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<document id="183C6C5B955B4BCE394CB555F852597F" ID-CLB-Dataset="28026" ID-DOI="10.11646/zootaxa.4512.1.1" ID-GBIF-Dataset="26e12e1b-2175-4007-871b-d0b73122e5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607575" ID-ZooBank="E752FB7B-F34C-4D12-B8A2-EA6C791DD6C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586644813" checkinUser="plazi" docAuthor="Kaiser, Christine M., Kaiser, Hinrich &amp; OShea, Mark" docDate="2018" docId="C80EBE29FFDDFFEDFF75FED00B7CFBE0" docLanguage="en" docName="zootaxa.4512.1.1.pdf" docOrigin="Zootaxa 4512 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Stegonotus australis Kaiser &amp; Kaiser &amp; OShea 2018, n. comb." docType="treatment" docVersion="8" lastPageNumber="28" masterDocId="3437C651FFC7FFF6FFE2FE470F12FFFE" masterDocTitle="The taxonomic history of Indo-Papuan groundsnakes, genus Stegonotus Duméril et al., 1854 (Colubridae), with some taxonomic revisions and the designation of a neotype for S. parvus (Meyer, 1874)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="27" updateTime="1698713152511" updateUser="ExternalLinkService">
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<mods:title id="1330940F635E6F6F6015C1B0719D32AA">The taxonomic history of Indo-Papuan groundsnakes, genus Stegonotus Duméril et al., 1854 (Colubridae), with some taxonomic revisions and the designation of a neotype for S. parvus (Meyer, 1874)</mods:title>
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<mods:namePart id="43938BCE03BBC259423790A18A0C4938">Kaiser, Christine M.</mods:namePart>
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<mods:namePart id="03EE50EF73B73CA5CE9D6E885CF0EDB6">Kaiser, Hinrich</mods:namePart>
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<mods:namePart id="B803CA78E0BE4D1A909A40A3FCDC50DA">OShea, Mark</mods:namePart>
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<treatment id="C80EBE29FFDDFFEDFF75FED00B7CFBE0" ID-DOI="http://doi.org/10.5281/zenodo.5997371" ID-GBIF-Taxon="154496297" ID-Zenodo-Dep="5997371" LSID="urn:lsid:plazi:treatment:C80EBE29FFDDFFEDFF75FED00B7CFBE0" httpUri="http://treatment.plazi.org/id/C80EBE29FFDDFFEDFF75FED00B7CFBE0" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<subSubSection id="08BD5CB4FFDDFFECFF75FED00A0FFD7D" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="40180F3FFFDDFFECFF75FED00DE2FF4C" blockId="26.[151,752,151,178]" box="[151,752,151,178]" pageId="26" pageNumber="27">
<heading id="1B50B853FFDDFFECFF75FED00DE2FF4C" bold="true" box="[151,752,151,178]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="72D3D32DFFDDFFECFF75FED00DE2FF4C" bold="true" box="[151,752,151,178]" pageId="26" pageNumber="27">
<taxonomicName id="87A774BCFFDDFFECFF75FED00D6DFF4C" authority="(Gunther, 1872: 21)" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityPageNumber="21" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityPageNumber="21" baseAuthorityYear="1872" box="[151,639,151,178]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="australis" status="comb. nov.">
<emphasis id="72D3D32DFFDDFFECFF75FED00E98FF4F" bold="true" box="[151,394,151,177]" italics="true" pageId="26" pageNumber="27">Stegonotus australis</emphasis>
(
<bibRefCitation id="243672CEFFDDFFECFE78FEDF0D67FF4C" author="Gunther, A." box="[410,629,152,178]" pageId="26" pageNumber="27" pagination="13 - 37" refId="ref44061" refString="Gunther, A. (1872) Seventh account of new species of snakes in the collection of the British Museum. Annals and Magazine of Natural History, 9 (4), 13 - 37. https: // doi. org / 10.1080 / 002229372011951771" type="journal article" year="1872">Günther, 1872:21</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="69E06E56FFDDFFECFD64FEDF0DE2FF4C" box="[646,752,152,178]" pageId="26" pageNumber="27" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="40180F3FFFDDFFECFF75FE990A0FFD7D" blockId="26.[151,1437,222,967]" pageId="26" pageNumber="27">
<emphasis id="72D3D32DFFDDFFECFF75FE990E7CFF09" bold="true" box="[151,366,222,247]" pageId="26" pageNumber="27">Taxonomic status.</emphasis>
<taxonomicName id="87A774BCFFDDFFECFE94FE980D4BFF09" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[374,601,222,247]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="australis">
<emphasis id="72D3D32DFFDDFFECFE94FE980D4BFF09" box="[374,601,222,247]" italics="true" pageId="26" pageNumber="27">Stegonotus australis</emphasis>
</taxonomicName>
has heretofore been considered as a synonym of
<taxonomicName id="87A774BCFFDDFFECFB61FE980A01FF09" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[1155,1299,222,247]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="cucullatus">
<emphasis id="72D3D32DFFDDFFECFB61FE980A01FF09" box="[1155,1299,222,247]" italics="true" pageId="26" pageNumber="27">S. cucullatus</emphasis>
</taxonomicName>
. We use the following characteristics to differentiate this species from
<taxonomicName id="87A774BCFFDDFFECFCA9FF440CCDFEE5" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[843,991,258,283]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="cucullatus">
<emphasis id="72D3D32DFFDDFFECFCA9FF440CCDFEE5" box="[843,991,258,283]" italics="true" pageId="26" pageNumber="27">S. cucullatus</emphasis>
</taxonomicName>
(whose characteristics we provide in parentheses): (1) the length of the internasal suture is half of internasal width (the rostral invades the internasal area and nearly separates the internasals, with the length of the internasal suture only one tenth of internasal width); (2) the suture formed by the prefrontals and the frontal lies posterior to or is level with the anterior edge of the eyes (anterior to eyes); (3) the posterior edge of the prefrontals is bent forward laterally, forming an angle of 100° (hardly bent, 165°); (4) the line drawn through the two posterolateral vertices of the frontal lies well posterior to the eyes (intersects the eyes); (5) the chin area is separated from true ventral scales (gastrosteges) by three scales in the ventral midline (two scales); (6) nine supralabials (eight); and (7) ten infralabials (nine). Based on these differences, and on the straight-line distance of ca.
<quantity id="875FA2DAFFDDFFECFCFAFC640C96FDC5" box="[792,900,546,571]" metricMagnitude="6" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="27" unit="km" value="1800.0">1800 km</quantity>
between
<typeStatus id="9F1CB19DFFDDFFECFBE2FC640B22FDC5" box="[1024,1072,547,571]" pageId="26" pageNumber="27">type</typeStatus>
localities, we hereby remove
<taxonomicName id="87A774BCFFDDFFECFF75FC010E99FDA1" authorityName=", Gunther" authorityYear="1872" box="[151,395,582,607]" class="Reptilia" family="Colubridae" genus="Zamenophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="australis">
<emphasis id="72D3D32DFFDDFFECFF75FC010E99FDA1" box="[151,395,582,607]" italics="true" pageId="26" pageNumber="27">Zamenophis australis</emphasis>
</taxonomicName>
from the synonymy of
<taxonomicName id="87A774BCFFDDFFECFD7DFC000C23FDA1" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[671,817,582,607]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="cucullatus">
<emphasis id="72D3D32DFFDDFFECFD7DFC000C23FDA1" box="[671,817,582,607]" italics="true" pageId="26" pageNumber="27">S. cucullatus</emphasis>
</taxonomicName>
. This population therefore takes the name
<taxonomicName id="87A774BCFFDDFFECFAC0FC000FEBFD7D" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityYear="1872" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="australis" status="comb. nov.">
<emphasis id="72D3D32DFFDDFFECFAC0FC000FEBFD7D" italics="true" pageId="26" pageNumber="27">Stegonotus australis</emphasis>
</taxonomicName>
<emphasis id="72D3D32DFFDDFFECFEE2FC2D0E74FD7D" bold="true" box="[256,358,618,643]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="69E06E56FFDDFFECFEE2FC2D0E74FD7D" box="[256,358,618,643]" pageId="26" pageNumber="27" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
The species
<taxonomicName id="87A774BCFFDDFFECFE15FC2D0D4BFD7D" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[503,601,618,643]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="australis">
<emphasis id="72D3D32DFFDDFFECFE15FC2D0D4BFD7D" box="[503,601,618,643]" italics="true" pageId="26" pageNumber="27">australis</emphasis>
</taxonomicName>
is the
<typeStatus id="9F1CB19DFFDDFFECFD41FC2C0DC1FD7D" box="[675,723,619,643]" pageId="26" pageNumber="27">type</typeStatus>
species of the genus
<taxonomicName id="87A774BCFFDDFFECFC20FC2D0A0BFD7D" authority="Gunther, 1872: 21" authorityName="Gunther" authorityPageNumber="21" authorityYear="1872" box="[962,1305,618,643]" class="Reptilia" family="Colubridae" genus="Zamenophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFDDFFECFC20FC2D0B59FD7D" box="[962,1099,618,643]" italics="true" pageId="26" pageNumber="27">Zamenophis</emphasis>
<bibRefCitation id="243672CEFFDDFFECFBB0FC2D0BEAFD7D" author="Gunther, A." box="[1106,1272,618,643]" pageId="26" pageNumber="27" pagination="13 - 37" refId="ref44061" refString="Gunther, A. (1872) Seventh account of new species of snakes in the collection of the British Museum. Annals and Magazine of Natural History, 9 (4), 13 - 37. https: // doi. org / 10.1080 / 002229372011951771" type="journal article" year="1872">Günther, 1872</bibRefCitation>
:21
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFDDFFECFF25FCC90C29FC39" pageId="26" pageNumber="27" type="description">
<paragraph id="40180F3FFFDDFFECFF25FCC90BABFD59" blockId="26.[151,1437,222,967]" box="[199,1209,654,679]" pageId="26" pageNumber="27">
<emphasis id="72D3D32DFFDDFFECFF25FCC90E5BFD59" bold="true" box="[199,329,654,679]" pageId="26" pageNumber="27">Synonyms.</emphasis>
<taxonomicName id="87A774BCFFDDFFECFEB2FCC90DE0FD59" authority="Macleay, 1877" authorityName="Macleay" authorityYear="1877" box="[336,754,654,679]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="darnleyensis">
<emphasis id="72D3D32DFFDDFFECFEB2FCC90D51FD59" box="[336,579,654,679]" italics="true" pageId="26" pageNumber="27">Lycodon darnleyensis</emphasis>
<bibRefCitation id="243672CEFFDDFFECFDA9FCC90DE0FD59" author="Macleay, W." box="[587,754,654,679]" pageId="26" pageNumber="27" pagination="33 - 41" refId="ref45137" refString="Macleay, W. (1877) The ophidians of the Chevert Expedition. Proceedings of the Linnean Society of New South Wales, 2, 33 - 41." type="journal article" year="1877">Macleay, 1877</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="87A774BCFFDDFFECFD1CFCC90BA7FD59" authority="Macleay, 1884" authorityName="Macleay" authorityYear="1884" box="[766,1205,654,679]" class="Reptilia" family="Colubridae" genus="Herbertophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="plumbeus">
<emphasis id="72D3D32DFFDDFFECFD1CFCC90B14FD59" box="[766,1030,654,679]" italics="true" pageId="26" pageNumber="27">Herbertophis plumbeus</emphasis>
