295 lines
44 KiB
XML
295 lines
44 KiB
XML
<document id="C1F4411147D39D7B16E96BB9AF2E995F" ID-CLB-Dataset="276639" ID-DOI="10.1111/j.1096-3642.2011.00712.x" ID-GBIF-Dataset="d8b97c83-d7fe-493d-93e2-b5e6bac16d1f" ID-ISSN="0024-4082" ID-Zenodo-Dep="5441546" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630451329536" checkinUser="valdenar" docAuthor="Kellner, Alexander W. A., Campos, Diogenes A., Riff, Douglas & Andrade, Marco Brandalise De" docDate="2011" docId="B66F7B1BC84B6F7E8E164DFD8F24F9BB" docLanguage="en" docName="j.1096-3642.2011.00712.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (5)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00712.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Caryonosuchus pricei Kellner & Campos & Riff & Andrade 2011" docType="treatment" docVersion="6" lastPageNumber="645" masterDocId="4A560363C84A6F7A8D784D3B8E60FFE0" masterDocTitle="A new crocodylomorph (Sphagesauridae, Notosuchia) with horn-like tubercles from Brazil" masterLastPageNumber="645" masterPageNumber="644" pageNumber="644" updateTime="1699287824389" updateUser="ExternalLinkService">
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<mods:titleInfo id="0B03ABCD435EFD93431A99DFD72E3993">
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<mods:title id="F980B2194A32BC2A5F8B7936DA34A004">A new crocodylomorph (Sphagesauridae, Notosuchia) with horn-like tubercles from Brazil</mods:title>
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<mods:namePart id="33F30CC513EC926BCFC62160D0377B42">Kellner, Alexander W. A.</mods:namePart>
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<mods:namePart id="970479771BF162FF16CF81A46ED3B58F">Campos, Diogenes A.</mods:namePart>
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<mods:namePart id="9967343E5CFFB09C8F1CF233066848D0">Riff, Douglas</mods:namePart>
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<mods:namePart id="2E0A43ECA7B65B929BCDD4C57B4BACAB">Andrade, Marco Brandalise De</mods:namePart>
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<mods:typeOfResource id="EA6CFF06099A94840D7F8742A5C51F09">text</mods:typeOfResource>
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<mods:title id="B85CC63365ACDC7C5F0B484CE7FBD1CA">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="7B40C27FADFEA689AD4E7609E6AAC77F">2011</mods:date>
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<treatment id="B66F7B1BC84B6F7E8E164DFD8F24F9BB" ID-DOI="http://doi.org/10.5281/zenodo.5492069" ID-GBIF-Taxon="217238319" ID-Zenodo-Dep="5492069" LSID="urn:lsid:plazi:treatment:B66F7B1BC84B6F7E8E164DFD8F24F9BB" httpUri="http://treatment.plazi.org/id/B66F7B1BC84B6F7E8E164DFD8F24F9BB" lastPageId="4" pageId="1">
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<subSubSection id="76DC9986C84B6F7B8E164DFD8B0EFF3B" box="[878,1390,198,219]" pageId="1" type="nomenclature">
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<paragraph id="3E79CA0DC84B6F7B8E164DFD8B0EFF3B" blockId="1.[878,1390,198,219]" box="[878,1390,198,219]" pageId="1">
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<heading id="65317D61C84B6F7B8E164DFD8B0EFF3B" box="[878,1390,198,219]" centered="true" fontSize="9" level="2" pageId="1" reason="4">
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<emphasis id="0CB2161FC84B6F7B8E164DFD8B0EFF3B" bold="true" box="[878,1390,198,219]" pageId="1">
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<taxonomicName id="F9C6B18EC84B6F7B8E164DFD8AF5FF3A" authorityName="Kellner & Campos & Riff & Andrade" authorityYear="2011" box="[878,1173,198,219]" class="Reptilia" family="Sphagesauridae" genus="Caryonosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="1" phylum="Chordata" rank="species" species="pricei">
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<emphasis id="0CB2161FC84B6F7B8E164DFD8AF5FF3A" bold="true" box="[878,1173,198,219]" italics="true" pageId="1">CARYONOSUCHUS PRICEI</emphasis>
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</taxonomicName>
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GEN. ET SP. NOV.
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="76DC9986C84B6F7E8E424DD78F24F9BB" lastPageId="4" pageId="1" type="materials_examined">
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<paragraph id="3E79CA0DC84B6F7B8E424DD78A68FE9D" blockId="1.[826,1441,236,381]" pageId="1">
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<materialsCitation id="8EAEC050C84B6F7B8E424DD78A68FE9D" ID-GBIF-Occurrence="4435725301" collectionCode="R, UFRJ" county="Departamento Nacional da Producao Mineral" location="Anterior" municipality="Earth Science Museum" pageId="1" specimenCode="DGM 1411" specimenCount="1" typeStatus="holotype">
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<emphasis id="0CB2161FC84B6F7B8E424DD78DC4FEE1" box="[826,932,236,257]" italics="true" pageId="1">
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<typeStatus id="E17D74AFC84B6F7B8E424DD78DFEFEE1" box="[826,926,236,257]" pageId="1" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<location id="3B199CD6C84B6F7B8ED54DD78A6EFEE2" LSID="urn:lsid:plazi:treatment:B66F7B1BC84B6F7E8E164DFD8F24F9BB:3B199CD6C84B6F7B8ED54DD78A6EFEE2" box="[941,1038,236,258]" county="Departamento Nacional da Producao Mineral" municipality="Earth Science Museum" name="Anterior" pageId="1">Anterior</location>
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portion of the skull and lower jaw housed at the
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<collectingMunicipality id="DE1D5077C84B6F7B8E904C308A99FEC0" box="[1000,1273,267,289]" pageId="1">Earth Science Museum</collectingMunicipality>
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of the
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<collectingCounty id="D718B281C84B6F7B882B4C308B7BFEA0" pageId="1">Departamento Nacional da Produção Mineral</collectingCounty>
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under the number
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<specimenCode id="6E606276C84B6F7B8EE74C738A7EFEBD" box="[927,1054,328,349]" collectionCode="DGM" country="Brazil" httpUri="http://grbio.org/cool/k5hr-hisv" name="Divisao de Geologia c Mineralogia" pageId="1">DGM 1411</specimenCode>
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-
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<collectionCode id="58D752C8C84B6F7B895D4C738A58FEBD" box="[1061,1080,328,349]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1">R</collectionCode>
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(cast at the Museu Nacional/
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<collectionCode id="58D752C8C84B6F7B8E424C5C8D1FFE9C" box="[826,895,359,380]" country="Brazil" name="Universidade Federal Rural do Rio de Janeiro" pageId="1" type="Herbarium">UFRJ</collectionCode>
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) (
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<figureCitation id="A6FDD688C84B6F7B8EE04C5C8D9BFE9D" box="[920,1019,359,381]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="2.[144,223,1853,1872]" captionStartId-1="3.[164,243,1012,1031]" captionStartId-2="4.[144,223,760,779]" captionStartId-3="5.[164,243,1486,1505]" captionTargetBox-0="[303,1264,1030,1823]" captionTargetBox-1="[168,777,196,982]" captionTargetBox-2="[303,1263,196,730]" captionTargetBox-3="[323,1283,195,1456]" captionTargetId-0="figure-374@2.[303,1264,1030,1823]" captionTargetId-1="figure-632@3.[168,777,196,982]" captionTargetId-2="figure-481@4.[303,1263,196,730]" captionTargetId-3="figure-0@5.[323,1283,195,1456]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Figure 1. Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), rostrum in lateral (top), medial (middle), and ventral (bottom) views. Note that the right side of the specimen (A) is better preserved than the left side (B). Scale bar = 20 mm." captionText-1="Figure 2. Ventral view of the right (A) and left (B) sections of the upper jaw of Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), showing the oblique disposition of posterior crowns. Scale bar = 20 mm." captionText-2="Figure 3. Symphysis of Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), in ventral (A) and dorsal (B) views, in comparative view with the symphysis of Sphagesaurus huenei (RCL-100) (C). Note the concave area on the anterior section of the symphysis of C. pricei and its wedge-like profile in ventral view. Scale bars = 10 mm." captionText-3="Figure 4. Dentition of Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), in detail. A, ventrolateral view of teeth of the left side, showing the labial face of the crowns with basiapical ridges; B, lingual face of the same teeth, where the pebbled ornamentation and a wear facet are evident. C, apex of symphyseal crown showing tuberous denticles. D, maxillary tooth of the right side, showing the denticulated carina and wear facet. E, symphysis of Sphagesaurus huenei (RCL-100) with the first four mandibular pairs of teeth, the last showing extensive wear. F, palate of S. huenei (RCL-100) showing the premaxillary hypertrophied caniniforms and the first maxillary teeth. In comparison, RCL-100 shows the same general features, but its ridges seem to be more numerous when compared with the dentition of C. pricei. Note that in (A) that the basiapical ridges are evident even in areas where the enamel was not preserved and the dentine is exposed; in (F) the crowns of the fifth dentary teeth are preserved, attached to the palate. White pointers indicate wear facets. Scale bars = 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5441548" figureDoi-1="http://doi.org/10.5281/zenodo.5441550" figureDoi-2="http://doi.org/10.5281/zenodo.5441554" figureDoi-3="http://doi.org/10.5281/zenodo.5441556" httpUri-0="https://zenodo.org/record/5441548/files/figure.png" httpUri-1="https://zenodo.org/record/5441550/files/figure.png" httpUri-2="https://zenodo.org/record/5441554/files/figure.png" httpUri-3="https://zenodo.org/record/5441556/files/figure.png" pageId="1">Figs 1–4</figureCitation>
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).
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</materialsCitation>
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||
</paragraph>
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<paragraph id="3E79CA0DC84B6F7B8E424C8C8DE0FDA7" blockId="1.[826,1441,439,583]" pageId="1">
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<emphasis id="0CB2161FC84B6F7B8E424C8C8DD8FE2C" box="[826,952,439,460]" italics="true" pageId="1">Etymology:</emphasis>
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The specific name honours Llewellyn Ivor Price, a palaeontologist who described the genus
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<taxonomicName id="F9C6B18EC84B6F7B88134CED8DCDFDEA" authorityName="Price" authorityYear="1950" class="Reptilia" family="Sphagesauridae" genus="Sphagesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="1" phylum="Chordata" rank="genus">
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<emphasis id="0CB2161FC84B6F7B88134CED8DCDFDEA" italics="true" pageId="1">Sphagesaurus</emphasis>
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</taxonomicName>
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and also had a partial (unpublished) manuscript on the specimen
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<collectionCode id="58D752C8C84B6F7B892C4F288AF2FDC8" box="[1108,1170,531,552]" country="Brazil" httpUri="http://grbio.org/cool/k5hr-hisv" name="Divisao de Geologia c Mineralogia" pageId="1">DGM</collectionCode>
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1411-
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<collectionCode id="58D752C8C84B6F7B89A24F288A8DFDC8" box="[1242,1261,531,552]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1">R</collectionCode>
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||
(
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<bibRefCitation id="5A57B7FCC84B6F7B88794F288D14FDA7" author="Kellner AWA & Campos DA & Price LI" pageId="1" pagination="70 - 71" refId="ref5055" refString="Kellner AWA, Campos DA, Price LI. 1995. Material of Sphagesaurus (Sphagesauridae, Crocodilia) from the Late Cretaceous of Brazil. In: Kellner AWA, Viana CF, eds. XIV Congresso Brasileiro de Paleontologia. Rio de Janeiro: Atas, 70 - 71." type="book chapter" year="1995">
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Kellner
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<emphasis id="0CB2161FC84B6F7B881A4F2F8BF3FDC8" box="[1378,1427,531,553]" italics="true" pageId="1">et al</emphasis>
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||
., 1995
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</bibRefCitation>
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).
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</paragraph>
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<paragraph id="3E79CA0DC84B6F7B8E424FB98AB9FD56" blockId="1.[826,1441,642,694]" pageId="1">
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<emphasis id="0CB2161FC84B6F7B8E424FB98DB6FD77" box="[826,982,642,663]" italics="true" pageId="1">Type locality:</emphasis>
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Vicinity of Presidente Prudente City, south-west
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<collectingRegion id="FC0204EFC84B6F7B8EB84F9B8A1DFD55" box="[960,1149,672,694]" country="Brazil" name="Sao Paulo" pageId="1">São Paulo State</collectingRegion>
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,
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<collectingCountry id="46D18A9DC84B6F7B89F44F9B8AB5FD56" box="[1164,1237,672,694]" name="Brazil" pageId="1">Brazil</collectingCountry>
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.
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</paragraph>
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<paragraph id="3E79CA0DC84B6F7B8E424FCA8A9EFCA4" blockId="1.[826,1441,753,836]" pageId="1">
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<emphasis id="0CB2161FC84B6F7B8E424FCA8DB4FCE6" box="[826,980,753,774]" italics="true" pageId="1">Type horizon:</emphasis>
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<taxonomicName id="F9C6B18EC84B6F7B8EA74FCA8B28FCE6" authority="Formation, Bauru" authorityName="Formation, Bauru" box="[991,1352,753,775]" class="Chondrichthyes" family="Jalodontidae" genus="Adamantina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="1" phylum="Chordata" rank="genus">Adamantina Formation, Bauru</taxonomicName>
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Group; Campanian to Maastrichtian (
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<bibRefCitation id="5A57B7FCC84B6F7B89EE4E348D93FCA3" author="Gobbo-Rodrigues SR & Petri S & Bertini RJ" pageId="1" pagination="3 - 13" refId="ref4887" refString="Gobbo-Rodrigues SR, Petri S, Bertini RJ. 1999. Ocorrencias de ostracodes na Formacao Adamantina do Grupo Bauru, Cretaceo Superior da Bacia do Parana e possibilidades de correlacao com depositos isocronos argentinos. Parte I - Familia Ilyocyprididae. Acta Geologica Leopoldensia 23: 3 - 13." type="journal article" year="1999">Gobbo-Rodrigues, Petri & Bertini, 1999</bibRefCitation>
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||
,
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<bibRefCitation id="5A57B7FCC84B6F7B89794E158A93FCA4" author="Batezelli A & Saad AR & Etchebehere MLC & Perinotto JAJ & Fulfaro VJ" box="[1025,1267,814,836]" pageId="1" pagination="5 - 19" refId="ref4469" refString="Batezelli A, Saad AR, Etchebehere MLC, Perinotto JAJ, Fulfaro VJ. 2003. Analise estratigrafica aplicada a Formacao Aracatuba (Grupo Bauru - Ks) no Centro-Oeste do Estado de Sao Paulo. Geociencias 22: 5 - 19." type="journal article" year="2003">
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||
Batezelli
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<emphasis id="0CB2161FC84B6F7B89084E158AC0FCA3" box="[1136,1184,814,835]" italics="true" pageId="1">et al</emphasis>
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||
., 2003
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||
</bibRefCitation>
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||
).
|
||
</paragraph>
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<paragraph id="3E79CA0DC84B6F7B8E424E448B49FB26" blockId="1.[826,1441,894,1222]" pageId="1">
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<emphasis id="0CB2161FC84B6F7B8E424E448DD1FC74" box="[826,945,895,916]" italics="true" pageId="1">Diagnosis:</emphasis>
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Sphagesaurid that can be separated from all other members of this clade based on the following combination of characters (autapomorphies are indicated with an asterisk): rostrum with horn-like tubercles on the premaxilla* and on the maxilla*; rostrum showing rough ornamentation with grooves and bony ridges*; first tooth posterior to the premaxillary hypercaniniform is supported exclusively by the premaxilla; palatine ramus of maxilla covered by low crests and shallow wrinkles; shallow fossa present at symphysis, medial to the first three teeth.
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</paragraph>
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<paragraph id="3E79CA0DC84B6F7B8E42483B8DA7FA4F" blockId="1.[826,1441,1280,1455]" pageId="1">
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<emphasis id="0CB2161FC84B6F7B8E42483B8DC6FAF5" box="[826,934,1280,1301]" italics="true" pageId="1">Remarks:</emphasis>
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<bibRefCitation id="5A57B7FCC84B6F7B8ED5483B8AB1FAF6" author="Kellner AWA & Campos DA" box="[941,1233,1280,1302]" pageId="1" pagination="238 - 251" refId="ref5032" refString="Kellner AWA, Campos DA. 1999. Vertebrate paleontology in Brazil - a review. Episodes 22: 238 - 251." type="journal article" year="1999">Kellner & Campos (1999)</bibRefCitation>
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||
briefly mentioned the specimen
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<collectionCode id="58D752C8C84B6F7B8EAF48248A75FAD4" box="[983,1045,1311,1332]" country="Brazil" httpUri="http://grbio.org/cool/k5hr-hisv" name="Divisao de Geologia c Mineralogia" pageId="1">DGM</collectionCode>
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1411-
|
||
<collectionCode id="58D752C8C84B6F7B892F48248A0AFAD4" box="[1111,1130,1311,1332]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1">R</collectionCode>
|
||
and tentatively assigned it to
|
||
<taxonomicName id="F9C6B18EC84B6F7B8E2348068A49FAB3" box="[859,1065,1341,1363]" class="Reptilia" family="Sphagesauridae" genus="Sphagesaurus" kingdom="Animalia" order="Crocodylia" pageId="1" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="0CB2161FC84B6F7B8E2348068D9CFAB2" box="[859,1020,1341,1362]" italics="true" pageId="1">Sphagesaurus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, but in the light of new sphagesaurid material recently described in the literature, we regard this specimen as representing a new genus and species.
|
||
</paragraph>
|
||
<paragraph id="3E79CA0DC84B6F7B8E4248D18DDBFA1F" blockId="1.[826,1441,1514,1903]" box="[826,955,1514,1535]" pageId="1">
|
||
<emphasis id="0CB2161FC84B6F7B8E4248D18DDBFA1F" box="[826,955,1514,1535]" italics="true" pageId="1">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="3E79CA0DC84B6F7B8E424B338D8DF88E" blockId="1.[826,1441,1514,1903]" pageId="1">
|
||
<materialsCitation id="8EAEC050C84B6F7B8E424B338D89F88E" ID-GBIF-Occurrence="4435725302" collectionCode="R" pageId="1" specimenCode="DGM 1411" specimenCount="1" typeStatus="holotype">
|
||
The
|
||
<typeStatus id="E17D74AFC84B6F7B8E084B338DB1F9FE" box="[880,977,1544,1566]" pageId="1" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="F9C6B18EC84B6F7B8E814B338A8FF9FE" authorityName="Kellner & Campos & Riff & Andrade" authorityYear="2011" box="[1017,1263,1544,1566]" class="Reptilia" family="Sphagesauridae" genus="Caryonosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="1" phylum="Chordata" rank="species" species="pricei">
|
||
<emphasis id="0CB2161FC84B6F7B8E814B338A8FF9FE" box="[1017,1263,1544,1566]" italics="true" pageId="1">Caryonosuchus pricei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<specimenCode id="6E606276C84B6F7B88794B338B1EF9FE" box="[1281,1406,1544,1566]" collectionCode="DGM" country="Brazil" httpUri="http://grbio.org/cool/k5hr-hisv" name="Divisao de Geologia c Mineralogia" pageId="1">DGM 1411</specimenCode>
|
||
-
|
||
<collectionCode id="58D752C8C84B6F7B88FE4B338BF9F9FE" box="[1414,1433,1544,1566]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1">R</collectionCode>
|
||
) consists of the anterior portion of the rostrum, whose bone surface is extremely well preserved (
|
||
<figureCitation id="A6FDD688C84B6F7B88654B7E8B3DF9BC" box="[1309,1373,1605,1628]" captionStart="Figure 1" captionStartId="2.[144,223,1853,1872]" captionTargetBox="[303,1264,1030,1823]" captionTargetId="figure-374@2.[303,1264,1030,1823]" captionTargetPageId="2" captionText="Figure 1. Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), rostrum in lateral (top), medial (middle), and ventral (bottom) views. Note that the right side of the specimen (A) is better preserved than the left side (B). Scale bar = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5441548" httpUri="https://zenodo.org/record/5441548/files/figure.png" pageId="1">Fig. 1</figureCitation>
|
||
). The upper jaw shows both premaxillae and maxillae, with opposite elements unfused. The nasal is missing, exposing the internal region of the external naris. The tip of the dentary that was originally preserved in close contact with the upper jaw was separated during mechanical preparation. The upper teeth are in their natural position whereas several teeth of the lower jaw were displaced anteriorly during the fossilization process
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3E79CA0DC8486F788DE84DFD8EB6FD68" blockId="2.[144,759,198,954]" pageId="2">
|
||
<emphasis id="0CB2161FC8486F788DE84DFD8F74FF3B" box="[144,276,198,219]" italics="true" pageId="2">Premaxilla:</emphasis>
|
||
The right premaxilla is more complete than the left and includes the palatine ramus (
|
||
<figureCitation id="A6FDD688C8486F788DE04C388EB8FEF9" box="[152,216,259,281]" captionStart="Figure 1" captionStartId="2.[144,223,1853,1872]" captionTargetBox="[303,1264,1030,1823]" captionTargetId="figure-374@2.[303,1264,1030,1823]" captionTargetPageId="2" captionText="Figure 1. Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), rostrum in lateral (top), medial (middle), and ventral (bottom) views. Note that the right side of the specimen (A) is better preserved than the left side (B). Scale bar = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5441548" httpUri="https://zenodo.org/record/5441548/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
||
). This bone has an anterodorsal flange partially covering the external naris that forms a lateral notch. This flange is strongly ornamented, as is the dorsal part of the premaxilla. Two well-developed and ornamented horn-like tubercles are present. On the area of the tubercles, the ornamentation is composed of semicircular well-delimited deep pits and differs from the remaining part of the rostrum. Dorsally, near the suture with the nasal, the pits become shallower, less defined, and elongated, turning into an area covered by elongated sulci and crests that are orientated diagonally in respect to the sagittal plane.
|
||
</paragraph>
|
||
<paragraph id="3E79CA0DC8486F788DD04FAA8C30FCE3" blockId="2.[144,759,198,954]" pageId="2">Part of the internal surface of the right nasal cavity is preserved, more extensively than its left counterpart. Although the bone surface is smooth, there is a small area that is lightly ornamented.</paragraph>
|
||
<paragraph id="3E79CA0DC8486F788DD04E308F9BFC5A" blockId="2.[144,759,198,954]" pageId="2">The alveolar margins lack ornamentation. The left premaxilla shows one neurovascular foramen located laterally, in an area above the middle point between the first and the second premaxillary teeth. A second and smaller foramen is located posterodorsally, close to the suture with the maxilla.</paragraph>
|
||
<paragraph id="3E79CA0DC8486F788E474DFD8A04FEF9" blockId="2.[806,1421,198,281]" pageId="2">The right palatine ramus of the premaxilla also shows light ornamentation, with a small foramen medial to the second tooth.</paragraph>
|
||
<paragraph id="3E79CA0DC8486F788E5E4C7B8DB8FDEE" blockId="2.[806,1421,320,526]" pageId="2">
|
||
<emphasis id="0CB2161FC8486F788E5E4C7B8A4FFEB5" box="[806,1071,320,341]" italics="true" pageId="2">Premaxillary dentition:</emphasis>
|
||
Two teeth are present in both premaxillae, all incomplete, showing oblique implantation as a result of paramesial rotation (
|
||
<figureCitation id="A6FDD688C8486F7888714C468B2CFE74" box="[1289,1356,381,404]" captionStart="Figure 2" captionStartId="3.[164,243,1012,1031]" captionTargetBox="[168,777,196,982]" captionTargetId="figure-632@3.[168,777,196,982]" captionTargetPageId="3" captionText="Figure 2. Ventral view of the right (A) and left (B) sections of the upper jaw of Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), showing the oblique disposition of posterior crowns. Scale bar = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5441550" httpUri="https://zenodo.org/record/5441550/files/figure.png" pageId="2">Fig. 2</figureCitation>
|
||
). The first tooth on the left side shows a partial crown and the root, and most probably was a hypertrophied caniniform. The second tooth is entirely supported by the premaxilla.
|
||
</paragraph>
|
||
<paragraph id="3E79CA0DC8486F788E5E4F0E8B3BFC9D" blockId="2.[806,1421,565,954]" pageId="2">
|
||
<emphasis id="0CB2161FC8486F788E5E4F0E8DE5FDAA" box="[806,901,565,586]" italics="true" pageId="2">Maxilla:</emphasis>
|
||
Both maxillae are incomplete, with the right one better preserved than the left (
|
||
<figureCitation id="A6FDD688C8486F7889BA4F6F8B63FD8A" box="[1218,1283,596,618]" captionStart="Figure 1" captionStartId="2.[144,223,1853,1872]" captionTargetBox="[303,1264,1030,1823]" captionTargetId="figure-374@2.[303,1264,1030,1823]" captionTargetPageId="2" captionText="Figure 1. Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), rostrum in lateral (top), medial (middle), and ventral (bottom) views. Note that the right side of the specimen (A) is better preserved than the left side (B). Scale bar = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5441548" httpUri="https://zenodo.org/record/5441548/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
||
). This bone partially overlies the premaxilla, a common feature in notosuchids (including other sphagesaurids). As in the premaxilla, the maxilla has the dorsal side heavily ornamented and the lateral side smooth, particularly close to the alveolar margin. At least two tubercles can be identified, which are smaller than the ones found in the premaxilla. The first is horizontally aligned with the two first ones whereas the second is located dorsally and medially in respect to those.
|
||
</paragraph>
|
||
<paragraph id="3E79CA0DC8486F798E474EBD8D6BFA3D" blockId="2.[806,1421,565,954]" lastBlockId="3.[164,779,1173,1501]" lastPageId="3" pageId="2">On the left maxilla there are five neurovascular foramina positioned above the alveolar margin, forming an anterodorsally arched line. The two anterior-most foramina are located dorsal to the first maxillary tooth and the following are dorsal to the second maxillary tooth. On the right maxilla, the neurovascular foramina are arranged in a more irregular way. The palatine ramus is laterodorsally displaced, and has light but evident ornamentation composed of low crests and shallow wrinkles. Another series of six neurovascular foramina can be identified next to the teeth, on the right maxilla. These foramina are located medially to the teeth, in couples.</paragraph>
|
||
<caption id="6AB99A85C8486F788DE84A068BEEF88E" ID-DOI="http://doi.org/10.5281/zenodo.5441548" ID-Zenodo-Dep="5441548" httpUri="https://zenodo.org/record/5441548/files/figure.png" pageId="2" startId="2.[144,223,1853,1872]" targetBox="[303,1264,1030,1823]" targetPageId="2">
|
||
<paragraph id="3E79CA0DC8486F788DE84A068BEEF88E" blockId="2.[144,1422,1853,1902]" pageId="2">
|
||
<emphasis id="0CB2161FC8486F788DE84A068C61F8B0" bold="true" box="[144,513,1853,1872]" pageId="2">
|
||
Figure 1.
|
||
<taxonomicName id="F9C6B18EC8486F788C7C4A068C61F8B0" authority="Kellner & Campos & Riff & Andrade, 2011" authorityName="Kellner & Campos & Riff & Andrade" authorityYear="2011" box="[260,513,1853,1872]" class="Reptilia" family="Sphagesauridae" genus="Caryonosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="2" phylum="Chordata" rank="species" species="pricei" status="gen. et sp. nov.">
|
||
<emphasis id="0CB2161FC8486F788C7C4A068C61F8B0" bold="true" box="[260,513,1853,1872]" italics="true" pageId="2">Caryonosuchus pricei</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="1781AB64C8486F788F704A068CD2F8B0" box="[520,690,1853,1873]" pageId="2" rank="species">
|
||
<emphasis id="0CB2161FC8486F788F704A068C59F8B0" bold="true" box="[520,569,1853,1872]" pageId="2">gen.</emphasis>
|
||
et
|
||
<emphasis id="0CB2161FC8486F788F234A068CD2F8B0" bold="true" box="[603,690,1853,1872]" pageId="2">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(DGM 1411-R), rostrum in lateral (top), medial (middle), and ventral (bottom) views. Note that the right side of the specimen (A) is better preserved than the left side (B). Scale bar = 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6AB99A85C8496F798DDC4ECF8C4EFB80" ID-DOI="http://doi.org/10.5281/zenodo.5441550" ID-Zenodo-Dep="5441550" httpUri="https://zenodo.org/record/5441550/files/figure.png" pageId="3" startId="3.[164,243,1012,1031]" targetBox="[168,777,196,982]" targetPageId="3">
|
||
<paragraph id="3E79CA0DC8496F798DDC4ECF8C4EFB80" blockId="3.[164,779,1012,1120]" pageId="3">
|
||
<emphasis id="0CB2161FC8496F798DDC4ECF8F6CFBE7" bold="true" box="[164,268,1012,1031]" pageId="3">Figure 2.</emphasis>
|
||
Ventral view of the right (A) and left (B) sections of the upper jaw of
|
||
<taxonomicName id="F9C6B18EC8496F798CCE492A8CD4FBC5" authority="Kellner & Campos & Riff & Andrade, 2011" authorityName="Kellner & Campos & Riff & Andrade" authorityYear="2011" box="[438,692,1041,1061]" class="Reptilia" family="Sphagesauridae" genus="Caryonosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="3" phylum="Chordata" rank="species" species="pricei" status="gen. et sp. nov.">Caryonosuchus pricei</taxonomicName>
|
||
<taxonomicNameLabel id="1781AB64C8496F798FC549298E9BFBA2" pageId="3" rank="species">
|
||
<emphasis id="0CB2161FC8496F798FC549298C8EFBC5" bold="true" box="[701,750,1042,1061]" pageId="3">gen.</emphasis>
|
||
et
|
||
<emphasis id="0CB2161FC8496F798DDC49148E9BFBA2" bold="true" box="[164,251,1071,1090]" pageId="3">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(DGM 1411-R), showing the oblique disposition of posterior crowns. Scale bar = 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3E79CA0DC8496F798DDC4B338D6BF936" blockId="3.[164,779,1544,1903]" pageId="3">
|
||
<emphasis id="0CB2161FC8496F798DDC4B338FEAF9FD" box="[164,394,1544,1565]" italics="true" pageId="3">Maxillary dentition:</emphasis>
|
||
There are three teeth preserved in each maxilla, but the third on the left side was damaged during the fossilization process (
|
||
<figureCitation id="A6FDD688C8496F798FE94B7E8CB4F9BC" box="[657,724,1605,1628]" captionStart="Figure 2" captionStartId="3.[164,243,1012,1031]" captionTargetBox="[168,777,196,982]" captionTargetId="figure-632@3.[168,777,196,982]" captionTargetPageId="3" captionText="Figure 2. Ventral view of the right (A) and left (B) sections of the upper jaw of Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), showing the oblique disposition of posterior crowns. Scale bar = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5441550" httpUri="https://zenodo.org/record/5441550/files/figure.png" pageId="3">Fig. 2</figureCitation>
|
||
). All are obliquely implanted with each tooth having the posterolingual portion of the crown ending in a carina and the anteriolabial margin rounded, resulting in a teardrop cross-section. None of the roots are exposed.
|
||
</paragraph>
|
||
<paragraph id="3E79CA0DC8496F798DC44BE58A95FE0F" blockId="3.[164,779,1544,1903]" lastBlockId="3.[826,1442,198,1321]" pageId="3">The first maxillary tooth on the left side is better preserved than the one on the right side. The apex and part of the lingual portion are not complete. On the labial face, seven undivided ridges that reach the apex of the tooth are present. Smaller ridges, restricted to the base of the crown can also be observed. On the lingual face there are remnants of only five ridges in the mesial-most section, whereas the remaining surface is destroyed. On both faces, the ridges are smooth, but the enamel is clearly ornamented in a pebbled pattern. Part of the root is dorsally exposed, showing an elliptic cross-section and oblique orientation, supporting the idea that this tooth has undergone paramesial rotation rather than a simple rearrangement of the crown.</paragraph>
|
||
<paragraph id="3E79CA0DC8496F798E2B4CC38AC8FB91" blockId="3.[826,1442,198,1321]" pageId="3">
|
||
Part of the enamel on the second maxillary tooth of the left side has been lost, exposing the dentine. The folded pattern shows that the ridges observed on the teeth express a structural characteristic of the crown, rather than a superficial ornamentation restricted to the enamel. As in the preceding tooth, seven main ridges are present in the labial face and five occur on the lingual side, again complemented by smaller ridges that do not reach the apex. The distance between ridges increases from the posterolingual to the anterolabial edges of the crown. The apex has a rounded shape. A wear surface is present, extending from the apex to the mid-length of the carina. No denticles or pebbled ornamentation can be identified on this surface. This worn down surface is neither a preservation artefact nor created during the collecting or preparation processes, but was produced
|
||
<emphasis id="0CB2161FC8496F7988564ED98B1AFC17" box="[1326,1402,994,1015]" italics="true" pageId="3">in vivo</emphasis>
|
||
, as the animal processed food. A similar wear surface is observed on the second maxillary tooth of the right side. This tooth has up to nine basiapical ridges on the labial surface of the crown.
|
||
</paragraph>
|
||
<paragraph id="3E79CA0DC8496F798E2B49408DA3FAC9" blockId="3.[826,1442,198,1321]" pageId="3">On the third maxillary tooth of the right side, six labial and up to three lingual ridges can be identified. This crown shows a posterior carina, with an undulated basal section that lacks denticles, and an apical section that displays three denticles. No wear surface is observed.</paragraph>
|
||
<paragraph id="3E79CA0DC8496F798E42485B8DD0F969" blockId="3.[826,1441,1376,1673]" pageId="3">
|
||
<emphasis id="0CB2161FC8496F798E42485B8A29FA95" box="[826,1097,1376,1397]" italics="true" pageId="3">Mandibular symphysis:</emphasis>
|
||
Only the anterior end of the mandibular symphysis is preserved (
|
||
<figureCitation id="A6FDD688C8496F79886948458B34FA75" box="[1297,1364,1406,1429]" captionStart="Figure 3" captionStartId="4.[144,223,760,779]" captionTargetBox="[303,1263,196,730]" captionTargetId="figure-481@4.[303,1263,196,730]" captionTargetPageId="4" captionText="Figure 3. Symphysis of Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), in ventral (A) and dorsal (B) views, in comparative view with the symphysis of Sphagesaurus huenei (RCL-100) (C). Note the concave area on the anterior section of the symphysis of C. pricei and its wedge-like profile in ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5441554" httpUri="https://zenodo.org/record/5441554/files/figure.png" pageId="3">Fig. 3</figureCitation>
|
||
). The ventral surface is eroded, suggesting that this portion of the specimen was exposed before collection. The anterior-most section is narrow and widens posterior to the third tooth. A concavity is present close to the anterior tip of the symphysis, positioned medial to the first three teeth. It is shallow, longer than wide, and unknown in other sphagesaurid specimens (including RCL 100).
|
||
</paragraph>
|
||
<paragraph id="3E79CA0DC8496F7E8E424BFB8E92FC17" blockId="3.[826,1442,1728,1903]" lastBlockId="4.[144,759,901,1627]" lastPageId="4" pageId="3">
|
||
<emphasis id="0CB2161FC8496F798E424BFB8A5AF935" box="[826,1082,1728,1749]" italics="true" pageId="3">Mandibular dentition:</emphasis>
|
||
The right side of the symphysis holds three conical teeth and part of one crown (
|
||
<figureCitation id="A6FDD688C8496F798E3A4BC68DE9F8F2" box="[834,905,1789,1811]" captionStart="Figure 3" captionStartId="4.[144,223,760,779]" captionTargetBox="[303,1263,196,730]" captionTargetId="figure-481@4.[303,1263,196,730]" captionTargetPageId="4" captionText="Figure 3. Symphysis of Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), in ventral (A) and dorsal (B) views, in comparative view with the symphysis of Sphagesaurus huenei (RCL-100) (C). Note the concave area on the anterior section of the symphysis of C. pricei and its wedge-like profile in ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5441554" httpUri="https://zenodo.org/record/5441554/files/figure.png" pageId="3">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="A6FDD688C8496F798EE14BC68DC9F8F3" box="[921,937,1789,1811]" captionStart="Figure 4" captionStartId="5.[164,243,1486,1505]" captionTargetBox="[323,1283,195,1456]" captionTargetId="figure-0@5.[323,1283,195,1456]" captionTargetPageId="5" captionText="Figure 4. Dentition of Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), in detail. A, ventrolateral view of teeth of the left side, showing the labial face of the crowns with basiapical ridges; B, lingual face of the same teeth, where the pebbled ornamentation and a wear facet are evident. C, apex of symphyseal crown showing tuberous denticles. D, maxillary tooth of the right side, showing the denticulated carina and wear facet. E, symphysis of Sphagesaurus huenei (RCL-100) with the first four mandibular pairs of teeth, the last showing extensive wear. F, palate of S. huenei (RCL-100) showing the premaxillary hypertrophied caniniforms and the first maxillary teeth. In comparison, RCL-100 shows the same general features, but its ridges seem to be more numerous when compared with the dentition of C. pricei. Note that in (A) that the basiapical ridges are evident even in areas where the enamel was not preserved and the dentine is exposed; in (F) the crowns of the fifth dentary teeth are preserved, attached to the palate. White pointers indicate wear facets. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5441556" httpUri="https://zenodo.org/record/5441556/files/figure.png" pageId="3">4</figureCitation>
|
||
). The first three pairs are mostly procumbent, as in other sphagesaurids and notosuchids. They are very inclined, which is attributed to the taphonomic compression suffered by the specimen during the fossilization process. All teeth bear isomorphic, smooth, and nonserrated basiapical ridges and an apical wear facet, which is small and inclined.
|
||
</paragraph>
|
||
<caption id="6AB99A85C84E6F7E8DE84FC38AD9FCA6" ID-DOI="http://doi.org/10.5281/zenodo.5441554" ID-Zenodo-Dep="5441554" httpUri="https://zenodo.org/record/5441554/files/figure.png" pageId="4" startId="4.[144,223,760,779]" targetBox="[303,1263,196,730]" targetPageId="4">
|
||
<paragraph id="3E79CA0DC84E6F7E8DE84FC38AD9FCA6" blockId="4.[144,1421,759,838]" pageId="4">
|
||
<emphasis id="0CB2161FC84E6F7E8DE84FC38E98FCEB" bold="true" box="[144,248,760,779]" pageId="4">Figure 3.</emphasis>
|
||
Symphysis of
|
||
<taxonomicName id="F9C6B18EC84E6F7E8CE14FCC8CF6FCEB" authority="Kellner & Campos & Riff & Andrade, 2011" authorityName="Kellner & Campos & Riff & Andrade" authorityYear="2011" box="[409,662,759,779]" class="Reptilia" family="Sphagesauridae" genus="Caryonosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" phylum="Chordata" rank="species" species="pricei" status="gen. et sp. nov.">Caryonosuchus pricei</taxonomicName>
|
||
<taxonomicNameLabel id="1781AB64C84E6F7E8FE64FC38D2AFCEB" box="[670,842,760,779]" pageId="4" rank="species">
|
||
<emphasis id="0CB2161FC84E6F7E8FE64FC38CAEFCEB" bold="true" box="[670,718,760,779]" pageId="4">gen.</emphasis>
|
||
et
|
||
<emphasis id="0CB2161FC84E6F7E8F8A4FC38D2AFCEB" bold="true" box="[754,842,760,779]" pageId="4">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(DGM 1411-R), in ventral (A) and dorsal (B) views, in comparative view with the symphysis of
|
||
<taxonomicName id="F9C6B18EC84E6F7E8F2B4E2E8D57FCC8" baseAuthorityName="Pol" baseAuthorityYear="2003" box="[595,823,789,808]" class="Reptilia" family="Sphagesauridae" genus="Sphagesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" phylum="Chordata" rank="species" species="huenei">
|
||
<emphasis id="0CB2161FC84E6F7E8F2B4E2E8D57FCC8" box="[595,823,789,808]" italics="true" pageId="4">Sphagesaurus huenei</emphasis>
|
||
</taxonomicName>
|
||
(RCL-100) (C). Note the concave area on the anterior section of the symphysis of
|
||
<taxonomicName id="F9C6B18EC84E6F7E8CC34E088C72FCA5" authorityName="Kellner & Campos & Riff & Andrade" authorityYear="2011" box="[443,530,819,838]" class="Reptilia" family="Sphagesauridae" genus="Caryonosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" phylum="Chordata" rank="species" species="pricei">
|
||
<emphasis id="0CB2161FC84E6F7E8CC34E088C72FCA5" box="[443,530,819,838]" italics="true" pageId="4">C. pricei</emphasis>
|
||
</taxonomicName>
|
||
and its wedge-like profile in ventral view. Scale bars = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3E79CA0DC84E6F7E8DD04EC48F24F9BB" blockId="4.[144,759,901,1627]" pageId="4">
|
||
Only the apical section of the fourth tooth of the right side is preserved, displaced from its original anatomical position (
|
||
<figureCitation id="A6FDD688C84E6F7E8C0649078FB4FBB3" box="[382,468,1084,1107]" captionStart="Figure 4" captionStartId="5.[164,243,1486,1505]" captionTargetBox="[323,1283,195,1456]" captionTargetId="figure-0@5.[323,1283,195,1456]" captionTargetPageId="5" captionText="Figure 4. Dentition of Caryonosuchus pricei gen. et sp. nov. (DGM 1411-R), in detail. A, ventrolateral view of teeth of the left side, showing the labial face of the crowns with basiapical ridges; B, lingual face of the same teeth, where the pebbled ornamentation and a wear facet are evident. C, apex of symphyseal crown showing tuberous denticles. D, maxillary tooth of the right side, showing the denticulated carina and wear facet. E, symphysis of Sphagesaurus huenei (RCL-100) with the first four mandibular pairs of teeth, the last showing extensive wear. F, palate of S. huenei (RCL-100) showing the premaxillary hypertrophied caniniforms and the first maxillary teeth. In comparison, RCL-100 shows the same general features, but its ridges seem to be more numerous when compared with the dentition of C. pricei. Note that in (A) that the basiapical ridges are evident even in areas where the enamel was not preserved and the dentine is exposed; in (F) the crowns of the fifth dentary teeth are preserved, attached to the palate. White pointers indicate wear facets. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5441556" httpUri="https://zenodo.org/record/5441556/files/figure.png" pageId="4">Fig. 4C</figureCitation>
|
||
). The apex is clearly compressed, consistent with the reversed teardrop morphology typical of sphagesaurids. Additionally, this tooth bears at least three lateral ridges and a carina. Although the medial and basal portions of the carina are not preserved, it is possible to identify three conspicuous denticles in the preserved section. These are proportionally large when compared with the denticles of other ziphosuchian crocodylians, and correspond to what
|
||
<bibRefCitation id="5A57B7FCC84E6F7E8C27486B8C0FFA86" author="Prasad GVR & Broin FL" box="[351,623,1360,1382]" pageId="4" pagination="19 - 71" refId="ref5442" refString="Prasad GVR, Broin FL. 2002. Late Cretaceous crocodile remains from Naskal (India): comparisons and biogeographic affinities. Annales de Paleontologie 88: 19 - 71." type="journal article" year="2002">Prasad & Broin (2002)</bibRefCitation>
|
||
classify as true denticles. However, these denticles are tuberous and do not correspond to the typical cuneiform morphology found in theropodomorph dentitions of highly predacious crocodylomorphs (e.g.
|
||
<taxonomicName id="F9C6B18EC84E6F7E8FF348F08C65FA1E" authority=", Stratiotosuchus, Sebecus" authorityName="Stratiotosuchus, Sebecus" class="Reptilia" family="Baurusuchidae" genus="Baurusuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="0CB2161FC84E6F7E8FF348F08C65FA1E" italics="true" pageId="4">Baurusuchus, Stratiotosuchus, Sebecus</emphasis>
|
||
</taxonomicName>
|
||
). They are rather similar to the denticles of
|
||
<taxonomicName id="F9C6B18EC84E6F7E8CC64B3C8C03F9FC" authorityName="Carvalho & Bertini" authorityYear="1999" box="[446,611,1543,1564]" class="Reptilia" family="Candidodontidae" genus="Mariliasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="0CB2161FC84E6F7E8CC64B3C8C03F9FC" box="[446,611,1543,1564]" italics="true" pageId="4">Mariliasuchus</emphasis>
|
||
</taxonomicName>
|
||
, corresponding to the ziphomorph pattern
|
||
<emphasis id="0CB2161FC84E6F7E8F584B1C8C3FF9DC" box="[544,607,1575,1596]" italics="true" pageId="4">sensu</emphasis>
|
||
<bibRefCitation id="5A57B7FCC84E6F7E8F174B1D8F20F9BB" author="Andrade MB & Bertini RJ" pageId="4" pagination="63 - 82" refId="ref4359" refString="Andrade MB, Bertini RJ. 2008 c. Morphology of the dental carinae in Mariliasuchus amarali (Crocodylomorpha, Notosuchia) and the pattern of tooth serration among basal Mesoeucrocodylia. Arquivos do Museu Nacional 66: 63 - 82." type="journal article" year="2008">Andrade & Bertini (2008c)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |