323 lines
39 KiB
XML
323 lines
39 KiB
XML
<document id="5CB83C499A8DE4E6FF1FA06F48CE552F" ID-CLB-Dataset="46687" ID-DOI="10.11646/zootaxa.3754.2.2" ID-GBIF-Dataset="92e56954-0445-4d78-b7f4-d54e656c0256" ID-ISSN="1175-5326" ID-Zenodo-Dep="226765" ID-ZooBank="AB51EB22-A9EA-432A-A84D-ED9A5B995BBF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460367787120" checkinUser="plazi" docAuthor="Aarvik, Leif & Agassiz, David J. L." docDate="2014" docId="497E87D53A6C3F5C3AF9FD0AFAFFACFB" docLanguage="en" docName="zt03754p132.pdf" docOrigin="Zootaxa 3754 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neaspasia Diakonoff 1989" docType="treatment" docVersion="12" lastPageNumber="119" masterDocId="B547FFAD3A6D3F5E3A6EFFBEFFA2AD34" masterDocTitle="Revision of African Neaspasia Diakonoff, 1989 and the related Conaspasia, n. gen. (Lepidoptera, Tortricidae)" masterLastPageNumber="132" masterPageNumber="117" pageNumber="118" updateTime="1698295405665" updateUser="ExternalLinkService">
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<mods:title id="8E6C9E4A67E3A6E480935A63F2665EEC">Revision of African Neaspasia Diakonoff, 1989 and the related Conaspasia, n. gen. (Lepidoptera, Tortricidae)</mods:title>
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<mods:namePart id="19CD2329253E7F05853F47940D62975A">Aarvik, Leif</mods:namePart>
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<mods:affiliation id="F86838AC83B6AE0CC75B95CCF5287DE8">Natural History Museum, University of Oslo, P. O. Box 1172 Blindern, NO- 0318 Oslo, Norway.</mods:affiliation>
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<mods:nameIdentifier id="3803F954F8882A35EF1CA57D5AC7126B" type="email">leif.aarvik@nhm.uio.no</mods:nameIdentifier>
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<mods:namePart id="0FE8BAB68A6015BD92F0A1A4ACA7BA7F">Agassiz, David J. L.</mods:namePart>
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<mods:affiliation id="A800707718CFB55452B10F1A9DEDF060">Dept. of Entomology, Natural History Museum, London SW 7 5 BD, United Kingdom.</mods:affiliation>
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<mods:nameIdentifier id="C6901F2E12B6024B5D6E426CC52CC3B5" type="email">D.Agassiz@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="CC1F0F478D43228AAF029B90813F3EB0">Zootaxa</mods:title>
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<mods:part id="893370C2801DB6AE5949700D71B839C7">
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<mods:date id="42FF10C0CAE008AC71204EEDFB399D89">2014</mods:date>
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<treatment id="497E87D53A6C3F5C3AF9FD0AFAFFACFB" ID-DOI="http://doi.org/10.5281/zenodo.6136196" ID-GBIF-Taxon="119411122" ID-Zenodo-Dep="6136196" LSID="urn:lsid:plazi:treatment:497E87D53A6C3F5C3AF9FD0AFAFFACFB" httpUri="http://treatment.plazi.org/id/497E87D53A6C3F5C3AF9FD0AFAFFACFB" lastPageId="2" lastPageNumber="119" pageId="1" pageNumber="118">
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<subSubSection id="89CD65483A6C3F5F3AF9FD0AFD87AFF9" box="[151,549,692,717]" pageId="1" pageNumber="118" type="nomenclature">
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<paragraph id="C16836C33A6C3F5F3AF9FD0AFD87AFF9" blockId="1.[151,549,692,717]" box="[151,549,692,717]" pageId="1" pageNumber="118">
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<heading id="9A2081AF3A6C3F5F3AF9FD0AFD87AFF9" bold="true" box="[151,549,692,717]" fontSize="10" level="2" pageId="1" pageNumber="118" reason="2">
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<emphasis id="F3A3EAD13A6C3F5F3AF9FD0AFD87AFF9" bold="true" box="[151,549,692,717]" pageId="1" pageNumber="118">
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Genus
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<taxonomicName id="06D74D403A6C3F5F3A84FD0AFD87AFF9" ID-CoL="94NR2" authority="Diakonoff, 1989" authorityName="Diakonoff" authorityYear="1989" box="[234,549,692,717]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
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<emphasis id="F3A3EAD13A6C3F5F3A84FD0AFEC2AFF8" bold="true" box="[234,352,692,716]" italics="true" pageId="1" pageNumber="118">Neaspasia</emphasis>
