargetId="figure-110@278.[176,1420,177,1860]" captionTargetPageId="278" captionText="FIGURE 33. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Parabezzia sp. (5 km. E. Danby, Vermont, USA). B. Clinohelea curriei. C. Pellucidomyia leei. D. Hebetula tonnoiri. E. Jenkinshelea sp. (from Ottawa, ON, Canada). F. Mallochohelea caudellii. G. Probezzia seminigra. H. Nilobezzia sp. (from Kruger NP, South Africa). I. Sphaeromias longipennis. J. Bezzia sp. (6 km E. Salmon Arm, BC, Canada) K. Palpomyia sp. (Black Lake, Quebec, Canada). L. Phaenobezzia sp. (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949184" httpUri="https://zenodo.org/record/4949184/files/figure.png" pageId="87" pageNumber="88">33H</figureCitation>
,
<figureCitation id="12E72A5ABD793021FCD81CB74E22E4BF" box="[800,850,1698,1723]" captionStart="FIGURE 39" captionStartId="284.[151,250,1973,1995]" captionTargetBox="[180,1379,215,1945]" captionTargetId="figure-65@284.[170,1401,210,1949]" captionTargetPageId="284" captionText="FIGURE 39. Legs, wings and apices of the antennae, in ventral view. A. Probezzia sabroskyi. B. Neobezzia fittkaui. C. Nilobezzia sp. (from Australia). D. Johannsenomyia argentata. E. Lanatomyia electra. F. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949198" httpUri="https://zenodo.org/record/4949198/files/figure.png" pageId="87" pageNumber="88">39C</figureCitation>
,
<figureCitation id="12E72A5ABD793021FCA51CB74EDBE4BF" box="[861,939,1698,1723]" captionStart="FIGURE 45" captionStartId="290.[151,250,1738,1760]" captionTargetBox="[222,1373,186,1678]" captionTargetId="figure-193@290.[202,1384,184,1720]" captionTargetPageId="290" captionText="FIGURE 45. Respiratory organs, in dorsal view, pattern on tracheal tubes shown only in some. A. Clinohelea tasmaniensis (from Elson-Harris &amp; Kettle 1986). B. Clinohelea bimaculata (from Wirth &amp; Grogan 1979, modified). C. Heteromyia wokei (from Wirth &amp; Grogan 1977). D. Heteromyia clavata (from Wirth &amp; Grogan 1977). E. Pellucidomyia leei (from Elson-Harris &amp; Kettle 1986). F. Pellucidomyia geari (from de Meillon &amp; Wirth 1981). G. Dibezzia prominens (from Mayer 1934c). H. Hebetula tonnoiri (from Debenham 1974). I. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). J. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). K. Mallochohelea albihalter (from Wirth &amp; Grogan 1979). L. Probezzia pallida (from Wirth &amp; Grogan 1979). M. Probezzia xanthogaster (from Wirth &amp; Grogan 1979). N. Neobezzia fittkaui (from Ronderos et al. 2011). O. Nilobezzia aranea (from Elson-Harris 1987). P. Nilobezzia virago (from Elson-Harris 1987). Q. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). R. Lanatomyia electra (from Debenham 1974). S. Lanatomyia miles (from Debenham 1974). T. Anebomyia atripes (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949212" httpUri="https://zenodo.org/record/4949212/files/figure.png" pageId="87" pageNumber="88">45O–P</figureCitation>
,
<figureCitation id="12E72A5ABD793021FC4E1CB74E97E4BF" box="[950,999,1698,1723]" captionStart="FIGURE 52" captionStartId="297.[151,250,1942,1964]" captionTargetBox="[190,1402,202,1928]" captionTargetId="figure-58@297.[181,1412,193,1932]" captionTargetPageId="297" captionText="FIGURE 52. Metathoraces and tergites 1, in dorsal view. A. Neobezzia fittkaui. B. Nilobezzia curticornis. C. Johannsenomyia argentata. D. Lanatomyia miles. E. Anebomyia atripes. F. Homohelea delanoe (reconstructed from distorted and dirty specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949234" httpUri="https://zenodo.org/record/4949234/files/figure.png" pageId="87" pageNumber="88">52B</figureCitation>
,
<figureCitation id="12E72A5ABD793021FC0A1CB74955E4BF" box="[1010,1061,1698,1723]" captionStart="FIGURE 67" captionStartId="312.[151,250,1975,1997]" captionTargetBox="[198,1410,217,1946]" captionTargetId="figure-53@312.[189,1420,211,1950]" captionTargetPageId="312" captionText="FIGURE 67. Abdominal segments four, in dorsal and ventral view (shagreen not shown). A. Nilobezzia curticornis. B. Johannsenomyia argentata. C. Lanatomyia miles." figureDoi="http://doi.org/10.5281/zenodo.4949288" httpUri="https://zenodo.org/record/4949288/files/figure.png" pageId="87" pageNumber="88">67A</figureCitation>
,
<figureCitation id="12E72A5ABD793021FBC91CB74913E4BF" box="[1073,1123,1698,1723]" captionStart="FIGURE 76" captionStartId="321.[151,250,1765,1787]" captionTargetBox="[225,1365,181,1745]" captionTargetId="figure-185@321.[215,1375,177,1746]" captionTargetPageId="321" captionText="FIGURE 76. Abdominal segments 9. A. Macropeza albitarsis, male, in ventral view (modified from Szadziewski &amp; Dominiak, 2007). B. Macropeza albitarsis, female, in ventral view (from Szadziewski &amp; Dominiak, 2007). C. Mallochohelea termophila, male, in ventral view (from Spinelli 1984). D. Mallochohelea termophila, female, in dorsal view (from Spinelli 1984). E. Probezzia pallida, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). F. Neobezzia fittkaui, male, in dorsal view (from Ronderos, et al. 2011). G. Nilobezzia virago, female, in dorsal view (from Elson-Harris 1987). H. Johannsenomyia argentata, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). I. Johannsenomyia argentata, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979). J. Lanatomyia miles, female, in dorsal view (from Elson- Harris 1987). K. Anebomyia atripes, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). L. Anebomyia atripes, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949324" httpUri="https://zenodo.org/record/4949324/files/figure.png" pageId="87" pageNumber="88">76G</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD793021FF6F1CF14E83E56C" pageId="87" pageNumber="88" type="diagnosis">
<paragraph id="8A6336DFBD793021FF6F1CF14E83E56C" blockId="87.[151,1437,1764,2004]" pageId="87" pageNumber="88">
<emphasis id="B8A8EACDBD793021FF6F1CF14C44E4F9" bold="true" box="[151,308,1764,1789]" pageId="87" pageNumber="88">DIAGNOSIS</emphasis>
: Only pupa of
<taxonomicName id="4DDC4D5CBD793021FE271CF14FEEE4F9" authorityName="Borkent" authorityYear="2014" box="[479,670,1764,1789]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
with the metathorax with only one campaniform sensillum (M-3-T) situated at least ⅓ the length of the metathorax from its anterior margin (
<figureCitation id="12E72A5ABD793021FC2E1D12494EE524" box="[982,1086,1799,1824]" captionStart="FIGURE 52" captionStartId="297.[151,250,1942,1964]" captionTargetBox="[190,1402,202,1928]" captionTargetId="figure-58@297.[181,1412,193,1932]" captionTargetPageId="297" captionText="FIGURE 52. Metathoraces and tergites 1, in dorsal view. A. Neobezzia fittkaui. B. Nilobezzia curticornis. C. Johannsenomyia argentata. D. Lanatomyia miles. E. Anebomyia atripes. F. Homohelea delanoe (reconstructed from distorted and dirty specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949234" httpUri="https://zenodo.org/record/4949234/files/figure.png" pageId="87" pageNumber="88">Fig. 52B</figureCitation>
), thorax with D-5-T a peg-like seta (as in
<figureCitation id="12E72A5ABD793021FEF51D394C1DE541" box="[269,365,1836,1861]" captionStart="FIGURE 30" captionStartId="275.[151,250,1839,1861]" captionTargetBox="[161,1425,193,1797]" captionTargetId="figure-119@275.[161,1426,193,1813]" captionTargetPageId="275" captionText="FIGURE 30. Dorsal setae from left side of thoraces. A. Serromyia sp. B. Allohelea japonica. C. Monohelea obscura. D. Atyphohelea macroneura. E. Parabezzia huberti. F. Echinohelea lanei. G. Clinohelea curriei. H. Heteromyia clavata. I. Pellucidomyia leei. J. Dibezzia debenhamae (dirty specimen, D-5-T perhaps broken setal socket). K. Hebetula tonnoiri. L. Jenkinshelea magnipennis. M. Macropeza natalensis. N. Mallochohelea caudellii. O. Probezzia sabroskyi. P. Neobezzia fittkaui (from Ronderos et al. 2011, modified). Q. Nilobezzia curticornis. R. Johannsenomyia argentata. S. Lanatomyia miles. T. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949174" httpUri="https://zenodo.org/record/4949174/files/figure.png" pageId="87" pageNumber="88">Fig. 30P</figureCitation>
) (or a short, stout seta, only in
<taxonomicName id="4DDC4D5CBD793021FD301D384E13E540" baseAuthorityName="Kieffer" baseAuthorityYear="1917" box="[712,867,1837,1860]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="curticornis">
<emphasis id="B8A8EACDBD793021FD301D384E13E540" box="[712,867,1837,1860]" italics="true" pageId="87" pageNumber="88">N. curticornis</emphasis>
</taxonomicName>
, then on a well-developed tubercle,
<figureCitation id="12E72A5ABD793021FB011D394810E541" box="[1273,1376,1836,1861]" captionStart="FIGURE 30" captionStartId="275.[151,250,1839,1861]" captionTargetBox="[161,1425,193,1797]" captionTargetId="figure-119@275.[161,1426,193,1813]" captionTargetPageId="275" captionText="FIGURE 30. Dorsal setae from left side of thoraces. A. Serromyia sp. B. Allohelea japonica. C. Monohelea obscura. D. Atyphohelea macroneura. E. Parabezzia huberti. F. Echinohelea lanei. G. Clinohelea curriei. H. Heteromyia clavata. I. Pellucidomyia leei. J. Dibezzia debenhamae (dirty specimen, D-5-T perhaps broken setal socket). K. Hebetula tonnoiri. L. Jenkinshelea magnipennis. M. Macropeza natalensis. N. Mallochohelea caudellii. O. Probezzia sabroskyi. P. Neobezzia fittkaui (from Ronderos et al. 2011, modified). Q. Nilobezzia curticornis. R. Johannsenomyia argentata. S. Lanatomyia miles. T. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949174" httpUri="https://zenodo.org/record/4949174/files/figure.png" pageId="87" pageNumber="88">Fig. 30Q</figureCitation>
), and abdominal segment 1 with D-2-I and D-3-I closely approximated (
<figureCitation id="12E72A5ABD793021FC781D5A4E97E56C" box="[896,999,1871,1896]" captionStart="FIGURE 52" captionStartId="297.[151,250,1942,1964]" captionTargetBox="[190,1402,202,1928]" captionTargetId="figure-58@297.[181,1412,193,1932]" captionTargetPageId="297" captionText="FIGURE 52. Metathoraces and tergites 1, in dorsal view. A. Neobezzia fittkaui. B. Nilobezzia curticornis. C. Johannsenomyia argentata. D. Lanatomyia miles. E. Anebomyia atripes. F. Homohelea delanoe (reconstructed from distorted and dirty specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949234" httpUri="https://zenodo.org/record/4949234/files/figure.png" pageId="87" pageNumber="88">Fig. 52B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD79302EFF3F1D6148EDE7DF" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88" type="description">
<paragraph id="8A6336DFBD79302EFF3F1D6148EDE7DF" blockId="87.[151,1437,1764,2004]" lastBlockId="88.[151,1437,151,2035]" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88">
<emphasis id="B8A8EACDBD793021FF3F1D614CFAE589" bold="true" box="[199,394,1908,1933]" pageId="87" pageNumber="88">DESCRIPTION</emphasis>
: Total length =
<quantity id="4D249B3ABD793021FDBD1D614F9DE58F" box="[581,749,1908,1933]" metricMagnitude="-3" metricUnit="m" metricValue="5.19" metricValueMax="7.16" metricValueMin="3.22" pageId="87" pageNumber="88" unit="mm" value="5.19" valueMax="7.16" valueMin="3.22">3.22–7.16 mm</quantity>
. Without larval exuviae retained on abdomen. Exuviae with flagellum appressed against lateral margin of midleg, wing (as in
<figureCitation id="12E72A5ABD793021FC741D824E8EE5B4" box="[908,1022,1943,1968]" captionStart="FIGURE 16" captionStartId="261.[151,250,1897,1919]" captionTargetBox="[172,1416,199,1871]" captionTargetId="figure-85@261.[172,1416,193,1876]" captionTargetPageId="261" captionText="FIGURE 16. Anterior portion of heads. A. Parabezzia huberti, in ventral view (distorted slide specimen). B. Parabezzia sp. (5 km. E. Danby, Vermont, USA), ventrolateral view. C. Clinohelea curriei, in ventral view. D. Heteromyia clavata, in ventral view, dorsal apotome not present. E. Heteromyia wokei, in ventral, slightly lateral view. F. Pellucidomyia leei, in ventral view, dorsal apotome not present." figureDoi="http://doi.org/10.5281/zenodo.4949118" httpUri="https://zenodo.org/record/4949118/files/figure.png" pageId="87" pageNumber="88">Figs. 16B</figureCitation>
,
<figureCitation id="12E72A5ABD793021FBF41D824930E5B4" box="[1036,1088,1943,1968]" captionStart="FIGURE 33" captionStartId="278.[151,250,1883,1905]" captionTargetBox="[176,1420,177,1860]" captionTargetId="figure-110@278.[176,1420,177,1860]" captionTargetPageId="278" captionText="FIGURE 33. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Parabezzia sp. (5 km. E. Danby, Vermont, USA). B. Clinohelea curriei. C. Pellucidomyia leei. D. Hebetula tonnoiri. E. Jenkinshelea sp. (from Ottawa, ON, Canada). F. Mallochohelea caudellii. G. Probezzia seminigra. H. Nilobezzia sp. (from Kruger NP, South Africa). I. Sphaeromias longipennis. J. Bezzia sp. (6 km E. Salmon Arm, BC, Canada) K. Palpomyia sp. (Black Lake, Quebec, Canada). L. Phaenobezzia sp. (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949184" httpUri="https://zenodo.org/record/4949184/files/figure.png" pageId="87" pageNumber="88">33H</figureCitation>
). Ecdysial tear around base of antenna, along lateral margin of face to palpus (as in
<figureCitation id="12E72A5ABD793021FD0A1DAE4E13E5D0" box="[754,867,1979,2004]" captionStart="FIGURE 17" captionStartId="262.[151,250,1910,1932]" captionTargetBox="[158,1430,178,1891]" captionTargetId="figure-91@262.[158,1431,175,1896]" captionTargetPageId="262" captionText="FIGURE 17. Anterior portion of heads. A. Jenkinshelea sp., in ventral view (from Ottawa, ON, Canada). B. Mallochohelea caudellii., in ventral view. C. Sphaeromias longipennis, in ventral view. D. Bezzia sp., in ventral view (6 km E Salmon Arm, BC, Canada). E. Palpomyia sp. (Black Lake, Quebec, Canada), in ventral view. F. Phaenobezzia sp., in ventral view (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949120" httpUri="https://zenodo.org/record/4949120/files/figure.png" pageId="87" pageNumber="88">Figs. 17C</figureCitation>
,
<figureCitation id="12E72A5ABD793021FC971DAE4ED4E5D0" box="[879,932,1979,2004]" captionStart="FIGURE 79" captionStartId="324.[151,250,1861,1883]" captionTargetBox="[192,1396,201,1846]" captionTargetId="figure-148@324.[184,1406,195,1847]" captionTargetPageId="324" captionText="FIGURE 79. Evolution of ecdysial sutures and splits in pupal exuviae of Ceratopogonidae, head and anterior portion of thorax in ventral view. Numbers refer to character states of characters 2 and 8 in text. Red indicates ecdysial splits. A. Leptoconopinae. B. Forcipomyiinae. C. Dasyheleinae. D. Most Ceratopogonini. E. Most Ceratopogonini, Clinohelea. F. Allohelea, Atyphohelea, Parabezzia. G. Heteromyia, Pellucidomyia, Hebetula. H. most Sphaeromiini s. lat., most Palpomyiini, Paryphoconus; some Bezzia are nearly completed fused (as in Fig. 79I), with only the dorsal apotome separate, along with an ecdysial split along the base of the antenna and the dorsal thoracic split. I. Jenkinshelea, Macropeza, Mallochohelea, Probezzia, Neobezzia." figureDoi="http://doi.org/10.5281/zenodo.4949336" httpUri="https://zenodo.org/record/4949336/files/figure.png" pageId="87" pageNumber="88">79H</figureCitation>
).
<emphasis id="B8A8EACDBD793021FC4F1DAE4E87E5D0" bold="true" box="[951,1015,1979,2004]" pageId="87" pageNumber="88">Head</emphasis>
: Dorsal apotome (
<figureCitation id="12E72A5ABD793021FB321DAE4826E5D0" box="[1226,1366,1979,2004]" captionStart="FIGURE 21" captionStartId="266.[151,250,1837,1859]" captionTargetBox="[182,1405,192,1792]" captionTargetId="figure-122@266.[160,1419,186,1818]" captionTargetPageId="266" captionText="FIGURE 21. Dorsal apotomes, anterior view. A. Dibezzia prominens (from Mayer 1934c). B. Hebetula tonnoiri (from Elson- Harris 1987). C. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). D. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). E. Mallochohelea termophila (from Spinelli 1984, modified). F. Probezzia pallida (from Wirth &amp; Grogan 1979). G. Neobezzia fittkaui (from Ronderos et al. 2011, modified). H. Nilobezzia basispinigera (from Elson-Harris 1987). I. Nilobezzia virago (from Elson-Harris 1987). J. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). K. Lanatomyia electra (from Elson-Harris 1987). L. Anebomyia atripes (from Wirth &amp; Grogan 1979, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949136" httpUri="https://zenodo.org/record/4949136/files/figure.png" pageId="87" pageNumber="88">Figs. 21H–I</figureCitation>
), with ventral line of weakness, without dorsomedial tubercle, without central dome; dorsolateral cephalic sclerite (as in
<figureCitation id="12E72A5ABD76302EFF6F1AAE4D8CE2D0" box="[151,252,187,212]" captionStart="FIGURE 13" captionStartId="258.[151,250,1897,1919]" captionTargetBox="[172,1416,193,1876]" captionTargetId="figure-99@258.[172,1416,193,1876]" captionTargetPageId="258" captionText="FIGURE 13. Anterior portion of heads and thoraces, in anterior or anterodorsal view. A. Leptoconops kerteszi, ecdysial sutures of head enhanced to show position. B. Culicoides denticulatus. C. Ceratopogon nr. abstrusus. D. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). E. Alluaudomyia bella, dorsal apotome absent. F. Clinohelea curriei. G. Pellucidomyia leei. H. Mallochohelea caudellii. I. Bezzia sp. (from Bolean Lake, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949104" httpUri="https://zenodo.org/record/4949104/files/figure.png" pageId="88" pageNumber="89">Fig. 13H</figureCitation>
) fused to scutum, each side separated medially by dorsal apotome in whole pupa; mouthparts (
<figureCitation id="12E72A5ABD76302EFAD61AAE48E4E2D0" box="[1326,1428,187,212]" captionStart="FIGURE 27" captionStartId="272.[151,250,1950,1972]" captionTargetBox="[198,1418,210,1940]" captionTargetId="figure-69@272.[190,1421,205,1944]" captionTargetPageId="272" captionText="FIGURE 27. Posterior portion of heads, in ventral view. A. Mallochohelea caudellii. B. Probezzia sabroskyi. C. Neobezzia fittkaui. D. Nilobezzia curticornis. E. Johannsenomyia argentata. F. Lanatomyia electra. G. Anebomyia atripes. H. Homohelea delanoe." figureDoi="http://doi.org/10.5281/zenodo.4949164" httpUri="https://zenodo.org/record/4949164/files/figure.png" pageId="88" pageNumber="89">Fig. 27D</figureCitation>
) with mandible well-developed, lacinia absent; palpus extending posterior to posterolateral margin of labium; labium separated medially by labrum, hypopharynx; apex of antenna (
<figureCitation id="12E72A5ABD76302EFC321B164942E318" box="[970,1074,259,284]" captionStart="FIGURE 39" captionStartId="284.[151,250,1973,1995]" captionTargetBox="[180,1379,215,1945]" captionTargetId="figure-65@284.[170,1401,210,1949]" captionTargetPageId="284" captionText="FIGURE 39. Legs, wings and apices of the antennae, in ventral view. A. Probezzia sabroskyi. B. Neobezzia fittkaui. C. Nilobezzia sp. (from Australia). D. Johannsenomyia argentata. E. Lanatomyia electra. F. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949198" httpUri="https://zenodo.org/record/4949198/files/figure.png" pageId="88" pageNumber="89">Fig. 39C</figureCitation>
) anterior to posterior extent of midlength portion of midleg (portion lateral to mesosternum), narrowed posteriorly; sensilla: dorsal apotomals (
<figureCitation id="12E72A5ABD76302EFF671B5F4C40E367" box="[159,304,330,355]" captionStart="FIGURE 21" captionStartId="266.[151,250,1837,1859]" captionTargetBox="[182,1405,192,1792]" captionTargetId="figure-122@266.[160,1419,186,1818]" captionTargetPageId="266" captionText="FIGURE 21. Dorsal apotomes, anterior view. A. Dibezzia prominens (from Mayer 1934c). B. Hebetula tonnoiri (from Elson- Harris 1987). C. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). D. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). E. Mallochohelea termophila (from Spinelli 1984, modified). F. Probezzia pallida (from Wirth &amp; Grogan 1979). G. Neobezzia fittkaui (from Ronderos et al. 2011, modified). H. Nilobezzia basispinigera (from Elson-Harris 1987). I. Nilobezzia virago (from Elson-Harris 1987). J. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). K. Lanatomyia electra (from Elson-Harris 1987). L. Anebomyia atripes (from Wirth &amp; Grogan 1979, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949136" httpUri="https://zenodo.org/record/4949136/files/figure.png" pageId="88" pageNumber="89">Figs. 21H–I</figureCitation>
)—1 elongate seta, 1 campaniform sensillum; dorsolateral cephalic sclerite sensilla—1 seta, 1 campaniform sensillum; clypeal-labrals (
<figureCitation id="12E72A5ABD76302EFD9F1B7B4FBFE383" box="[615,719,366,391]" captionStart="FIGURE 27" captionStartId="272.[151,250,1950,1972]" captionTargetBox="[198,1418,210,1940]" captionTargetId="figure-69@272.[190,1421,205,1944]" captionTargetPageId="272" captionText="FIGURE 27. Posterior portion of heads, in ventral view. A. Mallochohelea caudellii. B. Probezzia sabroskyi. C. Neobezzia fittkaui. D. Nilobezzia curticornis. E. Johannsenomyia argentata. F. Lanatomyia electra. G. Anebomyia atripes. H. Homohelea delanoe." figureDoi="http://doi.org/10.5281/zenodo.4949164" httpUri="https://zenodo.org/record/4949164/files/figure.png" pageId="88" pageNumber="89">Fig. 27D</figureCitation>
)—0–1 slender or 2 thick setae; oculars (
<figureCitation id="12E72A5ABD76302EFB5B1B7B4879E383" box="[1187,1289,366,391]" captionStart="FIGURE 27" captionStartId="272.[151,250,1950,1972]" captionTargetBox="[198,1418,210,1940]" captionTargetId="figure-69@272.[190,1421,205,1944]" captionTargetPageId="272" captionText="FIGURE 27. Posterior portion of heads, in ventral view. A. Mallochohelea caudellii. B. Probezzia sabroskyi. C. Neobezzia fittkaui. D. Nilobezzia curticornis. E. Johannsenomyia argentata. F. Lanatomyia electra. G. Anebomyia atripes. H. Homohelea delanoe." figureDoi="http://doi.org/10.5281/zenodo.4949164" httpUri="https://zenodo.org/record/4949164/files/figure.png" pageId="88" pageNumber="89">Fig. 27D</figureCitation>
)—2 setae, 1 campaniform sensillum.
<emphasis id="B8A8EACDBD76302EFE341B874F55E3AF" bold="true" box="[460,549,402,427]" pageId="88" pageNumber="89">Thorax</emphasis>
: Prothoracic extension (
<figureCitation id="12E72A5ABD76302EFC9E1B874EACE3AF" box="[870,988,402,427]" captionStart="FIGURE 27" captionStartId="272.[151,250,1950,1972]" captionTargetBox="[198,1418,210,1940]" captionTargetId="figure-69@272.[190,1421,205,1944]" captionTargetPageId="272" captionText="FIGURE 27. Posterior portion of heads, in ventral view. A. Mallochohelea caudellii. B. Probezzia sabroskyi. C. Neobezzia fittkaui. D. Nilobezzia curticornis. E. Johannsenomyia argentata. F. Lanatomyia electra. G. Anebomyia atripes. H. Homohelea delanoe." figureDoi="http://doi.org/10.5281/zenodo.4949164" httpUri="https://zenodo.org/record/4949164/files/figure.png" pageId="88" pageNumber="89">Fig. 27D</figureCitation>
) wide, well-developed but narrow dorsolaterally, extending from palpus to antenna; mesonotum with very short tubercles, not extending posteromedially, not dividing metathorax medially (
<figureCitation id="12E72A5ABD76302EFD0A1BCC4E2DE3F6" box="[754,861,473,498]" captionStart="FIGURE 52" captionStartId="297.[151,250,1942,1964]" captionTargetBox="[190,1402,202,1928]" captionTargetId="figure-58@297.[181,1412,193,1932]" captionTargetPageId="297" captionText="FIGURE 52. Metathoraces and tergites 1, in dorsal view. A. Neobezzia fittkaui. B. Nilobezzia curticornis. C. Johannsenomyia argentata. D. Lanatomyia miles. E. Anebomyia atripes. F. Homohelea delanoe (reconstructed from distorted and dirty specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949234" httpUri="https://zenodo.org/record/4949234/files/figure.png" pageId="88" pageNumber="89">Fig. 52B</figureCitation>
); respiratory organ (
<figureCitation id="12E72A5ABD76302EFBB61BCC4990E3F6" box="[1102,1248,473,498]" captionStart="FIGURE 45" captionStartId="290.[151,250,1738,1760]" captionTargetBox="[222,1373,186,1678]" captionTargetId="figure-193@290.[202,1384,184,1720]" captionTargetPageId="290" captionText="FIGURE 45. Respiratory organs, in dorsal view, pattern on tracheal tubes shown only in some. A. Clinohelea tasmaniensis (from Elson-Harris &amp; Kettle 1986). B. Clinohelea bimaculata (from Wirth &amp; Grogan 1979, modified). C. Heteromyia wokei (from Wirth &amp; Grogan 1977). D. Heteromyia clavata (from Wirth &amp; Grogan 1977). E. Pellucidomyia leei (from Elson-Harris &amp; Kettle 1986). F. Pellucidomyia geari (from de Meillon &amp; Wirth 1981). G. Dibezzia prominens (from Mayer 1934c). H. Hebetula tonnoiri (from Debenham 1974). I. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). J. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). K. Mallochohelea albihalter (from Wirth &amp; Grogan 1979). L. Probezzia pallida (from Wirth &amp; Grogan 1979). M. Probezzia xanthogaster (from Wirth &amp; Grogan 1979). N. Neobezzia fittkaui (from Ronderos et al. 2011). O. Nilobezzia aranea (from Elson-Harris 1987). P. Nilobezzia virago (from Elson-Harris 1987). Q. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). R. Lanatomyia electra (from Debenham 1974). S. Lanatomyia miles (from Debenham 1974). T. Anebomyia atripes (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949212" httpUri="https://zenodo.org/record/4949212/files/figure.png" pageId="88" pageNumber="89">Figs. 45O–P</figureCitation>
) length/width = 2.61–3.56, elongate, moderately slender, somewhat flattened apically, with pores closely abutting at apex of respiratory organ, arranged in single row, outer surface with a few to numbers of wrinkles, with short, wide pedicel, base with short posteromedial apodeme, membranous base of respiratory organ moderately elongate, annulated, tracheal tube straight to slightly curved along length, with spirals restricted to base, wrinkles to half length; wing (
<figureCitation id="12E72A5ABD76302EFF6718994C76E0A1" box="[159,262,652,677]" captionStart="FIGURE 39" captionStartId="284.[151,250,1973,1995]" captionTargetBox="[180,1379,215,1945]" captionTargetId="figure-65@284.[170,1401,210,1949]" captionTargetPageId="284" captionText="FIGURE 39. Legs, wings and apices of the antennae, in ventral view. A. Probezzia sabroskyi. B. Neobezzia fittkaui. C. Nilobezzia sp. (from Australia). D. Johannsenomyia argentata. E. Lanatomyia electra. F. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949198" httpUri="https://zenodo.org/record/4949198/files/figure.png" pageId="88" pageNumber="89">Fig. 39C</figureCitation>
) without apical tubercle or angle, separated medially by fore-, midlegs; halter apex and hind leg (
<figureCitation id="12E72A5ABD76302EFA9718994DB9E0CC" captionStart="FIGURE 33" captionStartId="278.[151,250,1883,1905]" captionTargetBox="[176,1420,177,1860]" captionTargetId="figure-110@278.[176,1420,177,1860]" captionTargetPageId="278" captionText="FIGURE 33. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Parabezzia sp. (5 km. E. Danby, Vermont, USA). B. Clinohelea curriei. C. Pellucidomyia leei. D. Hebetula tonnoiri. E. Jenkinshelea sp. (from Ottawa, ON, Canada). F. Mallochohelea caudellii. G. Probezzia seminigra. H. Nilobezzia sp. (from Kruger NP, South Africa). I. Sphaeromias longipennis. J. Bezzia sp. (6 km E. Salmon Arm, BC, Canada) K. Palpomyia sp. (Black Lake, Quebec, Canada). L. Phaenobezzia sp. (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949184" httpUri="https://zenodo.org/record/4949184/files/figure.png" pageId="88" pageNumber="89">Fig. 33H</figureCitation>
) broadly abutting; halter apex abutting anterolateral knob-like extension of tergite 2 or past tergite 2; legs (
<figureCitation id="12E72A5ABD76302EFA9718BA4DB6E0E8" captionStart="FIGURE 39" captionStartId="284.[151,250,1973,1995]" captionTargetBox="[180,1379,215,1945]" captionTargetId="figure-65@284.[170,1401,210,1949]" captionTargetPageId="284" captionText="FIGURE 39. Legs, wings and apices of the antennae, in ventral view. A. Probezzia sabroskyi. B. Neobezzia fittkaui. C. Nilobezzia sp. (from Australia). D. Johannsenomyia argentata. E. Lanatomyia electra. F. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949198" httpUri="https://zenodo.org/record/4949198/files/figure.png" pageId="88" pageNumber="89">Fig. 39C</figureCitation>
) with lateral margin of foreleg near midlength of wing evenly curved; hind leg visible at lateral margin of wing (
<figureCitation id="12E72A5ABD76302EFF6718E24C79E114" box="[159,265,759,784]" captionStart="FIGURE 33" captionStartId="278.[151,250,1883,1905]" captionTargetBox="[176,1420,177,1860]" captionTargetId="figure-110@278.[176,1420,177,1860]" captionTargetPageId="278" captionText="FIGURE 33. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Parabezzia sp. (5 km. E. Danby, Vermont, USA). B. Clinohelea curriei. C. Pellucidomyia leei. D. Hebetula tonnoiri. E. Jenkinshelea sp. (from Ottawa, ON, Canada). F. Mallochohelea caudellii. G. Probezzia seminigra. H. Nilobezzia sp. (from Kruger NP, South Africa). I. Sphaeromias longipennis. J. Bezzia sp. (6 km E. Salmon Arm, BC, Canada) K. Palpomyia sp. (Black Lake, Quebec, Canada). L. Phaenobezzia sp. (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949184" httpUri="https://zenodo.org/record/4949184/files/figure.png" pageId="88" pageNumber="89">Fig. 33H</figureCitation>
); with apex of foreleg moderately anterior to apex of midleg; apex of hind leg abutting apex of midleg laterally; sensilla: anteromedials—1 short peg, 1 elongate seta (as in
<figureCitation id="12E72A5ABD76302EFC67190E4977E130" box="[927,1031,795,820]" captionStart="FIGURE 31" captionStartId="276.[151,250,1885,1907]" captionTargetBox="[177,1424,178,1864]" captionTargetId="figure-106@276.[177,1424,178,1864]" captionTargetPageId="276" captionText="FIGURE 31. Dorsal setae from left side of thoraces; anteromedial setae. A. Homohelea delanoe. B. Leehelea hispida. C. Sphaeromias longipennis. D. Xenohelea galatea. E. Bezzia nobilis. F. Clastrieromyia dycei. G. Pachyhelea pachymera. H. Palpomyia jonesi. I. Phaenobezzia mashonensis. J. Paryphoconus oliveirai. K. Echinohelea lanei, partial lateral view (distorted slide specimen). L. Palpomyia (JAD685), dorsal view. M. Palpomyia (JAD685), dorsolateral view. N. Probezzia seminigra, dorsal view of left side." figureDoi="http://doi.org/10.5281/zenodo.4949178" httpUri="https://zenodo.org/record/4949178/files/figure.png" pageId="88" pageNumber="89">Fig. 31N</figureCitation>
); anterolaterals—1 moderately long seta; dorsal setae (
<figureCitation id="12E72A5ABD76302EFE9A192B4CB8E153" box="[354,456,830,855]" captionStart="FIGURE 30" captionStartId="275.[151,250,1839,1861]" captionTargetBox="[161,1425,193,1797]" captionTargetId="figure-119@275.[161,1426,193,1813]" captionTargetPageId="275" captionText="FIGURE 30. Dorsal setae from left side of thoraces. A. Serromyia sp. B. Allohelea japonica. C. Monohelea obscura. D. Atyphohelea macroneura. E. Parabezzia huberti. F. Echinohelea lanei. G. Clinohelea curriei. H. Heteromyia clavata. I. Pellucidomyia leei. J. Dibezzia debenhamae (dirty specimen, D-5-T perhaps broken setal socket). K. Hebetula tonnoiri. L. Jenkinshelea magnipennis. M. Macropeza natalensis. N. Mallochohelea caudellii. O. Probezzia sabroskyi. P. Neobezzia fittkaui (from Ronderos et al. 2011, modified). Q. Nilobezzia curticornis. R. Johannsenomyia argentata. S. Lanatomyia miles. T. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949174" httpUri="https://zenodo.org/record/4949174/files/figure.png" pageId="88" pageNumber="89">Fig. 30Q</figureCitation>
)—D-1-T, D-2-T peg-like or setae, D-4-T seta, D-5-T peg-like seta, D-3-T campaniform sensillum, D-3-T lateral to slightly posterolateral to D-4-T; supraalar 2—campaniform sensillum; metathoracics (
<figureCitation id="12E72A5ABD76302EFF6719934C7DE19B" box="[159,269,902,927]" captionStart="FIGURE 52" captionStartId="297.[151,250,1942,1964]" captionTargetBox="[190,1402,202,1928]" captionTargetId="figure-58@297.[181,1412,193,1932]" captionTargetPageId="297" captionText="FIGURE 52. Metathoraces and tergites 1, in dorsal view. A. Neobezzia fittkaui. B. Nilobezzia curticornis. C. Johannsenomyia argentata. D. Lanatomyia miles. E. Anebomyia atripes. F. Homohelea delanoe (reconstructed from distorted and dirty specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949234" httpUri="https://zenodo.org/record/4949234/files/figure.png" pageId="88" pageNumber="89">Fig. 52B</figureCitation>
)—1 campaniform sensillum; M-3-T distant from margin of metathorax (at least 1/3 length of metathorax).
<emphasis id="B8A8EACDBD76302EFED719BF4CD3E1C7" bold="true" box="[303,419,938,963]" pageId="88" pageNumber="89">Abdomen</emphasis>
: pigmentation light to dark brown, with tergite 1 with 3 medial spots, tergites 2–7 with medial area with stripe, 2 anterolateral spots, 2 pairs on tergite
<quantity id="4D249B3ABD76302EFCA819D84E0AE1E2" box="[848,890,973,998]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="88" pageNumber="89" unit="in" value="8.0">8 in</quantity>
some, sternites not pigmented or sternites 3–4 or sternites 3–7 (5–7 membranous and with single medial spot), 2 spots on sternite
<quantity id="4D249B3ABD76302EFBD419E44928E60E" box="[1068,1112,1009,1034]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="88" pageNumber="89" unit="in" value="8.0">8 in</quantity>
some, segment 2 as wide or slightly wider than segment 3, segments with undivided, thin to thick setae, with rounded to pointed, short to moderately elongate tubercles, tergites or sternites entire, not membranous or with sternites 5–7 each with membranous disc; segment 9 (
<figureCitation id="12E72A5ABD76302EFE0B1E494F2CE671" box="[499,604,1116,1141]" captionStart="FIGURE 76" captionStartId="321.[151,250,1765,1787]" captionTargetBox="[225,1365,181,1745]" captionTargetId="figure-185@321.[215,1375,177,1746]" captionTargetPageId="321" captionText="FIGURE 76. Abdominal segments 9. A. Macropeza albitarsis, male, in ventral view (modified from Szadziewski &amp; Dominiak, 2007). B. Macropeza albitarsis, female, in ventral view (from Szadziewski &amp; Dominiak, 2007). C. Mallochohelea termophila, male, in ventral view (from Spinelli 1984). D. Mallochohelea termophila, female, in dorsal view (from Spinelli 1984). E. Probezzia pallida, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). F. Neobezzia fittkaui, male, in dorsal view (from Ronderos, et al. 2011). G. Nilobezzia virago, female, in dorsal view (from Elson-Harris 1987). H. Johannsenomyia argentata, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). I. Johannsenomyia argentata, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979). J. Lanatomyia miles, female, in dorsal view (from Elson- Harris 1987). K. Anebomyia atripes, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). L. Anebomyia atripes, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949324" httpUri="https://zenodo.org/record/4949324/files/figure.png" pageId="88" pageNumber="89">Fig. 76G</figureCitation>
) not strongly modified, terminal processes closely approximated basally, each projecting posterodorsolaterally, tapering to pointed apex; apex of male genital lobe anterior or to level of base of terminal process (as in
<figureCitation id="12E72A5ABD76302EFE091EB14F01E6B9" box="[497,625,1188,1213]" captionStart="FIGURE 78" captionStartId="323.[151,250,1767,1789]" captionTargetBox="[223,1353,186,1749]" captionTargetId="figure-194@323.[213,1373,183,1752]" captionTargetPageId="323" captionText="FIGURE 78. Abdominal segments 9. A. Palpomyia lineata, male, in dorsal and ventral view (from Grogan &amp; Wirth 1979). B. Palpomyia lineata, female, in dorsal and ventral view (from Grogan &amp; Wirth 1979). C. Palpomyia jonesi, male, in dorsal and ventral view (from Grogan &amp; Wirth 1979. D. Palpomyia jonesi, female, in dorsal and ventral view (from Grogan &amp; Wirth 1979). E. Palpomyia belkini, male, in dorsal and ventral view (from Grogan &amp; Wirth 1979). F. Palpomyia belkini, female, in dorsal and ventral view (from Grogan &amp; Wirth 1979). G. Palpomyia rufa, male, in dorsal and ventral view (from Grogan &amp; Wirth 1979). H. Palpomyia flaviceps, female, in dorsal and ventral view (from Grogan &amp; Wirth 1979). I. Phaenobezzia vacunae, male, in ventral view (from Haeselbarth 1965). J. Phaenobezzia mashonensis, male, in ventral view (from Haeselbarth 1965). K. Phaenobezzia opaca, female, in dorsal and ventral view (from Wirth &amp; Grogan 1982). L. Paryphoconus oliveirai, female, in dorsal view (from Ronderos et al. 2007)." figureDoi="http://doi.org/10.5281/zenodo.4949332" httpUri="https://zenodo.org/record/4949332/files/figure.png" pageId="88" pageNumber="89">Figs. 78I–J</figureCitation>
), except for
<taxonomicName id="4DDC4D5CBD76302EFD041EB04EF7E6B9" box="[764,903,1188,1213]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" kingdom="Plantae" order="Caryophyllales" pageId="88" pageNumber="89" phylum="Tracheophyta" rank="species" species="schwarzii">
<emphasis id="B8A8EACDBD76302EFD041EB04EF7E6B9" box="[764,903,1188,1213]" italics="true" pageId="88" pageNumber="89">N. schwarzii</emphasis>
</taxonomicName>
where it extends to about half length of base of terminal process (as in
<figureCitation id="12E72A5ABD76302EFE5A1ED24F78E6E4" box="[418,520,1223,1248]" captionStart="FIGURE 78" captionStartId="323.[151,250,1767,1789]" captionTargetBox="[223,1353,186,1749]" captionTargetId="figure-194@323.[213,1373,183,1752]" captionTargetPageId="323" captionText="FIGURE 78. Abdominal segments 9. A. Palpomyia lineata, male, in dorsal and ventral view (from Grogan &amp; Wirth 1979). B. Palpomyia lineata, female, in dorsal and ventral view (from Grogan &amp; Wirth 1979). C. Palpomyia jonesi, male, in dorsal and ventral view (from Grogan &amp; Wirth 1979. D. Palpomyia jonesi, female, in dorsal and ventral view (from Grogan &amp; Wirth 1979). E. Palpomyia belkini, male, in dorsal and ventral view (from Grogan &amp; Wirth 1979). F. Palpomyia belkini, female, in dorsal and ventral view (from Grogan &amp; Wirth 1979). G. Palpomyia rufa, male, in dorsal and ventral view (from Grogan &amp; Wirth 1979). H. Palpomyia flaviceps, female, in dorsal and ventral view (from Grogan &amp; Wirth 1979). I. Phaenobezzia vacunae, male, in ventral view (from Haeselbarth 1965). J. Phaenobezzia mashonensis, male, in ventral view (from Haeselbarth 1965). K. Phaenobezzia opaca, female, in dorsal and ventral view (from Wirth &amp; Grogan 1982). L. Paryphoconus oliveirai, female, in dorsal view (from Ronderos et al. 2007)." figureDoi="http://doi.org/10.5281/zenodo.4949332" httpUri="https://zenodo.org/record/4949332/files/figure.png" pageId="88" pageNumber="89">Fig. 78E</figureCitation>
); sensilla: tergite 1 (
<figureCitation id="12E72A5ABD76302EFD0A1ED24E28E6E4" box="[754,856,1223,1248]" captionStart="FIGURE 52" captionStartId="297.[151,250,1942,1964]" captionTargetBox="[190,1402,202,1928]" captionTargetId="figure-58@297.[181,1412,193,1932]" captionTargetPageId="297" captionText="FIGURE 52. Metathoraces and tergites 1, in dorsal view. A. Neobezzia fittkaui. B. Nilobezzia curticornis. C. Johannsenomyia argentata. D. Lanatomyia miles. E. Anebomyia atripes. F. Homohelea delanoe (reconstructed from distorted and dirty specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949234" httpUri="https://zenodo.org/record/4949234/files/figure.png" pageId="88" pageNumber="89">Fig. 52B</figureCitation>
) with 8 setae, 2 campaniform sensilla, including 4 lateral sensilla, D-2-I, D-3-I closely approximated, D-7-I situated anterolaterally near L-1-I; segment 4 (
<figureCitation id="12E72A5ABD76302EFA971EFE4DB7E72C" captionStart="FIGURE 67" captionStartId="312.[151,250,1975,1997]" captionTargetBox="[198,1410,217,1946]" captionTargetId="figure-53@312.[189,1420,211,1950]" captionTargetPageId="312" captionText="FIGURE 67. Abdominal segments four, in dorsal and ventral view (shagreen not shown). A. Nilobezzia curticornis. B. Johannsenomyia argentata. C. Lanatomyia miles." figureDoi="http://doi.org/10.5281/zenodo.4949288" httpUri="https://zenodo.org/record/4949288/files/figure.png" pageId="88" pageNumber="89">Fig. 67A</figureCitation>
)—D-2-IV, D-3-IV short to moderately elongate setae on short tubercles; D-5-IV, D-8-IV, D-9-IV short to moderately elongate setae; D-5-IV on single tubercle, D-8-IV, D-9-IV on separate but closely approximated tubercles, posterior dorsal sensilla in transverse row, arranged medially to laterally: D-5-IV, D-4-IV, D-8-IV, D-9- IV; D-7-IV, if present, near D-3-IV; L-1-IV short seta on rounded tubercle, near L-3-IV; L-2-IV, L-3-IV, L-4-IV short setae on pointed tubercles, V-5-IV, V-6-IV, V-7-IV short setae on short tubercles, all closely approximated; segment 8 without D-3-VIII, without L-1-VIII; segment 9 (
<figureCitation id="12E72A5ABD76302EFCD31FD74EFEE7DF" box="[811,910,1474,1499]" captionStart="FIGURE 76" captionStartId="321.[151,250,1765,1787]" captionTargetBox="[225,1365,181,1745]" captionTargetId="figure-185@321.[215,1375,177,1746]" captionTargetPageId="321" captionText="FIGURE 76. Abdominal segments 9. A. Macropeza albitarsis, male, in ventral view (modified from Szadziewski &amp; Dominiak, 2007). B. Macropeza albitarsis, female, in ventral view (from Szadziewski &amp; Dominiak, 2007). C. Mallochohelea termophila, male, in ventral view (from Spinelli 1984). D. Mallochohelea termophila, female, in dorsal view (from Spinelli 1984). E. Probezzia pallida, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). F. Neobezzia fittkaui, male, in dorsal view (from Ronderos, et al. 2011). G. Nilobezzia virago, female, in dorsal view (from Elson-Harris 1987). H. Johannsenomyia argentata, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). I. Johannsenomyia argentata, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979). J. Lanatomyia miles, female, in dorsal view (from Elson- Harris 1987). K. Anebomyia atripes, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). L. Anebomyia atripes, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949324" httpUri="https://zenodo.org/record/4949324/files/figure.png" pageId="88" pageNumber="89">Fig. 76G</figureCitation>
)—with D-5-IX, D-6-IX campaniform sensilla.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD76302EFF3F1FF04E3BE442" pageId="88" pageNumber="89" type="distribution">
<paragraph id="8A6336DFBD76302EFF3F1FF04E3BE442" blockId="88.[151,1437,151,2035]" pageId="88" pageNumber="89">
<emphasis id="B8A8EACDBD76302EFF3F1FF04F15E7FA" bold="true" box="[199,613,1509,1534]" pageId="88" pageNumber="89">DISTRIBUTION AND HABITAT</emphasis>
: The genus
<taxonomicName id="4DDC4D5CBD76302EFD061FF04E04E7FA" box="[766,884,1509,1534]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD76302EFD061FF04E04E7FA" box="[766,884,1509,1534]" italics="true" pageId="88" pageNumber="89">Nilobezzia</emphasis>
</taxonomicName>
is known from 74 species from every Region worldwide (
<taxonomicName id="4DDC4D5CBD76302EFEE61C1C4CCDE426" authority="Borkent, 2014" authorityName="Borkent" authorityYear="2014" box="[286,445,1545,1570]" genus="Borkent" pageId="88" pageNumber="92" rank="genus" status="gen. nov.">
<bibRefCitation id="EE4D4B2EBD76302EFEE61C1C4CCDE426" author="Borkent" box="[286,445,1545,1570]" firstAuthor="Borkent" pageId="88" pageNumber="89" refId="ref177229" refString="Borkent, A. (2014) World Species of Biting Midges (Diptera: Ceratopogonidae). Available from: http: // www. inhs. illinois. edu / research / FLYTREE / Borkent. html (accessed 20 May 2014)" type="url" year="2014">Borkent 2014</bibRefCitation>
</taxonomicName>
; additional species below). Immatures are present in lakes, grassy wetlands, ponds, salt marshes (
<bibRefCitation id="EE4D4B2EBD76302EFEF91C384C9EE442" author="Knausenberger" box="[257,494,1581,1606]" firstAuthor="Knausenberger" pageId="88" pageNumber="89" refId="ref188128" refString="Knausenberger, W. I. (1987) Contributions to the autecology and ecosystematics of immature Ceratopogonidae (Diptera), with emphasis on the tribes Heteromyiini and Sphaeromiini in the Middle Atlantic United States. Ph. D. Dissertation, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, xxix + 729 pp." type="book" year="1987">Knausenberger 1987</bibRefCitation>
), reservoirs, creeks, and rivers.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD76302FFF3F1C444EA0E2F3" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89" type="discussion">
<paragraph id="8A6336DFBD76302EFF3F1C444F83E4FC" blockId="88.[151,1437,151,2035]" pageId="88" pageNumber="89">
<emphasis id="B8A8EACDBD76302EFF3F1C444F5DE46E" bold="true" box="[199,557,1617,1642]" pageId="88" pageNumber="89">TAXONOMIC DISCUSSION</emphasis>
: There are 17 species of
<taxonomicName id="4DDC4D5CBD76302EFCAA1C444EB8E46E" box="[850,968,1617,1642]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD76302EFCAA1C444EB8E46E" box="[850,968,1617,1642]" italics="true" pageId="88" pageNumber="89">Nilobezzia</emphasis>
</taxonomicName>
known as pupae (
<tableCitation id="C75E0364BD76302EFB621C44486BE46E" box="[1178,1307,1617,1642]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStartId-0="174.[147,245,486,512]" captionStartId-1="242.[151,239,152,175]" captionTargetBox-0="[150,1771,751,1679]" captionTargetBox-1="[159,1428,341,1978]" captionTargetPageId-0="174" captionTargetPageId-1="242" captionText-0="TABLE 2. References to species of Ceratopogonidae described as eggs, larvae and/or pupae. Species studied as pupae in the current study have their name highlighted in bold font. Stages studied are referred to as 'E' for egg, 'L' for larva and 'P' for pupa. Taxa are arrangement phyletically by subfamily and tribe (some of which are paraphyletic) and within tribes genera are listed alphabetically. Subfamilies are abbreviated as follows: Lept.— Leptoconopinae, Forc.—Forcipomyiinae, Dasy.—Dasyheleinae, Cera.—Ceratopogoninae. References highlighted in bold font are those with SEM photomicrographs. Theses are cited only when their results have not been published. Duplicated figures or repeated descriptions by subsequent authors are not listed here. Fossil taxa are identified with a cross (†). The species name in bold indicates only that named material of the pupa was studied, not that the cited author's specimens were reexamined (although in many instances they were). The larva of Jenkinshelea albaria was tentatively associated by Wirth &amp; Grogan 1979 (not reared); see discussion by Knausenberger, 1987:248." captionText-1="TABLE 3. Number of named extant species with described immatures for genera of Ceratopogonidae (taxa of uncertain identity or identified only as &quot;sp.&quot; are not included). Only genera with at least one immature stage known are listed. Percentages of described species within that genus are those for which those stages are known. Number of named species from Borkent (2014), with a few additions as noted in the Taxonomic Discussion under some genera. Taxa are arranged phyletically by subfamily and tribe and within tribes genera are listed alphabetically." httpUri-0="http://table.plazi.org/id/DEA36657BD8030D8FF6B1BF34923E0EC" httpUri-1="http://table.plazi.org/id/DEA36657BDDC3084FF6F1A8D496DE336" pageId="88" pageNumber="89" tableUuid-0="DEA36657BD8030D8FF6B1BF34923E0EC" tableUuid-1="DEA36657BDDC3084FF6F1A8D496DE336">Tables 2–3</tableCitation>
). Only two species,
<taxonomicName id="4DDC4D5CBD76302EFF0D1C604CF0E489" box="[245,384,1652,1677]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" kingdom="Plantae" order="Caryophyllales" pageId="88" pageNumber="89" phylum="Tracheophyta" rank="species" species="schwarzii">
<emphasis id="B8A8EACDBD76302EFF0D1C604CF0E489" box="[245,384,1652,1677]" italics="true" pageId="88" pageNumber="89">N. schwarzii</emphasis>
</taxonomicName>
and
<taxonomicName id="4DDC4D5CBD76302EFE4E1C604F48E489" authorityName="Wirth" authorityYear="1962" box="[438,568,1652,1677]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="mallochi">
<emphasis id="B8A8EACDBD76302EFE4E1C604F48E489" box="[438,568,1652,1677]" italics="true" pageId="88" pageNumber="89">N. mallochi</emphasis>
</taxonomicName>
, have abdominal membranous discs (see character 50). Nearly all male pupae have short genital lobes, not extending posteriorly past the base of the terminal processes, reflecting the short, modified male adult genitalia; only those of
<taxonomicName id="4DDC4D5CBD76302EFD761CA84E6BE4D1" box="[654,795,1724,1749]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" kingdom="Plantae" order="Caryophyllales" pageId="88" pageNumber="89" phylum="Tracheophyta" rank="species" species="schwarzii">
<emphasis id="B8A8EACDBD76302EFD761CA84E6BE4D1" box="[654,795,1724,1749]" italics="true" pageId="88" pageNumber="89">N. schwarzii</emphasis>
</taxonomicName>
have more elongate genital lobes (even though the male adult genitalia doesn't seem disproportionally longer).
</paragraph>
<paragraph id="8A6336DFBD76302EFF3F1D164969E544" blockId="88.[151,1437,151,2035]" pageId="88" pageNumber="89">
<taxonomicName id="4DDC4D5CBD76302EFF3F1D164FF9E518" authority="(de Meillon &amp; Wirth)" baseAuthorityName="de Meillon &amp; Wirth" baseAuthorityYear="1981" box="[199,649,1795,1820]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="theileri">
<emphasis id="B8A8EACDBD76302EFF3F1D164CE5E518" box="[199,405,1795,1820]" italics="true" pageId="88" pageNumber="89">Nilobezzia theileri</emphasis>
(de Meillon &amp; Wirth)
</taxonomicName>
was originally described as a
<taxonomicName id="4DDC4D5CBD76302EFC2F1D1648ECE518" authority="(de Meillon &amp; Wirth 1981)" baseAuthorityName="de Meillon &amp; Wirth" baseAuthorityYear="1981" box="[983,1436,1795,1820]" class="Insecta" family="Ceratopogonidae" genus="Sphaeromias" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD76302EFC2F1D164918E518" box="[983,1128,1795,1820]" italics="true" pageId="88" pageNumber="89">Sphaeromias</emphasis>
(
<bibRefCitation id="EE4D4B2EBD76302EFB8F1D1648E4E518" author="de Meillon &amp; Wirth" box="[1143,1428,1795,1820]" firstAuthor="de Meillon" pageId="88" pageNumber="89" pagination="525 - 561" refId="ref181048" refString="De Meillon, B. &amp; Wirth, W. W. (1981) Subsaharan Ceratopogonidae (Diptera) VI. New species and records of South African biting midges collected by A. L. Dyce. Annals of the Natal Museum, 24, 525 - 561." type="journal article" year="1981">de Meillon &amp; Wirth 1981</bibRefCitation>
)
</taxonomicName>
but is here considered a member of
<taxonomicName id="4DDC4D5CBD76302EFDDF1D324FEDE544" box="[551,669,1831,1856]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD76302EFDDF1D324FEDE544" box="[551,669,1831,1856]" italics="true" pageId="88" pageNumber="89">Nilobezzia</emphasis>
</taxonomicName>
as a new combination as follows:
</paragraph>
<paragraph id="8A6336DFBD76302EFF6F1D5E4911E560" blockId="88.[151,1437,151,2035]" box="[151,1121,1867,1892]" pageId="88" pageNumber="89">
<taxonomicName id="4DDC4D5CBD76302EFF6F1D5E4FA5E560" authority="(de Meillon &amp; Wirth), 1981:552" baseAuthorityName="de Meillon &amp; Wirth" baseAuthorityPageNumber="552" baseAuthorityYear="1981" box="[151,725,1867,1892]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="theileri">
<emphasis id="B8A8EACDBD76302EFF6F1D5E4C15E560" box="[151,357,1867,1892]" italics="true" pageId="88" pageNumber="89">Nilobezzia theileri</emphasis>
(de Meillon &amp; Wirth), 1981:552
</taxonomicName>
(
<taxonomicName id="4DDC4D5CBD76302EFD1C1D5E4E05E560" baseAuthorityName="de Meillon &amp; Wirth" baseAuthorityYear="1981" box="[740,885,1867,1892]" class="Insecta" family="Ceratopogonidae" genus="Sphaeromias" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD76302EFD1C1D5E4E05E560" box="[740,885,1867,1892]" italics="true" pageId="88" pageNumber="89">Sphaeromias</emphasis>
</taxonomicName>
).
<emphasis id="B8A8EACDBD76302EFC751D5E4911E560" bold="true" box="[909,1121,1867,1892]" pageId="88" pageNumber="89">new combination.</emphasis>
</paragraph>
<paragraph id="8A6336DFBD76302EFF3F1D7B4C3DE5F7" blockId="88.[151,1437,151,2035]" pageId="88" pageNumber="89">
The pupa has the diagnostic features of this genus and reexamination of the adult
<typeStatus id="5567887DBD76302EFBAC1D7B49C6E583" box="[1108,1206,1902,1927]" pageId="88" pageNumber="89" type="holotype">holotype</typeStatus>
shows that the basal tooth of the claws of the adult female are actually outer and not inner, as described and inaccurately illustrated by
<bibRefCitation id="EE4D4B2EBD76302EFF6F1DA34CB7E5CB" author="de Meillon &amp; Wirth" box="[151,455,1974,1999]" firstAuthor="de Meillon" pageId="88" pageNumber="89" pagination="525 - 561" refId="ref181048" refString="De Meillon, B. &amp; Wirth, W. W. (1981) Subsaharan Ceratopogonidae (Diptera) VI. New species and records of South African biting midges collected by A. L. Dyce. Annals of the Natal Museum, 24, 525 - 561." type="journal article" year="1981">de Meillon &amp; Wirth (1981)</bibRefCitation>
. The adult actually keys to
<taxonomicName id="4DDC4D5CBD76302EFD0E1DA34E1CE5CB" box="[758,876,1974,1999]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD76302EFD0E1DA34E1CE5CB" box="[758,876,1974,1999]" italics="true" pageId="88" pageNumber="89">Nilobezzia</emphasis>
</taxonomicName>
in the key to the Afrotropical fauna by
<bibRefCitation id="EE4D4B2EBD76302EFADB1DA34C39E5F7" author="de Meillon &amp; Wirth" firstAuthor="de Meillon" pageId="88" pageNumber="89" pagination="27 - 147" refId="ref181099" refString="De Meillon, B. &amp; Wirth, W. W. (1991) The genera and subgenera (excluding Culicoides) of the Afrotropical Biting Midges (Diptera: Ceratopogonidae). Annals of the Natal Museum, 32, 27 - 147." type="journal article" year="1991">de Meillon &amp; Wirth (1991)</bibRefCitation>
.
</paragraph>
<paragraph id="8A6336DFBD77302FFF3F1A824EA0E2F3" blockId="89.[151,1437,151,1149]" pageId="89" pageNumber="90">
<bibRefCitation id="EE4D4B2EBD77302FFF3F1A824CD7E2B4" author="Elson-Harris" box="[199,423,151,176]" firstAuthor="Elson-Harris" pageId="89" pageNumber="90" pagination="45 - 61" refId="ref182239" refString="Elson-Harris, M. M. (1987) The immature stages of some Australian Sphaeromiini (Diptera: Ceratopogonidae). Journal of the Australian Entomological Society, 26, 45 - 61. http: // dx. doi. org / 10.1111 / j. 1440 - 6055.1987. tb 00258. x" type="journal article" year="1987">Elson-Harris (1987)</bibRefCitation>
keyed the pupae of six Australian species.
<bibRefCitation id="EE4D4B2EBD77302FFC7E1A824916E2B4" author="Elson-Harris" box="[902,1126,151,176]" firstAuthor="Elson-Harris" pageId="89" pageNumber="90" pagination="267 - 275" refId="ref182295" refString="Elson-Harris, M. M. (1990) Keys to the immature stages of some Australian Ceratopogonidae (Diptera). Journal of the Australian Entomological Society, 29, 267 - 275. http: // dx. doi. org / 10.1111 / j. 1440 - 6055.1990. tb 00361. x" type="journal article" year="1990">Elson-Harris (1990)</bibRefCitation>
used the position of L-3-IV ventral to L-1-IV (her lpm ii &quot;in line with&quot; lasm) to distinguish Australian
<taxonomicName id="4DDC4D5CBD77302FFC2A1AA94938E2D1" box="[978,1096,188,213]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD77302FFC2A1AA94938E2D1" box="[978,1096,188,213]" italics="true" pageId="89" pageNumber="90">Nilobezzia</emphasis>
</taxonomicName>
from Australian
<taxonomicName id="4DDC4D5CBD77302FFAFE1AA9481DE2D1" box="[1286,1389,188,213]" class="Insecta" family="Ceratopogonidae" genus="Hebetula" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD77302FFAFE1AA9481DE2D1" box="[1286,1389,188,213]" italics="true" pageId="89" pageNumber="90">Hebetula</emphasis>
</taxonomicName>
and
<taxonomicName id="4DDC4D5CBD77302FFF6F1AF54C6DE2F3" box="[151,285,224,247]" class="Insecta" family="Ceratopogonidae" genus="Lanatomyia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD77302FFF6F1AF54C6DE2F3" box="[151,285,224,247]" italics="true" pageId="89" pageNumber="90">Lanatomyia</emphasis>
</taxonomicName>
and this feature does indeed work for known Australian taxa.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD77302FFF3F1B114F11E679" pageId="89" pageNumber="90" type="materials_examined">
<paragraph id="8A6336DFBD77302FFF3F1B114F11E679" blockId="89.[151,1437,151,1149]" pageId="89" pageNumber="90">
<emphasis id="B8A8EACDBD77302FFF3F1B114F70E319" bold="true" box="[199,512,260,285]" pageId="89" pageNumber="90">MATERIAL EXAMINED</emphasis>
:
<taxonomicName id="4DDC4D5CBD77302FFDF71B104F0EE318" authorityName="Debenham" authorityYear="1974" box="[527,638,261,284]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="aranea">
<emphasis id="B8A8EACDBD77302FFDF71B104F0EE318" box="[527,638,261,284]" italics="true" pageId="89" pageNumber="90">N. aranea</emphasis>
</taxonomicName>
: 1 pupal exuviae (of
<typeStatus id="5567887DBD77302FFC921B114EA0E319" box="[874,976,260,285]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
), 1 pupal exuviae (of
<typeStatus id="5567887DBD77302FFB381B11486FE319" box="[1216,1311,260,285]" pageId="89" pageNumber="90" type="allotype">allotype</typeStatus>
), Breakfast Creek, Spencer,
<collectingRegion id="4818F83DBD77302FFEA31B3D4F42E344" box="[347,562,295,320]" country="Australia" name="New South Wales" pageId="89" pageNumber="90">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFDBB1B324FDDE344" box="[579,685,295,320]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFD461B324E59E344" box="[702,809,295,320]" pageId="89" pageNumber="90" value="1965-02-04">4-II-1965</date>
(ANIC); 1 pupal exuviae (of
<typeStatus id="5567887DBD77302FFB611B32498EE344" box="[1177,1278,295,320]" pageId="89" pageNumber="90" type="paratype">paratype</typeStatus>
), as previous locality,
<date id="FE62101FBD77302FFF001B594C1DE361" box="[248,365,332,357]" pageId="89" pageNumber="90" value="1969-03-03">3-III-1969</date>
(ANIC); 1 pupal exuviae, McCarrs Creek,
<collectingRegion id="4818F83DBD77302FFCA61B59495CE361" box="[862,1068,332,357]" country="Australia" name="New South Wales" pageId="89" pageNumber="90">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFBC01B5949D1E361" box="[1080,1185,332,357]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFB561B59486EE361" box="[1198,1310,332,357]" pageId="89" pageNumber="90" value="1969-01-22">22-I-1969</date>
(ANIC); 1 pupal exuviae, Oystershell Road,
<collectingRegion id="4818F83DBD77302FFDD11B654E70E38C" box="[553,768,367,392]" country="Australia" name="New South Wales" pageId="89" pageNumber="90">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFCE91B7A4E0BE38C" box="[785,891,367,392]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFC731B7A4E87E38C" box="[907,1015,367,392]" pageId="89" pageNumber="90" value="1965-02-04">4-II-1965</date>
(ANIC).
<taxonomicName id="4DDC4D5CBD77302FFB971B64485DE38C" authorityName="Debenham" authorityYear="1974" box="[1135,1325,367,392]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="basispinigera">
<emphasis id="B8A8EACDBD77302FFB971B64485DE38C" box="[1135,1325,367,392]" italics="true" pageId="89" pageNumber="90">N. basispinigera</emphasis>
</taxonomicName>
: 1 pupal exuviae, Warregor, Engonia,
<collectingRegion id="4818F83DBD77302FFE1A1B814FC0E3A9" box="[482,688,404,429]" country="Australia" name="New South Wales" pageId="89" pageNumber="90">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFD451B814E56E3A9" box="[701,806,404,429]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFCCB1B814EB0E3A9" box="[819,960,404,429]" pageId="89" pageNumber="90" value="1963-12-11">11-XII-1963</date>
(ANIC).
<taxonomicName id="4DDC4D5CBD77302FFBD71B8049BFE3A9" authorityName="Waugh &amp; Wirth" authorityYear="1976" box="[1071,1231,404,429]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="B8A8EACDBD77302FFBD71B8049BFE3A9" box="[1071,1231,404,429]" italics="true" pageId="89" pageNumber="90">N. brevicornis</emphasis>
</taxonomicName>
: 1 pupal exuviae, Ft. Summer #1, DeBaca County,
<collectingRegion id="4818F83DBD77302FFDDE1BAD4FCFE3D4" box="[550,703,439,464]" country="United States of America" name="New Mexico" pageId="89" pageNumber="90">New Mexico</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFD281BA24E7BE3D4" box="[720,779,439,464]" name="United States of America" pageId="89" pageNumber="90">USA</collectingCountry>
,
<date id="FE62101FBD77302FFCE31BA24EC1E3D4" box="[795,945,439,464]" pageId="89" pageNumber="90" value="1967-08-21">21-VIII-1967</date>
(USNM); 2 pupal exuviae, Bitter Lakes, Chaues County,
<collectingRegion id="4818F83DBD77302FFEB71BC94C93E3F1" box="[335,483,476,501]" country="United States of America" name="New Mexico" pageId="89" pageNumber="90">New Mexico</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFE161BC94F59E3F1" box="[494,553,476,501]" name="United States of America" pageId="89" pageNumber="90">USA</collectingCountry>
,
<date id="FE62101FBD77302FFDCC1BC94FBBE3F1" box="[564,715,476,501]" pageId="89" pageNumber="90" value="1967-08-17">17-VIII-1967</date>
(USNM).
<taxonomicName id="4DDC4D5CBD77302FFCBB1BC84EAFE3F0" baseAuthorityName="Kieffer" baseAuthorityYear="1917" box="[835,991,477,500]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="curticornis">
<emphasis id="B8A8EACDBD77302FFCBB1BC84EAFE3F0" box="[835,991,477,500]" italics="true" pageId="89" pageNumber="90">N. curticornis</emphasis>
</taxonomicName>
(as
<taxonomicName id="4DDC4D5CBD77302FFBF51BC849A3E3F0" authorityName="Debenham" authorityYear="1974" box="[1037,1235,477,500]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="triquetrinotata">
<emphasis id="B8A8EACDBD77302FFBF51BC849A3E3F0" box="[1037,1235,477,500]" italics="true" pageId="89" pageNumber="90">N. triquetrinotata</emphasis>
</taxonomicName>
): 1 pupal exuviae (of
<typeStatus id="5567887DBD77302FFF461BEA4C50E01C" box="[190,288,511,536]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4DDC4D5CBD77302FFEBE18144F7CE013" authorityName="Debenham" authorityYear="1974" box="[326,524,512,536]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="triquetrinotata">
<emphasis id="B8A8EACDBD77302FFEBE18144F7CE013" box="[326,524,512,536]" italics="true" pageId="89" pageNumber="90">N. triquetrinotata</emphasis>
</taxonomicName>
), Griffith,
<collectingRegion id="4818F83DBD77302FFD7E18154E22E01C" box="[646,850,511,536]" country="Australia" name="New South Wales" pageId="89" pageNumber="90">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFCA51BEA4EB7E01C" box="[861,967,511,536]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFC291BEA4926E01C" box="[977,1110,511,536]" pageId="89" pageNumber="90" value="1956-11-15">15-XI-1956</date>
(ANIC); 1 pupal exuviae (of
<typeStatus id="5567887DBD77302FFF6F18314D81E039" box="[151,241,548,573]" pageId="89" pageNumber="90" type="allotype">allotype</typeStatus>
of
<taxonomicName id="4DDC4D5CBD77302FFEE218304C91E038" authorityName="Debenham" authorityYear="1974" box="[282,481,549,572]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="triquetrinotata">
<emphasis id="B8A8EACDBD77302FFEE218304C91E038" box="[282,481,549,572]" italics="true" pageId="89" pageNumber="90">N. triquetrinotata</emphasis>
</taxonomicName>
), 6 pupal exuviae (of
<typeStatus id="5567887DBD77302FFD1918314E3DE038" box="[737,845,548,572]" pageId="89" pageNumber="90" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4DDC4D5CBD77302FFC8D1830494DE038" authorityName="Debenham" authorityYear="1974" box="[885,1085,549,572]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="triquetrinotata">
<emphasis id="B8A8EACDBD77302FFC8D1830494DE038" box="[885,1085,549,572]" italics="true" pageId="89" pageNumber="90">N. triquetrinotata</emphasis>
</taxonomicName>
), previous locality (ANIC).
<taxonomicName id="4DDC4D5CBD77302FFA7A18304D83E064" baseAuthorityName="Loew" baseAuthorityYear="1869" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="formosa">
<emphasis id="B8A8EACDBD77302FFA7A18304D83E064" italics="true" pageId="89" pageNumber="90">N. formosa</emphasis>
</taxonomicName>
: 1 pupal exuviae, Kozennyi Torets river, Raigorodok,
<collectingRegion id="4818F83DBD77302FFCAE18524950E064" box="[854,1056,583,608]" country="Ukraine" name="Donets'k" pageId="89" pageNumber="90">Donetsk Province</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFBD4185249F9E064" box="[1068,1161,583,608]" name="Ukraine" pageId="89" pageNumber="90">Ukraine</collectingCountry>
,
<date id="FE62101FBD77302FFB6C18524847E064" box="[1172,1335,583,608]" pageId="89" pageNumber="90" value="1970-05-19" valueMax="1970-05-30" valueMin="1970-05-19">19–30-V-1970</date>
(ZIN); 2 pupal, as previous locality,
<date id="FE62101FBD77302FFE3018794F4BE081" box="[456,571,620,645]" pageId="89" pageNumber="90" value="1970-04-01">1-IV-1970</date>
(ZIN).
<taxonomicName id="4DDC4D5CBD77302FFD6918784E5BE081" box="[657,811,620,645]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="fuscitarsus">
<emphasis id="B8A8EACDBD77302FFD6918784E5BE081" box="[657,811,620,645]" italics="true" pageId="89" pageNumber="90">N. fuscitarsus</emphasis>
</taxonomicName>
: 1 pupal exuviae, Darban,
<collectingRegion id="4818F83DBD77302FFBA018794853E081" box="[1112,1315,620,645]" country="Australia" name="Western Australia" pageId="89" pageNumber="90">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFAD7187948E8E081" box="[1327,1432,620,645]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFF6F189A4C61E0AC" box="[151,273,655,680]" pageId="89" pageNumber="90" value="1985-10-28">28-X-1985</date>
(ANIC); 2 pupal exuviae, Humpty Doo,
<collectingRegion id="4818F83DBD77302FFCE3189A4E84E0AC" box="[795,1012,655,680]" country="Australia" name="Northern Territory" pageId="89" pageNumber="90">Northern Territory</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFBF0189A4902E0AC" box="[1032,1138,655,680]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFB7E189A4875E0AC" box="[1158,1285,655,680]" pageId="89" pageNumber="90" value="1958-12-05">5-XII-1958</date>
(ANIC).
<taxonomicName id="4DDC4D5CBD77302FFA7A18844C7FE0C8" baseAuthorityName="Macfie" baseAuthorityYear="1940" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="neotropica">
<emphasis id="B8A8EACDBD77302FFA7A18844C7FE0C8" italics="true" pageId="89" pageNumber="90">N. neotropica</emphasis>
</taxonomicName>
: 1 pupal exuviae, Lago Monte Aleque,
<collectingRegion id="4818F83DBD77302FFD3418A14E0EE0C9" box="[716,894,692,717]" country="Brazil" name="Sao Paulo" pageId="89" pageNumber="90">Sao Paulo State</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFC7118A14EA0E0C9" box="[905,976,692,717]" name="Brazil" pageId="89" pageNumber="90">Brazil</collectingCountry>
,
<date id="FE62101FBD77302FFC2318A1492BE0C9" box="[987,1115,692,717]" pageId="89" pageNumber="90" value="2000-07-05">5-VII-2000</date>
(CNCI).
<taxonomicName id="4DDC4D5CBD77302FFB3D18A0484AE0C9" baseAuthorityName="Meillon" baseAuthorityYear="1937" box="[1221,1338,692,717]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B8A8EACDBD77302FFB3D18A0484AE0C9" box="[1221,1338,692,717]" italics="true" pageId="89" pageNumber="90">N. robusta</emphasis>
</taxonomicName>
: 1 pupal exuviae; Douglas,
<collectingRegion id="4818F83DBD77302FFE9518C24F66E0F4" box="[365,534,727,752]" country="South Africa" name="Western Cape" pageId="89" pageNumber="90">Cape Province</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFDDC18C24FCAE0F4" box="[548,698,727,752]" name="South Africa" pageId="89" pageNumber="90">South Africa</collectingCountry>
,
<date id="FE62101FBD77302FFD3F18C24E47E0F4" box="[711,823,727,752]" pageId="89" pageNumber="90" value="1974-01-23">23-I-1974</date>
(NMSA); 1 pupal exuviae (of
<typeStatus id="5567887DBD77302FFB6718C24874E0F4" box="[1183,1284,727,752]" pageId="89" pageNumber="90" type="paratype">paratype</typeStatus>
), Berg River, Piquetberg,
<collectingCountry id="F2CB764FBD77302FFEE718E94CC2E111" box="[287,434,764,789]" name="South Africa" pageId="89" pageNumber="90">South Africa</collectingCountry>
,
<date id="FE62101FBD77302FFE4718E94F32E111" box="[447,578,764,789]" pageId="89" pageNumber="90" value="1953-03-16">16-III-1953</date>
(SAIM).
<taxonomicName id="4DDC4D5CBD77302FFD4A18E84E4FE111" box="[690,831,764,789]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" kingdom="Plantae" order="Caryophyllales" pageId="89" pageNumber="90" phylum="Tracheophyta" rank="species" species="schwarzii">
<emphasis id="B8A8EACDBD77302FFD4A18E84E4FE111" box="[690,831,764,789]" italics="true" pageId="89" pageNumber="90">N. schwarzii</emphasis>
</taxonomicName>
: 3 pupal exuviae, Lake Providence,
<collectingRegion id="4818F83DBD77302FFB2718E94820E111" box="[1247,1360,764,789]" country="United States of America" name="Louisiana" pageId="89" pageNumber="90">Louisiana</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFAA518E948E8E111" box="[1373,1432,764,789]" name="United States of America" pageId="89" pageNumber="90">USA</collectingCountry>
,
<date id="FE62101FBD77302FFF6F190A4D8FE13C" box="[151,255,799,824]" pageId="89" pageNumber="90" value="1953-07">VII-1953</date>
(USNM); 3 pupal exuviae, Sanibel Island, Lee County, Florida,
<collectingCountry id="F2CB764FBD77302FFBF9190A494CE13C" box="[1025,1084,799,824]" name="United States of America" pageId="89" pageNumber="90">USA</collectingCountry>
,
<date id="FE62101FBD77302FFBB3190A49B2E13C" box="[1099,1218,799,824]" pageId="89" pageNumber="90" value="1973-05-11">11-V-1973</date>
(USNM); 1 pupal exuviae, Lake Munson, Leon County, Florida,
<collectingCountry id="F2CB764FBD77302FFD5519514F98E159" box="[685,744,836,861]" name="United States of America" pageId="89" pageNumber="90">USA</collectingCountry>
,
<date id="FE62101FBD77302FFD0C19514EFBE159" box="[756,907,836,861]" pageId="89" pageNumber="90" value="1987-08-17">17-VIII-1987</date>
(
<collectionCode id="ECCDAE1ABD77302FFC6419514E95E159" box="[924,997,836,861]" pageId="89" pageNumber="90">JHEC</collectionCode>
); 1 pupal exuviae,
<quantity id="4D249B3ABD77302FFB4619514989E159" box="[1214,1273,836,861]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="89" pageNumber="90" unit="km" value="3.0">3 km</quantity>
N of Caldera,
<collectingCountry id="F2CB764FBD77302FFF6F19724C63E184" box="[151,275,871,896]" name="Costa Rica" pageId="89" pageNumber="90">Costa Rica</collectingCountry>
,
<date id="FE62101FBD77302FFEE619724CDCE184" box="[286,428,871,896]" pageId="89" pageNumber="90" value="1993-12-17">17-XII-1993</date>
(CNCI).
<taxonomicName id="4DDC4D5CBD77302FFDED197C4FF6E184" baseAuthorityName="de Meillon &amp; Wirth" baseAuthorityYear="1981" box="[533,646,871,896]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="theileri">
<emphasis id="B8A8EACDBD77302FFDED197C4FF6E184" box="[533,646,871,896]" italics="true" pageId="89" pageNumber="90">N. theileri</emphasis>
</taxonomicName>
: 1 pupal exuviae (of
<typeStatus id="5567887DBD77302FFC8B19724EA8E184" box="[883,984,871,896]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
; with adult female), Loutpan, Transvaal,
<collectingCountry id="F2CB764FBD77302FFF6F19994C5CE1A1" box="[151,300,908,933]" name="South Africa" pageId="89" pageNumber="90">South Africa</collectingCountry>
,
<date id="FE62101FBD77302FFEC119994CDAE1A1" box="[313,426,908,933]" pageId="89" pageNumber="90" value="1974-01-10">10-I-1974</date>
(NMSA).
<taxonomicName id="4DDC4D5CBD77302FFDDE19984FE2E1A0" authorityName="Debenham" authorityYear="1974" box="[550,658,909,932]" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="virago">
<emphasis id="B8A8EACDBD77302FFDDE19984FE2E1A0" box="[550,658,909,932]" italics="true" pageId="89" pageNumber="90">N. virago</emphasis>
</taxonomicName>
: 2 pupal exuviae, Darwin,
<collectingRegion id="4818F83DBD77302FFC37199949D1E1A1" box="[975,1185,908,933]" country="Australia" name="Northern Territory" pageId="89" pageNumber="90">Northern Territory</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFB5619994868E1A1" box="[1198,1304,908,933]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFADE199948ECE1A1" box="[1318,1436,908,933]" pageId="89" pageNumber="90" value="1958-06-07">7-VI-1958</date>
(ANIC); 1 pupal exuviae, 1 pupal exuviae (of
<typeStatus id="5567887DBD77302FFD5219BA4E78E1CC" box="[682,776,943,968]" pageId="89" pageNumber="90" type="allotype">allotype</typeStatus>
), Gove,
<collectingRegion id="4818F83DBD77302FFC9219BA494CE1CC" box="[874,1084,943,968]" country="Australia" name="Northern Territory" pageId="89" pageNumber="90">Northern Territory</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFBB019BA49C2E1CC" box="[1096,1202,943,968]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFB4719BA4863E1CC" box="[1215,1299,943,968]" pageId="89" pageNumber="90" value="1970-02">II-1970</date>
(ANIC).
<taxonomicName id="4DDC4D5CBD77302FFA7A19A44D8CE1E9" authorityName="Borkent" authorityYear="2014" class="Insecta" family="Ceratopogonidae" genus="Nilobezzia" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="whartoni">
<emphasis id="B8A8EACDBD77302FFA7A19A44D8CE1E9" italics="true" pageId="89" pageNumber="90">N. whartoni</emphasis>
</taxonomicName>
: 1 pupal exuviae, Spencer,
<collectingRegion id="4818F83DBD77302FFDC419C14E7AE1E9" box="[572,778,980,1005]" country="Australia" name="New South Wales" pageId="89" pageNumber="90">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFCE019C14EF1E1E9" box="[792,897,980,1005]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFC7719C14E8AE1E9" box="[911,1018,980,1005]" pageId="89" pageNumber="90" value="1965-02-04">4-II-1965</date>
(ANIC); 8 pupal exuviae, Breakfast Creek, Spencer,
<collectingRegion id="4818F83DBD77302FFEAA19ED4F50E614" box="[338,544,1015,1040]" country="Australia" name="New South Wales" pageId="89" pageNumber="90">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFDD519E24FE6E614" box="[557,662,1015,1040]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFD5B19E24E7FE614" box="[675,783,1015,1040]" pageId="89" pageNumber="90" value="1965-02-04">4-II-1965</date>
(ANIC); 1 pupal exuviae, Oystershell Road.,
<collectingRegion id="4818F83DBD77302FFAE519ED4DAEE631" country="Australia" name="New South Wales" pageId="89" pageNumber="90">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFF111E094C22E631" box="[233,338,1052,1077]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFEA61E094CB9E631" box="[350,457,1052,1077]" pageId="89" pageNumber="90" value="1965-02-04">4-II-1965</date>
(ANIC).
<emphasis id="B8A8EACDBD77302FFDCD1E084F38E630" box="[565,584,1053,1076]" italics="true" pageId="89" pageNumber="90">N</emphasis>
. sp.: 1 pupal exuviae, Constance Bay,
<collectingRegion id="4818F83DBD77302FFC031E094923E631" box="[1019,1107,1052,1077]" country="Canada" name="Ontario" pageId="89" pageNumber="90">Ontario</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFBA71E0949C6E631" box="[1119,1206,1052,1077]" name="Canada" pageId="89" pageNumber="90">Canada</collectingCountry>
,
<date id="FE62101FBD77302FFB391E094848E631" box="[1217,1336,1052,1077]" pageId="89" pageNumber="90" value="1979-06-08">8-VI-1979</date>
(CNCI); 1 pupal exuviae (in glycerin), Kruger National Park,
<collectingCountry id="F2CB764FBD77302FFCF81E2A4EE4E65C" box="[768,916,1087,1112]" name="South Africa" pageId="89" pageNumber="90">South Africa</collectingCountry>
(ANIC); 9 pupal exuviae, Bindoon,
<collectingRegion id="4818F83DBD77302FFAB91E554C71E679" country="Australia" name="Western Australia" pageId="89" pageNumber="90">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD77302FFEF41E714C05E679" box="[268,373,1124,1149]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
,
<date id="FE62101FBD77302FFE781E714C8BE679" box="[384,507,1124,1149]" pageId="89" pageNumber="90" value="1985-10-27">27-X-1985</date>
(ANIC).
</paragraph>
</subSubSection>
</treatment>
</document>
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