<bibRefCitation id="243672CEFFDDFFECFBEFFCC90BA7FD59" author="Macleay, W." box="[1037,1205,654,679]" pageId="26" pageNumber="27" pagination="432 - 436" refId="ref45167" refString="Macleay, W. (1884) Notes on some reptiles from the Herbert River, Queensland. Proceedings of the Linnean Society of New South Wales, 8, 432 - 436. https: // doi. org / 10.5962 / bhl. part. 28670" type="journal article" year="1884">Macleay, 1884</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="40180F3FFFDDFFECFF25FCF50C29FC39" blockId="26.[151,1437,222,967]" pageId="26" pageNumber="27">
<emphasis id="72D3D32DFFDDFFECFF25FCF50E6DFD35" bold="true" box="[199,383,690,715]" pageId="26" pageNumber="27">Original name.</emphasis>
<taxonomicName id="87A774BCFFDDFFECFE69FCF50C4AFD35" authority="Gunther, 1872: 21" authorityName=", Gunther" authorityPageNumber="21" authorityYear="1872" box="[395,856,690,715]" class="Reptilia" family="Colubridae" genus="Zamenophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="australis">
<emphasis id="72D3D32DFFDDFFECFE69FCF50D92FD35" box="[395,640,690,715]" italics="true" pageId="26" pageNumber="27">Zamenophis australis</emphasis>
<bibRefCitation id="243672CEFFDDFFECFD6EFCF50C24FD35" author="Gunther, A." box="[652,822,690,715]" pageId="26" pageNumber="27" pagination="13 - 37" refId="ref44061" refString="Gunther, A. (1872) Seventh account of new species of snakes in the collection of the British Museum. Annals and Magazine of Natural History, 9 (4), 13 - 37. https: // doi. org / 10.1080 / 002229372011951771" type="journal article" year="1872">Günther, 1872</bibRefCitation>
:21
</taxonomicName>
. The genus name references
<taxonomicName id="87A774BCFFDDFFECFB4DFCF30A1CFD35" authorityName=", Gunther" authorityYear="1872" box="[1199,1294,692,715]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFDDFFECFB4DFCF30A1CFD35" box="[1199,1294,692,715]" italics="true" pageId="26" pageNumber="27">Zamenis</emphasis>
</taxonomicName>
, a genus of colubrid snakes from Europe and the Middle East, which Günther considered to possess the same suite of characters as his new form. Given the well-defined distribution of
<taxonomicName id="87A774BCFFDDFFECFC70FCBB0BA2FCED" authority=", Gunther (1872)" authorityName=", Gunther" authorityYear="1872" box="[914,1200,762,787]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFDDFFECFC70FCBB0CE3FCED" box="[914,1009,764,787]" italics="true" pageId="26" pageNumber="27">Zamenis</emphasis>
,
<bibRefCitation id="243672CEFFDDFFECFBE3FCBD0BA2FCED" author="Gunther, A." box="[1025,1200,762,787]" pageId="26" pageNumber="27" pagination="13 - 37" refId="ref44061" refString="Gunther, A. (1872) Seventh account of new species of snakes in the collection of the British Museum. Annals and Magazine of Natural History, 9 (4), 13 - 37. https: // doi. org / 10.1080 / 002229372011951771" type="journal article" year="1872">Günther (1872)</bibRefCitation>
</taxonomicName>
decided to place his new Australian species into a separate genus, using “the entire anal shield” as the single defining characteristic. The genus name is a composite noun, including the genus name
<taxonomicName id="87A774BCFFDDFFECFC95FD030CC4FCA5" authorityName=", Gunther" authorityYear="1872" box="[887,982,836,859]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFDDFFECFC95FD030CC4FCA5" box="[887,982,836,859]" italics="true" pageId="26" pageNumber="27">Zamenis</emphasis>
</taxonomicName>
and the Greek
<emphasis id="72D3D32DFFDDFFECFB64FD050BD1FCA5" box="[1158,1219,834,859]" italics="true" pageId="26" pageNumber="27">ophis</emphasis>
, meaning “a snake like
<taxonomicName id="87A774BCFFDDFFECFF2EFD200E39FC80" authorityName=", Gunther" authorityYear="1872" box="[204,299,871,894]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFDDFFECFF2EFD200E39FC80" box="[204,299,871,894]" italics="true" pageId="26" pageNumber="27">Zamenis</emphasis>
</taxonomicName>
. The species epithet could reflect the collecting locality,
<collectingCountry id="38B04FAFFFDDFFECFC31FD210B2FFC81" box="[979,1085,870,895]" name="Australia" pageId="26" pageNumber="27">Australia</collectingCountry>
, or it may have been used in recognition that this was the species of
<taxonomicName id="87A774BCFFDDFFECFD63FDCC0DE9FC5C" authorityName=", McDowell" authorityYear="1972" box="[641,763,907,930]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFDDFFECFD63FDCC0DE9FC5C" box="[641,763,907,930]" italics="true" pageId="26" pageNumber="27">Stegonotus</emphasis>
</taxonomicName>
with the most southerly distribution (
<taxonomicName id="87A774BCFFDDFFECFB2EFDCD0A3CFC5D" authority="Latin" authorityName=", Gunther" authorityYear="1872" box="[1228,1326,906,931]" class="Reptilia" family="Colubridae" genus="Zamenophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="australis">
<emphasis id="72D3D32DFFDDFFECFB2EFDCD0A3CFC5D" box="[1228,1326,906,931]" italics="true" pageId="26" pageNumber="27">australis</emphasis>
</taxonomicName>
= Latin, southern). The species description was presented in English.
</paragraph>
</subSubSection>
<caption id="14D85FB7FFDDFFECFF75F9100C16F855" httpUri="https://zenodo.org/record/2607602/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,1879,1901]" targetBox="[151,1436,1012,1868]" targetPageId="26">
<paragraph id="40180F3FFFDDFFECFF75F9100C16F855" blockId="26.[151,1435,1879,1963]" pageId="26" pageNumber="27">
<emphasis id="72D3D32DFFDDFFECFF75F9100E33F892" bold="true" box="[151,289,1879,1901]" pageId="26" pageNumber="27">FIGURE 14.</emphasis>
Holotype of
<taxonomicName id="87A774BCFFDDFFECFE4EF91F0C3AF893" authority="(Gunther, 1872)" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[428,808,1879,1901]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="australis" status="comb. nov.">
<emphasis id="72D3D32DFFDDFFECFE4EF91F0D6EF893" box="[428,636,1879,1901]" italics="true" pageId="26" pageNumber="27">Stegonotus australis</emphasis>
(Günther, 1872)
</taxonomicName>
<taxonomicNameLabel id="69E06E56FFDDFFECFCD2F91F0CF1F893" box="[816,995,1879,1901]" pageId="26" pageNumber="27" rank="species">new combination</taxonomicNameLabel>
, BMNH 1946.1.14.93, an adult male from “Cape York,” Queensland, Australia. Shown are the entire specimen (TTL = 638 mm) in (A) dorsal and (B) ventral views, and its head in (C) dorsal, (D) right lateral, and (E) ventral views.
</paragraph>
</caption>
<subSubSection id="08BD5CB4FFDCFFEDFF25FED00EE8FD9E" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="40180F3FFFDCFFEDFF25FED00BF0FF4E" blockId="27.[151,1436,151,1054]" box="[199,1250,151,176]" pageId="27" pageNumber="28">
<materialsCitation id="F0CF0562FFDCFFEDFF25FED00BF0FF4E" ID-GBIF-Occurrence="2028140009" box="[199,1250,151,176]" collectionCode="BMNH" pageId="27" pageNumber="28" specimenCode="BMNH 1946.1, BMNH 68.6" specimenCount="1" typeStatus="holotype">
<emphasis id="72D3D32DFFDCFFEDFF25FED00E29FF4E" bold="true" box="[199,315,151,176]" pageId="27" pageNumber="28">
<typeStatus id="9F1CB19DFFDCFFEDFF25FED00E24FF4E" box="[199,310,151,176]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="1001A744FFDCFFEDFEA0FEDF0EFFFF4E" box="[322,493,151,176]" collectionCode="BMNH" pageId="27" pageNumber="28">BMNH 1946.1</specimenCode>
.14.93, formerly
<specimenCode id="1001A744FFDCFFEDFD45FEDF0C25FF4E" box="[679,823,151,176]" collectionCode="BMNH" pageId="27" pageNumber="28">BMNH 68.6</specimenCode>
.16.3 (
<figureCitation id="D89C13BAFFDCFFEDFC98FED00CDFFF4E" box="[890,973,151,176]" captionStart="FIGURE 14" captionStartId="26.[151,250,1879,1901]" captionTargetBox="[151,1436,1012,1868]" captionTargetId="figure@26.[151,1436,1012,1868]" captionTargetPageId="26" captionText="FIGURE 14. Holotype of Stegonotus australis (Günther, 1872) new combination, BMNH 1946.1.14.93, an adult male from “Cape York,” Queensland, Australia. Shown are the entire specimen (TTL = 638 mm) in (A) dorsal and (B) ventral views, and its head in (C) dorsal, (D) right lateral, and (E) ventral views." httpUri="https://zenodo.org/record/2607602/files/figure.png" pageId="27" pageNumber="28">Fig. 14</figureCitation>
;
<tableCitation id="0D253A84FFDCFFEDFC3BFED00B39FF4E" box="[985,1067,151,176]" captionStart="TABLE 1" captionStartId="60.[147,229,603,624]" captionText="TABLE 1. Name-bearing types of species in the genus Stegonotus. In this list we present basic data, including common measurements and selected scale counts, for all namebearing types (i.e., holotypes—HT, lectotypes—LT, neotypes—NT, syntypes—ST) to provide an easy overview of species. Additional details and the information for paratypes and paralectotypes can be found in the text. Character abbreviations are explained in the main text. For specimens where sex is indicated in red, we used the SCR value and not direct observation to estimate sex. Missing values are indicated by O. Specimen Type SCR SubC Species Synonym Sex SVL TL TTL V SC Dorsals SCE SL IL ILG Number Status (%) Condition LSUMZ admiraltiensis HT M 658 185 843 208 98 32 paired 17-19-15 4+5 8 10 10 93958 MSNG aruensis HT M 582 233 815 190 103 35 paired 17-17-15 3+4 7 8 4 30186 BMNH australis HT M 503 135 638 205 77 27 paired 17-17-15 4+5 8 9 5 1946.1.14.93 AMS australis darnleyensis HT F 720 O O 198 53+ O paired 17-17-15 4+5 9 10 5 R31894 AMS australis plumbeus HT M 1025 250 1275 220 74 25 paired 17-17-15 3+4+5 9 10 5 R31895 BMNH batjanensis HT M 1239 321 1560 235 86 27 paired 17-17-15 3+4+5 8 10 5 1946.1.11.36 FMNH borneensis HT F 783 221 1004 195 79 29 paired 17-17-15 4+5 10 11 5 164746 MNHN cucullatus HT M 769 254 1023 209 91 30 paired 17-17-15 4+5 8 9 5 3412 MTKD a cucullatus magnus ST F O O O 207 74 26 paired O-19-O 4+5 8 O O 573 ZMB b cucullatus magnus ST M 975 265 1240 211 87 29 paired 19-19-17 3+4 8 10 5 8794 MZB derooijae HT M 455 193 648 181 94 34 paired 17-17-15 3+4 7 8 8 Ophi.3288 MWNH diehli HT juv 196 56 252 179 82 31 paired 15-15-15 3+4 7 8 4 1244 NMW dorsalis HT M 805 245 1050 208 88 30 paired 15-15-15 4+5 8 10 5 14861 ZMA 9R florensis HT M 447 109 556 217 65 23 single 21-21-19 3+4+5 10 5 11080 8L BMNH guentheri LT F 683 165 848 192 64 25 paired 15-15-13 4+5 8 9 4 1946.1.11.38 ……continued on the next page" pageId="27" pageNumber="28">Table 1</tableCitation>
), an adult male.
</materialsCitation>
</paragraph>
<paragraph id="40180F3FFFDCFFEDFF25FEFB0D40FE9A" blockId="27.[151,1436,151,1054]" pageId="27" pageNumber="28">
<emphasis id="72D3D32DFFDCFFEDFF25FEFB0E7EFF2B" bold="true" box="[199,364,188,213]" pageId="27" pageNumber="28">Type locality.</emphasis>
“Cape York” [
<collectingRegion id="8263C1DDFFDCFFEDFDC5FEFB0DA3FF2B" box="[551,689,188,213]" country="Australia" name="Queensland" pageId="27" pageNumber="28">Queensland</collectingRegion>
,
<collectingCountry id="38B04FAFFFDCFFEDFD21FEFB0C22FF2B" box="[707,816,188,213]" name="Australia" pageId="27" pageNumber="28">Australia</collectingCountry>
]. This locality could either refer to the Cape York Peninsula, a large cone-shaped landmass that juts northwards from the northeastern side of the Australian continent to give its east coast a characteristic horn pointing towards New
<collectingCountry id="38B04FAFFFDCFFEDFC87FF430CABFEE3" box="[869,953,260,285]" name="Guinea-Bissau" pageId="27" pageNumber="28">Guinea</collectingCountry>
, or to Cape York, which is the name for the tip of the peninsula. This tropical peninsula was connected to New
<collectingCountry id="38B04FAFFFDCFFEDFC66FF600CC7FEBE" box="[900,981,295,320]" name="Guinea-Bissau" pageId="27" pageNumber="28">Guinea</collectingCountry>
across the Torres Strait for 40,000 of the last 2 million years (
<bibRefCitation id="243672CEFFDCFFEDFE99FF0B0D54FE9A" author="Harris, P. T. &amp; Heap, A. D. &amp; Marshall, J. F. &amp; McCulloch, M. T." box="[379,582,332,357]" pageId="27" pageNumber="28" pagination="85 - 97" refId="ref44182" refString="Harris, P. T., Heap, A. D., Marshall, J. F. &amp; McCulloch, M. T. (2008) A new coral reef province in the Gulf of Carpentaria, Australia: colonisation, growth and submergence during the early Holocene. Marine Geology, 251, 85 - 97. https: // doi. org / 10.1016 / j. margeo. 2008.02.010" type="journal article" year="2008">
Harris
<emphasis id="72D3D32DFFDCFFEDFE28FF0A0EE9FE9B" box="[458,507,332,357]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. 2008
</bibRefCitation>
).
</paragraph>
<paragraph id="40180F3FFFDCFFEDFF25FF280EE8FD9E" blockId="27.[151,1436,151,1054]" pageId="27" pageNumber="28">
<emphasis id="72D3D32DFFDCFFEDFF25FF280E55FE76" bold="true" box="[199,327,367,392]" pageId="27" pageNumber="28">Collection.</emphasis>
In his account, Günther provided a listing of “the new Species procured and described since
<date id="341929FFFFDCFFEDFA8BFF370FC3FE53" pageId="27" pageNumber="28" value="1868-06">June 1868</date>
” (
<bibRefCitation id="243672CEFFDCFFEDFF08FFD30EBCFE52" author="Gunther, A." box="[234,430,404,429]" pageId="27" pageNumber="28" pagination="13 - 37" refId="ref44061" refString="Gunther, A. (1872) Seventh account of new species of snakes in the collection of the British Museum. Annals and Magazine of Natural History, 9 (4), 13 - 37. https: // doi. org / 10.1080 / 002229372011951771" type="journal article" year="1872">Günther 1872:14</bibRefCitation>
), which included their provenance. Under the name
<taxonomicName id="87A774BCFFDCFFEDFC19FFD30A4AFE53" authority=", Gunther" authorityName=", Gunther" authorityYear="1872" box="[1019,1368,404,429]" class="Reptilia" family="Colubridae" genus="Zamenophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="australis">
<emphasis id="72D3D32DFFDCFFEDFC19FFD30BFFFE53" box="[1019,1261,404,429]" italics="true" pageId="27" pageNumber="28">Zamenophis australis</emphasis>
, Günther
</taxonomicName>
listed the specimen as having been purchased. Based on information on the original label, the
<typeStatus id="9F1CB19DFFDCFFEDFB6DFFF00BE3FE2E" box="[1167,1265,439,464]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
of
<taxonomicName id="87A774BCFFDCFFEDFAF8FFFE0A8EFE2E" authorityName=", Gunther" authorityYear="1872" box="[1306,1436,439,464]" class="Reptilia" family="Colubridae" genus="Zamenophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="australis">
<emphasis id="72D3D32DFFDCFFEDFAF8FFFE0A8EFE2E" box="[1306,1436,439,464]" italics="true" pageId="27" pageNumber="28">Z. australis</emphasis>
</taxonomicName>
was collected by a “Mr. Higgins,” but no further information is provided. Edmund Thomas Higgins (ca.
<date id="341929FFFFDCFFEDFAB4FF9B0FC2FDE6" pageId="27" pageNumber="28" value="1816" valueMax="1891">1816 1891</date>
) was a London-based dealer in natural history and geological specimens, who traded in Australian wildlife in the 1860s and visited
<collectingCountry id="38B04FAFFFDCFFEDFE77FC630EEEFDC3" box="[405,508,548,573]" name="Australia" pageId="27" pageNumber="28">Australia</collectingCountry>
himself twice (
<bibRefCitation id="243672CEFFDCFFEDFD52FC630C5EFDC2" author="Torrens, H. S." box="[688,844,548,572]" pageId="27" pageNumber="28" pagination="200 - 208" refId="ref47155" refString="Torrens, H. S. (1994) E. T. Higgins (c. 1816 - 1891): geological collector and natural history dealer, Bengal to England and then Australia (twice). In: Branagan, D. F. &amp; McNally, G. (Eds.), Useful and Curious Geological Enquiries Beyond the World. Conference Publications, 19 th International Symposium of the International Commission on the History of Geological Sciences, Sydney, Australia, pp. 200 - 208." type="proceedings paper" year="1994">Torrens 1994</bibRefCitation>
;
<bibRefCitation id="243672CEFFDCFFEDFCB8FC620B26FDC3" author="Horton, P. &amp; van Grouw, H. &amp; Johnstone, R. E." box="[858,1076,548,573]" pageId="27" pageNumber="28" pagination="129 - 138" refId="ref44468" refString="Horton, P., van Grouw, H. &amp; Johnstone, R. E. (2016) Clarifying collection details of specimens from Champion Bay, Western Australia, held in the Natural History Museum, Tring. Bulletin of the British Ornithologists Club, 136 (2), 129 - 138." type="journal article" year="2016">
Horton
<emphasis id="72D3D32DFFDCFFEDFC51FC620CF4FDC3" box="[947,998,548,573]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. 2016
</bibRefCitation>
). However, it is not possible to restrict the type locality further.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFDCFFEDFF25FC2B0CFBFC05" pageId="27" pageNumber="28" type="key">
<paragraph id="40180F3FFFDCFFEDFF25FC2B0A87FD55" blockId="27.[151,1436,151,1054]" pageId="27" pageNumber="28">
<emphasis id="72D3D32DFFDCFFEDFF25FC2B0D77FD7B" bold="true" box="[199,613,620,645]" pageId="27" pageNumber="28">
Key characteristics of the
<materialsCitation id="F0CF0562FFDCFFEDFE18FC2B0D77FD7B" ID-GBIF-Occurrence="2028140006" box="[506,613,620,645]" collectionCode="SVL" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
<typeStatus id="9F1CB19DFFDCFFEDFE18FC2B0D72FD7B" box="[506,608,620,645]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</emphasis>
470 (503) mm SVL + 140 (135) mm TL = 610 (638) mm TTL.
<materialsCitation id="F0CF0562FFDCFFEDFAD7FC2A0EA9FD55" ID-GBIF-Occurrence="2028140025" collectionCode="SCR" pageId="27" pageNumber="28" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
V
<subScript id="DC230D7AFFDCFFEDFAAAFC3D0A46FD76" attach="left" box="[1352,1364,634,648]" fontSize="6" pageId="27" pageNumber="28">
<specimenCount id="56A1C4B6FFDCFFEDFAAAFC3D0A46FD76" box="[1352,1364,634,648]" pageId="27" pageNumber="28" type="male"></specimenCount>
</subScript>
= 204 (205), SC
<subScript id="DC230D7AFFDCFFEDFEE3FCE60E1FFD51" attach="left" box="[257,269,673,687]" fontSize="6" pageId="27" pageNumber="28">
<specimenCount id="56A1C4B6FFDCFFEDFEE3FCE60E1FFD51" box="[257,269,673,687]" pageId="27" pageNumber="28" type="male"></specimenCount>
</subScript>
= 79 (77),
<collectionCode id="26B697FAFFDCFFEDFE65FCD40EA9FD55" box="[391,443,659,683]" country="USA" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography, University of California" pageId="27" pageNumber="28" type="Herbarium">SCR</collectionCode>
</materialsCitation>
<materialsCitation id="F0CF0562FFDCFFEDFE59FCE60B85FD54" ID-GBIF-Occurrence="2028140028" box="[443,1175,658,687]" collectionCode="IL" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<subScript id="DC230D7AFFDCFFEDFE59FCE60ED5FD51" attach="left" box="[443,455,673,687]" fontSize="6" pageId="27" pageNumber="28">
<specimenCount id="56A1C4B6FFDCFFEDFE59FCE60ED5FD51" box="[443,455,673,687]" pageId="27" pageNumber="28" type="male"></specimenCount>
</subScript>
= 0.28 (0.27), D = O (17-17-15), SL
<subScript id="DC230D7AFFDCFFEDFCBEFCE60C77FD51" attach="left" box="[860,869,673,687]" fontSize="6" pageId="27" pageNumber="28">E</subScript>
= 4+5 (4+5), SL = 9 (9),
<collectionCode id="26B697FAFFDCFFEDFB9FFCD40B85FD54" box="[1149,1175,659,682]" pageId="27" pageNumber="28">IL</collectionCode>
</materialsCitation>
= O (10), IL
<subScript id="DC230D7AFFDCFFEDFAC5FCE60A20FD51" attach="left" box="[1319,1330,673,687]" fontSize="6" pageId="27" pageNumber="28">G</subScript>
= O (5).
</paragraph>
<paragraph id="40180F3FFFDCFFEDFF25FCFE0CFBFC05" blockId="27.[151,1436,151,1054]" pageId="27" pageNumber="28">
<emphasis id="72D3D32DFFDCFFEDFF25FCFE0D77FD2C" bold="true" box="[199,613,697,722]" pageId="27" pageNumber="28">Key characteristics of the species.</emphasis>
In order to avoid any confusion with populations of
<taxonomicName id="87A774BCFFDCFFEDFB37FCFD0A5DFD2F" authorityName=", McDowell" authorityYear="1972" box="[1237,1359,698,721]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFDCFFEDFB37FCFD0A5DFD2F" box="[1237,1359,698,721]" italics="true" pageId="27" pageNumber="28">Stegonotus</emphasis>
</taxonomicName>
in the Northern Territories of
<collectingCountry id="38B04FAFFFDCFFEDFE79FC9A0D10FD08" box="[411,514,733,758]" name="Australia" pageId="27" pageNumber="28">Australia</collectingCountry>
(the subject of a separate study), we limit our discussion of the species
<taxonomicName id="87A774BCFFDCFFEDFAFFFC9A0A6DFD08" authorityName=", Gunther" authorityYear="1872" box="[1309,1407,733,758]" class="Reptilia" family="Colubridae" genus="Zamenophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="australis">
<emphasis id="72D3D32DFFDCFFEDFAFFFC9A0A6DFD08" box="[1309,1407,733,758]" italics="true" pageId="27" pageNumber="28">australis</emphasis>
</taxonomicName>
to
<collectingRegion id="8263C1DDFFDCFFEDFF75FD460E0CFCE4" box="[151,286,769,794]" country="Australia" name="Queensland" pageId="27" pageNumber="28">Queensland</collectingRegion>
, the state in which the
<typeStatus id="9F1CB19DFFDCFFEDFDC4FD450D44FCE4" box="[550,598,770,794]" pageId="27" pageNumber="28">type</typeStatus>
locality is located. We were able to examine
<specimenCount id="56A1C4B6FFDCFFEDFBBCFD460BE5FCE4" box="[1118,1271,769,794]" pageId="27" pageNumber="28" type="generic">35 specimens</specimenCount>
that, based on their collecting locality and morphological characteristics, could be members of
<taxonomicName id="87A774BCFFDCFFEDFBC7FD610BB6FCC0" authorityName=", Gunther" authorityYear="1872" box="[1061,1188,805,830]" class="Reptilia" family="Colubridae" genus="Zamenophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="australis">
<emphasis id="72D3D32DFFDCFFEDFBC7FD610BB6FCC0" box="[1061,1188,805,830]" italics="true" pageId="27" pageNumber="28">S. australis</emphasis>
</taxonomicName>
(
<specimenCount id="56A1C4B6FFDCFFEDFB56FD610A11FCC3" box="[1204,1283,806,830]" pageId="27" pageNumber="28" type="female">11 ♀♀</specimenCount>
,
<specimenCount id="56A1C4B6FFDCFFEDFAF3FD610A73FCC3" box="[1297,1377,806,830]" pageId="27" pageNumber="28" type="male">20 ♂♂</specimenCount>
, four of unknown sex). Characteristics include V
<subScript id="DC230D7AFFDCFFEDFD9FFD1F0D9BFC98" attach="left" box="[637,649,856,870]" fontSize="6" pageId="27" pageNumber="28">
<specimenCount id="56A1C4B6FFDCFFEDFD9FFD1F0D9BFC98" box="[637,649,856,870]" pageId="27" pageNumber="28" type="female"></specimenCount>
</subScript>
= 178215 (200 ± 11.2), V
<subScript id="DC230D7AFFDCFFEDFC5CFD1F0CD8FC98" attach="left" box="[958,970,856,870]" fontSize="6" pageId="27" pageNumber="28">
<specimenCount id="56A1C4B6FFDCFFEDFC5CFD1F0CD8FC98" box="[958,970,856,870]" pageId="27" pageNumber="28" type="male"></specimenCount>
</subScript>
= 191220 (211 ± 6.9); SC
<subScript id="DC230D7AFFDCFFEDFB1DFD1F0A19FC98" attach="left" box="[1279,1291,856,870]" fontSize="6" pageId="27" pageNumber="28">
<specimenCount id="56A1C4B6FFDCFFEDFB1DFD1F0A19FC98" box="[1279,1291,856,870]" pageId="27" pageNumber="28" type="female"></specimenCount>
</subScript>
= 6987 (78 ± 5.7), SC
<subScript id="DC230D7AFFDCFFEDFEE8FD390E04FC72" attach="left" box="[266,278,894,908]" fontSize="6" pageId="27" pageNumber="28">
<specimenCount id="56A1C4B6FFDCFFEDFEE8FD390E04FC72" box="[266,278,894,908]" pageId="27" pageNumber="28" type="male"></specimenCount>
</subScript>
= 7788 (83 ± 5.0); SCR
<subScript id="DC230D7AFFDCFFEDFDD1FD390D2DFC72" attach="left" box="[563,575,894,908]" fontSize="6" pageId="27" pageNumber="28">
<specimenCount id="56A1C4B6FFDCFFEDFDD1FD390D2DFC72" box="[563,575,894,908]" pageId="27" pageNumber="28" type="female"></specimenCount>
</subScript>
= 0.28, SCR
<subScript id="DC230D7AFFDCFFEDFD33FD390DCFFC72" attach="left" box="[721,733,894,908]" fontSize="6" pageId="27" pageNumber="28">
<specimenCount id="56A1C4B6FFDCFFEDFD33FD390DCFFC72" box="[721,733,894,908]" pageId="27" pageNumber="28" type="male"></specimenCount>
</subScript>
= 0.28; D = 17-17-15 (100%); SL
<subScript id="DC230D7AFFDCFFEDFBBFFD390B74FC72" attach="left" box="[1117,1126,894,908]" fontSize="6" pageId="27" pageNumber="28">E</subScript>
= 4+5 (100%); SL = 8 (n = 21, 57%) or 9 (n = 16, 43%); IL = 8 (n = 9, 24%), or 9 (n = 28, 76%); IL
<subScript id="DC230D7AFFDCFFEDFC28FDE20CC7FC4D" attach="left" box="[970,981,933,947]" fontSize="6" pageId="27" pageNumber="28">G</subScript>
= 5. We note that the standard deviation of the ventral count in females is quite high compared with other species of
<taxonomicName id="87A774BCFFDCFFEDFBE3FDF90B69FC2B" authorityName=", McDowell" authorityYear="1972" box="[1025,1147,958,981]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFDCFFEDFBE3FDF90B69FC2B" box="[1025,1147,958,981]" italics="true" pageId="27" pageNumber="28">Stegonotus</emphasis>
</taxonomicName>
, and we wonder whether this could indicate that two distinct species may be lumped into our sample.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFDCFFEDFF25FA420B7CFBE0" box="[199,1134,1029,1054]" pageId="27" pageNumber="28" type="discussion">
<paragraph id="40180F3FFFDCFFEDFF25FA420B7CFBE0" blockId="27.[151,1436,151,1054]" box="[199,1134,1029,1054]" pageId="27" pageNumber="28">
<emphasis id="72D3D32DFFDCFFEDFF25FA420E5DFBE0" bold="true" box="[199,335,1029,1054]" pageId="27" pageNumber="28">Comments.</emphasis>
The species was synonymized with
<taxonomicName id="87A774BCFFDCFFEDFD0AFA410C65FBE0" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[744,887,1029,1054]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="cucullatus">
<emphasis id="72D3D32DFFDCFFEDFD0AFA410C65FBE0" box="[744,887,1029,1054]" italics="true" pageId="27" pageNumber="28">S. cucullatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="243672CEFFDCFFEDFC43FA420B79FBE0" author="Boulenger, G. A." box="[929,1131,1029,1054]" pageId="27" pageNumber="28" refId="ref42396" refString="Boulenger, G. A. (1893) Catalogue of the snakes in the British Museum (Nat. Hist.) Folume I. Containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae, and Colubridae Aglyphae, part. Taylor &amp; Francis, London, United Kingdom, 448 pp." type="book" year="1893">Boulenger (1893)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>