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<bibRefCitation id="A5464B323A6C3F5F3B09FD0AFD87AFF9" author="Diakonoff" box="[359,549,692,717]" pageId="1" pageNumber="118" refString="Diakonoff, A. (1989) Tortricidae from Madagascar. Part 2. Olethreutinae, 6 (Lepidoptera). Annales de la Societe Entomologique de France (new series), 25 (4), 431 - 460." type="journal article" year="1989">Diakonoff, 1989</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="89CD65483A6C3F5F3AF9FD43FB13AE04" pageId="1" pageNumber="118" type="reference_group">
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<paragraph id="C16836C33A6C3F5F3AF9FD43FBF0AE26" blockId="1.[151,1201,764,817]" box="[151,1106,764,786]" pageId="1" pageNumber="118">
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<treatmentCitationGroup id="E1C711ED3A6C3F5F3AF9FD43FBF0AE26" box="[151,1106,764,786]" pageId="1" pageNumber="118">
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<taxonomicName id="06D74D403A6C3F5F3AF9FD43FE46AE26" ID-CoL="94NR2" authority="Diakonoff, 1989: 445" authorityName="Diakonoff" authorityPageNumber="445" authorityYear="1989" box="[151,484,764,786]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
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<emphasis id="F3A3EAD13A6C3F5F3AF9FD43FEA3AE26" box="[151,257,765,786]" italics="true" pageId="1" pageNumber="118">Neaspasia</emphasis>
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<treatmentCitation id="407610D23A6C3F5F3B69FD42FE46AE26" author="Diakonoff" box="[263,484,764,786]" page="445" pageId="1" pageNumber="118" year="1989">
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<bibRefCitation id="A5464B323A6C3F5F3B69FD42FE13AE26" author="Diakonoff" box="[263,433,764,786]" pageId="1" pageNumber="118" refString="Diakonoff, A. (1989) Tortricidae from Madagascar. Part 2. Olethreutinae, 6 (Lepidoptera). Annales de la Societe Entomologique de France (new series), 25 (4), 431 - 460." type="journal article" year="1989">Diakonoff, 1989</bibRefCitation>
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: 445
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||
</treatmentCitation>
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</taxonomicName>
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.
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<typeStatus id="1E6C88613A6C3F5F3B83FD43FDBDAE26" box="[493,543,765,786]" pageId="1" pageNumber="118">Type</typeStatus>
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species:
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<taxonomicName id="06D74D403A6C3F5F3812FD43FBECAE26" ID-CoL="94V7Z" authority="Diakonoff, 1989: 445" authorityName="Diakonoff" authorityPageNumber="445" authorityYear="1989" box="[636,1102,764,786]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="loxochlamys">
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<emphasis id="F3A3EAD13A6C3F5F3812FD43FCC9AE26" box="[636,875,764,786]" italics="true" pageId="1" pageNumber="118">Neaspasia loxochlamys</emphasis>
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<treatmentCitation id="407610D23A6C3F5F391FFD42FBECAE26" author="Diakonoff" box="[881,1102,764,786]" page="445" pageId="1" pageNumber="118" year="1989">
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<bibRefCitation id="A5464B323A6C3F5F391FFD42FBB9AE26" author="Diakonoff" box="[881,1051,764,786]" pageId="1" pageNumber="118" refString="Diakonoff, A. (1989) Tortricidae from Madagascar. Part 2. Olethreutinae, 6 (Lepidoptera). Annales de la Societe Entomologique de France (new series), 25 (4), 431 - 460." type="journal article" year="1989">Diakonoff, 1989</bibRefCitation>
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: 445
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="C16836C33A6C3F5F3AF9FCA4FB13AE04" blockId="1.[151,1201,764,817]" box="[151,1201,794,817]" pageId="1" pageNumber="118">
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<emphasis id="F3A3EAD13A6C3F5F3AF9FCA4FEB9AE04" box="[151,283,794,816]" italics="true" pageId="1" pageNumber="118">Genetancylis</emphasis>
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<bibRefCitation id="A5464B323A6C3F5F3B4FFCA4FE6BAE04" author="Razowski" box="[289,457,794,816]" pageId="1" pageNumber="118" refString="Razowski, J. (1995) Some Tortricidae from the Arabian Peninsula, with description of one new genus and four new species (Insecta: Lepidoptera). SHILAP Revista de lepidopterologia, 23, 129 - 139." type="journal article" year="1995">Razowski, 1995</bibRefCitation>
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: 134.
