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<document id="A317A86319C6EF09483CE48D687F49CA" ID-CLB-Dataset="3916" ID-DOI="10.11646/zootaxa.3879.1.1" ID-GBIF-Dataset="4b976ba3-1a7b-4dd5-8afb-630b7cc8a695" ID-ISSN="1175-5326" ID-Zenodo-Dep="4949051" ID-ZooBank="6423894B-97D9-4286-ABB9-D4AF072B57FD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623702926238" checkinUser="felipe" docAuthor="Borkent, Art" docDate="2014" docId="027587C9BD023046FD521ACB4E15E1CC" docLanguage="en" docName="zt03879p327.pdf.imf" docOrigin="Zootaxa 3879 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dasyhelea Kieffer" docType="treatment" docVersion="11" lastPageNumber="47" masterDocId="FE4CFFB1BD2E3076FFF81A154D70E204" masterDocTitle="The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera" masterLastPageNumber="327" masterPageNumber="1" pageNumber="45" updateTime="1699015668130" updateUser="ExternalLinkService">
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<mods:title id="E48A95A8447C6D54A9E5218C4311D113">The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera</mods:title>
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<mods:namePart id="F332AB8A3B214D26445A20BDFEE9137C">Borkent, Art</mods:namePart>
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<mods:date id="04D2EA3B8C244953F759EF8A8593CF65">2014</mods:date>
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<mods:number id="A068C754CAA9FD807A8CF5C5A5F63D79">2014-10-31</mods:number>
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<treatment id="027587C9BD023046FD521ACB4E15E1CC" ID-DOI="http://doi.org/10.5281/zenodo.5592974" ID-GBIF-Taxon="189161069" ID-Zenodo-Dep="5592974" LSID="urn:lsid:plazi:treatment:027587C9BD023046FD521ACB4E15E1CC" httpUri="http://treatment.plazi.org/id/027587C9BD023046FD521ACB4E15E1CC" lastPageId="48" lastPageNumber="47" pageId="44" pageNumber="45">
<subSubSection id="C2C66554BD02305AFD521ACB4EF8E2FC" box="[682,904,222,248]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8A6336DFBD02305AFD521ACB4EF8E2FC" blockId="44.[299,1288,222,282]" box="[682,904,222,248]" pageId="44" pageNumber="45">
<taxonomicName id="4DDC4D5CBD02305AFD521ACB4EF8E2FC" ID-CoL="4224" ID-ENA="41822" authority="Kieffer" authorityName="Kieffer" box="[682,904,222,248]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD02305AFD521ACB4EF8E2FC" bold="true" box="[682,904,222,248]" pageId="44" pageNumber="45">
<emphasis id="B8A8EACDBD02305AFD521ACB4E57E2FC" bold="true" box="[682,807,222,248]" italics="true" pageId="44" pageNumber="45">Dasyhelea</emphasis>
Kieffer
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD02305AFED31B144878E31E" box="[299,1288,257,282]" pageId="44" pageNumber="45" type="description">
<paragraph id="8A6336DFBD02305AFED31B144878E31E" blockId="44.[299,1288,222,282]" box="[299,1288,257,282]" pageId="44" pageNumber="45">
(
<figureCitation id="12E72A5ABD02305AFECB1B144CD1E31E" box="[307,417,257,282]" captionStart="FIGURE 11" captionStartId="256.[151,250,1845,1867]" captionTargetBox="[213,1374,203,1806]" captionTargetId="figure-145@256.[191,1386,193,1810]" captionTargetPageId="256" captionText="FIGURE 11. Habitus. A. Forcipomyia (Thyridomyia) rugosa, male, in dorsal view (from Chan &amp; LeRoux 1970). B. Forcipomyia (Pterobosca) crinume, female, in ventral view (from Tokunaga 1932). C. Forcipomyia (Lasiohelea) rugosa, male, in dorsal view (from Saunders 1964). D. Atrichopogon hexastichus, male, in dorsal view (from Nielsen 1951). E. Atrichopogon wirthi, female, in dorsal view (from Chan &amp; Linley 1988). F. Atrichopogon muelleri, sex unknown, in dorsal view, with larval exuviae on posterior of abdomen (from Szadziewski et al. 1997). G. Dasyhelea vericolor, sex unknown, in dorsal view (from Szadziewski et al. 1997). H. Schizonyxhelea bulla, sex unknown, in dorsal view (from Thomsen 1937)." figureDoi="http://doi.org/10.5281/zenodo.4949094" httpUri="https://zenodo.org/record/4949094/files/figure.png" pageId="44" pageNumber="45">Figs. 11G</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFE541B144CA8E31E" box="[428,472,257,282]" captionStart="FIGURE 14" captionStartId="259.[151,250,1923,1945]" captionTargetBox="[152,1436,177,1909]" captionTargetId="figure-97@259.[151,1436,175,1914]" captionTargetPageId="259" captionText="FIGURE 14. Anterior portion of heads. A. Austroconops mcmillani, in ventral view, dorsal apotome not present. B. Leptoconops bequaerti, in anterior view (distorted slide specimen). C. Forcipomyia (Phytohelea) sp. in anterior view (Florida, USA). D. Forcipomyia (Phytohelea) sp. in anteroventral view (Florida, USA). E. Atrichopogon sp., in anterior view (from 6.5 km NW Enderby, BC, Canada). F. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949108" httpUri="https://zenodo.org/record/4949108/files/figure.png" pageId="44" pageNumber="45">14F</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFE1C1B144F40E31E" box="[484,560,257,282]" captionStart="FIGURE 18" captionStartId="263.[151,250,1850,1872]" captionTargetBox="[164,1422,191,1811]" captionTargetId="figure-119@263.[151,1436,185,1842]" captionTargetPageId="263" captionText="FIGURE 18. Dorsal apotomes, anterior view. A. Austroconops mcmillani (from Borkent &amp; Craig 2004). B. Leptoconops bequaerti. C. Forcipomyia (Phytohelea) n. sp. near F. antiguensis. D. Forcipomyia (Euprojoannisia) sp. (undescribed, from Peru). E. Atrichopogon sp. (undescribed, from Costa Rica). F. Dasyhelea insons (from Mayer 1934c). G. Dasyhelea tersa (from Mayer 1934c). H. Dasyhelea assimilis (from Mayer 1934c). I. Dasyhelea pseudocincta (from Waugh &amp; Wirth 1976, modified). J. Dasyhelea pseudoincisurata (from Waugh &amp; Wirth 1976, modified). K. Dasyhelea messersmithi (from Waugh &amp; Wirth 1976, modified). L. Dasyhelea atlantis (from Waugh &amp; Wirth 1976, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949124" httpUri="https://zenodo.org/record/4949124/files/figure.png" pageId="44" pageNumber="45">18F–L</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFDC41B144FF9E31E" box="[572,649,257,282]" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="44" pageNumber="45">23F–G</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFD6D1B144F90E31E" box="[661,736,257,282]" captionStart="FIGURE 29" captionStartId="274.[151,250,1822,1844]" captionTargetBox="[165,1429,184,1785]" captionTargetId="figure-146@274.[165,1430,181,1801]" captionTargetPageId="274" captionText="FIGURE 29. Dorsal setae from left side of thoraces. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Leptoconops linleyi. D. Forcipomyia (Euprojoannisia) sp. (from Peru). E. Forcipomyia (Forcipomyia) bipunctata. F. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. G. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). H. Dasyhelea paracincta. I. Dasyhelea atlantis. J. Dasyhelea pseudoincisurata. K. Culicoides haematopotus. L. Culicoides phlebotomus. M. Paradasyhelea albipunctata. N. Ceratopogon boomerangus. O. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). P. Alluaudomyia needhami. Q. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). R. Austrohelea campbellensis. S. Stilobezzia antennalis. T. Schizonyxhelea forattinii (from Borkent 2000a)." figureDoi="http://doi.org/10.5281/zenodo.4949172" httpUri="https://zenodo.org/record/4949172/files/figure.png" pageId="44" pageNumber="45">29H–J</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFD141B144E47E31E" box="[748,823,257,282]" captionStart="FIGURE 32" captionStartId="277.[151,250,1887,1909]" captionTargetBox="[174,1419,183,1869]" captionTargetId="figure-104@277.[173,1420,183,1869]" captionTargetPageId="277" captionText="FIGURE 32. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Austroconops mcmillani. B. Leptoconops linleyi. C. Forcipomyia (Phytohelea) sp. (Florida, USA). D. Atrichopogon sp. (from 6.5 km NW Enderby, BC, Canada). E. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada). F. Dasyhelea oppresa. G. Culicoides denticulatus. H. Ceratopogon nr. abstrusus. I. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). J. Alluaudomyia bella. K. Baeodasymyia michaeli. L. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949182" httpUri="https://zenodo.org/record/4949182/files/figure.png" pageId="44" pageNumber="45">32E–F</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFCBA1B144E01E31E" box="[834,881,257,282]" captionStart="FIGURE 34" captionStartId="279.[151,250,1931,1953]" captionTargetBox="[195,1385,213,1901]" captionTargetId="figure-69@279.[187,1395,201,1905]" captionTargetPageId="279" captionText="FIGURE 34. Legs, wings and apices of the antennae, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). E. Dasyhelea paracincta. F. Culicoides sp. (from 3 km E. Carp, Ontario, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949186" httpUri="https://zenodo.org/record/4949186/files/figure.png" pageId="44" pageNumber="45">34E</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFC851B144EBFE31E" box="[893,975,257,282]" captionStart="FIGURE 43" captionStartId="288.[151,250,1735,1757]" captionTargetBox="[241,1367,200,1690]" captionTargetId="figure-197@288.[196,1378,196,1723]" captionTargetPageId="288" captionText="FIGURE 43. Respiratory organs, most in dorsal view, Fig. 43T in lateral view, pattern on tracheal tubes shown only in some. A. Dasyhelea pseudoincisurata (from Waugh &amp; Wirth 1976). B. Dasyhelea grisea (from Waugh &amp; Wirth 1976). C. Dasyhelea traverae (from Waugh &amp; Wirth 1976). D. Dasyhelea pseudocincta (from Waugh &amp; Wirth 1976). E. Dasyhelea leptobranchia (from Waugh &amp; Wirth 1976). F. Dasyhelea brevicornis (from Waugh &amp; Wirth 1976). G. Dasyhelea hitchcocki (from Wirth 1976a). H. Culicoides chiopterus (from Jamnback 1965). I. Culicoides furens (from Jamnback 1965). J. Culicoides guttipennis (from Jamnback 1965). K. Culicoides dewulfi (from Kettle &amp; Lawson 1952, as C. pseudochiopterus). L. Culicoides punctatus (from Kettle &amp; Lawson 1952). M. Culicoides sanguisuga (from Jamnback 1965). N. Culicoides variipennis (from Jamnback 1965). O. Paradasyhelea albipunctata (from Elson-Harris &amp; Kettle 1985a). P. Ceratopogon inverecundus (from Borkent &amp; Grogan 1995). Q. Ceratopogon willisi (from Borkent &amp; Grogan 1995). R. Brachypogon sp. (from Virginia, USA). S. Alluaudomyia needhami (from Wirth &amp; Grogan 1981). T. Alluaudomyia fumosipennis (from Debenham 1971)." figureDoi="http://doi.org/10.5281/zenodo.4949206" httpUri="https://zenodo.org/record/4949206/files/figure.png" pageId="44" pageNumber="45">43A–G</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFC221B144977E31E" box="[986,1031,257,282]" captionStart="FIGURE 47" captionStartId="292.[151,250,1952,1974]" captionTargetBox="[209,1375,191,1853]" captionTargetId="figure-74@292.[201,1383,188,1927]" captionTargetPageId="292" captionText="FIGURE 47. Metathoraces and tergites 1 (in dorsal view). A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Forcipomyia) bipunctata. D. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. E. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). F. Dasyhelea pseudoincisuata." figureDoi="http://doi.org/10.5281/zenodo.4949218" httpUri="https://zenodo.org/record/4949218/files/figure.png" pageId="44" pageNumber="45">47F</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFBEA1B144933E31E" box="[1042,1091,257,282]" captionStart="FIGURE 56" captionStartId="301.[151,250,1953,1975]" captionTargetBox="[221,1382,193,1855]" captionTargetId="figure-49@301.[213,1395,188,1927]" captionTargetPageId="301" captionText="FIGURE 56. Abdominal segment four, in dorsal and ventral view (shagreen not shown). A. Forcipomyia dominicana. B. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). C. Dasyhelea paracincta." figureDoi="http://doi.org/10.5281/zenodo.4949250" httpUri="https://zenodo.org/record/4949250/files/figure.png" pageId="44" pageNumber="45">56C</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFBB61B1449D1E31E" box="[1102,1185,257,282]" captionStart="FIGURE 72" captionStartId="317.[151,250,1765,1787]" captionTargetBox="[228,1373,193,1750]" captionTargetId="figure-192@317.[227,1378,193,1750]" captionTargetPageId="317" captionText="FIGURE 72. Abdominal segments 9. A. Austroconops mcmillani, male, in dorsal and ventral view (from Borkent &amp; Craig 2004). B. Leptoconops whitseli, male, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). C. Leptoconops whitseli, female, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). D. Leptoconops asilomar, male, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). E. Leptoconops asilomar, female, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). F. Forcipomyia anabaenae, female, in dorsal view (from Chan &amp; Saunders 1965). G. Forcipomyia eques, male, in ventral view (from Chan &amp; LeRoux 1965). H. Forcipomyia rugosa, female, in dorsal view (from Chan &amp; LeRoux 1970). I. Forcipomyia brevis, female, in dorsal view (from Mayer 1934a). J. Atrichopogon wirthi, male, in dorsal view, arrow indicating anteriorly directed spines (from Chan &amp; Linley 1988). K. Dasyhelea contigua, female, in ventral view (from Mayer 1934c). L. Dasyhelea grata, male, in ventral view (from Mayer 1934c)." figureDoi="http://doi.org/10.5281/zenodo.4949308" httpUri="https://zenodo.org/record/4949308/files/figure.png" pageId="44" pageNumber="45">72K–L</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFB541B144870E31E" box="[1196,1280,257,282]" captionStart="FIGURE 73" captionStartId="318.[151,250,1765,1787]" captionTargetBox="[213,1371,197,1757]" captionTargetId="figure-189@318.[211,1371,188,1757]" captionTargetPageId="318" captionText="FIGURE 73. Abdominal segments 9. A. Dasyhelea laeta, female, in ventral view (from Mayer 1934c). B. Dasyhelea assimilis, male, in lateral view with ventral to the right (from Mayer 1934c). C. Dasyhelea assimilis, male, in lateral view with ventral to the right (from Mayer 1934c). D. Dasyhelea grisea, male, in dorsal view (from Waugh &amp; Wirth 1976). E. Dasyhelea chani, female, in dorsal view (from Wirth &amp; Linley 1990). F. Dasyhelea atlantis, female, in ventral view (from Waugh &amp; Wirth 1976). G. Dasyhelea gluchovae, female, in ventral view (from Brodskaya 1996). H. Culicoides haematopotus, female, in dorsal and ventral view (shagreen not shown). I. Paradasyhelea albipunctata, female, in dorsal view (Elson-Harris &amp; Kettle 1985a). J. Ceratopogon boomerangus, female, in ventral view (modified from Borkent &amp; Grogan 1995). K. Brachypogon artemis, female, in dorsal view (from Debenham 1991). L. Alluaudomyia appendiculata, female, in dorsal view (from Elson-Harris &amp; Kettle 1985b)." figureDoi="http://doi.org/10.5281/zenodo.4949314" httpUri="https://zenodo.org/record/4949314/files/figure.png" pageId="44" pageNumber="45">73A–G</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD02305AFF6F1B5C4922E3CA" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="8A6336DFBD02305AFF6F1B5C4922E3CA" blockId="44.[151,1437,329,2010]" pageId="44" pageNumber="45">
<emphasis id="B8A8EACDBD02305AFF6F1B5C4C44E366" bold="true" box="[151,308,329,354]" pageId="44" pageNumber="45">DIAGNOSIS</emphasis>
: Only pupa of
<taxonomicName id="4DDC4D5CBD02305AFE141B5C4FDBE366" authorityName="Borkent" authorityYear="2014" box="[492,683,329,354]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
with the labium clearly split medially by an elongate suture (or overlapping halves) (
<figureCitation id="12E72A5ABD02305AFE7D1B784F67E382" box="[389,535,365,390]" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="44" pageNumber="45">Figs. 23F–G</figureCitation>
); most species have uniquely complex terminal processes each of which is bifid or forked and bearing two setae; also most species have the posterior sensilla of abdominal segments 3–8 arranged on broad shelf-like tubercles, forming a partial and unique ring around each segment.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD02305AFF3F1BCC4CEFE5B2" pageId="44" pageNumber="45" type="description">
<paragraph id="8A6336DFBD02305AFF3F1BCC4CEFE5B2" blockId="44.[151,1437,329,2010]" pageId="44" pageNumber="45">
<emphasis id="B8A8EACDBD02305AFF3F1BCC4CFAE3F6" bold="true" box="[199,394,473,498]" pageId="44" pageNumber="45">DESCRIPTION</emphasis>
: Habitus as in
<figureCitation id="12E72A5ABD02305AFDC31BCC4FD0E3F6" box="[571,672,473,498]" captionStart="FIGURE 11" captionStartId="256.[151,250,1845,1867]" captionTargetBox="[213,1374,203,1806]" captionTargetId="figure-145@256.[191,1386,193,1810]" captionTargetPageId="256" captionText="FIGURE 11. Habitus. A. Forcipomyia (Thyridomyia) rugosa, male, in dorsal view (from Chan &amp; LeRoux 1970). B. Forcipomyia (Pterobosca) crinume, female, in ventral view (from Tokunaga 1932). C. Forcipomyia (Lasiohelea) rugosa, male, in dorsal view (from Saunders 1964). D. Atrichopogon hexastichus, male, in dorsal view (from Nielsen 1951). E. Atrichopogon wirthi, female, in dorsal view (from Chan &amp; Linley 1988). F. Atrichopogon muelleri, sex unknown, in dorsal view, with larval exuviae on posterior of abdomen (from Szadziewski et al. 1997). G. Dasyhelea vericolor, sex unknown, in dorsal view (from Szadziewski et al. 1997). H. Schizonyxhelea bulla, sex unknown, in dorsal view (from Thomsen 1937)." figureDoi="http://doi.org/10.5281/zenodo.4949094" httpUri="https://zenodo.org/record/4949094/files/figure.png" pageId="44" pageNumber="45">Fig. 11G</figureCitation>
. Total length =
<quantity id="4D249B3ABD02305AFCA11BCC4971E3F5" box="[857,1025,473,498]" metricMagnitude="-3" metricUnit="m" metricValue="2.895" metricValueMax="4.16" metricValueMin="1.63" pageId="44" pageNumber="45" unit="mm" value="2.895" valueMax="4.16" valueMin="1.63">1.63–4.16 mm</quantity>
. Without larval exuviae retained on abdomen. Exuviae with flagellum appressed against lateral margin of face (
<figureCitation id="12E72A5ABD02305AFC161BE84923E012" box="[1006,1107,509,534]" captionStart="FIGURE 14" captionStartId="259.[151,250,1923,1945]" captionTargetBox="[152,1436,177,1909]" captionTargetId="figure-97@259.[151,1436,175,1914]" captionTargetPageId="259" captionText="FIGURE 14. Anterior portion of heads. A. Austroconops mcmillani, in ventral view, dorsal apotome not present. B. Leptoconops bequaerti, in anterior view (distorted slide specimen). C. Forcipomyia (Phytohelea) sp. in anterior view (Florida, USA). D. Forcipomyia (Phytohelea) sp. in anteroventral view (Florida, USA). E. Atrichopogon sp., in anterior view (from 6.5 km NW Enderby, BC, Canada). F. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949108" httpUri="https://zenodo.org/record/4949108/files/figure.png" pageId="44" pageNumber="45">Fig. 14F</figureCitation>
). Ecdysial tear not extending at all (
<figureCitation id="12E72A5ABD02305AFF1B18344C21E03E" box="[227,337,545,570]" captionStart="FIGURE 14" captionStartId="259.[151,250,1923,1945]" captionTargetBox="[152,1436,177,1909]" captionTargetId="figure-97@259.[151,1436,175,1914]" captionTargetPageId="259" captionText="FIGURE 14. Anterior portion of heads. A. Austroconops mcmillani, in ventral view, dorsal apotome not present. B. Leptoconops bequaerti, in anterior view (distorted slide specimen). C. Forcipomyia (Phytohelea) sp. in anterior view (Florida, USA). D. Forcipomyia (Phytohelea) sp. in anteroventral view (Florida, USA). E. Atrichopogon sp., in anterior view (from 6.5 km NW Enderby, BC, Canada). F. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949108" httpUri="https://zenodo.org/record/4949108/files/figure.png" pageId="44" pageNumber="45">Figs. 14F</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFEA718344CE2E03E" box="[351,402,545,570]" captionStart="FIGURE 79" captionStartId="324.[151,250,1861,1883]" captionTargetBox="[192,1396,201,1846]" captionTargetId="figure-148@324.[184,1406,195,1847]" captionTargetPageId="324" captionText="FIGURE 79. Evolution of ecdysial sutures and splits in pupal exuviae of Ceratopogonidae, head and anterior portion of thorax in ventral view. Numbers refer to character states of characters 2 and 8 in text. Red indicates ecdysial splits. A. Leptoconopinae. B. Forcipomyiinae. C. Dasyheleinae. D. Most Ceratopogonini. E. Most Ceratopogonini, Clinohelea. F. Allohelea, Atyphohelea, Parabezzia. G. Heteromyia, Pellucidomyia, Hebetula. H. most Sphaeromiini s. lat., most Palpomyiini, Paryphoconus; some Bezzia are nearly completed fused (as in Fig. 79I), with only the dorsal apotome separate, along with an ecdysial split along the base of the antenna and the dorsal thoracic split. I. Jenkinshelea, Macropeza, Mallochohelea, Probezzia, Neobezzia." figureDoi="http://doi.org/10.5281/zenodo.4949336" httpUri="https://zenodo.org/record/4949336/files/figure.png" pageId="44" pageNumber="45">79C</figureCitation>
); posterolateral portion of eye.
<emphasis id="B8A8EACDBD02305AFD0618344E4EE03E" bold="true" box="[766,830,545,570]" pageId="44" pageNumber="45">Head</emphasis>
: Dorsal apotome (
<figureCitation id="12E72A5ABD02305AFBEF183449D8E03E" box="[1047,1192,545,570]" captionStart="FIGURE 18" captionStartId="263.[151,250,1850,1872]" captionTargetBox="[164,1422,191,1811]" captionTargetId="figure-119@263.[151,1436,185,1842]" captionTargetPageId="263" captionText="FIGURE 18. Dorsal apotomes, anterior view. A. Austroconops mcmillani (from Borkent &amp; Craig 2004). B. Leptoconops bequaerti. C. Forcipomyia (Phytohelea) n. sp. near F. antiguensis. D. Forcipomyia (Euprojoannisia) sp. (undescribed, from Peru). E. Atrichopogon sp. (undescribed, from Costa Rica). F. Dasyhelea insons (from Mayer 1934c). G. Dasyhelea tersa (from Mayer 1934c). H. Dasyhelea assimilis (from Mayer 1934c). I. Dasyhelea pseudocincta (from Waugh &amp; Wirth 1976, modified). J. Dasyhelea pseudoincisurata (from Waugh &amp; Wirth 1976, modified). K. Dasyhelea messersmithi (from Waugh &amp; Wirth 1976, modified). L. Dasyhelea atlantis (from Waugh &amp; Wirth 1976, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949124" httpUri="https://zenodo.org/record/4949124/files/figure.png" pageId="44" pageNumber="45">Figs. 18F–L</figureCitation>
), with ventral line of weakness, with or without dorsomedial tubercle, without central dome; dorsolateral cephalic sclerite (as in
<figureCitation id="12E72A5ABD02305AFA8818504DB6E086" captionStart="FIGURE 13" captionStartId="258.[151,250,1897,1919]" captionTargetBox="[172,1416,193,1876]" captionTargetId="figure-99@258.[172,1416,193,1876]" captionTargetPageId="258" captionText="FIGURE 13. Anterior portion of heads and thoraces, in anterior or anterodorsal view. A. Leptoconops kerteszi, ecdysial sutures of head enhanced to show position. B. Culicoides denticulatus. C. Ceratopogon nr. abstrusus. D. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). E. Alluaudomyia bella, dorsal apotome absent. F. Clinohelea curriei. G. Pellucidomyia leei. H. Mallochohelea caudellii. I. Bezzia sp. (from Bolean Lake, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949104" httpUri="https://zenodo.org/record/4949104/files/figure.png" pageId="44" pageNumber="45">Fig. 13B</figureCitation>
) separated from scutum by thin cuticle, separate from scutum upon emergence, each side separated medially by dorsal apotome in whole pupa; mouthparts (
<figureCitation id="12E72A5ABD02305AFD3018984E2CE0A2" box="[712,860,653,678]" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="44" pageNumber="45">Figs. 23F–G</figureCitation>
) with mandible, lacinia absent; palpus extending posterior to posterolateral margin of labium; labium divided medially by longitudinal suture or two halves slightly overlapping; apex of antenna (
<figureCitation id="12E72A5ABD02305AFE1518C04F20E0EA" box="[493,592,725,750]" captionStart="FIGURE 34" captionStartId="279.[151,250,1931,1953]" captionTargetBox="[195,1385,213,1901]" captionTargetId="figure-69@279.[187,1395,201,1905]" captionTargetPageId="279" captionText="FIGURE 34. Legs, wings and apices of the antennae, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). E. Dasyhelea paracincta. F. Culicoides sp. (from 3 km E. Carp, Ontario, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949186" httpUri="https://zenodo.org/record/4949186/files/figure.png" pageId="44" pageNumber="45">Fig. 34E</figureCitation>
) anterior to posterior extent of midlength portion of midleg (portion lateral to mesosternum); sensilla: dorsal apotomals (
<figureCitation id="12E72A5ABD02305AFD5818EC4E41E116" box="[672,817,761,786]" captionStart="FIGURE 18" captionStartId="263.[151,250,1850,1872]" captionTargetBox="[164,1422,191,1811]" captionTargetId="figure-119@263.[151,1436,185,1842]" captionTargetPageId="263" captionText="FIGURE 18. Dorsal apotomes, anterior view. A. Austroconops mcmillani (from Borkent &amp; Craig 2004). B. Leptoconops bequaerti. C. Forcipomyia (Phytohelea) n. sp. near F. antiguensis. D. Forcipomyia (Euprojoannisia) sp. (undescribed, from Peru). E. Atrichopogon sp. (undescribed, from Costa Rica). F. Dasyhelea insons (from Mayer 1934c). G. Dasyhelea tersa (from Mayer 1934c). H. Dasyhelea assimilis (from Mayer 1934c). I. Dasyhelea pseudocincta (from Waugh &amp; Wirth 1976, modified). J. Dasyhelea pseudoincisurata (from Waugh &amp; Wirth 1976, modified). K. Dasyhelea messersmithi (from Waugh &amp; Wirth 1976, modified). L. Dasyhelea atlantis (from Waugh &amp; Wirth 1976, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949124" httpUri="https://zenodo.org/record/4949124/files/figure.png" pageId="44" pageNumber="45">Figs. 18F–L</figureCitation>
)—1 short to elongate seta, 1 campaniform sensillum; dorsolateral cephalic sclerite sensilla—2 small setae, 1 campaniform sensillum; clypeal-labrals (
<figureCitation id="12E72A5ABD02305AFB2219084819E132" box="[1242,1385,797,822]" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="44" pageNumber="45">Figs. 23F–G</figureCitation>
)—2 minute setae; oculars (
<figureCitation id="12E72A5ABD02305AFE6219544F5DE15E" box="[410,557,833,858]" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="44" pageNumber="45">Figs. 23F–G</figureCitation>
)—0–1 seta, 1 campaniform sensillum.
<emphasis id="B8A8EACDBD02305AFC0A1954493BE15E" bold="true" box="[1010,1099,833,858]" pageId="44" pageNumber="45">Thorax</emphasis>
: Prothoracic extension (
<figureCitation id="12E72A5ABD02305AFA9C19544D96E17A" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="44" pageNumber="45">Figs. 23F–G</figureCitation>
) absent; mesonotum with or without short tubercles, not extending posteromedially to extending posteromedially, completely dividing metathorax medially (
<figureCitation id="12E72A5ABD02305AFCB3199C4EC4E1A6" box="[843,948,905,930]" captionStart="FIGURE 47" captionStartId="292.[151,250,1952,1974]" captionTargetBox="[209,1375,191,1853]" captionTargetId="figure-74@292.[201,1383,188,1927]" captionTargetPageId="292" captionText="FIGURE 47. Metathoraces and tergites 1 (in dorsal view). A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Forcipomyia) bipunctata. D. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. E. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). F. Dasyhelea pseudoincisuata." figureDoi="http://doi.org/10.5281/zenodo.4949218" httpUri="https://zenodo.org/record/4949218/files/figure.png" pageId="44" pageNumber="45">Fig. 47F</figureCitation>
); respiratory organ (
<figureCitation id="12E72A5ABD02305AFB5C199C484BE1A6" box="[1188,1339,905,930]" captionStart="FIGURE 43" captionStartId="288.[151,250,1735,1757]" captionTargetBox="[241,1367,200,1690]" captionTargetId="figure-197@288.[196,1378,196,1723]" captionTargetPageId="288" captionText="FIGURE 43. Respiratory organs, most in dorsal view, Fig. 43T in lateral view, pattern on tracheal tubes shown only in some. A. Dasyhelea pseudoincisurata (from Waugh &amp; Wirth 1976). B. Dasyhelea grisea (from Waugh &amp; Wirth 1976). C. Dasyhelea traverae (from Waugh &amp; Wirth 1976). D. Dasyhelea pseudocincta (from Waugh &amp; Wirth 1976). E. Dasyhelea leptobranchia (from Waugh &amp; Wirth 1976). F. Dasyhelea brevicornis (from Waugh &amp; Wirth 1976). G. Dasyhelea hitchcocki (from Wirth 1976a). H. Culicoides chiopterus (from Jamnback 1965). I. Culicoides furens (from Jamnback 1965). J. Culicoides guttipennis (from Jamnback 1965). K. Culicoides dewulfi (from Kettle &amp; Lawson 1952, as C. pseudochiopterus). L. Culicoides punctatus (from Kettle &amp; Lawson 1952). M. Culicoides sanguisuga (from Jamnback 1965). N. Culicoides variipennis (from Jamnback 1965). O. Paradasyhelea albipunctata (from Elson-Harris &amp; Kettle 1985a). P. Ceratopogon inverecundus (from Borkent &amp; Grogan 1995). Q. Ceratopogon willisi (from Borkent &amp; Grogan 1995). R. Brachypogon sp. (from Virginia, USA). S. Alluaudomyia needhami (from Wirth &amp; Grogan 1981). T. Alluaudomyia fumosipennis (from Debenham 1971)." figureDoi="http://doi.org/10.5281/zenodo.4949206" httpUri="https://zenodo.org/record/4949206/files/figure.png" pageId="44" pageNumber="45">Figs. 43A–G</figureCitation>
) length/ width = 2.81–29.14, highly variable, moderately to very elongate, apex pointed to truncate, somewhat flattened or circular in cross-section, with pores closely abutting or slightly separated at apex of respiratory organ, arranged in single line of varying shapes, with or without additional, more basal pores, outer surface with or without annulations, with or without spicules, without pedicel, base with short, strung out posteromedial apodeme, membranous base of respiratory organ short, tracheal tube straight to slightly curved along length, smooth or with spirals restricted to base; wing (
<figureCitation id="12E72A5ABD02305AFDFE1E744F1BE67E" box="[518,619,1121,1146]" captionStart="FIGURE 34" captionStartId="279.[151,250,1931,1953]" captionTargetBox="[195,1385,213,1901]" captionTargetId="figure-69@279.[187,1395,201,1905]" captionTargetPageId="279" captionText="FIGURE 34. Legs, wings and apices of the antennae, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). E. Dasyhelea paracincta. F. Culicoides sp. (from 3 km E. Carp, Ontario, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949186" httpUri="https://zenodo.org/record/4949186/files/figure.png" pageId="44" pageNumber="45">Fig. 34E</figureCitation>
) with apical tubercle lateral to apex of hind leg, separated medially by fore-, midlegs; halter apex and hind leg (
<figureCitation id="12E72A5ABD02305AFD881E904F8EE69A" box="[624,766,1157,1182]" captionStart="FIGURE 32" captionStartId="277.[151,250,1887,1909]" captionTargetBox="[174,1419,183,1869]" captionTargetId="figure-104@277.[173,1420,183,1869]" captionTargetPageId="277" captionText="FIGURE 32. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Austroconops mcmillani. B. Leptoconops linleyi. C. Forcipomyia (Phytohelea) sp. (Florida, USA). D. Atrichopogon sp. (from 6.5 km NW Enderby, BC, Canada). E. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada). F. Dasyhelea oppresa. G. Culicoides denticulatus. H. Ceratopogon nr. abstrusus. I. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). J. Alluaudomyia bella. K. Baeodasymyia michaeli. L. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949182" httpUri="https://zenodo.org/record/4949182/files/figure.png" pageId="44" pageNumber="45">Figs. 32E–F</figureCitation>
) just separate; halter apex abutting anterolateral knob-like extension of tergite 2; legs (
<figureCitation id="12E72A5ABD02305AFE341EBC4F5FE6C6" box="[460,559,1193,1218]" captionStart="FIGURE 34" captionStartId="279.[151,250,1931,1953]" captionTargetBox="[195,1385,213,1901]" captionTargetId="figure-69@279.[187,1395,201,1905]" captionTargetPageId="279" captionText="FIGURE 34. Legs, wings and apices of the antennae, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). E. Dasyhelea paracincta. F. Culicoides sp. (from 3 km E. Carp, Ontario, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949186" httpUri="https://zenodo.org/record/4949186/files/figure.png" pageId="44" pageNumber="45">Fig. 34E</figureCitation>
) with lateral margin of foreleg near midlength of wing evenly curved; hind leg visible at lateral margin of wing (
<figureCitation id="12E72A5ABD02305AFDF31ED84FE9E6E2" box="[523,665,1229,1254]" captionStart="FIGURE 32" captionStartId="277.[151,250,1887,1909]" captionTargetBox="[174,1419,183,1869]" captionTargetId="figure-104@277.[173,1420,183,1869]" captionTargetPageId="277" captionText="FIGURE 32. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Austroconops mcmillani. B. Leptoconops linleyi. C. Forcipomyia (Phytohelea) sp. (Florida, USA). D. Atrichopogon sp. (from 6.5 km NW Enderby, BC, Canada). E. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada). F. Dasyhelea oppresa. G. Culicoides denticulatus. H. Ceratopogon nr. abstrusus. I. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). J. Alluaudomyia bella. K. Baeodasymyia michaeli. L. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949182" httpUri="https://zenodo.org/record/4949182/files/figure.png" pageId="44" pageNumber="45">Figs. 32E–F</figureCitation>
); with apex of foreleg moderately anterior to apex of midleg; apex of hind leg slightly dorsal to, partially abutting apex of midleg laterally; sensilla: anteromedials—2 setae; anterolaterals—1 seta, 1 campaniform sensillum; dorsal setae (
<figureCitation id="12E72A5ABD02305AFC5C1F00494BE72A" box="[932,1083,1301,1326]" captionStart="FIGURE 29" captionStartId="274.[151,250,1822,1844]" captionTargetBox="[165,1429,184,1785]" captionTargetId="figure-146@274.[165,1430,181,1801]" captionTargetPageId="274" captionText="FIGURE 29. Dorsal setae from left side of thoraces. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Leptoconops linleyi. D. Forcipomyia (Euprojoannisia) sp. (from Peru). E. Forcipomyia (Forcipomyia) bipunctata. F. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. G. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). H. Dasyhelea paracincta. I. Dasyhelea atlantis. J. Dasyhelea pseudoincisurata. K. Culicoides haematopotus. L. Culicoides phlebotomus. M. Paradasyhelea albipunctata. N. Ceratopogon boomerangus. O. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). P. Alluaudomyia needhami. Q. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). R. Austrohelea campbellensis. S. Stilobezzia antennalis. T. Schizonyxhelea forattinii (from Borkent 2000a)." figureDoi="http://doi.org/10.5281/zenodo.4949172" httpUri="https://zenodo.org/record/4949172/files/figure.png" pageId="44" pageNumber="45">Figs. 29H–J</figureCitation>
)—D-1-T, D-2-T seta, D-3-T campaniform sensillum, D-1-T well anterior of D-2-T; supraalar 2—campaniform sensillum; metathoracics (
<figureCitation id="12E72A5ABD02305AFA971F2C4DB3E772" captionStart="FIGURE 47" captionStartId="292.[151,250,1952,1974]" captionTargetBox="[209,1375,191,1853]" captionTargetId="figure-74@292.[201,1383,188,1927]" captionTargetPageId="292" captionText="FIGURE 47. Metathoraces and tergites 1 (in dorsal view). A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Forcipomyia) bipunctata. D. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. E. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). F. Dasyhelea pseudoincisuata." figureDoi="http://doi.org/10.5281/zenodo.4949218" httpUri="https://zenodo.org/record/4949218/files/figure.png" pageId="44" pageNumber="45">Fig. 47F</figureCitation>
)—2 campaniform sensilla; M-3-T near anterior margin of metathorax.
<emphasis id="B8A8EACDBD02305AFBE41F4849E0E772" bold="true" box="[1052,1168,1373,1398]" pageId="44" pageNumber="45">Abdomen</emphasis>
: without pigmentation pattern or some with two light brown pigment spots near middle of tergites 1–7, segment 2 as wide or slightly wider than segment 3, segments with undivided, thin to thick setae, with rounded to pointed, short, wide, shelf-like tubercles, rarely with elongate tubercles, tergites or sternites entire, each without membranous disc; segment 9 (
<figureCitation id="12E72A5ABD02305AFF671FF84C45E402" box="[159,309,1517,1542]" captionStart="FIGURE 72" captionStartId="317.[151,250,1765,1787]" captionTargetBox="[228,1373,193,1750]" captionTargetId="figure-192@317.[227,1378,193,1750]" captionTargetPageId="317" captionText="FIGURE 72. Abdominal segments 9. A. Austroconops mcmillani, male, in dorsal and ventral view (from Borkent &amp; Craig 2004). B. Leptoconops whitseli, male, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). C. Leptoconops whitseli, female, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). D. Leptoconops asilomar, male, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). E. Leptoconops asilomar, female, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). F. Forcipomyia anabaenae, female, in dorsal view (from Chan &amp; Saunders 1965). G. Forcipomyia eques, male, in ventral view (from Chan &amp; LeRoux 1965). H. Forcipomyia rugosa, female, in dorsal view (from Chan &amp; LeRoux 1970). I. Forcipomyia brevis, female, in dorsal view (from Mayer 1934a). J. Atrichopogon wirthi, male, in dorsal view, arrow indicating anteriorly directed spines (from Chan &amp; Linley 1988). K. Dasyhelea contigua, female, in ventral view (from Mayer 1934c). L. Dasyhelea grata, male, in ventral view (from Mayer 1934c)." figureDoi="http://doi.org/10.5281/zenodo.4949308" httpUri="https://zenodo.org/record/4949308/files/figure.png" pageId="44" pageNumber="45">Figs. 72K–L</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFEBD1FF84CE9E402" box="[325,409,1517,1542]" captionStart="FIGURE 73" captionStartId="318.[151,250,1765,1787]" captionTargetBox="[213,1371,197,1757]" captionTargetId="figure-189@318.[211,1371,188,1757]" captionTargetPageId="318" captionText="FIGURE 73. Abdominal segments 9. A. Dasyhelea laeta, female, in ventral view (from Mayer 1934c). B. Dasyhelea assimilis, male, in lateral view with ventral to the right (from Mayer 1934c). C. Dasyhelea assimilis, male, in lateral view with ventral to the right (from Mayer 1934c). D. Dasyhelea grisea, male, in dorsal view (from Waugh &amp; Wirth 1976). E. Dasyhelea chani, female, in dorsal view (from Wirth &amp; Linley 1990). F. Dasyhelea atlantis, female, in ventral view (from Waugh &amp; Wirth 1976). G. Dasyhelea gluchovae, female, in ventral view (from Brodskaya 1996). H. Culicoides haematopotus, female, in dorsal and ventral view (shagreen not shown). I. Paradasyhelea albipunctata, female, in dorsal view (Elson-Harris &amp; Kettle 1985a). J. Ceratopogon boomerangus, female, in ventral view (modified from Borkent &amp; Grogan 1995). K. Brachypogon artemis, female, in dorsal view (from Debenham 1991). L. Alluaudomyia appendiculata, female, in dorsal view (from Elson-Harris &amp; Kettle 1985b)." figureDoi="http://doi.org/10.5281/zenodo.4949314" httpUri="https://zenodo.org/record/4949314/files/figure.png" pageId="44" pageNumber="45">73A–G</figureCitation>
) highly variable in shape, terminal processes widely separated to closely approximated basally, each projecting posteriorly to laterally, generally shelf-like with two apices or undivided but modified and tapering to pointed apex, in some elongate and slender; sensilla: tergite 1 (
<figureCitation id="12E72A5ABD02305AFC1E1C204938E44A" box="[998,1096,1589,1614]" captionStart="FIGURE 47" captionStartId="292.[151,250,1952,1974]" captionTargetBox="[209,1375,191,1853]" captionTargetId="figure-74@292.[201,1383,188,1927]" captionTargetPageId="292" captionText="FIGURE 47. Metathoraces and tergites 1 (in dorsal view). A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Forcipomyia) bipunctata. D. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. E. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). F. Dasyhelea pseudoincisuata." figureDoi="http://doi.org/10.5281/zenodo.4949218" httpUri="https://zenodo.org/record/4949218/files/figure.png" pageId="44" pageNumber="45">Fig. 47F</figureCitation>
) with 4 setae, 2 campaniform sensilla, including 3 lateral sensilla, D-2-I well posterior from anterior margin of tergite 1, D-3-I absent, D-7-I situated posteriorly; segment 4 (
<figureCitation id="12E72A5ABD02305AFE051C684F2FE492" box="[509,607,1661,1686]" captionStart="FIGURE 56" captionStartId="301.[151,250,1953,1975]" captionTargetBox="[221,1382,193,1855]" captionTargetId="figure-49@301.[213,1395,188,1927]" captionTargetPageId="301" captionText="FIGURE 56. Abdominal segment four, in dorsal and ventral view (shagreen not shown). A. Forcipomyia dominicana. B. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). C. Dasyhelea paracincta." figureDoi="http://doi.org/10.5281/zenodo.4949250" httpUri="https://zenodo.org/record/4949250/files/figure.png" pageId="44" pageNumber="45">Fig. 56C</figureCitation>
)—D-2-IV short to elongate seta, without tubercle, D-3-IV absent; D-8-IV short to (rarely) elongate seta, D-5-IV, D-9-IV absent; D-8-IV without or on wide, low tubercle, D-4-IV-D-7-IV on wide, low, slightly separate tubercles, rarely all on elongate tubercles, posterior dorsal sensilla in transverse row, arranged medially to laterally: D-4-IV, D-8-IV, D-7-IV; L-1-IV short to moderately elongate seta, in transverse row with other lateral setae, arranged L-2-IV, L-1-IV, L-3-IV, L-4-IV; L-2-IV, L-3-IV, L-4-IV short setae on low, shelflike tubercles or (rarely) on elongate tubercles, V-6-IV, V-7-IV short setae on wide, low tubercles or (rarely) on elongate tubercles, V-5-IV absent; segment 8 without D-3-VIII, without L-1-VIII; segment 9 (
<figureCitation id="12E72A5ABD02305AFAF81D4048E8E56A" box="[1280,1432,1877,1902]" captionStart="FIGURE 72" captionStartId="317.[151,250,1765,1787]" captionTargetBox="[228,1373,193,1750]" captionTargetId="figure-192@317.[227,1378,193,1750]" captionTargetPageId="317" captionText="FIGURE 72. Abdominal segments 9. A. Austroconops mcmillani, male, in dorsal and ventral view (from Borkent &amp; Craig 2004). B. Leptoconops whitseli, male, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). C. Leptoconops whitseli, female, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). D. Leptoconops asilomar, male, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). E. Leptoconops asilomar, female, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). F. Forcipomyia anabaenae, female, in dorsal view (from Chan &amp; Saunders 1965). G. Forcipomyia eques, male, in ventral view (from Chan &amp; LeRoux 1965). H. Forcipomyia rugosa, female, in dorsal view (from Chan &amp; LeRoux 1970). I. Forcipomyia brevis, female, in dorsal view (from Mayer 1934a). J. Atrichopogon wirthi, male, in dorsal view, arrow indicating anteriorly directed spines (from Chan &amp; Linley 1988). K. Dasyhelea contigua, female, in ventral view (from Mayer 1934c). L. Dasyhelea grata, male, in ventral view (from Mayer 1934c)." figureDoi="http://doi.org/10.5281/zenodo.4949308" httpUri="https://zenodo.org/record/4949308/files/figure.png" pageId="44" pageNumber="45">Figs. 72K–L</figureCitation>
,
<figureCitation id="12E72A5ABD02305AFF6F1D6C4D99E596" box="[151,233,1913,1938]" captionStart="FIGURE 73" captionStartId="318.[151,250,1765,1787]" captionTargetBox="[213,1371,197,1757]" captionTargetId="figure-189@318.[211,1371,188,1757]" captionTargetPageId="318" captionText="FIGURE 73. Abdominal segments 9. A. Dasyhelea laeta, female, in ventral view (from Mayer 1934c). B. Dasyhelea assimilis, male, in lateral view with ventral to the right (from Mayer 1934c). C. Dasyhelea assimilis, male, in lateral view with ventral to the right (from Mayer 1934c). D. Dasyhelea grisea, male, in dorsal view (from Waugh &amp; Wirth 1976). E. Dasyhelea chani, female, in dorsal view (from Wirth &amp; Linley 1990). F. Dasyhelea atlantis, female, in ventral view (from Waugh &amp; Wirth 1976). G. Dasyhelea gluchovae, female, in ventral view (from Brodskaya 1996). H. Culicoides haematopotus, female, in dorsal and ventral view (shagreen not shown). I. Paradasyhelea albipunctata, female, in dorsal view (Elson-Harris &amp; Kettle 1985a). J. Ceratopogon boomerangus, female, in ventral view (modified from Borkent &amp; Grogan 1995). K. Brachypogon artemis, female, in dorsal view (from Debenham 1991). L. Alluaudomyia appendiculata, female, in dorsal view (from Elson-Harris &amp; Kettle 1985b)." figureDoi="http://doi.org/10.5281/zenodo.4949314" httpUri="https://zenodo.org/record/4949314/files/figure.png" pageId="44" pageNumber="45">73A–G</figureCitation>
)—with D-5-IX, D-6-IX campaniform sensilla, V-1-IX, V-2-IX setae, in some one seta or one campaniform sensillum likely absent.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD02305BFF3F1DD4483CE3D4" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="distribution">
<paragraph id="8A6336DFBD02305BFF3F1DD4483CE3D4" blockId="44.[151,1437,329,2010]" lastBlockId="45.[151,1437,151,2013]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis id="B8A8EACDBD02305AFF3F1DD44F13E5DE" bold="true" box="[199,611,1985,2010]" pageId="44" pageNumber="45">DISTRIBUTION AND HABITAT</emphasis>
: The genus
<taxonomicName id="4DDC4D5CBD02305AFD011DD44E1FE5DE" box="[761,879,1985,2010]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD02305AFD011DD44E1FE5DE" box="[761,879,1985,2010]" italics="true" pageId="44" pageNumber="45">Dasyhelea</emphasis>
</taxonomicName>
is known from 609 species from every Region worldwide (
<bibRefCitation id="EE4D4B2EBD03305BFEE71A824CCFE2B4" author="Borkent" box="[287,447,151,176]" firstAuthor="Borkent" pageId="45" pageNumber="46" refId="ref177229" refString="Borkent, A. (2014) World Species of Biting Midges (Diptera: Ceratopogonidae). Available from: http: // www. inhs. illinois. edu / research / FLYTREE / Borkent. html (accessed 20 May 2014)" type="url" year="2014">Borkent 2014</bibRefCitation>
, Díaz, et al. 2014). Immatures are known from a wide array of small aquatic habitats, primarily from algal growth on damp soil or other wet substrates, wet moss, algae or moss at the margins of standing or running water and amongst floating vegetation (including algal mats). As one would expect with a species rich genus, immatures occur in a wide array of other microhabitats but all of these are small aquatic (or soaking wet) habitats such as from
<taxonomicName id="4DDC4D5CBD03305BFDD01B3D4F19E33B" box="[552,617,296,319]" class="Liliopsida" family="Araceae" genus="Pistia" kingdom="Plantae" order="Alismatales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="genus">
<emphasis id="B8A8EACDBD03305BFDD01B3D4F19E33B" box="[552,617,296,319]" italics="true" pageId="45" pageNumber="46">Pistia</emphasis>
</taxonomicName>
, wet decomposing vegetation, ant refuse piles (of
<taxonomicName id="4DDC4D5CBD03305BFB5E1B3D49A5E33B" box="[1190,1237,296,319]" class="Insecta" family="Formicidae" genus="Atta" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD03305BFB5E1B3D49A5E33B" box="[1190,1237,296,319]" italics="true" pageId="45" pageNumber="46">Atta</emphasis>
</taxonomicName>
), mining floating leaves of
<taxonomicName id="4DDC4D5CBD03305BFEF11B594C15E361" box="[265,357,332,357]" class="Polypodiopsida" family="Salviniaceae" genus="Salvinia" kingdom="Plantae" order="Salviniales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="genus">
<emphasis id="B8A8EACDBD03305BFEF11B594C15E361" box="[265,357,332,357]" italics="true" pageId="45" pageNumber="46">Salvinia</emphasis>
</taxonomicName>
, sap from tree wounds, wet woodland leaf packs, small ground or rock pools, phytotelmata, margins of extreme saline pools, hot springs, intertidal zone, marine splash-zone, salt marshes, small artificial habitats such as rain gutters, and attacking tobacco seedlings grown hydroponically. Larvae appear to require water on their bodies to survive, while pupae either remain immersed in media or crawl upward to drier substrate.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD033058FF3F1BC94C6DE2D1" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="discussion">
<paragraph id="8A6336DFBD03305BFF3F1BC94952E0A3" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
<bibRefCitation id="EE4D4B2EBD03305BFF3F1BC94CEDE3F1" author="Munchberg" box="[199,413,476,501]" firstAuthor="Munchberg" pageId="45" pageNumber="46" pagination="24 - 40" refId="ref191303" refString="Munchberg, P. (1961) Zur Bedeutung der Prothorakalhorner der Puppen der Heleidae. Deutsche Entomologische Zeitschrift, 8, 24 - 40." type="journal article" year="1961">Münchberg (1961)</bibRefCitation>
studied the function of the respiratory organs of
<taxonomicName id="4DDC4D5CBD03305BFC1F1BC84908E3F1" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1833" box="[999,1144,476,501]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavifrons">
<emphasis id="B8A8EACDBD03305BFC1F1BC84908E3F1" box="[999,1144,476,501]" italics="true" pageId="45" pageNumber="46">D. flavifrons</emphasis>
</taxonomicName>
(as
<taxonomicName id="4DDC4D5CBD03305BFB571BC84837E3F1" baseAuthorityName="Winnertz" baseAuthorityYear="1852" box="[1199,1351,476,501]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B8A8EACDBD03305BFB571BC84837E3F1" box="[1199,1351,476,501]" italics="true" pageId="45" pageNumber="46">D. versicolor</emphasis>
</taxonomicName>
) under experimental conditions, removing either one or both of the respiratory organs or applying sealant to the pores. His questionable approach resulted in death for those with both respiratory organs removed and very little survivorship in those with one removed. He suggested that some respiration may occur through the terminal processes but this is clearly not possible (there is no opening posteriorly). Some young pupae could remain submerged for substantial periods of time indicating they likely extract oxygen from the surrounding water.
</paragraph>
<paragraph id="8A6336DFBD03305BFF3F18A14E43E13C" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
<emphasis id="B8A8EACDBD03305BFF3F18A14F5BE0C9" bold="true" box="[199,555,692,717]" pageId="45" pageNumber="46">TAXONOMIC DISCUSSION</emphasis>
: There are 90 species of
<taxonomicName id="4DDC4D5CBD03305BFCB918A14EC7E0C9" box="[833,951,692,717]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD03305BFCB918A14EC7E0C9" box="[833,951,692,717]" italics="true" pageId="45" pageNumber="46">Dasyhelea</emphasis>
</taxonomicName>
known as pupae (
<tableCitation id="C75E0364BD03305BFB8718A1498AE0C9" box="[1151,1274,692,717]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStartId-0="174.[147,245,486,512]" captionStartId-1="242.[151,239,152,175]" captionTargetBox-0="[150,1771,751,1679]" captionTargetBox-1="[159,1428,341,1978]" captionTargetPageId-0="174" captionTargetPageId-1="242" captionText-0="TABLE 2. References to species of Ceratopogonidae described as eggs, larvae and/or pupae. Species studied as pupae in the current study have their name highlighted in bold font. Stages studied are referred to as 'E' for egg, 'L' for larva and 'P' for pupa. Taxa are arrangement phyletically by subfamily and tribe (some of which are paraphyletic) and within tribes genera are listed alphabetically. Subfamilies are abbreviated as follows: Lept.— Leptoconopinae, Forc.—Forcipomyiinae, Dasy.—Dasyheleinae, Cera.—Ceratopogoninae. References highlighted in bold font are those with SEM photomicrographs. Theses are cited only when their results have not been published. Duplicated figures or repeated descriptions by subsequent authors are not listed here. Fossil taxa are identified with a cross (†). The species name in bold indicates only that named material of the pupa was studied, not that the cited author's specimens were reexamined (although in many instances they were). The larva of Jenkinshelea albaria was tentatively associated by Wirth &amp; Grogan 1979 (not reared); see discussion by Knausenberger, 1987:248." captionText-1="TABLE 3. Number of named extant species with described immatures for genera of Ceratopogonidae (taxa of uncertain identity or identified only as &quot;sp.&quot; are not included). Only genera with at least one immature stage known are listed. Percentages of described species within that genus are those for which those stages are known. Number of named species from Borkent (2014), with a few additions as noted in the Taxonomic Discussion under some genera. Taxa are arranged phyletically by subfamily and tribe and within tribes genera are listed alphabetically." httpUri-0="http://table.plazi.org/id/DEA36657BD8030D8FF6B1BF34923E0EC" httpUri-1="http://table.plazi.org/id/DEA36657BDDC3084FF6F1A8D496DE336" pageId="45" pageNumber="46" tableUuid-0="DEA36657BD8030D8FF6B1BF34923E0EC" tableUuid-1="DEA36657BDDC3084FF6F1A8D496DE336">Tables 2–3</tableCitation>
) and there is a high level of morphological differentiation between many of them. Clearly, there are numerous synapomorphies waiting to be interpreted. Furthermore, this genus is extremely species rich, with many undescribed species, and pupal morphology will certainly assist in interpreting these.
</paragraph>
<paragraph id="8A6336DFBD03305BFF3F19514C07E65C" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
Even though there are a goodly number of named species with described pupae, it is clear that the identity of some of them is uncertain. For example,
<bibRefCitation id="EE4D4B2EBD03305BFD9119724E38E184" author="Thienemann" box="[617,840,871,896]" firstAuthor="Thienemann" pageId="45" pageNumber="46" pagination="102 - 126" refId="ref197494" refString="Thienemann, A. (1925) Dipteren aus den Salzgewassern von Oldesloe. Mitteilungen der Geographischen Gesellschaft und des Naturhistorischen Museums in Lubeck, Series 2, 31, 102 - 126." type="journal article" year="1925">Thienemann (1925)</bibRefCitation>
recognized the pupae of
<taxonomicName id="4DDC4D5CBD03305BFB90197C4986E184" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1833" box="[1128,1270,871,896]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavifrons">
<emphasis id="B8A8EACDBD03305BFB90197C4986E184" box="[1128,1270,871,896]" italics="true" pageId="45" pageNumber="46">D. flavifrons</emphasis>
</taxonomicName>
and five other species (
<taxonomicName id="4DDC4D5CBD03305BFF0019984CD0E1A1" authorityName="Goetghebuer" authorityYear="1919" box="[248,416,908,933]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="brevitibialis">
<emphasis id="B8A8EACDBD03305BFF0019984CD0E1A1" box="[248,416,908,933]" italics="true" pageId="45" pageNumber="46">D. brevitibialis</emphasis>
</taxonomicName>
,
<taxonomicName id="4DDC4D5CBD03305BFE5519984F43E1A1" authorityName="Kieffer" authorityYear="1919" box="[429,563,908,933]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="lignicola">
<emphasis id="B8A8EACDBD03305BFE5519984F43E1A1" box="[429,563,908,933]" italics="true" pageId="45" pageNumber="46">D. lignicola</emphasis>
</taxonomicName>
,
<taxonomicName id="4DDC4D5CBD03305BFDB919984FCDE1A1" box="[577,701,908,933]" class="Insecta" family="Ceratopogonidae" genus="Schizonyxhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B8A8EACDBD03305BFDB919984FCDE1A1" box="[577,701,908,933]" italics="true" pageId="45" pageNumber="46">D. obscura</emphasis>
</taxonomicName>
,
<taxonomicName id="4DDC4D5CBD03305BFD3319984E22E1A1" authorityName="Kieffer" authorityYear="1919" box="[715,850,908,933]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="sensualis">
<emphasis id="B8A8EACDBD03305BFD3319984E22E1A1" box="[715,850,908,933]" italics="true" pageId="45" pageNumber="46">D. sensualis</emphasis>
</taxonomicName>
,
<taxonomicName id="4DDC4D5CBD03305BFC9919984E85E1A1" baseAuthorityName="Winnertz" baseAuthorityYear="1852" box="[865,1013,908,933]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B8A8EACDBD03305BFC9919984E85E1A1" box="[865,1013,908,933]" italics="true" pageId="45" pageNumber="46">D. versicolor</emphasis>
</taxonomicName>
) now all considered to be one species (the latter as synonyms of
<taxonomicName id="4DDC4D5CBD03305BFE2419A44F1EE1CC" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1833" box="[476,622,943,968]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavifrons">
<emphasis id="B8A8EACDBD03305BFE2419A44F1EE1CC" box="[476,622,943,968]" italics="true" pageId="45" pageNumber="46">D. flavifrons</emphasis>
</taxonomicName>
). Similarly,
<bibRefCitation id="EE4D4B2EBD03305BFCFD19A54EE4E1CC" author="Lenz" box="[773,916,943,968]" firstAuthor="Lenz" pageId="45" pageNumber="46" refId="ref189178" refString="Lenz, F. (1934) B. Die Metamorphose der Heleidae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. Vol. 3. Stuttgart, pp. 95 - 128." type="book" year="1934">Lenz (1934)</bibRefCitation>
noted differences between the pupae of
<taxonomicName id="4DDC4D5CBD03305BFA7919A44C71E1E9" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1833" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavifrons">
<emphasis id="B8A8EACDBD03305BFA7919A44C71E1E9" italics="true" pageId="45" pageNumber="46">D. flavifrons</emphasis>
</taxonomicName>
and some of these same taxa. Some of the differences noted are likely valid indicators of species differences, although it is now uncertain what adults were actually reared and associated.
<bibRefCitation id="EE4D4B2EBD03305BFB4119ED4814E614" author="Mayer" box="[1209,1380,1015,1040]" firstAuthor="Mayer" pageId="45" pageNumber="46" pagination="205 - 288" refId="ref190389" refString="Mayer, K. (1934 a) Die Metamorphose der Ceratopogonidae (Dipt.). Ein Beitrag zur Morphologie, Systematik. Okologie und Biologie der Jugendstadien dieser Dipterenfamilie. Archiv fur Naturgeschicthe, 3, 205 - 288." type="journal article" year="1934" yearSuffix="a">Mayer (1934a)</bibRefCitation>
also describes differences between species now considered synonyms (e.g.
<taxonomicName id="4DDC4D5CBD03305BFC391E084933E631" baseAuthorityName="Winnertz" baseAuthorityYear="1852" box="[961,1091,1052,1077]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B8A8EACDBD03305BFC391E084933E631" box="[961,1091,1052,1077]" italics="true" pageId="45" pageNumber="46">D. modesta</emphasis>
</taxonomicName>
with its synonyms
<taxonomicName id="4DDC4D5CBD03305BFADE1E0848ECE631" authorityName="Kieffer" authorityYear="1918" box="[1318,1436,1052,1077]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="inclusa">
<emphasis id="B8A8EACDBD03305BFADE1E0848ECE631" box="[1318,1436,1052,1077]" italics="true" pageId="45" pageNumber="46">D. inclusa</emphasis>
</taxonomicName>
and
<taxonomicName id="4DDC4D5CBD03305BFF3E1E544C16E65C" authorityName="Kieffer" authorityYear="1913" box="[198,358,1087,1112]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="B8A8EACDBD03305BFF3E1E544C16E65C" box="[198,358,1087,1112]" italics="true" pageId="45" pageNumber="46">D. longipalpis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8A6336DFBD03305BFF3F1E714C31E7C4" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
<bibRefCitation id="EE4D4B2EBD03305BFF3F1E714C06E679" author="Mayer" box="[199,374,1124,1149]" firstAuthor="Mayer" pageId="45" pageNumber="46" pagination="205 - 288" refId="ref190389" refString="Mayer, K. (1934 a) Die Metamorphose der Ceratopogonidae (Dipt.). Ein Beitrag zur Morphologie, Systematik. Okologie und Biologie der Jugendstadien dieser Dipterenfamilie. Archiv fur Naturgeschicthe, 3, 205 - 288." type="journal article" year="1934" yearSuffix="a">Mayer (1934a)</bibRefCitation>
and
<bibRefCitation id="EE4D4B2EBD03305BFE441E714F3DE679" author="Lenz" box="[444,589,1124,1149]" firstAuthor="Lenz" pageId="45" pageNumber="46" refId="ref189178" refString="Lenz, F. (1934) B. Die Metamorphose der Heleidae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. Vol. 3. Stuttgart, pp. 95 - 128." type="book" year="1934">Lenz (1934)</bibRefCitation>
provided a key to the 19 European species known at that time and
<bibRefCitation id="EE4D4B2EBD03305BFF6F1E924C07E6A4" author="Thienemann" box="[151,375,1159,1184]" firstAuthor="Thienemann" pageId="45" pageNumber="46" pagination="102 - 126" refId="ref197494" refString="Thienemann, A. (1925) Dipteren aus den Salzgewassern von Oldesloe. Mitteilungen der Geographischen Gesellschaft und des Naturhistorischen Museums in Lubeck, Series 2, 31, 102 - 126." type="journal article" year="1925">Thienemann (1925)</bibRefCitation>
has a key to some species from northern
<collectingCountry id="F2CB764FBD03305BFC9A1E924EBCE6A4" box="[866,972,1159,1184]" name="Germany" pageId="45" pageNumber="46">Germany</collectingCountry>
. The pupae of 11 Nearctic species were described and illustrated by
<bibRefCitation id="EE4D4B2EBD03305BFE2A1EB94FA8E6C1" author="Waugh &amp; Wirth" box="[466,728,1196,1221]" firstAuthor="Waugh" pageId="45" pageNumber="46" pagination="219 - 247" refId="ref198494" refString="Waugh, W. T. &amp; Wirth, W. W. (1976) A revision of the genus Dasyhelea Kieffer of the eastern United States north of Florida (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 69, 219 - 247." type="journal article" year="1976">Waugh &amp; Wirth (1976)</bibRefCitation>
, showing significant differences between them but they did not provide a key. The campaniform sensilla DA-2-H is present in these species but were not illustrated.
<bibRefCitation id="EE4D4B2EBD03305BFAAB1EC54D9EE709" author="Mayer" firstAuthor="Mayer" pageId="45" pageNumber="46" pagination="166 - 202" refId="ref190459" refString="Mayer, K. (1934 c) Ceratopogoniden-Metamorphosen (C. Intermediae und C. Vermiformes) der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement 13, 166 - 202." type="journal article" year="1934" yearSuffix="c">Mayer (1934c)</bibRefCitation>
described and keyed 13 species from
<collectingCountry id="F2CB764FBD03305BFD571EE14E6CE709" box="[687,796,1268,1293]" name="Indonesia" pageId="45" pageNumber="46">Indonesia</collectingCountry>
indicating and illustrating detailed differences between their dorsal apotomes, respiratory organs, and the shape and chaetotaxy of segment 9.
<bibRefCitation id="EE4D4B2EBD03305BFB941F0D4864E734" author="Mayer" box="[1132,1300,1303,1328]" firstAuthor="Mayer" pageId="45" pageNumber="46" pagination="166 - 202" refId="ref190459" refString="Mayer, K. (1934 c) Ceratopogoniden-Metamorphosen (C. Intermediae und C. Vermiformes) der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement 13, 166 - 202." type="journal article" year="1934" yearSuffix="c">Mayer (1934c)</bibRefCitation>
showed the dorsal apotome of
<taxonomicName id="4DDC4D5CBD03305BFE941F284CA7E750" authorityName="Mayer" authorityYear="1934" box="[364,471,1341,1364]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="insons">
<emphasis id="B8A8EACDBD03305BFE941F284CA7E750" box="[364,471,1341,1364]" italics="true" pageId="45" pageNumber="46">D. insons</emphasis>
</taxonomicName>
with DA-2-H as a seta (so there are two setae on a single tubercle), which would be unique within the genus. All other species I studied had one seta and one campaniform sensillum.
<bibRefCitation id="EE4D4B2EBD03305BFB101F4A4D91E799" author="Wirth &amp; Beaver" firstAuthor="Wirth" pageId="45" pageNumber="46" pagination="41 - 52" refId="ref199490" refString="Wirth, W. W. &amp; Beaver, R. A. (1979) The Dasyhelea biting midges living in pitchers of Nepenthes in southeast Asia (Diptera, Ceratopogonidae). Annales de la Societe Entomologique de France, 15, 41 - 52." type="journal article" year="1979">Wirth &amp; Beaver (1979)</bibRefCitation>
described and keyed the unusual pupae of five species of
<taxonomicName id="4DDC4D5CBD03305BFC841F914E82E799" box="[892,1010,1412,1437]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD03305BFC841F914E82E799" box="[892,1010,1412,1437]" italics="true" pageId="45" pageNumber="46">Dasyhelea</emphasis>
</taxonomicName>
found in
<taxonomicName id="4DDC4D5CBD03305BFB9B1F9149A8E799" box="[1123,1240,1412,1437]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" kingdom="Plantae" order="Caryophyllales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="genus">
<emphasis id="B8A8EACDBD03305BFB9B1F9149A8E799" box="[1123,1240,1412,1437]" italics="true" pageId="45" pageNumber="46">Nepenthes</emphasis>
</taxonomicName>
(monkey cup) in southeast Asia.
</paragraph>
<paragraph id="8A6336DFBD03305BFF3F1FD94FADE470" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
Fossil pupae of six species were described (or redescribed) by
<bibRefCitation id="EE4D4B2EBD03305BFC791FD9496CE7E1" author="Pierce" box="[897,1052,1484,1509]" firstAuthor="Pierce" pageId="45" pageNumber="46" pagination="81 - 98" refId="ref192715" refString="Pierce, W. D. (1966) Fossil arthropods of California. 29. Silicified Miocene pupae of ceratopogonid flies. Bulletin of the Southern California Academy of Sciences, 65 (2), 81 - 98." type="journal article" year="1966">Pierce (1966)</bibRefCitation>
. Although the specimens were not reexamined here, based on the form of the terminal processes (divided), they are clearly correctly assigned to genus.
<bibRefCitation id="EE4D4B2EBD03305BFF131C014F49E429" author="Grimaldi &amp; Engel" box="[235,569,1556,1581]" firstAuthor="Grimaldi" page="45" pageId="45" pageNumber="46" refId="ref184374" refString="Grimaldi, D. &amp; Engel, M. S. (2005) Evolution of the insects. Cambridge University Press, New York, xv + 755 pp." type="book" year="2005">Grimaldi &amp; Engel (2005:45)</bibRefCitation>
include a photograph of two silicified pupae identified as 'ceratopogonid midges' and which are likely members of
<taxonomicName id="4DDC4D5CBD03305BFD8A1C224F98E454" box="[626,744,1591,1616]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD03305BFD8A1C224F98E454" box="[626,744,1591,1616]" italics="true" pageId="45" pageNumber="46">Dasyhelea</emphasis>
</taxonomicName>
, based on their size and the presence of distinctive shelf-like tubercles nearly encircling the abdominal segments.
</paragraph>
<paragraph id="8A6336DFBD03305BFF3F1C6A4FC9E501" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
Scanning electron photomicrographs of pupae were published by
<bibRefCitation id="EE4D4B2EBD03305BFC5C1C6A49D0E49C" author="Ronderos, M. M. &amp; Spinelli, G. R. &amp; Diaz, F." box="[932,1184,1663,1688]" pageId="45" pageNumber="46" pagination="45 - 54" refId="ref194018" refString="Ronderos, M. M., Spinelli, G. R. &amp; Diaz, F. (2004) Description of larva and redescription of pupa and adult of Palpomyia guarani (Diptera: Ceratopogonidae). Revista de la Sociedad Entomologica Argentina, 63, 45 - 54." type="journal article" year="2004">Ronderos et al. (2004)</bibRefCitation>
,
<bibRefCitation id="EE4D4B2EBD03305BFB541C6A481DE49C" author="Dominiak" box="[1196,1389,1663,1688]" firstAuthor="Dominiak" pageId="45" pageNumber="46" pagination="211 - 304" refId="ref181487" refString="Dominiak, P. (2012) Biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae) in Poland. Polish Journal of Entomology, 81, 211 - 304." type="journal article" year="2012">Dominiak (2012)</bibRefCitation>
and
<bibRefCitation id="EE4D4B2EBD03305BFF6F1CB14C1DE4B9" author="Diaz" box="[151,365,1700,1725]" etAl="et al." firstAuthor="Diaz" pageId="45" pageNumber="46" pagination="85 - 93" refId="ref181334" refString="Diaz, F., Ronderos, M. M., Spinelli, G. R., Ferreira-Keppler, R. L. &amp; Torreias, S. R. S. (2013) A new species of Dasyhelea Kieffer (Diptera: Ceratopogonidae) from Brazilian Amazonia. Zootaxa, 3686 (1), 85 - 93. http: // dx. doi. org / 10.11646 / zootaxa. 3686.1.5" type="journal article" year="2013">Díaz et al. (2013)</bibRefCitation>
.
<bibRefCitation id="EE4D4B2EBD03305BFE851CB14FEEE4B9" author="Park &amp; Downing" box="[381,670,1700,1725]" firstAuthor="Park" pageId="45" pageNumber="46" pagination="175 - 184" refId="ref192289" refString="Park, L. E. &amp; Downing, K. F. (2001) Paleoecology of an exceptionally preserved arthropod fauna from lake deposits of the Miocene Barstow Formation, southern California, U. S. A. Palaios, 16, 175 - 184." type="journal article" year="2001">Park &amp; Downing (2001)</bibRefCitation>
published an scanning electron photomicrograph of the terminal abdominal segments of the fossil Miocene species
<taxonomicName id="4DDC4D5CBD03305BFD2E1CDC4E2BE4E4" box="[726,859,1735,1760]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B8A8EACDBD03305BFD2E1CDC4E2BE4E4" box="[726,859,1735,1760]" italics="true" pageId="45" pageNumber="46">D. australis</emphasis>
</taxonomicName>
, showing how a similar approach could be used to better understand other fossil
<taxonomicName id="4DDC4D5CBD03305BFE1C1CF94F2AE501" box="[484,602,1772,1797]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B8A8EACDBD03305BFE1C1CF94F2AE501" box="[484,602,1772,1797]" italics="true" pageId="45" pageNumber="46">Dasyhelea</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="8A6336DFBD03305BFF3F1D0549ACE549" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
Pupae of
<taxonomicName id="4DDC4D5CBD03305BFECA1D1A4CD8E52C" box="[306,424,1807,1832]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD03305BFECA1D1A4CD8E52C" box="[306,424,1807,1832]" italics="true" pageId="45" pageNumber="46">Dasyhelea</emphasis>
</taxonomicName>
have a reduced number of thoracic sensilla and the naming of the two setae as D-1-T and D-2-T is arbitrary as they may not be homologous to those specific setae in other ceratopogonids.
</paragraph>
<paragraph id="8A6336DFBD033058FF3F1D4D4C6DE2D1" blockId="45.[151,1437,151,2013]" lastBlockId="46.[151,1437,151,2013]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
Most species of
<taxonomicName id="4DDC4D5CBD03305BFE781D424C86E574" box="[384,502,1879,1904]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD03305BFE781D424C86E574" box="[384,502,1879,1904]" italics="true" pageId="45" pageNumber="46">Dasyhelea</emphasis>
</taxonomicName>
have abdominal segments with a typical and characteristic transverse arrangement of lower, broad tubercles (
<figureCitation id="12E72A5ABD03305BFE3A1D694F5AE591" box="[450,554,1916,1941]" captionStart="FIGURE 56" captionStartId="301.[151,250,1953,1975]" captionTargetBox="[221,1382,193,1855]" captionTargetId="figure-49@301.[213,1395,188,1927]" captionTargetPageId="301" captionText="FIGURE 56. Abdominal segment four, in dorsal and ventral view (shagreen not shown). A. Forcipomyia dominicana. B. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). C. Dasyhelea paracincta." figureDoi="http://doi.org/10.5281/zenodo.4949250" httpUri="https://zenodo.org/record/4949250/files/figure.png" pageId="45" pageNumber="46">Fig. 56C</figureCitation>
). A few species found in
<taxonomicName id="4DDC4D5CBD03305BFCB61D694EB3E591" box="[846,963,1916,1941]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" kingdom="Plantae" order="Caryophyllales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="genus">
<emphasis id="B8A8EACDBD03305BFCB61D694EB3E591" box="[846,963,1916,1941]" italics="true" pageId="45" pageNumber="46">Nepenthes</emphasis>
</taxonomicName>
in southeast Asia, however, have sensilla on somewhat to markedly elongate tubercles (
<bibRefCitation id="EE4D4B2EBD03305BFD4E1DB54ECDE5BD" author="Wirth &amp; Beaver" box="[694,957,1952,1977]" firstAuthor="Wirth" pageId="45" pageNumber="46" pagination="41 - 52" refId="ref199490" refString="Wirth, W. W. &amp; Beaver, R. A. (1979) The Dasyhelea biting midges living in pitchers of Nepenthes in southeast Asia (Diptera, Ceratopogonidae). Annales de la Societe Entomologique de France, 15, 41 - 52." type="journal article" year="1979">Wirth &amp; Beaver 1979</bibRefCitation>
). An undescribed species from
<collectingCountry id="F2CB764FBD03305BFACD1DB548ECE5BD" box="[1333,1436,1952,1977]" name="Australia" pageId="45" pageNumber="46">Australia</collectingCountry>
(Running Creek) also had abdominal sensilla D-4 IV, D-8 IV, L-2 IV, L-3 IV, L-4 IV, V-7 IV on markedly elongate tubercles. D-4-IV in these taxa was represented by a seta (rather than a campaniform sensillum present in other
<taxonomicName id="4DDC4D5CBD003058FF6F1AA94C7DE2D1" box="[151,269,188,213]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD003058FF6F1AA94C7DE2D1" box="[151,269,188,213]" italics="true" pageId="46" pageNumber="47">Dasyhelea</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD003046FF3F1ACA4E15E1CC" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8A6336DFBD003046FF3F1ACA4E15E1CC" blockId="46.[151,1437,151,2013]" lastBlockId="48.[151,1436,151,968]" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
<emphasis id="B8A8EACDBD003058FF3F1ACA4F71E2FC" bold="true" box="[199,513,223,248]" pageId="46" pageNumber="47">MATERIAL EXAMINED</emphasis>
:
<taxonomicName id="4DDC4D5CBD003058FDE91AF44FCDE2FC" authorityName="Macfie" authorityYear="1934" box="[529,701,223,248]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ampullariae">
<emphasis id="B8A8EACDBD003058FDE91AF44FCDE2FC" box="[529,701,223,248]" italics="true" pageId="46" pageNumber="47">D. ampullariae</emphasis>
</taxonomicName>
: 2 pupal exuviae,
<collectingCountry id="F2CB764FBD003058FC751ACA4972E2FC" box="[909,1026,223,248]" name="Singapore" pageId="46" pageNumber="47">Singapore</collectingCountry>
,
<date id="FE62101FBD003058FBF51ACA49D3E2FC" box="[1037,1187,223,248]" pageId="46" pageNumber="47" value="1952-08-22">22-VIII-1952</date>
(USNM).
<taxonomicName id="4DDC4D5CBD003058FAE51AF448E4E2FC" authorityName="Wirth &amp; Williams" authorityYear="1957" box="[1309,1428,223,248]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B8A8EACDBD003058FAE51AF448E4E2FC" box="[1309,1428,223,248]" italics="true" pageId="46" pageNumber="47">D. atlantis</emphasis>
</taxonomicName>
: 1 pupal exuviae, Oak Island, Long Island,
<collectingRegion id="4818F83DBD003058FD851B114F84E319" box="[637,756,260,285]" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FCF81B114E4BE319" box="[768,827,260,285]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FCBF1B114ECFE319" box="[839,959,260,285]" pageId="46" pageNumber="47" value="1956-05-15">15-V-1956</date>
(NYSM); 1 pupal exuviae, Orient Beach, Long Island,
<collectingRegion id="4818F83DBD003058FED21B3D4CEFE344" box="[298,415,295,320]" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FE521B324C95E344" box="[426,485,295,320]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FE171B324F17E344" box="[495,615,295,320]" pageId="46" pageNumber="47" value="1963-05-25">25-V-1963</date>
(USNM); 1 pupal exuviae,
<quantity id="4D249B3ABD003058FC661B324EA6E344" box="[926,982,295,320]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="km" value="2.0">2 km</quantity>
E of
<collectingCountry id="F2CB764FBD003058FBEA1B32494AE344" box="[1042,1082,295,320]" name="Iceland" pageId="46" pageNumber="47">Isla</collectingCountry>
Santa Cruz,
<collectingRegion id="4818F83DBD003058FB311B3248E8E344" box="[1225,1432,295,320]" country="Ecuador" name="Galapagos" pageId="46" pageNumber="47">Galapagos Islands</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FF6F1B594D86E361" box="[151,246,332,357]" name="Ecuador" pageId="46" pageNumber="47">Ecuador</collectingCountry>
,
<date id="FE62101FBD003058FEF91B594C01E361" box="[257,369,332,357]" pageId="46" pageNumber="47" value="1989-01-18">18-I-1989</date>
(CNCI); 1 pupal exuviae, Pto.
<collectingCountry id="F2CB764FBD003058FD371B594F87E361" box="[719,759,332,357]" name="Iceland" pageId="46" pageNumber="47">Isla</collectingCountry>
Isabela,
<collectingRegion id="4818F83DBD003058FCA51B59495CE361" box="[861,1068,332,357]" country="Ecuador" name="Galapagos" pageId="46" pageNumber="47">Galapagos Islands</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FBCF1B5949E6E361" box="[1079,1174,332,357]" name="Ecuador" pageId="46" pageNumber="47">Ecuador</collectingCountry>
,
<date id="FE62101FBD003058FB591B594866E361" box="[1185,1302,332,357]" pageId="46" pageNumber="47" value="1989-03-06">6-III-1989</date>
(CNCI).
<taxonomicName id="4DDC4D5CBD003058FA791B584D8BE38C" authorityName="Goetghebuer" authorityYear="1920" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="bilineata">
<emphasis id="B8A8EACDBD003058FA791B584D8BE38C" italics="true" pageId="46" pageNumber="47">D. bilineata</emphasis>
</taxonomicName>
:
<specimenCount id="9CDAFD56BD003058FEF31B7A4C18E383" box="[267,360,367,392]" pageId="46" pageNumber="47" type="pupa">3 pupae</specimenCount>
, Harz,
<collectingCountry id="F2CB764FBD003058FE431B7A4F54E38C" box="[443,548,367,392]" name="Germany" pageId="46" pageNumber="47">Germany</collectingCountry>
,
<date id="FE62101FBD003058FDC91B7A4FD4E38C" box="[561,676,367,392]" pageId="46" pageNumber="47" value="1931-04-04">4-IV-1931</date>
(ZSMC); 3 pupal exuviae, (Lunz collection), 1941-1943 (ZSMC).
<taxonomicName id="4DDC4D5CBD003058FF6F1B804C53E3A9" authorityName="Wirth &amp; Beaver" authorityYear="1979" box="[151,291,404,429]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B8A8EACDBD003058FF6F1B804C53E3A9" box="[151,291,404,429]" italics="true" pageId="46" pageNumber="47">D. biseriata</emphasis>
</taxonomicName>
: 1 pupal exuviae (of
<typeStatus id="5567887DBD003058FDD51B814FE2E3A9" box="[557,658,404,429]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
),
<collectingCountry id="F2CB764FBD003058FD531B814E6FE3A9" box="[683,799,404,429]" name="Singapore" pageId="46" pageNumber="47">Singapore</collectingCountry>
,
<date id="FE62101FBD003058FCC81B814EC9E3A9" box="[816,953,404,429]" pageId="46" pageNumber="47" value="1952-08-06">6-VIII-1952</date>
(USNM).
<taxonomicName id="4DDC4D5CBD003058FBC51B8049B4E3A9" authorityName="Kieffer" authorityYear="1915" box="[1085,1220,404,429]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="bistriata">
<emphasis id="B8A8EACDBD003058FBC51B8049B4E3A9" box="[1085,1220,404,429]" italics="true" pageId="46" pageNumber="47">D. bistriata</emphasis>
</taxonomicName>
: 6 pupal exuviae, probably
<collectingRegion id="4818F83DBD003058FEF31BA24C0CE3D4" box="[267,380,439,464]" country="Germany" name="Thueringen" pageId="46" pageNumber="47">Thuringia</collectingRegion>
, from potash works at Heringen on Werra River,
<collectingCountry id="F2CB764FBD003058FC011BA24912E3D4" box="[1017,1122,439,464]" name="Germany" pageId="46" pageNumber="47">Germany</collectingCountry>
,
<date id="FE62101FBD003058FB8F1BA2498AE3D4" box="[1143,1274,439,464]" pageId="46" pageNumber="47" value="1914-03-23">23-III-1914</date>
(ZSMC).
<taxonomicName id="4DDC4D5CBD003058FA791BAC4C65E3F1" authorityName="Waugh &amp; Wirth" authorityYear="1976" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="B8A8EACDBD003058FA791BAC4C65E3F1" italics="true" pageId="46" pageNumber="47">D. brevicornis</emphasis>
</taxonomicName>
: 1 pupal exuviae (of
<typeStatus id="5567887DBD003058FDEE1BC94F0BE3F1" box="[534,635,476,501]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
), Salmon River, Blue Mountain Lake,
<collectingRegion id="4818F83DBD003058FBBA1BC949CBE3F1" box="[1090,1211,476,501]" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FB321BC94875E3F1" box="[1226,1285,476,501]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FAEB1BC948ECE3F1" box="[1299,1436,476,501]" pageId="46" pageNumber="47" value="1959-08-06">6-VIII-1959</date>
(USNM).
<taxonomicName id="4DDC4D5CBD003058FEF118144C19E01C" authorityName="Wirth &amp; Linley" authorityYear="1990" box="[265,361,511,536]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="chani">
<emphasis id="B8A8EACDBD003058FEF118144C19E01C" box="[265,361,511,536]" italics="true" pageId="46" pageNumber="47">D. chani</emphasis>
</taxonomicName>
: 1 pupal exuviae (of
<typeStatus id="5567887DBD003058FDA21BEA4FCFE01C" box="[602,703,511,536]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
), Chinese Farm, Ft. Pierce, Florida,
<collectingCountry id="F2CB764FBD003058FBAE1BEA49E1E01C" box="[1110,1169,511,536]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FB651BEA485AE01C" box="[1181,1322,511,536]" pageId="46" pageNumber="47" value="1987-07-24">24-VII-1987</date>
(USNM).
<taxonomicName id="4DDC4D5CBD003058FF6F18304C70E038" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[151,256,549,572]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="B8A8EACDBD003058FF6F18304C70E038" box="[151,256,549,572]" italics="true" pageId="46" pageNumber="47">D. cincta</emphasis>
</taxonomicName>
: 8 pupal exuviae, Poncha Hot Springs, Chaffee County
<collectingRegion id="4818F83DBD003058FC7718314E8AE039" box="[911,1018,548,573]" country="United States of America" name="Colorado" pageId="46" pageNumber="47">Colorado</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FBF018314933E039" box="[1032,1091,548,573]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FBA9183149A5E039" box="[1105,1237,548,573]" pageId="46" pageNumber="47" value="1991-09-21">21-IX-1991</date>
(CNCI); 1 pupal exuviae, Los Talas,
<collectingCountry id="F2CB764FBD003058FE8D18524C97E064" box="[373,487,583,608]" name="Argentina" pageId="46" pageNumber="47">Argentina</collectingCountry>
,
<date id="FE62101FBD003058FE0A18524F05E064" box="[498,629,583,608]" pageId="46" pageNumber="47" value="1981-09-25">25-IX-1981</date>
(MLPA).
<taxonomicName id="4DDC4D5CBD003058FD10185C4E1CE05B" baseAuthorityName="Johannsen" baseAuthorityYear="1931" box="[744,876,584,608]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B8A8EACDBD003058FD10185C4E1CE05B" box="[744,876,584,608]" italics="true" pageId="46" pageNumber="47">D. contigua</emphasis>
</taxonomicName>
: 1 pupal exuviae,
<collectingRegion id="4818F83DBD003058FBC0185249E8E064" box="[1080,1176,583,608]" country="Indonesia" name="Sumatera Utara" pageId="46" pageNumber="47">Sumatra</collectingRegion>
, Toba area, creek north of Balige, in moss of a waterfall,
<collectingCountry id="F2CB764FBD003058FDD618794FEEE081" box="[558,670,620,645]" name="Indonesia" pageId="46" pageNumber="47">Indonesia</collectingCountry>
,
<date id="FE62101FBD003058FD5618794E55E081" box="[686,805,620,645]" pageId="46" pageNumber="47" value="1929-04-01">1-IV-1929</date>
. (ZSMC).
<taxonomicName id="4DDC4D5CBD003058FC5D18784953E081" authorityName="Kieffer" authorityYear="1914" box="[933,1059,620,645]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="diplosis">
<emphasis id="B8A8EACDBD003058FC5D18784953E081" box="[933,1059,620,645]" italics="true" pageId="46" pageNumber="47">D. diplosis</emphasis>
</taxonomicName>
:
<specimenCount id="9CDAFD56BD003058FBCF187949E6E087" box="[1079,1174,620,645]" pageId="46" pageNumber="47" type="pupa">3 pupae</specimenCount>
, ditches at Sassendorf saltworks (E of Soest),
<collectingRegion id="4818F83DBD003058FE65189A4FC1E0AC" box="[413,689,655,680]" country="Germany" name="Nordrhein-Westfalen" pageId="46" pageNumber="47">North Rhine-Westphalia</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FD44189A4E55E0AC" box="[700,805,655,680]" name="Germany" pageId="46" pageNumber="47">Germany</collectingCountry>
, mid-April to late May, 1912 (ZSMC).
<taxonomicName id="4DDC4D5CBD003058FB271884484AE0AC" authorityName="Carter, Ingram &amp; Macfie" authorityYear="1921" box="[1247,1338,655,680]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="B8A8EACDBD003058FB271884484AE0AC" box="[1247,1338,655,680]" italics="true" pageId="46" pageNumber="47">D. flava</emphasis>
</taxonomicName>
: 1 pupal exuviae, Burgeshall, Hazyview, East Transvaal,
<collectingCountry id="F2CB764FBD003058FD2018A14E02E0C9" box="[728,882,692,717]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
,
<date id="FE62101FBD003058FC7D18A14975E0C9" box="[901,1029,692,717]" pageId="46" pageNumber="47" value="1973-12-03">3-XII-1973</date>
(NMSA).
<taxonomicName id="4DDC4D5CBD003058FB7318A0485BE0C9" authorityName="Macfie" authorityYear="1939" box="[1163,1323,692,717]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="flavicauda">
<emphasis id="B8A8EACDBD003058FB7318A0485BE0C9" box="[1163,1323,692,717]" italics="true" pageId="46" pageNumber="47">D. flavicauda</emphasis>
</taxonomicName>
: 1 pupal exuviae, Laguna “La Brava”,
<collectingRegion id="4818F83DBD003058FE1218C24F11E0F4" box="[490,609,727,752]" country="Argentina" name="Corrientes" pageId="46" pageNumber="47">Corrientes</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FD9618C24FADE0F4" box="[622,733,727,752]" name="Argentina" pageId="46" pageNumber="47">Argentina</collectingCountry>
(MLPA).
<taxonomicName id="4DDC4D5CBD003058FCAB18CC4E90E0F4" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1833" box="[851,992,727,752]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="flavifrons">
<emphasis id="B8A8EACDBD003058FCAB18CC4E90E0F4" box="[851,992,727,752]" italics="true" pageId="46" pageNumber="47">D. flavifrons</emphasis>
</taxonomicName>
: 2 pupal exuviae (in glycerin), locality uncertain (CNCI); 4 pupal exuviae, Dolina Radości, Gdańsk,
<collectingCountry id="F2CB764FBD003058FCBE18E94EE7E111" box="[838,919,764,789]" name="Poland" pageId="46" pageNumber="47">Poland</collectingCountry>
,
<date id="FE62101FBD003058FC5918E94965E111" box="[929,1045,764,789]" pageId="46" pageNumber="47" value="1989-04-03">3-IV-1989</date>
(
<collectionCode id="ECCDAE1ABD003058FBDB18E9491CE111" box="[1059,1132,764,789]" country="Italy" httpUri="http://grbio.org/cool/26db-tnj6" name="Istituto di Zoologia dell'Universita" pageId="46" pageNumber="47">IZUG</collectionCode>
); 2 pupal exuviae, Loiterau river valley nr Taarstedt,
<collectingRegion id="4818F83DBD003058FE39190A4F0FE13C" box="[449,639,799,824]" country="Germany" name="Schleswig-Holstein" pageId="46" pageNumber="47">Slesvig-Holstein</collectingRegion>
, NE of Schleswig,
<collectingCountry id="F2CB764FBD003058FC9D190A4EBEE13C" box="[869,974,799,824]" name="Germany" pageId="46" pageNumber="47">Germany</collectingCountry>
,
<date id="FE62101FBD003058FC26190A4910E13C" box="[990,1120,799,824]" pageId="46" pageNumber="47" value="1939-04-27">27-IV-1939</date>
(ZSMC); 4 pupal exuviae,
<collectingRegion id="4818F83DBD003058FF6F19514D81E159" box="[151,241,836,861]" country="Germany" name="Bayern" pageId="46" pageNumber="47">Bavaria</collectingRegion>
, Munich,
<collectingCountry id="F2CB764FBD003058FE9019514CA1E159" box="[360,465,836,861]" name="Germany" pageId="46" pageNumber="47">Germany</collectingCountry>
,
<date id="FE62101FBD003058FE1819514F38E159" box="[480,584,836,861]" pageId="46" pageNumber="47" value="1922-12">XII-1922</date>
(ZSMC); 1 pupal exuviae, Buda Park,
<collectingRegion id="4818F83DBD003058FBE0195149F3E159" box="[1048,1155,836,861]" country="Hungary" name="Budapest" pageId="46" pageNumber="47">Budapest</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FB6A19514985E158" box="[1170,1269,836,860]" name="Hungary" pageId="46" pageNumber="47">Hungary</collectingCountry>
, VIII-IX-1986 (CNCI).
<taxonomicName id="4DDC4D5CBD003058FEF8197C4C0EE184" authorityName="Meillon &amp; Wirth" authorityYear="1981" box="[256,382,871,896]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fontana">
<emphasis id="B8A8EACDBD003058FEF8197C4C0EE184" box="[256,382,871,896]" italics="true" pageId="46" pageNumber="47">D. fontana</emphasis>
</taxonomicName>
: 1 pupal exuviae, 1 pupal exuviae (of
<typeStatus id="5567887DBD003058FCA519724EB3E184" box="[861,963,871,896]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
),
<specimenCount id="9CDAFD56BD003058FC2419724952E17B" box="[988,1058,871,896]" pageId="46" pageNumber="47" type="generic">
<quantity id="4D249B3ABD003058FC241972496EE17B" box="[988,1054,871,896]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="47" unit="m" value="25.0">25 m</quantity>
.
</specimenCount>
E of Middelburg, Sewefontein, Transvaal,
<collectingCountry id="F2CB764FBD003058FEE119994CC2E1A1" box="[281,434,908,933]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
,
<date id="FE62101FBD003058FE3B19994F32E1A1" box="[451,578,908,933]" pageId="46" pageNumber="47" value="1973-12-05">5-XII-1973</date>
(NMSA).
<taxonomicName id="4DDC4D5CBD003058FD3F19984E5BE1A1" authorityName="Carter, Ingram &amp; Macfie" authorityYear="1921" box="[711,811,908,933]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="B8A8EACDBD003058FD3F19984E5BE1A1" box="[711,811,908,933]" italics="true" pageId="46" pageNumber="47">D. fusca</emphasis>
</taxonomicName>
: 1 pupal exuviae, Zoutpan, Transvaal,
<collectingCountry id="F2CB764FBD003058FAFE199948ECE1A1" box="[1286,1436,908,933]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
(NMSA); 1 pupal exuviae,
<quantity id="4D249B3ABD003058FE3F19BA4F76E1CC" box="[455,518,943,968]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="46" pageNumber="47" unit="mi" value="15.0">15 mi</quantity>
SE of Potgietersrus, Transvaal,
<collectingCountry id="F2CB764FBD003058FC9519BA4E8FE1CC" box="[877,1023,943,968]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
,
<date id="FE62101FBD003058FBF219BA491CE1CC" box="[1034,1132,943,968]" pageId="46" pageNumber="47" value="1974-01-03">3-I-1974</date>
(NMSA); 1 pupal exuviae, Lodwigslust, Hectorspruit, East Transvaal,
<collectingCountry id="F2CB764FBD003058FD8619C14E60E1E9" box="[638,784,980,1005]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
,
<date id="FE62101FBD003058FCE419C14ED0E1E9" box="[796,928,980,1005]" pageId="46" pageNumber="47" value="1973-11-29">29-XI-1973</date>
(NMSA); 2 pupal exuviae, Doornpan, Bulge River IV., Transvaal,
<collectingCountry id="F2CB764FBD003058FE7D19E24F67E614" box="[389,535,1015,1040]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
,
<date id="FE62101FBD003058FDDA19E24FE8E614" box="[546,664,1015,1040]" pageId="46" pageNumber="47" value="1973-11-06">6-XI-1973</date>
(NMSA); 1 pupal exuviae, Mamba River, Zululand,
<collectingCountry id="F2CB764FBD003058FB1019E2480BE614" box="[1256,1403,1015,1040]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
,
<date id="FE62101FBD003058FA7D19E24D91E631" pageId="46" pageNumber="47" value="1937-01-05">5- I-1937</date>
(SAIM).
<taxonomicName id="4DDC4D5CBD003058FEB61E084F66E631" authorityName="Meillon" authorityYear="1937" box="[334,534,1052,1077]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="gigantosalpinx">
<emphasis id="B8A8EACDBD003058FEB61E084F66E631" box="[334,534,1052,1077]" italics="true" pageId="46" pageNumber="47">D. gigantosalpinx</emphasis>
</taxonomicName>
: 1 pupal exuviae, Jam Tin Creek, Maklane, East Transvaal,
<collectingCountry id="F2CB764FBD003058FB4F1E09483AE631" box="[1207,1354,1052,1077]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
,
<date id="FE62101FBD003058FAAD1E094DBFE65C" pageId="46" pageNumber="47" value="1973-12-02">2-XII- 1973</date>
(NMSA); 1 pupal exuviae (of
<typeStatus id="5567887DBD003058FDBD1E2A4FDAE65C" box="[581,682,1087,1112]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
), Umlalazi River, Eshowe, Zululand,
<collectingCountry id="F2CB764FBD003058FB9F1E2A498EE65C" box="[1127,1278,1087,1112]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
,
<date id="FE62101FBD003058FAF71E2A48ECE65C" box="[1295,1436,1087,1112]" pageId="46" pageNumber="47" value="1936-12-19">19-XII-1936</date>
(SAIM).
<taxonomicName id="4DDC4D5CBD003058FF061E704C17E678" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[254,359,1125,1148]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="grisea">
<emphasis id="B8A8EACDBD003058FF061E704C17E678" box="[254,359,1125,1148]" italics="true" pageId="46" pageNumber="47">D. grisea</emphasis>
</taxonomicName>
: 4 pupal exuviae,
<collectingRegion id="4818F83DBD003058FDC11E714FDBE679" box="[569,683,1124,1149]" country="United States of America" name="California" pageId="46" pageNumber="47">California</collectingRegion>
(USNM); 1 pupal exuviae, Hamilton-Essex Line, Newcomb,
<collectingRegion id="4818F83DBD003058FA9F1E714DA1E6A4" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FF181E924C6BE6A4" box="[224,283,1159,1184]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FED21E924CD2E6A4" box="[298,418,1159,1184]" pageId="46" pageNumber="47" value="1960-05-24">24-V-1960</date>
(NYSM); 1 pupal exuviae, Grandview Park Natural Preserve, Hampton City,
<collectingRegion id="4818F83DBD003058FAC21E9248E8E6A4" box="[1338,1432,1159,1184]" country="United States of America" name="Virginia" pageId="46" pageNumber="47">Virginia</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FF6F1EB94DA2E6C1" box="[151,210,1196,1221]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FF1B1EB94C17E6C1" box="[227,359,1196,1221]" pageId="46" pageNumber="47" value="1976-11-27">27-XI-1976</date>
(VPIC).
<taxonomicName id="4DDC4D5CBD003058FE231EB84F1EE6C1" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1833" box="[475,622,1196,1221]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="flavifrons">
<emphasis id="B8A8EACDBD003058FE231EB84F1EE6C1" box="[475,622,1196,1221]" italics="true" pageId="46" pageNumber="47">D. flavifrons</emphasis>
</taxonomicName>
: 1 pupal exuviae, Vancouver,
<collectingRegion id="4818F83DBD003058FC181EB949DAE6C1" box="[992,1194,1196,1221]" country="Canada" name="British Columbia" pageId="46" pageNumber="47">British Columbia</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FB441EB94863E6C1" box="[1212,1299,1196,1221]" name="Canada" pageId="46" pageNumber="47">Canada</collectingCountry>
,
<date id="FE62101FBD003058FADC1EB948ECE6C1" box="[1316,1436,1196,1221]" pageId="46" pageNumber="47" value="1979-05-15">15-V-1979</date>
(CNCI).
<taxonomicName id="4DDC4D5CBD003058FEF81EC44C0DE6EC" authorityName="Kieffer" authorityYear="1924" box="[256,381,1231,1256]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="halobia">
<emphasis id="B8A8EACDBD003058FEF81EC44C0DE6EC" box="[256,381,1231,1256]" italics="true" pageId="46" pageNumber="47">D. halobia</emphasis>
</taxonomicName>
: 8 pupal exuviae, Brenner Moor on Trave river NW of Bad Oldesloe,
<collectingRegion id="4818F83DBD003058FB221EDA48E8E6EC" box="[1242,1432,1231,1256]" country="Germany" name="Schleswig-Holstein" pageId="46" pageNumber="47">Slesvig-Holstein</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FF6F1EE14C70E709" box="[151,256,1268,1293]" name="Germany" pageId="46" pageNumber="47">Germany</collectingCountry>
,
<date id="FE62101FBD003058FEF51EE14CEBE709" box="[269,411,1268,1293]" pageId="46" pageNumber="47" value="1922-07-15">15-VII-1922</date>
(ZSMC).
<taxonomicName id="4DDC4D5CBD003058FDEB1EE04FE7E709" authorityName="Ingram &amp; Macfie" authorityYear="1931" box="[531,663,1268,1293]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lacustris">
<emphasis id="B8A8EACDBD003058FDEB1EE04FE7E709" box="[531,663,1268,1293]" italics="true" pageId="46" pageNumber="47">D. lacustris</emphasis>
</taxonomicName>
: 1 pupal exuviae, Lago Ramos Mexia,
<collectingRegion id="4818F83DBD003058FB981EE149B8E70F" box="[1120,1224,1268,1291]" country="Argentina" name="Neuquen" pageId="46" pageNumber="47">Neuquen</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FB2D1EE14837E709" box="[1237,1351,1268,1293]" name="Argentina" pageId="46" pageNumber="47">Argentina</collectingCountry>
,
<date id="FE62101FBD003058FAAD1EE14DBFE734" pageId="46" pageNumber="47" value="1983-12-02">2-XII- 1983</date>
(MLPA).
<taxonomicName id="4DDC4D5CBD003058FEA91F0C4F67E734" authorityName="Waugh &amp; Wirth" authorityYear="1976" box="[337,535,1303,1328]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="leptobranchia">
<emphasis id="B8A8EACDBD003058FEA91F0C4F67E734" box="[337,535,1303,1328]" italics="true" pageId="46" pageNumber="47">D. leptobranchia</emphasis>
</taxonomicName>
: 1 pupal exuviae, Fishing Creek, Newcomb,
<collectingRegion id="4818F83DBD003058FBBF1F0D49B4E734" box="[1095,1220,1303,1328]" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FB2E1F024861E734" box="[1238,1297,1303,1328]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FADC1F0248EDE734" box="[1316,1437,1303,1328]" pageId="46" pageNumber="47" value="1958-05-28">28-V-1958</date>
(USNM).
<taxonomicName id="4DDC4D5CBD003058FEF11F284CCEE751" authorityName="Waugh &amp; Wirth" authorityYear="1976" box="[265,446,1340,1365]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="messersmithi">
<emphasis id="B8A8EACDBD003058FEF11F284CCEE751" box="[265,446,1340,1365]" italics="true" pageId="46" pageNumber="47">D. messersmithi</emphasis>
</taxonomicName>
: 1 pupal exuviae, Res. Sta., Newcomb,
<collectingRegion id="4818F83DBD003058FC791F294E88E751" box="[897,1016,1340,1365]" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FBFB1F29494EE751" box="[1027,1086,1340,1365]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FBB21F2949BEE751" box="[1098,1230,1340,1365]" pageId="46" pageNumber="47" value="1958-06-26">26-VI-1958</date>
(NYSM);
<specimenCount id="9CDAFD56BD003058FAB11F2948E8E757" box="[1353,1432,1340,1365]" pageId="46" pageNumber="47" type="pupa">1 pupa</specimenCount>
, 1 pupal exuviae, Quogue, Long Island,
<collectingRegion id="4818F83DBD003058FDAD1F754FBCE77C" box="[597,716,1375,1400]" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FD2F1F4A4E62E77C" box="[727,786,1375,1400]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FCE61F4A4EF8E77C" box="[798,904,1375,1400]" pageId="46" pageNumber="47" value="1957-05-09">9-V-1957</date>
(NYSM); 2 pupal exuviae (of
<typeStatus id="5567887DBD003058FB1E1F4A4826E77C" box="[1254,1366,1375,1400]" pageId="46" pageNumber="47" type="paratype">paratypes</typeStatus>
), Kerr County,
<collectingRegion id="4818F83DBD003058FF0E1F914C4AE79F" box="[246,314,1412,1435]" country="United States of America" name="Texas" pageId="46" pageNumber="47">Texas</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FEBD1F914CF0E799" box="[325,384,1412,1437]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
, coll.
<date id="FE62101FBD003058FE3C1F914F23E799" box="[452,595,1412,1437]" pageId="46" pageNumber="47" value="1956-07-12">12-VII-1956</date>
, emerged
<date id="FE62101FBD003058FD3F1F914E4DE799" box="[711,829,1412,1437]" pageId="46" pageNumber="47" value="1956-09-03">3-IX-1956</date>
(USNM).
<taxonomicName id="4DDC4D5CBD003058FC4D1F904945E799" baseAuthorityName="Winnertz" baseAuthorityYear="1852" box="[949,1077,1412,1437]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B8A8EACDBD003058FC4D1F904945E799" box="[949,1077,1412,1437]" italics="true" pageId="46" pageNumber="47">D. modesta</emphasis>
</taxonomicName>
:
<specimenCount id="9CDAFD56BD003058FBBC1F9149D0E79F" box="[1092,1184,1412,1437]" pageId="46" pageNumber="47" type="pupa">6 pupae</specimenCount>
, at least four localities in or NW of Bad Oldesloe,
<collectingRegion id="4818F83DBD003058FE351FB24FFBE7C4" box="[461,651,1447,1472]" country="Germany" name="Schleswig-Holstein" pageId="46" pageNumber="47">Slesvig-Holstein</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FD6E1FB24F8FE7C4" box="[662,767,1447,1472]" name="Germany" pageId="46" pageNumber="47">Germany</collectingCountry>
, late April to June (ZSMC);
<specimenCount id="9CDAFD56BD003058FBBB1FB249DBE7BB" box="[1091,1195,1447,1472]" pageId="46" pageNumber="47" type="pupa">11 pupae</specimenCount>
, northern
<collectingCountry id="F2CB764FBD003058FAE41FB24820E7C4" box="[1308,1360,1447,1472]" name="Italy" pageId="46" pageNumber="47">Italy</collectingCountry>
,
<date id="FE62101FBD003058FAA41FB24DBFE7E1" pageId="46" pageNumber="47" value="1950-05-24">24-V- 1950</date>
(ZSMC).
<taxonomicName id="4DDC4D5CBD003058FEB41FD84CACE7E1" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[332,476,1484,1509]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="B8A8EACDBD003058FEB41FD84CACE7E1" box="[332,476,1484,1509]" italics="true" pageId="46" pageNumber="47">D. mutabilis</emphasis>
</taxonomicName>
: 3 pupal exuviae, Rusk County,
<collectingRegion id="4818F83DBD003058FC9F1FD94EAFE7E1" box="[871,991,1484,1509]" country="United States of America" name="Wisconsin" pageId="46" pageNumber="47">Wisconsin</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FC031FD94946E7E1" box="[1019,1078,1484,1509]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FBBE1FD949B0E7E1" box="[1094,1216,1484,1509]" pageId="46" pageNumber="47" value="1953-10-17">17-X-1953</date>
(USNM); 8 pupal exuviae, High Cr. Fen Preserve,
<quantity id="4D249B3ABD003058FDF51FFA4F31E40C" box="[525,577,1519,1544]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="46" pageNumber="47" unit="mi" value="9.0">9 mi</quantity>
S of Fairplay Park,
<collectingRegion id="4818F83DBD003058FCD41FFA4EE7E40C" box="[812,919,1519,1544]" country="United States of America" name="Colorado" pageId="46" pageNumber="47">Colorado</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FC5D1FFA4E90E40C" box="[933,992,1519,1544]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FC141FFA4900E40C" box="[1004,1136,1519,1544]" pageId="46" pageNumber="47" value="1995-06-18">18-VI-1995</date>
(CNCI); 1 pupal exuviae, Hamilton-Essex Line, Newcomb,
<collectingRegion id="4818F83DBD003058FDE21C014FE2E429" box="[538,658,1556,1581]" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FD671C014FA9E429" box="[671,729,1556,1581]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FD251C014E26E429" box="[733,854,1556,1581]" pageId="46" pageNumber="47" value="1959-05-29">29-V-1959</date>
(NSYM); 1 pupal exuviae, Cow Neck Salt Marsh, North Sea, Long Island,
<collectingRegion id="4818F83DBD003058FE3B1C2D4F4FE454" box="[451,575,1591,1616]" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FDA81C224FFBE454" box="[592,651,1591,1616]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FD641C224E6CE454" box="[668,796,1591,1616]" pageId="46" pageNumber="47" value="1957-07-09">9-VII-1957</date>
(NSYM); 2 pupal exuviae, Dyke Marsh, Alexandria,
<collectingRegion id="4818F83DBD003058FF6F1C494D85E471" box="[151,245,1628,1653]" country="United States of America" name="Virginia" pageId="46" pageNumber="47">Virginia</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FEF91C494C4CE471" box="[257,316,1628,1653]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FEB11C494CB1E471" box="[329,449,1628,1653]" pageId="46" pageNumber="47" value="1977-05-18">18-V-1977</date>
(USNM); 1 pupal exuviae,
<quantity id="4D249B3ABD003058FCF91C494E4BE471" box="[769,827,1628,1653]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="46" pageNumber="47" unit="km" value="7.0">7 km</quantity>
W of Luray, Page County,
<collectingRegion id="4818F83DBD003058FB8E1C4949A4E471" box="[1142,1236,1628,1653]" country="United States of America" name="Virginia" pageId="46" pageNumber="47">Virginia</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FB181C49486AE471" box="[1248,1306,1628,1653]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FAD01C4948ECE471" box="[1320,1436,1628,1653]" pageId="46" pageNumber="47" value="1977-03-06">6-III-1977</date>
(VPIC).
<taxonomicName id="4DDC4D5CBD003058FF001C944CDCE49C" authorityName="Wirth &amp; Beaver" authorityYear="1979" box="[248,428,1663,1688]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="nepenthicola">
<emphasis id="B8A8EACDBD003058FF001C944CDCE49C" box="[248,428,1663,1688]" italics="true" pageId="46" pageNumber="47">D. nepenthicola</emphasis>
</taxonomicName>
: 3 pupal exuviae, Telok Bahang,
<collectingRegion id="4818F83DBD003058FCD01C954E0DE49C" box="[808,893,1664,1688]" country="Malaysia" name="Pulau Pinang" pageId="46" pageNumber="47">Penang</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FC711C6A4E82E49C" box="[905,1010,1663,1688]" name="Malaysia" pageId="46" pageNumber="47">Malaysia</collectingCountry>
,
<date id="FE62101FBD003058FC061C6A4939E49C" box="[1022,1097,1663,1688]" pageId="46" pageNumber="47" value="1977-01">I-1977</date>
(USNM); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD003058FE991CB14C90E4B9" box="[353,480,1700,1725]" pageId="46" pageNumber="47" value="1976-12-04">4-XII-1976</date>
(USNM); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD003058FBFF1CB14916E4B9" box="[1031,1126,1700,1725]" pageId="46" pageNumber="47" value="1977-11">XI-1977</date>
(USNM); 1 pupal exuviae, Ayer Itam,
<collectingRegion id="4818F83DBD003058FEEE1CDD4C1BE4E4" box="[278,363,1736,1760]" country="Malaysia" name="Pulau Pinang" pageId="46" pageNumber="47">Penang</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FE8F1CD24C90E4E4" box="[375,480,1735,1760]" name="Malaysia" pageId="46" pageNumber="47">Malaysia</collectingCountry>
,
<date id="FE62101FBD003058FE151CD24F3CE4E4" box="[493,588,1735,1760]" pageId="46" pageNumber="47" value="1976-11">XI-1976</date>
(USNM).
<taxonomicName id="4DDC4D5CBD003058FD3D1CDC4E47E4E4" baseAuthorityName="Skuse" baseAuthorityYear="1889" box="[709,823,1735,1760]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="nigella">
<emphasis id="B8A8EACDBD003058FD3D1CDC4E47E4E4" box="[709,823,1735,1760]" italics="true" pageId="46" pageNumber="47">D. nigella</emphasis>
</taxonomicName>
: 7 pupal exuviae, Berowra Waters,
<collectingRegion id="4818F83DBD003058FB341CDD48E8E4E4" box="[1228,1432,1735,1760]" country="Australia" name="New South Wales" pageId="46" pageNumber="47">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FF6F1CF94C71E501" box="[151,257,1772,1797]" name="Australia" pageId="46" pageNumber="47">Australia</collectingCountry>
,
<date id="FE62101FBD003058FEF71CF94CE3E501" box="[271,403,1772,1797]" pageId="46" pageNumber="47" value="1956-09-18">18-IX-1956</date>
(ANIC).
<taxonomicName id="4DDC4D5CBD003058FDFD1CF84FC7E501" authorityName="Carter, Ingram &amp; Macfie" authorityYear="1921" box="[517,695,1772,1797]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pallidihalter">
<emphasis id="B8A8EACDBD003058FDFD1CF84FC7E501" box="[517,695,1772,1797]" italics="true" pageId="46" pageNumber="47">D. pallidihalter</emphasis>
</taxonomicName>
: 2 pupal exuviae, Lodwig’s Folly, Hectorspruit, East Transvaal,
<collectingCountry id="F2CB764FBD003058FF6F1D1A4C59E52C" box="[151,297,1807,1832]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
,
<date id="FE62101FBD003058FECD1D1A4CD4E52C" box="[309,420,1807,1832]" pageId="46" pageNumber="47" value="1974-01-21">21-I-1974</date>
(NMSA).
<taxonomicName id="4DDC4D5CBD003058FDE51D044FC8E523" authorityName="Wirth" authorityYear="1969" box="[541,696,1808,1832]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="paracincta">
<emphasis id="B8A8EACDBD003058FDE51D044FC8E523" box="[541,696,1808,1832]" italics="true" pageId="46" pageNumber="47">D. paracincta</emphasis>
</taxonomicName>
: 1 pupal exuviae,
<collectingCountry id="F2CB764FBD003058FC7E1D1A4EDEE52C" box="[902,942,1807,1832]" name="Iceland" pageId="46" pageNumber="47">Isla</collectingCountry>
Isabella,
<collectingRegion id="4818F83DBD003058FBE31D1A499AE52C" box="[1051,1258,1807,1832]" country="Ecuador" name="Galapagos" pageId="46" pageNumber="47">Galapagos Islands</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FB0D1D1A4824E52C" box="[1269,1364,1807,1832]" name="Ecuador" pageId="46" pageNumber="47">Ecuador</collectingCountry>
,
<date id="FE62101FBD003058FA981D1A4DBFE549" pageId="46" pageNumber="47" value="1989-03-09">9-III- 1989</date>
(CNCI); 4 pupal exuviae, Pto.
<collectingCountry id="F2CB764FBD003058FDB81D214F18E549" box="[576,616,1844,1869]" name="Iceland" pageId="46" pageNumber="47">Isla</collectingCountry>
Isabela,
<collectingRegion id="4818F83DBD003058FD2B1D214ED5E549" box="[723,933,1844,1869]" country="Ecuador" name="Galapagos" pageId="46" pageNumber="47">Galapagos Islands</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FC4C1D214963E549" box="[948,1043,1844,1869]" name="Ecuador" pageId="46" pageNumber="47">Ecuador</collectingCountry>
,
<date id="FE62101FBD003058FBD91D2149E5E549" box="[1057,1173,1844,1869]" pageId="46" pageNumber="47" value="1989-03-06">6-III-1989</date>
(CNCI).
<taxonomicName id="4DDC4D5CBD003058FAFE1D20480CE549" baseAuthorityName="Johannsen" baseAuthorityYear="1931" box="[1286,1404,1844,1869]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="perfida">
<emphasis id="B8A8EACDBD003058FAFE1D20480CE549" box="[1286,1404,1844,1869]" italics="true" pageId="46" pageNumber="47">D. perfida</emphasis>
</taxonomicName>
: 2 pupal exuviae, Ranoe Bedali,
<collectingRegion id="4818F83DBD003058FDF91D4D4F45E56B" box="[513,565,1880,1903]" country="Indonesia" name="Jawa Timur" pageId="46" pageNumber="47">Java</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FDB01D424FC5E574" box="[584,693,1879,1904]" name="Indonesia" pageId="46" pageNumber="47">Indonesia</collectingCountry>
(ZSMC).
<taxonomicName id="4DDC4D5CBD003058FCCD1D4C4EB6E574" baseAuthorityName="Wirth" baseAuthorityYear="1952" box="[821,966,1879,1904]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pollinosa">
<emphasis id="B8A8EACDBD003058FCCD1D4C4EB6E574" box="[821,966,1879,1904]" italics="true" pageId="46" pageNumber="47">D. pollinosa</emphasis>
</taxonomicName>
: 2 pupal exuviae, 1 pupal exuviae (of
<typeStatus id="5567887DBD003058FF6F1D694D8CE591" box="[151,252,1916,1941]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
), Kern County,
<collectingRegion id="4818F83DBD003058FE4B1D694F58E591" box="[435,552,1916,1941]" country="United States of America" name="California" pageId="46" pageNumber="47">California</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FDCD1D694F1DE591" box="[565,621,1916,1941]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
(USNM).
<taxonomicName id="4DDC4D5CBD003058FD131D684ED3E591" authorityName="Waugh &amp; Wirth" authorityYear="1976" box="[747,931,1916,1941]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudocincta">
<emphasis id="B8A8EACDBD003058FD131D684ED3E591" box="[747,931,1916,1941]" italics="true" pageId="46" pageNumber="47">D. pseudocincta</emphasis>
</taxonomicName>
: 3 pupal exuviae, Hotsprings, Thermopolis,
<collectingRegion id="4818F83DBD003058FF6F1DB54C78E5BD" box="[151,264,1952,1977]" country="United States of America" name="Wyoming" pageId="46" pageNumber="47">Wyoming</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FEEC1DB54C3FE5BD" box="[276,335,1952,1977]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FEA31DB54CAFE5BD" box="[347,479,1952,1977]" pageId="46" pageNumber="47" value="1960-06-13">13-VI-1960</date>
(USNM); 3 pupal exuviae, Oak Island, Long Island,
<collectingRegion id="4818F83DBD003058FBC41DB449C2E5BD" box="[1084,1202,1952,1977]" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FB461DB54989E5BD" box="[1214,1273,1952,1977]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FAFD1DB5480DE5BD" box="[1285,1405,1952,1977]" pageId="46" pageNumber="47" value="1956-05-15">15-V-1956</date>
(1 USNM, 2 NYSM); 2 pupal exuviae (of
<typeStatus id="5567887DBD003058FD861DD14F93E5D9" box="[638,739,1988,2013]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
), Orient Beach, Long Island,
<collectingRegion id="4818F83DBD003058FBB31DD149B7E5D9" box="[1099,1223,1988,2013]" country="United States of America" name="New York" pageId="46" pageNumber="47">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD003058FB201DD14863E5D9" box="[1240,1299,1988,2013]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
,
<date id="FE62101FBD003058FADC1DD148ECE5D9" box="[1316,1436,1988,2013]" pageId="46" pageNumber="47" value="1963-05-25">25-V-1963</date>
(USNM); 1 pupal exuviae, Cow Neck Salt Marsh, Long Island,
<collectingCountry id="F2CB764FBD013059FC961A824ED9E2B4" box="[878,937,151,176]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FC4D1A824950E2B4" box="[949,1056,151,176]" pageId="47" pageNumber="48" value="1956-05-01">1-V-1956</date>
(NYSM).
<taxonomicName id="4DDC4D5CBD013059FB621A8C480EE2B4" authorityName="Waugh &amp; Wirth" authorityYear="1976" box="[1178,1406,151,176]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pseudoincisurata">
<emphasis id="B8A8EACDBD013059FB621A8C480EE2B4" box="[1178,1406,151,176]" italics="true" pageId="47" pageNumber="48">D. pseudoincisurata</emphasis>
</taxonomicName>
: 3 pupal exuviae, Iroquois Falls,
<collectingRegion id="4818F83DBD013059FE121AA94F32E2D1" box="[490,578,188,213]" country="Canada" name="Ontario" pageId="47" pageNumber="48">Ontario</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FDB51AA94FD4E2D1" box="[589,676,188,213]" name="Canada" pageId="47" pageNumber="48">Canada</collectingCountry>
,
<date id="FE62101FBD013059FD571AA94E43E2D1" box="[687,819,188,213]" pageId="47" pageNumber="48" value="1987-06-21">21-VI-1987</date>
(CNCI); 5 pupal exuviae, Plummers Island,
<collectingRegion id="4818F83DBD013059FAD01AA948E8E2D1" box="[1320,1432,188,213]" country="United States of America" name="Maryland" pageId="47" pageNumber="48">Maryland</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FF6F1ACA4DA2E2FC" box="[151,210,223,248]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FF261ACA4C12E2FC" box="[222,354,223,248]" pageId="47" pageNumber="48" value="1977-06-23">23-VI-1977</date>
(USNM); 1 pupal exuviae, Mt. Solon,
<collectingRegion id="4818F83DBD013059FCD81ACA4E0EE2FC" box="[800,894,223,248]" country="United States of America" name="Virginia" pageId="47" pageNumber="48">Virginia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FC711ACA4EB4E2FC" box="[905,964,223,248]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FC281ACA4917E2FC" box="[976,1127,223,248]" pageId="47" pageNumber="48" value="1951-08-22">22-VIII-1951</date>
(USNM).
<taxonomicName id="4DDC4D5CBD013059FB1A1AF4480EE2FC" authorityName="Waugh &amp; Wirth" authorityYear="1976" box="[1250,1406,223,248]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="spiniforma">
<emphasis id="B8A8EACDBD013059FB1A1AF4480EE2FC" box="[1250,1406,223,248]" italics="true" pageId="47" pageNumber="48">D. spiniforma</emphasis>
</taxonomicName>
: 1 pupal exuviae, Towd Point, Long Island,
<collectingRegion id="4818F83DBD013059FD821B114F84E319" box="[634,756,260,285]" country="United States of America" name="New York" pageId="47" pageNumber="48">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FCFA1B114E4DE319" box="[770,829,260,285]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FCB31B114EBFE319" box="[843,975,260,285]" pageId="47" pageNumber="48" value="1955-06-30">30-VI-1955</date>
(NYSM); 2 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FF061B324CF2E344" box="[254,386,295,320]" pageId="47" pageNumber="48" value="1955-06-28">28-VI-1955</date>
(NYSM).
<taxonomicName id="4DDC4D5CBD013059FDF01B3C4F19E344" baseAuthorityName="Johannsen" baseAuthorityYear="1931" box="[520,617,297,320]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tersa">
<emphasis id="B8A8EACDBD013059FDF01B3C4F19E344" box="[520,617,297,320]" italics="true" pageId="47" pageNumber="48">D. tersa</emphasis>
</taxonomicName>
: 1 pupal exuviae, between Tjibodas and the Gedeh, Kuripan,
<collectingRegion id="4818F83DBD013059FA9B1B3D48E7E33B" box="[1379,1431,296,319]" country="Indonesia" name="Jawa Timur" pageId="47" pageNumber="48">Java</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FF6F1B594C77E361" box="[151,263,332,357]" name="Indonesia" pageId="47" pageNumber="48">Indonesia</collectingCountry>
,
<date id="FE62101FBD013059FEE91B594CEFE361" box="[273,415,332,357]" pageId="47" pageNumber="48" value="1929-07-13">13-VII-1929</date>
(ZSMC).
<taxonomicName id="4DDC4D5CBD013059FDEA1B584FD7E361" authorityName="Meillon" authorityYear="1936" box="[530,679,332,357]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis id="B8A8EACDBD013059FDEA1B584FD7E361" box="[530,679,332,357]" italics="true" pageId="47" pageNumber="48">D. thompsoni</emphasis>
</taxonomicName>
: 1 pupal exuviae, Louis Fichardt, North Transvaal,
<collectingCountry id="F2CB764FBD013059FB1F1B594809E361" box="[1255,1401,332,357]" name="South Africa" pageId="47" pageNumber="48">South Africa</collectingCountry>
,
<date id="FE62101FBD013059FA7C1B594D91E38C" pageId="47" pageNumber="48" value="1974-01-02">2- I-1974</date>
(NMSA); 1 pupal exuviae, Sabi River, East Transvaal,
<collectingCountry id="F2CB764FBD013059FCAB1B7A4E92E38C" box="[851,994,367,392]" name="South Africa" pageId="47" pageNumber="48">South Africa</collectingCountry>
<date id="FE62101FBD013059FC121B7A491AE38C" box="[1002,1130,367,392]" pageId="47" pageNumber="48" value="1973-12-03">3-XII-1973</date>
(NMSA); 1 pupal exuviae, Doornpan, Bulge River, North Transvaal,
<collectingCountry id="F2CB764FBD013059FD6F1B814E41E3A9" box="[663,817,404,429]" name="South Africa" pageId="47" pageNumber="48">South Africa</collectingCountry>
,
<date id="FE62101FBD013059FCBD1B814ECBE3A9" box="[837,955,404,429]" pageId="47" pageNumber="48" value="1973-11-06">6-XI-1973</date>
(NMSA); 1 pupal exuviae, Die Tuine, Transvaal,
<collectingCountry id="F2CB764FBD013059FEEA1BA24CD3E3D4" box="[274,419,439,464]" name="South Africa" pageId="47" pageNumber="48">South Africa</collectingCountry>
,
<date id="FE62101FBD013059FE561BA24F60E3D4" box="[430,528,439,464]" pageId="47" pageNumber="48" value="1983-01-06">6-I-1983</date>
(SAIM).
<taxonomicName id="4DDC4D5CBD013059FD851BAC4F80E3D4" authorityName="Meillon" authorityYear="1936" box="[637,752,439,464]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tugelae">
<emphasis id="B8A8EACDBD013059FD851BAC4F80E3D4" box="[637,752,439,464]" italics="true" pageId="47" pageNumber="48">D. tugelae</emphasis>
</taxonomicName>
: 3 pupal exuviae,
<quantity id="4D249B3ABD013059FC441BA24E85E3CB" box="[956,1013,439,464]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="47" pageNumber="48" unit="m" value="15.0">15 m</quantity>
SE of Potgietersrus, Transvaal,
<collectingCountry id="F2CB764FBD013059FAA21BA24D92E3F1" name="South Africa" pageId="47" pageNumber="48">South Africa</collectingCountry>
,
<date id="FE62101FBD013059FF091BC94C23E3F1" box="[241,339,476,501]" pageId="47" pageNumber="48" value="1974-01-03">3-I-1974</date>
(NMSA); 4 pupal exuviae,
<quantity id="4D249B3ABD013059FD5B1BC94F91E3F1" box="[675,737,476,501]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="47" pageNumber="48" unit="km" value="7.0">7 km</quantity>
S of Messina, North Transvaal,
<collectingCountry id="F2CB764FBD013059FB9F1BC9498EE3F1" box="[1127,1278,476,501]" name="South Africa" pageId="47" pageNumber="48">South Africa</collectingCountry>
,
<date id="FE62101FBD013059FAF71BC948ECE3F1" box="[1295,1436,476,501]" pageId="47" pageNumber="48" value="1973-12-26">26-XII-1973</date>
(NMSA); 2 pupal exuviae, Onderstepoort, Transvaal,
<collectingCountry id="F2CB764FBD013059FCF81BEA4EE4E01C" box="[768,916,511,536]" name="South Africa" pageId="47" pageNumber="48">South Africa</collectingCountry>
,
<date id="FE62101FBD013059FC5A1BEA4952E01C" box="[930,1058,511,536]" pageId="47" pageNumber="48" value="1973-12-08">8-XII-1973</date>
(NMSA); 1 pupal exuviae, Glen Allen Farm, Tzaneen,
<collectingCountry id="F2CB764FBD013059FE6A18314F55E039" box="[402,549,548,573]" name="South Africa" pageId="47" pageNumber="48">South Africa</collectingCountry>
, Jan–April, 1974 (SAIM).
<emphasis id="B8A8EACDBD013059FCAB18304E1EE038" box="[851,878,549,572]" italics="true" pageId="47" pageNumber="48">D.</emphasis>
nr.
<taxonomicName id="4DDC4D5CBD013059FC6118314961E039" box="[921,1041,548,573]" class="Insecta" family="Ceratopogonidae" genus="Ceratopogon" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="brookmani">
<emphasis id="B8A8EACDBD013059FC6118314961E039" box="[921,1041,548,573]" italics="true" pageId="47" pageNumber="48">brookmani</emphasis>
</taxonomicName>
:
<specimenCount id="9CDAFD56BD013059FBD81831490BE03F" box="[1056,1147,548,573]" pageId="47" pageNumber="48" type="pupa">6 pupae</specimenCount>
, High Cr. Fen Preserve,
<quantity id="4D249B3ABD013059FA7618314DC5E064" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="47" pageNumber="48" unit="mi" value="9.0">9 mi</quantity>
S of Fairplay Park,
<collectingRegion id="4818F83DBD013059FE5318524F66E064" box="[427,534,583,608]" country="United States of America" name="Colorado" pageId="47" pageNumber="48">Colorado</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FDDD18524F10E064" box="[549,608,583,608]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FD8818524F8DE064" box="[624,765,583,608]" pageId="47" pageNumber="48" value="1995-07-17">17-VII-1995</date>
(CNCI).
<taxonomicName id="4DDC4D5CBD013059FC88185C4EA9E05B" box="[880,985,584,608]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B8A8EACDBD013059FC88185C4EF4E064" box="[880,900,585,608]" italics="true" pageId="47" pageNumber="48">D</emphasis>
.
<emphasis id="B8A8EACDBD013059FC6E185D4EA9E05B" box="[918,985,584,607]" italics="true" pageId="47" pageNumber="48">major</emphasis>
</taxonomicName>
(?): 1 pupal exuviae, Horsepen Creek (Poanoke R. trib.), Charlotte County,
<collectingRegion id="4818F83DBD013059FDC018794FE6E081" box="[568,662,620,645]" country="United States of America" name="Virginia" pageId="47" pageNumber="48">Virginia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FD5818794FABE081" box="[672,731,620,645]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
, 15-VI-975 (VPIC).
<emphasis id="B8A8EACDBD013059FC3A18784EA6E080" box="[962,982,621,644]" italics="true" pageId="47" pageNumber="48">D</emphasis>
. nr.
<taxonomicName id="4DDC4D5CBD013059FBFF1879491FE081" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[1031,1135,620,645]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="B8A8EACDBD013059FBFF1879491FE081" box="[1031,1135,620,645]" italics="true" pageId="47" pageNumber="48">mutabilis</emphasis>
</taxonomicName>
: 1 pupal exuviae, Res. Sta. Stream, Newcomb,
<collectingRegion id="4818F83DBD013059FE8118854C81E0AC" box="[377,497,655,680]" country="United States of America" name="New York" pageId="47" pageNumber="48">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FE05189A4F48E0AC" box="[509,568,655,680]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FDBD189A4FCCE0AC" box="[581,700,655,680]" pageId="47" pageNumber="48" value="1958-06-06">6-VI-1958</date>
(NYSM); 1 pupal exuviae, Salmon River, Blue Mountain Lake,
<collectingRegion id="4818F83DBD013059FF6F18A14C7CE0C9" box="[151,268,692,717]" country="United States of America" name="New York" pageId="47" pageNumber="48">New York</collectingRegion>
<collectingCountry id="F2CB764FBD013059FEED18A14C20E0C9" box="[277,336,692,717]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FEA618A14CA6E0C9" box="[350,470,692,717]" pageId="47" pageNumber="48" value="1959-05-14">14-V-1959</date>
(NYSM).
<emphasis id="B8A8EACDBD013059FDAB18A04F17E0C8" box="[595,615,693,716]" italics="true" pageId="47" pageNumber="48">D</emphasis>
. sp.: 4 pupal exuviae (in glycerin), Herman Lake,
<quantity id="4D249B3ABD013059FB4A18A1499DE0C9" box="[1202,1261,692,717]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="47" pageNumber="48" unit="km" value="4.0">4 km</quantity>
SE Eagle Bay,
<collectingRegion id="4818F83DBD013059FF6F18C24C11E0F4" box="[151,353,727,752]" country="Canada" name="British Columbia" pageId="47" pageNumber="48">British Columbia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FE8918C24CB8E0F4" box="[369,456,727,752]" name="Canada" pageId="47" pageNumber="48">Canada</collectingCountry>
,
<date id="FE62101FBD013059FE2018C24F20E0F4" box="[472,592,727,752]" pageId="47" pageNumber="48" value="2013-05-26">26-V-2013</date>
(CNCI); 5 pupal exuviae, Bolean Lake,
<quantity id="4D249B3ABD013059FBC318C2490AE0F4" box="[1083,1146,727,752]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="47" pageNumber="48" unit="km" value="6.0">6 km</quantity>
NE of Falkland,
<collectingRegion id="4818F83DBD013059FAA818C24C78E111" country="Canada" name="British Columbia" pageId="47" pageNumber="48">British Columbia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FEED18E94C1CE111" box="[277,364,764,789]" name="Canada" pageId="47" pageNumber="48">Canada</collectingCountry>
,
<date id="FE62101FBD013059FE8018E94F76E111" box="[376,518,764,789]" pageId="47" pageNumber="48" value="1989-07-12">12-VII-1989</date>
(CNCI); 5 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FBDA18E949A9E111" box="[1058,1241,764,789]" pageId="47" pageNumber="48" value="1989-07-12" valueMax="1989-07-13" valueMin="1989-07-12">12–13-VII-1989</date>
(CNCI); 1 pupal exuviae,
<quantity id="4D249B3ABD013059FF06190A4C49E13C" box="[254,313,799,824]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="47" pageNumber="48" unit="km" value="3.0">3 km</quantity>
E of Salmon Arm,
<collectingRegion id="4818F83DBD013059FDE7190A4F96E13C" box="[543,742,799,824]" country="Canada" name="British Columbia" pageId="47" pageNumber="48">British Columbia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FD0B190A4E3AE13C" box="[755,842,799,824]" name="Canada" pageId="47" pageNumber="48">Canada</collectingCountry>
,
<date id="FE62101FBD013059FCA0190A4EBEE13C" box="[856,974,799,824]" pageId="47" pageNumber="48" value="1988-06-09">9-VI-1988</date>
(CNIC); 3 pupal exuviae, Salmon Arm,
<collectingRegion id="4818F83DBD013059FF6F19514C2CE159" box="[151,348,836,861]" country="Canada" name="British Columbia" pageId="47" pageNumber="48">British Columbia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FE9019514CCFE159" box="[360,447,836,861]" name="Canada" pageId="47" pageNumber="48">Canada</collectingCountry>
,
<date id="FE62101FBD013059FE3319514F28E159" box="[459,600,836,861]" pageId="47" pageNumber="48" value="1988-07-19">19-VII-1988</date>
(CNIC); 5 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FB97195149ABE159" box="[1135,1243,836,861]" pageId="47" pageNumber="48" value="2006-10-02">2-X-2006</date>
(CNCI); 2 pupal exuviae, Gibson Lake, Kokanee Glacier Provincial Park,
<collectingRegion id="4818F83DBD013059FCE419724E91E184" box="[796,993,871,896]" country="Canada" name="British Columbia" pageId="47" pageNumber="48">British Columbia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FC1419724933E184" box="[1004,1091,871,896]" name="Canada" pageId="47" pageNumber="48">Canada</collectingCountry>
,
<date id="FE62101FBD013059FBB7197249BFE184" box="[1103,1231,871,896]" pageId="47" pageNumber="48" value="2008-07-09">9-VII-2008</date>
(CNCI); 18 pupal exuviae, Dark Canyon Creek, Eddy County,
<collectingRegion id="4818F83DBD013059FD5A19994E4AE1A1" box="[674,826,908,933]" country="United States of America" name="New Mexico" pageId="47" pageNumber="48">New Mexico</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FCB219994EF5E1A1" box="[842,901,908,933]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FC6C19994968E1A1" box="[916,1048,908,933]" pageId="47" pageNumber="48" value="1991-06-13">13-VI-1991</date>
(CNCI); 1 pupal exuviae, Chosa Draw, Eddy County,
<collectingRegion id="4818F83DBD013059FE6A19A54F5CE1CC" box="[402,556,943,968]" country="United States of America" name="New Mexico" pageId="47" pageNumber="48">New Mexico</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FDC519BA4F08E1CC" box="[573,632,943,968]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FD7019BA4E6EE1CC" box="[648,798,943,968]" pageId="47" pageNumber="48" value="1997-08-26">26-VIII-1997</date>
(CNCI); 1 pupal exuviae, Midfork Gila River, Grant County,
<collectingRegion id="4818F83DBD013059FF0019C14CFEE1E9" box="[248,398,980,1005]" country="United States of America" name="New Mexico" pageId="47" pageNumber="48">New Mexico</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FE6419C14CA6E1E9" box="[412,470,980,1005]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FE1C19C14F0AE1E9" box="[484,634,980,1005]" pageId="47" pageNumber="48" value="1991-08-15">15-VIII-1991</date>
(CNCI); 2 pupal exuviae, Gila River above Gila, Grant County,
<collectingRegion id="4818F83DBD013059FA9F19C14D9FE614" country="United States of America" name="New Mexico" pageId="47" pageNumber="48">New Mexico</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FF0419E24C47E614" box="[252,311,1015,1040]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FEBB19E24CAAE614" box="[323,474,1015,1040]" pageId="47" pageNumber="48" value="1991-08-16">16-VIII-1991</date>
(CNCI); 2 pupal exuviae, Towd Point, Southampton, Long Island,
<collectingRegion id="4818F83DBD013059FB2219ED4821E614" box="[1242,1361,1015,1040]" country="United States of America" name="New York" pageId="47" pageNumber="48">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FAA519E248E8E614" box="[1373,1432,1015,1040]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FF6F1E094C54E631" box="[151,292,1052,1077]" pageId="47" pageNumber="48" value="1954-07-29">29-VII-1954</date>
(NYSM); 2 pupal exuviae, Salt Marsh, Towd Point, Southampton, Long Island,
<collectingRegion id="4818F83DBD013059FB481E094855E631" box="[1200,1317,1052,1077]" country="United States of America" name="New York" pageId="47" pageNumber="48">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FAC91E09481CE631" box="[1329,1388,1052,1077]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FA8F1E094C78E65C" pageId="47" pageNumber="48" value="1954-08-10">10- VIII-1954</date>
(NYSM); 1 pupal exuviae, Dune Rd. Salt Marsh, H. Bays, Long Island,
<collectingRegion id="4818F83DBD013059FBA01E5549A0E65C" box="[1112,1232,1087,1112]" country="United States of America" name="New York" pageId="47" pageNumber="48">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FB251E2A4868E65C" box="[1245,1304,1087,1112]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FADE1E2A48ECE65C" box="[1318,1436,1087,1112]" pageId="47" pageNumber="48" value="1956-06-04">4-VI-1956</date>
(NYSM); 3 pupal exuviae, Cow Neck, Long Island,
<collectingRegion id="4818F83DBD013059FD111E714E10E679" box="[745,864,1124,1149]" country="United States of America" name="New York" pageId="47" pageNumber="48">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FC931E714ED6E679" box="[875,934,1124,1149]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FC4A1E71495AE679" box="[946,1066,1124,1149]" pageId="47" pageNumber="48" value="1956-05-28">28-V-1956</date>
(NYSM); 1 pupal exuviae, New Salem,
<collectingRegion id="4818F83DBD013059FF131E9D4C11E6A4" box="[235,353,1159,1184]" country="United States of America" name="New York" pageId="47" pageNumber="48">New York</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FE941E924CD7E6A4" box="[364,423,1159,1184]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FE4A1E924F5BE6A4" box="[434,555,1159,1184]" pageId="47" pageNumber="48" value="1963-05-13">13-V-1963</date>
(NYSM); 1 pupal exuviae, Dunaley Bog,
<quantity id="4D249B3ABD013059FBFF1E9D4949E6A4" box="[1031,1081,1159,1184]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="47" pageNumber="48" unit="mi" value="7.0">7 mi</quantity>
S of Cashiers, Jackson County,
<collectingRegion id="4818F83DBD013059FF6F1EB94C34E6C1" box="[151,324,1196,1221]" country="United States of America" name="North Carolina" pageId="47" pageNumber="48">North Carolina</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FEA91EB94CFCE6C1" box="[337,396,1196,1221]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FE611EB94F56E6C1" box="[409,550,1196,1221]" pageId="47" pageNumber="48" value="1987-07-13">13-VII-1987</date>
(USNM); 1 pupal exuviae, Montgomery County,
<collectingRegion id="4818F83DBD013059FB9D1EB949B3E6C1" box="[1125,1219,1196,1221]" country="United States of America" name="Virginia" pageId="47" pageNumber="48">Virginia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FB281EB9487BE6C1" box="[1232,1291,1196,1221]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FAE01EB948ECE6C1" box="[1304,1436,1196,1221]" pageId="47" pageNumber="48" value="1975-06-22">22-VI-1975</date>
(VPIC); 3 pupal exuviae, Assateague Is., Chincoteague NWR, Accomack County,
<collectingRegion id="4818F83DBD013059FBA41EDA49CAE6EC" box="[1116,1210,1231,1256]" country="United States of America" name="Virginia" pageId="47" pageNumber="48">Virginia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FB321EDA4874E6EC" box="[1226,1284,1231,1256]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FAEC1EDA48ECE6EC" box="[1300,1436,1231,1256]" pageId="47" pageNumber="48" value="1975-08-01">1-VIII-1975</date>
(VPIC); 1 pupal exuviae, Waycross,
<collectingRegion id="4818F83DBD013059FDC21EE14FE7E709" box="[570,663,1268,1293]" country="United States of America" name="Georgia" pageId="47" pageNumber="48">Georgia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FD5C1EE14FAFE709" box="[676,735,1268,1293]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
,
<date id="FE62101FBD013059FD131EE14E00E709" box="[747,880,1268,1293]" pageId="47" pageNumber="48" value="1972-06-26">26-VI-1972</date>
(USNM); 1 pupal exuviae, Chiemsee,
<collectingCountry id="F2CB764FBD013059FAD71EE148E8E709" box="[1327,1432,1268,1293]" name="Germany" pageId="47" pageNumber="48">Germany</collectingCountry>
,
<date id="FE62101FBD013059FF6F1F024C5EE734" box="[151,302,1303,1328]" pageId="47" pageNumber="48" value="1990-08-13">13-VIII-1990</date>
(CNCI); 1 pupal exuviae, Grimswald,
<collectingCountry id="F2CB764FBD013059FD151F024E25E734" box="[749,853,1303,1328]" name="Germany" pageId="47" pageNumber="48">Germany</collectingCountry>
(CNCI); 1 pupal exuviae, Brunnsee,
<collectingCountry id="F2CB764FBD013059FAF91F02481AE734" box="[1281,1386,1303,1328]" name="Germany" pageId="47" pageNumber="48">Germany</collectingCountry>
,
<date id="FE62101FBD013059FA8F1F024D8FE751" pageId="47" pageNumber="48" value="1990-07-12">12- VII-1990</date>
(CNCI); 3 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FCE91F294EEFE751" box="[785,927,1340,1365]" pageId="47" pageNumber="48" value="1990-07-13">13-VII-1990</date>
(CNCI); 1 pupal exuviae, Lustsee,
<collectingCountry id="F2CB764FBD013059FAD71F2948E8E751" box="[1327,1432,1340,1365]" name="Germany" pageId="47" pageNumber="48">Germany</collectingCountry>
,
<date id="FE62101FBD013059FF6F1F4A4C7CE77C" box="[151,268,1375,1400]" pageId="47" pageNumber="48" value="1990-06-01">1-VI-1990</date>
(CNCI); 2 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FCBA1F4A4EA7E77C" box="[834,983,1375,1400]" pageId="47" pageNumber="48" value="1990-08-11">11-VIII-1990</date>
(CNCI); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FF021F914C0DE799" box="[250,381,1412,1437]" pageId="47" pageNumber="48" value="1990-06-10">10-VI-1990</date>
(CNCI); 2 pupal exuviae, La Gomera, Canary Islands,
<date id="FE62101FBD013059FBFA1F91491EE799" box="[1026,1134,1412,1437]" pageId="47" pageNumber="48" value="1993-02-08">8-II-1993</date>
(CNCI); 5 pupal exuviae, Skukusa, Kruger National Park,
<collectingCountry id="F2CB764FBD013059FDFA1FB24FE4E7C4" box="[514,660,1447,1472]" name="South Africa" pageId="47" pageNumber="48">South Africa</collectingCountry>
, II,
<date id="FE62101FBD013059FD381FB24E6DE7C4" box="[704,797,1447,1472]" pageId="47" pageNumber="48" value="1993-03">III-1993</date>
(ANIC); 1 pupal exuviae, Moung Sayaboury, Sayaboury Province,
<collectingCountry id="F2CB764FBD013059FEF21FD94C33E7E7" box="[266,323,1484,1507]" name="Laos" pageId="47" pageNumber="48">Laos</collectingCountry>
,
<date id="FE62101FBD013059FEA81FD94CA1E7E1" box="[336,465,1484,1509]" pageId="47" pageNumber="48" value="1967-04-16">16-IV-1967</date>
(BPBM); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FC001FD94917E7E1" box="[1016,1127,1484,1509]" pageId="47" pageNumber="48" value="1968-01-14">14-I-1968</date>
(BPBM); 2 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FE681FFA4F70E40C" box="[400,512,1519,1544]" pageId="47" pageNumber="48" value="1967-01-22">22-I-1967</date>
(BPBM); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FBB71FFA49B0E40C" box="[1103,1216,1519,1544]" pageId="47" pageNumber="48" value="1968-01-13">13-I-1968</date>
(BPBM); 2 pupal exuviae, Dumoga Bone National Park,
<collectingRegion id="4818F83DBD013059FD911C014FA1E429" box="[617,721,1556,1581]" country="Indonesia" name="Sulawesi Utara" pageId="47" pageNumber="48">Sulawesi</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FD191C014E21E429" box="[737,849,1556,1581]" name="Indonesia" pageId="47" pageNumber="48">Indonesia</collectingCountry>
,
<date id="FE62101FBD013059FC991C014EA1E429" box="[865,977,1556,1581]" pageId="47" pageNumber="48" value="1985-01-28">28-I-1985</date>
(ANIC); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FF031C224C1BE454" box="[251,363,1591,1616]" pageId="47" pageNumber="48" value="1985-01-30">30-I-1985</date>
(ANIC); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FC621C224975E454" box="[922,1029,1591,1616]" pageId="47" pageNumber="48" value="1985-02-01">1-II-1985</date>
(ANIC);
<specimenCount id="9CDAFD56BD013059FB811C2249BBE44B" box="[1145,1227,1591,1616]" pageId="47" pageNumber="48" type="pupa">1 pupa</specimenCount>
, Darwin Harbour, Darwin,
<collectingRegion id="4818F83DBD013059FF071C494CA5E471" box="[255,469,1628,1653]" country="Australia" name="Northern Territory" pageId="47" pageNumber="48">Northern Territory</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FE1E1C494F20E471" box="[486,592,1628,1653]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
,
<date id="FE62101FBD013059FD991C494FA9E471" box="[609,729,1628,1653]" pageId="47" pageNumber="48" value="2006-05-12">12-V-2006</date>
(CNCI); 2 pupal exuviae, Collie Rd., W. Darban,
<collectingRegion id="4818F83DBD013059FAB91C494C71E49C" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FEF51C6A4C04E49C" box="[269,372,1663,1688]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
(ANIC); 5 pupal exuviae, Darban,
<collectingRegion id="4818F83DBD013059FCF31C954EA9E49C" box="[779,985,1663,1688]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FC1D1C6A493FE49C" box="[997,1103,1663,1688]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
,
<date id="FE62101FBD013059FBA41C6A49A6E49C" box="[1116,1238,1663,1688]" pageId="47" pageNumber="48" value="1985-10-29">29-X-1985</date>
(ANIC); 1 pupal exuviae, Gnieraoora Pool, Robe River, Pilbara,
<collectingRegion id="4818F83DBD013059FD221CB14EDDE4B9" box="[730,941,1700,1725]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FC381CB1495AE4B9" box="[960,1066,1700,1725]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
,
<date id="FE62101FBD013059FBC41CB149C7E4B9" box="[1084,1207,1700,1725]" pageId="47" pageNumber="48" value="1995-10-30">30-X-1995</date>
(ANIC); 12 pupal exuviae, as previous locality (ANIC); 1 pupal exuviae, Upper Durack River, Kimberley,
<collectingRegion id="4818F83DBD013059FB3E1CDD48E8E4E4" box="[1222,1432,1735,1760]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FF6F1CF94D8EE501" box="[151,254,1772,1797]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
(ANIC); 1 pupal exuviae, Richenda Gorge, Kimberley,
<collectingRegion id="4818F83DBD013059FC8A1CF9494EE501" box="[882,1086,1772,1797]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FBB11CF949C3E501" box="[1097,1203,1772,1797]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
,
<date id="FE62101FBD013059FB461CF94846E501" box="[1214,1334,1772,1797]" pageId="47" pageNumber="48" value="1995-05-10">10-V-1995</date>
(ANIC); 1 pupal exuviae, Palm Springs, Fitzroy, Kimberley,
<collectingRegion id="4818F83DBD013059FD041D054EBCE52C" box="[764,972,1807,1832]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FC241D1A4935E52C" box="[988,1093,1807,1832]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
,
<date id="FE62101FBD013059FBAE1D1A49BEE52C" box="[1110,1230,1807,1832]" pageId="47" pageNumber="48" value="1995-05-12">12-V-1995</date>
(ANIC); 1 pupal exuviae, King Edward River, Kalumburu, Kimberley,
<collectingRegion id="4818F83DBD013059FCE01D214E99E549" box="[792,1001,1844,1869]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FC021D214910E549" box="[1018,1120,1844,1869]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
(ANIC); 9 pupal exuviae, Cangan Pool, Pilbara,
<collectingRegion id="4818F83DBD013059FE591D4D4F01E574" box="[417,625,1879,1904]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FD791D424F97E574" box="[641,743,1879,1904]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
(ANIC); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FAD91D4248ECE574" box="[1313,1436,1879,1904]" pageId="47" pageNumber="48" value="1995-10-15">15-X-1995</date>
(ANIC); 2 pupal exuviae, Whale Back Creek, Fortescu River, Pilbara,
<collectingRegion id="4818F83DBD013059FC371D6949EDE591" box="[975,1181,1916,1941]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FB541D694866E591" box="[1196,1302,1916,1941]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
,
<date id="FE62101FBD013059FADC1D6948ECE591" box="[1316,1436,1916,1941]" pageId="47" pageNumber="48" value="1995-05-16">16-V-1995</date>
(ANIC); 1 pupal exuviae, Erewallana Pool, Pilbara,
<collectingRegion id="4818F83DBD013059FCFC1DB44EA5E5BD" box="[772,981,1952,1977]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD013059FC1E1DB5493FE5BD" box="[998,1103,1952,1977]" name="Australia" pageId="47" pageNumber="48">Australia</collectingCountry>
,
<date id="FE62101FBD013059FB981DB549BCE5BD" box="[1120,1228,1952,1977]" pageId="47" pageNumber="48" value="1995-10-01">1-X-1995</date>
(ANIC); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FE1F1DD14F21E5D9" box="[487,593,1988,2013]" pageId="47" pageNumber="48" value="1995-05-01">1-V-1995</date>
(ANIC); 2 pupal exuviae, as previous locality,
<date id="FE62101FBD013059FB961DD149A8E5D9" box="[1134,1240,1988,2013]" pageId="47" pageNumber="48" value="1995-05-09">9-V-1995</date>
(ANIC); 5 pupal exuviae, Whiskey Pool, Ashburton River, Pilbara,
<collectingRegion id="4818F83DBD1E3046FD161A8D4ECFE2B4" box="[750,959,151,176]" country="Australia" name="Western Australia" pageId="48" pageNumber="49">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FC281A82494AE2B4" box="[976,1082,151,176]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FBB21A8249BBE2B4" box="[1098,1227,151,176]" pageId="48" pageNumber="49" value="1995-04-28">28-IV-1995</date>
(ANIC); 1 pupal exuviae, Mooka Ruins, Gascoyne River,
<collectingRegion id="4818F83DBD1E3046FD8F1AA94E37E2D1" box="[631,839,188,213]" country="Australia" name="Western Australia" pageId="48" pageNumber="49">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FCAE1AA94ECFE2D1" box="[854,959,188,213]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FC371AA94916E2D1" box="[975,1126,188,213]" pageId="48" pageNumber="49" value="1995-08-15">15-VIII-1995</date>
(ANIC); 3 pupal exuviae, Yanchep National Park,
<collectingRegion id="4818F83DBD1E3046FE521AF54F06E2FC" box="[426,630,223,248]" country="Australia" name="Western Australia" pageId="48" pageNumber="49">Western Australia</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FD7B1ACA4F9DE2FC" box="[643,749,223,248]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FD001ACA4E03E2FC" box="[760,883,223,248]" pageId="48" pageNumber="49" value="1985-10-24">24-X-1985</date>
(ANIC); 1 pupal exuviae, Receptably Cemetery, Townsville,
<collectingRegion id="4818F83DBD1E3046FEDA1B114CD9E319" box="[290,425,260,285]" country="Australia" name="Queensland" pageId="48" pageNumber="49">Queensland</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FE4D1B114F6FE319" box="[437,543,260,285]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FDD31B114FD3E319" box="[555,675,260,285]" pageId="48" pageNumber="49" value="1950-05-31">31-V-1950</date>
(ANIC); 1 pupal exuviae, Southport,
<collectingRegion id="4818F83DBD1E3046FBAB1B1149AAE319" box="[1107,1242,260,285]" country="Australia" name="Queensland" pageId="48" pageNumber="49">Queensland</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FB1E1B114820E319" box="[1254,1360,260,285]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FAA41B114DBFE344" pageId="48" pageNumber="49" value="1953-05-21">21-V- 1953</date>
(ANIC); 3 pupal exuviae, Basin, Southport,
<collectingRegion id="4818F83DBD1E3046FD2D1B324E2CE344" box="[725,860,295,320]" country="Australia" name="Queensland" pageId="48" pageNumber="49">Queensland</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FC911B324EA2E344" box="[873,978,295,320]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FC191B32491DE344" box="[993,1133,295,320]" pageId="48" pageNumber="49" value="1950-12-31">31-XII-1950</date>
(ANIC); 1 pupal exuviae, Gladstone,
<collectingRegion id="4818F83DBD1E3046FEE01B594CEFE361" box="[280,415,332,357]" country="Australia" name="Queensland" pageId="48" pageNumber="49">Queensland</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FE531B594F65E361" box="[427,533,332,357]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FDD91B594FE1E361" box="[545,657,332,357]" pageId="48" pageNumber="49" value="1947-01-28">28-I-1947</date>
(ANIC); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FB571B59486EE361" box="[1199,1310,332,357]" pageId="48" pageNumber="49" value="1947-01-27">27-I-1947</date>
(ANIC); 1 pupal exuviae, FNQS Running Creek, Weipa,
<collectingRegion id="4818F83DBD1E3046FD621B7A4E51E38C" box="[666,801,367,392]" country="Australia" name="Queensland" pageId="48" pageNumber="49">Queensland</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FCD41B7A4EE6E38C" box="[812,918,367,392]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FC581B7A497DE38C" box="[928,1037,367,392]" pageId="48" pageNumber="49" value="1985-10-09">9-X-1985</date>
(ANIC); 6 pupal exuviae, Hornsby,
<collectingRegion id="4818F83DBD1E3046FF6F1B814C12E3A9" box="[151,354,404,429]" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FE951B814CA6E3A9" box="[365,470,404,429]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FE191B814F2BE3A9" box="[481,603,404,429]" pageId="48" pageNumber="49" value="1966-02-22">22-II-1966</date>
(ANIC); 1 pupal exuviae, Careel Bay,
<collectingRegion id="4818F83DBD1E3046FBE81B8149ABE3A9" box="[1040,1243,404,429]" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FB1E1B81483FE3A9" box="[1254,1359,404,429]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FAA31B814DBFE3D4" pageId="48" pageNumber="49" value="1966-02-16">16-II- 1966</date>
(ANIC); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FD1D1BA24E10E3D4" box="[741,864,439,464]" pageId="48" pageNumber="49" value="1956-10-29">29-X-1956</date>
(ANIC); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FA8F1BA24D85E3F1" pageId="48" pageNumber="49" value="1965-09-20">20- IX-1965</date>
(ANIC); 2 pupal exuviae, Colo Vale,
<collectingRegion id="4818F83DBD1E3046FD631BC94E15E3F1" box="[667,869,476,501]" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FC971BC94EA9E3F1" box="[879,985,476,501]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FC1B1BC9492CE3F1" box="[995,1116,476,501]" pageId="48" pageNumber="49" value="1958-02-24">24-II-1958</date>
(ANIC); 6 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FE981BEA4CABE01C" box="[352,475,511,536]" pageId="48" pageNumber="49" value="1956-10-18">18-X-1956</date>
(ANIC); 2 pupal exuviae, Stockyard Creek, Colo Vale,
<collectingRegion id="4818F83DBD1E3046FBAE18154853E01C" box="[1110,1315,511,536]" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FAD71BEA48E8E01C" box="[1327,1432,511,536]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FF6F18314C69E039" box="[151,281,548,573]" pageId="48" pageNumber="49" value="1957-03-10">10-III-1957</date>
(ANIC); 2 pupal exuviae, Nattai River, Mittagong,
<collectingRegion id="4818F83DBD1E3046FC981831495BE039" box="[864,1067,548,573]" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FBCE183149D0E039" box="[1078,1184,548,573]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FB5318314851E039" box="[1195,1313,548,573]" pageId="48" pageNumber="49" value="1964-11-04">4-XI-1964</date>
(ANIC); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FDD218524FD5E064" box="[554,677,583,608]" pageId="48" pageNumber="49" value="1968-10-18">18-X-1968</date>
(ANIC); 3 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FB4318524846E064" box="[1211,1334,583,608]" pageId="48" pageNumber="49" value="1968-10-25">25-X-1968</date>
(ANIC); 1 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FDA718794F92E081" box="[607,738,620,645]" pageId="48" pageNumber="49" value="1968-11-15">15-XI-1968</date>
(ANIC); 2 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FADB187948ECE081" box="[1315,1436,620,645]" pageId="48" pageNumber="49" value="1969-02-18">18-II-1969</date>
(ANIC); 8 pupal exuviae, Nattai River, The Crags, Mittagong,
<collectingRegion id="4818F83DBD1E3046FC9B18854941E0AC" box="[867,1073,655,680]" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FBC5189A49D6E0AC" box="[1085,1190,655,680]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FB4B189A486FE0AC" box="[1203,1311,655,680]" pageId="48" pageNumber="49" value="1966-02-09">9-II-1966</date>
(ANIC); 1 pupal exuviae, Picton Lakes,
<collectingRegion id="4818F83DBD1E3046FE1018A14FC7E0C9" box="[488,695,692,717]" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FD3C18A14E5DE0C9" box="[708,813,692,717]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FCC218A14ED5E0C9" box="[826,933,692,717]" pageId="48" pageNumber="49" value="1966-02-02">2-II-1966</date>
(ANIC); 4 pupal exuviae, Mona Vale,
<collectingRegion id="4818F83DBD1E3046FA9F18A14C59E0F4" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FECF18C24CD0E0F4" box="[311,416,727,752]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FE5718C24F5AE0F4" box="[431,554,727,752]" pageId="48" pageNumber="49" value="1956-10-10">10-X-1956</date>
(ANIC); 1 pupal exuviae, Oxford Falls,
<collectingRegion id="4818F83DBD1E3046FBFF18CD49A8E0F4" box="[1031,1240,727,752]" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FB1E18C2483FE0F4" box="[1254,1359,727,752]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FAA618C24DBFE111" pageId="48" pageNumber="49" value="1947-11-06">6-XI- 1947</date>
(ANIC); 4 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FD1318E94E2CE111" box="[747,860,764,789]" pageId="48" pageNumber="49" value="1967-01-18">18-I-1967</date>
(ANIC); 3 pupal exuviae, Wattamolla,
<collectingRegion id="4818F83DBD1E3046FAE618E94DAEE13C" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FF13190A4C24E13C" box="[235,340,799,824]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FE99190A4CABE13C" box="[353,475,799,824]" pageId="48" pageNumber="49" value="1969-02-24">24-II-1969</date>
(ANIC); 3 pupal exuviae, McCarrs Creek,
<collectingRegion id="4818F83DBD1E3046FC28193549EFE13C" box="[976,1183,799,824]" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
,
<collectingCountry id="F2CB764FBD1E3046FB54190A4866E13C" box="[1196,1302,799,824]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FADB190A48EBE13C" box="[1315,1435,799,824]" pageId="48" pageNumber="49" value="1971-02-28">28-II-1971</date>
(ANIC); 2 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FD5C19514E67E159" box="[676,791,836,861]" pageId="48" pageNumber="49" value="1971-03-04">4-III-1971</date>
(ANIC); 2 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FAD4195148ECE159" box="[1324,1436,836,861]" pageId="48" pageNumber="49" value="1969-01-14">14-I-1969</date>
(ANIC); 6 pupal exuviae, as previous locality,
<date id="FE62101FBD1E3046FD5419724E40E184" box="[684,816,871,896]" pageId="48" pageNumber="49" value="1956-09-20">20-IX-1956</date>
(ANIC); 1 pupal exuviae, Anson Bay,
<collectingCountry id="F2CB764FBD1E3046FB08197248E8E184" box="[1264,1432,871,896]" name="Norfolk Island" pageId="48" pageNumber="49">Norfolk Island</collectingCountry>
,
<collectingCountry id="F2CB764FBD1E3046FF6F19994C71E1A1" box="[151,257,908,933]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FEF519994CFFE1A1" box="[269,399,908,933]" pageId="48" pageNumber="49" value="1972-04-14">14-IV-1972</date>
(ANIC); 9 pupal exuviae,
<collectingCountry id="F2CB764FBD1E3046FD3B19994E1BE1A1" box="[707,875,908,933]" name="Norfolk Island" pageId="48" pageNumber="49">Norfolk Island</collectingCountry>
,
<collectingCountry id="F2CB764FBD1E3046FC8019994E92E1A1" box="[888,994,908,933]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FC1719994901E1A1" box="[1007,1137,908,933]" pageId="48" pageNumber="49" value="1972-04-19">19-IV-1972</date>
(ANIC); 3 pupal exuviae, no data (ANIC); 1 pupal exuviae,
<collectingCountry id="F2CB764FBD1E3046FDED19BA4F0FE1CC" box="[533,639,943,968]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
,
<date id="FE62101FBD1E3046FD7119BA4E70E1CC" box="[649,768,943,968]" pageId="48" pageNumber="49" value="1971-06-09">9-VI-1971</date>
(ANIC).
</paragraph>
</subSubSection>
</treatment>
</document>
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