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<typeStatus id="1E6C88613A6C3F5F386AFCA5FD94AE04" box="[516,566,795,816]" pageId="1" pageNumber="118">Type</typeStatus>
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species:
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<emphasis id="F3A3EAD13A6C3F5F38FDFCA4FCB5AE04" box="[659,791,794,816]" italics="true" pageId="1" pageNumber="118">Genetancylis</emphasis>
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<taxonomicName id="06D74D403A6C3F5F3973FCA4FBFFAE04" ID-CoL="B3XPX" authority="Razowski, 1995: 134" authorityName="Razowski" authorityPageNumber="134" authorityYear="1995" box="[797,1117,794,816]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="homalota">
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<emphasis id="F3A3EAD13A6C3F5F3973FCA4FCDFAE04" box="[797,893,794,816]" italics="true" pageId="1" pageNumber="118">homalota</emphasis>
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<bibRefCitation id="A5464B323A6C3F5F39EAFCA4FB89AE04" author="Razowski" box="[900,1067,794,816]" pageId="1" pageNumber="118" refString="Razowski, J. (1995) Some Tortricidae from the Arabian Peninsula, with description of one new genus and four new species (Insecta: Lepidoptera). SHILAP Revista de lepidopterologia, 23, 129 - 139." type="journal article" year="1995">Razowski, 1995</bibRefCitation>
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: 134
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</taxonomicName>
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.
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<emphasis id="F3A3EAD13A6C3F5F3E09FCA4FB13AE04" bold="true" box="[1127,1201,794,817]" pageId="1" pageNumber="118">Syn. n.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="89CD65483A6C3F5F3AF9FCE3FC43A879" pageId="1" pageNumber="118" type="diagnosis">
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<paragraph id="C16836C33A6C3F5F3AF9FCE3FC43A879" blockId="1.[151,1436,861,2005]" pageId="1" pageNumber="118">
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<emphasis id="F3A3EAD13A6C3F5F3AF9FCE3FEB3AE42" bold="true" box="[151,273,861,886]" pageId="1" pageNumber="118">Diagnosis.</emphasis>
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Most species of
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<taxonomicName id="06D74D403A6C3F5F3BB4FCE0FDEDAE41" box="[474,591,862,885]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
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<emphasis id="F3A3EAD13A6C3F5F3BB4FCE0FDEDAE41" box="[474,591,862,885]" italics="true" pageId="1" pageNumber="118">Neaspasia</emphasis>
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</taxonomicName>
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share a characteristic forewing pattern consisting of a dark, more or less blackish basal area and the rest of the wing surface with a white or beige ground colour, dark suffusion is usually present along the termen. A similar wing pattern is found in Holarctic species of
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<taxonomicName id="06D74D403A6C3F5F3E1CFC1BFAC5AE8A" authority="Hubner, 1825" authorityName="Hubner" authorityYear="1825" box="[1138,1383,933,958]" class="Insecta" family="Tortricidae" genus="Hedya" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
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<emphasis id="F3A3EAD13A6C3F5F3E1CFC1BFB19AE8A" box="[1138,1211,933,958]" italics="true" pageId="1" pageNumber="118">Hedya</emphasis>
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Hübner, 1825
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</taxonomicName>
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and
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<taxonomicName id="06D74D403A6C3F5F3AF9FC74FE0AAED6" authority="Hubner, 1825" authorityName="Hubner" authorityYear="1825" box="[151,424,969,994]" class="Insecta" family="Tortricidae" genus="Apotomis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
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<emphasis id="F3A3EAD13A6C3F5F3AF9FC74FEA3AED5" box="[151,257,970,993]" italics="true" pageId="1" pageNumber="118">Apotomis</emphasis>
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Hübner, 1825
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</taxonomicName>
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. In the forewing all veins are present and separate; in the hindwing M2, M3 and Cu1 are trifurcate with their bases close together. The male hindwing has an anal roll. In the hind tibia of males there is a dorsal groove concealing a long hairpencil. Tergum
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<quantity id="062F9B263A6C3F5F388FFBAAFCAFA919" box="[737,781,1044,1069]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="118" unit="in" value="2.0">2 in</quantity>
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both sexes has a pair of subdorsal pits (
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<figureCitation id="59EC2A463A6C3F5F3EA0FBAAFAB2A918" box="[1230,1296,1044,1069]" captionStart="FIGURE 1" captionStartId="2.[151,250,1198,1220]" captionTargetBox="[343,1241,522,1167]" captionTargetId="figure@2.[338,1248,513,1173]" captionTargetPageId="2" captionText="FIGURE 1. Anterior part of male abdomen of Neaspasia orthacta (Meyrick) showing pair of circular depressions on tergum 2 (slide NHMO 2247). Scale 1 mm." httpUri="https://zenodo.org/record/226766/files/figure.png" pageId="1" pageNumber="118">Fig. 1</figureCitation>
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); on tergum 3 there are similar, but indistinct depressions, in some cases not discernible. The male genitalia are of a generalized Olethreutinae structure, with well demarcated sacculus and cucullus of the valva, and a setose uncus and socii. The sacculus has a group of spines between the caudal edge of the sacculus and basal excavation. The phallus lacks cornuti, but has internal sclerotised folds. In the female genitalia the ostium is surrounded by a large sclerite, and the sterigma is species specific in shape. The ductus bursae is tubular, sclerotised posteriorly, widened and membranous before its junction with the corpus bursae. The ductus seminalis arises closer to corpus bursae than to the ostium; the ductus bursae has a sclerite where the ductus seminalis arises. The shape of the paired signa is unique, each a curved, oval plate with a large, rounded opening at the base.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="89CD65483A6C3F5C3AA9FAE6FAFFACFB" lastPageId="2" lastPageNumber="119" pageId="1" pageNumber="118" type="discussion">
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<paragraph id="C16836C33A6C3F5F3AA9FAE6FEE2AA14" blockId="1.[151,1436,861,2005]" pageId="1" pageNumber="118">
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<emphasis id="F3A3EAD13A6C3F5F3AA9FAE6FE14A845" bold="true" box="[199,438,1368,1393]" pageId="1" pageNumber="118">Systematic position.</emphasis>
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The hindwing venation, the presence of an anal roll, and modifications of the hind tibia in males indicate a position within
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<taxonomicName id="06D74D403A6C3F5F384DFAC2FD0CA8A1" box="[547,686,1404,1429]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="tribe" tribe="Olethreutini">Olethreutini</taxonomicName>
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. Abdominal depressions or ‘dorsal pits’ as found in
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<taxonomicName id="06D74D403A6C3F5F3E93FAC3FAD0A8A0" box="[1277,1394,1405,1428]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
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<emphasis id="F3A3EAD13A6C3F5F3E93FAC3FAD0A8A0" box="[1277,1394,1405,1428]" italics="true" pageId="1" pageNumber="118">Neaspasia</emphasis>
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</taxonomicName>
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are present in several genera of
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<taxonomicName id="06D74D403A6C3F5F3B87FA1EFDC5A88D" box="[489,615,1440,1465]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="family">Tortricidae</taxonomicName>
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.
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<bibRefCitation id="A5464B323A6C3F5F381BFA1EFCB0A88D" author="Horak" box="[629,786,1440,1465]" pageId="1" pageNumber="118" refString="Horak, M. (2006) Olethreutine moths of Australia (Lepidoptera: Tortricidae). Monographs on Australian Lepidoptera. Vol. 10. CSIRO Publishing, Canberra, 522 pp." type="book" year="2006">Horak (2006)</bibRefCitation>
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recognized them as present in the following genera of
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<taxonomicName id="06D74D403A6C3F5F3AF9FA7AFE81A8E9" box="[151,291,1476,1501]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="tribe" tribe="Olethreutini">Olethreutini</taxonomicName>
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:
|
||
<taxonomicName id="06D74D403A6C3F5F3B5CFA7AFDFFA8E8" authority="Lower, 1901" authorityName="Lower" authorityYear="1901" box="[306,605,1476,1501]" class="Insecta" family="Tortricidae" genus="Temnolopha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3B5CFA7AFE1EA8E9" box="[306,444,1476,1501]" italics="true" pageId="1" pageNumber="118">Temnolopha</emphasis>
|
||
Lower, 1901
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="06D74D403A6C3F5F3800FA7AFC4EA8E9" authority="Diakonoff, 1966" authorityName="Diakonoff" authorityYear="1966" box="[622,1004,1476,1501]" class="Insecta" family="Tortricidae" genus="Gatesclarkeana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3800FA7AFC82A8E9" box="[622,800,1476,1501]" italics="true" pageId="1" pageNumber="118">Gatesclarkeana</emphasis>
|
||
Diakonoff, 1966
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="06D74D403A6C3F5F3992FA7AFAC5A8E9" authority="Diakonoff, 1966" authorityName="Diakonoff" authorityYear="1966" box="[1020,1383,1476,1501]" class="Insecta" family="Tortricidae" genus="Ophiorrhabda" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3992FA7AFB3FA8E9" box="[1020,1181,1476,1501]" italics="true" pageId="1" pageNumber="118">Ophiorrhabda</emphasis>
|
||
Diakonoff, 1966
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="06D74D403A6C3F5F3AF9FA57FE75AB34" authority="Diakonoff, 1984" authorityName="Diakonoff" authorityYear="1984" box="[151,471,1512,1537]" class="Insecta" family="Tortricidae" genus="Zomariana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3AF9FA57FEB6AB34" box="[151,276,1513,1536]" italics="true" pageId="1" pageNumber="118">Zomariana</emphasis>
|
||
Diakonoff, 1984
|
||
</taxonomicName>
|
||
. Dorsal pits are also present in several tribes of
|
||
<taxonomicName id="06D74D403A6C3F5F399EFA56FBCCAB35" box="[1008,1134,1512,1537]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="subFamily" subFamily="Tortricinae">Tortricinae</taxonomicName>
|
||
,
|
||
<emphasis id="F3A3EAD13A6C3F5F3E17FA57FB3AAB34" box="[1145,1176,1513,1536]" italics="true" pageId="1" pageNumber="118">viz</emphasis>
|
||
.
|
||
<taxonomicName id="06D74D403A6C3F5F3EC8FA56FAB6AB35" box="[1190,1300,1512,1537]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="tribe" tribe="Archipini">Archipini</taxonomicName>
|
||
,
|
||
<taxonomicName id="06D74D403A6C3F5F3F4EFA56FACEAB35" box="[1312,1388,1512,1537]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="tribe" tribe="Euliini">Euliini</taxonomicName>
|
||
and
|
||
<taxonomicName id="06D74D403A6C3F5F3AF9F9B2FE96AB11" box="[151,308,1548,1573]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="tribe" tribe="Sparganothini">Sparganothini</taxonomicName>
|
||
(
|
||
<bibRefCitation id="A5464B323A6C3F5F3B2DF9B2FDE1AB10" author="Brown" box="[323,579,1548,1573]" pageId="1" pageNumber="118" refString="Brown, J. W. & Miller, S. E. (1999) A new species of Coelostathma Clemens (Lepidoptera: Tortricidae) from Cocos Island, Costa Rica, with comments on the phylogenetic significance of abdominal dorsal pits in Sparganothini. Proceedings of the Entomological Society of Washington, 101, 701 - 707." type="journal article" year="1999">Brown & Miller 1999</bibRefCitation>
|
||
). The four genera mentioned by
|
||
<bibRefCitation id="A5464B323A6C3F5F39D7F9B2FBF0AB11" author="Horak" box="[953,1106,1548,1573]" pageId="1" pageNumber="118" refString="Horak, M. (2006) Olethreutine moths of Australia (Lepidoptera: Tortricidae). Monographs on Australian Lepidoptera. Vol. 10. CSIRO Publishing, Canberra, 522 pp." type="book" year="2006">Horak (2006)</bibRefCitation>
|
||
all belong to different genus groups within
|
||
<taxonomicName id="06D74D403A6C3F5F3B53F98EFE6AAB7D" box="[317,456,1584,1609]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="tribe" tribe="Olethreutini">Olethreutini</taxonomicName>
|
||
. The female genitalia of
|
||
<taxonomicName id="06D74D403A6C3F5F3889F98EFCD3AB7D" box="[743,881,1584,1609]" class="Insecta" family="Tortricidae" genus="Temnolopha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3889F98EFCD3AB7D" box="[743,881,1584,1609]" italics="true" pageId="1" pageNumber="118">Temnolopha</emphasis>
|
||
</taxonomicName>
|
||
shows similarities with
|
||
<taxonomicName id="06D74D403A6C3F5F3EE7F98FFB5FAB7C" box="[1161,1277,1585,1608]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3EE7F98FFB5FAB7C" box="[1161,1277,1585,1608]" italics="true" pageId="1" pageNumber="118">Neaspasia</emphasis>
|
||
</taxonomicName>
|
||
, especially in the ductus bursae which in both genera is a posteriorly sclerotised tube, with a sclerite in the widened anterior part at the junction of the ductus seminalis (compare figure of
|
||
<taxonomicName id="06D74D403A6C3F5F3973F9C6FB3DABA4" authority="Lower, 1901" authorityName="Lower" authorityYear="1901" box="[797,1183,1656,1681]" class="Insecta" family="Tortricidae" genus="Temnolopha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="mosaica">
|
||
<emphasis id="F3A3EAD13A6C3F5F3973F9C6FBA8ABA4" box="[797,1034,1656,1681]" italics="true" pageId="1" pageNumber="118">Temnolopha mosaica</emphasis>
|
||
Lower,
|
||
<quantity id="062F9B263A6C3F5F3E09F9C6FB3DABA4" box="[1127,1183,1656,1680]" metricMagnitude="1" metricUnit="m" metricValue="4.828539999999999" pageId="1" pageNumber="118" unit="in" value="1901.0">1901</quantity>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="A5464B323A6C3F5F3EACF9C6FAC4ABA5" author="Horak" box="[1218,1382,1656,1681]" pageId="1" pageNumber="118" refString="Horak, M. (2006) Olethreutine moths of Australia (Lepidoptera: Tortricidae). Monographs on Australian Lepidoptera. Vol. 10. CSIRO Publishing, Canberra, 522 pp." type="book" year="2006">Horak (2006))</bibRefCitation>
|
||
. The next genus in Horak’s (2006) treatment is
|
||
<taxonomicName id="06D74D403A6C3F5F38FAF922FC7BAB80" authority="Kocak, 1981" authorityName="Kocak" authorityYear="1981" box="[660,985,1692,1717]" class="Insecta" family="Tortricidae" genus="Diakonoffiana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F38FAF922FC94AB81" box="[660,822,1692,1717]" italics="true" pageId="1" pageNumber="118">Diakonoffiana</emphasis>
|
||
Koçak, 1981
|
||
</taxonomicName>
|
||
. This genus possesses a pair of signa resembling those found in
|
||
<taxonomicName id="06D74D403A6C3F5F3BA7F97FFD9FABEC" box="[457,573,1729,1752]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3BA7F97FFD9FABEC" box="[457,573,1729,1752]" italics="true" pageId="1" pageNumber="118">Neaspasia</emphasis>
|
||
</taxonomicName>
|
||
. Both
|
||
<taxonomicName id="06D74D403A6C3F5F38E3F97EFCB5ABED" box="[653,791,1728,1753]" class="Insecta" family="Tortricidae" genus="Temnolopha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F38E3F97EFCB5ABED" box="[653,791,1728,1753]" italics="true" pageId="1" pageNumber="118">Temnolopha</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="06D74D403A6C3F5F393EF97EFC50ABED" box="[848,1010,1728,1753]" class="Insecta" family="Tortricidae" genus="Diakonoffiana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F393EF97EFC50ABED" box="[848,1010,1728,1753]" italics="true" pageId="1" pageNumber="118">Diakonoffiana</emphasis>
|
||
</taxonomicName>
|
||
are placed in the
|
||
<taxonomicName id="06D74D403A6C3F5F3EACF97FFAE9ABEC" box="[1218,1355,1729,1752]" class="Insecta" family="Tortricidae" genus="Neopotamia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3EACF97FFAE9ABEC" box="[1218,1355,1729,1752]" italics="true" pageId="1" pageNumber="118">Neopotamia</emphasis>
|
||
</taxonomicName>
|
||
group. From these facts it is concluded that
|
||
<taxonomicName id="06D74D403A6C3F5F385CF95BFD05ABC8" box="[562,679,1765,1788]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F385CF95BFD05ABC8" box="[562,679,1765,1788]" italics="true" pageId="1" pageNumber="118">Neaspasia</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="06D74D403A6C3F5F393CF95BFC79ABC8" box="[850,987,1765,1788]" class="Insecta" family="Tortricidae" genus="Neopotamia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F393CF95BFC79ABC8" box="[850,987,1765,1788]" italics="true" pageId="1" pageNumber="118">Neopotamia</emphasis>
|
||
</taxonomicName>
|
||
group and is related to
|
||
<taxonomicName id="06D74D403A6C3F5F3E8DF95AFACEABC9" box="[1251,1388,1764,1789]" class="Insecta" family="Tortricidae" genus="Temnolopha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3E8DF95AFACEABC9" box="[1251,1388,1764,1789]" italics="true" pageId="1" pageNumber="118">Temnolopha</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="06D74D403A6C3F5F3AF9F8B6FE9BAA15" box="[151,313,1800,1825]" class="Insecta" family="Tortricidae" genus="Diakonoffiana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3AF9F8B6FE9BAA15" box="[151,313,1800,1825]" italics="true" pageId="1" pageNumber="118">Diakonoffiana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C16836C33A6C3F5C3AA9F893FDCDADE0" blockId="1.[151,1436,861,2005]" lastBlockId="2.[151,1436,151,464]" lastPageId="2" lastPageNumber="119" pageId="1" pageNumber="118">
|
||
In the description of
|
||
<taxonomicName id="06D74D403A6C3F5F3BDEF893FD87AA70" box="[432,549,1837,1860]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6C3F5F3BDEF893FD87AA70" box="[432,549,1837,1860]" italics="true" pageId="1" pageNumber="118">Neaspasia</emphasis>
|
||
</taxonomicName>
|
||
and its
|
||
<typeStatus id="1E6C88613A6C3F5F3813F893FD0FAA71" box="[637,685,1837,1861]" pageId="1" pageNumber="118">type</typeStatus>
|
||
species,
|
||
<bibRefCitation id="A5464B323A6C3F5F397DF892FC75AA70" author="Diakonoff" box="[787,983,1836,1861]" pageId="1" pageNumber="118" refString="Diakonoff, A. (1989) Tortricidae from Madagascar. Part 2. Olethreutinae, 6 (Lepidoptera). Annales de la Societe Entomologique de France (new series), 25 (4), 431 - 460." type="journal article" year="1989">Diakonoff (1989)</bibRefCitation>
|
||
did not mention the anal roll in the hind wing and the modification of the hind leg tibia. He stated the venation to be “characteristically eucosmine”. It is possible that the characters of the hind wing and hind tibia were overlooked, or that they have been secondarily lost in
|
||
<taxonomicName id="06D74D403A6C3F5F3ADAF827FEC2AA85" box="[180,352,1944,1969]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="loxochlamys">
|
||
<emphasis id="F3A3EAD13A6C3F5F3ADAF827FEC2AA85" box="[180,352,1944,1969]" italics="true" pageId="1" pageNumber="118">N. loxochlamys</emphasis>
|
||
</taxonomicName>
|
||
. This is difficult to judge from the photo of the specimen. The genitalia of
|
||
<taxonomicName id="06D74D403A6C3F5F3EC8F827FAF1AA85" box="[1190,1363,1944,1969]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="loxochlamys">
|
||
<emphasis id="F3A3EAD13A6C3F5F3EC8F827FAF1AA85" box="[1190,1363,1944,1969]" italics="true" pageId="1" pageNumber="118">N. loxochlamys</emphasis>
|
||
</taxonomicName>
|
||
are so similar to those of
|
||
<taxonomicName id="06D74D403A6C3F5F3B04F803FE49AAE1" box="[362,491,1980,2005]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="orthacta">
|
||
<emphasis id="F3A3EAD13A6C3F5F3B04F803FE49AAE1" box="[362,491,1980,2005]" italics="true" pageId="1" pageNumber="118">N. orthacta</emphasis>
|
||
</taxonomicName>
|
||
that it is possible they are synonyms. This similarity is strong evidence that the two species are congeneric, and that Diakonoff’s placement of the genus in
|
||
<taxonomicName id="06D74D403A6F3F5C3983FF29FBC7AD84" box="[1005,1125,151,176]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="tribe" tribe="Eucosmini">Eucosmini</taxonomicName>
|
||
is incorrect. We have not examined the
|
||
<typeStatus id="1E6C88613A6F3F5C3B5AFF02FE34ADE1" box="[308,406,188,213]" pageId="2" pageNumber="119" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="06D74D403A6F3F5C3BD5FF03FDCAADE1" box="[443,616,188,213]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="loxochlamys">
|
||
<emphasis id="F3A3EAD13A6F3F5C3BD5FF03FDCAADE1" box="[443,616,188,213]" italics="true" pageId="2" pageNumber="119">N. loxochlamys</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C16836C33A6F3F5C3AA9FF61FAFFACFB" blockId="2.[151,1436,151,464]" pageId="2" pageNumber="119">
|
||
<bibRefCitation id="A5464B323A6F3F5C3AA9FF61FE2FADCC" author="Razowski" box="[199,397,223,248]" pageId="2" pageNumber="119" refString="Razowski, J. (1995) Some Tortricidae from the Arabian Peninsula, with description of one new genus and four new species (Insecta: Lepidoptera). SHILAP Revista de lepidopterologia, 23, 129 - 139." type="journal article" year="1995">Razowski (1995)</bibRefCitation>
|
||
proposed the genus
|
||
<emphasis id="F3A3EAD13A6F3F5C38E2FF61FCBCADCC" box="[652,798,223,248]" italics="true" pageId="2" pageNumber="119">Genetancylis</emphasis>
|
||
based on a male from
|
||
<collectingCountry id="B9C076533A6F3F5C3E2CFF5EFB24ADCC" box="[1090,1158,224,248]" name="Oman" pageId="2" pageNumber="119">Oman</collectingCountry>
|
||
which he described as
|
||
<emphasis id="F3A3EAD13A6F3F5C3AF9FEBAFE8BAC29" box="[151,297,260,285]" italics="true" pageId="2" pageNumber="119">Genetancylis</emphasis>
|
||
<taxonomicName id="06D74D403A6F3F5C3B5EFEBAFDFAAC29" authority="Razowski, 1995" authorityName="Razowski" authorityYear="1995" box="[304,600,260,285]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="homalota">
|
||
<emphasis id="F3A3EAD13A6F3F5C3B5EFEBAFE38AC29" box="[304,410,260,285]" italics="true" pageId="2" pageNumber="119">homalota</emphasis>
|
||
<bibRefCitation id="A5464B323A6F3F5C3BCEFEBAFDFAAC29" author="Razowski" box="[416,600,260,285]" pageId="2" pageNumber="119" refString="Razowski, J. (1995) Some Tortricidae from the Arabian Peninsula, with description of one new genus and four new species (Insecta: Lepidoptera). SHILAP Revista de lepidopterologia, 23, 129 - 139." type="journal article" year="1995">Razowski, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
. Based on Razowski’s figure of the male genitalia, we transfer this species to
|
||
<taxonomicName id="06D74D403A6F3F5C3ADBFE97FE8BAC74" box="[181,297,297,320]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6F3F5C3ADBFE97FE8BAC74" box="[181,297,297,320]" italics="true" pageId="2" pageNumber="119">Neaspasia</emphasis>
|
||
</taxonomicName>
|
||
. This leads to the new generic synonymy.
|
||
<taxonomicName id="06D74D403A6F3F5C397DFE97FB6FAC74" authority="Razowski, 1995" authorityName="Razowski" authorityYear="1995" box="[787,1229,295,320]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="homalota" status="comb. nov.">
|
||
<emphasis id="F3A3EAD13A6F3F5C397DFE97FC58AC74" box="[787,1018,295,320]" italics="true" pageId="2" pageNumber="119">Neaspasia homalota</emphasis>
|
||
(
|
||
<bibRefCitation id="A5464B323A6F3F5C3E64FE99FB67AC74" author="Razowski" box="[1034,1221,295,320]" pageId="2" pageNumber="119" refString="Razowski, J. (1995) Some Tortricidae from the Arabian Peninsula, with description of one new genus and four new species (Insecta: Lepidoptera). SHILAP Revista de lepidopterologia, 23, 129 - 139." type="journal article" year="1995">Razowski, 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="E89057AA3A6F3F5C3EB4FE99FA9CAC0B" box="[1242,1342,295,320]" pageId="2" pageNumber="119">comb. n.</taxonomicNameLabel>
|
||
, closely resembles the African congeners in the male genitalia, but appears to be specifically distinct.
|
||
<bibRefCitation id="A5464B323A6F3F5C3EC6FEF2FACAAC51" author="Razowski" box="[1192,1384,332,357]" pageId="2" pageNumber="119" refString="Razowski, J. (1995) Some Tortricidae from the Arabian Peninsula, with description of one new genus and four new species (Insecta: Lepidoptera). SHILAP Revista de lepidopterologia, 23, 129 - 139." type="journal article" year="1995">Razowski (1995)</bibRefCitation>
|
||
also described
|
||
<taxonomicName id="06D74D403A6F3F5C3B7EFED1FD1AACBC" authority="Razowski, 1995" authorityName="Razowski" authorityYear="1995" box="[272,696,367,392]" class="Insecta" family="Tortricidae" genus="Rhopobota" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="cornuta">
|
||
<emphasis id="F3A3EAD13A6F3F5C3B7EFED1FE4DACBC" box="[272,495,367,392]" italics="true" pageId="2" pageNumber="119">Rhopobota cornuta</emphasis>
|
||
<bibRefCitation id="A5464B323A6F3F5C3B93FED1FD1AACBC" author="Razowski" box="[509,696,367,392]" pageId="2" pageNumber="119" refString="Razowski, J. (1995) Some Tortricidae from the Arabian Peninsula, with description of one new genus and four new species (Insecta: Lepidoptera). SHILAP Revista de lepidopterologia, 23, 129 - 139." type="journal article" year="1995">Razowski, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="B9C076533A6F3F5C3967FECEFCF3ACBC" box="[777,849,368,392]" name="Oman" pageId="2" pageNumber="119">Oman</collectingCountry>
|
||
. This species was described from a female. The genitalia are typical for
|
||
<taxonomicName id="06D74D403A6F3F5C3BD3FE2BFD90AC98" box="[445,562,405,428]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6F3F5C3BD3FE2BFD90AC98" box="[445,562,405,428]" italics="true" pageId="2" pageNumber="119">Neaspasia</emphasis>
|
||
</taxonomicName>
|
||
species, e.g., showing the very characteristic signa. This species is also transferred to
|
||
<taxonomicName id="06D74D403A6F3F5C3B5AFE07FE0AACE4" box="[308,424,441,464]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3A3EAD13A6F3F5C3B5AFE07FE0AACE4" box="[308,424,441,464]" italics="true" pageId="2" pageNumber="119">Neaspasia</emphasis>
|
||
</taxonomicName>
|
||
, resulting in the new combination
|
||
<taxonomicName id="06D74D403A6F3F5C3940FE07FB73ACE4" authority="Razowski, 1995" authorityName="Razowski" authorityYear="1995" box="[814,1233,439,464]" class="Insecta" family="Tortricidae" genus="Neaspasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="cornuta" status="comb. nov.">
|
||
<emphasis id="F3A3EAD13A6F3F5C3940FE07FBA3ACE4" box="[814,1025,441,464]" italics="true" pageId="2" pageNumber="119">Neaspasia cornuta</emphasis>
|
||
(
|
||
<bibRefCitation id="A5464B323A6F3F5C3E61FE09FB6AACE4" author="Razowski" box="[1039,1224,439,464]" pageId="2" pageNumber="119" refString="Razowski, J. (1995) Some Tortricidae from the Arabian Peninsula, with description of one new genus and four new species (Insecta: Lepidoptera). SHILAP Revista de lepidopterologia, 23, 129 - 139." type="journal article" year="1995">Razowski, 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="E89057AA3A6F3F5C3EB3FE09FAFFACFB" box="[1245,1373,439,464]" pageId="2" pageNumber="119">
|
||
<emphasis id="F3A3EAD13A6F3F5C3EB3FE09FAF4ACE4" bold="true" box="[1245,1366,439,464]" pageId="2" pageNumber="119">comb. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |